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<document ID-DOI="10.11646/zootaxa.5256.5.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7758872" ID-ZooBank="E5B9708E-D626-4923-825A-A9FD59AE4024" approvalRequired="3" approvalRequired_for_textStreams="2" approvalRequired_for_treatments="1" checkinTime="1679468565563" checkinUser="plazi" docAuthor="Matalin, Andrey V. &amp; Wiesner, Jürgen" docDate="2023" docId="050B87D78D462469E1B81A10FB2875A4" docLanguage="en" docName="zootaxa.5256.5.1.pdf" docOrigin="Zootaxa 5256 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5256.5.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Therates pseudovitalisi Matalin et Wiesner 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="408" masterDocId="F932FFAF8D40246EE12F1976FFC07601" masterDocTitle="Revision of the Therates fruhstorferi complex (Coleoptera, Cicindelidae)" masterLastPageNumber="433" masterPageNumber="401" pageNumber="407" updateTime="1679469078842" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the Therates fruhstorferi complex (Coleoptera, Cicindelidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Matalin, Andrey V.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-7790-8709</mods:nameIdentifier>
<mods:affiliation>Moscow State Pedagogical University, Department of Zoology &amp; Ecology, Kibalchicha str. 6, build. 3, Moscow 129164, Russia. andrei-matalin @ yandex. ru; https: // orcid. org / 0000 - 0002 - 7790 - 8709 &amp; Pirogov National Research Medical University, Department of Biology, Ostrovitianova str. 1, Moscow 117997, Russia.</mods:affiliation>
<mods:nameIdentifier type="email">andrei-matalin@yandex.ru</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wiesner, Jürgen</mods:namePart>
<mods:affiliation>Dresdener Ring 11, D- 38444 Wolfsburg, Germany.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5256</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>5</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>401</mods:start>
<mods:end>433</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5256.5.1</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5256.5.1</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7758872</mods:identifier>
<mods:identifier type="ZooBank">E5B9708E-D626-4923-825A-A9FD59AE4024</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7758886" ID-Zenodo-Dep="7758886" LSID="urn:lsid:plazi:treatment:050B87D78D462469E1B81A10FB2875A4" httpUri="http://treatment.plazi.org/id/050B87D78D462469E1B81A10FB2875A4" lastPageId="7" lastPageNumber="408" pageId="6" pageNumber="407">
<subSubSection box="[151,755,870,897]" pageId="6" pageNumber="407" type="nomenclature">
<paragraph blockId="6.[151,755,870,933]" box="[151,755,870,897]" pageId="6" pageNumber="407">
<heading bold="true" box="[151,755,870,897]" fontSize="11" level="1" pageId="6" pageNumber="407" reason="1">
<emphasis bold="true" box="[151,755,870,897]" pageId="6" pageNumber="407">
<taxonomicName authority="Matalin et Wiesner" authorityName="Matalin et Wiesner" authorityYear="2023" box="[151,676,870,897]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="pseudovitalisi" status="sp. n.">
<emphasis bold="true" box="[151,429,870,897]" italics="true" pageId="6" pageNumber="407">Therates pseudovitalisi</emphasis>
Matalin et Wiesner
</taxonomicName>
,
<taxonomicNameLabel box="[688,755,871,897]" pageId="6" pageNumber="407" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,634,906,933]" pageId="6" pageNumber="407" type="description">
<paragraph blockId="6.[151,755,870,933]" box="[151,634,906,933]" pageId="6" pageNumber="407">
<figureCitation box="[151,264,907,933]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="6" pageNumber="407">Figs 9, 10</figureCitation>
,
<figureCitation box="[276,348,907,933]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="6" pageNumber="407">36, 37</figureCitation>
,
<figureCitation box="[360,432,906,933]" captionStart="FIGURES 58–64" captionStartId="16.[152,270,1909,1933]" captionTargetBox="[356,1241,203,1829]" captionTargetId="figure-28@16.[314,1272,181,1885]" captionTargetPageId="16" captionText="FIGURES 58–64. Therates spp., labrum: 58, 59—T. pseudovitalisi pseudovitalisi ssp. n. (58—Laos, Ban Song Cha; 59— Laos, Ban Bun Neua); 60, 61—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 62, 63—T. motoensis (China, Hanmi); 64—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 58, 60, 62, 64—males; 59, 61, 63—females; 58, 60, 64—holotypes; 59, 61—paratypes." figureDoi="http://doi.org/10.5281/zenodo.7758884" httpUri="https://zenodo.org/record/7758884/files/figure.png" pageId="6" pageNumber="407">58, 59</figureCitation>
,
<figureCitation box="[444,517,907,933]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="6" pageNumber="407">90–93</figureCitation>
,
<figureCitation box="[528,630,907,933]" captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="6" pageNumber="407">132–134</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[151,1410,975,999]" pageId="6" pageNumber="407" type="reference_group">
<paragraph blockId="6.[151,1410,975,999]" box="[151,1410,975,999]" pageId="6" pageNumber="407">
<treatmentCitationGroup box="[151,1410,975,999]" pageId="6" pageNumber="407">
<taxonomicName ID-CoL="8MKSW" authority="W. Horn, 1913" authorityName="W. Horn" authorityYear="1913" box="[151,592,975,999]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="vitalisi">
<emphasis box="[151,438,975,999]" italics="true" pageId="6" pageNumber="407">Therates fruhstorferi vitalisi</emphasis>
W.
<bibRefCitation author="Horn, W." box="[478,592,975,999]" pageId="6" pageNumber="407" pagination="362 - 366" refId="ref25772" refString="Horn, W. (1913) Materiaux pour servir a l'etude de la faune entomologique de l'Indochine. Cicindelinae. Annales de la Societe Entomologique de Belgique, 57, 362 - 366. https: // doi. org / 10.5962 / bhl. part. 4597" type="journal article" year="1913">Horn, 1913</bibRefCitation>
</taxonomicName>
—
<treatmentCitation author="Sawada, H. &amp; Wiesner, J." box="[618,949,975,999]" page="63" pageId="6" pageNumber="407" year="2000">
<bibRefCitation author="Sawada, H. &amp; Wiesner, J." box="[618,949,975,999]" pageId="6" pageNumber="407" pagination="59 - 68" refId="ref26747" refString="Sawada, H. &amp; Wiesner, J. (2000) New records of tiger beetles collected in Laos (Coleoptera: Cicindelidae). Entomological Revue of Japan, 55 (1), 59 - 68." type="journal article" year="2000">Sawada &amp; Wiesner 2000: 63–64</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation author="Sawada, H. &amp; Wiesner, J." box="[960,1116,975,999]" page="263" pageId="6" pageNumber="407" year="2004">
<bibRefCitation author="Sawada, H. &amp; Wiesner, J." box="[960,1116,975,999]" pageId="6" pageNumber="407" pagination="255 - 274" refId="ref26828" refString="Sawada, H. &amp; Wiesner, J. (2004) Enumeration of the tiger beetles from Laos with description of the three new species (Coleoptera: Cicindelidae). Entomological Revue of Japan, 59 (2), 255 - 274." type="journal article" year="2004">2004: 263–264</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Wiesner, J. &amp; Geiser, M." box="[1125,1406,975,999]" page="88" pageId="6" pageNumber="407" year="2016">
<bibRefCitation author="Wiesner, J. &amp; Geiser, M." box="[1125,1406,975,999]" pageId="6" pageNumber="407" pagination="61 - 117" refId="ref27796" refString="Wiesner, J. &amp; Geiser, M. (2016) Faunistic survey of the Tiger Beetles (Coleoptera, Carabidae, Cicindelinae) of Laos. Entomologica Basiliensia et Collectionis Frey, 35, 61 - 117." type="journal article" year="2016">Wiesner &amp; Geiser 2016: 88</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="407" type="materials_examined">
<paragraph blockId="6.[151,1437,1045,2008]" pageId="6" pageNumber="407">
<emphasis bold="true" box="[151,328,1045,1071]" pageId="6" pageNumber="407">Type material.</emphasis>
<materialsCitation collectingDate="1999-05-16" collectionCode="JW" collectorName="Vit Kuban &amp; C. Holzschuh" country="Laos" elevation="1200" location="Ban Song Cha" longitude="102.23333" pageId="6" pageNumber="407" specimenCount="1" stateProvince="Louangphabang" typeStatus="holotype">
<typeStatus box="[337,455,1045,1071]" pageId="6" pageNumber="407">HOLOTYPE</typeStatus>
, 1Ô—
<collectingCountry box="[533,592,1047,1074]" name="Laos" pageId="6" pageNumber="407">Laos</collectingCountry>
,
<collectingRegion box="[605,849,1045,1072]" country="Laos" name="Louangphabang" pageId="6" pageNumber="407">Luang Prabang Prov.</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:050B87D78D462469E1B81A10FB2875A4:887D601A8D462468E2711D60FBC5722E" box="[862,1029,1045,1072]" country="Laos" longitude="102.23333" name="Ban Song Cha" pageId="6" pageNumber="407" stateProvince="Louangphabang">Ban Song Cha</location>
(
<quantity box="[1048,1108,1045,1072]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="407" unit="km" value="5.0">5 km</quantity>
W), 20
<superScript attach="none" box="[1197,1205,1045,1059]" fontSize="6" pageId="6" pageNumber="407">o</superScript>
33–4’N
<geoCoordinate box="[1300,1407,1045,1071]" degrees="102" direction="east" minutes="14" orientation="longitude" pageId="6" pageNumber="407" precision="925" value="102.23333">
102
<superScript attach="none" box="[1342,1350,1045,1059]" fontSize="6" pageId="6" pageNumber="407">o</superScript>
14’E
</geoCoordinate>
, ±
<quantity box="[151,242,1081,1108]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="407" unit="m" value="1200.0">
<elevation box="[151,242,1081,1108]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="407" unit="m" value="1200.0">1200 m</elevation>
</quantity>
, 24.4–
<date box="[321,434,1081,1108]" pageId="6" pageNumber="407" value="1999-05-16">
<collectingDate box="[321,434,1081,1108]" pageId="6" pageNumber="407" value="1999-05-16">16.5.1999</collectingDate>
</date>
, leg.
<collectorName box="[500,620,1081,1107]" pageId="6" pageNumber="407">Vit Kubaň</collectorName>
&amp;
<collectorName box="[662,818,1081,1107]" pageId="6" pageNumber="407">C. Holzschuh</collectorName>
(
<collectionCode box="[837,880,1081,1107]" pageId="6" pageNumber="407">JW</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectionCode="JW" country="Laos" pageId="6" pageNumber="407" specimenCount="1" specimenCount-female="1" stateProvince="Louangphabang" typeStatus="paratype">
<typeStatus box="[903,1025,1081,1107]" pageId="6" pageNumber="407">PARATYPES</typeStatus>
:
<specimenCount box="[1038,1061,1081,1107]" pageId="6" pageNumber="407" type="female">1♀</specimenCount>
—same labeled as the
<typeStatus box="[1338,1436,1081,1107]" pageId="6" pageNumber="407">holotype</typeStatus>
(
<collectionCode box="[159,202,1117,1143]" pageId="6" pageNumber="407">JW</collectionCode>
)
</materialsCitation>
; 3ÔÔ
<materialsCitation box="[282,733,1117,1146]" collectionCode="NEE" country="Laos" location="Oudom Xai" pageId="6" pageNumber="407" specimenCount="6" specimenCount-female="6" typeStatus="paratype">
<specimenCount box="[282,326,1119,1146]" pageId="6" pageNumber="407" type="female">6♀♀</specimenCount>
—
<collectingCountry box="[359,420,1119,1146]" name="Laos" pageId="6" pageNumber="407">Laos</collectingCountry>
,
<location LSID="urn:lsid:plazi:treatment:050B87D78D462469E1B81A10FB2875A4:887D601A8D462468E09D1D2BFDF67276" box="[434,566,1117,1143]" country="Laos" name="Oudom Xai" pageId="6" pageNumber="407">Oudom Xai</location>
(
<quantity box="[584,657,1117,1143]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="407" unit="km" value="17.0">17 km</quantity>
<collectionCode box="[666,725,1117,1143]" pageId="6" pageNumber="407">NEE</collectionCode>
)
</materialsCitation>
,
<materialsCitation collectingDate="1998-05-01" collectionCode="JW, MSPU" collectorName="Vit Kuban" country="Laos" elevation="1100" latitude="20.75" longLatPrecision="1266" longitude="102.15" pageId="6" pageNumber="407" specimenCount="7" specimenCount-female="7" typeStatus="paratype">
<geoCoordinate box="[745,839,1117,1144]" degrees="20" direction="north" minutes="45" orientation="latitude" pageId="6" pageNumber="407" precision="925" value="20.75">
20
<superScript attach="none" box="[773,781,1117,1131]" fontSize="6" pageId="6" pageNumber="407">o</superScript>
45’N
</geoCoordinate>
<geoCoordinate box="[848,955,1117,1143]" degrees="102" direction="east" minutes="09" orientation="longitude" pageId="6" pageNumber="407" precision="925" value="102.15">
102
<superScript attach="none" box="[890,898,1117,1131]" fontSize="6" pageId="6" pageNumber="407">o</superScript>
09’E
</geoCoordinate>
, ~
<quantity box="[992,1079,1117,1144]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="407" unit="m" value="1100.0">
<elevation box="[992,1079,1117,1144]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="407" unit="m" value="1100.0">1100 m</elevation>
</quantity>
, 18.4–
<date box="[1156,1256,1117,1144]" pageId="6" pageNumber="407" value="1998-05-01">
<collectingDate box="[1156,1256,1117,1144]" pageId="6" pageNumber="407" value="1998-05-01">1.5.1998</collectingDate>
</date>
, leg.
<collectorName box="[1319,1437,1117,1143]" pageId="6" pageNumber="407">Vit Kubaň</collectorName>
(2ÔÔ
<specimenCount box="[219,262,1155,1182]" pageId="6" pageNumber="407" type="female">5♀♀</specimenCount>
—
<collectionCode box="[295,338,1155,1182]" pageId="6" pageNumber="407">JW</collectionCode>
, 1Ô
<specimenCount box="[383,428,1155,1182]" pageId="6" pageNumber="407" type="female">2♀♀</specimenCount>
—
<collectionCode box="[461,550,1155,1182]" pageId="6" pageNumber="407">MSPU</collectionCode>
)
</materialsCitation>
;
<materialsCitation box="[573,1305,1153,1182]" collectingDate="2011-07" collectionCode="MSPU" country="Laos" location="Phou Pane Mt" pageId="6" pageNumber="407" specimenCount="1" stateProvince="Houaphan" typeStatus="paratype">
1Ô—
<collectingCountry box="[634,693,1155,1182]" name="Laos" pageId="6" pageNumber="407">Laos</collectingCountry>
,
<collectingRegion box="[706,906,1153,1180]" country="Laos" name="Houaphan" pageId="6" pageNumber="407">Houaphanh Prov.</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:050B87D78D462469E1B81A10FB2875A4:887D601A8D462468E2B81DF7FBFC729D" box="[919,1084,1153,1180]" country="Laos" name="Phou Pane Mt" pageId="6" pageNumber="407" stateProvince="Houaphan">Phou Pane Mt</location>
,
<date box="[1097,1198,1153,1179]" pageId="6" pageNumber="407" value="2011-07">
<collectingDate box="[1097,1198,1153,1179]" pageId="6" pageNumber="407" value="2011-07">VII.2011</collectingDate>
</date>
(
<collectionCode box="[1215,1298,1153,1179]" pageId="6" pageNumber="407">MSPU</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2002-05-17" collectionCode="V, JW" country="Laos" location="Neua" municipality="Sam" pageId="6" pageNumber="407" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount box="[1319,1342,1153,1179]" pageId="6" pageNumber="407" type="female">1♀</specimenCount>
—
<collectingCountry box="[1375,1436,1153,1179]" name="Laos" pageId="6" pageNumber="407">Laos</collectingCountry>
NE,
<collectingMunicipality box="[203,253,1189,1215]" pageId="6" pageNumber="407">Sam</collectingMunicipality>
[
<emphasis box="[269,346,1190,1215]" italics="true" pageId="6" pageNumber="407">= Xam</emphasis>
]
<location LSID="urn:lsid:plazi:treatment:050B87D78D462469E1B81A10FB2875A4:887D601A8D462468E0451DD0FE6872C1" box="[362,424,1190,1216]" country="Laos" municipality="Sam" name="Neua" pageId="6" pageNumber="407">Neua</location>
,
<date box="[436,550,1189,1215]" pageId="6" pageNumber="407" value="2002-05-17">
<collectingDate box="[436,550,1189,1215]" pageId="6" pageNumber="407" value="2002-05-17">
17.
<collectionCode box="[465,486,1189,1215]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="407" type="Museum">V</collectionCode>
.2002
</collectingDate>
</date>
(
<collectionCode box="[566,609,1189,1215]" pageId="6" pageNumber="407">JW</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2003-06-20" collectionCode="VI, JW" collectorName="P. Pacholatko" country="Laos" elevation="1100" latitude="21.633333" location="Ban Bun Neua" longLatPrecision="1263" longitude="101.95" pageId="6" pageNumber="407" specimenCount="1" stateProvince="Phongsali" typeStatus="paratype">
1Ô—Laos-N,
<collectingRegion box="[789,973,1189,1216]" country="Laos" name="Phongsali" pageId="6" pageNumber="407">Phongsaly prov.</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:050B87D78D462469E1B81A10FB2875A4:887D601A8D462468E2F71DD0FBBD72C1" box="[984,1149,1190,1216]" country="Laos" latitude="21.633333" longLatPrecision="1263" longitude="101.95" name="Ban Bun Neua" pageId="6" pageNumber="407" stateProvince="Phongsali">Ban Bun Neua</location>
[=
<emphasis box="[1188,1313,1190,1215]" italics="true" pageId="6" pageNumber="407">Boun Neua</emphasis>
] (
<quantity box="[1338,1396,1189,1215]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="407" unit="km" value="4.0">4 km</quantity>
E),
<date box="[151,282,1225,1251]" pageId="6" pageNumber="407" value="2003-06-20">
<collectingDate box="[151,282,1225,1251]" pageId="6" pageNumber="407" value="2003-06-20">
20.
<collectionCode box="[185,217,1225,1251]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="6" pageNumber="407">VI</collectionCode>
.2003
</collectingDate>
</date>
, ~
<quantity box="[307,394,1225,1252]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="407" unit="m" value="1100.0">
<elevation box="[307,394,1225,1252]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="407" unit="m" value="1100.0">1100 m</elevation>
</quantity>
,
<geoCoordinate box="[406,499,1225,1251]" degrees="21" direction="north" minutes="38" orientation="latitude" pageId="6" pageNumber="407" precision="925" value="21.633333">
21
<superScript attach="none" box="[434,442,1225,1239]" fontSize="6" pageId="6" pageNumber="407">o</superScript>
38’N
</geoCoordinate>
<geoCoordinate box="[506,613,1225,1252]" degrees="101" direction="east" minutes="57" orientation="longitude" pageId="6" pageNumber="407" precision="925" value="101.95">
101
<superScript attach="none" box="[548,556,1225,1239]" fontSize="6" pageId="6" pageNumber="407">o</superScript>
57’E
</geoCoordinate>
,
<collectorName box="[625,775,1225,1252]" pageId="6" pageNumber="407">P. Pacholátko</collectorName>
leg. (
<collectionCode box="[839,882,1225,1251]" pageId="6" pageNumber="407">JW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="407" type="reference_group">
<paragraph blockId="6.[151,1437,1045,2008]" pageId="6" pageNumber="407">
<emphasis bold="true" box="[199,336,1261,1287]" pageId="6" pageNumber="407">References.</emphasis>
<materialsCitation collectingDate="2002-07-15" collectingDateMax="2003-06-21" collectingDateMin="2002-07-15" collectorName="Xam Nua Distr &amp; Phu Pan" country="Laos" elevation="1350" location="Ban Saleui" municipality="Xam Nua Distr" pageId="6" pageNumber="407" specimenCount="2" specimenCount-female="2" stateProvince="Houaphan">
<collectingCountry box="[343,401,1261,1287]" name="Laos" pageId="6" pageNumber="407">LAOS</collectingCountry>
,
<emphasis box="[415,614,1261,1287]" italics="true" pageId="6" pageNumber="407">
<collectingRegion box="[415,614,1261,1287]" country="Laos" name="Houaphan" pageId="6" pageNumber="407">Houaphanh Prov.</collectingRegion>
</emphasis>
: 1Ô
<specimenCount box="[659,705,1263,1290]" pageId="6" pageNumber="407" type="female">2♀♀</specimenCount>
—
<collectingMunicipality box="[738,920,1261,1290]" pageId="6" pageNumber="407">Xam Nua Distr</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:050B87D78D462469E1B81A10FB2875A4:887D601A8D462468E28C1D98FBE17306" box="[931,1057,1261,1288]" country="Laos" municipality="Xam Nua Distr" name="Ban Saleui" pageId="6" pageNumber="407" stateProvince="Houaphan">Ban Saleui</location>
,
<quantity box="[1068,1155,1261,1288]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="6" pageNumber="407" unit="m" value="1350.0">
<elevation box="[1068,1155,1261,1288]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="6" pageNumber="407" unit="m" value="1350.0">1350 m</elevation>
</quantity>
,
<date box="[1167,1307,1261,1288]" pageId="6" pageNumber="407" value="2002-07-15">
<collectingDate box="[1167,1307,1261,1288]" pageId="6" pageNumber="407" value="2002-07-15">15.VII.2002</collectingDate>
</date>
; 1Ô—
<collectorName pageId="6" pageNumber="407">Xam Nua Distr</collectorName>
,
<collectorName box="[275,371,1297,1324]" pageId="6" pageNumber="407">Phu Pan</collectorName>
,
<quantity box="[382,470,1297,1324]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="407" unit="m" value="1800.0">
<elevation box="[382,470,1297,1324]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="407" unit="m" value="1800.0">1800 m</elevation>
</quantity>
,
<date box="[482,652,1297,1323]" pageId="6" pageNumber="407" value="2003-06-17" valueMax="2003-06-21" valueMin="2003-06-17">
<collectingDate box="[482,652,1297,1323]" pageId="6" pageNumber="407" value="2003-06-17" valueMax="2003-06-21" valueMin="2003-06-17">17–21.VI.2003</collectingDate>
</date>
(both
<bibRefCitation author="Sawada, H. &amp; Wiesner, J." box="[726,1010,1297,1323]" pageId="6" pageNumber="407" pagination="255 - 274" refId="ref26828" refString="Sawada, H. &amp; Wiesner, J. (2004) Enumeration of the tiger beetles from Laos with description of the three new species (Coleoptera: Cicindelidae). Entomological Revue of Japan, 59 (2), 255 - 274." type="journal article" year="2004">Sawada &amp; Wiesner 2004</bibRefCitation>
)
</materialsCitation>
;
<materialsCitation collectingDate="2009-06-01" collectingDateMax="2009-06-16" collectingDateMin="2009-06-01" collectorName="Z. Kraus" country="Laos" elevation="1515" location="Phou Pane Mt." longitude="103.99833" pageId="6" pageNumber="407" specimenCount="1" specimenCount-female="1" stateProvince="Houaphanh Prov.">
<specimenCount box="[1030,1053,1297,1323]" pageId="6" pageNumber="407" type="female">1♀</specimenCount>
—
<location LSID="urn:lsid:plazi:treatment:050B87D78D462469E1B81A10FB2875A4:887D601A8D462468E5121C67FB2E732D" box="[1085,1262,1297,1324]" country="Laos" longitude="103.99833" name="Phou Pane Mt." pageId="6" pageNumber="407" stateProvince="Houaphanh Prov.">Phou Pane Mt.</location>
,
<quantity box="[1274,1432,1297,1324]" metricMagnitude="3" metricUnit="m" metricValue="1.515" metricValueMax="1.55" metricValueMin="1.48" pageId="6" pageNumber="407" unit="m" value="1515.0" valueMax="1550.0" valueMin="1480.0">
<elevation box="[1274,1432,1297,1324]" metricMagnitude="3" metricUnit="m" metricValue="1.515" metricValueMax="1.55" metricValueMin="1.48" pageId="6" pageNumber="407" unit="m" value="1515.0" valueMax="1550.0" valueMin="1480.0">1480–1550 m</elevation>
</quantity>
, 20
<superScript attach="none" box="[179,187,1333,1347]" fontSize="6" pageId="6" pageNumber="407">o</superScript>
13’09–19’’N
<geoCoordinate box="[338,491,1333,1360]" degrees="103" direction="east" minutes="59" orientation="longitude" pageId="6" pageNumber="407" precision="15" seconds="54" value="103.99833">103º59’54’’E</geoCoordinate>
,
<date box="[502,661,1333,1359]" pageId="6" pageNumber="407" value="2009-06-01" valueMax="2009-06-16" valueMin="2009-06-01">
<collectingDate box="[502,661,1333,1359]" pageId="6" pageNumber="407" value="2009-06-01" valueMax="2009-06-16" valueMin="2009-06-01">1–16.VI.2009</collectingDate>
</date>
, leg.
<collectorName box="[720,818,1334,1360]" pageId="6" pageNumber="407">Z. Kraus</collectorName>
(
<bibRefCitation author="Wiesner, J. &amp; Geiser, M." box="[832,1103,1333,1359]" pageId="6" pageNumber="407" pagination="61 - 117" refId="ref27796" refString="Wiesner, J. &amp; Geiser, M. (2016) Faunistic survey of the Tiger Beetles (Coleoptera, Carabidae, Cicindelinae) of Laos. Entomologica Basiliensia et Collectionis Frey, 35, 61 - 117." type="journal article" year="2016">Wiesner &amp; Geiser 2016</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="407" type="diagnosis">
<paragraph blockId="6.[151,1437,1045,2008]" pageId="6" pageNumber="407">
<emphasis bold="true" box="[199,322,1369,1395]" pageId="6" pageNumber="407">Diagnosis.</emphasis>
This new species is readily distinguished from
<taxonomicName baseAuthorityName="Wiesner &amp; Bi" baseAuthorityYear="2014" box="[880,987,1369,1395]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis box="[880,987,1369,1395]" italics="true" pageId="6" pageNumber="407">T. vitalisi</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Tan, Mo, Liang" authorityYear="1991" box="[1004,1149,1369,1395]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="biserratus">
<emphasis box="[1004,1149,1369,1395]" italics="true" pageId="6" pageNumber="407">T. biserratus</emphasis>
</taxonomicName>
,
<taxonomicName authority="Matalin &amp; Wiesner, 2023" authorityName="Matalin &amp; Wiesner" authorityYear="2023" box="[1165,1322,1369,1395]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="hunanensis" status="sp. n.">
<emphasis box="[1165,1322,1369,1395]" italics="true" pageId="6" pageNumber="407">T. hunanensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1332,1399,1369,1395]" pageId="6" pageNumber="407">
<taxonomicNameLabel box="[1332,1399,1369,1395]" pageId="6" pageNumber="407" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName authorityName="W. Horn" authorityYear="1912" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis italics="true" pageId="6" pageNumber="407">T. sauteri</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Tan" authorityYear="1981" box="[283,422,1406,1431]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="motoensis">
<emphasis box="[283,422,1406,1431]" italics="true" pageId="6" pageNumber="407">T. motoensis</emphasis>
</taxonomicName>
by lacking lateral carinae on the apex of aedeagus (
<figureCitation box="[1001,1155,1405,1431]" captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="6" pageNumber="407">Figs 132–134</figureCitation>
<emphasis box="[1162,1193,1406,1431]" italics="true" pageId="6" pageNumber="407">vs.</emphasis>
<figureCitation box="[1199,1357,1405,1432]" captionStart="FIGURES 152–174" captionStartId="24.[152,270,1210,1234]" captionTargetBox="[163,1428,181,1165]" captionTargetId="figure-28@24.[151,1435,181,1186]" captionTargetPageId="24" captionText="FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758906" httpUri="https://zenodo.org/record/7758906/files/figure.png" pageId="6" pageNumber="407">Figs 152–174</figureCitation>
). From
<taxonomicName authorityName="W. Horn" authorityYear="1902" box="[151,307,1441,1467]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="fruhstorferi">
<emphasis box="[151,307,1441,1467]" italics="true" pageId="6" pageNumber="407">T. fruhstorferi</emphasis>
</taxonomicName>
and
<taxonomicName authority="Matalin &amp; Wiesner, 2023" authorityName="Matalin &amp; Wiesner" authorityYear="2023" box="[360,547,1441,1468]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="ziyardamensis" status="sp. n.">
T.
<emphasis box="[388,547,1441,1467]" italics="true" pageId="6" pageNumber="407">ziyardamensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[553,616,1441,1467]" pageId="6" pageNumber="407">
<taxonomicNameLabel box="[553,616,1441,1467]" pageId="6" pageNumber="407" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
a new species is easily distinguished by the shape of aedeagus (
<figureCitation captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="6" pageNumber="407">Figs 132– 134</figureCitation>
<emphasis box="[200,230,1478,1503]" italics="true" pageId="6" pageNumber="407">vs.</emphasis>
<figureCitation box="[237,392,1477,1504]" captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="6" pageNumber="407">Figs 115–131</figureCitation>
), from
<taxonomicName authorityName="W. Horn" authorityYear="1902" box="[473,629,1477,1503]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="fruhstorferi">
<emphasis box="[473,629,1477,1503]" italics="true" pageId="6" pageNumber="407">T. fruhstorferi</emphasis>
</taxonomicName>
additionally by wider apices of elytra (
<figureCitation box="[1066,1180,1477,1503]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="6" pageNumber="407">Figs 9–11</figureCitation>
,
<figureCitation box="[1191,1261,1477,1504]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="6" pageNumber="407">90–95</figureCitation>
<emphasis box="[1268,1298,1478,1503]" italics="true" pageId="6" pageNumber="407">vs.</emphasis>
<figureCitation box="[1305,1431,1477,1503]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="6" pageNumber="407">Figs 1, 3–8</figureCitation>
,
<figureCitation box="[151,223,1513,1539]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="6" pageNumber="407">81–89</figureCitation>
) and by shorter hind tarsus (
<figureCitation box="[531,688,1513,1540]" captionStart="FIGURES 175–178" captionStartId="25.[152,270,1739,1763]" captionTargetBox="[151,1435,181,1716]" captionTargetId="figure-17@25.[151,1435,181,1716]" captionTargetPageId="25" captionText="FIGURES 175–178. Morphometric features of Therates spp: 175, 177—relation of the proportions EL/PL and HTbL/HTaL (green rhombus—T. pseudovitalisi sp. n.; blue circles—T. fruhstorferi; open symbols—males; solid symbols—females); 176, 178—relation of the proportions HTbL/HTaL (small quadrates—medians; rectangles—25–75 percentiles; whiskers—min–max; significantly different samples are marked with different lower letters, according to Mann–Whitney U-test, p &lt;0.05 (176—U = 112.0, z = 5.08, p = 0.000; 178—U = 23.5, z = 3.37, p = 0.0007)); 175, 176—all specimens; 177, 178—males." figureDoi="http://doi.org/10.5281/zenodo.7758910" httpUri="https://zenodo.org/record/7758910/files/figure.png" pageId="6" pageNumber="407">Figs 175–178</figureCitation>
). From
<taxonomicName authorityName="Matalin et Wiesner" authorityYear="2023" box="[771,1050,1513,1539]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="subSpecies" species="pseudovitalisi" subSpecies="tibetana">
<emphasis box="[771,1050,1513,1539]" italics="true" pageId="6" pageNumber="407">T. pseudovitalisi tibetana</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1055,1127,1513,1539]" pageId="6" pageNumber="407">ssp. n.</emphasis>
a nominotypical subspecies is clearly differentiated by the shape of antennomeres 9–11 (
<figureCitation box="[826,952,1549,1575]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="6" pageNumber="407">Figs 36, 37</figureCitation>
<emphasis box="[959,989,1550,1575]" italics="true" pageId="6" pageNumber="407">vs.</emphasis>
<figureCitation box="[996,1124,1549,1575]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="6" pageNumber="407">Figs 38, 39</figureCitation>
), by the proportions of hind tibiae and tarsi—mean HTbL/HTaL = 1.32
<emphasis box="[638,668,1586,1611]" italics="true" pageId="6" pageNumber="407">vs.</emphasis>
HTbL/HTaL = 1.25, by narrow, elongate, clearly oblique central dot (
<figureCitation box="[159,274,1621,1647]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="6" pageNumber="407">Figs 9, 10</figureCitation>
,
<figureCitation box="[286,356,1621,1647]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="6" pageNumber="407">90–93</figureCitation>
<emphasis box="[363,393,1622,1647]" italics="true" pageId="6" pageNumber="407">vs.</emphasis>
<figureCitation box="[400,484,1621,1647]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="6" pageNumber="407">Figs 11</figureCitation>
,
<figureCitation box="[496,569,1621,1648]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="6" pageNumber="407">94, 95</figureCitation>
), and by the longer elytra—mean EL/AL = 2.6
<emphasis box="[1101,1132,1622,1647]" italics="true" pageId="6" pageNumber="407">vs.</emphasis>
mean EL/AL = 2.48.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="408" pageId="6" pageNumber="407" type="description">
<paragraph blockId="6.[151,1437,1045,2008]" pageId="6" pageNumber="407">
<emphasis bold="true" box="[199,345,1657,1683]" pageId="6" pageNumber="407">Description.</emphasis>
TL =
<quantity box="[414,590,1657,1684]" metricMagnitude="-2" metricUnit="m" metricValue="1.0815" metricValueMax="1.1099999999999999" metricValueMin="1.053" pageId="6" pageNumber="407" unit="mm" value="10.815" valueMax="11.1" valueMin="10.53">10.53–11.1 mm</quantity>
in males (mean =
<quantity box="[796,912,1657,1684]" metricMagnitude="-2" metricUnit="m" metricValue="1.074" pageId="6" pageNumber="407" unit="mm" value="10.74">10.74 mm</quantity>
, n = 5),
<quantity box="[1004,1167,1657,1684]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" metricValueMax="1.21" metricValueMin="1.05" pageId="6" pageNumber="407" unit="mm" value="11.3" valueMax="12.1" valueMin="10.5">10.5–12.1 mm</quantity>
(mean =
<quantity box="[1273,1388,1657,1684]" metricMagnitude="-2" metricUnit="m" metricValue="1.117" pageId="6" pageNumber="407" unit="mm" value="11.17">11.17 mm</quantity>
, n = 7) in females.
</paragraph>
<paragraph blockId="6.[151,1437,1045,2008]" lastBlockId="7.[151,1437,151,933]" lastPageId="7" lastPageNumber="408" pageId="6" pageNumber="407">
Head shining black-blue with purple-violet reflection; orbital plates finely striated in posterior third; frons convex, semi-circular (see above), slightly downward anteriorly (see at the side), smooth or slightly rugose, frontal sulci deep and slightly convergent in anterior 2/3, shallow and distinctly divergent in posterior third. Mandibles in males brown underside, yellow-brown topside with brown teeth and apical molar; in females dark-brown except pale basal quarter topside. Labial palpi yellow-brown or brown except dark-brown apical palpomeres; maxillary palpi dark brown with narrow light-brown apices in all palpomeres. Antennae barely extend shoulders, black, scape dark brown on anterior margin and black on posterior margin, in some females brown-black, antennomeres 2–5 with light violet tinge and narrow dark-brown apices, antennomeres
<quantity box="[854,937,1981,2007]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" metricValueMax="2.794" metricValueMin="2.286" pageId="6" pageNumber="407" unit="in" value="10.0" valueMax="11.0" valueMin="9.0">9–11 in</quantity>
males distinctly dilated, antennomeres 9 and 10 with wide protruding downward anterior lower margins (
<figureCitation box="[822,905,151,177]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="7" pageNumber="408">Fig. 37</figureCitation>
), in female only slightly delated and protruding (
<figureCitation box="[159,246,187,213]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="7" pageNumber="408">Fig. 36</figureCitation>
). Labrum indistinctly transverse, LW/LL = 1.0–1.08 (mean = 1.04, n = 12), in males black-brown with medium-sized yellow-brown apical spot indistinctly separated from central apical teeth (
<figureCitation box="[1129,1211,222,249]" captionStart="FIGURES 58–64" captionStartId="16.[152,270,1909,1933]" captionTargetBox="[356,1241,203,1829]" captionTargetId="figure-28@16.[314,1272,181,1885]" captionTargetPageId="16" captionText="FIGURES 58–64. Therates spp., labrum: 58, 59—T. pseudovitalisi pseudovitalisi ssp. n. (58—Laos, Ban Song Cha; 59— Laos, Ban Bun Neua); 60, 61—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 62, 63—T. motoensis (China, Hanmi); 64—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 58, 60, 62, 64—males; 59, 61, 63—females; 58, 60, 64—holotypes; 59, 61—paratypes." figureDoi="http://doi.org/10.5281/zenodo.7758884" httpUri="https://zenodo.org/record/7758884/files/figure.png" pageId="7" pageNumber="408">Fig. 58</figureCitation>
), in females entirely black (
<figureCitation box="[227,312,258,285]" captionStart="FIGURES 58–64" captionStartId="16.[152,270,1909,1933]" captionTargetBox="[356,1241,203,1829]" captionTargetId="figure-28@16.[314,1272,181,1885]" captionTargetPageId="16" captionText="FIGURES 58–64. Therates spp., labrum: 58, 59—T. pseudovitalisi pseudovitalisi ssp. n. (58—Laos, Ban Song Cha; 59— Laos, Ban Bun Neua); 60, 61—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 62, 63—T. motoensis (China, Hanmi); 64—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 58, 60, 62, 64—males; 59, 61, 63—females; 58, 60, 64—holotypes; 59, 61—paratypes." figureDoi="http://doi.org/10.5281/zenodo.7758884" httpUri="https://zenodo.org/record/7758884/files/figure.png" pageId="7" pageNumber="408">Fig. 59</figureCitation>
).
</paragraph>
<paragraph blockId="7.[151,1437,151,933]" pageId="7" pageNumber="408">Pronotum shining blue with violet reflection; indistinctly transverse, PW/PL = 1.0–1.11 (mean = 1.06, n = 12), apical lobe practically equal wide throughout; thorax shining black-blue with light golden-violet reflection.</paragraph>
<paragraph blockId="7.[151,1437,151,933]" pageId="7" pageNumber="408">
Femora bicolored, fore femora pale-yellow with brown violet-tinged anterior side, black with violet tinge on posterior side, middle femora brown-black with light violet tinge except pale-yellow posterior basal third, hind femora pale-yellow in basal third or quarter and black with violet tinge in apical 2/3 or 3/4; tibiae brown-black with blue-violet lustre; tarsi black with blue-violet tinge; HTbL/HTaL = 1.25–1.42 (mean = 1.32, n = 12) (
<figureCitation captionStart="FIGURES 175–178" captionStartId="25.[152,270,1739,1763]" captionTargetBox="[151,1435,181,1716]" captionTargetId="figure-17@25.[151,1435,181,1716]" captionTargetPageId="25" captionText="FIGURES 175–178. Morphometric features of Therates spp: 175, 177—relation of the proportions EL/PL and HTbL/HTaL (green rhombus—T. pseudovitalisi sp. n.; blue circles—T. fruhstorferi; open symbols—males; solid symbols—females); 176, 178—relation of the proportions HTbL/HTaL (small quadrates—medians; rectangles—25–75 percentiles; whiskers—min–max; significantly different samples are marked with different lower letters, according to Mann–Whitney U-test, p &lt;0.05 (176—U = 112.0, z = 5.08, p = 0.000; 178—U = 23.5, z = 3.37, p = 0.0007)); 175, 176—all specimens; 177, 178—males." figureDoi="http://doi.org/10.5281/zenodo.7758910" httpUri="https://zenodo.org/record/7758910/files/figure.png" pageId="7" pageNumber="408">Figs 175–178</figureCitation>
).
</paragraph>
<paragraph blockId="7.[151,1437,151,933]" pageId="7" pageNumber="408">
Elytra shining black-blue with purple-violet or blue-violet reflection; indistinctly divergent toward apex, EL/ EW = 1.92–2.06 (mean = 1.98, n = 12); punctuation deep and regular between basal humps and central dot, in apical third shallow and sparse; scutellum black with blue or violet tinge; apical margin shortly cut. Elytral pattern presented by well visible pale-yellow small basal portion of humeral lunule (see in front) and elongate slightly or clearly oblique narrow wide-uniformed or slightly dilated in lateral portion central dot; basal dot very small, in some specimens indistinct or entirely absent (
<figureCitation box="[595,710,727,753]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="7" pageNumber="408">Figs 9, 10</figureCitation>
,
<figureCitation box="[722,797,727,753]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="7" pageNumber="408">90–93</figureCitation>
).
</paragraph>
<paragraph blockId="7.[151,1437,151,933]" pageId="7" pageNumber="408">
Aedeagus gradually sinuate with elongate rounded apex, lacking lateral carinae (
<figureCitation box="[1107,1266,763,789]" captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="7" pageNumber="408">Figs 132–134</figureCitation>
); AL =
<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="7" pageNumber="408" unit="mm" value="2.6" valueMax="2.7" valueMin="2.5">2.5–2.7 mm</quantity>
(mean = 2.62, n = 5), EL/AL = 2.56–2.72 (mean = 2.6, n = 5), in one aberrant male EL/AL = 2.45.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="408" type="etymology">
<paragraph blockId="7.[151,1437,151,933]" pageId="7" pageNumber="408">
<emphasis bold="true" box="[199,334,835,861]" pageId="7" pageNumber="408">Etymology.</emphasis>
The new species is named “
<taxonomicName authorityName="Matalin et Wiesner" authorityYear="2023" box="[676,832,835,861]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="pseudovitalisi">
<emphasis box="[676,832,835,861]" italics="true" pageId="7" pageNumber="408">pseudovitalisi</emphasis>
</taxonomicName>
” due to it very similar outward appearance with
<taxonomicName authority="Horn, 1913" authorityName="W. Horn" authorityYear="1913" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis italics="true" pageId="7" pageNumber="408">T. vitalisi</emphasis>
<bibRefCitation author="Horn, W." box="[234,365,871,897]" pageId="7" pageNumber="408" pagination="362 - 366" refId="ref25772" refString="Horn, W. (1913) Materiaux pour servir a l'etude de la faune entomologique de l'Indochine. Cicindelinae. Annales de la Societe Entomologique de Belgique, 57, 362 - 366. https: // doi. org / 10.5962 / bhl. part. 4597" type="journal article" year="1913">Horn, 1913</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection box="[199,1256,906,933]" pageId="7" pageNumber="408" type="distribution">
<paragraph blockId="7.[151,1437,151,933]" box="[199,1256,906,933]" pageId="7" pageNumber="408">
<emphasis bold="true" box="[199,353,906,932]" pageId="7" pageNumber="408">Distribution.</emphasis>
<collectingCountry box="[359,422,907,933]" name="Laos" pageId="7" pageNumber="408">LAOS</collectingCountry>
:
<collectingRegion box="[432,558,907,933]" country="Laos" name="Oudomxai" pageId="7" pageNumber="408">Oudomxay</collectingRegion>
,
<collectingRegion box="[568,742,907,933]" country="Laos" name="Louangphabang" pageId="7" pageNumber="408">Luang Prabang</collectingRegion>
,
<collectingRegion box="[753,885,907,933]" country="Laos" name="Houaphan" pageId="7" pageNumber="408">Houaphanh</collectingRegion>
,
<collectingRegion box="[896,1013,907,933]" country="Laos" name="Phongsali" pageId="7" pageNumber="408">Phongsaly</collectingRegion>
Provinces (
<figureCitation box="[1145,1244,907,933]" captionStart="FIGURES 184" captionStartId="28.[152,270,1579,1603]" captionTargetBox="[151,1435,597,1556]" captionTargetId="figure-226@28.[151,1435,597,1556]" captionTargetPageId="28" captionText="FIGURES 184. Distribution of Therates spp.: dark blue solid circle—T. vitalisi vitalisi; dark blue half-solid circle—T. vitalisi ida; lilac solid circle—T. hunnanensis sp. n.; orange solid circle—T. biserratus; pink solid circle—T. motoensis; red open circle—T. sauteri; red solid quadrat—T. fruhstorferi fruhstorferi; red half-solid quadrat—T. fruhstorferi australis ssp. n.; blue solid quadrat—T. ziyardamensis sp. n.; green solid quadrat—T. pseudovitalisi sp. n. (nominotypical subspecies); green halfsolid quadrat—T. pseudovitalisi tibetana ssp. n.; L in the symbols—according only to the literature sources. Map source— https://commons.wikimedia.org/wiki/File:East_Asia_topographic_map.png." figureDoi="http://doi.org/10.5281/zenodo.7758916" httpUri="https://zenodo.org/record/7758916/files/figure.png" pageId="7" pageNumber="408">Fig. 184</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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