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<document id="841F66442F1131C4D64574D28937BA12" ID-CLB-Dataset="39033" ID-DOI="10.11646/zootaxa.3658.1.1" ID-GBIF-Dataset="0adf09ba-5301-4828-abf6-8dbe89f041ca" ID-ISSN="1175-5326" ID-Zenodo-Dep="246551" ID-ZooBank="C02D1C74-25C0-41DD-B098-62098EB7B62A" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1463647879570" checkinUser="plazi" docAuthor="Dragan P. Chobanov, Beata Grzywacz, Ionuţ Ş. Iorgu, Battal Cιplak, Maya B. Ilieva &amp; Elżbieta Warchałowska-Śliwa" docDate="2013" docId="F26F3128393DFFA5B1B00A94FA489AAF" docLanguage="en" docName="zt03658p081.pdf.imf" docOrigin="Zootaxa 3658 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Isophya" docType="key" docVersion="12" lastPageNumber="56" masterDocId="0E5649503908FF92B1270A03FFD39875" masterDocTitle="Review of the Balkan Isophya (Orthoptera: Phaneropteridae) with particular emphasis on the Isophya modesta group and remarks on the systematics of the genus based on morphological and acoustic data" masterLastPageNumber="81" masterPageNumber="1" pageNumber="54" updateTime="1698662571276" updateUser="ExternalLinkService">
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<mods:title id="33A0F185B546D3DA384145208FF57978">Review of the Balkan Isophya (Orthoptera: Phaneropteridae) with particular emphasis on the Isophya modesta group and remarks on the systematics of the genus based on morphological and acoustic data</mods:title>
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<mods:name id="01479A6C6DA8A81A73092BDC3B4C5676" type="personal">
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<mods:namePart id="0102F3DA29D436B5CB64B2FA2876680E">Dragan P. Chobanov</mods:namePart>
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</mods:role>
<mods:namePart id="3F5313B76C4F5E5B5702C674B89EAC08">Beata Grzywacz</mods:namePart>
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<mods:namePart id="4219D619CFD3D440EC02BDE5FD5A9AAF">Ionuţ Ş. Iorgu</mods:namePart>
</mods:name>
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<mods:namePart id="23FFA7792FC08731A56865EA5BC21473">Battal Cιplak</mods:namePart>
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<mods:namePart id="B23953F5A2326546A058B1A701368C00">Maya B. Ilieva</mods:namePart>
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<mods:roleTerm id="FFEB2A45B18AF2B9C9520F38B4B24EEE">Author</mods:roleTerm>
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<mods:namePart id="54F5386AEF7ED6D4C1279C091879FE50">Elżbieta Warchałowska-Śliwa</mods:namePart>
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<mods:title id="74C1AC9B4EAC9A60C560D20793C65BBB">Zootaxa</mods:title>
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<mods:date id="AC8B86AB6BB18752E38575EB3E1939AD">2013</mods:date>
<mods:detail id="DCF2D7F050774C638AA7214C21F6314C" type="volume">
<mods:number id="DCF32651E6EE818F22D1A5E60BF0E20E">3658</mods:number>
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<mods:number id="E99D69B4488085D5F423169744999AA2">1</mods:number>
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<mods:classification id="53BD1FDEAA0ACFDA970DF8C167BEF101">journal article</mods:classification>
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<mods:identifier id="DB6079627C30F5C32F165D6564A29736" type="DOI">10.11646/zootaxa.3658.1.1</mods:identifier>
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<subSection id="134999D5393DFFA5B1B00A94FA489AAF" lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54" type="multiple">
<treatment id="F26F3128393DFFA5B1B00A94FA489AAF" ID-DOI="http://doi.org/10.5281/zenodo.5617404" ID-GBIF-Taxon="120682285" ID-Zenodo-Dep="5617404" LSID="urn:lsid:plazi:treatment:F26F3128393DFFA5B1B00A94FA489AAF" httpUri="http://treatment.plazi.org/id/F26F3128393DFFA5B1B00A94FA489AAF" lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54">
<subSubSection id="32DCD3B5393DFFA7B1B00A94FCD398C4" box="[151,768,151,177]" pageId="53" pageNumber="54" type="nomenclature">
<paragraph id="7A79803E393DFFA7B1B00A94FCD398C4" blockId="53.[151,768,151,177]" box="[151,768,151,177]" pageId="53" pageNumber="54">
<heading id="21313752393DFFA7B1B00A94FCD398C4" bold="true" box="[151,768,151,177]" fontSize="11" level="1" pageId="53" pageNumber="54" reason="1">
<emphasis id="48B25C2C393DFFA7B1B00A94FCD398C4" bold="true" box="[151,768,151,177]" pageId="53" pageNumber="54">
A key to identification the Balkan taxa of
<taxonomicName id="BDC6FBBD393DFFA7B3870A94FCD398C4" box="[672,768,151,177]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="48B25C2C393DFFA7B3870A94FCD398C4" bold="true" box="[672,768,151,177]" italics="true" pageId="53" pageNumber="54">Isophya</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="32DCD3B5393DFFA7B1B00ADCFB94996F" pageId="53" pageNumber="54" type="discussion">
<paragraph id="7A79803E393DFFA7B1B00ADCFB94996F" blockId="53.[151,1436,222,282]" pageId="53" pageNumber="54">
Note:
<taxonomicName id="BDC6FBBD393DFFA7B1FB0AE3FEA39883" box="[220,368,222,247]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="brevicauda">
<emphasis id="48B25C2C393DFFA7B1FB0AE3FEA39883" box="[220,368,222,247]" italics="true" pageId="53" pageNumber="54">I. brevicauda</emphasis>
</taxonomicName>
was recorded in
<collectingCountry id="02D1C0AE393DFFA7B3090ADDFD539883" box="[558,640,222,246]" name="Croatia" pageId="53" pageNumber="54">Croatia</collectingCountry>
not far from the geographic border of the Balkan Peninsula. Therefore, its occurrence on the Balkans is not excluded and we include it in the following key.
</paragraph>
</subSubSection>
<subSubSection id="32DCD3B5393DFFA5B1B00B49FA489AAF" lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54" type="key">
<key id="0E57D70F393DFFA5B1B00B49FA489AAF" lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54">
<keyStep id="C132989B393DFFA7B1B00B49FA4F9A59" pageId="53" pageNumber="54">
<paragraph id="7A79803E393DFFA7B1B00B49FA4F99A1" blockId="53.[151,1437,330,2023]" pageId="53" pageNumber="54">
<keyLead id="C137230B393DFFA7B1B00B49FA4F99A1" pageId="53" pageNumber="54">
1. Fastigium verticis wider, as wide as or slightly narrower than scapus (do not confuse with outside Balkan taxa
<emphasis id="48B25C2C393DFFA7B4020B49FE4C9908" italics="true" pageId="53" pageNumber="54">
<taxonomicName id="BDC6FBBD393DFFA7B4020B49FA50992A" box="[1317,1411,330,351]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="costata">I. costata</taxonomicName>
,
<taxonomicName id="BDC6FBBD393DFFA7B4A90B49FEEE9908" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="boldyrevi">I. boldyrevi</taxonomicName>
,
<taxonomicName id="BDC6FBBD393DFFA7B0610B6BFE4C9908" box="[326,415,360,381]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="taurica">I. taurica</taxonomicName>
</emphasis>
, which do not fit the following description);
<emphasis id="48B25C2C393DFFA7B26B0B64FC889909" bold="true" box="[844,859,359,380]" pageId="53" pageNumber="54">if</emphasis>
fastigium ±3/4 of scapus and song unknown then male cerci long and evenly roundly incurved
<emphasis id="48B25C2C393DFFA7B3080B86FD8699EF" bold="true" box="[559,597,389,410]" pageId="53" pageNumber="54">and</emphasis>
subgenital plate short with a deep triangle excision.
<emphasis id="48B25C2C393DFFA7B5700B86FB4299EF" box="[1111,1169,389,410]" italics="true" pageId="53" pageNumber="54">Males</emphasis>
: tegmina flat, much or distinctly longer than (sometimes in
<taxonomicName id="BDC6FBBD393DFFA7B3140BA1FD6799C2" box="[563,692,418,439]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rectipennis">
<emphasis id="48B25C2C393DFFA7B3140BA1FD6799C2" box="[563,692,418,439]" italics="true" pageId="53" pageNumber="54">I. rectipennis</emphasis>
</taxonomicName>
equal to) pronotum; tegmina not wider (usually narrower) than pronotum;
<emphasis id="48B25C2C393DFFA7B1C60BC3FEE199A0" box="[225,306,448,469]" italics="true" pageId="53" pageNumber="54">Females</emphasis>
: tegmina with clearly parallel venation.
<emphasis id="48B25C2C393DFFA7B3910BC3FD3599A0" box="[694,742,448,469]" italics="true" pageId="53" pageNumber="54">Song</emphasis>
consists of phrases of syllables (
<figureCitation id="E2FD9CBB393DFFA7B5070BBCFB7599A1" box="[1056,1190,447,468]" captionStart="FIGURES 135 138" captionStartId="20.[151,264,1762,1785]" captionTargetBox="[151,1436,445,1741]" captionTargetId="figure@20.[151,1436,445,1742]" captionTargetPageId="20" captionText="FIGURES 135 138. Male calling song at two (136 138) or three (135) different speeds (air temperature at recording and time scale below each figure): 135 — I. hospodar (RO: Gura Dobrogei); 136 — I. rectipennis (BG: Karandila Locality); 137 — I. pavelii (BG: Chernogorovo Locality); 138 — I. thracica (TR: Elmali Village)." httpUri="https://zenodo.org/record/246558/files/figure.png" pageId="53" pageNumber="54">Figs 135137</figureCitation>
).................. 2
</keyLead>
</paragraph>
<paragraph id="7A79803E393DFFA7B1B00BDEFA4F9A59" blockId="53.[151,1437,330,2023]" pageId="53" pageNumber="54">
<keyLead id="C137230B393DFFA7B1B00BDEFA4F9A59" pageId="53" pageNumber="54">
Fastigium verticis 1/4 to 3/4 as wide as scapus. Male and female tegmina with diverse shape and venation.
<emphasis id="48B25C2C393DFFA7B5DA0BDEFAFE9987" box="[1277,1325,477,498]" italics="true" pageId="53" pageNumber="54">Song</emphasis>
consists of isolated syllables, sparse groups of syllables or long sequences of syllables that are not arranged into phrases (if fastigium ±3/4 of scapus and song unknown compare
<emphasis id="48B25C2C393DFFA7B373081BFAB29A58" box="[596,1377,536,557]" italics="true" pageId="53" pageNumber="54">
<taxonomicName id="BDC6FBBD393DFFA7B373081BFD079A58" box="[596,724,536,557]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rectipennis">I. rectipennis</taxonomicName>
,
<taxonomicName id="BDC6FBBD393DFFA7B3F9081BFCA19A58" box="[734,882,536,557]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="longicaudata">I. longicaudata</taxonomicName>
,
<taxonomicName id="BDC6FBBD393DFFA7B25B081BFB809A58" box="[892,1107,536,557]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="subSpecies" species="rhodopensis" subSpecies="petkovi">I. rhodopensis petkovi</taxonomicName>
,
<taxonomicName id="BDC6FBBD393DFFA7B57B081BFB039A58" box="[1116,1232,536,557]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="modestior">I. modestior</taxonomicName>
,
<taxonomicName id="BDC6FBBD393DFFA7B5FE081BFAB29A58" box="[1241,1377,536,557]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="dobrogensis">I. dobrogensis</taxonomicName>
</emphasis>
)... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C132989B393DFFA7B1B00836FA4F9B1A" pageId="53" pageNumber="54">
<paragraph id="7A79803E393DFFA7B1B00836FA4F9AA8" blockId="53.[151,1437,330,2023]" pageId="53" pageNumber="54">
<keyLead id="C137230B393DFFA7B1B00836FA4F9AA8" pageId="53" pageNumber="54">
2. Body short, massive, pronotum almost quadrate, as broad as or broader than long, no thin lateral (red or brownish) stripes on pronotum developed.
<emphasis id="48B25C2C393DFFA7B0950851FE3E9A12" box="[434,493,594,615]" italics="true" pageId="53" pageNumber="54">Males</emphasis>
: tegmina slightly to distinctly (1.5 times) longer than pronotum with the medial (Medio-Anal) part (or disc) as large as the lateral (Costo-Medial) part; CuP weak and short; stridulatory file apically reduced (
<figureCitation id="E2FD9CBB393DFFA7B428086CFAB49AF1" box="[1295,1383,623,644]" captionStart="FIGURES 130 134" captionStartId="19.[151,264,1657,1680]" captionTargetBox="[151,1435,193,1634]" captionTargetId="figure@19.[151,1436,193,1636]" captionTargetPageId="19" captionText="FIGURES 130 134. SEM of taxonomically important structures of Isophya: 130, 131 — I. hospodar (130 — BG: Dolni Glavanak; 131 — BG: Sofia Kettle); 132 — I. rectipennis (132 A, C, BG: Karandila Locality; 132 B, TR: “ TURKEY: Bolu. | Ala Dagi. 2000 m. | Kartal Kaya Tepe. | 15. VII. 1962 | Guichard &amp; Harvey. | B. M. 1962 299. ”); 133 — I. pavelii (BG: Chernogorovo Locality); 134 — I. thracica (TR: Elmali Village). A—male stridulatory file (scale 200 μm); B—apical part of male stridulatory file (scale 20 μm); C—apex of male cerci (scale 100 μm) (130, 132 134 — ventro-apical view; 131 — dorsal view); D—female stridulatory apparatus (scale 100 μm)." httpUri="https://zenodo.org/record/246557/files/figure.png" pageId="53" pageNumber="54">Figs 130</figureCitation>
A, B, 131A, B); cerci (
<figureCitation id="E2FD9CBB393DFFA7B0A2088EFE1F9AD7" box="[389,460,653,674]" captionStart="FIGURES 80 104" captionStartId="14.[151,264,1835,1858]" captionTargetBox="[226,1360,681,1815]" captionTargetId="figure@14.[226,1362,681,1816]" captionTargetPageId="14" captionText="FIGURES 80 104. Male abdominal terminalia of the Balkan Isophya and I. yaraligozi (localities as in Figs 31 55): 80 — I. hospodar; 81 — I. rectipennis; 82 — I. pavelii; 83 — I. thracica; 84 — I. bureschi; 85 — I. yaraligozi; 86 — I. tosevski; 87 — I. andreevae; 88 — I. clara; 89 — I. miksici; 90 — I. plevnensis; 91 — I. longicaudata adamovici; 92 — I. l. longicaudata; 93 — I. m. modesta (= intermedia); 94 — I. rhodopensis leonorae; 95 — I. rh. rhodopensis; 96 — I. rh. petkovi; 97 — I. modestior; 98 — I. dobrogensis; 99 — I. zubowskii; 100 — I. aff. camptoxypha; 101 — I. gulae; 102 — I. obtusa; 103 — I. amplipennis; 104 — I. speciosa. Scale (if present) = 2 mm." httpUri="https://zenodo.org/record/246555/files/figure.png" pageId="53" pageNumber="54">Fig. 80</figureCitation>
) short, stout, almost straight over the basal 2/3.
<emphasis id="48B25C2C393DFFA7B286088EFC219AD7" box="[929,1010,653,674]" italics="true" pageId="53" pageNumber="54">Females</emphasis>
: tegmina apically blunt; female stridulatory apparatus with 34 closely situated rows of basal teeth (
<figureCitation id="E2FD9CBB393DFFA7B3D008A9FC999ACA" box="[759,842,682,703]" captionStart="FIGURES 130 134" captionStartId="19.[151,264,1657,1680]" captionTargetBox="[151,1435,193,1634]" captionTargetId="figure@19.[151,1436,193,1636]" captionTargetPageId="19" captionText="FIGURES 130 134. SEM of taxonomically important structures of Isophya: 130, 131 — I. hospodar (130 — BG: Dolni Glavanak; 131 — BG: Sofia Kettle); 132 — I. rectipennis (132 A, C, BG: Karandila Locality; 132 B, TR: “ TURKEY: Bolu. | Ala Dagi. 2000 m. | Kartal Kaya Tepe. | 15. VII. 1962 | Guichard &amp; Harvey. | B. M. 1962 299. ”); 133 — I. pavelii (BG: Chernogorovo Locality); 134 — I. thracica (TR: Elmali Village). A—male stridulatory file (scale 200 μm); B—apical part of male stridulatory file (scale 20 μm); C—apex of male cerci (scale 100 μm) (130, 132 134 — ventro-apical view; 131 — dorsal view); D—female stridulatory apparatus (scale 100 μm)." httpUri="https://zenodo.org/record/246557/files/figure.png" pageId="53" pageNumber="54">Fig. 130</figureCitation>
D).
<emphasis id="48B25C2C393DFFA7B24A08A9FC4E9ACA" box="[877,925,682,703]" italics="true" pageId="53" pageNumber="54">Song</emphasis>
(
<figureCitation id="E2FD9CBB393DFFA7B28D08A9FC2F9ACA" box="[938,1020,682,703]" captionStart="FIGURES 135 138" captionStartId="20.[151,264,1762,1785]" captionTargetBox="[151,1436,445,1741]" captionTargetId="figure@20.[151,1436,445,1742]" captionTargetPageId="20" captionText="FIGURES 135 138. Male calling song at two (136 138) or three (135) different speeds (air temperature at recording and time scale below each figure): 135 — I. hospodar (RO: Gura Dobrogei); 136 — I. rectipennis (BG: Karandila Locality); 137 — I. pavelii (BG: Chernogorovo Locality); 138 — I. thracica (TR: Elmali Village)." httpUri="https://zenodo.org/record/246558/files/figure.png" pageId="53" pageNumber="54">Fig. 135</figureCitation>
) of grouped phrases of very short (&lt;50 ms) syllables consisting of well detached impulses.......................................................
<taxonomicName id="BDC6FBBD393DFFA7B40D08CBFA4F9AA8" box="[1322,1436,712,733]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="hospodar">
<emphasis id="48B25C2C393DFFA7B40D08CBFA4F9AA8" box="[1322,1436,712,733]" italics="true" pageId="53" pageNumber="54">I. hospodar</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7A79803E393DFFA7B1B008E6FA4F9B1A" blockId="53.[151,1437,330,2023]" pageId="53" pageNumber="54">
<keyLead id="C137230B393DFFA7B1B008E6FA4F9B1A" pageId="53" pageNumber="54">
Body not apparently massive, pronotum elongated, usually longer than broad, and has two reddish lateral stripes over the light bands in the metazone.
<emphasis id="48B25C2C393DFFA7B0EE0901FDD09B62" box="[457,515,770,791]" italics="true" pageId="53" pageNumber="54">Males</emphasis>
: tegmina about as long as pronotum, the Medio-Anal part (disc) larger than the Costo-Medial part; stridulatory file normally developed; cerci long and gradually tapering towards apex.
<emphasis id="48B25C2C393DFFA7B5450923FB609B40" box="[1122,1203,800,821]" italics="true" pageId="53" pageNumber="54">Females</emphasis>
: tegmina apically oval; stridulatory apparatus with two separated rows of basal teeth.
<emphasis id="48B25C2C393DFFA7B209093EFC8D9B27" box="[814,862,829,850]" italics="true" pageId="53" pageNumber="54">Song</emphasis>
of grouped or isolate phrases of longer (usually 50120 ms) syllables of dense impulses.............................................................................. 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C132989B393DFFA7B1B00974FA489C2F" pageId="53" pageNumber="54">
<paragraph id="7A79803E393DFFA7B1B00974FA4F9B90" blockId="53.[151,1437,330,2023]" pageId="53" pageNumber="54">
<keyLead id="C137230B393DFFA7B1B00974FA4F9B90" pageId="53" pageNumber="54">
3. Slender species, fastigium moderately protuberant with a deep groove, as wide as or slightly narrower than scape.
<emphasis id="48B25C2C393DFFA7B40B097BFAB59BF8" box="[1324,1382,888,909]" italics="true" pageId="53" pageNumber="54">Males</emphasis>
: pronotum slightly widened in metazone; CuP is short (about 1/2 of the width of metazone), weak; stridulatory file with about 50 teeth.
<emphasis id="48B25C2C393DFFA7B03B09B1FEBE9BB2" box="[284,365,946,967]" italics="true" pageId="53" pageNumber="54">Females</emphasis>
: pronotum with almost parallel sides; lateral pit between gonangulum and lamella strongly closed by lamella; ovipositor &lt;
<quantity id="BD3E2DDB393DFFA7B07109CCFE439B91" box="[342,400,975,996]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="53" pageNumber="54" unit="mm" value="9.0">9 mm</quantity>
.
<emphasis id="48B25C2C393DFFA7B0BE09D3FE1A9B90" box="[409,457,976,997]" italics="true" pageId="53" pageNumber="54">Song</emphasis>
of phrases of 2040 syllables that last 1350 ms...............................
<taxonomicName id="BDC6FBBD393DFFA7B43909D3FA4F9B90" box="[1310,1436,976,997]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rectipennis">
<emphasis id="48B25C2C393DFFA7B43909D3FA4F9B90" box="[1310,1436,976,997]" italics="true" pageId="53" pageNumber="54">I. rectipennis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7A79803E393DFFA7B1B009EEFA489C2F" blockId="53.[151,1437,330,2023]" pageId="53" pageNumber="54">
<keyLead id="C137230B393DFFA7B1B009EEFA489C2F" pageId="53" pageNumber="54">
Larger and stouter species, fastigium flattened, as wide as or wider than scape.
<emphasis id="48B25C2C393DFFA7B2C909EEFBFB9C77" box="[1006,1064,1005,1026]" italics="true" pageId="53" pageNumber="54">Males</emphasis>
: pronotum strongly widened in metazone; CuP is long (2/33/4 of the width of metazone), wide; stridulatory file with about 100 teeth.
<emphasis id="48B25C2C393DFFA7B5940E09FAD79C6A" box="[1203,1284,1034,1055]" italics="true" pageId="53" pageNumber="54">Females</emphasis>
: pronotum distinctly widened in metazone; lateral pit between gonangulum and lamella partly closed by lamella; ovipositor&gt;
<quantity id="BD3E2DDB393DFFA7B4280E24FA9A9C49" box="[1295,1353,1063,1084]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="53" pageNumber="54" unit="mm" value="9.0">9 mm</quantity>
.
<emphasis id="48B25C2C393DFFA7B4750E2BFA519C48" box="[1362,1410,1064,1085]" italics="true" pageId="53" pageNumber="54">Song</emphasis>
of phrases of 513 syllables that last (excl. the first one) over 50 ms..........................................
<taxonomicName id="BDC6FBBD393DFFA7B4610E46FA489C2F" box="[1350,1435,1093,1114]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pavelii">
<emphasis id="48B25C2C393DFFA7B4610E46FA489C2F" box="[1350,1435,1093,1114]" italics="true" pageId="53" pageNumber="54">I. pavelii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C132989B393DFFA7B1B00E61FA4F9DCF" pageId="53" pageNumber="54">
<paragraph id="7A79803E393DFFA7B1B00E61FA4F9C98" blockId="53.[151,1437,330,2023]" pageId="53" pageNumber="54">
<keyLead id="C137230B393DFFA7B1B00E61FA4F9C98" pageId="53" pageNumber="54">
4.
<emphasis id="48B25C2C393DFFA7B1C60E61FECF9C02" box="[225,284,1122,1143]" italics="true" pageId="53" pageNumber="54">Males</emphasis>
: tegmina distinctly elongated, 1.21.5 times longer than pronotum but narrower than the width of metazone; Medio- Anal part (disc)
<emphasis id="48B25C2C393DFFA7B0A60E7CFE719CE1" bold="true" box="[385,418,1151,1172]" pageId="53" pageNumber="54">not</emphasis>
larger than the Costo-Medial part; CuP is short—1/22/3 of the width of metazone.
<emphasis id="48B25C2C393DFFA7B5FC0E83FAFF9CE0" box="[1243,1324,1152,1173]" italics="true" pageId="53" pageNumber="54">Females</emphasis>
: lamella of ovipositor short (slightly protruding laterally) with an excision in its distal part (
<figureCitation id="E2FD9CBB393DFFA7B2C20E9EFBEA9CC7" box="[997,1081,1181,1202]" captionStart="FIGURES 105 129" captionStartId="17.[151,264,1173,1196]" captionTargetBox="[151,1436,195,1152]" captionTargetId="figure@17.[151,1436,195,1152]" captionTargetPageId="17" captionText="FIGURES 105 129. Female abdominal terminalia of the Balkan Isophya and I. yaraligozi (localities as in Figs 31 55 unless specifically given); figures of the whole ovipositor are followed by enlarged figure of the ovipositor base: 105 — I. hospodar (BG: Dolni Glavanak, 25.04. 2005, CC); 106 — I. rectipennis (BG: Byala Vill., 27.06. 2002, CC); 107 — I. pavelii (BG: Chernogorovo, 29.05. 2006, CC); 108 — I. thracica; 109 — I. bureschi (BG: Bansko, 8.08. 2006, CC); 110 — I. yaraligozi; 111 — I. tosevski; 112 — I. andreevae (BG: Kresnensko Hanche, 19.04. 2006, CC); 113 — I. clara; 114 — I. miksici (BG: Gorski Dom Lodge, 24.06. 2006, CC); 115 — I. plevnensis (BG: Levishte, 14.07. 2009, CC); 116 — I. longicaudata adamovici (BG: Karandila, 24.06. 2008, CC); 117 — I. l. longicaudata (BG: Sofia University Botanical garden, 26.06. 1.07.2004, CC); 118 — I. m. modesta (= intermedia); 119 — I. rhodopensis leonorae (BG: Livade, 9.08. 2006, CC); 120 — I. rh. rhodopensis (BG: Bachkovo, 600 m, 23.05. 2004, CC); 121 — I. rh. petkovi (BG: Gluhite Kamani, 23.06. 2008, CC); 122 — I. modestior (BG: Bankya, 24.05. 2006, CC); 123 — I. dobrogensis; 124 — I. zubowskii; 125 — I. aff. camptoxypha (data as in Fig. 75); 126 — I. gulae; 127 — I. obtusa (BG: Ravnets Ridge, 14.07. 2009, CC); 128 — I. amplipennis (as in Fig. 78); 129 — I. speciosa (BG: Bankya, 24.05. 2006, CC). Indications on Fig. 127 show, respectively: G—gonangulum; L—lamella; P—lateral pit between lamella and gonangulum. Scale (if present) = 5 mm." httpUri="https://zenodo.org/record/246556/files/figure.png" pageId="53" pageNumber="54">Fig. 108</figureCitation>
); ovipositor is stout,
<quantity id="BD3E2DDB393DFFA7B4230E9EFAB69CC7" box="[1284,1381,1181,1202]" metricMagnitude="-3" metricUnit="m" metricValue="8.25" metricValueMax="9.0" metricValueMin="7.5" pageId="53" pageNumber="54" unit="mm" value="8.25" valueMax="9.0" valueMin="7.5">7.59 mm</quantity>
long, slightly upcurved (its lower edge almost sraight in the middle); pronotal disc not constricted in the middle; female tegmina shorter than half the pronotum lenght.
<emphasis id="48B25C2C393DFFA7B3750EDBFD519C98" box="[594,642,1240,1261]" italics="true" pageId="53" pageNumber="54">Song</emphasis>
of isolated syllables lasting &lt;50 ms and followed by 12 after-clicks...
<taxonomicName id="BDC6FBBD393DFFA7B4120EDBFA4F9C98" box="[1333,1436,1240,1261]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="thracica">
<emphasis id="48B25C2C393DFFA7B4120EDBFA4F9C98" box="[1333,1436,1240,1261]" italics="true" pageId="53" pageNumber="54">I. thracica</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7A79803E393DFFA7B1B00EF6FA4F9DCF" blockId="53.[151,1437,330,2023]" pageId="53" pageNumber="54">
<keyLead id="C137230B393DFFA7B1B00EF6FA4F9DCF" pageId="53" pageNumber="54">
<emphasis id="48B25C2C393DFFA7B1C60EF6FECF9D7F" box="[225,284,1269,1290]" italics="true" pageId="53" pageNumber="54">Males</emphasis>
: tegmina
<emphasis id="48B25C2C393DFFA7B0580EF6FE739D7F" bold="true" box="[383,416,1269,1290]" pageId="53" pageNumber="54">not</emphasis>
distinctly elongated, less than 1.2 times longer than pronotum;
<emphasis id="48B25C2C393DFFA7B5300EF6FBF59D7F" bold="true" box="[1047,1062,1269,1290]" pageId="53" pageNumber="54">if</emphasis>
longer then the tegmen is not significantly narrower than the width of metazone; Medio-Anal part (disc) always larger than the Costo-Medial part; CuP usually&gt;2/ 3 of the width of metazone.
<emphasis id="48B25C2C393DFFA7B0DF0F33FD9A9D30" box="[504,585,1328,1349]" italics="true" pageId="53" pageNumber="54">Females</emphasis>
: lamella of ovipositor without an excision distally;
<emphasis id="48B25C2C393DFFA7B5620F2CFB879D31" bold="true" box="[1093,1108,1327,1348]" pageId="53" pageNumber="54">if</emphasis>
with excision and ovipositor less than
<quantity id="BD3E2DDB393DFFA7B0370F4EFE999D17" box="[272,330,1357,1378]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="53" pageNumber="54" unit="mm" value="9.0">9 mm</quantity>
, then the ovipositor is strongly upcurved
<emphasis id="48B25C2C393DFFA7B3F20F4EFD2F9D17" bold="true" box="[725,764,1357,1378]" pageId="53" pageNumber="54">and</emphasis>
distinctly constricted in the middle (
<figureCitation id="E2FD9CBB393DFFA7B57C0F4EFB319D17" box="[1115,1250,1357,1378]" captionStart="FIGURES 80 104" captionStartId="14.[151,264,1835,1858]" captionTargetBox="[226,1360,681,1815]" captionTargetId="figure@14.[226,1362,681,1816]" captionTargetPageId="14" captionText="FIGURES 80 104. Male abdominal terminalia of the Balkan Isophya and I. yaraligozi (localities as in Figs 31 55): 80 — I. hospodar; 81 — I. rectipennis; 82 — I. pavelii; 83 — I. thracica; 84 — I. bureschi; 85 — I. yaraligozi; 86 — I. tosevski; 87 — I. andreevae; 88 — I. clara; 89 — I. miksici; 90 — I. plevnensis; 91 — I. longicaudata adamovici; 92 — I. l. longicaudata; 93 — I. m. modesta (= intermedia); 94 — I. rhodopensis leonorae; 95 — I. rh. rhodopensis; 96 — I. rh. petkovi; 97 — I. modestior; 98 — I. dobrogensis; 99 — I. zubowskii; 100 — I. aff. camptoxypha; 101 — I. gulae; 102 — I. obtusa; 103 — I. amplipennis; 104 — I. speciosa. Scale (if present) = 2 mm." httpUri="https://zenodo.org/record/246555/files/figure.png" pageId="53" pageNumber="54">Figs 103, 104</figureCitation>
),
<emphasis id="48B25C2C393DFFA7B5D40F4EFAC99D17" bold="true" box="[1267,1306,1357,1378]" pageId="53" pageNumber="54">and</emphasis>
pronotal disc is constricted in the middle,
<emphasis id="48B25C2C393DFFA7B0C80F69FDC59D0A" bold="true" box="[495,534,1386,1407]" pageId="53" pageNumber="54">and</emphasis>
tegmina longer than half the pronotum lenght.
<emphasis id="48B25C2C393DFFA7B2FE0F69FBDA9D0A" box="[985,1033,1386,1407]" italics="true" pageId="53" pageNumber="54">Song</emphasis>
of syllables lasting&gt;100 ms or
<emphasis id="48B25C2C393DFFA7B41D0F69FA9A9D0A" bold="true" box="[1338,1353,1386,1407]" pageId="53" pageNumber="54">if</emphasis>
&lt;50 ms, then the syllable is followed by well defined group of after-clicks (
<taxonomicName id="BDC6FBBD393DFFA7B2430F8BFC189DE8" box="[868,971,1416,1437]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="bureschi">
<emphasis id="48B25C2C393DFFA7B2430F8BFC189DE8" box="[868,971,1416,1437]" italics="true" pageId="53" pageNumber="54">I. bureschi</emphasis>
</taxonomicName>
)
<emphasis id="48B25C2C393DFFA7B2FE0F84FC3C9DE9" bold="true" box="[985,1007,1415,1436]" pageId="53" pageNumber="54">or</emphasis>
the syllables are arranged in long sequences (
<taxonomicName id="BDC6FBBD393DFFA7B1CE0FA6FEAE9DCF" box="[233,381,1445,1466]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="camptoxypha">
<emphasis id="48B25C2C393DFFA7B1CE0FA6FEAE9DCF" box="[233,381,1445,1466]" italics="true" pageId="53" pageNumber="54">I. camptoxypha</emphasis>
</taxonomicName>
)...................................................................................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C132989B393DFFA7B1B00FC1FA489F07" pageId="53" pageNumber="54">
<paragraph id="7A79803E393DFFA7B1B00FC1FA4F9ED1" blockId="53.[151,1437,330,2023]" pageId="53" pageNumber="54">
<keyLead id="C137230B393DFFA7B1B00FC1FA4F9ED1" pageId="53" pageNumber="54">
5. Body colouration may be variable but never with black spots (melanism not developed).
<emphasis id="48B25C2C393DFFA7B5670FC1FBA99DA2" box="[1088,1146,1474,1495]" italics="true" pageId="53" pageNumber="54">Males</emphasis>
: pronotum not distinctly constricted in the middle (even if metazone is much wider than prozone);
<emphasis id="48B25C2C393DFFA7B2A50FDCFC429D81" bold="true" box="[898,913,1503,1524]" pageId="53" pageNumber="54">if</emphasis>
constricted (e.g.
<taxonomicName id="BDC6FBBD393DFFA7B5110FE3FB599D80" box="[1078,1162,1504,1525]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="obtusa">
<emphasis id="48B25C2C393DFFA7B5110FE3FB599D80" box="[1078,1162,1504,1525]" italics="true" pageId="53" pageNumber="54">I. obtusa</emphasis>
</taxonomicName>
,
<taxonomicName id="BDC6FBBD393DFFA7B5B10FE3FAF99D80" box="[1174,1322,1504,1525]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="camptoxypha">
<emphasis id="48B25C2C393DFFA7B5B10FE3FAF99D80" box="[1174,1322,1504,1525]" italics="true" pageId="53" pageNumber="54">I. camptoxypha</emphasis>
</taxonomicName>
), then male cerci ±stout, usually with blunt apex and female ovipositor longer than
<quantity id="BD3E2DDB393DFFA7B2B60FFEFC079E67" box="[913,980,1533,1554]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="53" pageNumber="54" unit="mm" value="10.0">10 mm</quantity>
<emphasis id="48B25C2C393DFFA7B2FC0FFEFC219E67" bold="true" box="[987,1010,1533,1554]" pageId="53" pageNumber="54">or</emphasis>
tegmina are ±shorter than pronotum
<emphasis id="48B25C2C393DFFA7B47F0FFEFA4F9E67" bold="true" box="[1368,1436,1533,1554]" pageId="53" pageNumber="54">and/or</emphasis>
CuA and CuP are closely attached;
<emphasis id="48B25C2C393DFFA7B3640C19FD479E5A" box="[579,660,1562,1583]" italics="true" pageId="53" pageNumber="54">Females</emphasis>
: pronotum not constricted in the middle; tegmina with more or less expressed reticulate venation; lateral pit between gonangulum and lamella widely opened, lamella without distinct excision; ovipositor longer than
<quantity id="BD3E2DDB393DFFA7B0710C56FE419E1F" box="[342,402,1621,1642]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="53" pageNumber="54" unit="mm" value="9.0">9 mm</quantity>
;
<emphasis id="48B25C2C393DFFA7B0BA0C56FE7F9E1F" bold="true" box="[413,428,1621,1642]" pageId="53" pageNumber="54">if</emphasis>
shorter (
<emphasis id="48B25C2C393DFFA7B3220C56FCFB9E1F" box="[517,808,1621,1642]" italics="true" pageId="53" pageNumber="54">
<taxonomicName id="BDC6FBBD393DFFA7B3220C56FD5B9E1F" box="[517,648,1621,1642]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="brevicauda">I. brevicauda</taxonomicName>
,
<taxonomicName id="BDC6FBBD393DFFA7B3B50C56FCFB9E1F" box="[658,808,1621,1642]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="camptoxypha">I. camptoxypha</taxonomicName>
</emphasis>
), then see lateral pit of gonangulum.
<emphasis id="48B25C2C393DFFA7B5BF0C56FB1B9E1F" box="[1176,1224,1621,1642]" italics="true" pageId="53" pageNumber="54">Song</emphasis>
with diverse
<typeStatus id="A57D3E9C393DFFA7B4690C56FA529E1F" box="[1358,1409,1621,1642]" pageId="53" pageNumber="54">types</typeStatus>
of syllables lasting &lt;1 s;
<emphasis id="48B25C2C393DFFA7B09D0C71FE1B9EF2" bold="true" box="[442,456,1650,1671]" pageId="53" pageNumber="54">if</emphasis>
of single syllables lasting&gt;12 s, then the syllable has two parts is produced by single closing wing movement (but cf.
<taxonomicName id="BDC6FBBD393DFFA7B0B00C93FDCC9ED0" box="[407,543,1680,1701]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="dobrogensis">
<emphasis id="48B25C2C393DFFA7B0B00C93FDCC9ED0" box="[407,543,1680,1701]" italics="true" pageId="53" pageNumber="54">I. dobrogensis</emphasis>
</taxonomicName>
)........................................................................ 6
</keyLead>
</paragraph>
<paragraph id="7A79803E393DFFA7B1B00CAEFA489F07" blockId="53.[151,1437,330,2023]" pageId="53" pageNumber="54">
<keyLead id="C137230B393DFFA7B1B00CAEFA489F07" pageId="53" pageNumber="54">
Body colouration frequently (but not always!) variegated with black spots at least on pronotum or apices of male cerci (melanism developed).
<emphasis id="48B25C2C393DFFA7B0B70CC9FE199EAA" box="[400,458,1738,1759]" italics="true" pageId="53" pageNumber="54">Males</emphasis>
: pronotum distinctly constricted in the middle (saddle shaped); tegmina bulged, usually longer and wider than pronotum; CuA and CuP separated; cerci long, slim, fery thin or, alternatively, widened at apex.
<emphasis id="48B25C2C393DFFA7B5C60CEBFAE19E88" box="[1249,1330,1768,1789]" italics="true" pageId="53" pageNumber="54">Females</emphasis>
: pronotum visibly constricted in the middle; tegmina with well expressed reticulate venation; lateral pit between gonangulum and lamella partly closed by lamella, the latter with excision distally; ovipositor shorter than
<quantity id="BD3E2DDB393DFFA7B2D30D21FBFC9F42" box="[1012,1071,1826,1847]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="53" pageNumber="54" unit="mm" value="9.0">9 mm</quantity>
, strongly upcurved and constricted in the middle.
<emphasis id="48B25C2C393DFFA7B0710D43FE559F20" box="[342,390,1856,1877]" italics="true" pageId="53" pageNumber="54">Song</emphasis>
of single syllables lasting&gt;2 s and consisting of at least three parts produced by two opening-and-closing wing movements..................................................................................... 21
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C132989B393DFFA4B1B00D79FA4F9957" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54">
<paragraph id="7A79803E393DFFA7B1B00D79FA4F9F92" blockId="53.[151,1437,330,2023]" pageId="53" pageNumber="54">
<keyLead id="C137230B393DFFA7B1B00D79FA4F9F92" pageId="53" pageNumber="54">
6.
<emphasis id="48B25C2C393DFFA7B1C60D79FECF9FFA" box="[225,284,1914,1935]" italics="true" pageId="53" pageNumber="54">Males</emphasis>
: tegmina equal or longer than pronotum (if slightly shorter see below); CuA and CuP well separated or moderately attached; stridulatory file usually with &lt;180 teeth; if CuA and CuP distinctly attached and file with ±180 teeth, then very large species (str.file&gt;
<quantity id="BD3E2DDB393DFFA7B0A30DB6FE029FBF" box="[388,465,1973,1994]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="53" pageNumber="54" unit="mm" value="3.8">3.8 mm</quantity>
, hind femur&gt;
<quantity id="BD3E2DDB393DFFA7B3700DB6FD4E9FBF" box="[599,669,1973,1994]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="53" pageNumber="54" unit="mm" value="22.0">22 mm</quantity>
) and/or the cerci bear large long tooth (
<figureCitation id="E2FD9CBB393DFFA7B53E0DB6FB729FBF" box="[1049,1185,1973,1994]" captionStart="FIGURES 162 167" captionStartId="30.[151,264,1332,1355]" captionTargetBox="[151,1435,218,1310]" captionTargetId="figure@30.[151,1436,218,1311]" captionTargetPageId="30" captionText="FIGURES 162 167. SEM of taxonomically important structures of Isophya: 162, 163 — I. rhodopensis leonorae (162 — BG: N Pirin Mts; 163 — BG: Alibotoush Mt.); 164 166 — I. rhodopensis rhodopensis (164 — typical form, BG: above Smolyan; 165 — northwestern form, BG: Batak Lake; 166 — typical form or transitional to I. rh. petkovi, BG: near Assenovgrad); 167 — I. rhodopensis petkovi (BG: Glouhite Kamani Locality). A—male stridulatory file (162, 166 — scale 200 μm; the rest—scale 1 mm); B—ventral / ventro-apical view of apex of male cerci (scale 100 μm); C—female stridulatory apparatus (162 — scale 100 μm; 166 — scale 200 μm; the rest—scale 500 μm)." httpUri="https://zenodo.org/record/246563/files/figure.png" pageId="53" pageNumber="54">Figs 162167</figureCitation>
B).
<emphasis id="48B25C2C393DFFA7B5E60DB6FAC19FBF" box="[1217,1298,1973,1994]" italics="true" pageId="53" pageNumber="54">Females</emphasis>
: tegmina usually with oval or slightly blunt apex;
<emphasis id="48B25C2C393DFFA7B36E0DD1FD849F92" bold="true" box="[585,599,2002,2023]" pageId="53" pageNumber="54">if</emphasis>
apex considerably blunt, then very large species (hind femur&gt;
<quantity id="BD3E2DDB393DFFA7B59B0DD1FAD79F92" box="[1212,1284,2002,2023]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="53" pageNumber="54" unit="mm" value="22.0">22 mm</quantity>
),
<emphasis id="48B25C2C393DFFA7B4320DD1FA8A9F92" bold="true" box="[1301,1369,2002,2023]" pageId="53" pageNumber="54">and/or</emphasis>
lateral
</keyLead>
</paragraph>
<paragraph id="7A79803E393EFFA4B1C60A94FA4F9957" blockId="54.[151,1436,151,1992]" pageId="54" pageNumber="55">
stripe in metazone brownish or black (some
<taxonomicName id="BDC6FBBD393EFFA4B3AD0A9BFCC098D8" box="[650,787,152,173]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="rhodopensis">
<emphasis id="48B25C2C393EFFA4B3AD0A9BFCC098D8" box="[650,787,152,173]" italics="true" pageId="54" pageNumber="55">I. rhodopensis</emphasis>
</taxonomicName>
),
<emphasis id="48B25C2C393EFFA4B2000A94FCED98D9" bold="true" box="[807,830,151,172]" pageId="54" pageNumber="55">or</emphasis>
tibiae reddish
<emphasis id="48B25C2C393EFFA4B2EB0A94FC3098D9" bold="true" box="[972,995,151,172]" pageId="54" pageNumber="55">or</emphasis>
tegmina darker than body (
<emphasis id="48B25C2C393EFFA4B5CB0A9BFEFE98BF" italics="true" pageId="54" pageNumber="55">
<taxonomicName id="BDC6FBBD393EFFA4B5CB0A9BFA9198D8" box="[1260,1346,152,173]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="obtusa">I. obtusa</taxonomicName>
,
<taxonomicName id="BDC6FBBD393EFFA4B46C0A9BFEFE98BF" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="camptoxypha">I. camptoxypha</taxonomicName>
</emphasis>
)
<emphasis id="48B25C2C393EFFA4B01C0AB6FE8198BF" bold="true" box="[315,338,181,202]" pageId="54" pageNumber="55">or</emphasis>
refer to a conspecific male (e.g. some
<emphasis id="48B25C2C393EFFA4B3E30AB6FC7E98BF" box="[708,941,181,202]" italics="true" pageId="54" pageNumber="55">
<taxonomicName id="BDC6FBBD393EFFA4B3E30AB6FCC398BF" box="[708,784,181,202]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="gulae">I. gulae</taxonomicName>
,
<taxonomicName id="BDC6FBBD393EFFA4B23D0AB6FC7E98BF" box="[794,941,181,202]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="camptoxypha">I. camptoxypha</taxonomicName>
</emphasis>
)....................................... 7
<keyLead id="C137230B393EFFA4B1B00AD1FA4F9957" pageId="54" pageNumber="55">
<emphasis id="48B25C2C393EFFA4B1C60AD1FECF9892" box="[225,284,210,231]" italics="true" pageId="54" pageNumber="55">Males</emphasis>
: tegmina shorter than pronotum (if other see below); CuA and CuP closely attached (
<figureCitation id="E2FD9CBB393EFFA4B54E0AD1FB089892" box="[1129,1243,210,231]" captionStart="FIGURES 31 55" captionStartId="12.[151,264,244,267]" captionText="FIGURES 31 55. Morphology of male head, pronotum and tegmina of the Balkan Isophya and I. yaraligozi: 31 — I. hospodar (BG: Momchilgrad, 31.05. 1965, NMNHS); 32 — I. rectipennis (BG: Ropotamo, 7.07. 1965, NMNHS); 33 — I. pavelii (holotype of I. rammei Peshev, BG: Chernogorovo, 19.06. 1973, NMNHS); 34 — I. thracica (TR: Elmali, 3.07. 2011, CC); 35 — I. bureschi (BG: “ Pirin, 1300 1450 m ”, 17.08. 1961, NMNHS); 36 — I. yaraligozi (TR: Yaraligoz Pass, 9.07. 2011, CC); 37 — I. tosevski (MK: Moklishte, 30.05. 2010, CC); 38 — I. andreevae (holotype, BG: Kresna Gorge, 2.06. 1961, NMNHS); 39 — I. clara (BA: Sarajevo, NMW); 40 — I. miksici (holotype, BG: Berkovitsa, 700 m, 21.06. 1958, NMNHS); 41 — I. plevnensis (holotype, BG: Chernelka, 30.07. 1973, NMNHS); 42 — I. longicaudata adamovici (holotype, BG: “ Sliven ”, NMNHS); 43 — I. l. longicaudata (BG: Batova Gora, 26.06. 1956, NMNHS); 44 — I. m. modesta (= intermedia) (RO: Dobrogea, Babadag, CI); 45 — I. rhodopensis leonorae (BG: Popovi Livadi, 1300 m, 4.07. 1961, NMNHS); 46 — I. rh. rhodopensis (BG: Chepelare, 16.07. 1956, NMNHS); 47 — I. rh. petkovi (BG: Mandritsa, 4.06. 1965, NMNHS); 48 — I. modestior (BG: Kopitoto, 17.07. 1958, NMNHS); 49 — I. dobrogensis (RO: Popina Island, CI); 50 — I. zubowskii (RO: “ Jasi pād, Birnari, 25.06. 1956, NMNHS); 51 — I. aff. camptoxypha (“ Montenegro ”, NMW); 52 — I. gulae (BG: Dolna Topchiya, 23.06. 2008, NMNHS); 53 — I. obtusa (BG: Vezhen Peak, 1900 m, 1.09. 1960, NMNHS); 54 — I. amplipennis (TR: “ Bursa | Gursu, | Kazanci &amp; | Eskibaglar | 14. vi. 1959 ”, “ by Dr. E. Can. | B. M. 1960 173 ”, NHM); 55 — I. speciosa (BG: Plevoun, 5.06. 1965, NMNHS)." pageId="54" pageNumber="55">Figs 4850</figureCitation>
), sometimes almost touching each other, CuA frequently disposed higher than CuP; stridulatory file with&gt;180 teeth (only in
<taxonomicName id="BDC6FBBD393EFFA4B5EF0AF3FAEF9970" box="[1224,1340,240,261]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="modestior">
<emphasis id="48B25C2C393EFFA4B5EF0AF3FAEF9970" box="[1224,1340,240,261]" italics="true" pageId="54" pageNumber="55">I. modestior</emphasis>
</taxonomicName>
out of the Balkans less).
<emphasis id="48B25C2C393EFFA4B04B0B0EFE6E9957" box="[364,445,269,290]" italics="true" pageId="54" pageNumber="55">Females</emphasis>
: tegmina with distinctly blunt or even concave apex...................................... 18
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C132989B393EFFA4B1B00B29FA4F9AF7" pageId="54" pageNumber="55">
<paragraph id="7A79803E393EFFA4B1B00B29FA4F999A" blockId="54.[151,1436,151,1992]" pageId="54" pageNumber="55">
<keyLead id="C137230B393EFFA4B1B00B29FA4F999A" pageId="54" pageNumber="55">
7.
<emphasis id="48B25C2C393EFFA4B1C60B29FECF994A" box="[225,284,298,319]" italics="true" pageId="54" pageNumber="55">Males</emphasis>
: CuP lighter than the disc of tegmen, much wider than CuA; if CuP thin and green (
<taxonomicName id="BDC6FBBD393EFFA4B57E0B29FB13994A" box="[1113,1216,298,319]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="bureschi">
<emphasis id="48B25C2C393EFFA4B57E0B29FB13994A" box="[1113,1216,298,319]" italics="true" pageId="54" pageNumber="55">I. bureschi</emphasis>
</taxonomicName>
), then disc of tegmina green, with a darkened area behind the CuA; cercal apex is wide, blunt or cut with a massive subapical tooth; if CuP is thin and the cercal apex is thin with long almost apical tooth (
<taxonomicName id="BDC6FBBD393EFFA4B3C30B66FC99990F" box="[740,842,357,378]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="bureschi">
<emphasis id="48B25C2C393EFFA4B3C30B66FC99990F" box="[740,842,357,378]" italics="true" pageId="54" pageNumber="55">I. bureschi</emphasis>
</taxonomicName>
), then CuP is short (1/22/3 of the hind margin of metazone) and the syllable has short main part (&lt;100 ms) followed by few after-clicks.
<emphasis id="48B25C2C393EFFA4B2E20B81FBC599E2" box="[965,1046,386,407]" italics="true" pageId="54" pageNumber="55">Females</emphasis>
: distinction complicated; general colouration fresh-, or yellowish-, or bluish-, or greyish-green; either very large species (hind femur&gt;
<quantity id="BD3E2DDB393EFFA4B5A40B9CFB1999C1" box="[1155,1226,415,436]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="54" pageNumber="55" unit="mm" value="21.0">21 mm</quantity>
, ovipositor&gt;
<quantity id="BD3E2DDB393EFFA4B46D0B9CFA4199C1" box="[1354,1426,415,436]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="54" pageNumber="55" unit="mm" value="14.0">14 mm</quantity>
),
<emphasis id="48B25C2C393EFFA4B1C60BBEFEF699A7" bold="true" box="[225,293,445,466]" pageId="54" pageNumber="55">and/or</emphasis>
tegmina at apex oval,
<emphasis id="48B25C2C393EFFA4B0D80BBEFD9099A7" bold="true" box="[511,579,445,466]" pageId="54" pageNumber="55">and/or</emphasis>
tegmina longer than half the pronotal length
<emphasis id="48B25C2C393EFFA4B2D00BBEFBE899A7" bold="true" box="[1015,1083,445,466]" pageId="54" pageNumber="55">and/or</emphasis>
lateral stripes in metazone brownish or black............................................................................................. 8
</keyLead>
</paragraph>
<paragraph id="7A79803E393EFFA4B1B00BF4FA4F9AF7" blockId="54.[151,1436,151,1992]" pageId="54" pageNumber="55">
<keyLead id="C137230B393EFFA4B1B00BF4FA4F9AF7" pageId="54" pageNumber="55">
<emphasis id="48B25C2C393EFFA4B1C60BFBFECF9A78" box="[225,284,504,525]" italics="true" pageId="54" pageNumber="55">Males</emphasis>
: CuP darker or the same as the colouration of disc of tegmen, CuP not conspicuously widened; disc of tegmina brownish-green, brown or brownish-violet (coloured different that body) without well distinguishable darkened area behind the CuA; cercal apex is wide, rounded or thin (
<emphasis id="48B25C2C393EFFA4B36E0831FDB89A32" bold="true" box="[585,619,562,583]" pageId="54" pageNumber="55">not</emphasis>
blunt or cut); the tooth is not distinctly subapical.
<emphasis id="48B25C2C393EFFA4B5730831FB769A32" box="[1108,1189,562,583]" italics="true" pageId="54" pageNumber="55">Females</emphasis>
: general colouration may be dark green or violet-green; moderately large species (hind femur &lt;
<quantity id="BD3E2DDB393EFFA4B259084CFC179A11" box="[894,964,591,612]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="54" pageNumber="55" unit="mm" value="21.0">21 mm</quantity>
, ovipositor &lt;
<quantity id="BD3E2DDB393EFFA4B564084CFB5A9A11" box="[1091,1161,591,612]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="54" pageNumber="55" unit="mm" value="14.0">14 mm</quantity>
),
<emphasis id="48B25C2C393EFFA4B5BE084CFB6C9A11" bold="true" box="[1177,1215,591,612]" pageId="54" pageNumber="55">and</emphasis>
tegmina at apex blunt,
<emphasis id="48B25C2C393EFFA4B1C6086EFEDB9AF7" bold="true" box="[225,264,621,642]" pageId="54" pageNumber="55">and</emphasis>
tegmina shorter or about half the pronotal length
<emphasis id="48B25C2C393EFFA4B3C2086EFCD89AF7" bold="true" box="[741,779,621,642]" pageId="54" pageNumber="55">and</emphasis>
lateral stripes in metazone (if visible) reddish...............16
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C132989B393EFFA4B1B00889FA489AC9" pageId="54" pageNumber="55">
<paragraph id="7A79803E393EFFA4B1B00889FA4F9AEA" blockId="54.[151,1436,151,1992]" box="[151,1436,650,671]" pageId="54" pageNumber="55">
<keyLead id="C137230B393EFFA4B1B00889FA4F9AEA" box="[151,1436,650,671]" pageId="54" pageNumber="55">
8.
<emphasis id="48B25C2C393EFFA4B1C60889FECF9AEA" box="[225,284,650,671]" italics="true" pageId="54" pageNumber="55">Males</emphasis>
: Stridulatory teeth &lt;100 (as an exception up to
<quantity id="BD3E2DDB393EFFA4B3C30889FCF39AEA" box="[740,800,650,671]" metricMagnitude="0" metricUnit="m" metricValue="2.7939999999999996" pageId="54" pageNumber="55" unit="in" value="110.0">110 in</quantity>
<taxonomicName id="BDC6FBBD393EFFA4B2010889FC559AEA" box="[806,902,650,671]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="tosevski">
<emphasis id="48B25C2C393EFFA4B2010889FC559AEA" box="[806,902,650,671]" italics="true" pageId="54" pageNumber="55">I. tosevski</emphasis>
</taxonomicName>
)........................................... 9
</keyLead>
</paragraph>
<paragraph id="7A79803E393EFFA4B1B008A4FA489AC9" blockId="54.[151,1436,151,1992]" box="[151,1435,679,701]" pageId="54" pageNumber="55">
<keyLead id="C137230B393EFFA4B1B008A4FA489AC9" box="[151,1435,679,701]" pageId="54" pageNumber="55">
<emphasis id="48B25C2C393EFFA4B1C608ABFECF9AC8" box="[225,284,680,701]" italics="true" pageId="54" pageNumber="55">Males</emphasis>
: Stridulatory teeth&gt;100.......................................................................... 13
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C132989B393EFFA4B1B008C6FA4F9B3A" pageId="54" pageNumber="55">
<paragraph id="7A79803E393EFFA4B1B008C6FA4F9B60" blockId="54.[151,1436,151,1992]" pageId="54" pageNumber="55">
<keyLead id="C137230B393EFFA4B1B008C6FA4F9B60" pageId="54" pageNumber="55">
9.
<emphasis id="48B25C2C393EFFA4B1C608C6FECF9AAF" box="[225,284,709,730]" italics="true" pageId="54" pageNumber="55">Males</emphasis>
: CuP weak and short (&lt;2/3 of hind edge of pronotum), well separated from CuA (
<figureCitation id="E2FD9CBB393EFFA4B50D08C6FBA29AAF" box="[1066,1137,709,730]" captionStart="FIGURES 31 55" captionStartId="12.[151,264,244,267]" captionText="FIGURES 31 55. Morphology of male head, pronotum and tegmina of the Balkan Isophya and I. yaraligozi: 31 — I. hospodar (BG: Momchilgrad, 31.05. 1965, NMNHS); 32 — I. rectipennis (BG: Ropotamo, 7.07. 1965, NMNHS); 33 — I. pavelii (holotype of I. rammei Peshev, BG: Chernogorovo, 19.06. 1973, NMNHS); 34 — I. thracica (TR: Elmali, 3.07. 2011, CC); 35 — I. bureschi (BG: “ Pirin, 1300 1450 m ”, 17.08. 1961, NMNHS); 36 — I. yaraligozi (TR: Yaraligoz Pass, 9.07. 2011, CC); 37 — I. tosevski (MK: Moklishte, 30.05. 2010, CC); 38 — I. andreevae (holotype, BG: Kresna Gorge, 2.06. 1961, NMNHS); 39 — I. clara (BA: Sarajevo, NMW); 40 — I. miksici (holotype, BG: Berkovitsa, 700 m, 21.06. 1958, NMNHS); 41 — I. plevnensis (holotype, BG: Chernelka, 30.07. 1973, NMNHS); 42 — I. longicaudata adamovici (holotype, BG: “ Sliven ”, NMNHS); 43 — I. l. longicaudata (BG: Batova Gora, 26.06. 1956, NMNHS); 44 — I. m. modesta (= intermedia) (RO: Dobrogea, Babadag, CI); 45 — I. rhodopensis leonorae (BG: Popovi Livadi, 1300 m, 4.07. 1961, NMNHS); 46 — I. rh. rhodopensis (BG: Chepelare, 16.07. 1956, NMNHS); 47 — I. rh. petkovi (BG: Mandritsa, 4.06. 1965, NMNHS); 48 — I. modestior (BG: Kopitoto, 17.07. 1958, NMNHS); 49 — I. dobrogensis (RO: Popina Island, CI); 50 — I. zubowskii (RO: “ Jasi pād, Birnari, 25.06. 1956, NMNHS); 51 — I. aff. camptoxypha (“ Montenegro ”, NMW); 52 — I. gulae (BG: Dolna Topchiya, 23.06. 2008, NMNHS); 53 — I. obtusa (BG: Vezhen Peak, 1900 m, 1.09. 1960, NMNHS); 54 — I. amplipennis (TR: “ Bursa | Gursu, | Kazanci &amp; | Eskibaglar | 14. vi. 1959 ”, “ by Dr. E. Can. | B. M. 1960 173 ”, NHM); 55 — I. speciosa (BG: Plevoun, 5.06. 1965, NMNHS)." pageId="54" pageNumber="55">Fig. 35</figureCitation>
); tip of cerci narrow-oval (
<figureCitation id="E2FD9CBB393EFFA4B45608C6FF289A82" captionStart="FIGURES 80 104" captionStartId="14.[151,264,1835,1858]" captionTargetBox="[226,1360,681,1815]" captionTargetId="figure@14.[226,1362,681,1816]" captionTargetPageId="14" captionText="FIGURES 80 104. Male abdominal terminalia of the Balkan Isophya and I. yaraligozi (localities as in Figs 31 55): 80 — I. hospodar; 81 — I. rectipennis; 82 — I. pavelii; 83 — I. thracica; 84 — I. bureschi; 85 — I. yaraligozi; 86 — I. tosevski; 87 — I. andreevae; 88 — I. clara; 89 — I. miksici; 90 — I. plevnensis; 91 — I. longicaudata adamovici; 92 — I. l. longicaudata; 93 — I. m. modesta (= intermedia); 94 — I. rhodopensis leonorae; 95 — I. rh. rhodopensis; 96 — I. rh. petkovi; 97 — I. modestior; 98 — I. dobrogensis; 99 — I. zubowskii; 100 — I. aff. camptoxypha; 101 — I. gulae; 102 — I. obtusa; 103 — I. amplipennis; 104 — I. speciosa. Scale (if present) = 2 mm." httpUri="https://zenodo.org/record/246555/files/figure.png" pageId="54" pageNumber="55">Figs 84</figureCitation>
,
<figureCitation id="E2FD9CBB393EFFA4B02308E1FEF89A82" box="[260,299,738,759]" captionStart="FIGURES 139 144" captionStartId="22.[151,264,1519,1542]" captionTargetBox="[151,1435,301,1497]" captionTargetId="figure@22.[151,1436,301,1498]" captionTargetPageId="22" captionText="FIGURES 139 144. SEM of taxonomically important structures of Isophya: 139 — I. bureschi (BG: Pirin Mts, Chalin Valog); 140 — I. yaraligozi (TR: Yaraligoz Mt.); 141 — I. tosevski (MK: Dojran Lake); 142, 143 — I. andreevae (142 — BG: Kresna Gorge; 143 — BG: Eleshnitsa Locality); 144 — I. clara (BA: Sarajevo). A—male stridulatory file (139 — scale 200 μm; othersscale 1 mm); B—ventro-apical view of apex of male cerci (scale 100 μm); C—female stridulatory apparatus (139 — scale 200 μm; 141, 144 — scale 1 mm; 142 — scale 500 μm)." httpUri="https://zenodo.org/record/246559/files/figure.png" pageId="54" pageNumber="55">139</figureCitation>
B).
<emphasis id="48B25C2C393EFFA4B06B08E1FEAF9A82" box="[332,380,738,759]" italics="true" pageId="54" pageNumber="55">Song</emphasis>
(
<figureCitation id="E2FD9CBB393EFFA4B0AF08E1FE0A9A82" box="[392,473,738,759]" captionStart="FIGURES 145 149" captionStartId="23.[151,264,1555,1578]" captionTargetBox="[151,1436,409,1534]" captionTargetId="figure@23.[151,1436,409,1534]" captionTargetPageId="23" captionText="FIGURES 145 149. Male calling song at two different speeds (air temperature at recording shown; time scale below): 145 — I. bureschi (BG: Pirin Mts, Chalin Valog); 146 — I. yaraligozi (TR: Yaraligoz Mt.); 147 — I. tosevski (MK: Moklishte Village); 148, 149 — I. andreevae (148 — BG: Eleshnitsa Locality; 149 — BG: Kresna Gorge)." httpUri="https://zenodo.org/record/246560/files/figure.png" pageId="54" pageNumber="55">Fig. 145</figureCitation>
) of syllables with very short main part (3060 ms) frequently followed by a group of after-clicks following at&gt;500 ms interval......................................................................
<taxonomicName id="BDC6FBBD393EFFA4B4110903FA4F9B60" box="[1334,1436,768,789]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="bureschi">
<emphasis id="48B25C2C393EFFA4B4110903FA4F9B60" box="[1334,1436,768,789]" italics="true" pageId="54" pageNumber="55">I. bureschi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7A79803E393EFFA4B1B0091EFA4F9B3A" blockId="54.[151,1436,151,1992]" pageId="54" pageNumber="55">
<keyLead id="C137230B393EFFA4B1B0091EFA4F9B3A" pageId="54" pageNumber="55">
<emphasis id="48B25C2C393EFFA4B1C6091EFECF9B47" box="[225,284,797,818]" italics="true" pageId="54" pageNumber="55">Males</emphasis>
: CuP variable, &lt;2/33/4 of the hind edge of pronotum, moderately or strongly attached to CuA; tip of cerci wide—blunt or cut.
<emphasis id="48B25C2C393EFFA4B0030939FE879B3A" box="[292,340,826,847]" italics="true" pageId="54" pageNumber="55">Song</emphasis>
: main part of syllables longer (100450 ms), sometimes followed by less separated after-clicks............. 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C132989B393EFFA4B1B00954FA489B8A" pageId="54" pageNumber="55">
<paragraph id="7A79803E393EFFA4B1B00954FA489BD2" blockId="54.[151,1436,151,1992]" pageId="54" pageNumber="55">
<keyLead id="C137230B393EFFA4B1B00954FA489BD2" pageId="54" pageNumber="55">
10.
<emphasis id="48B25C2C393EFFA4B1C6095BFECF9B18" box="[225,284,856,877]" italics="true" pageId="54" pageNumber="55">Males</emphasis>
: CuP weaker and shorter (±2/3 of hind edge of pronotum), moderately attached to CuA (
<figureCitation id="E2FD9CBB393EFFA4B5530954FB379B19" box="[1140,1252,855,876]" captionStart="FIGURES 31 55" captionStartId="12.[151,264,244,267]" captionText="FIGURES 31 55. Morphology of male head, pronotum and tegmina of the Balkan Isophya and I. yaraligozi: 31 — I. hospodar (BG: Momchilgrad, 31.05. 1965, NMNHS); 32 — I. rectipennis (BG: Ropotamo, 7.07. 1965, NMNHS); 33 — I. pavelii (holotype of I. rammei Peshev, BG: Chernogorovo, 19.06. 1973, NMNHS); 34 — I. thracica (TR: Elmali, 3.07. 2011, CC); 35 — I. bureschi (BG: “ Pirin, 1300 1450 m ”, 17.08. 1961, NMNHS); 36 — I. yaraligozi (TR: Yaraligoz Pass, 9.07. 2011, CC); 37 — I. tosevski (MK: Moklishte, 30.05. 2010, CC); 38 — I. andreevae (holotype, BG: Kresna Gorge, 2.06. 1961, NMNHS); 39 — I. clara (BA: Sarajevo, NMW); 40 — I. miksici (holotype, BG: Berkovitsa, 700 m, 21.06. 1958, NMNHS); 41 — I. plevnensis (holotype, BG: Chernelka, 30.07. 1973, NMNHS); 42 — I. longicaudata adamovici (holotype, BG: “ Sliven ”, NMNHS); 43 — I. l. longicaudata (BG: Batova Gora, 26.06. 1956, NMNHS); 44 — I. m. modesta (= intermedia) (RO: Dobrogea, Babadag, CI); 45 — I. rhodopensis leonorae (BG: Popovi Livadi, 1300 m, 4.07. 1961, NMNHS); 46 — I. rh. rhodopensis (BG: Chepelare, 16.07. 1956, NMNHS); 47 — I. rh. petkovi (BG: Mandritsa, 4.06. 1965, NMNHS); 48 — I. modestior (BG: Kopitoto, 17.07. 1958, NMNHS); 49 — I. dobrogensis (RO: Popina Island, CI); 50 — I. zubowskii (RO: “ Jasi pād, Birnari, 25.06. 1956, NMNHS); 51 — I. aff. camptoxypha (“ Montenegro ”, NMW); 52 — I. gulae (BG: Dolna Topchiya, 23.06. 2008, NMNHS); 53 — I. obtusa (BG: Vezhen Peak, 1900 m, 1.09. 1960, NMNHS); 54 — I. amplipennis (TR: “ Bursa | Gursu, | Kazanci &amp; | Eskibaglar | 14. vi. 1959 ”, “ by Dr. E. Can. | B. M. 1960 173 ”, NHM); 55 — I. speciosa (BG: Plevoun, 5.06. 1965, NMNHS)." pageId="54" pageNumber="55">Figs 39, 40</figureCitation>
); tip of cerci blunt, bearing short, wide tooth (
<figureCitation id="E2FD9CBB393EFFA4B0C10976FDED9BFF" box="[486,574,885,906]" captionStart="FIGURES 139 144" captionStartId="22.[151,264,1519,1542]" captionTargetBox="[151,1435,301,1497]" captionTargetId="figure@22.[151,1436,301,1498]" captionTargetPageId="22" captionText="FIGURES 139 144. SEM of taxonomically important structures of Isophya: 139 — I. bureschi (BG: Pirin Mts, Chalin Valog); 140 — I. yaraligozi (TR: Yaraligoz Mt.); 141 — I. tosevski (MK: Dojran Lake); 142, 143 — I. andreevae (142 — BG: Kresna Gorge; 143 — BG: Eleshnitsa Locality); 144 — I. clara (BA: Sarajevo). A—male stridulatory file (139 — scale 200 μm; othersscale 1 mm); B—ventro-apical view of apex of male cerci (scale 100 μm); C—female stridulatory apparatus (139 — scale 200 μm; 141, 144 — scale 1 mm; 142 — scale 500 μm)." httpUri="https://zenodo.org/record/246559/files/figure.png" pageId="54" pageNumber="55">Figs 144</figureCitation>
B, 150B). Colouration usually fresh-green; disc of tegmina dark-green with dark-brown stridulatory area.
<emphasis id="48B25C2C393EFFA4B0AE0991FE6A9BD2" box="[393,441,914,935]" italics="true" pageId="54" pageNumber="55">Song</emphasis>
: syllables usually without after-clicks..................................................11
</keyLead>
</paragraph>
<paragraph id="7A79803E393EFFA4B1B009ACFA489B8A" blockId="54.[151,1436,151,1992]" pageId="54" pageNumber="55">
<keyLead id="C137230B393EFFA4B1B009ACFA489B8A" pageId="54" pageNumber="55">
<emphasis id="48B25C2C393EFFA4B1C609B3FECF9BB0" box="[225,284,944,965]" italics="true" pageId="54" pageNumber="55">Males</emphasis>
: CuP wider and longer (2/33/4 of hind edge of pronotum), strongly attached to CuA (
<figureCitation id="E2FD9CBB393EFFA4B57809ACFB1C9BB1" box="[1119,1231,943,964]" captionStart="FIGURES 31 55" captionStartId="12.[151,264,244,267]" captionText="FIGURES 31 55. Morphology of male head, pronotum and tegmina of the Balkan Isophya and I. yaraligozi: 31 — I. hospodar (BG: Momchilgrad, 31.05. 1965, NMNHS); 32 — I. rectipennis (BG: Ropotamo, 7.07. 1965, NMNHS); 33 — I. pavelii (holotype of I. rammei Peshev, BG: Chernogorovo, 19.06. 1973, NMNHS); 34 — I. thracica (TR: Elmali, 3.07. 2011, CC); 35 — I. bureschi (BG: “ Pirin, 1300 1450 m ”, 17.08. 1961, NMNHS); 36 — I. yaraligozi (TR: Yaraligoz Pass, 9.07. 2011, CC); 37 — I. tosevski (MK: Moklishte, 30.05. 2010, CC); 38 — I. andreevae (holotype, BG: Kresna Gorge, 2.06. 1961, NMNHS); 39 — I. clara (BA: Sarajevo, NMW); 40 — I. miksici (holotype, BG: Berkovitsa, 700 m, 21.06. 1958, NMNHS); 41 — I. plevnensis (holotype, BG: Chernelka, 30.07. 1973, NMNHS); 42 — I. longicaudata adamovici (holotype, BG: “ Sliven ”, NMNHS); 43 — I. l. longicaudata (BG: Batova Gora, 26.06. 1956, NMNHS); 44 — I. m. modesta (= intermedia) (RO: Dobrogea, Babadag, CI); 45 — I. rhodopensis leonorae (BG: Popovi Livadi, 1300 m, 4.07. 1961, NMNHS); 46 — I. rh. rhodopensis (BG: Chepelare, 16.07. 1956, NMNHS); 47 — I. rh. petkovi (BG: Mandritsa, 4.06. 1965, NMNHS); 48 — I. modestior (BG: Kopitoto, 17.07. 1958, NMNHS); 49 — I. dobrogensis (RO: Popina Island, CI); 50 — I. zubowskii (RO: “ Jasi pād, Birnari, 25.06. 1956, NMNHS); 51 — I. aff. camptoxypha (“ Montenegro ”, NMW); 52 — I. gulae (BG: Dolna Topchiya, 23.06. 2008, NMNHS); 53 — I. obtusa (BG: Vezhen Peak, 1900 m, 1.09. 1960, NMNHS); 54 — I. amplipennis (TR: “ Bursa | Gursu, | Kazanci &amp; | Eskibaglar | 14. vi. 1959 ”, “ by Dr. E. Can. | B. M. 1960 173 ”, NHM); 55 — I. speciosa (BG: Plevoun, 5.06. 1965, NMNHS)." pageId="54" pageNumber="55">Figs 37, 38</figureCitation>
); tip of cerci cut and/ or incised, bearing long tooth (
<figureCitation id="E2FD9CBB393EFFA4B33709CEFD509B97" box="[528,643,973,994]" captionStart="FIGURES 80 104" captionStartId="14.[151,264,1835,1858]" captionTargetBox="[226,1360,681,1815]" captionTargetId="figure@14.[226,1362,681,1816]" captionTargetPageId="14" captionText="FIGURES 80 104. Male abdominal terminalia of the Balkan Isophya and I. yaraligozi (localities as in Figs 31 55): 80 — I. hospodar; 81 — I. rectipennis; 82 — I. pavelii; 83 — I. thracica; 84 — I. bureschi; 85 — I. yaraligozi; 86 — I. tosevski; 87 — I. andreevae; 88 — I. clara; 89 — I. miksici; 90 — I. plevnensis; 91 — I. longicaudata adamovici; 92 — I. l. longicaudata; 93 — I. m. modesta (= intermedia); 94 — I. rhodopensis leonorae; 95 — I. rh. rhodopensis; 96 — I. rh. petkovi; 97 — I. modestior; 98 — I. dobrogensis; 99 — I. zubowskii; 100 — I. aff. camptoxypha; 101 — I. gulae; 102 — I. obtusa; 103 — I. amplipennis; 104 — I. speciosa. Scale (if present) = 2 mm." httpUri="https://zenodo.org/record/246555/files/figure.png" pageId="54" pageNumber="55">Figs 86, 87</figureCitation>
). Colouration usually yellowish-green; disc of tegmina green with both dark and lighter (yellowish-brown) areas.
<emphasis id="48B25C2C393EFFA4B33C09E9FD989B8A" box="[539,587,1002,1023]" italics="true" pageId="54" pageNumber="55">Song</emphasis>
: syllables usually with after-clicks......................................... 12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C132989B393EFFA4B1B00E04FA4F9C4F" pageId="54" pageNumber="55">
<paragraph id="7A79803E393EFFA4B1B00E04FA4F9C68" blockId="54.[151,1436,151,1992]" box="[151,1436,1031,1053]" pageId="54" pageNumber="55">
<keyLead id="C137230B393EFFA4B1B00E04FA4F9C68" box="[151,1436,1031,1053]" pageId="54" pageNumber="55">
11. Western and Central Balkan Peninsula from
<collectingCountry id="02D1C0AE393EFFA4B3AB0E04FCD29C69" box="[652,769,1031,1052]" name="Serbia" pageId="54" pageNumber="55">Montenegro</collectingCountry>
to Belgrade.......................................
<taxonomicName id="BDC6FBBD393EFFA4B4720E0BFA4F9C68" box="[1365,1436,1032,1053]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="clara">
<emphasis id="48B25C2C393EFFA4B4720E0BFA4F9C68" box="[1365,1436,1032,1053]" italics="true" pageId="54" pageNumber="55">I. clara</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7A79803E393EFFA4B1B00E26FA4F9C4F" blockId="54.[151,1436,151,1992]" box="[151,1436,1061,1082]" pageId="54" pageNumber="55">
<keyLead id="C137230B393EFFA4B1B00E26FA4F9C4F" box="[151,1436,1061,1082]" pageId="54" pageNumber="55">
NW
<collectingCountry id="02D1C0AE393EFFA4B0280E26FEB19C4F" box="[271,354,1061,1082]" name="Bulgaria" pageId="54" pageNumber="55">Bulgaria</collectingCountry>
and E
<collectingCountry id="02D1C0AE393EFFA4B0830E26FE379C4F" box="[420,484,1061,1082]" name="Serbia" pageId="54" pageNumber="55">Serbia</collectingCountry>
.........................................................................
<taxonomicName id="BDC6FBBD393EFFA4B4660E26FA4F9C4F" box="[1345,1436,1061,1082]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="miksici">
<emphasis id="48B25C2C393EFFA4B4660E26FA4F9C4F" box="[1345,1436,1061,1082]" italics="true" pageId="54" pageNumber="55">I. miksici</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C132989B393EFFA4B1B00E41FA4F9C9F" pageId="54" pageNumber="55">
<paragraph id="7A79803E393EFFA4B1B00E41FA4F9CE7" blockId="54.[151,1436,151,1992]" pageId="54" pageNumber="55">
<keyLead id="C137230B393EFFA4B1B00E41FA4F9CE7" pageId="54" pageNumber="55">
12.
<emphasis id="48B25C2C393EFFA4B1C60E41FECF9C22" box="[225,284,1090,1111]" italics="true" pageId="54" pageNumber="55">Males</emphasis>
: tip of cerci notably incised (
<figureCitation id="E2FD9CBB393EFFA4B3660E41FD459C22" box="[577,662,1090,1111]" captionStart="FIGURES 139 144" captionStartId="22.[151,264,1519,1542]" captionTargetBox="[151,1435,301,1497]" captionTargetId="figure@22.[151,1436,301,1498]" captionTargetPageId="22" captionText="FIGURES 139 144. SEM of taxonomically important structures of Isophya: 139 — I. bureschi (BG: Pirin Mts, Chalin Valog); 140 — I. yaraligozi (TR: Yaraligoz Mt.); 141 — I. tosevski (MK: Dojran Lake); 142, 143 — I. andreevae (142 — BG: Kresna Gorge; 143 — BG: Eleshnitsa Locality); 144 — I. clara (BA: Sarajevo). A—male stridulatory file (139 — scale 200 μm; othersscale 1 mm); B—ventro-apical view of apex of male cerci (scale 100 μm); C—female stridulatory apparatus (139 — scale 200 μm; 141, 144 — scale 1 mm; 142 — scale 500 μm)." httpUri="https://zenodo.org/record/246559/files/figure.png" pageId="54" pageNumber="55">Fig. 141</figureCitation>
B).
<emphasis id="48B25C2C393EFFA4B39C0E41FCDF9C22" box="[699,780,1090,1111]" italics="true" pageId="54" pageNumber="55">Females</emphasis>
: ovipositor (
<figureCitation id="E2FD9CBB393EFFA4B2AC0E41FC0E9C22" box="[907,989,1090,1111]" captionStart="FIGURES 105 129" captionStartId="17.[151,264,1173,1196]" captionTargetBox="[151,1436,195,1152]" captionTargetId="figure@17.[151,1436,195,1152]" captionTargetPageId="17" captionText="FIGURES 105 129. Female abdominal terminalia of the Balkan Isophya and I. yaraligozi (localities as in Figs 31 55 unless specifically given); figures of the whole ovipositor are followed by enlarged figure of the ovipositor base: 105 — I. hospodar (BG: Dolni Glavanak, 25.04. 2005, CC); 106 — I. rectipennis (BG: Byala Vill., 27.06. 2002, CC); 107 — I. pavelii (BG: Chernogorovo, 29.05. 2006, CC); 108 — I. thracica; 109 — I. bureschi (BG: Bansko, 8.08. 2006, CC); 110 — I. yaraligozi; 111 — I. tosevski; 112 — I. andreevae (BG: Kresnensko Hanche, 19.04. 2006, CC); 113 — I. clara; 114 — I. miksici (BG: Gorski Dom Lodge, 24.06. 2006, CC); 115 — I. plevnensis (BG: Levishte, 14.07. 2009, CC); 116 — I. longicaudata adamovici (BG: Karandila, 24.06. 2008, CC); 117 — I. l. longicaudata (BG: Sofia University Botanical garden, 26.06. 1.07.2004, CC); 118 — I. m. modesta (= intermedia); 119 — I. rhodopensis leonorae (BG: Livade, 9.08. 2006, CC); 120 — I. rh. rhodopensis (BG: Bachkovo, 600 m, 23.05. 2004, CC); 121 — I. rh. petkovi (BG: Gluhite Kamani, 23.06. 2008, CC); 122 — I. modestior (BG: Bankya, 24.05. 2006, CC); 123 — I. dobrogensis; 124 — I. zubowskii; 125 — I. aff. camptoxypha (data as in Fig. 75); 126 — I. gulae; 127 — I. obtusa (BG: Ravnets Ridge, 14.07. 2009, CC); 128 — I. amplipennis (as in Fig. 78); 129 — I. speciosa (BG: Bankya, 24.05. 2006, CC). Indications on Fig. 127 show, respectively: G—gonangulum; L—lamella; P—lateral pit between lamella and gonangulum. Scale (if present) = 5 mm." httpUri="https://zenodo.org/record/246556/files/figure.png" pageId="54" pageNumber="55">Fig. 111</figureCitation>
) shorter, &lt;
<quantity id="BD3E2DDB393EFFA4B56D0E41FB5C9C22" box="[1098,1167,1090,1111]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="54" pageNumber="55" unit="mm" value="15.0">15 mm</quantity>
(usually
<quantity id="BD3E2DDB393EFFA4B5CB0E41FA899C22" box="[1260,1370,1090,1111]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="54" pageNumber="55" unit="mm" value="13.0" valueMax="14.0" valueMin="12.0">1214 mm</quantity>
).
<emphasis id="48B25C2C393EFFA4B44B0E41FA4F9C22" box="[1388,1436,1090,1111]" italics="true" pageId="54" pageNumber="55">Song</emphasis>
(
<figureCitation id="E2FD9CBB393EFFA4B1CF0E5CFEEE9C01" box="[232,317,1119,1140]" captionStart="FIGURES 145 149" captionStartId="23.[151,264,1555,1578]" captionTargetBox="[151,1436,409,1534]" captionTargetId="figure@23.[151,1436,409,1534]" captionTargetPageId="23" captionText="FIGURES 145 149. Male calling song at two different speeds (air temperature at recording shown; time scale below): 145 — I. bureschi (BG: Pirin Mts, Chalin Valog); 146 — I. yaraligozi (TR: Yaraligoz Mt.); 147 — I. tosevski (MK: Moklishte Village); 148, 149 — I. andreevae (148 — BG: Eleshnitsa Locality; 149 — BG: Kresna Gorge)." httpUri="https://zenodo.org/record/246560/files/figure.png" pageId="54" pageNumber="55">Fig. 147</figureCitation>
): syllables usually with 510 after-clicks. SE
<collectingCountry id="02D1C0AE393EFFA4B3D10E5CFC0B9C01" box="[758,984,1119,1140]" name="Macedonia" pageId="54" pageNumber="55">Republic of Macedonia</collectingCountry>
and N
<collectingCountry id="02D1C0AE393EFFA4B5380E5CFBB59C01" box="[1055,1126,1119,1140]" name="Greece" pageId="54" pageNumber="55">Greece</collectingCountry>
, mostly along the Lower Vardar Valley.......................................................................................
<taxonomicName id="BDC6FBBD393EFFA4B41B0E7EFA4F9CE7" box="[1340,1436,1149,1170]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="tosevski">
<emphasis id="48B25C2C393EFFA4B41B0E7EFA4F9CE7" box="[1340,1436,1149,1170]" italics="true" pageId="54" pageNumber="55">I. tosevski</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7A79803E393EFFA4B1B00E99FA4F9C9F" blockId="54.[151,1436,151,1992]" pageId="54" pageNumber="55">
<keyLead id="C137230B393EFFA4B1B00E99FA4F9C9F" pageId="54" pageNumber="55">
<emphasis id="48B25C2C393EFFA4B1C60E99FECF9CDA" box="[225,284,1178,1199]" italics="true" pageId="54" pageNumber="55">Males</emphasis>
: tip of cerci not or slightly incised (
<figureCitation id="E2FD9CBB393EFFA4B3A50E99FD089CDA" box="[642,731,1178,1199]" captionStart="FIGURES 139 144" captionStartId="22.[151,264,1519,1542]" captionTargetBox="[151,1435,301,1497]" captionTargetId="figure@22.[151,1436,301,1498]" captionTargetPageId="22" captionText="FIGURES 139 144. SEM of taxonomically important structures of Isophya: 139 — I. bureschi (BG: Pirin Mts, Chalin Valog); 140 — I. yaraligozi (TR: Yaraligoz Mt.); 141 — I. tosevski (MK: Dojran Lake); 142, 143 — I. andreevae (142 — BG: Kresna Gorge; 143 — BG: Eleshnitsa Locality); 144 — I. clara (BA: Sarajevo). A—male stridulatory file (139 — scale 200 μm; othersscale 1 mm); B—ventro-apical view of apex of male cerci (scale 100 μm); C—female stridulatory apparatus (139 — scale 200 μm; 141, 144 — scale 1 mm; 142 — scale 500 μm)." httpUri="https://zenodo.org/record/246559/files/figure.png" pageId="54" pageNumber="55">Figs 142</figureCitation>
B, 143B).
<emphasis id="48B25C2C393EFFA4B2660E99FCA29CDA" box="[833,881,1178,1199]" italics="true" pageId="54" pageNumber="55">Song</emphasis>
(
<figureCitation id="E2FD9CBB393EFFA4B2580E99FBDF9CDA" box="[895,1036,1178,1199]" captionStart="FIGURES 145 149" captionStartId="23.[151,264,1555,1578]" captionTargetBox="[151,1436,409,1534]" captionTargetId="figure@23.[151,1436,409,1534]" captionTargetPageId="23" captionText="FIGURES 145 149. Male calling song at two different speeds (air temperature at recording shown; time scale below): 145 — I. bureschi (BG: Pirin Mts, Chalin Valog); 146 — I. yaraligozi (TR: Yaraligoz Mt.); 147 — I. tosevski (MK: Moklishte Village); 148, 149 — I. andreevae (148 — BG: Eleshnitsa Locality; 149 — BG: Kresna Gorge)." httpUri="https://zenodo.org/record/246560/files/figure.png" pageId="54" pageNumber="55">Figs 148, 149</figureCitation>
): syllables without or with up to 5 afterclicks.
<emphasis id="48B25C2C393EFFA4B00F0EBBFEAA9CB8" box="[296,377,1208,1229]" italics="true" pageId="54" pageNumber="55">Females</emphasis>
: ovipositor (
<figureCitation id="E2FD9CBB393EFFA4B0DC0EB4FD9C9CB9" box="[507,591,1207,1228]" captionStart="FIGURES 105 129" captionStartId="17.[151,264,1173,1196]" captionTargetBox="[151,1436,195,1152]" captionTargetId="figure@17.[151,1436,195,1152]" captionTargetPageId="17" captionText="FIGURES 105 129. Female abdominal terminalia of the Balkan Isophya and I. yaraligozi (localities as in Figs 31 55 unless specifically given); figures of the whole ovipositor are followed by enlarged figure of the ovipositor base: 105 — I. hospodar (BG: Dolni Glavanak, 25.04. 2005, CC); 106 — I. rectipennis (BG: Byala Vill., 27.06. 2002, CC); 107 — I. pavelii (BG: Chernogorovo, 29.05. 2006, CC); 108 — I. thracica; 109 — I. bureschi (BG: Bansko, 8.08. 2006, CC); 110 — I. yaraligozi; 111 — I. tosevski; 112 — I. andreevae (BG: Kresnensko Hanche, 19.04. 2006, CC); 113 — I. clara; 114 — I. miksici (BG: Gorski Dom Lodge, 24.06. 2006, CC); 115 — I. plevnensis (BG: Levishte, 14.07. 2009, CC); 116 — I. longicaudata adamovici (BG: Karandila, 24.06. 2008, CC); 117 — I. l. longicaudata (BG: Sofia University Botanical garden, 26.06. 1.07.2004, CC); 118 — I. m. modesta (= intermedia); 119 — I. rhodopensis leonorae (BG: Livade, 9.08. 2006, CC); 120 — I. rh. rhodopensis (BG: Bachkovo, 600 m, 23.05. 2004, CC); 121 — I. rh. petkovi (BG: Gluhite Kamani, 23.06. 2008, CC); 122 — I. modestior (BG: Bankya, 24.05. 2006, CC); 123 — I. dobrogensis; 124 — I. zubowskii; 125 — I. aff. camptoxypha (data as in Fig. 75); 126 — I. gulae; 127 — I. obtusa (BG: Ravnets Ridge, 14.07. 2009, CC); 128 — I. amplipennis (as in Fig. 78); 129 — I. speciosa (BG: Bankya, 24.05. 2006, CC). Indications on Fig. 127 show, respectively: G—gonangulum; L—lamella; P—lateral pit between lamella and gonangulum. Scale (if present) = 5 mm." httpUri="https://zenodo.org/record/246556/files/figure.png" pageId="54" pageNumber="55">Fig. 112</figureCitation>
) longer, usually over
<quantity id="BD3E2DDB393EFFA4B2090EB4FCA49CB9" box="[814,887,1207,1228]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="54" pageNumber="55" unit="mm" value="15.0">15 mm</quantity>
. SW
<collectingCountry id="02D1C0AE393EFFA4B2960EB4FBD79CB9" box="[945,1028,1207,1228]" name="Bulgaria" pageId="54" pageNumber="55">Bulgaria</collectingCountry>
and N
<collectingCountry id="02D1C0AE393EFFA4B5730EB4FB489CB9" box="[1108,1179,1207,1228]" name="Greece" pageId="54" pageNumber="55">Greece</collectingCountry>
, mostly along the Middle Strouma Valley..............................................................................
<taxonomicName id="BDC6FBBD393EFFA4B4020ED6FA4F9C9F" box="[1317,1436,1237,1258]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="andreevae">
<emphasis id="48B25C2C393EFFA4B4020ED6FA4F9C9F" box="[1317,1436,1237,1258]" italics="true" pageId="54" pageNumber="55">I. andreevae</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C132989B393EFFA4B1B00EF1FA489D2A" pageId="54" pageNumber="55">
<paragraph id="7A79803E393EFFA4B1B00EF1FA4F9D51" blockId="54.[151,1436,151,1992]" pageId="54" pageNumber="55">
<keyLead id="C137230B393EFFA4B1B00EF1FA4F9D51" pageId="54" pageNumber="55">
13.
<emphasis id="48B25C2C393EFFA4B1C60EF1FECF9D72" box="[225,284,1266,1287]" italics="true" pageId="54" pageNumber="55">Males</emphasis>
: Tip of cerci cut or incised, bearing large, long, showel- or crest-like tooth (
<figureCitation id="E2FD9CBB393EFFA4B2D10EF1FB539D72" box="[1014,1152,1266,1287]" captionStart="FIGURES 162 167" captionStartId="30.[151,264,1332,1355]" captionTargetBox="[151,1435,218,1310]" captionTargetId="figure@30.[151,1436,218,1311]" captionTargetPageId="30" captionText="FIGURES 162 167. SEM of taxonomically important structures of Isophya: 162, 163 — I. rhodopensis leonorae (162 — BG: N Pirin Mts; 163 — BG: Alibotoush Mt.); 164 166 — I. rhodopensis rhodopensis (164 — typical form, BG: above Smolyan; 165 — northwestern form, BG: Batak Lake; 166 — typical form or transitional to I. rh. petkovi, BG: near Assenovgrad); 167 — I. rhodopensis petkovi (BG: Glouhite Kamani Locality). A—male stridulatory file (162, 166 — scale 200 μm; the rest—scale 1 mm); B—ventral / ventro-apical view of apex of male cerci (scale 100 μm); C—female stridulatory apparatus (162 — scale 100 μm; 166 — scale 200 μm; the rest—scale 500 μm)." httpUri="https://zenodo.org/record/246563/files/figure.png" pageId="54" pageNumber="55">Figs 162167</figureCitation>
B). Colouration usually freshgreen. S
<collectingCountry id="02D1C0AE393EFFA4B0120F0CFE5B9D51" box="[309,392,1295,1316]" name="Bulgaria" pageId="54" pageNumber="55">Bulgaria</collectingCountry>
and NE
<collectingCountry id="02D1C0AE393EFFA4B0FA0F0CFDF29D51" box="[477,545,1295,1316]" name="Greece" pageId="54" pageNumber="55">Greece</collectingCountry>
on the territory of Rila-Rhodopean Mountain group......
<taxonomicName id="BDC6FBBD393EFFA4B54F0F13FB229D50" box="[1128,1265,1296,1317]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="rhodopensis">
<emphasis id="48B25C2C393EFFA4B54F0F13FB229D50" box="[1128,1265,1296,1317]" italics="true" pageId="54" pageNumber="55">I. rhodopensis</emphasis>
</taxonomicName>
(for ssp. see text)
</keyLead>
</paragraph>
<paragraph id="7A79803E393EFFA4B1B00F2EFA489D2A" blockId="54.[151,1436,151,1992]" pageId="54" pageNumber="55">
<keyLead id="C137230B393EFFA4B1B00F2EFA489D2A" pageId="54" pageNumber="55">
<emphasis id="48B25C2C393EFFA4B1C60F2EFECF9D37" box="[225,284,1325,1346]" italics="true" pageId="54" pageNumber="55">Males</emphasis>
: Tip of cerci widely blunt, bearing short, wide tooth (
<figureCitation id="E2FD9CBB393EFFA4B2010F2EFC7C9D37" box="[806,943,1325,1346]" captionStart="FIGURES 150 155" captionStartId="27.[151,264,1462,1485]" captionTargetBox="[151,1435,193,1440]" captionTargetId="figure@27.[151,1436,193,1441]" captionTargetPageId="27" captionText="FIGURES 150 155. SEM of taxonomically important structures of Isophya: 150, 151 — I. miksici (150 — BG: Gorski Dom Lodge; 151 — BG: Iskar); 152 — I. plevnensis (152 A—BG: Chernelka Canyon; 152 B, C—BG: Apriltsi); 153 — I. longicaudata adamovici (BG: Karandila Locality); 154 — I. longicaudata longicaudata (BG: Bolata Valley); 155 — I. m. modesta (= intermedia) (RO: Babadag). A—male stridulatory file (scale 1 mm); B—ventro-apical view of apex of male cerci (152 B deformed) (scale 100 μm); C—female stridulatory apparatus (scale 500 μm)." httpUri="https://zenodo.org/record/246561/files/figure.png" pageId="54" pageNumber="55">Figs 152154</figureCitation>
B). Colouration usually bluish- or greyish-green. N
<collectingCountry id="02D1C0AE393EFFA4B1C60F49FEE79D2A" box="[225,308,1354,1375]" name="Bulgaria" pageId="54" pageNumber="55">Bulgaria</collectingCountry>
and West Black Sea coast....................................................................... 14
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C132989B393EFFA4B1B00F64FA489DC2" pageId="54" pageNumber="55">
<paragraph id="7A79803E393EFFA4B1B00F64FA4F9D09" blockId="54.[151,1436,151,1992]" box="[151,1436,1383,1405]" pageId="54" pageNumber="55">
<keyLead id="C137230B393EFFA4B1B00F64FA4F9D09" box="[151,1436,1383,1405]" pageId="54" pageNumber="55">
14.
<emphasis id="48B25C2C393EFFA4B1C60F6BFECF9D08" box="[225,284,1384,1405]" italics="true" pageId="54" pageNumber="55">Males</emphasis>
: Stridulatory teeth&gt;145.
<emphasis id="48B25C2C393EFFA4B3350F6BFD919D08" box="[530,578,1384,1405]" italics="true" pageId="54" pageNumber="55">Song</emphasis>
(
<figureCitation id="E2FD9CBB393EFFA4B3680F64FD0B9D09" box="[591,728,1383,1404]" captionStart="FIGURES 156 161" captionStartId="28.[151,264,1682,1705]" captionTargetBox="[151,1436,331,1661]" captionTargetId="figure@28.[151,1436,331,1662]" captionTargetPageId="28" captionText="FIGURES 156 161. Male calling song at two different speeds (air temperature at recording shown; time scale below): 156 — I. miksici (BG: Gorski Dom Lodge); 157 — I. plevnensis (BG: Apriltsi); 158, 159 — I. longicaudata adamovici (BG: Karandila Locality, two males); 160 — I. longicaudata longicaudata (BG: Bolata Valley); 161 — I. modesta modesta (RO: Babadag). Break-signs in Fig. 161 B indicate the missing silent interval of the syllable shown in Fig. 161 A." httpUri="https://zenodo.org/record/246562/files/figure.png" pageId="54" pageNumber="55">Figs 158160</figureCitation>
): syllables of
<quantity id="BD3E2DDB393EFFA4B27B0F64FC069D09" box="[860,981,1383,1404]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="15.0" metricValueMin="4.0" pageId="54" pageNumber="55" unit="m" value="950.0" valueMax="1500.0" valueMin="400.0">4001500 m</quantity>
...........
<taxonomicName id="BDC6FBBD393EFFA4B5790F6BFB239D08" box="[1118,1264,1384,1405]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="longicaudata">
<emphasis id="48B25C2C393EFFA4B5790F6BFB239D08" box="[1118,1264,1384,1405]" italics="true" pageId="54" pageNumber="55">I. longicaudata</emphasis>
</taxonomicName>
(for ssp. see text)
</keyLead>
</paragraph>
<paragraph id="7A79803E393EFFA4B1B00F86FA489DC2" blockId="54.[151,1436,151,1992]" pageId="54" pageNumber="55">
<keyLead id="C137230B393EFFA4B1B00F86FA489DC2" pageId="54" pageNumber="55">
<emphasis id="48B25C2C393EFFA4B1C60F86FECF9DEF" box="[225,284,1413,1434]" italics="true" pageId="54" pageNumber="55">Males</emphasis>
: Stridulatory teeth 100145.
<emphasis id="48B25C2C393EFFA4B31D0F86FDB99DEF" box="[570,618,1413,1434]" italics="true" pageId="54" pageNumber="55">Song</emphasis>
(
<figureCitation id="E2FD9CBB393EFFA4B35F0F86FCD19DEF" box="[632,770,1413,1434]" captionStart="FIGURES 156 161" captionStartId="28.[151,264,1682,1705]" captionTargetBox="[151,1436,331,1661]" captionTargetId="figure@28.[151,1436,331,1662]" captionTargetPageId="28" captionText="FIGURES 156 161. Male calling song at two different speeds (air temperature at recording shown; time scale below): 156 — I. miksici (BG: Gorski Dom Lodge); 157 — I. plevnensis (BG: Apriltsi); 158, 159 — I. longicaudata adamovici (BG: Karandila Locality, two males); 160 — I. longicaudata longicaudata (BG: Bolata Valley); 161 — I. modesta modesta (RO: Babadag). Break-signs in Fig. 161 B indicate the missing silent interval of the syllable shown in Fig. 161 A." httpUri="https://zenodo.org/record/246562/files/figure.png" pageId="54" pageNumber="55">Figs 157, 161</figureCitation>
): syllables either short (up to 400 ms), either of two distinctly separated parts, together lasting over 5000 ms.................................................................. 15
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C132989B393EFFA4B1B00FBCFA4F9D87" pageId="54" pageNumber="55">
<paragraph id="7A79803E393EFFA4B1B00FBCFA4F9DA0" blockId="54.[151,1436,151,1992]" box="[151,1436,1471,1493]" pageId="54" pageNumber="55">
<keyLead id="C137230B393EFFA4B1B00FBCFA4F9DA0" box="[151,1436,1471,1493]" pageId="54" pageNumber="55">
15.
<emphasis id="48B25C2C393EFFA4B1C60FC3FEC29DA0" box="[225,273,1472,1493]" italics="true" pageId="54" pageNumber="55">Song</emphasis>
(
<figureCitation id="E2FD9CBB393EFFA4B0390FBCFEA09DA1" box="[286,371,1471,1492]" captionStart="FIGURES 156 161" captionStartId="28.[151,264,1682,1705]" captionTargetBox="[151,1436,331,1661]" captionTargetId="figure@28.[151,1436,331,1662]" captionTargetPageId="28" captionText="FIGURES 156 161. Male calling song at two different speeds (air temperature at recording shown; time scale below): 156 — I. miksici (BG: Gorski Dom Lodge); 157 — I. plevnensis (BG: Apriltsi); 158, 159 — I. longicaudata adamovici (BG: Karandila Locality, two males); 160 — I. longicaudata longicaudata (BG: Bolata Valley); 161 — I. modesta modesta (RO: Babadag). Break-signs in Fig. 161 B indicate the missing silent interval of the syllable shown in Fig. 161 A." httpUri="https://zenodo.org/record/246562/files/figure.png" pageId="54" pageNumber="55">Fig. 157</figureCitation>
): syllables compact, &lt;400 ms long. Central N
<collectingCountry id="02D1C0AE393EFFA4B2090FBCFC529DA1" box="[814,897,1471,1492]" name="Bulgaria" pageId="54" pageNumber="55">Bulgaria</collectingCountry>
...................................
<taxonomicName id="BDC6FBBD393EFFA4B4030FC3FA4F9DA0" box="[1316,1436,1472,1493]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="plevnensis">
<emphasis id="48B25C2C393EFFA4B4030FC3FA4F9DA0" box="[1316,1436,1472,1493]" italics="true" pageId="54" pageNumber="55">I. plevnensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7A79803E393EFFA4B1B00FDEFA4F9D87" blockId="54.[151,1436,151,1992]" box="[151,1436,1501,1522]" pageId="54" pageNumber="55">
<keyLead id="C137230B393EFFA4B1B00FDEFA4F9D87" box="[151,1436,1501,1522]" pageId="54" pageNumber="55">
<emphasis id="48B25C2C393EFFA4B1C60FDEFEC29D87" box="[225,273,1501,1522]" italics="true" pageId="54" pageNumber="55">Song</emphasis>
(
<figureCitation id="E2FD9CBB393EFFA4B0390FDEFEA19D87" box="[286,370,1501,1522]" captionStart="FIGURES 156 161" captionStartId="28.[151,264,1682,1705]" captionTargetBox="[151,1436,331,1661]" captionTargetId="figure@28.[151,1436,331,1662]" captionTargetPageId="28" captionText="FIGURES 156 161. Male calling song at two different speeds (air temperature at recording shown; time scale below): 156 — I. miksici (BG: Gorski Dom Lodge); 157 — I. plevnensis (BG: Apriltsi); 158, 159 — I. longicaudata adamovici (BG: Karandila Locality, two males); 160 — I. longicaudata longicaudata (BG: Bolata Valley); 161 — I. modesta modesta (RO: Babadag). Break-signs in Fig. 161 B indicate the missing silent interval of the syllable shown in Fig. 161 A." httpUri="https://zenodo.org/record/246562/files/figure.png" pageId="54" pageNumber="55">Fig. 161</figureCitation>
): syllables of two parts, 59 s long. N Dobrogea........................
<taxonomicName id="BDC6FBBD393EFFA4B5500FDEFB0F9D87" box="[1143,1244,1501,1522]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="modesta">
<emphasis id="48B25C2C393EFFA4B5500FDEFB0F9D87" box="[1143,1244,1501,1522]" italics="true" pageId="54" pageNumber="55">I. modesta</emphasis>
</taxonomicName>
(for ssp. see above)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C132989B393EFFA4B1B00FF9FA489ED7" pageId="54" pageNumber="55">
<paragraph id="7A79803E393EFFA4B1B00FF9FA4F9E3F" blockId="54.[151,1436,151,1992]" pageId="54" pageNumber="55">
<keyLead id="C137230B393EFFA4B1B00FF9FA4F9E3F" pageId="54" pageNumber="55">
16. Smaller species, hind femora without ventral spines.
<emphasis id="48B25C2C393EFFA4B3CC0FF9FCF69E7A" box="[747,805,1530,1551]" italics="true" pageId="54" pageNumber="55">Males</emphasis>
: tegmina small (
<figureCitation id="E2FD9CBB393EFFA4B2EB0FF9FBC29E7A" box="[972,1041,1530,1551]" captionStart="FIGURES 31 55" captionStartId="12.[151,264,244,267]" captionText="FIGURES 31 55. Morphology of male head, pronotum and tegmina of the Balkan Isophya and I. yaraligozi: 31 — I. hospodar (BG: Momchilgrad, 31.05. 1965, NMNHS); 32 — I. rectipennis (BG: Ropotamo, 7.07. 1965, NMNHS); 33 — I. pavelii (holotype of I. rammei Peshev, BG: Chernogorovo, 19.06. 1973, NMNHS); 34 — I. thracica (TR: Elmali, 3.07. 2011, CC); 35 — I. bureschi (BG: “ Pirin, 1300 1450 m ”, 17.08. 1961, NMNHS); 36 — I. yaraligozi (TR: Yaraligoz Pass, 9.07. 2011, CC); 37 — I. tosevski (MK: Moklishte, 30.05. 2010, CC); 38 — I. andreevae (holotype, BG: Kresna Gorge, 2.06. 1961, NMNHS); 39 — I. clara (BA: Sarajevo, NMW); 40 — I. miksici (holotype, BG: Berkovitsa, 700 m, 21.06. 1958, NMNHS); 41 — I. plevnensis (holotype, BG: Chernelka, 30.07. 1973, NMNHS); 42 — I. longicaudata adamovici (holotype, BG: “ Sliven ”, NMNHS); 43 — I. l. longicaudata (BG: Batova Gora, 26.06. 1956, NMNHS); 44 — I. m. modesta (= intermedia) (RO: Dobrogea, Babadag, CI); 45 — I. rhodopensis leonorae (BG: Popovi Livadi, 1300 m, 4.07. 1961, NMNHS); 46 — I. rh. rhodopensis (BG: Chepelare, 16.07. 1956, NMNHS); 47 — I. rh. petkovi (BG: Mandritsa, 4.06. 1965, NMNHS); 48 — I. modestior (BG: Kopitoto, 17.07. 1958, NMNHS); 49 — I. dobrogensis (RO: Popina Island, CI); 50 — I. zubowskii (RO: “ Jasi pād, Birnari, 25.06. 1956, NMNHS); 51 — I. aff. camptoxypha (“ Montenegro ”, NMW); 52 — I. gulae (BG: Dolna Topchiya, 23.06. 2008, NMNHS); 53 — I. obtusa (BG: Vezhen Peak, 1900 m, 1.09. 1960, NMNHS); 54 — I. amplipennis (TR: “ Bursa | Gursu, | Kazanci &amp; | Eskibaglar | 14. vi. 1959 ”, “ by Dr. E. Can. | B. M. 1960 173 ”, NHM); 55 — I. speciosa (BG: Plevoun, 5.06. 1965, NMNHS)." pageId="54" pageNumber="55">Fig. 51</figureCitation>
)—shorter and narrower than pronotum; stridulatory teeth (
<figureCitation id="E2FD9CBB393EFFA4B0B50C14FE309E59" box="[402,483,1559,1580]" captionStart="FIGURES 174 181" captionStartId="36.[151,264,1078,1101]" captionTargetBox="[151,1436,193,1056]" captionTargetId="figure@36.[151,1436,193,1057]" captionTargetPageId="36" captionText="FIGURES 174 181. SEM of taxonomically important structures of Isophya: 174 — I. dobrogensis (RO: Popina Island); 175 — I. zubowskii (RO: Babadag, NMB); 176 — I. camptoxypha (RO: “ Bucec b. Sinaia, 13 1800 m, 25.8. 41, Ramme ”, NMB); 177 — I. gulae (BG: Elhovo, Dolna Topchiya Resrve); 178, 179 — I. obtusa (178 — BG: Lyulin Mt.; 179 — Pleven Lodge); 180 — I. amplipennis (TR: near Sogut); 181 — I. speciosa (BG: Kovach Locality). A—male stridulatory file (174 179 — scale 200 μm; 180, 181 — scale 100 μm); B—ventral / ventro-apical view of apex of male cerci (scale 100 μm); C—female stridulatory apparatus (scale 100 μm; for 175 C— 200 μm)." httpUri="https://zenodo.org/record/246565/files/figure.png" pageId="54" pageNumber="55">Fig. 176</figureCitation>
A) &lt;90; subgenital plate with deep excision.
<emphasis id="48B25C2C393EFFA4B2B20C1BFC359E58" box="[917,998,1560,1581]" italics="true" pageId="54" pageNumber="55">Females</emphasis>
: subgenital plate with apical elongation; ovipositor ±2 times the length of pronotum.
<emphasis id="48B25C2C393EFFA4B34C0C36FD489E3F" box="[619,667,1589,1610]" italics="true" pageId="54" pageNumber="55">Song</emphasis>
(
<figureCitation id="E2FD9CBB393EFFA4B38F0C36FD299E3F" box="[680,762,1589,1610]" captionStart="FIGURES 182 189" captionStartId="38.[151,264,1858,1881]" captionTargetBox="[151,1436,555,1837]" captionTargetId="figure@38.[151,1436,555,1837]" captionTargetPageId="38" captionText="FIGURES 182 189. Male calling song at two different speeds (air temperature at recording shown; time scale below): 182, 183 — I. modestior (182 — BG: Vitosha Mt.; 183 — Serbia: Novi Sad); 184 — I. dobrogensis (RO: Popina Island); 185 — I. zubowskii (RO: Dobrogea, Niculiţel); 186 — I. camptoxypha (RO: Ciucaş Mt.); 187 — I. gulae (BG: Elhovo, Dolna Topchiya Reserve); 188, 189 — I. obtusa (188 — Middle Stara Planina Mts, Pleven Lodge; 189 — BG: Vitosha Mt.). Arrow and a “ Ƥ ” symbol on fig. 187 indicate the response of a female I. gulae." httpUri="https://zenodo.org/record/246566/files/figure.png" pageId="54" pageNumber="55">Fig. 186</figureCitation>
) consists of continuous sequence of syllables........
<taxonomicName id="BDC6FBBD393EFFA4B4220C36FA4F9E3F" box="[1285,1436,1589,1610]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="camptoxypha">
<emphasis id="48B25C2C393EFFA4B4220C36FA4F9E3F" box="[1285,1436,1589,1610]" italics="true" pageId="54" pageNumber="55">I. camptoxypha</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7A79803E393EFFA4B1B00C51FA489ED7" blockId="54.[151,1436,151,1992]" pageId="54" pageNumber="55">
<keyLead id="C137230B393EFFA4B1B00C51FA489ED7" pageId="54" pageNumber="55">
Larger species, hind femora frequently with 12 ventral spines.
<emphasis id="48B25C2C393EFFA4B26A0C51FC549E12" box="[845,903,1618,1639]" italics="true" pageId="54" pageNumber="55">Males</emphasis>
: tegmina large, strongly bulged—as wide as and equal or longer than pronotum; stridulatory teeth&gt;100; subgenital plate with shallow excision.
<emphasis id="48B25C2C393EFFA4B5760C73FB719EF0" box="[1105,1186,1648,1669]" italics="true" pageId="54" pageNumber="55">Females</emphasis>
: subgenital plate without apical elongation; ovipositor&gt;2.3 times the length of pronotum.
<emphasis id="48B25C2C393EFFA4B26A0C8EFCAE9ED7" box="[845,893,1677,1698]" italics="true" pageId="54" pageNumber="55">Song</emphasis>
consists of groups of or isolated syllables......... 17
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<keyStep id="C132989B393EFFA4B1B00CA9FA4F9F27" pageId="54" pageNumber="55">
<paragraph id="7A79803E393EFFA4B1B00CA9FA469E8F" blockId="54.[151,1436,151,1992]" pageId="54" pageNumber="55">
<keyLead id="C137230B393EFFA4B1B00CA9FA469E8F" pageId="54" pageNumber="55">
17.
<emphasis id="48B25C2C393EFFA4B1C60CA9FECF9ECA" box="[225,284,1706,1727]" italics="true" pageId="54" pageNumber="55">Males</emphasis>
: Body colouration dark-green; pronotum less saddle-shaped; tegmina (
<figureCitation id="E2FD9CBB393EFFA4B2F70CA9FBCB9ECA" box="[976,1048,1706,1727]" captionStart="FIGURES 31 55" captionStartId="12.[151,264,244,267]" captionText="FIGURES 31 55. Morphology of male head, pronotum and tegmina of the Balkan Isophya and I. yaraligozi: 31 — I. hospodar (BG: Momchilgrad, 31.05. 1965, NMNHS); 32 — I. rectipennis (BG: Ropotamo, 7.07. 1965, NMNHS); 33 — I. pavelii (holotype of I. rammei Peshev, BG: Chernogorovo, 19.06. 1973, NMNHS); 34 — I. thracica (TR: Elmali, 3.07. 2011, CC); 35 — I. bureschi (BG: “ Pirin, 1300 1450 m ”, 17.08. 1961, NMNHS); 36 — I. yaraligozi (TR: Yaraligoz Pass, 9.07. 2011, CC); 37 — I. tosevski (MK: Moklishte, 30.05. 2010, CC); 38 — I. andreevae (holotype, BG: Kresna Gorge, 2.06. 1961, NMNHS); 39 — I. clara (BA: Sarajevo, NMW); 40 — I. miksici (holotype, BG: Berkovitsa, 700 m, 21.06. 1958, NMNHS); 41 — I. plevnensis (holotype, BG: Chernelka, 30.07. 1973, NMNHS); 42 — I. longicaudata adamovici (holotype, BG: “ Sliven ”, NMNHS); 43 — I. l. longicaudata (BG: Batova Gora, 26.06. 1956, NMNHS); 44 — I. m. modesta (= intermedia) (RO: Dobrogea, Babadag, CI); 45 — I. rhodopensis leonorae (BG: Popovi Livadi, 1300 m, 4.07. 1961, NMNHS); 46 — I. rh. rhodopensis (BG: Chepelare, 16.07. 1956, NMNHS); 47 — I. rh. petkovi (BG: Mandritsa, 4.06. 1965, NMNHS); 48 — I. modestior (BG: Kopitoto, 17.07. 1958, NMNHS); 49 — I. dobrogensis (RO: Popina Island, CI); 50 — I. zubowskii (RO: “ Jasi pād, Birnari, 25.06. 1956, NMNHS); 51 — I. aff. camptoxypha (“ Montenegro ”, NMW); 52 — I. gulae (BG: Dolna Topchiya, 23.06. 2008, NMNHS); 53 — I. obtusa (BG: Vezhen Peak, 1900 m, 1.09. 1960, NMNHS); 54 — I. amplipennis (TR: “ Bursa | Gursu, | Kazanci &amp; | Eskibaglar | 14. vi. 1959 ”, “ by Dr. E. Can. | B. M. 1960 173 ”, NHM); 55 — I. speciosa (BG: Plevoun, 5.06. 1965, NMNHS)." pageId="54" pageNumber="55">Fig. 52</figureCitation>
) narrower; tip of cerci (
<figureCitation id="E2FD9CBB393EFFA4B4230CA9FA849ECA" box="[1284,1367,1706,1727]" captionStart="FIGURES 174 181" captionStartId="36.[151,264,1078,1101]" captionTargetBox="[151,1436,193,1056]" captionTargetId="figure@36.[151,1436,193,1057]" captionTargetPageId="36" captionText="FIGURES 174 181. SEM of taxonomically important structures of Isophya: 174 — I. dobrogensis (RO: Popina Island); 175 — I. zubowskii (RO: Babadag, NMB); 176 — I. camptoxypha (RO: “ Bucec b. Sinaia, 13 1800 m, 25.8. 41, Ramme ”, NMB); 177 — I. gulae (BG: Elhovo, Dolna Topchiya Resrve); 178, 179 — I. obtusa (178 — BG: Lyulin Mt.; 179 — Pleven Lodge); 180 — I. amplipennis (TR: near Sogut); 181 — I. speciosa (BG: Kovach Locality). A—male stridulatory file (174 179 — scale 200 μm; 180, 181 — scale 100 μm); B—ventral / ventro-apical view of apex of male cerci (scale 100 μm); C—female stridulatory apparatus (scale 100 μm; for 175 C— 200 μm)." httpUri="https://zenodo.org/record/246565/files/figure.png" pageId="54" pageNumber="55">Fig. 177</figureCitation>
B) narrow, having few hairs and a long tooth.
<emphasis id="48B25C2C393EFFA4B34F0CCBFD4B9EA8" box="[616,664,1736,1757]" italics="true" pageId="54" pageNumber="55">Song</emphasis>
(
<figureCitation id="E2FD9CBB393EFFA4B3810CC4FD299EA9" box="[678,762,1735,1756]" captionStart="FIGURES 182 189" captionStartId="38.[151,264,1858,1881]" captionTargetBox="[151,1436,555,1837]" captionTargetId="figure@38.[151,1436,555,1837]" captionTargetPageId="38" captionText="FIGURES 182 189. Male calling song at two different speeds (air temperature at recording shown; time scale below): 182, 183 — I. modestior (182 — BG: Vitosha Mt.; 183 — Serbia: Novi Sad); 184 — I. dobrogensis (RO: Popina Island); 185 — I. zubowskii (RO: Dobrogea, Niculiţel); 186 — I. camptoxypha (RO: Ciucaş Mt.); 187 — I. gulae (BG: Elhovo, Dolna Topchiya Reserve); 188, 189 — I. obtusa (188 — Middle Stara Planina Mts, Pleven Lodge; 189 — BG: Vitosha Mt.). Arrow and a “ Ƥ ” symbol on fig. 187 indicate the response of a female I. gulae." httpUri="https://zenodo.org/record/246566/files/figure.png" pageId="54" pageNumber="55">Fig. 187</figureCitation>
) of isolated syllables typically separated in a shorter main part and a second part of 1520 after-clicks altogether lasting 14002500 ms.........................................
<taxonomicName id="BDC6FBBD393EFFA4B46B0CE6FA469E8F" box="[1356,1429,1765,1786]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="gulae">
<emphasis id="48B25C2C393EFFA4B46B0CE6FA469E8F" box="[1356,1429,1765,1786]" italics="true" pageId="54" pageNumber="55">I. gulae</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7A79803E393EFFA4B1B00D01FA4F9F27" blockId="54.[151,1436,151,1992]" pageId="54" pageNumber="55">
<keyLead id="C137230B393EFFA4B1B00D01FA4F9F27" pageId="54" pageNumber="55">
<emphasis id="48B25C2C393EFFA4B1C60D01FECF9F62" box="[225,284,1794,1815]" italics="true" pageId="54" pageNumber="55">Males</emphasis>
: Body colouration dark-green to violet-green; pronotum (
<figureCitation id="E2FD9CBB393EFFA4B26A0D01FC409F62" box="[845,915,1794,1815]" captionStart="FIGURES 31 55" captionStartId="12.[151,264,244,267]" captionText="FIGURES 31 55. Morphology of male head, pronotum and tegmina of the Balkan Isophya and I. yaraligozi: 31 — I. hospodar (BG: Momchilgrad, 31.05. 1965, NMNHS); 32 — I. rectipennis (BG: Ropotamo, 7.07. 1965, NMNHS); 33 — I. pavelii (holotype of I. rammei Peshev, BG: Chernogorovo, 19.06. 1973, NMNHS); 34 — I. thracica (TR: Elmali, 3.07. 2011, CC); 35 — I. bureschi (BG: “ Pirin, 1300 1450 m ”, 17.08. 1961, NMNHS); 36 — I. yaraligozi (TR: Yaraligoz Pass, 9.07. 2011, CC); 37 — I. tosevski (MK: Moklishte, 30.05. 2010, CC); 38 — I. andreevae (holotype, BG: Kresna Gorge, 2.06. 1961, NMNHS); 39 — I. clara (BA: Sarajevo, NMW); 40 — I. miksici (holotype, BG: Berkovitsa, 700 m, 21.06. 1958, NMNHS); 41 — I. plevnensis (holotype, BG: Chernelka, 30.07. 1973, NMNHS); 42 — I. longicaudata adamovici (holotype, BG: “ Sliven ”, NMNHS); 43 — I. l. longicaudata (BG: Batova Gora, 26.06. 1956, NMNHS); 44 — I. m. modesta (= intermedia) (RO: Dobrogea, Babadag, CI); 45 — I. rhodopensis leonorae (BG: Popovi Livadi, 1300 m, 4.07. 1961, NMNHS); 46 — I. rh. rhodopensis (BG: Chepelare, 16.07. 1956, NMNHS); 47 — I. rh. petkovi (BG: Mandritsa, 4.06. 1965, NMNHS); 48 — I. modestior (BG: Kopitoto, 17.07. 1958, NMNHS); 49 — I. dobrogensis (RO: Popina Island, CI); 50 — I. zubowskii (RO: “ Jasi pād, Birnari, 25.06. 1956, NMNHS); 51 — I. aff. camptoxypha (“ Montenegro ”, NMW); 52 — I. gulae (BG: Dolna Topchiya, 23.06. 2008, NMNHS); 53 — I. obtusa (BG: Vezhen Peak, 1900 m, 1.09. 1960, NMNHS); 54 — I. amplipennis (TR: “ Bursa | Gursu, | Kazanci &amp; | Eskibaglar | 14. vi. 1959 ”, “ by Dr. E. Can. | B. M. 1960 173 ”, NHM); 55 — I. speciosa (BG: Plevoun, 5.06. 1965, NMNHS)." pageId="54" pageNumber="55">Fig. 53</figureCitation>
) strongly saddle-shaped; tegmina very wide, strongly bulged; tip of cerci (
<figureCitation id="E2FD9CBB393EFFA4B0850D1CFE209F41" box="[418,499,1823,1844]" captionStart="FIGURES 174 181" captionStartId="36.[151,264,1078,1101]" captionTargetBox="[151,1436,193,1056]" captionTargetId="figure@36.[151,1436,193,1057]" captionTargetPageId="36" captionText="FIGURES 174 181. SEM of taxonomically important structures of Isophya: 174 — I. dobrogensis (RO: Popina Island); 175 — I. zubowskii (RO: Babadag, NMB); 176 — I. camptoxypha (RO: “ Bucec b. Sinaia, 13 1800 m, 25.8. 41, Ramme ”, NMB); 177 — I. gulae (BG: Elhovo, Dolna Topchiya Resrve); 178, 179 — I. obtusa (178 — BG: Lyulin Mt.; 179 — Pleven Lodge); 180 — I. amplipennis (TR: near Sogut); 181 — I. speciosa (BG: Kovach Locality). A—male stridulatory file (174 179 — scale 200 μm; 180, 181 — scale 100 μm); B—ventral / ventro-apical view of apex of male cerci (scale 100 μm); C—female stridulatory apparatus (scale 100 μm; for 175 C— 200 μm)." httpUri="https://zenodo.org/record/246565/files/figure.png" pageId="54" pageNumber="55">Fig. 178</figureCitation>
B) very wide, obtuse, bearing many hairs and a very short tooth.
<emphasis id="48B25C2C393EFFA4B57C0D23FB589F40" box="[1115,1163,1824,1845]" italics="true" pageId="54" pageNumber="55">Song</emphasis>
(
<figureCitation id="E2FD9CBB393EFFA4B5B00D1CFAC89F41" box="[1175,1307,1823,1844]" captionStart="FIGURES 182 189" captionStartId="38.[151,264,1858,1881]" captionTargetBox="[151,1436,555,1837]" captionTargetId="figure@38.[151,1436,555,1837]" captionTargetPageId="38" captionText="FIGURES 182 189. Male calling song at two different speeds (air temperature at recording shown; time scale below): 182, 183 — I. modestior (182 — BG: Vitosha Mt.; 183 — Serbia: Novi Sad); 184 — I. dobrogensis (RO: Popina Island); 185 — I. zubowskii (RO: Dobrogea, Niculiţel); 186 — I. camptoxypha (RO: Ciucaş Mt.); 187 — I. gulae (BG: Elhovo, Dolna Topchiya Reserve); 188, 189 — I. obtusa (188 — Middle Stara Planina Mts, Pleven Lodge; 189 — BG: Vitosha Mt.). Arrow and a “ Ƥ ” symbol on fig. 187 indicate the response of a female I. gulae." httpUri="https://zenodo.org/record/246566/files/figure.png" pageId="54" pageNumber="55">Figs 188, 189</figureCitation>
) of groups of syllables lasting 100500 ms with few or without after-clicks............................................
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<emphasis id="48B25C2C393EFFA4B46F0D3EFA4F9F27" box="[1352,1436,1853,1874]" italics="true" pageId="54" pageNumber="55">I. obtusa</emphasis>
</taxonomicName>
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</paragraph>
</keyStep>
<paragraph id="7A79803E393EFFA4B1B00D59FA4F9FDF" blockId="54.[151,1436,151,1992]" pageId="54" pageNumber="55">
<keyLead id="C137230B393EFFA4B1B00D59FA4F9FDF" pageId="54" pageNumber="55">
<keyStep id="C132989B393EFFA4B1B00D59FA4F9FDF" pageId="54" pageNumber="55">
18.
<collectingCountry id="02D1C0AE393EFFA4B1C60D59FEC29F1A" box="[225,273,1882,1903]" name="India" pageId="54" pageNumber="55">Hind</collectingCountry>
femora always without ventral spines.
<emphasis id="48B25C2C393EFFA4B3A80D59FD199F1A" box="[655,714,1882,1903]" italics="true" pageId="54" pageNumber="55">Males</emphasis>
: Cerci slender, tip of cerci attenuated or almost pointed, bearing an apical tooth.
<emphasis id="48B25C2C393EFFA4B0390D7BFEBC9FF8" box="[286,367,1912,1933]" italics="true" pageId="54" pageNumber="55">Females</emphasis>
: subgenital plate rounded; lateral pit at the base of ovipositor almost closed by the upcurved lamella; ovipositor very short (&lt;
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), less than 2 times longer than pronotum..........................................
<taxonomicName id="BDC6FBBD393EFFA4B43A0D96FA4F9FDF" box="[1309,1436,1941,1962]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="brevicauda">
<emphasis id="48B25C2C393EFFA4B43A0D96FA4F9FDF" box="[1309,1436,1941,1962]" italics="true" pageId="54" pageNumber="55">I. brevicauda</emphasis>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="7A79803E393EFFA4B1B00DB1FA499FB2" blockId="54.[151,1436,151,1992]" box="[151,1434,1970,1992]" pageId="54" pageNumber="55">
<keyLead id="C137230B393EFFA4B1B00DB1FA499FB2" box="[151,1434,1970,1992]" pageId="54" pageNumber="55">
<collectingCountry id="02D1C0AE393EFFA4B1C60DB1FEC29FB2" box="[225,273,1970,1991]" name="India" pageId="54" pageNumber="55">Hind</collectingCountry>
femora without or with few ventral spines.
<emphasis id="48B25C2C393EFFA4B3920DB0FD3C9FBD" box="[693,751,1971,1992]" italics="true" pageId="54" pageNumber="55">Males</emphasis>
: Cerci stout, tip of cerci widely blunt or cut, bearing very short
<emphasis id="48B25C2C393EFFA4B4730DB1FAB99FB2" bold="true" box="[1364,1386,1970,1991]" pageId="54" pageNumber="55">or</emphasis>
sub-
</keyLead>
</paragraph>
<paragraph id="7A79803E393FFFA5B1C60A94FA4F994A" blockId="55.[151,1437,151,730]" pageId="55" pageNumber="56">
apical tooth.
<emphasis id="48B25C2C393FFFA5B07A0A9BFE7D98D8" box="[349,430,152,173]" italics="true" pageId="55" pageNumber="56">Females</emphasis>
: subgenital plate with an apical elongation; lateral pit at the base of ovipositor widely opened; ovipositor longer than
<quantity id="BD3E2DDB393FFFA5B0740AB6FE5E98BF" box="[339,397,181,202]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="55" pageNumber="56" unit="mm" value="9.0">9 mm</quantity>
, over 2 times longer than pronotum........................................................ 19
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<keyStep id="C132989B393FFFA5B1B00AD1FA4F994A" pageId="55" pageNumber="56">
19. Fastigium of vertex about 1/3 the width of scapus.
<emphasis id="48B25C2C393FFFA5B3EE0AD1FCD09892" box="[713,771,210,231]" italics="true" pageId="55" pageNumber="56">Males</emphasis>
: CuP thinner, about as wide as second antennal limb; pronotum (
<figureCitation id="E2FD9CBB393FFFA5B4520AD1FF299971" captionStart="FIGURES 31 55" captionStartId="12.[151,264,244,267]" captionText="FIGURES 31 55. Morphology of male head, pronotum and tegmina of the Balkan Isophya and I. yaraligozi: 31 — I. hospodar (BG: Momchilgrad, 31.05. 1965, NMNHS); 32 — I. rectipennis (BG: Ropotamo, 7.07. 1965, NMNHS); 33 — I. pavelii (holotype of I. rammei Peshev, BG: Chernogorovo, 19.06. 1973, NMNHS); 34 — I. thracica (TR: Elmali, 3.07. 2011, CC); 35 — I. bureschi (BG: “ Pirin, 1300 1450 m ”, 17.08. 1961, NMNHS); 36 — I. yaraligozi (TR: Yaraligoz Pass, 9.07. 2011, CC); 37 — I. tosevski (MK: Moklishte, 30.05. 2010, CC); 38 — I. andreevae (holotype, BG: Kresna Gorge, 2.06. 1961, NMNHS); 39 — I. clara (BA: Sarajevo, NMW); 40 — I. miksici (holotype, BG: Berkovitsa, 700 m, 21.06. 1958, NMNHS); 41 — I. plevnensis (holotype, BG: Chernelka, 30.07. 1973, NMNHS); 42 — I. longicaudata adamovici (holotype, BG: “ Sliven ”, NMNHS); 43 — I. l. longicaudata (BG: Batova Gora, 26.06. 1956, NMNHS); 44 — I. m. modesta (= intermedia) (RO: Dobrogea, Babadag, CI); 45 — I. rhodopensis leonorae (BG: Popovi Livadi, 1300 m, 4.07. 1961, NMNHS); 46 — I. rh. rhodopensis (BG: Chepelare, 16.07. 1956, NMNHS); 47 — I. rh. petkovi (BG: Mandritsa, 4.06. 1965, NMNHS); 48 — I. modestior (BG: Kopitoto, 17.07. 1958, NMNHS); 49 — I. dobrogensis (RO: Popina Island, CI); 50 — I. zubowskii (RO: “ Jasi pād, Birnari, 25.06. 1956, NMNHS); 51 — I. aff. camptoxypha (“ Montenegro ”, NMW); 52 — I. gulae (BG: Dolna Topchiya, 23.06. 2008, NMNHS); 53 — I. obtusa (BG: Vezhen Peak, 1900 m, 1.09. 1960, NMNHS); 54 — I. amplipennis (TR: “ Bursa | Gursu, | Kazanci &amp; | Eskibaglar | 14. vi. 1959 ”, “ by Dr. E. Can. | B. M. 1960 173 ”, NHM); 55 — I. speciosa (BG: Plevoun, 5.06. 1965, NMNHS)." pageId="55" pageNumber="56">Fig. 50</figureCitation>
) short, saddle-shaped, with strongly sinuate side borders of metazone; cerci (
<figureCitation id="E2FD9CBB393FFFA5B2C60AECFBF49971" box="[993,1063,239,260]" captionStart="FIGURES 80 104" captionStartId="14.[151,264,1835,1858]" captionTargetBox="[226,1360,681,1815]" captionTargetId="figure@14.[226,1362,681,1816]" captionTargetPageId="14" captionText="FIGURES 80 104. Male abdominal terminalia of the Balkan Isophya and I. yaraligozi (localities as in Figs 31 55): 80 — I. hospodar; 81 — I. rectipennis; 82 — I. pavelii; 83 — I. thracica; 84 — I. bureschi; 85 — I. yaraligozi; 86 — I. tosevski; 87 — I. andreevae; 88 — I. clara; 89 — I. miksici; 90 — I. plevnensis; 91 — I. longicaudata adamovici; 92 — I. l. longicaudata; 93 — I. m. modesta (= intermedia); 94 — I. rhodopensis leonorae; 95 — I. rh. rhodopensis; 96 — I. rh. petkovi; 97 — I. modestior; 98 — I. dobrogensis; 99 — I. zubowskii; 100 — I. aff. camptoxypha; 101 — I. gulae; 102 — I. obtusa; 103 — I. amplipennis; 104 — I. speciosa. Scale (if present) = 2 mm." httpUri="https://zenodo.org/record/246555/files/figure.png" pageId="55" pageNumber="56">Fig. 99</figureCitation>
) slightly incurved, their base not more than twice the diameter of the apex; apex of cerci slightly widened.
<emphasis id="48B25C2C393FFFA5B25D0B0EFC189957" box="[890,971,269,290]" italics="true" pageId="55" pageNumber="56">Females</emphasis>
: ovipositor slender,
<quantity id="BD3E2DDB393FFFA5B5B40B0EFB289957" box="[1171,1275,269,290]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="55" pageNumber="56" unit="mm" value="13.0" valueMax="14.0" valueMin="12.0">1214 mm</quantity>
long (
<figureCitation id="E2FD9CBB393FFFA5B41B0B0EFA429957" box="[1340,1425,269,290]" captionStart="FIGURES 105 129" captionStartId="17.[151,264,1173,1196]" captionTargetBox="[151,1436,195,1152]" captionTargetId="figure@17.[151,1436,195,1152]" captionTargetPageId="17" captionText="FIGURES 105 129. Female abdominal terminalia of the Balkan Isophya and I. yaraligozi (localities as in Figs 31 55 unless specifically given); figures of the whole ovipositor are followed by enlarged figure of the ovipositor base: 105 — I. hospodar (BG: Dolni Glavanak, 25.04. 2005, CC); 106 — I. rectipennis (BG: Byala Vill., 27.06. 2002, CC); 107 — I. pavelii (BG: Chernogorovo, 29.05. 2006, CC); 108 — I. thracica; 109 — I. bureschi (BG: Bansko, 8.08. 2006, CC); 110 — I. yaraligozi; 111 — I. tosevski; 112 — I. andreevae (BG: Kresnensko Hanche, 19.04. 2006, CC); 113 — I. clara; 114 — I. miksici (BG: Gorski Dom Lodge, 24.06. 2006, CC); 115 — I. plevnensis (BG: Levishte, 14.07. 2009, CC); 116 — I. longicaudata adamovici (BG: Karandila, 24.06. 2008, CC); 117 — I. l. longicaudata (BG: Sofia University Botanical garden, 26.06. 1.07.2004, CC); 118 — I. m. modesta (= intermedia); 119 — I. rhodopensis leonorae (BG: Livade, 9.08. 2006, CC); 120 — I. rh. rhodopensis (BG: Bachkovo, 600 m, 23.05. 2004, CC); 121 — I. rh. petkovi (BG: Gluhite Kamani, 23.06. 2008, CC); 122 — I. modestior (BG: Bankya, 24.05. 2006, CC); 123 — I. dobrogensis; 124 — I. zubowskii; 125 — I. aff. camptoxypha (data as in Fig. 75); 126 — I. gulae; 127 — I. obtusa (BG: Ravnets Ridge, 14.07. 2009, CC); 128 — I. amplipennis (as in Fig. 78); 129 — I. speciosa (BG: Bankya, 24.05. 2006, CC). Indications on Fig. 127 show, respectively: G—gonangulum; L—lamella; P—lateral pit between lamella and gonangulum. Scale (if present) = 5 mm." httpUri="https://zenodo.org/record/246556/files/figure.png" pageId="55" pageNumber="56">Fig. 124</figureCitation>
).
<emphasis id="48B25C2C393FFFA5B1C60B29FEC2994A" box="[225,273,298,319]" italics="true" pageId="55" pageNumber="56">Song</emphasis>
(
<figureCitation id="E2FD9CBB393FFFA5B0390B29FEA3994A" box="[286,368,298,319]" captionStart="FIGURES 182 189" captionStartId="38.[151,264,1858,1881]" captionTargetBox="[151,1436,555,1837]" captionTargetId="figure@38.[151,1436,555,1837]" captionTargetPageId="38" captionText="FIGURES 182 189. Male calling song at two different speeds (air temperature at recording shown; time scale below): 182, 183 — I. modestior (182 — BG: Vitosha Mt.; 183 — Serbia: Novi Sad); 184 — I. dobrogensis (RO: Popina Island); 185 — I. zubowskii (RO: Dobrogea, Niculiţel); 186 — I. camptoxypha (RO: Ciucaş Mt.); 187 — I. gulae (BG: Elhovo, Dolna Topchiya Reserve); 188, 189 — I. obtusa (188 — Middle Stara Planina Mts, Pleven Lodge; 189 — BG: Vitosha Mt.). Arrow and a “ Ƥ ” symbol on fig. 187 indicate the response of a female I. gulae." httpUri="https://zenodo.org/record/246566/files/figure.png" pageId="55" pageNumber="56">Fig. 185</figureCitation>
) consists of long series of syllables...................................................
<taxonomicName id="BDC6FBBD393FFFA5B40E0B29FA4F994A" box="[1321,1436,298,319]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="zubowskii">
<emphasis id="48B25C2C393FFFA5B40E0B29FA4F994A" box="[1321,1436,298,319]" italics="true" pageId="55" pageNumber="56">I. zubowskii</emphasis>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="7A79803E393FFFA5B1B00B44FA4F99C1" blockId="55.[151,1437,151,730]" pageId="55" pageNumber="56">
<keyLead id="C137230B393FFFA5B1B00B44FA4F99C1" pageId="55" pageNumber="56">
Fastigium of vertex 2/3 to almost equal to the width of scapus.
<emphasis id="48B25C2C393FFFA5B2710B4BFC439928" box="[854,912,328,349]" italics="true" pageId="55" pageNumber="56">Males</emphasis>
: CuP wider that the second antennal limb; pronotum long, not saddle-shaped but metazone much wider than prozone; cerci strongly incurved in their apical third, their base more than twice the diameter of the apex; apex of cerci not widened, blunt or cut.
<emphasis id="48B25C2C393FFFA5B2990B81FBDC99E2" box="[958,1039,386,407]" italics="true" pageId="55" pageNumber="56">Females</emphasis>
: ovipositor wider (
<figureCitation id="E2FD9CBB393FFFA5B5E00B81FA9C99E2" box="[1223,1359,386,407]" captionStart="FIGURES 105 129" captionStartId="17.[151,264,1173,1196]" captionTargetBox="[151,1436,195,1152]" captionTargetId="figure@17.[151,1436,195,1152]" captionTargetPageId="17" captionText="FIGURES 105 129. Female abdominal terminalia of the Balkan Isophya and I. yaraligozi (localities as in Figs 31 55 unless specifically given); figures of the whole ovipositor are followed by enlarged figure of the ovipositor base: 105 — I. hospodar (BG: Dolni Glavanak, 25.04. 2005, CC); 106 — I. rectipennis (BG: Byala Vill., 27.06. 2002, CC); 107 — I. pavelii (BG: Chernogorovo, 29.05. 2006, CC); 108 — I. thracica; 109 — I. bureschi (BG: Bansko, 8.08. 2006, CC); 110 — I. yaraligozi; 111 — I. tosevski; 112 — I. andreevae (BG: Kresnensko Hanche, 19.04. 2006, CC); 113 — I. clara; 114 — I. miksici (BG: Gorski Dom Lodge, 24.06. 2006, CC); 115 — I. plevnensis (BG: Levishte, 14.07. 2009, CC); 116 — I. longicaudata adamovici (BG: Karandila, 24.06. 2008, CC); 117 — I. l. longicaudata (BG: Sofia University Botanical garden, 26.06. 1.07.2004, CC); 118 — I. m. modesta (= intermedia); 119 — I. rhodopensis leonorae (BG: Livade, 9.08. 2006, CC); 120 — I. rh. rhodopensis (BG: Bachkovo, 600 m, 23.05. 2004, CC); 121 — I. rh. petkovi (BG: Gluhite Kamani, 23.06. 2008, CC); 122 — I. modestior (BG: Bankya, 24.05. 2006, CC); 123 — I. dobrogensis; 124 — I. zubowskii; 125 — I. aff. camptoxypha (data as in Fig. 75); 126 — I. gulae; 127 — I. obtusa (BG: Ravnets Ridge, 14.07. 2009, CC); 128 — I. amplipennis (as in Fig. 78); 129 — I. speciosa (BG: Bankya, 24.05. 2006, CC). Indications on Fig. 127 show, respectively: G—gonangulum; L—lamella; P—lateral pit between lamella and gonangulum. Scale (if present) = 5 mm." httpUri="https://zenodo.org/record/246556/files/figure.png" pageId="55" pageNumber="56">Figs 122123</figureCitation>
),
<quantity id="BD3E2DDB393FFFA5B4470B81FED599C1" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="55" pageNumber="56" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">1013 mm</quantity>
long.
<emphasis id="48B25C2C393FFFA5B0640BA3FEA099C0" box="[323,371,416,437]" italics="true" pageId="55" pageNumber="56">Song</emphasis>
(
<figureCitation id="E2FD9CBB393FFFA5B0580B9CFDD699C1" box="[383,517,415,436]" captionStart="FIGURES 182 189" captionStartId="38.[151,264,1858,1881]" captionTargetBox="[151,1436,555,1837]" captionTargetId="figure@38.[151,1436,555,1837]" captionTargetPageId="38" captionText="FIGURES 182 189. Male calling song at two different speeds (air temperature at recording shown; time scale below): 182, 183 — I. modestior (182 — BG: Vitosha Mt.; 183 — Serbia: Novi Sad); 184 — I. dobrogensis (RO: Popina Island); 185 — I. zubowskii (RO: Dobrogea, Niculiţel); 186 — I. camptoxypha (RO: Ciucaş Mt.); 187 — I. gulae (BG: Elhovo, Dolna Topchiya Reserve); 188, 189 — I. obtusa (188 — Middle Stara Planina Mts, Pleven Lodge; 189 — BG: Vitosha Mt.). Arrow and a “ Ƥ ” symbol on fig. 187 indicate the response of a female I. gulae." httpUri="https://zenodo.org/record/246566/files/figure.png" pageId="55" pageNumber="56">Figs 182184</figureCitation>
) consists of loose groups of syllables............................................. 20
</keyLead>
</paragraph>
<keyStep id="C132989B393FFFA5B1B00BBEFA489A10" pageId="55" pageNumber="56">
<paragraph id="7A79803E393FFFA5B1B00BBEFA4F9A78" blockId="55.[151,1437,151,730]" pageId="55" pageNumber="56">
<keyLead id="C137230B393FFFA5B1B00BBEFA4F9A78" pageId="55" pageNumber="56">
20.
<emphasis id="48B25C2C393FFFA5B1C60BBEFECF99A7" box="[225,284,445,466]" italics="true" pageId="55" pageNumber="56">Males</emphasis>
: tegmen slightly longer than pronotum; stridulatory teeth &lt;260; cerci with a stout, wide short apical tooth.
<emphasis id="48B25C2C393FFFA5B43F0BBEFABA99A7" box="[1304,1385,445,466]" italics="true" pageId="55" pageNumber="56">Females</emphasis>
: ovipositor&gt;2.3 times longer than pronotum.
<emphasis id="48B25C2C393FFFA5B3560BD9FD72999A" box="[625,673,474,495]" italics="true" pageId="55" pageNumber="56">Song</emphasis>
(
<figureCitation id="E2FD9CBB393FFFA5B3890BD9FCE0999A" box="[686,819,474,495]" captionStart="FIGURES 182 189" captionStartId="38.[151,264,1858,1881]" captionTargetBox="[151,1436,555,1837]" captionTargetId="figure@38.[151,1436,555,1837]" captionTargetPageId="38" captionText="FIGURES 182 189. Male calling song at two different speeds (air temperature at recording shown; time scale below): 182, 183 — I. modestior (182 — BG: Vitosha Mt.; 183 — Serbia: Novi Sad); 184 — I. dobrogensis (RO: Popina Island); 185 — I. zubowskii (RO: Dobrogea, Niculiţel); 186 — I. camptoxypha (RO: Ciucaş Mt.); 187 — I. gulae (BG: Elhovo, Dolna Topchiya Reserve); 188, 189 — I. obtusa (188 — Middle Stara Planina Mts, Pleven Lodge; 189 — BG: Vitosha Mt.). Arrow and a “ Ƥ ” symbol on fig. 187 indicate the response of a female I. gulae." httpUri="https://zenodo.org/record/246566/files/figure.png" pageId="55" pageNumber="56">Figs 182, 183</figureCitation>
) consists of groups of short simple syllables (&lt;400 ms) containing or lacking after-clicks.......................................................................
<taxonomicName id="BDC6FBBD393FFFA5B40F0BFBFA4F9A78" box="[1320,1436,504,525]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="modestior">
<emphasis id="48B25C2C393FFFA5B40F0BFBFA4F9A78" box="[1320,1436,504,525]" italics="true" pageId="55" pageNumber="56">I. modestior</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7A79803E393FFFA5B1B00816FA489A10" blockId="55.[151,1437,151,730]" pageId="55" pageNumber="56">
<keyLead id="C137230B393FFFA5B1B00816FA489A10" pageId="55" pageNumber="56">
<emphasis id="48B25C2C393FFFA5B1C60816FECF9A5F" box="[225,284,533,554]" italics="true" pageId="55" pageNumber="56">Males</emphasis>
: tegmen slightly shorter than pronotum; stridulatory teeth&gt;260; cerci with a wide short apical tooth.
<emphasis id="48B25C2C393FFFA5B5C80816FA939A5F" box="[1263,1344,533,554]" italics="true" pageId="55" pageNumber="56">Females</emphasis>
: ovipositor &lt;2.3 times longer than pronotum.
<emphasis id="48B25C2C393FFFA5B37D0831FD599A32" box="[602,650,562,583]" italics="true" pageId="55" pageNumber="56">Song</emphasis>
(
<figureCitation id="E2FD9CBB393FFFA5B3BD0831FD3D9A32" box="[666,750,562,583]" captionStart="FIGURES 182 189" captionStartId="38.[151,264,1858,1881]" captionTargetBox="[151,1436,555,1837]" captionTargetId="figure@38.[151,1436,555,1837]" captionTargetPageId="38" captionText="FIGURES 182 189. Male calling song at two different speeds (air temperature at recording shown; time scale below): 182, 183 — I. modestior (182 — BG: Vitosha Mt.; 183 — Serbia: Novi Sad); 184 — I. dobrogensis (RO: Popina Island); 185 — I. zubowskii (RO: Dobrogea, Niculiţel); 186 — I. camptoxypha (RO: Ciucaş Mt.); 187 — I. gulae (BG: Elhovo, Dolna Topchiya Reserve); 188, 189 — I. obtusa (188 — Middle Stara Planina Mts, Pleven Lodge; 189 — BG: Vitosha Mt.). Arrow and a “ Ƥ ” symbol on fig. 187 indicate the response of a female I. gulae." httpUri="https://zenodo.org/record/246566/files/figure.png" pageId="55" pageNumber="56">Fig. 184</figureCitation>
) consists of groups of long elaborate syllables (&gt;800 ms) containing shorter main part and few groups of additional impulses............................................
<taxonomicName id="BDC6FBBD393FFFA5B4340853FA489A10" box="[1299,1435,592,613]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="dobrogensis">
<emphasis id="48B25C2C393FFFA5B4340853FA489A10" box="[1299,1435,592,613]" italics="true" pageId="55" pageNumber="56">I. dobrogensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C132989B393FFFA5B1B0086EFA489AAF" pageId="55" pageNumber="56">
<paragraph id="7A79803E393FFFA5B1B0086EFA489AEA" blockId="55.[151,1437,151,730]" pageId="55" pageNumber="56">
<keyLead id="C137230B393FFFA5B1B0086EFA489AEA" pageId="55" pageNumber="56">
21.
<emphasis id="48B25C2C393FFFA5B1C6086EFECF9AF7" box="[225,284,621,642]" italics="true" pageId="55" pageNumber="56">Males</emphasis>
: tip of cerci conspicuosly widened and obtuse at end, bearing short tooth (
<figureCitation id="E2FD9CBB393FFFA5B2C4086EFBE49AF7" box="[995,1079,621,642]" captionStart="FIGURES 174 181" captionStartId="36.[151,264,1078,1101]" captionTargetBox="[151,1436,193,1056]" captionTargetId="figure@36.[151,1436,193,1057]" captionTargetPageId="36" captionText="FIGURES 174 181. SEM of taxonomically important structures of Isophya: 174 — I. dobrogensis (RO: Popina Island); 175 — I. zubowskii (RO: Babadag, NMB); 176 — I. camptoxypha (RO: “ Bucec b. Sinaia, 13 1800 m, 25.8. 41, Ramme ”, NMB); 177 — I. gulae (BG: Elhovo, Dolna Topchiya Resrve); 178, 179 — I. obtusa (178 — BG: Lyulin Mt.; 179 — Pleven Lodge); 180 — I. amplipennis (TR: near Sogut); 181 — I. speciosa (BG: Kovach Locality). A—male stridulatory file (174 179 — scale 200 μm; 180, 181 — scale 100 μm); B—ventral / ventro-apical view of apex of male cerci (scale 100 μm); C—female stridulatory apparatus (scale 100 μm; for 175 C— 200 μm)." httpUri="https://zenodo.org/record/246565/files/figure.png" pageId="55" pageNumber="56">Fig. 180</figureCitation>
B); stridulatory teeth (
<figureCitation id="E2FD9CBB393FFFA5B42F086EFA8B9AF7" box="[1288,1368,621,642]" captionStart="FIGURES 174 181" captionStartId="36.[151,264,1078,1101]" captionTargetBox="[151,1436,193,1056]" captionTargetId="figure@36.[151,1436,193,1057]" captionTargetPageId="36" captionText="FIGURES 174 181. SEM of taxonomically important structures of Isophya: 174 — I. dobrogensis (RO: Popina Island); 175 — I. zubowskii (RO: Babadag, NMB); 176 — I. camptoxypha (RO: “ Bucec b. Sinaia, 13 1800 m, 25.8. 41, Ramme ”, NMB); 177 — I. gulae (BG: Elhovo, Dolna Topchiya Resrve); 178, 179 — I. obtusa (178 — BG: Lyulin Mt.; 179 — Pleven Lodge); 180 — I. amplipennis (TR: near Sogut); 181 — I. speciosa (BG: Kovach Locality). A—male stridulatory file (174 179 — scale 200 μm; 180, 181 — scale 100 μm); B—ventral / ventro-apical view of apex of male cerci (scale 100 μm); C—female stridulatory apparatus (scale 100 μm; for 175 C— 200 μm)." httpUri="https://zenodo.org/record/246565/files/figure.png" pageId="55" pageNumber="56">Fig. 180</figureCitation>
A) less than 120.
<emphasis id="48B25C2C393FFFA5B0670889FE5B9AEA" box="[320,392,650,671]" italics="true" pageId="55" pageNumber="56">Female</emphasis>
: subgenital plate usually rounded...............................................
<taxonomicName id="BDC6FBBD393FFFA5B4340889FA489AEA" box="[1299,1435,650,671]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="amplipennis">
<emphasis id="48B25C2C393FFFA5B4340889FA489AEA" box="[1299,1435,650,671]" italics="true" pageId="55" pageNumber="56">I. amplipennis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7A79803E393FFFA5B1B008A4FA489AAF" blockId="55.[151,1437,151,730]" pageId="55" pageNumber="56">
<keyLead id="C137230B393FFFA5B1B008A4FA489AAF" pageId="55" pageNumber="56">
<emphasis id="48B25C2C393FFFA5B1C608ABFECF9AC8" box="[225,284,680,701]" italics="true" pageId="55" pageNumber="56">Males</emphasis>
: tip of cerci pointed, bearing longer tooth apically or slightly subapically (
<figureCitation id="E2FD9CBB393FFFA5B52608A4FB859AC9" box="[1025,1110,679,700]" captionStart="FIGURES 174 181" captionStartId="36.[151,264,1078,1101]" captionTargetBox="[151,1436,193,1056]" captionTargetId="figure@36.[151,1436,193,1057]" captionTargetPageId="36" captionText="FIGURES 174 181. SEM of taxonomically important structures of Isophya: 174 — I. dobrogensis (RO: Popina Island); 175 — I. zubowskii (RO: Babadag, NMB); 176 — I. camptoxypha (RO: “ Bucec b. Sinaia, 13 1800 m, 25.8. 41, Ramme ”, NMB); 177 — I. gulae (BG: Elhovo, Dolna Topchiya Resrve); 178, 179 — I. obtusa (178 — BG: Lyulin Mt.; 179 — Pleven Lodge); 180 — I. amplipennis (TR: near Sogut); 181 — I. speciosa (BG: Kovach Locality). A—male stridulatory file (174 179 — scale 200 μm; 180, 181 — scale 100 μm); B—ventral / ventro-apical view of apex of male cerci (scale 100 μm); C—female stridulatory apparatus (scale 100 μm; for 175 C— 200 μm)." httpUri="https://zenodo.org/record/246565/files/figure.png" pageId="55" pageNumber="56">Fig. 181</figureCitation>
B); stridulatory teeth (
<figureCitation id="E2FD9CBB393FFFA5B40808A4FA529AC9" box="[1327,1409,679,700]" captionStart="FIGURES 174 181" captionStartId="36.[151,264,1078,1101]" captionTargetBox="[151,1436,193,1056]" captionTargetId="figure@36.[151,1436,193,1057]" captionTargetPageId="36" captionText="FIGURES 174 181. SEM of taxonomically important structures of Isophya: 174 — I. dobrogensis (RO: Popina Island); 175 — I. zubowskii (RO: Babadag, NMB); 176 — I. camptoxypha (RO: “ Bucec b. Sinaia, 13 1800 m, 25.8. 41, Ramme ”, NMB); 177 — I. gulae (BG: Elhovo, Dolna Topchiya Resrve); 178, 179 — I. obtusa (178 — BG: Lyulin Mt.; 179 — Pleven Lodge); 180 — I. amplipennis (TR: near Sogut); 181 — I. speciosa (BG: Kovach Locality). A—male stridulatory file (174 179 — scale 200 μm; 180, 181 — scale 100 μm); B—ventral / ventro-apical view of apex of male cerci (scale 100 μm); C—female stridulatory apparatus (scale 100 μm; for 175 C— 200 μm)." httpUri="https://zenodo.org/record/246565/files/figure.png" pageId="55" pageNumber="56">Fig. 181</figureCitation>
A) more than 150.
<emphasis id="48B25C2C393FFFA5B05008C6FE6C9AAF" box="[375,447,709,730]" italics="true" pageId="55" pageNumber="56">Female</emphasis>
: subgenital plate usually with an apical elongation..................................
<taxonomicName id="BDC6FBBD393FFFA5B41208C6FA489AAF" box="[1333,1435,709,730]" class="Insecta" family="Phaneropteridae" genus="Odontura" kingdom="Animalia" order="Orthoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="speciosa">
<emphasis id="48B25C2C393FFFA5B41208C6FA489AAF" box="[1333,1435,709,730]" italics="true" pageId="55" pageNumber="56">I. speciosa</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</subSection>
</document>