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<document id="CFC65687AAC31EC5E55E9B49FFF95DBC" ID-CLB-Dataset="26687" ID-DOI="http://dx.doi.org/10.3897/evolsyst.2.25200" ID-GBIF-Dataset="a86a1df2-cbc2-4667-a53f-8684c0a50fa2" ID-Pensoft-Pub="2535-0730-1-65" ID-ZooBank="ACAEC71D964C4314AAA7A404F23A6569" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="2535-0730-1-65" ModsDocOrigin="Evolutionary Systematics 2 (1)" ModsDocTitle="Taxonomic descriptions of nine new species of the goblin spider genera Cavisternum, Grymeus, Ischnothyreus, Opopaea, Pelicinus and Silhouettella (Araneae, Oonopidae) from Sri Lanka" checkinTime="1558621782066" checkinUser="pensoft" docAuthor="Ranasinghe, U. G. S. L. &amp; Benjamin, Suresh P." docDate="2018" docId="BE1A24A959F2C48303E6B8CCF0CD67E7" docLanguage="en" docName="EvolutSyst 2(1): 65-80" docOrigin="Evolutionary Systematics 2 (1)" docSource="http://dx.doi.org/10.3897/evolsyst.2.25200" docTitle="Grymeus dharmapriyai n. sp." docType="treatment" docVersion="5" lastPageNumber="66" masterDocId="BD018C0FFFB0FFECC95EAF12A033FFD9" masterDocTitle="Taxonomic descriptions of nine new species of the goblin spider genera Cavisternum, Grymeus, Ischnothyreus, Opopaea, Pelicinus and Silhouettella (Araneae, Oonopidae) from Sri Lanka" masterLastPageNumber="80" masterPageNumber="65" pageNumber="65" updateTime="1701379503719" updateUser="ExternalLinkService">
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<mods:title id="CDA101C092570EDC8B0CA1304007360E">Taxonomic descriptions of nine new species of the goblin spider genera Cavisternum, Grymeus, Ischnothyreus, Opopaea, Pelicinus and Silhouettella (Araneae, Oonopidae) from Sri Lanka</mods:title>
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<mods:name id="2C278DD487A337DEEEF8E752E62D1660" type="personal">
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<mods:namePart id="B242789636D27772F3AFFBB00F7061C2">Ranasinghe, U. G. S. L.</mods:namePart>
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<mods:namePart id="E11A2EAB40288ACB1BFDA3214641A147">Benjamin, Suresh P.</mods:namePart>
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<mods:title id="11CBEBEEFBB2A70BC9ED4268F0C1DEFF">Evolutionary Systematics</mods:title>
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<mods:date id="93B0807E86864B89799BDB490DCD0532">2018</mods:date>
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<mods:classification id="C7F8EAB42E7E80BE1633173D8239EB3F">journal article</mods:classification>
<mods:identifier id="AA79F45FA7566A011E7216FF28CEEC7D" type="DOI">http://dx.doi.org/10.3897/evolsyst.2.25200</mods:identifier>
<mods:identifier id="744AC765EA56311F4609EED046365B1B" type="Pensoft-Pub">2535-0730-1-65</mods:identifier>
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<treatment id="BE1A24A959F2C48303E6B8CCF0CD67E7" ID-GBIF-Taxon="157251956" LSID="urn:lsid:plazi:treatment:BE1A24A959F2C48303E6B8CCF0CD67E7" httpUri="http://treatment.plazi.org/id/BE1A24A959F2C48303E6B8CCF0CD67E7" lastPageId="1" lastPageNumber="66" pageId="0" pageNumber="65">
<subSubSection id="1315EBD4CE3051F677576DEF495CD040" pageId="0" pageNumber="65" type="nomenclature">
<paragraph id="2A75EAC77F265F32C45F8EFBFF554EE3" pageId="0" pageNumber="65">
<taxonomicName id="325822729C30F7411FDEDC008711FFC1" ID-CoL="3HHBK" LSID="http://zoobank.org/CDD21286-ED78-48F5-B758-63243C16F4F1" authority="n. sp." class="Arachnida" family="Oonopidae" genus="Grymeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Grymeus dharmapriyai" order="Araneae" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="dharmapriyai">Grymeus dharmapriyai n. sp.</taxonomicName>
Figs 3
<normalizedToken id="300D7FA9351CB4DC0AB4A596CBD4DD35" originalValue="AJ">A-J</normalizedToken>
, 4
<normalizedToken id="5E55C288D61933DD96C3E5839EC3BF0E" originalValue="AE">A-E</normalizedToken>
, 5
<normalizedToken id="84BC0D635F538F236F47B89F0710708E" originalValue="AB">A-B</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection id="9B914D00195409FE94404B25C34CDEB7" pageId="1" pageNumber="66" type="type material">
<paragraph id="84F357390B4F04177A975607BDC5538E" pageId="1" pageNumber="66">
<pageBreakToken id="FC87CD3F9B26EFF7B00B068DC63FE964" pageId="1" pageNumber="66" start="start">Type</pageBreakToken>
material.
</paragraph>
<paragraph id="EFF30A7B22E2DCE799CAB8CC352A5591" pageId="1" pageNumber="66">
Male holotype from Sri Lanka, North Western Province, Puttalam District, Wanathavilluwa,
<geoCoordinate id="1C50616C427AC52FFD2B402C7763E544" direction="north" orientation="latitude" precision="15" value="8.170834">08°10'15&quot;N</geoCoordinate>
,
<geoCoordinate id="C29C52629EA3362379EED694569A2325" direction="east" orientation="longitude" precision="15" value="79.875">79°52'30&quot;E</geoCoordinate>
, 30 m, litter; 24 May 2010; leg. N. Athukorala (IFS_Oon_007) (ZFMK).
</paragraph>
</subSubSection>
<subSubSection id="8318DD8D3D759647EA0EFCD699D7E5D3" pageId="1" pageNumber="66" type="paratypes">
<paragraph id="22C45AF865236C7B5F86BFDEEF926F9C" pageId="1" pageNumber="66">Paratypes.</paragraph>
<paragraph id="62AB1A7E5DC620FC661B746A68C54BAD" pageId="1" pageNumber="66">1 male and 3 females; same locality data (IFS_Oon_008-011) (ZFMK).</paragraph>
</subSubSection>
<subSubSection id="EC84E5753A53E55F4E55118B0E3D55F9" pageId="1" pageNumber="66" type="materials_examined">
<paragraph id="4BC32421EC98045E0C5C8DC5A406DCA4" pageId="1" pageNumber="66">Other material examined.</paragraph>
<paragraph id="BC0801B6268311346460529B8E4C4D8A" pageId="1" pageNumber="66">
2 males; Sri Lanka, Central Province, Kandy District, Gannoruwa FR,
<geoCoordinate id="914B06DE8A7EFA695781605009E0F6A5" direction="north" orientation="latitude" precision="1" value="7.282459">07°16'56.85&quot;N</geoCoordinate>
,
<geoCoordinate id="E155A91E2907AAEB53D8E92998586398" direction="east" orientation="longitude" precision="1" value="80.599236">80°35'57.25&quot;E</geoCoordinate>
, 575m, litter; 25 September 2013; leg. N. Athukorala et al. (IFS_Oon_156, 166). 1 male; Sri Lanka, Northern Province, Mannar District, surroundings of Vayu resorts-kitesurfing, Thaleimannar,
<geoCoordinate id="3A9DA1C57C377CD7DFCB92D7D1FE848C" direction="north" orientation="latitude" precision="15" value="9.082222">09°04'56&quot;N</geoCoordinate>
,
<geoCoordinate id="935644A259FD3A886DF45ED991B42758" direction="east" orientation="longitude" precision="15" value="79.70111">79°42'4&quot;E</geoCoordinate>
, 4m, litter; 03 April 2018; leg. S.P. Benjamin et al. (IFS_Oon_502).
</paragraph>
</subSubSection>
<subSubSection id="13063D4CA6E6E9C917717AA134BD12A8" pageId="1" pageNumber="66" type="diagnosis">
<paragraph id="28417BE9675F26FD7050C3A5194A6BDF" pageId="1" pageNumber="66">Diagnosis.</paragraph>
<paragraph id="712D1CC42F83B66C36C1E80B2A8B0BD9" pageId="1" pageNumber="66">
The male of
<taxonomicName id="FB3EE144243A779D41773A4057437D7D" lsidName="G. dharmapriyai" pageId="1" pageNumber="66" rank="species" species="dharmapriyai">G. dharmapriyai</taxonomicName>
n. sp. resembles to
<taxonomicName id="494C0BC33B50755C62745B87CE6EEEC0" lsidName="G. robertsi" pageId="1" pageNumber="66" rank="species" species="robertsi">G. robertsi</taxonomicName>
Harvey, 1987 by carapace without longitudinal rows of long, stout dorsal setae (Fig. 3A) and sperm pore situated in anterior third of abdomen (Fig. 3H), but differ in having the embolic accessory appendage with a dorsal, tooth-like projection (to) and a sharp, pointed embolus (Fig. 5A). Females are recognized by the stout and rounded receptaculum (Fig. 5B).
</paragraph>
<caption id="DBD793DDADF561CB0212B41C1D56C61A" pageId="1" pageNumber="66">
<paragraph id="7E66472B4763EB8B7895D8A17116F07B" pageId="1" pageNumber="66">
Figure 3.
<taxonomicName id="9D0001324F313F810E5A78CD5DEA25E5" class="Arachnida" family="Oonopidae" genus="Grymeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Grymeus dharmapriyai" order="Araneae" pageId="1" pageNumber="66" phylum="Arthropoda" rank="species" species="dharmapriyai">Grymeus dharmapriyai</taxonomicName>
n. sp., male from Wanathavilluwa: A, carapace, dorsal view; B, sternum, ventral view; C, carapace, lateral view; D, same, anterior view; E, same, posterior view; F, abdomen, anterior view; G, same, dorsal view; H, same, ventral view; I, same, lateral view; J, same, posterior view. Scale bars: 0.2 mm (
<normalizedToken id="A87339821BA086DB9B17322C3FD1674E" originalValue="AF">A-F</normalizedToken>
, J), 0.5 mm (
<normalizedToken id="0AAD29A81BC8D40D76B7CD5ABB4EA948" originalValue="GI">G-I</normalizedToken>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="4ECC7A3BD69F13B9FF9F373E3EAA3EF0" pageId="1" pageNumber="66" type="etymology">
<paragraph id="CBD9AA1D6E21829B9D3B5482C709E474" pageId="1" pageNumber="66">Etymology.</paragraph>
<paragraph id="ED7D29C3A7103D79ED2C3E3BE528C127" pageId="1" pageNumber="66">
The species is named for the first
<normalizedToken id="D205C176076B31E81314DEB15F036417" originalValue="authors">author's</normalizedToken>
husband P. L. Dharmapriya.
</paragraph>
</subSubSection>
<subSubSection id="B87D81A79B2F2BADBB1875BB234C3FCF" pageId="1" pageNumber="66" type="description">
<paragraph id="BB1C1B38BBEADA7A0301812AF881E740" pageId="1" pageNumber="66">Description.</paragraph>
<paragraph id="C8FF7B17008FEE284225044FA173C8A8" pageId="1" pageNumber="66">Male: Total length 2.12 (Carapace, L: 0.84, W: 0.34. Abdomen, L: 1.28, W; 0.92). Coloration: uniformly orange brown colored. Carapace pyriform in dorsal view (Fig. 3A), pars cephalica gradually elevated and sudden elevation at middle (Fig. 3C), posterior circular shaped in dorsal view, but anterior rectangular shaped narrowed to less than 0.5 times its maximum width (Fig. 3A), posterior lateral spikes absent, lateral margin undulated. Clypeus straight in front view, margin unmodified, ALE separated from edge of carapace by about their diameter, setae absent. Six eyes, well developed (Fig. 3D), ALE largest, oval, separated each other by less than their diameter, posterior eyes circular, subequal, touching each other. Sternum heart-shaped (Fig. 3B), sharp margin visible, fused to carapace, radial furrows absent, decorated with rounded spots, lateral margin without indented extensions between coxae, posterior margin not extend. Cheliceral fangs unmodified, labium rounded, anterior invagination of sternum form a pouch (Fig. 3B). Abdomen elongated (Fig. 3G), evenly scattered with short setae, dorsal scutum covering full length of abdomen, no soft tissue visible from above, not fused to epigastric scutum, spinnerets not visible in dorsal view. Epigastric scutum sclertized, book lung covers pale brown, oval shaped, setaceous and not pointed (Fig. 3F). Postepigastric scutum long, fused to epigastric scutum (Fig. 3H). Spinneret scutum present as an incomplete ring. Leg spines absent, short brush like setae on all segments in leg I, II, III and IV. Genitalia: Palp of normal size, not strongly sclerotized, cymbium not extending beyond bulb, covered 2/3 of the bulb. Embolus (em), embolic accessory appendage (ma) and conductor (co) sharp and pointed, distal part of embolic accessory appendage bears a tooth-like projection (to) dorsally, slightly shorter than embolus, conductor short, curved ventrally (Fig. 5A).</paragraph>
<paragraph id="C64FBFA3630748AA0998D428223B41E0" pageId="1" pageNumber="66">Female: Total length 3.10 (Carapace, L:1.22, W:0.82; Abdomen, L: 1.80, W: 1.24). In general similar to males, larger than male, pars cephalica elevated from in lateral view, surface sides granulated (Figs 4A, 4C), lateral margin slightly undulated, sternum with hairs, not decorated with rounded spots (Fig. 4D), setae presence on chelicera, book lung covers small, narrowed, differ from male (Fig. 4B). Genitalia: Receptaculum (re) stout, rounded shaped, secretory sac (ssa) oval, pore field of receptaculum (pof) concentrated at middle of secretory sac, globular appendix (gap) short, less than 0.5 times the length of receptaculum, with an anterior paddle like sclerite (psc) and a nail like process (na), with lateral sclerites, with lateral apodemes (lap) (Fig. 5B).</paragraph>
</subSubSection>
<subSubSection id="B70920DE2FFA72E3A78F6255962340D5" pageId="1" pageNumber="66" type="distribution">
<paragraph id="87876EA4EF6A1579FC621D5B32587509" pageId="1" pageNumber="66">Distribution.</paragraph>
<paragraph id="4337CA34EAC68E39D03EA2B7A0931C23" pageId="1" pageNumber="66">Wanathavilluwa, Gannoruwa FR, Thaleimannar (Fig. 21).</paragraph>
<caption id="AE8C18109567EF183E91DFC996BB5693" pageId="1" pageNumber="66">
<paragraph id="7921E72F5C8FF3C00C217E8DD0723152" pageId="1" pageNumber="66">
Figure 4.
<taxonomicName id="7D0ACA85A3CEBAB2FCE13B31CF9AD1A7" class="Arachnida" family="Oonopidae" genus="Grymeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Grymeus dharmapriyai" order="Araneae" pageId="1" pageNumber="66" phylum="Arthropoda" rank="species" species="dharmapriyai">Grymeus dharmapriyai</taxonomicName>
n. sp., female from Wanathavilluwa: A, habitus, dorsal view; B, same, ventral view; C, carapace, dorsal view; D, sternum, ventral view; E, abdomen, ventral view. Scale bars: 1 mm (
<normalizedToken id="DC4E47E4434C7FA1B91504BA1BE532EE" originalValue="AB">A-B</normalizedToken>
), 0.5 mm (
<normalizedToken id="43F697CA216B887C098AB7B0B0B4642A" originalValue="CE">C-E</normalizedToken>
).
</paragraph>
</caption>
<caption id="3AB0946251ED414373155EF420151BF2" pageId="1" pageNumber="66">
<paragraph id="EAB219A19B8257BC16547DB6C09E6374" pageId="1" pageNumber="66">
Figure 5.
<taxonomicName id="EECB540BE61AC42420BF9AC23CD20C6C" class="Arachnida" family="Oonopidae" genus="Grymeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Grymeus dharmapriyai" order="Araneae" pageId="1" pageNumber="66" phylum="Arthropoda" rank="species" species="dharmapriyai">Grymeus dharmapriyai</taxonomicName>
n. sp., from Wanathavilluwa: A, male left palp, distal part, retrolateral view; B, female epigastric region, dorsal view. Abbreviations: co, conductor; em, embolus; gap, globular appendix; lap, lateral apodemes; ma, embolic accessory appendage; na, nail like process; psc, paddle-like sclerite; re, receptaculum; ssa, secretory sac; to, tooth like projection. Scale bars: 0.1 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>