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<document id="6417CE9B502C6B8AD164EFBA2E4575C6" ID-DOI="10.11646/zootaxa.5383.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10361564" ID-ZooBank="20988E42-C14D-4878-A716-8CEAB5E5EF92" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1702374886811" checkinUser="plazi" docAuthor="Tasaku, Yuto, Ono, Hiroki &amp; Maruyama, Munetoshi" docDate="2023" docId="03FE87D6425CFFEC82AA3C894FCCFE0F" docLanguage="en" docName="zootaxa.5383.3.1.pdf" docOrigin="Zootaxa 5383 (3)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5383.3.1/52454" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amblopusa brevipes Casey 1893" docType="treatment" docVersion="2" lastPageNumber="280" masterDocId="FFC7FFAE4247FFF1823D3A504E1BFFA2" masterDocTitle="Review of the intertidal rove beetle tribe Liparocephalini Fenyes (Coleoptera, Staphylinidae, Aleocharinae) from Japan" masterLastPageNumber="296" masterPageNumber="251" pageNumber="278" updateTime="1702375545460" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="45143130F729C86092E28A4FEEF99A9D">Review of the intertidal rove beetle tribe Liparocephalini Fenyes (Coleoptera, Staphylinidae, Aleocharinae) from Japan</mods:title>
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<mods:roleTerm id="94A4B4DB2F5BB183EB405AD1349B0F2D">Author</mods:roleTerm>
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<mods:namePart id="7E7B055273F04A537859F4EB03EF9CE9">Tasaku, Yuto</mods:namePart>
<mods:affiliation id="0A6E27EF487C8AEED7C8AD428E2F729F">Department of Biology, School of Biological Sciences, Tokai University, Hokkaido, 005 - 8601, Japan. &amp; Corresponding author</mods:affiliation>
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<mods:namePart id="80508D468B9A0AA884066E1654AA36EB">Ono, Hiroki</mods:namePart>
<mods:affiliation id="C4E9A02268E83B91224B4297B0A0CEA8">Sounan, Minami-ku, Sagamihara-shi, Kanagawa, 252 - 0312, Japan.</mods:affiliation>
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<mods:namePart id="EA9F6D2C7BA85EE2280EE344B0BD13A7">Maruyama, Munetoshi</mods:namePart>
<mods:affiliation id="8C7F028EF9039135D4B08B869F97A542">The Kyushu University Museum, Hakozaki 6 - 10 - 1, Higashi-ku, Fukuoka, 812 - 8581, Japan.</mods:affiliation>
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<mods:title id="F3C7BE79983D43980E93AC97B228F1DD">Zootaxa</mods:title>
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<mods:part id="4E8894934F6A96CB8E6DCD15AB720AE5">
<mods:date id="D271EE706DBE61F9A7A8B48AA037617D">2023</mods:date>
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<mods:number id="FCEBD94FC5168A4A9E6ADF8BF4BDCB8B">2023-12-12</mods:number>
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<mods:number id="4671D19FB3978FDA8BAE8DDC6A10BEFE">5383</mods:number>
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<treatment id="03FE87D6425CFFEC82AA3C894FCCFE0F" ID-DOI="http://doi.org/10.5281/zenodo.10361661" ID-Zenodo-Dep="10361661" LSID="urn:lsid:plazi:treatment:03FE87D6425CFFEC82AA3C894FCCFE0F" httpUri="http://treatment.plazi.org/id/03FE87D6425CFFEC82AA3C894FCCFE0F" lastPageId="29" lastPageNumber="280" pageId="27" pageNumber="278">
<subSubSection id="C34D654B425CFFEA82AA3C894CADF8B5" pageId="27" pageNumber="278" type="nomenclature">
<paragraph id="8BE836C0425CFFEA82AA3C894C3EF951" blockId="27.[151,694,1753,1851]" box="[151,549,1753,1780]" pageId="27" pageNumber="278">
<heading id="D0A081AC425CFFEA82AA3C894C3EF951" bold="true" box="[151,549,1753,1780]" fontSize="11" level="1" pageId="27" pageNumber="278" reason="1">
<taxonomicName id="4C574D43425CFFEA82AA3C894C3EF951" ID-CoL="65ZWV" authority="Casey, 1893" authorityName="Casey" authorityYear="1893" box="[151,549,1753,1780]" class="Insecta" family="Staphylinidae" genus="Amblopusa" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="278" phylum="Arthropoda" rank="species" species="brevipes">
<emphasis id="B923EAD2425CFFEA82AA3C894C3EF951" bold="true" box="[151,549,1753,1780]" pageId="27" pageNumber="278">
<emphasis id="B923EAD2425CFFEA82AA3C894F91F956" bold="true" box="[151,394,1753,1780]" italics="true" pageId="27" pageNumber="278">Amblopusa brevipes</emphasis>
Casey, 1893
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE836C0425CFFEA82AA3CAE4CADF8B5" blockId="27.[151,694,1753,1851]" box="[151,694,1789,1816]" pageId="27" pageNumber="278">[Japanese name: Kita-nokoba-umi-hanekakushi]</paragraph>
</subSubSection>
<subSubSection id="C34D654B425CFFEA82AA3D714FC4F899" box="[151,479,1825,1851]" pageId="27" pageNumber="278" type="description">
<paragraph id="8BE836C0425CFFEA82AA3D714FC4F899" blockId="27.[151,694,1753,1851]" box="[151,479,1825,1851]" pageId="27" pageNumber="278">
(
<figureCitation id="136C2A45425CFFEA82A23D714F0BF899" box="[159,272,1825,1851]" captionStart="FIGURE 12" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[189,1398,191,1909]" captionTargetId="figure-19@19.[180,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 12 Diaulota orientalis sp. n. A, habitus, male (holotype); B, ditto, female (paratype); C, labium; D, mentum; E, median lode of aedeagus, lateral view; F, internal structure of median lobe, ventral view; G, spermatheca. Scale bars: 1.0 mm for A &amp; B; 0.1 mm for C–G." figureDoi="http://doi.org/10.5281/zenodo.10361610" httpUri="https://zenodo.org/record/10361610/files/figure.png" pageId="27" pageNumber="278">Figs. 12C</figureCitation>
;
<figureCitation id="136C2A45425CFFEA83213D714F74F899" box="[284,367,1825,1851]" captionStart="FIGURE 18" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[183,1404,191,1911]" captionTargetId="figure-22@28.[173,1413,181,1920]" captionTargetPageId="28" captionText="FIGURE 18 Genitalia of Amblopusa spp. A &amp; B, A. magna (A, median lobe of aedeagus; B, spermatheca); C &amp; D, A. pacifica (C, median lobe of aedeagus; D, spermatheca); E–G, A. brevipes (E, median lobe of aedeagus; F, spermatheca; G, apical lobe of paramere). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10361639" httpUri="https://zenodo.org/record/10361639/files/figure.png" pageId="27" pageNumber="278">18E–G</figureCitation>
;
<figureCitation id="136C2A45425CFFEA83473D714F82F899" box="[378,409,1825,1851]" captionStart="FIGURE 19" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[196,1393,192,1908]" captionTargetId="figure-22@30.[185,1403,181,1919]" captionTargetPageId="30" captionText="FIGURE 19 Mouth parts of Amblopusa brevipes. A, labrum (right side, dorsal view; left side, ventral view (epipharynx)); B, mentum; C, labium; D, maxilla; E, right mandible. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10361643" httpUri="https://zenodo.org/record/10361643/files/figure.png" pageId="27" pageNumber="278">19</figureCitation>
;
<figureCitation id="136C2A45425CFFEA83983D714FCCF899" box="[421,471,1825,1851]" captionStart="FIGURE 26" captionStartId="38.[151,250,1588,1612]" captionTargetBox="[151,1436,181,1564]" captionTargetId="figure-22@38.[151,1436,181,1564]" captionTargetPageId="38" captionText="FIGURE 26 Distribution and habitat of Amblopusa spp. of Japan. A, known localities in Japan (red circle, A. magna; blue square, A. pacifica; yellow triangle, A. brevipes; green inverted triangle, A. hokkaidona); B, habitat of A. magna and A. pacifica (Wakkainai-shi, Hokkaidô, photographed by Daisuke Sumikawa); C, habitat of A. magna (Abashiri-shi, Hokkaidô)." figureDoi="http://doi.org/10.5281/zenodo.10361675" httpUri="https://zenodo.org/record/10361675/files/figure.png" pageId="27" pageNumber="278">26A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34D654B425CFFEA82AA3D364A8DF878" pageId="27" pageNumber="278" type="reference_group">
<paragraph id="8BE836C0425CFFEA82AA3D364FA9F819" blockId="27.[151,1437,1893,2010]" pageId="27" pageNumber="278">
<taxonomicName id="4C574D43425CFFEA82AA3D364C0DF8DF" ID-CoL="65ZWV" authority="Casey, 1893: 356" authorityName="Casey" authorityPageNumber="356" authorityYear="1893" box="[151,534,1893,1918]" class="Insecta" family="Staphylinidae" genus="Amblopusa" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="278" phylum="Arthropoda" rank="species" species="brevipes">
<emphasis id="B923EAD2425CFFEA82AA3D364F7FF8DC" box="[151,356,1894,1918]" italics="true" pageId="27" pageNumber="278">Amblopusa brevipes</emphasis>
Casey, 1893: 356
</taxonomicName>
(original description;
<typeStatus id="54EC8862425CFFEA80C83D364D3AF8DC" box="[757,801,1894,1918]" pageId="27" pageNumber="278">type</typeStatus>
locality: Fort Wrangel,
<collectingRegion id="4993F822425CFFEA86333D364A42F8DC" box="[1038,1113,1894,1918]" country="United States of America" name="Alaska" pageId="27" pageNumber="278">Alaska</collectingRegion>
);
<bibRefCitation id="EFC64B31425CFFEA86573D364B3AF8DF" author="Zerche, L." box="[1130,1313,1893,1918]" pageId="27" pageNumber="278" pagination="103 - 113" refId="ref23977" refString="Zerche, L. (1998) Amblopusa magna sp. n. - eine neue Art der amphipazifischen Gattung Amblopusa Casey, 1893 aus dem Fernen Osten RuSSlands und ihre Stellung im phylogenetischen System (Coleoptera: Staphylinidae, Aleocharinae, Liparocephalini). Beitrage zur Entomologie, 48, 103 - 113." type="journal article" year="1998">Zerche, 1998: 106</bibRefCitation>
(characters; habitus photograph; phylogenetic analysis);
<bibRefCitation id="EFC64B31425CFFEA80423DD54D45F83E" author="Yoo, I. - S. &amp; Ahn, K. - J." box="[639,862,1924,1948]" pageId="27" pageNumber="278" pagination="505 - 514" refId="ref23780" refString="Yoo, I. - S. &amp; Ahn, K. - J. (2020) Discovery of a new intertidal species, Amblopusa vancouverensis from Vancouver Island, Canada with notes on Amblopusa brevipes Casey (Coleoptera, Staphylinidae, Aleocharinae). Zootaxa, 4803 (3), 505 - 514. https: // doi. org / 10.11646 / zootaxa. 4803.3.6" type="journal article" year="2020">Yoo &amp;Ahn, 2020: 506</bibRefCitation>
(redescription; habitus photograph). For other references, see
<bibRefCitation id="EFC64B31425CFFEA82D03DF34FB6F819" author="Yoo, I. - S. &amp; Ahn, K. - J." box="[237,429,1955,1979]" pageId="27" pageNumber="278" pagination="505 - 514" refId="ref23780" refString="Yoo, I. - S. &amp; Ahn, K. - J. (2020) Discovery of a new intertidal species, Amblopusa vancouverensis from Vancouver Island, Canada with notes on Amblopusa brevipes Casey (Coleoptera, Staphylinidae, Aleocharinae). Zootaxa, 4803 (3), 505 - 514. https: // doi. org / 10.11646 / zootaxa. 4803.3.6" type="journal article" year="2020">Yoo &amp; Ahn (2020)</bibRefCitation>
.
</paragraph>
<paragraph id="8BE836C0425CFFEA82AA3D924A8DF878" blockId="27.[151,1437,1893,2010]" box="[151,1174,1986,2010]" pageId="27" pageNumber="278">
<taxonomicName id="4C574D43425CFFEA82AA3D924C17F878" ID-CoL="CLB4" authority="Casey, 1911: 212" authorityName="Casey" authorityPageNumber="212" authorityYear="1911" box="[151,524,1986,2010]" class="Insecta" family="Staphylinidae" genus="Amblopusa" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="278" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B923EAD2425CFFEA82AA3D924F43F878" box="[151,344,1986,2010]" italics="true" pageId="27" pageNumber="278">Amblopusa pallida</emphasis>
Casey, 1911: 212
</taxonomicName>
(original description;
<typeStatus id="54EC8862425CFFEA80D33D924D01F878" box="[750,794,1986,2010]" pageId="27" pageNumber="278">type</typeStatus>
locality: Victoria, Vancouver Island).
</paragraph>
</subSubSection>
<caption id="DF286648425BFFED82AA3DC74FF4F855" ID-DOI="http://doi.org/10.5281/zenodo.10361639" ID-Zenodo-Dep="10361639" httpUri="https://zenodo.org/record/10361639/files/figure.png" pageId="28" pageNumber="279" startId="28.[151,250,1943,1967]" targetBox="[183,1404,191,1911]" targetPageId="28" targetType="figure">
<paragraph id="8BE836C0425BFFED82AA3DC74FF4F855" blockId="28.[151,1437,1943,2040]" pageId="28" pageNumber="279">
<emphasis id="B923EAD2425BFFED82AA3DC74F02F80D" bold="true" box="[151,281,1943,1967]" pageId="28" pageNumber="279">FIGURE 18</emphasis>
Genitalia of
<taxonomicName id="4C574D43425BFFED83A23DC74C4DF812" box="[415,598,1943,1968]" class="Insecta" family="Staphylinidae" genus="Amblopusa" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="B923EAD2425BFFED83A23DC74C0AF80D" box="[415,529,1943,1967]" italics="true" pageId="28" pageNumber="279">Amblopusa</emphasis>
spp. A
</taxonomicName>
&amp; B,
<taxonomicName id="4C574D43425BFFED80A93DC84CEDF80D" authorityName="Zerche" authorityYear="1998" box="[660,758,1944,1967]" class="Insecta" family="Staphylinidae" genus="Amblopusa" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="species" species="magna">
<emphasis id="B923EAD2425BFFED80A93DC84CB8F80D" box="[660,675,1944,1967]" italics="true" pageId="28" pageNumber="279">A</emphasis>
.
<emphasis id="B923EAD2425BFFED808C3DC84CEDF80D" box="[689,758,1944,1967]" italics="true" pageId="28" pageNumber="279">magna</emphasis>
</taxonomicName>
(A, median lobe of aedeagus; B, spermatheca); C &amp; D,
<taxonomicName id="4C574D43425BFFED87133DC84B86F812" baseAuthorityName="Sawada" baseAuthorityYear="1991" box="[1326,1437,1944,1968]" class="Insecta" family="Staphylinidae" genus="Amblopusa" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="species" species="pacifica">
<emphasis id="B923EAD2425BFFED87133DC84B26F80D" box="[1326,1341,1944,1967]" italics="true" pageId="28" pageNumber="279">A</emphasis>
.
<emphasis id="B923EAD2425BFFED87763DC84B86F812" box="[1355,1437,1944,1968]" italics="true" pageId="28" pageNumber="279">pacifica</emphasis>
</taxonomicName>
(C, median lobe of aedeagus; D, spermatheca); E–G,
<taxonomicName id="4C574D43425BFFED80813DEC4D35F871" authorityName="Casey" authorityYear="1893" box="[700,814,1979,2003]" class="Insecta" family="Staphylinidae" genus="Amblopusa" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="species" species="brevipes">
<emphasis id="B923EAD2425BFFED80813DEC4CD0F871" box="[700,715,1980,2003]" italics="true" pageId="28" pageNumber="279">A</emphasis>
.
<emphasis id="B923EAD2425BFFED80E43DEB4D35F871" box="[729,814,1979,2003]" italics="true" pageId="28" pageNumber="279">brevipes</emphasis>
</taxonomicName>
(E, median lobe of aedeagus; F, spermatheca; G, apical lobe of paramere). Scale bars: 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C34D654B425AFFEC82AA3AC64DA2FF5B" pageId="29" pageNumber="280" type="materials_examined">
<paragraph id="8BE836C0425AFFEC82AA3AC64DA2FF5B" blockId="29.[151,1437,150,429]" pageId="29" pageNumber="280">
<emphasis id="B923EAD2425AFFEC82AA3AC64F9FFF12" bold="true" box="[151,388,150,176]" pageId="29" pageNumber="280">Material examined.</emphasis>
<materialsCitation id="3B3F3C9D425AFFEC83B03AC74F2BFF77" collectingDate="2020-03-22" collectionCode="KUM" collectorName="Y. Tasaku" country="Japan" county="Ishikari-shi" location="Gokibiru" municipality="Atsuta-ku" pageId="29" pageNumber="280" specimenCount="1" stateProvince="Hokkaido">
<emphasis id="B923EAD2425AFFEC83B03AC74C72FF12" bold="true" box="[397,617,150,177]" pageId="29" pageNumber="280">
<collectingCountry id="F3407650425AFFEC83B03AC74FC1FF13" box="[397,474,151,177]" name="Japan" pageId="29" pageNumber="280">Japan</collectingCountry>
:
<collectingRegion id="4993F822425AFFEC83D73AC64C7EFF12" box="[490,613,150,176]" country="Japan" name="Hokkaido" pageId="29" pageNumber="280">Hokkaidô</collectingRegion>
:
</emphasis>
58 unsexed,
<collectingCounty id="62894E4C425AFFEC813B3AC74D93FF13" box="[774,904,151,177]" pageId="29" pageNumber="280">Ishikari-shi</collectingCounty>
,
<collectingMunicipality id="6B8CACBA425AFFEC81A83AC74A1DFF13" box="[917,1030,151,177]" pageId="29" pageNumber="280">Atsuta-ku</collectingMunicipality>
,
<location id="8E88601B425AFFEC86293AC74A67FF13" LSID="urn:lsid:plazi:treatment:03FE87D6425CFFEC82AA3C894FCCFE0F:8E88601B425AFFEC86293AC74A67FF13" box="[1044,1148,151,177]" country="Japan" county="Ishikari-shi" municipality="Atsuta-ku" name="Gokibiru" pageId="29" pageNumber="280" stateProvince="Hokkaido">Gokibiru</location>
,
<date id="FFE91000425AFFEC86B73AC74B04FF13" box="[1162,1311,151,177]" pageId="29" pageNumber="280" value="2020-03-22">
<collectingDate id="EFADE9E8425AFFEC86B73AC74B04FF13" box="[1162,1311,151,177]" pageId="29" pageNumber="280" value="2020-03-22">22. III. 2020</collectingDate>
</date>
,
<collectorName id="26A25316425AFFEC87113AC74B87FF13" box="[1324,1436,151,177]" pageId="29" pageNumber="280">Y. Tasaku</collectorName>
(
<collectionCode id="ED46AE05425AFFEC82A23AEB4EFEFF77" box="[159,229,187,213]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="29" pageNumber="280" type="Herbarium">KUM</collectionCode>
, cYT)
</materialsCitation>
;
<materialsCitation id="3B3F3C9D425AFFEC83013AEB4A32FF77" box="[316,1065,186,213]" collectingDate="2020-05-02" collectionCode="KUM" collectorName="Y. Tasaku" country="Japan" county="Ishikari-shi" location="Gokibiru" municipality="Atsuta-ku" pageId="29" pageNumber="280" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Hokkaido">
<specimenCount id="9D51FD49425AFFEC83013AEB4F8CFF77" box="[316,407,187,213]" count="2" pageId="29" pageNumber="280" type="male">2 males</specimenCount>
,
<specimenCount id="9D51FD49425AFFEC839E3AEB4C12FF77" box="[419,521,187,213]" count="1" pageId="29" pageNumber="280" type="female">1 female</specimenCount>
, 15 unsexed, ditto,
<date id="FFE91000425AFFEC80DA3AEB4D44FF77" box="[743,863,187,213]" pageId="29" pageNumber="280" value="2020-05-02">
<collectingDate id="EFADE9E8425AFFEC80DA3AEB4D44FF77" box="[743,863,187,213]" pageId="29" pageNumber="280" value="2020-05-02">2. V. 2020</collectingDate>
</date>
,
<collectorName id="26A25316425AFFEC81563AEB4DC2FF77" box="[875,985,187,213]" pageId="29" pageNumber="280">Y. Tasaku</collectorName>
(cYT)
</materialsCitation>
;
<materialsCitation id="3B3F3C9D425AFFEC860B3AEB4DAEFF5B" collectingDate="2020-05-12" collectorName="D. Sumikawa" country="Japan" county="Shimamakimura" location="Taihei-gawa" municipality="Taihei" pageId="29" pageNumber="280" specimenCount="1" specimenCount-male="1" stateProvince="Hokkaido">
<specimenCount id="9D51FD49425AFFEC860B3AEB4A9EFF77" box="[1078,1157,187,213]" count="1" pageId="29" pageNumber="280" type="male">1 male</specimenCount>
, 4 unsexed,
<collectingCounty id="62894E4C425AFFEC872E3AEB4EC8FF5B" pageId="29" pageNumber="280">Shimamakimura</collectingCounty>
,
<collectingMunicipality id="6B8CACBA425AFFEC82E33A8F4F3CFF5B" box="[222,295,223,249]" pageId="29" pageNumber="280">Taihei</collectingMunicipality>
, estuary of
<location id="8E88601B425AFFEC839A3A8F4C2FFF5B" LSID="urn:lsid:plazi:treatment:03FE87D6425CFFEC82AA3C894FCCFE0F:8E88601B425AFFEC839A3A8F4C2FFF5B" box="[423,564,223,249]" country="Japan" county="Shimamakimura" municipality="Taihei" name="Taihei-gawa" pageId="29" pageNumber="280" stateProvince="Hokkaido">Taihei-gawa</location>
,
<date id="FFE91000425AFFEC80023A8F4CD8FF5B" box="[575,707,223,249]" pageId="29" pageNumber="280" value="2020-05-12">
<collectingDate id="EFADE9E8425AFFEC80023A8F4CD8FF5B" box="[575,707,223,249]" pageId="29" pageNumber="280" value="2020-05-12">12. V. 2020</collectingDate>
</date>
,
<collectorName id="26A25316425AFFEC80F33A8F4D7DFF5B" box="[718,870,223,249]" pageId="29" pageNumber="280">D. Sumikawa</collectorName>
(cYT)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34D654B425AFFEC82FA3B524B39FEC7" pageId="29" pageNumber="280" type="diagnosis">
<paragraph id="8BE836C0425AFFEC82FA3B524B39FEC7" blockId="29.[151,1437,150,429]" pageId="29" pageNumber="280">
<emphasis id="B923EAD2425AFFEC82FA3B524F59FEBE" bold="true" box="[199,322,258,284]" pageId="29" pageNumber="280">Diagnosis.</emphasis>
This species can be distinguished from other
<taxonomicName id="4C574D43425AFFEC81243B534DF8FEBF" box="[793,995,259,285]" pageId="29" pageNumber="280">
<emphasis id="B923EAD2425AFFEC81243B534D89FEBF" box="[793,914,259,285]" italics="true" pageId="29" pageNumber="280">Amblopusa</emphasis>
species
</taxonomicName>
by the body size (about
<quantity id="4CAF9B25425AFFEC86E03B524B34FEBF" box="[1245,1327,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="29" pageNumber="280" unit="mm" value="2.5">2.5 mm</quantity>
) except
<taxonomicName id="4C574D43425AFFEC87B83B534EF7FEE3" baseAuthorityName="Sawada" baseAuthorityYear="1991" class="Insecta" family="Staphylinidae" genus="Amblopusa" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="280" phylum="Arthropoda" rank="species" species="pacifica">
<emphasis id="B923EAD2425AFFEC87B83B534B8EFEBF" box="[1413,1429,259,285]" italics="true" pageId="29" pageNumber="280">A</emphasis>
.
<emphasis id="B923EAD2425AFFEC82AA3B774EF7FEE3" box="[151,236,295,321]" italics="true" pageId="29" pageNumber="280">pacifica</emphasis>
</taxonomicName>
. From
<taxonomicName id="4C574D43425AFFEC830B3B774FBCFEE3" baseAuthorityName="Sawada" baseAuthorityYear="1991" box="[310,423,295,321]" class="Insecta" family="Staphylinidae" genus="Amblopusa" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="280" phylum="Arthropoda" rank="species" species="pacifica">
<emphasis id="B923EAD2425AFFEC830B3B774F5DFEE3" box="[310,326,295,321]" italics="true" pageId="29" pageNumber="280">A</emphasis>
.
<emphasis id="B923EAD2425AFFEC836F3B774FBCFEE3" box="[338,423,295,321]" italics="true" pageId="29" pageNumber="280">pacifica</emphasis>
</taxonomicName>
, it can be distinguished the following characteristics: pronotum slightly convex with anterolateral margin rounded (
<figureCitation id="136C2A45425AFFEC83733B1B4FB4FEC7" box="[334,431,331,357]" captionStart="FIGURE 17" captionStartId="27.[151,250,1647,1671]" captionTargetBox="[163,1424,559,1611]" captionTargetId="figure-163@27.[151,1436,548,1623]" captionTargetPageId="27" captionText="FIGURE 17 Habitus of Amblopusa spp. A, A. magna; B, A. pacifica; C, A. brevipes; D, A.hokkaidona. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10361635" httpUri="https://zenodo.org/record/10361635/files/figure.png" pageId="29" pageNumber="280">Fig. 17C</figureCitation>
); median lobe of aedeagus with apical lobe acute at apex in lateral view (
<figureCitation id="136C2A45425AFFEC86883B1B4B0EFEC7" box="[1205,1301,331,357]" captionStart="FIGURE 18" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[183,1404,191,1911]" captionTargetId="figure-22@28.[173,1413,181,1920]" captionTargetPageId="28" captionText="FIGURE 18 Genitalia of Amblopusa spp. A &amp; B, A. magna (A, median lobe of aedeagus; B, spermatheca); C &amp; D, A. pacifica (C, median lobe of aedeagus; D, spermatheca); E–G, A. brevipes (E, median lobe of aedeagus; F, spermatheca; G, apical lobe of paramere). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10361639" httpUri="https://zenodo.org/record/10361639/files/figure.png" pageId="29" pageNumber="280">Fig. 18E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34D654B425AFFEC82FA3B3E4FCCFE0F" pageId="29" pageNumber="280" type="distribution">
<paragraph id="8BE836C0425AFFEC82FA3B3E4FCCFE0F" blockId="29.[151,1437,150,429]" pageId="29" pageNumber="280">
<emphasis id="B923EAD2425AFFEC82FA3B3E4F7AFE2A" bold="true" box="[199,353,366,392]" pageId="29" pageNumber="280">Distribution.</emphasis>
<collectingCountry id="F3407650425AFFEC83563B3F4FB4FE2B" box="[363,431,367,393]" name="Japan" pageId="29" pageNumber="280">Japan</collectingCountry>
: western
<collectingRegion id="4993F822425AFFEC80203B3F4C94FE2B" box="[541,655,367,393]" country="Japan" name="Hokkaido" pageId="29" pageNumber="280">Hokkaidô</collectingRegion>
;
<collectingCountry id="F3407650425AFFEC80A33B3F4CEDFE2B" box="[670,758,367,393]" name="Canada" pageId="29" pageNumber="280">Canada</collectingCountry>
;
<collectingCountry id="F3407650425AFFEC813E3B3F4D24FE2B" box="[771,831,367,393]" name="United States of America" pageId="29" pageNumber="280">USA</collectingCountry>
:
<collectingRegion id="4993F822425AFFEC81713B3F4D86FE2B" box="[844,925,367,393]" country="United States of America" name="Alaska" pageId="29" pageNumber="280">Alaska</collectingRegion>
, the Aleutian Islands,
<collectingRegion id="4993F822425AFFEC869D3B3F4B09FE2B" box="[1184,1298,367,393]" country="United States of America" name="California" pageId="29" pageNumber="280">California</collectingRegion>
(
<figureCitation id="136C2A45425AFFEC87193B3F4B8BFE2B" box="[1316,1424,367,393]" captionStart="FIGURE 26" captionStartId="38.[151,250,1588,1612]" captionTargetBox="[151,1436,181,1564]" captionTargetId="figure-22@38.[151,1436,181,1564]" captionTargetPageId="38" captionText="FIGURE 26 Distribution and habitat of Amblopusa spp. of Japan. A, known localities in Japan (red circle, A. magna; blue square, A. pacifica; yellow triangle, A. brevipes; green inverted triangle, A. hokkaidona); B, habitat of A. magna and A. pacifica (Wakkainai-shi, Hokkaidô, photographed by Daisuke Sumikawa); C, habitat of A. magna (Abashiri-shi, Hokkaidô)." figureDoi="http://doi.org/10.5281/zenodo.10361675" httpUri="https://zenodo.org/record/10361675/files/figure.png" pageId="29" pageNumber="280">Fig. 26A</figureCitation>
). Newly recorded from
<collectingCountry id="F3407650425AFFEC83B23BC34FC9FE0F" box="[399,466,403,429]" name="Japan" pageId="29" pageNumber="280">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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