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<document id="8E350F5F2F7E1AD6D272CB321DF3BFD2" ID-DOI="10.5252/g2012n4a1" ID-ISSN="1638-9395" ID-Zenodo-Dep="5380890" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630291299389" checkinUser="marcus" docAuthor="Ernst, Andrej, Fernández, Luis Pedro, Fernández-Martínez, Esperanza &amp; Vera, Carmen" docDate="2012" docId="03A6D074FFF87E04FF1AF9EEFC25F93A" docLanguage="en" docName="Geodiversitas.2012.34.4.693-738.pdf" docOrigin="Geodiversitas 34 (4)" docSource="http://dx.doi.org/10.5252/g2012n4a1" docStyle="DocumentStyle:DF8BFA78045876A77BB343B212438C9A.3:Geodiversitas.2011-2012.journal_article.type1" docStyleId="DF8BFA78045876A77BB343B212438C9A" docStyleName="Geodiversitas.2011-2012.journal_article.type1" docStyleVersion="3" docTitle="Orthopora spinosa Ernst &amp; Fernández &amp; Fernández-Martínez &amp; Vera 2012, n. sp." docType="treatment" docVersion="5" lastPageNumber="716" masterDocId="FF9FA80CFFEF7E13FFEDFFC7FF96FF9E" masterDocTitle="Description of a bryozoan fauna from mud mounds of the Lebanza Formation (Lower Devonian) in the Arauz area (Pisuerga-Carrión Province, Cantabrian Zone, NW Spain)" masterLastPageNumber="738" masterPageNumber="693" pageNumber="716" updateTime="1699216830824" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="EA2586547D889DD0A53DAC096936E414">Description of a bryozoan fauna from mud mounds of the Lebanza Formation (Lower Devonian) in the Arauz area (Pisuerga-Carrión Province, Cantabrian Zone, NW Spain)</mods:title>
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<mods:namePart id="5D4B1495F5607B6B91DBBC2EA8195897">Ernst, Andrej</mods:namePart>
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<mods:namePart id="3D4830F89948DAF9BEC77CF3F5F411C7">Fernández, Luis Pedro</mods:namePart>
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<mods:namePart id="C7FF91CBF5E7CED8819C610530B8CAD4">Fernández-Martínez, Esperanza</mods:namePart>
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<mods:title id="5D6C5A6FEA673398F98013F3B2B0055A">Geodiversitas</mods:title>
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<subSubSection id="C31532E9FFF87E04FF1AF9EEFDB1F9FB" pageId="23" pageNumber="716" type="nomenclature">
<paragraph id="8BB06162FFF87E04FF1AF9EEFDB1F9FB" blockId="23.[208,551,1577,1637]" pageId="23" pageNumber="716">
<heading id="D0F8D60EFFF87E04FF1AF9EEFDB1F9FB" centered="true" fontSize="11" level="2" pageId="23" pageNumber="716" reason="2">
<taxonomicName id="4C0F1AE1FFF87E04FF1AF9EEFE29F9DA" authority="Ernst &amp; Fernández &amp; Fernández-Martínez &amp; Vera, 2012" authorityName="Ernst &amp; Fernández &amp; Fernández-Martínez &amp; Vera" authorityYear="2012" box="[247,447,1577,1604]" class="Stenolaemata" family="Rhabdomesidae" genus="Orthopora" kingdom="Animalia" order="Cryptostomida" pageId="23" pageNumber="716" phylum="Bryozoa" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="B97BBD70FFF87E04FF1AF9EEFE29F9DA" bold="true" box="[247,447,1577,1604]" italics="true" pageId="23" pageNumber="716">Orthopora spinosa</emphasis>
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<taxonomicNameLabel id="A248000BFFF87E04FE28F9ECFD97F9DB" box="[453,513,1579,1605]" pageId="23" pageNumber="716" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="13347DE7FFF87E04FF3AF98DFEB1F9FA" box="[215,295,1610,1636]" captionStart="FIG" captionStartId="24.[113,123,1589,1606]" captionTargetBox="[144,1180,215,1551]" captionTargetId="figure-140@24.[144,1181,215,1552]" captionTargetPageId="24" captionText="FIG.14.— A-F, Leptotrypella armata n.sp.: A, B, tangential section showing autozooecial apertures and acanthostyles,holotype SMF 21.238; C, tangential section showing autozooecial walls with abundant mural spines,paratype SMF 21.239;D, E, transverse section showing autozooecia with abundant diaphragms and autozooecial wall with mural spines,paratype SMF 21.239;F, longitudinal section,paratype SMF 21.252; G-I, Orthopora spinosa n. sp.; G, H, longitudinal section showing autozooecia with hemisepta, holotype SMF 21.254; I, tangential section showing autozooecial apertures and acanthostyles,holotype SMF 21.254.Scale bars:A-C,E, F,0.2 mm; D, G, H, 0.5 mm;I, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5380922" httpUri="https://zenodo.org/record/5380922/files/figure.png" pageId="23" pageNumber="716">Figs 14</figureCitation>
G-I; 15A-C; Appendix)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31532E9FFF87E04FF9CF94BFBD7FF71" box="[113,1089,215,1699]" pageId="23" pageNumber="716" type="materials_examined">
<paragraph id="8BB06162FFF87E04FF9CF94BFE1DF93D" blockId="23.[113,395,1675,1699]" box="[113,395,1675,1699]" pageId="23" pageNumber="716">
<materialsCitation id="3B676B3FFFF87E04FF9CF94BFE1DF93D" box="[113,395,1675,1699]" collectionCode="SMF" pageId="23" pageNumber="716" specimenCode="SMF 21.254" specimenCount="1" typeStatus="holotype">
<typeStatus id="54B4DFC0FFF87E04FF9CF94BFF4DF93C" box="[113,219,1676,1699]" pageId="23" pageNumber="716" type="holotype">HOLOTYPE</typeStatus>
. —
<specimenCode id="DBA9C919FFF87E04FEEAF94CFE11F93D" box="[263,391,1675,1699]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="23" pageNumber="716">SMF 21.254</specimenCode>
.
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</paragraph>
<paragraph id="8BB06162FFF87E04FD48FF1FFBD7FF71" blockId="23.[677,1089,215,239]" box="[677,1089,215,239]" pageId="23" pageNumber="716">
<typeStatus id="54B4DFC0FFF87E04FD48FF1FFC9DFF70" box="[677,779,216,239]" pageId="23" pageNumber="716" type="paratype">PARATYPES</typeStatus>
. —
<collectionCode id="ED1EF9A7FFF87E04FCD5FF10FCFFFF71" box="[824,873,215,239]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="23" pageNumber="716">SMF</collectionCode>
21.255-
<collectionCode id="ED1EF9A7FFF87E04FC56FF10FC78FF71" box="[955,1006,215,239]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="23" pageNumber="716">SMF</collectionCode>
21.293.
</paragraph>
</subSubSection>
<subSubSection id="C31532E9FFF87E04FD48FEC0FCF7FECA" pageId="23" pageNumber="716" type="etymology">
<paragraph id="8BB06162FFF87E04FD48FEC0FCF7FECA" blockId="23.[677,1210,263,340]" pageId="23" pageNumber="716">
ETYMOLOGY. — The species name refers to the presence of abundant acanthostyles (derived from Latin “
<emphasis id="B97BBD70FFF87E04FD5CFEFBFD6DFECD" box="[689,763,316,339]" italics="true" pageId="23" pageNumber="716">spinosus</emphasis>
spiny).
</paragraph>
</subSubSection>
<subSubSection id="C31532E9FFF87E04FD48FEABFCA1FE76" pageId="23" pageNumber="716" type="materials_examined">
<paragraph id="8BB06162FFF87E04FD48FEABFB14FE00" blockId="23.[675,1210,363,414]" pageId="23" pageNumber="716">
<collectionCode id="ED1EF9A7FFF87E04FD48FEABFCA2FE1C" box="[677,820,364,387]" pageId="23" pageNumber="716">TYPE LOCALITY</collectionCode>
. — Arauz Sur (Arroyo section), Province of Palencia, NW-Spain (Cantabrian Mountains).
</paragraph>
<paragraph id="8BB06162FFF87E04FD48FE71FCA1FE76" blockId="23.[677,1209,437,488]" pageId="23" pageNumber="716">
<collectionCode id="ED1EF9A7FFF87E04FD48FE71FCAFFE52" box="[677,825,438,461]" pageId="23" pageNumber="716">TYPE HORIZON</collectionCode>
. — Lebanza Formation, Lower Devonian (Pragian).
</paragraph>
</subSubSection>
<subSubSection id="C31532E9FFF87E04FD48FDC7FBE2FDF9" pageId="23" pageNumber="716" type="diagnosis">
<paragraph id="8BB06162FFF87E04FD48FDC7FBE2FDF9" blockId="23.[675,1212,511,615]" pageId="23" pageNumber="716">DIAGNOSIS. — Branched colonies; bifurcations common; both superior and inferior hemisepta well developed; paurostyles absent; acanthostyles abundant, arranged in single row between autozooecia, varying in size.</paragraph>
</subSubSection>
<subSubSection id="C31532E9FFF87E04FD48FD53FC25F93A" pageId="23" pageNumber="716" type="description">
<paragraph id="8BB06162FFF87E04FD48FD53FCA3FD32" blockId="23.[674,1212,660,1287]" box="[677,821,660,686]" pageId="23" pageNumber="716">DESCRIPTION</paragraph>
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Branches
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in diameter, with
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wide endozones and
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wide exozones. Branch bifurcations common. Transverse sections of branches circular.Autozooecia short, growing in spiral pattern from the distinct median axis at angles of 35-51° in endozones, abruptly bending in exozones and intersecting colony surface at angles of 51-71°; having polygonal,tear-drop shape in transverse sections of endozone. Autozooecial diaphragms rare to absent. Long superior hemisepta present, curved proximally; inferior hemisepta long, positioned beneath superior hemisepta, curved distally.Autozooecial apertures oval, arranged regularly in alternating rows on the colony surface. Walls in the endozone granular,
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mm
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thick; in exozone laminated. Acanthostyles abundant, arranged in longitudinal rows between apertures, slightly varying in size, having narrow hyaline cores and wide laminated sheaths. Heterozooecia absent.
</paragraph>
<paragraph id="8BB06162FFF87E04FD48FAE9FCA2FADB" blockId="23.[675,1211,1326,1700]" box="[677,820,1326,1352]" pageId="23" pageNumber="716">COMPARISON</paragraph>
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<taxonomicName id="4C0F1AE1FFF87E04FD48FA8BFCFCFAF8" authority="Ernst &amp; Fernández &amp; Fernández-Martínez &amp; Vera, 2012" authorityName="Ernst &amp; Fernández &amp; Fernández-Martínez &amp; Vera" authorityYear="2012" box="[677,874,1356,1382]" class="Stenolaemata" family="Rhabdomesidae" genus="Orthopora" kingdom="Animalia" order="Cryptostomida" pageId="23" pageNumber="716" phylum="Bryozoa" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="B97BBD70FFF87E04FD48FA8BFCFCFAF8" box="[677,874,1356,1382]" italics="true" pageId="23" pageNumber="716">Orthopora spinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A248000BFFF87E04FC9EFA8AFC24FAF9" box="[883,946,1357,1383]" pageId="23" pageNumber="716" rank="species">n. sp.</taxonomicNameLabel>
is similar to
<taxonomicName id="4C0F1AE1FFF87E04FBA6FA8AFCBBFA18" authority="Ernst, 2011" authorityName="Ernst" authorityYear="2011" class="Stenolaemata" family="Rhabdomesidae" genus="Orthopora" kingdom="Animalia" order="Cryptostomida" pageId="23" pageNumber="716" phylum="Bryozoa" rank="species" species="sincera">
<emphasis id="B97BBD70FFF87E04FBA6FA8AFB2FFAF8" box="[1099,1209,1357,1383]" italics="true" pageId="23" pageNumber="716">O. sincera</emphasis>
Ernst, 2011
</taxonomicName>
from the Lower to Middle Devonian of NW Spain but differs in having thinner branches (average branch width
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vs
<quantity id="4CF7CC87FFF87E04FD48FA0CFC9BFA78" box="[677,781,1483,1510]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" pageId="23" pageNumber="716" unit="mm" value="0.74">0.74 mm</quantity>
in
<taxonomicName id="4C0F1AE1FFF87E04FCDBFA0CFC35FA7B" authorityName="Ernst" authorityYear="2011" box="[822,931,1483,1509]" class="Stenolaemata" family="Rhabdomesidae" genus="Orthopora" kingdom="Animalia" order="Cryptostomida" pageId="23" pageNumber="716" phylum="Bryozoa" rank="species" species="sincera">
<emphasis id="B97BBD70FFF87E04FCDBFA0CFC35FA7B" box="[822,931,1483,1509]" italics="true" pageId="23" pageNumber="716">O. sincera</emphasis>
</taxonomicName>
), and in the smaller and more widely spaced apertures (average aperture spacing along branch
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vs
<quantity id="4CF7CC87FFF87E04FBFBF9CDFBECF9BB" box="[1046,1146,1546,1573]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="23" pageNumber="716" unit="mm" value="0.36">0.36 mm</quantity>
in
<taxonomicName id="4C0F1AE1FFF87E04FB71F9CCFD7BF9DA" authorityName="Ernst" authorityYear="2011" class="Stenolaemata" family="Rhabdomesidae" genus="Orthopora" kingdom="Animalia" order="Cryptostomida" pageId="23" pageNumber="716" phylum="Bryozoa" rank="species" species="sincera">
<emphasis id="B97BBD70FFF87E04FB71F9CCFD7BF9DA" italics="true" pageId="23" pageNumber="716">O. sincera</emphasis>
</taxonomicName>
).
<taxonomicName id="4C0F1AE1FFF87E04FCE8F9EDFC51F9DA" authority="Ernst &amp; Fernández &amp; Fernández-Martínez &amp; Vera, 2012" authorityName="Ernst &amp; Fernández &amp; Fernández-Martínez &amp; Vera" authorityYear="2012" box="[773,967,1578,1604]" class="Stenolaemata" family="Rhabdomesidae" genus="Orthopora" kingdom="Animalia" order="Cryptostomida" pageId="23" pageNumber="716" phylum="Bryozoa" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="B97BBD70FFF87E04FCE8F9EDFC51F9DA" box="[773,967,1578,1604]" italics="true" pageId="23" pageNumber="716">Orthopora spinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A248000BFFF87E04FC22F9ECFB9DF9DB" box="[975,1035,1579,1605]" pageId="23" pageNumber="716" rank="species">n. sp.</taxonomicNameLabel>
differs from
<taxonomicName id="4C0F1AE1FFF87E04FB73F9EDFCFCF9FA" authority="Ernst, 2008" authorityName="Ernst" authorityYear="2008" class="Stenolaemata" family="Trematoporidae" genus="Neotrematopora" higherTaxonomySource="CoL" kingdom="Animalia" order="Trepostomatida" pageId="23" pageNumber="716" phylum="Bryozoa" rank="species" species="tenuis">
<emphasis id="B97BBD70FFF87E04FB73F9EDFD75F9FA" italics="true" pageId="23" pageNumber="716">O. tenuis</emphasis>
Ernst,
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in having smaller autozooecial apertures (average aperture width
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vs
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in
<taxonomicName id="4C0F1AE1FFF87E04FCADF94DFC34F93D" authorityName="Ernst" authorityYear="2008" box="[832,930,1674,1700]" class="Stenolaemata" family="Trematoporidae" genus="Neotrematopora" higherTaxonomySource="CoL" kingdom="Animalia" order="Trepostomatida" pageId="23" pageNumber="716" phylum="Bryozoa" rank="species" species="tenuis">
<emphasis id="B97BBD70FFF87E04FCADF94DFC34F93D" box="[832,930,1674,1700]" italics="true" pageId="23" pageNumber="716">O. tenuis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>