Revision of the Therates fruhstorferi complex (Coleoptera, Cicindelidae) Author Matalin, Andrey V. 0000-0002-7790-8709 Moscow State Pedagogical University, Department of Zoology & Ecology, Kibalchicha str. 6, build. 3, Moscow 129164, Russia. andrei-matalin @ yandex. ru; https: // orcid. org / 0000 - 0002 - 7790 - 8709 & Pirogov National Research Medical University, Department of Biology, Ostrovitianova str. 1, Moscow 117997, Russia. andrei-matalin@yandex.ru Author Wiesner, Jürgen Dresdener Ring 11, D- 38444 Wolfsburg, Germany. text Zootaxa 2023 2023-03-22 5256 5 401 433 http://dx.doi.org/10.11646/zootaxa.5256.5.1 journal article 10.11646/zootaxa.5256.5.1 1175-5326 7758872 E5B9708E-D626-4923-825A-A9FD59AE4024 Therates pseudovitalisi Matalin et Wiesner , sp. n. Figs 9, 10 , 36, 37 , 58, 59 , 90–93 , 132–134 . Therates fruhstorferi vitalisi W. Horn, 1913 — Sawada & Wiesner 2000: 63–64 , 2004: 263–264 ; Wiesner & Geiser 2016: 88 . Type material. HOLOTYPE , 1Ô— Laos , Luang Prabang Prov. , Ban Song Cha ( 5 km W), 20 o 33–4’N 102 o 14’E , ± 1200 m , 24.4– 16.5.1999 , leg. Vit Kubaň & C. Holzschuh ( JW ) ; PARATYPES : 1♀ —same labeled as the holotype ( JW ) ; 3ÔÔ 6♀♀ — Laos , Oudom Xai ( 17 km NEE ) , 20 o 45’N 102 o 09’E , ~ 1100 m , 18.4– 1.5.1998 , leg. Vit Kubaň (2ÔÔ 5♀♀ — JW , 1Ô 2♀♀ — MSPU ) ; 1Ô— Laos , Houaphanh Prov. , Phou Pane Mt , VII.2011 ( MSPU ) ; 1♀ — Laos NE, Sam [ = Xam ] Neua , 17. V .2002 ( JW ) ; 1Ô—Laos-N, Phongsaly prov. , Ban Bun Neua [= Boun Neua ] ( 4 km E), 20. VI .2003 , ~ 1100 m , 21 o 38’N 101 o 57’E , P. Pacholátko leg. ( JW ) . References. LAOS , Houaphanh Prov. : 1Ô 2♀♀ — Xam Nua Distr , Ban Saleui , 1350 m , 15.VII.2002 ; 1Ô— Xam Nua Distr , Phu Pan , 1800 m , 17–21.VI.2003 (both Sawada & Wiesner 2004 ) ; 1♀ — Phou Pane Mt. , 1480–1550 m , 20 o 13’09–19’’N 103º59’54’’E , 1–16.VI.2009 , leg. Z. Kraus ( Wiesner & Geiser 2016 ) . Diagnosis. This new species is readily distinguished from T. vitalisi , T. biserratus , T. hunanensis sp. n. , T. sauteri and T. motoensis by lacking lateral carinae on the apex of aedeagus ( Figs 132–134 vs. Figs 152–174 ). From T. fruhstorferi and T. ziyardamensis sp. n. a new species is easily distinguished by the shape of aedeagus ( Figs 132– 134 vs. Figs 115–131 ), from T. fruhstorferi additionally by wider apices of elytra ( Figs 9–11 , 90–95 vs. Figs 1, 3–8 , 81–89 ) and by shorter hind tarsus ( Figs 175–178 ). From T. pseudovitalisi tibetana ssp. n. a nominotypical subspecies is clearly differentiated by the shape of antennomeres 9–11 ( Figs 36, 37 vs. Figs 38, 39 ), by the proportions of hind tibiae and tarsi—mean HTbL/HTaL = 1.32 vs. HTbL/HTaL = 1.25, by narrow, elongate, clearly oblique central dot ( Figs 9, 10 , 90–93 vs. Figs 11 , 94, 95 ), and by the longer elytra—mean EL/AL = 2.6 vs. mean EL/AL = 2.48. Description. TL = 10.53–11.1 mm in males (mean = 10.74 mm , n = 5), 10.5–12.1 mm (mean = 11.17 mm , n = 7) in females. Head shining black-blue with purple-violet reflection; orbital plates finely striated in posterior third; frons convex, semi-circular (see above), slightly downward anteriorly (see at the side), smooth or slightly rugose, frontal sulci deep and slightly convergent in anterior 2/3, shallow and distinctly divergent in posterior third. Mandibles in males brown underside, yellow-brown topside with brown teeth and apical molar; in females dark-brown except pale basal quarter topside. Labial palpi yellow-brown or brown except dark-brown apical palpomeres; maxillary palpi dark brown with narrow light-brown apices in all palpomeres. Antennae barely extend shoulders, black, scape dark brown on anterior margin and black on posterior margin, in some females brown-black, antennomeres 2–5 with light violet tinge and narrow dark-brown apices, antennomeres 9–11 in males distinctly dilated, antennomeres 9 and 10 with wide protruding downward anterior lower margins ( Fig. 37 ), in female only slightly delated and protruding ( Fig. 36 ). Labrum indistinctly transverse, LW/LL = 1.0–1.08 (mean = 1.04, n = 12), in males black-brown with medium-sized yellow-brown apical spot indistinctly separated from central apical teeth ( Fig. 58 ), in females entirely black ( Fig. 59 ). Pronotum shining blue with violet reflection; indistinctly transverse, PW/PL = 1.0–1.11 (mean = 1.06, n = 12), apical lobe practically equal wide throughout; thorax shining black-blue with light golden-violet reflection. Femora bicolored, fore femora pale-yellow with brown violet-tinged anterior side, black with violet tinge on posterior side, middle femora brown-black with light violet tinge except pale-yellow posterior basal third, hind femora pale-yellow in basal third or quarter and black with violet tinge in apical 2/3 or 3/4; tibiae brown-black with blue-violet lustre; tarsi black with blue-violet tinge; HTbL/HTaL = 1.25–1.42 (mean = 1.32, n = 12) ( Figs 175–178 ). Elytra shining black-blue with purple-violet or blue-violet reflection; indistinctly divergent toward apex, EL/ EW = 1.92–2.06 (mean = 1.98, n = 12); punctuation deep and regular between basal humps and central dot, in apical third shallow and sparse; scutellum black with blue or violet tinge; apical margin shortly cut. Elytral pattern presented by well visible pale-yellow small basal portion of humeral lunule (see in front) and elongate slightly or clearly oblique narrow wide-uniformed or slightly dilated in lateral portion central dot; basal dot very small, in some specimens indistinct or entirely absent ( Figs 9, 10 , 90–93 ). Aedeagus gradually sinuate with elongate rounded apex, lacking lateral carinae ( Figs 132–134 ); AL = 2.5–2.7 mm (mean = 2.62, n = 5), EL/AL = 2.56–2.72 (mean = 2.6, n = 5), in one aberrant male EL/AL = 2.45. Etymology. The new species is named “ pseudovitalisi ” due to it very similar outward appearance with T. vitalisi Horn, 1913 . Distribution. LAOS : Oudomxay , Luang Prabang , Houaphanh , Phongsaly Provinces ( Fig. 184 ).