161 lines
18 KiB
XML
161 lines
18 KiB
XML
<document id="7B677DE47BD76C0A2FF70C00182C234D" ID-DOI="10.5281/zenodo.281291" ID-GBIF-Dataset="afd83487-f54f-4143-8481-4bf90f9aa1d7" ID-ISSN="1175-5326" ID-Zenodo-Dep="281291" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460486325359" checkinUser="plazi" docAuthor="Cruz, Estevam L., Silva, Da, Viquez, Carlos & Lise, Arno A." docDate="2012" docId="03FF87FB7F014908FF44FA0C1AABFE64" docLanguage="en" docName="zt03334p062.pdf" docOrigin="Zootaxa 3334" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Enna osaensis Cruz, Silva, Viquez & Lise, 2012, new species" docType="treatment" docVersion="8" lastPageNumber="59" masterDocId="FFC6FF837F02490CFFD3FFC61938FFAB" masterDocTitle="On the Neotropical spider genera Enna O. Pickard-Cambridge, 1897 and Syntrechalea F. O. Pickard-Cambridge, 1902 (Araneae, Lycosoidea, Trechaleidae): descriptions, taxonomic notes and new records" masterLastPageNumber="62" masterPageNumber="55" pageNumber="58" updateTime="1698309220701" updateUser="plazi">
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<mods:title id="9D2C76E86D344162BCE784C00E0A3C11">On the Neotropical spider genera Enna O. Pickard-Cambridge, 1897 and Syntrechalea F. O. Pickard-Cambridge, 1902 (Araneae, Lycosoidea, Trechaleidae): descriptions, taxonomic notes and new records</mods:title>
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<mods:namePart id="0A0567411BA9060668AF05929BF74789">Cruz, Estevam L.</mods:namePart>
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<mods:namePart id="67D9D3AD22A02C6B424E7333C6FB1207">Silva, Da</mods:namePart>
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<mods:namePart id="404DB42927C6CFE03976097D238DEB09">Viquez, Carlos</mods:namePart>
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</mods:name>
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<mods:name id="BE2C0AC869C3F4A8D2094153A6F9864C" type="personal">
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<mods:roleTerm id="EEAABC5BA2E04DE7B41BD9C190488A4B">Author</mods:roleTerm>
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<mods:namePart id="6809BEAD5D794B0449C0A7FB920FA2CC">Lise, Arno A.</mods:namePart>
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<mods:typeOfResource id="B428FFC1475561F61BCF97D6107A56FF">text</mods:typeOfResource>
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<mods:title id="93D83F3DAE6F8B270760D9193B1F71D0">Zootaxa</mods:title>
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<mods:date id="8B8DA46D101E57A87DB8412E4789F76C">2012</mods:date>
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<mods:number id="82C160FE0662BCB110E55A5578DAC956">3334</mods:number>
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<mods:identifier id="D15AA6A461FB5CC232001DE0692950E5" type="DOI">10.5281/zenodo.281291</mods:identifier>
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<treatment id="03FF87FB7F014908FF44FA0C1AABFE64" ID-DOI="http://doi.org/10.5281/zenodo.6173856" ID-GBIF-Taxon="119518403" ID-Zenodo-Dep="6173856" LSID="urn:lsid:plazi:treatment:03FF87FB7F014908FF44FA0C1AABFE64" httpUri="http://treatment.plazi.org/id/03FF87FB7F014908FF44FA0C1AABFE64" lastPageId="4" lastPageNumber="59" pageId="3" pageNumber="58">
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<subSubSection id="C34C65667F01490FFF44FA0C19C0F9AE" pageId="3" pageNumber="58" type="nomenclature">
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<paragraph id="8BE936ED7F01490FFF44FA0C18E5FA4F" blockId="3.[151,477,1482,1541]" box="[151,477,1482,1508]" pageId="3" pageNumber="58">
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<heading id="D0A181817F01490FFF44FA0C18E5FA4F" bold="true" box="[151,477,1482,1508]" fontSize="11" level="1" pageId="3" pageNumber="58" reason="1">
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<emphasis id="B922EAFF7F01490FFF44FA0C18E5FA4F" bold="true" box="[151,477,1482,1508]" pageId="3" pageNumber="58">
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<taxonomicName id="4C564D6E7F01490FFF44FA0C187EFA4F" ID-CoL="39VKR" box="[151,326,1482,1508]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="osaensis" status="sp. nov.">
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<emphasis id="B922EAFF7F01490FFF44FA0C187EFA4F" bold="true" box="[151,326,1482,1508]" italics="true" pageId="3" pageNumber="58">Enna osaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21157847F01490FFE9EFA0C18E5FA4F" box="[333,477,1482,1508]" pageId="3" pageNumber="58" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE936ED7F01490FFF44FA2B19C0F9AE" blockId="3.[151,477,1482,1541]" box="[151,248,1517,1541]" pageId="3" pageNumber="58">
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<figureCitation id="136D2A687F01490FFF44FA2B19C0F9AE" box="[151,248,1517,1541]" captionStart="FIGURES 1 – 6. 1, 2" captionStartId="2.[151,264,1620,1643]" captionTargetBox="[165,1420,194,1582]" captionTargetId="figure@2.[151,1436,171,1596]" captionTargetPageId="2" captionText="FIGURES 1 – 6. 1, 2. Enna chickeringi Silva, Lise & Carico, 2008. Male palpus (1 ventral, 2 retrolateral). 3, 4 Enna moyobamba sp. nov. Female epigynum (3 ventral, 4 dorsal). 5, 6. Enna osaensis sp. nov. Female epigynum (5 ventral, 6 dorsal). (AS = accessory spermathecae, D = duct, DD = dorsal division of median apophysis, ECD = ectal division of the retrolateral tibial apophysis, END = ental division of the retrolateral tibial apophysis, HS = head of spermathecae, ST = subtegulum, T = tegulum)." httpUri="https://zenodo.org/record/281292/files/figure.png" pageId="3" pageNumber="58">Figs 5, 6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34C65667F01490FFF44F9F31BF7F9DA" pageId="3" pageNumber="58" type="materials_examined">
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<paragraph id="8BE936ED7F01490FFF44F9F31BF7F9DA" blockId="3.[151,1436,1589,2009]" pageId="3" pageNumber="58">
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<emphasis id="B922EAFF7F01490FFF44F9F318FBF9E6" bold="true" box="[151,451,1589,1614]" pageId="3" pageNumber="58">
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<typeStatus id="54ED884F7F01490FFF44F9F319EAF9E6" box="[151,210,1589,1613]" pageId="3" pageNumber="58">Type</typeStatus>
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material:
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<typeStatus id="54ED884F7F01490FFE9EF9F31886F9E6" box="[333,446,1589,1613]" pageId="3" pageNumber="58" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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female from Peninsula de Osa, Rio Agujas, Quebrada Salitre [
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<geoCoordinate id="EE62502A7F01490FFBAEF9F31DDDF9E6" box="[1149,1253,1589,1613]" direction="north" orientation="latitude" pageId="3" pageNumber="58" precision="925" value="9.75">09°45”N</geoCoordinate>
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,
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<geoCoordinate id="EE62502A7F01490FFB22F9F31C62F9E6" box="[1265,1370,1589,1613]" direction="west" orientation="longitude" pageId="3" pageNumber="58" precision="925" value="-83.75">83°45’W</geoCoordinate>
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] Puntarenas,
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<collectingCountry id="F341767D7F01490FFF27F99F1849F9DA" box="[244,369,1625,1649]" name="Costa Rica" pageId="3" pageNumber="58">Costa Rica</collectingCountry>
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,
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<date id="FFE8102D7F01490FFEAFF99F18CCF9DA" box="[380,500,1625,1649]" pageId="3" pageNumber="58" value="1998-02-14">14.II.1998</date>
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, M. Lobo, (
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<collectionCode id="ED47AE287F01490FFDA4F99F1BFBF9DA" box="[631,707,1625,1649]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="3" pageNumber="58">INBio</collectionCode>
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).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C34C65667F01490FFF1AF9BB1ADCF93E" box="[201,996,1661,1686]" pageId="3" pageNumber="58" type="etymology">
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<paragraph id="8BE936ED7F01490FFF1AF9BB1ADCF93E" blockId="3.[151,1436,1589,2009]" box="[201,996,1661,1686]" pageId="3" pageNumber="58">
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<emphasis id="B922EAFF7F01490FFF1AF9BB1868F93E" bold="true" box="[201,336,1661,1685]" pageId="3" pageNumber="58">Etymology.</emphasis>
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The specific name is a noun and refers to the
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<typeStatus id="54ED884F7F01490FFC80F9B81ABBF93D" box="[851,899,1662,1686]" pageId="3" pageNumber="58">type</typeStatus>
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locality.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34C65667F01490FFF1AF9671B47F88E" pageId="3" pageNumber="58" type="diagnosis">
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<paragraph id="8BE936ED7F01490FFF1AF9671B47F88E" blockId="3.[151,1436,1589,2009]" pageId="3" pageNumber="58">
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<emphasis id="B922EAFF7F01490FFF1AF967187BF911" bold="true" box="[201,323,1697,1722]" pageId="3" pageNumber="58">Diagnosis.</emphasis>
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The female of
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<taxonomicName id="4C564D6E7F01490FFE38F9641B51F912" box="[491,617,1698,1721]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="osaensis" status="sp. nov.">
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<emphasis id="B922EAFF7F01490FFE38F9641B51F912" box="[491,617,1698,1721]" italics="true" pageId="3" pageNumber="58">E. osaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21157847F01490FFDBCF9641BF0F911" box="[623,712,1698,1722]" pageId="3" pageNumber="58" rank="species">
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<emphasis id="B922EAFF7F01490FFDBCF9641BF0F911" bold="true" box="[623,712,1698,1722]" pageId="3" pageNumber="58">sp. nov.</emphasis>
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</taxonomicNameLabel>
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||
(
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||
<figureCitation id="136D2A687F01490FFD05F9671A20F912" box="[726,792,1697,1721]" captionStart="FIGURES 1 – 6. 1, 2" captionStartId="2.[151,264,1620,1643]" captionTargetBox="[165,1420,194,1582]" captionTargetId="figure@2.[151,1436,171,1596]" captionTargetPageId="2" captionText="FIGURES 1 – 6. 1, 2. Enna chickeringi Silva, Lise & Carico, 2008. Male palpus (1 ventral, 2 retrolateral). 3, 4 Enna moyobamba sp. nov. Female epigynum (3 ventral, 4 dorsal). 5, 6. Enna osaensis sp. nov. Female epigynum (5 ventral, 6 dorsal). (AS = accessory spermathecae, D = duct, DD = dorsal division of median apophysis, ECD = ectal division of the retrolateral tibial apophysis, END = ental division of the retrolateral tibial apophysis, HS = head of spermathecae, ST = subtegulum, T = tegulum)." httpUri="https://zenodo.org/record/281292/files/figure.png" pageId="3" pageNumber="58">Fig. 5</figureCitation>
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) resembles those of
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<taxonomicName id="4C564D6E7F01490FFC2EF9641C42F912" authority="Silva, Lise & Carico 2008" authorityName="Silva, Lise & Carico" authorityYear="2008" box="[1021,1402,1697,1721]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="rothi">
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<emphasis id="B922EAFF7F01490FFC2EF9641D68F912" box="[1021,1104,1697,1721]" italics="true" pageId="3" pageNumber="58">E. rothi</emphasis>
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<bibRefCitation id="EFC74B1C7F01490FFB84F9671C42F912" author="Silva" box="[1111,1402,1697,1721]" pageId="3" pageNumber="58" refString="Silva, E. L. C., Lise, A. A. & Carico, J. E. (2008) Revision of the Neotropical spider genus Enna (Araneae, Lycosoidea, Trechaleidae). Journal of Arachnology, 36, 76 - 110." type="journal article" year="2008">Silva, Lise & Carico 2008</bibRefCitation>
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</taxonomicName>
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by the shape of the median field and anterior margin of the epigynum apophysis (Silva
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<emphasis id="B922EAFF7F01490FFB99F9001D43F976" box="[1098,1147,1733,1757]" italics="true" pageId="3" pageNumber="58">et al</emphasis>
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. 2008, p. 87, fig. 48), but can be distinguished by the presence of large accessory spermathecae (
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<collectingCountry id="F341767D7F01490FFC62F92F1AEFF8AA" box="[945,983,1769,1793]" name="American Samoa" pageId="3" pageNumber="58">AS</collectingCountry>
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) and by the rounded head of spermathecae (HS) without distal projections (
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<figureCitation id="136D2A687F01490FFDFEF8CB1B49F88E" box="[557,625,1805,1829]" captionStart="FIGURES 1 – 6. 1, 2" captionStartId="2.[151,264,1620,1643]" captionTargetBox="[165,1420,194,1582]" captionTargetId="figure@2.[151,1436,171,1596]" captionTargetPageId="2" captionText="FIGURES 1 – 6. 1, 2. Enna chickeringi Silva, Lise & Carico, 2008. Male palpus (1 ventral, 2 retrolateral). 3, 4 Enna moyobamba sp. nov. Female epigynum (3 ventral, 4 dorsal). 5, 6. Enna osaensis sp. nov. Female epigynum (5 ventral, 6 dorsal). (AS = accessory spermathecae, D = duct, DD = dorsal division of median apophysis, ECD = ectal division of the retrolateral tibial apophysis, END = ental division of the retrolateral tibial apophysis, HS = head of spermathecae, ST = subtegulum, T = tegulum)." httpUri="https://zenodo.org/record/281292/files/figure.png" pageId="3" pageNumber="58">Fig. 6</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C34C65667F014908FF1AF8F718B2FE07" lastPageId="4" lastPageNumber="59" pageId="3" pageNumber="58" type="description">
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||
<paragraph id="8BE936ED7F01490FFF1AF8F71CA4F872" blockId="3.[151,1436,1589,2009]" pageId="3" pageNumber="58">
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||
<emphasis id="B922EAFF7F01490FFF1AF8F71863F8E1" bold="true" box="[201,347,1841,1866]" pageId="3" pageNumber="58">Description.</emphasis>
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Female (
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<typeStatus id="54ED884F7F01490FFE14F8F71B0CF8E2" box="[455,564,1841,1865]" pageId="3" pageNumber="58" type="holotype">Holotype</typeStatus>
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, INBio). Total length 11.97. Carapace 7.05 long and 5.05 wide, dark brown with a median light brown band. Clypeus light brown, 0.46 high. Anterior eye row straight, 1.24 wide; posterior 2.32 wide. Eye diameters, interdistances, and median ocular quadrangle: AME 0.22, ALE 0.16, PME 0.26, PLE 0.15; AME-AME 0.22, AME-ALE 0.14, PME-PME 0.66, PME-PLE 0.50, OQA 0.56, OQP 1.22, OQH 0.68. Chelicerae dark brown, bristly; promargin and retromargin of fang furrow with three teeth equidistant and equal in size.
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</paragraph>
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<paragraph id="8BE936ED7F064908FF44FF511AA7FE2C" blockId="4.[151,1437,151,464]" pageId="4" pageNumber="59">
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Sternum light brown, bristly; 2.66 long, 2.79 wide. Labium light brown, slightly lighter anteriorly, 0.79 long, 1.19 wide. Legs dark brown, unmarked, relative length: IV-I-I-III, I – femur 5.98/ tibia-patella 8.91/ metatarsus 5.71/ tarsus 2.26/ total 22.86; II – 5.88/ 9.04/ 5.45/ 2.12/ 22.49; III – 5.17/ 6.91/ 4.65/ 1.72/ 18.45; IV – 6.11/ 7.98/ 6.65/ 2.66/ 23.40. Ventral pairs of macrosetae on tibiae:
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<date id="FFE8102D7F064908FD33FEC21A3CFEB7" box="[736,772,260,284]" pageId="4" pageNumber="59">I-4</date>
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;
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<date id="FFE8102D7F064908FCC1FEC21A78FEB7" box="[786,832,260,284]" pageId="4" pageNumber="59">II-4</date>
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||
;
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<date id="FFE8102D7F064908FC9DFEC21ABCFEB7" box="[846,900,260,284]" pageId="4" pageNumber="59">III-3</date>
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;
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<date id="FFE8102D7F064908FC40FEC21AF4FEB7" box="[915,972,260,284]" pageId="4" pageNumber="59">IV-3.</date>
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Abdomen, 7.71 long, dorsum dark gray with three light brown median bands; bristly, with three pairs of sigilla. Venter light gray, with scattered setae. Epigynum with two lateral rounded projections on the middle field of epigynum (
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<figureCitation id="136D2A687F064908FB92FE8A1DBFFECF" box="[1089,1159,332,356]" captionStart="FIGURES 1 – 6. 1, 2" captionStartId="2.[151,264,1620,1643]" captionTargetBox="[165,1420,194,1582]" captionTargetId="figure@2.[151,1436,171,1596]" captionTargetPageId="2" captionText="FIGURES 1 – 6. 1, 2. Enna chickeringi Silva, Lise & Carico, 2008. Male palpus (1 ventral, 2 retrolateral). 3, 4 Enna moyobamba sp. nov. Female epigynum (3 ventral, 4 dorsal). 5, 6. Enna osaensis sp. nov. Female epigynum (5 ventral, 6 dorsal). (AS = accessory spermathecae, D = duct, DD = dorsal division of median apophysis, ECD = ectal division of the retrolateral tibial apophysis, END = ental division of the retrolateral tibial apophysis, HS = head of spermathecae, ST = subtegulum, T = tegulum)." httpUri="https://zenodo.org/record/281292/files/figure.png" pageId="4" pageNumber="59">Fig. 5</figureCitation>
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). Head of spermathecae rounded with conspicuous and large accessory spermathecae (
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<figureCitation id="136D2A687F064908FC9EFEA91AAAFE2C" box="[845,914,367,391]" captionStart="FIGURES 1 – 6. 1, 2" captionStartId="2.[151,264,1620,1643]" captionTargetBox="[165,1420,194,1582]" captionTargetId="figure@2.[151,1436,171,1596]" captionTargetPageId="2" captionText="FIGURES 1 – 6. 1, 2. Enna chickeringi Silva, Lise & Carico, 2008. Male palpus (1 ventral, 2 retrolateral). 3, 4 Enna moyobamba sp. nov. Female epigynum (3 ventral, 4 dorsal). 5, 6. Enna osaensis sp. nov. Female epigynum (5 ventral, 6 dorsal). (AS = accessory spermathecae, D = duct, DD = dorsal division of median apophysis, ECD = ectal division of the retrolateral tibial apophysis, END = ental division of the retrolateral tibial apophysis, HS = head of spermathecae, ST = subtegulum, T = tegulum)." httpUri="https://zenodo.org/record/281292/files/figure.png" pageId="4" pageNumber="59">Fig. 6</figureCitation>
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).
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</paragraph>
|
||
<paragraph id="8BE936ED7F064908FF1AFE5218B2FE07" blockId="4.[151,1437,151,464]" box="[201,394,404,428]" pageId="4" pageNumber="59">
|
||
<emphasis id="B922EAFF7F064908FF1AFE521835FE07" bold="true" box="[201,269,404,428]" pageId="4" pageNumber="59">Male.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C65667F064908FF1AFE711AABFE64" box="[201,915,439,464]" pageId="4" pageNumber="59" type="distribution">
|
||
<paragraph id="8BE936ED7F064908FF1AFE711AABFE64" blockId="4.[151,1437,151,464]" box="[201,915,439,464]" pageId="4" pageNumber="59">
|
||
<emphasis id="B922EAFF7F064908FF1AFE71185AFE7B" bold="true" box="[201,354,439,464]" pageId="4" pageNumber="59">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54ED884F7F064908FDB1FE7E1BAAFE7B" box="[610,658,440,464]" pageId="4" pageNumber="59">type</typeStatus>
|
||
locality in
|
||
<collectingCountry id="F341767D7F064908FCC2FE711AB7FE64" box="[785,911,439,463]" name="Costa Rica" pageId="4" pageNumber="59">Costa Rica</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |