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<document id="AECC30C639BDBC4BA8C10D36EAED1171" ID-CLB-Dataset="6015" ID-DOI="10.11646/zootaxa.4980.2.1" ID-GBIF-Dataset="f94c88c7-ef33-4171-a0d2-23832db1e2ba" ID-ISSN="1175-5326" ID-Zenodo-Dep="4888860" ID-ZooBank="344CA83A-6C54-4425-8A1F-1BA540611214" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622531830003" checkinUser="plazi" docAuthor="Abé, Hiroshi" docDate="2021" docId="022487BFFFC1FF82FF24F943FE74FDDF" docLanguage="en" docName="zootaxa.4980.2.1.pdf" docOrigin="Zootaxa 4980 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scaptognathus hallezi Trouessart 1894" docType="treatment" docVersion="6" lastPageNumber="230" masterDocId="FE1DFFC7FFDDFF9FFFB3FFE1FFB6FFE2" masterDocTitle="Taxonomic review of the subfamily Lohmannellinae (Acari: Trombidiformes Halacaridae) with morphological comparisons among genera" masterLastPageNumber="255" masterPageNumber="201" pageNumber="229" updateTime="1698995727304" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5B387D380907868573ECDF0F4BDCF3FF">Taxonomic review of the subfamily Lohmannellinae (Acari: Trombidiformes Halacaridae) with morphological comparisons among genera</mods:title>
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<treatment id="022487BFFFC1FF82FF24F943FE74FDDF" ID-DOI="http://doi.org/10.5281/zenodo.4985127" ID-GBIF-Taxon="182958168" ID-Zenodo-Dep="4985127" LSID="urn:lsid:plazi:treatment:022487BFFFC1FF82FF24F943FE74FDDF" httpUri="http://treatment.plazi.org/id/022487BFFFC1FF82FF24F943FE74FDDF" lastPageId="29" lastPageNumber="230" pageId="28" pageNumber="229">
<subSubSection id="C2976522FFC1FF83FF24F943FDD4F95E" box="[151,610,1698,1725]" pageId="28" pageNumber="229" type="nomenclature">
<paragraph id="8A3236A9FFC1FF83FF24F943FDD4F95E" blockId="28.[151,1437,1698,2013]" box="[151,610,1698,1725]" pageId="28" pageNumber="229">
<heading id="D17A81C5FFC1FF83FF24F943FDD4F95E" bold="true" box="[151,610,1698,1725]" fontSize="11" level="1" pageId="28" pageNumber="229" reason="1">
<taxonomicName id="4D8D4D2AFFC1FF83FF24F943FDD4F95E" authority="Trouessart, 1894" authorityName="Trouessart" authorityYear="1894" box="[151,610,1698,1725]" class="Arachnida" family="Halacaridae" genus="Scaptognathus" kingdom="Animalia" order="Trombidiformes" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="hallezi">
<emphasis id="B8F9EABBFFC1FF83FF24F943FDD4F95E" bold="true" box="[151,610,1698,1725]" pageId="28" pageNumber="229">
<emphasis id="B8F9EABBFFC1FF83FF24F943FE23F95F" bold="true" box="[151,405,1698,1725]" italics="true" pageId="28" pageNumber="229">Scaptognathus hallezi</emphasis>
Trouessart, 1894
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C2976522FFC1FF82FF74F926FE74FDDF" lastPageId="29" lastPageNumber="230" pageId="28" pageNumber="229" type="diagnosis">
<paragraph id="8A3236A9FFC1FF83FF74F926FD6BF83F" blockId="28.[151,1437,1698,2013]" pageId="28" pageNumber="229">
Diagnosis. Glp-1 same level as Ds-1. Ds-2 on
<collectingRegion id="4849F84BFFC1FF83FD52F926FCBBF903" box="[737,781,1735,1761]" country="Serbia" name="Ada" pageId="28" pageNumber="229">AD</collectingRegion>
. PE with 1 dorsal seta. GA bipartite in female and male. Female with 3 or 4 Pgs and 4 Sgs. Male with 30–40 Pgs and 6 Sgs. Tr I–IV with 1-1-1-1 setae; Bf I–IV with 2-(1,2)- (1,2)-(1,2) setae; Tf I–IV with (4,5)-4-2-2 setae, 2-0-0-0 Bp setae; Gn I–IV with (2,4)-(3,5)-3-3 setae, 2-1-0-0 Bp setae; Tb I–IV with (7,9)-6-(6,7)-(5,7) setae, 6-3-4-4 Bp setae; Ta I–IV with 3-3-3-3 dorsal setae, 1-1-0-0 Bp setae. Lc I with accessory process.
<taxonomicName id="4D8D4D2AFFC1FF83FE6FF8B6FDF2F893" authorityName="Ausserer" authorityYear="1871" box="[476,580,1879,1905]" class="Arachnida" family="Idiopidae" genus="Idiosoma" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="240" phylum="Arthropoda" rank="genus">Idiosoma</taxonomicName>
length
<quantity id="4D759B4CFFC1FF83FD2AF8B6FCAEF893" box="[665,792,1879,1905]" metricMagnitude="0" metricUnit="m" metricValue="6.134099999999999" metricValueMax="6.603999999999999" metricValueMin="5.664199999999999" pageId="28" pageNumber="229" unit="in" value="241.5" valueMax="260.0" valueMin="223.0">223–260 in</quantity>
female,
<quantity id="4D759B4CFFC1FF83FCCAF8B6FC4FF893" box="[889,1017,1879,1905]" metricMagnitude="0" metricUnit="m" metricValue="5.892799999999999" metricValueMax="6.1213999999999995" metricValueMin="5.664199999999999" pageId="28" pageNumber="229" unit="in" value="232.0" valueMax="241.0" valueMin="223.0">223–241 in</quantity>
male.
<taxonomicName id="4D8D4D2AFFC1FF83FBF7F8B6FB1AF893" authorityName="Ausserer" authorityYear="1871" box="[1092,1196,1879,1905]" class="Arachnida" family="Idiopidae" genus="Idiosoma" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="240" phylum="Arthropoda" rank="genus">Idiosoma</taxonomicName>
width 126 (
<taxonomicName id="4D8D4D2AFFC1FF83FA87F8B6FA2BF893" authorityName="Ausserer" authorityYear="1871" box="[1332,1437,1879,1905]" class="Arachnida" family="Idiopidae" genus="Idiosoma" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="240" phylum="Arthropoda" rank="genus">idiosoma</taxonomicName>
length 231) in female, 145–148 (
<taxonomicName id="4D8D4D2AFFC1FF83FDB6F89AFDD8F877" authorityName="Ausserer" authorityYear="1871" box="[517,622,1915,1941]" class="Arachnida" family="Idiopidae" genus="Idiosoma" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="240" phylum="Arthropoda" rank="genus">idiosoma</taxonomicName>
length 237–240) in male.
<taxonomicName id="4D8D4D2AFFC1FF83FC26F89AFB95F877" authorityName="Krefft" authorityYear="1871" box="[917,1059,1915,1941]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="28" pageNumber="203" phylum="Nematoda" rank="genus">Gnathosoma</taxonomicName>
length 148–167 (
<taxonomicName id="4D8D4D2AFFC1FF83FB54F89AFAE6F877" authorityName="Ausserer" authorityYear="1871" box="[1255,1360,1915,1941]" class="Arachnida" family="Idiopidae" genus="Idiosoma" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="240" phylum="Arthropoda" rank="genus">idiosoma</taxonomicName>
length 223–260) in female, 148–155 (
<taxonomicName id="4D8D4D2AFFC1FF83FE45F87EFDE9F85B" authorityName="Ausserer" authorityYear="1871" box="[502,607,1951,1977]" class="Arachnida" family="Idiopidae" genus="Idiosoma" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="240" phylum="Arthropoda" rank="genus">idiosoma</taxonomicName>
length 223–241) in male. Ratio of
<taxonomicName id="4D8D4D2AFFC1FF83FC42F87EFBCEF85B" authorityName="Krefft" authorityYear="1871" box="[1009,1144,1951,1977]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="28" pageNumber="203" phylum="Nematoda" rank="genus">gnathosoma</taxonomicName>
to
<taxonomicName id="4D8D4D2AFFC1FF83FB2CF87EFAB0F85B" authorityName="Ausserer" authorityYear="1871" box="[1183,1286,1951,1977]" class="Arachnida" family="Idiopidae" genus="Idiosoma" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="240" phylum="Arthropoda" rank="genus">idiosoma</taxonomicName>
0.66–0.71 in female, 0.62–0.69 in male. Ratio of P-2 to P-1 3.17.
</paragraph>
<paragraph id="8A3236A9FFC0FF82FF63FF76FB9DFF1B" blockId="29.[151,1437,151,573]" pageId="29" pageNumber="230">
Remarks. This species is similar to
<taxonomicName id="4D8D4D2AFFC0FF82FDD2FF76FD42FF53" authorityName="Andre" authorityYear="1961" box="[609,756,151,177]" class="Arachnida" family="Halacaridae" genus="Scaptognathus" kingdom="Animalia" order="Trombidiformes" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="sabularius">
<emphasis id="B8F9EABBFFC0FF82FDD2FF76FD42FF53" box="[609,756,151,177]" italics="true" pageId="29" pageNumber="230">S. sabularius</emphasis>
</taxonomicName>
and
<taxonomicName id="4D8D4D2AFFC0FF82FC9FFF76FC06FF52" authorityName="Bartsch" authorityYear="1986" box="[812,944,151,177]" class="Arachnida" family="Halacaridae" genus="Scaptognathus" kingdom="Animalia" order="Trombidiformes" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="tereninus">
<emphasis id="B8F9EABBFFC0FF82FC9FFF76FC06FF52" box="[812,944,151,177]" italics="true" pageId="29" pageNumber="230">S. tereninus</emphasis>
</taxonomicName>
in having Ds-2 on
<collectingRegion id="4849F84BFFC0FF82FB3AFF76FB03FF53" box="[1161,1205,151,177]" country="Serbia" name="Ada" pageId="29" pageNumber="230">AD</collectingRegion>
. However,
<taxonomicName id="4D8D4D2AFFC0FF82FA86FF76FA2BFF53" authorityName="Trouessart" authorityYear="1894" box="[1333,1437,151,177]" class="Arachnida" family="Halacaridae" genus="Scaptognathus" kingdom="Animalia" order="Trombidiformes" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="hallezi">
<emphasis id="B8F9EABBFFC0FF82FA86FF76FA2BFF53" box="[1333,1437,151,177]" italics="true" pageId="29" pageNumber="230">S. hallezi</emphasis>
</taxonomicName>
differs from the latter two species in the shape of GA in female. Bartsch (1986) suggested that
<taxonomicName id="4D8D4D2AFFC0FF82FB07FF5AFE86FF1B" authority="Morselli &amp; Mari, 1981" authorityName="Morselli &amp; Mari" authorityYear="1981" class="Arachnida" family="Halacaridae" genus="Scaptognathus" kingdom="Animalia" order="Trombidiformes" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="neretinus">
<emphasis id="B8F9EABBFFC0FF82FB07FF5AFA81FF36" box="[1204,1335,187,213]" italics="true" pageId="29" pageNumber="230">S. neretinus</emphasis>
Morselli &amp; Mari, 1981
</taxonomicName>
would be synonymous with
<taxonomicName id="4D8D4D2AFFC0FF82FDC0FF3EFD6CFF1B" authorityName="Trouessart" authorityYear="1894" box="[627,730,223,249]" class="Arachnida" family="Halacaridae" genus="Scaptognathus" kingdom="Animalia" order="Trombidiformes" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="hallezi">
<emphasis id="B8F9EABBFFC0FF82FDC0FF3EFD6CFF1B" box="[627,730,223,249]" italics="true" pageId="29" pageNumber="230">S. hallezi</emphasis>
</taxonomicName>
. Female and male are known.
</paragraph>
<paragraph id="8A3236A9FFC0FF82FF63FEE2FE8CFEA3" blockId="29.[151,1437,151,573]" pageId="29" pageNumber="230">
Habitat. Littoral–Sublittoral zones: Medium to coarse sandy sediment, algae, and bryozoans at intertidal zone to
<quantity id="4D759B4CFFC0FF82FF07FEC7FF5BFEA3" box="[180,237,294,321]" metricMagnitude="1" metricUnit="m" metricValue="5.8" pageId="29" pageNumber="230" unit="m" value="58.0">58 m</quantity>
depth.
</paragraph>
<paragraph id="8A3236A9FFC0FF82FF63FEAAFC0BFE4F" blockId="29.[151,1437,151,573]" pageId="29" pageNumber="230">
Distribution. North Atlantic Ocean: Roscoff, Granville [
<collectingCountry id="F29A7639FFC0FF82FCE4FEABFC1AFE87" box="[855,940,330,357]" name="France" pageId="29" pageNumber="230">France</collectingCountry>
]. Mediterranean: Île If, Île Ratonneau, Île Riou, Île Jarre, Plateau des Chevres, Envau, Port-Pin [
<collectingCountry id="F29A7639FFC0FF82FCB7FE8FFCEFFE6B" box="[772,857,366,393]" name="France" pageId="29" pageNumber="230">France</collectingCountry>
]. Lido Frascone (
<collectingRegion id="4849F84BFFC0FF82FB93FE8EFBDBFE68" box="[1056,1133,367,394]" country="Italy" name="Puglia" pageId="29" pageNumber="230">Puglia</collectingRegion>
), Lido dell’Ancora [
<collectingCountry id="F29A7639FFC0FF82FAE5FE8FFA27FE6B" box="[1366,1425,366,393]" name="Italy" pageId="29" pageNumber="230">Italy</collectingCountry>
]. Kaþ, Hamitbey Plajý (
<collectingRegion id="4849F84BFFC0FF82FE22FE72FE5BFE4C" box="[401,493,403,430]" country="Turkey" name="Antalya" pageId="29" pageNumber="230">Antalya</collectingRegion>
) [
<collectingCountry id="F29A7639FFC0FF82FDB0FE73FDEAFE4F" box="[515,604,402,429]" name="Turkey" pageId="29" pageNumber="230">Turkey</collectingCountry>
]. Indian Ocean: Waltair [
<collectingCountry id="F29A7639FFC0FF82FCC2FE73FC05FE4F" box="[881,947,402,429]" name="India" pageId="29" pageNumber="230">India</collectingCountry>
].
</paragraph>
<paragraph id="8A3236A9FFC0FF82FF63FE56FE08FDF8" blockId="29.[151,1437,151,573]" pageId="29" pageNumber="230">
References.
<bibRefCitation id="EE1C4B58FFC0FF82FEE4FE56FE53FE33" author="Abe, H." box="[343,485,439,465]" pageId="29" pageNumber="230" pagination="349 - 363" refId="ref41924" refString="Abe, H. (1990 c) Two new species of the genus Scaptognathus (Acari, Halacaridae) from Hokkaido, Japan. Cahiers de Biologie Marine, 31, 349 - 363." type="journal article" year="1990">Abé (1990c)</bibRefCitation>
,
<bibRefCitation id="EE1C4B58FFC0FF82FE5DFE56FD62FE33" author="Abe, H. &amp; Green, J." box="[494,724,439,465]" pageId="29" pageNumber="230" pagination="169 - 178" refId="ref42059" refString="Abe, H. &amp; Green, J. (1994) A new species of the genus Scaptognathus (Acari: Halacaridae) from Tasmania. Acta Arachnologica, 43, 169 - 178." type="journal article" year="1994">Abé &amp; Green (1994)</bibRefCitation>
,
<bibRefCitation id="EE1C4B58FFC0FF82FD6EFE56FCC0FE30" author="Andre, M." box="[733,886,439,466]" pageId="29" pageNumber="230" pagination="1 - 152" refId="ref42268" refString="Andre, M. (1946) Halacariens marins. Faune de France, 46, 1 - 152." type="journal article" year="1946">André (1946)</bibRefCitation>
,
<bibRefCitation id="EE1C4B58FFC0FF82FC33FE56FB98FE33" author="Bartsch, I." box="[896,1070,439,465]" pageId="29" pageNumber="230" pagination="231 - 244" refId="ref43737" refString="Bartsch, I. (1986 b) Lohmannellinae (Halacaroidea, Acari) aus dem Mittelmeer. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 8, 231 - 244." type="journal article" year="1986">Bartsch (1986b</bibRefCitation>
,
<bibRefCitation id="EE1C4B58FFC0FF82FB84FE56FBC8FE33" author="Bartsch, I." box="[1079,1150,439,465]" pageId="29" pageNumber="230" pagination="417 - 422" refId="ref43861" refString="Bartsch, I. (1988 c) Halacaroidea. In: Higgins, R. P. &amp; Thiel, H. (Eds.), Introduction to the study of Meiofauna. Smithsonian Institution Press, Washington, D. C., pp. 417 - 422." type="book chapter" year="1988">1988c</bibRefCitation>
,
<bibRefCitation id="EE1C4B58FFC0FF82FB3AFE56FB65FE30" author="Bartsch, I." box="[1161,1235,439,466]" pageId="29" pageNumber="230" pagination="61 - 74" refId="ref44260" refString="Bartsch, I. (1994 b) Sexualdimorphismus bei Meeresmilben (Halacaridae, Acari). Ubersicht und Beschreibung des Mannchens von Phacacarus flavellus. Zoologischer Anzeiger, 232, 61 - 74." type="journal article" year="1994">1994b</bibRefCitation>
),
<bibRefCitation id="EE1C4B58FFC0FF82FB56FE56FA2EFE33" author="Durucan, F." box="[1253,1432,439,465]" pageId="29" pageNumber="230" pagination="499 - 507" refId="ref46189" refString="Durucan, F. (2018) New record of the genus Scaptognathus (Acari: Halacaridae) from Antalya with a checklist of marine halacarid mites of Turkey. Turkish Journal of Zoology, 42, 499 - 507. https: // doi: 10.3906 / zoo- 1803 - 6" type="journal article" year="2018">Durucan (2018)</bibRefCitation>
,
<bibRefCitation id="EE1C4B58FFC0FF82FF24FE3AFE89FE17" author="Halbert, J. N." box="[151,319,474,501]" pageId="29" pageNumber="230" pagination="45 - 136" refId="ref47196" refString="Halbert, J. N. (1915) Clare Island Survey 39, Acarinida. Section II. Terrestrial and marine Acarina. Proceedings of the Royal Irish Academy, 31, 45 - 136." type="journal article" year="1915">Halbert (1915)</bibRefCitation>
,
<bibRefCitation id="EE1C4B58FFC0FF82FEFFFE3AFDBAFE17" author="Lohmann, H." box="[332,524,475,501]" pageId="29" pageNumber="230" pagination="273 - 305" refId="ref48316" refString="Lohmann, H. (1901) Fam. Halacaridae. Das Tierreich, 13, 273 - 305." type="journal article" year="1901">Lohmann (1901)</bibRefCitation>
,
<bibRefCitation id="EE1C4B58FFC0FF82FDABFE3AFD7AFE14" author="Monniot, F." box="[536,716,475,502]" pageId="29" pageNumber="230" pagination="491 - 498" refId="ref48974" refString="Monniot, F. (1964) Sur deux especes du genre Scaptognathus presentes a Roscoff. Acarologia, 6, 491 - 498." type="journal article" year="1964">Monniot (1964)</bibRefCitation>
, Morselli &amp; Mari (1984),
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</paragraph>
<paragraph id="8A3236A9FFC0FF82FF63FDC2FE74FDDF" blockId="29.[151,1437,151,573]" box="[208,450,547,573]" pageId="29" pageNumber="230">Depository. MNHNP.</paragraph>
</subSubSection>
</treatment>
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