179 lines
19 KiB
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179 lines
19 KiB
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<document id="8E350F5F2F7E1AD6D272CB321DF3BFD2" ID-DOI="10.5252/g2012n4a1" ID-ISSN="1638-9395" ID-Zenodo-Dep="5380890" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630291299389" checkinUser="marcus" docAuthor="Ernst, Andrej, Fernández, Luis Pedro, Fernández-Martínez, Esperanza & Vera, Carmen" docDate="2012" docId="03A6D074FFF87E04FF1AF9EEFC25F93A" docLanguage="en" docName="Geodiversitas.2012.34.4.693-738.pdf" docOrigin="Geodiversitas 34 (4)" docSource="http://dx.doi.org/10.5252/g2012n4a1" docStyle="DocumentStyle:DF8BFA78045876A77BB343B212438C9A.3:Geodiversitas.2011-2012.journal_article.type1" docStyleId="DF8BFA78045876A77BB343B212438C9A" docStyleName="Geodiversitas.2011-2012.journal_article.type1" docStyleVersion="3" docTitle="Orthopora spinosa Ernst & Fernández & Fernández-Martínez & Vera 2012, n. sp." docType="treatment" docVersion="5" lastPageNumber="716" masterDocId="FF9FA80CFFEF7E13FFEDFFC7FF96FF9E" masterDocTitle="Description of a bryozoan fauna from mud mounds of the Lebanza Formation (Lower Devonian) in the Arauz area (Pisuerga-Carrión Province, Cantabrian Zone, NW Spain)" masterLastPageNumber="738" masterPageNumber="693" pageNumber="716" updateTime="1699216830824" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="EA2586547D889DD0A53DAC096936E414">Description of a bryozoan fauna from mud mounds of the Lebanza Formation (Lower Devonian) in the Arauz area (Pisuerga-Carrión Province, Cantabrian Zone, NW Spain)</mods:title>
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<mods:namePart id="5D4B1495F5607B6B91DBBC2EA8195897">Ernst, Andrej</mods:namePart>
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</mods:name>
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</mods:role>
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<mods:namePart id="3D4830F89948DAF9BEC77CF3F5F411C7">Fernández, Luis Pedro</mods:namePart>
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</mods:name>
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<mods:name id="2E0EB4B091D6F1493D32172087C16EEB" type="personal">
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</mods:role>
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<mods:namePart id="C7FF91CBF5E7CED8819C610530B8CAD4">Fernández-Martínez, Esperanza</mods:namePart>
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</mods:name>
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<mods:name id="85C2D2A518E7B5C28737760FB7B41A7E" type="personal">
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<mods:namePart id="4C097D9EFD229138A719C90B0C8C09D3">Vera, Carmen</mods:namePart>
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<mods:typeOfResource id="E77C66B40837B78ECA13D05EEAFC6C16">text</mods:typeOfResource>
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<mods:title id="5D6C5A6FEA673398F98013F3B2B0055A">Geodiversitas</mods:title>
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<mods:part id="D7211CE88FF788EBB90A25CCE50CF3AA">
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<mods:date id="392FF5B8F5908C9025C4A65A32B85822">2012</mods:date>
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<mods:number id="EF56BDE413F7374AFBE9404A1D6B0326">2012-12-31</mods:number>
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<mods:number id="270ED88BEEEC254918C13E68BFF0CF7A">34</mods:number>
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<mods:number id="D2D77E75292E8EF95AA19E88B0F3AFBA">4</mods:number>
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<mods:url id="9EEDC5231DA9A4C9025D02864C5404F5">http://dx.doi.org/10.5252/g2012n4a1</mods:url>
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<mods:classification id="0A66CCC7EB78DC62C27884A51670CE85">journal article</mods:classification>
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<mods:identifier id="34A593B5E41466EFD43BA9899D1E0ECA" type="DOI">10.5252/g2012n4a1</mods:identifier>
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<mods:identifier id="E3AD232B025023A341507135356FA601" type="ISSN">1638-9395</mods:identifier>
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<mods:identifier id="863DBCE641B56868F3CEB64573137C9D" type="Zenodo-Dep">5380890</mods:identifier>
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<treatment id="03A6D074FFF87E04FF1AF9EEFC25F93A" LSID="urn:lsid:plazi:treatment:03A6D074FFF87E04FF1AF9EEFC25F93A" httpUri="http://treatment.plazi.org/id/03A6D074FFF87E04FF1AF9EEFC25F93A" lastPageNumber="716" pageId="23" pageNumber="716">
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<subSubSection id="C31532E9FFF87E04FF1AF9EEFDB1F9FB" pageId="23" pageNumber="716" type="nomenclature">
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<paragraph id="8BB06162FFF87E04FF1AF9EEFDB1F9FB" blockId="23.[208,551,1577,1637]" pageId="23" pageNumber="716">
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<heading id="D0F8D60EFFF87E04FF1AF9EEFDB1F9FB" centered="true" fontSize="11" level="2" pageId="23" pageNumber="716" reason="2">
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<taxonomicName id="4C0F1AE1FFF87E04FF1AF9EEFE29F9DA" authority="Ernst & Fernández & Fernández-Martínez & Vera, 2012" authorityName="Ernst & Fernández & Fernández-Martínez & Vera" authorityYear="2012" box="[247,447,1577,1604]" class="Stenolaemata" family="Rhabdomesidae" genus="Orthopora" kingdom="Animalia" order="Cryptostomida" pageId="23" pageNumber="716" phylum="Bryozoa" rank="species" species="spinosa" status="sp. nov.">
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<emphasis id="B97BBD70FFF87E04FF1AF9EEFE29F9DA" bold="true" box="[247,447,1577,1604]" italics="true" pageId="23" pageNumber="716">Orthopora spinosa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A248000BFFF87E04FE28F9ECFD97F9DB" box="[453,513,1579,1605]" pageId="23" pageNumber="716" rank="species">n. sp.</taxonomicNameLabel>
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(
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<figureCitation id="13347DE7FFF87E04FF3AF98DFEB1F9FA" box="[215,295,1610,1636]" captionStart="FIG" captionStartId="24.[113,123,1589,1606]" captionTargetBox="[144,1180,215,1551]" captionTargetId="figure-140@24.[144,1181,215,1552]" captionTargetPageId="24" captionText="FIG.14.— A-F, Leptotrypella armata n.sp.: A, B, tangential section showing autozooecial apertures and acanthostyles,holotype SMF 21.238; C, tangential section showing autozooecial walls with abundant mural spines,paratype SMF 21.239;D, E, transverse section showing autozooecia with abundant diaphragms and autozooecial wall with mural spines,paratype SMF 21.239;F, longitudinal section,paratype SMF 21.252; G-I, Orthopora spinosa n. sp.; G, H, longitudinal section showing autozooecia with hemisepta, holotype SMF 21.254; I, tangential section showing autozooecial apertures and acanthostyles,holotype SMF 21.254.Scale bars:A-C,E, F,0.2 mm; D, G, H, 0.5 mm;I, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5380922" httpUri="https://zenodo.org/record/5380922/files/figure.png" pageId="23" pageNumber="716">Figs 14</figureCitation>
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G-I; 15A-C; Appendix)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31532E9FFF87E04FF9CF94BFBD7FF71" box="[113,1089,215,1699]" pageId="23" pageNumber="716" type="materials_examined">
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<paragraph id="8BB06162FFF87E04FF9CF94BFE1DF93D" blockId="23.[113,395,1675,1699]" box="[113,395,1675,1699]" pageId="23" pageNumber="716">
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<materialsCitation id="3B676B3FFFF87E04FF9CF94BFE1DF93D" box="[113,395,1675,1699]" collectionCode="SMF" pageId="23" pageNumber="716" specimenCode="SMF 21.254" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54B4DFC0FFF87E04FF9CF94BFF4DF93C" box="[113,219,1676,1699]" pageId="23" pageNumber="716" type="holotype">HOLOTYPE</typeStatus>
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. —
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<specimenCode id="DBA9C919FFF87E04FEEAF94CFE11F93D" box="[263,391,1675,1699]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="23" pageNumber="716">SMF 21.254</specimenCode>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BB06162FFF87E04FD48FF1FFBD7FF71" blockId="23.[677,1089,215,239]" box="[677,1089,215,239]" pageId="23" pageNumber="716">
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<typeStatus id="54B4DFC0FFF87E04FD48FF1FFC9DFF70" box="[677,779,216,239]" pageId="23" pageNumber="716" type="paratype">PARATYPES</typeStatus>
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. —
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<collectionCode id="ED1EF9A7FFF87E04FCD5FF10FCFFFF71" box="[824,873,215,239]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="23" pageNumber="716">SMF</collectionCode>
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21.255-
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<collectionCode id="ED1EF9A7FFF87E04FC56FF10FC78FF71" box="[955,1006,215,239]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="23" pageNumber="716">SMF</collectionCode>
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21.293.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31532E9FFF87E04FD48FEC0FCF7FECA" pageId="23" pageNumber="716" type="etymology">
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<paragraph id="8BB06162FFF87E04FD48FEC0FCF7FECA" blockId="23.[677,1210,263,340]" pageId="23" pageNumber="716">
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ETYMOLOGY. — The species name refers to the presence of abundant acanthostyles (derived from Latin “
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<emphasis id="B97BBD70FFF87E04FD5CFEFBFD6DFECD" box="[689,763,316,339]" italics="true" pageId="23" pageNumber="716">spinosus</emphasis>
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” – spiny).
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</paragraph>
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</subSubSection>
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<subSubSection id="C31532E9FFF87E04FD48FEABFCA1FE76" pageId="23" pageNumber="716" type="materials_examined">
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<paragraph id="8BB06162FFF87E04FD48FEABFB14FE00" blockId="23.[675,1210,363,414]" pageId="23" pageNumber="716">
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<collectionCode id="ED1EF9A7FFF87E04FD48FEABFCA2FE1C" box="[677,820,364,387]" pageId="23" pageNumber="716">TYPE LOCALITY</collectionCode>
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. — Arauz Sur (Arroyo section), Province of Palencia, NW-Spain (Cantabrian Mountains).
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</paragraph>
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<paragraph id="8BB06162FFF87E04FD48FE71FCA1FE76" blockId="23.[677,1209,437,488]" pageId="23" pageNumber="716">
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<collectionCode id="ED1EF9A7FFF87E04FD48FE71FCAFFE52" box="[677,825,438,461]" pageId="23" pageNumber="716">TYPE HORIZON</collectionCode>
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. — Lebanza Formation, Lower Devonian (Pragian).
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</paragraph>
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</subSubSection>
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<subSubSection id="C31532E9FFF87E04FD48FDC7FBE2FDF9" pageId="23" pageNumber="716" type="diagnosis">
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<paragraph id="8BB06162FFF87E04FD48FDC7FBE2FDF9" blockId="23.[675,1212,511,615]" pageId="23" pageNumber="716">DIAGNOSIS. — Branched colonies; bifurcations common; both superior and inferior hemisepta well developed; paurostyles absent; acanthostyles abundant, arranged in single row between autozooecia, varying in size.</paragraph>
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</subSubSection>
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<subSubSection id="C31532E9FFF87E04FD48FD53FC25F93A" pageId="23" pageNumber="716" type="description">
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<paragraph id="8BB06162FFF87E04FD48FD53FCA3FD32" blockId="23.[674,1212,660,1287]" box="[677,821,660,686]" pageId="23" pageNumber="716">DESCRIPTION</paragraph>
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<paragraph id="8BB06162FFF87E04FD48FD74FC11FA99" blockId="23.[674,1212,660,1287]" pageId="23" pageNumber="716">
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Branches
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<quantity id="4CF7CC87FFF87E04FCE2FD74FC24FD50" box="[783,946,691,718]" metricMagnitude="-4" metricUnit="m" metricValue="6.15" metricValueMax="7.5" metricValueMin="4.8" pageId="23" pageNumber="716" unit="mm" value="0.615" valueMax="0.75" valueMin="0.48">0.48-0.75 mm</quantity>
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in diameter, with
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<quantity id="4CF7CC87FFF87E04FB6CFD74FC91FD73" metricMagnitude="-4" metricUnit="m" metricValue="2.7" metricValueMax="3.5" metricValueMin="1.9" pageId="23" pageNumber="716" unit="mm" value="0.27" valueMax="0.35" valueMin="0.19">0.19- 0.35 mm</quantity>
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wide endozones and
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<quantity id="4CF7CC87FFF87E04FC0AFD14FB14FD73" box="[999,1154,723,749]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="2.2" metricValueMin="1.4" pageId="23" pageNumber="716" unit="mm" value="0.18" valueMax="0.22" valueMin="0.14">0.14-0.22 mm</quantity>
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wide exozones. Branch bifurcations common. Transverse sections of branches circular.Autozooecia short, growing in spiral pattern from the distinct median axis at angles of 35-51° in endozones, abruptly bending in exozones and intersecting colony surface at angles of 51-71°; having polygonal,tear-drop shape in transverse sections of endozone. Autozooecial diaphragms rare to absent. Long superior hemisepta present, curved proximally; inferior hemisepta long, positioned beneath superior hemisepta, curved distally.Autozooecial apertures oval, arranged regularly in alternating rows on the colony surface. Walls in the endozone granular,
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<quantity id="4CF7CC87FFF87E04FD21FBA9FC13FB17" box="[716,901,1134,1161]" metricMagnitude="-6" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="23" pageNumber="716" unit="mm" value="0.0075" valueMax="0.01" valueMin="0.005">
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<geoCoordinate id="EE3B07A5FFF87E04FD21FBA9FC9CFB17" box="[716,778,1134,1161]" degrees="0.005" direction="north" orientation="latitude" pageId="23" pageNumber="716" precision="55" value="0.005">0.005</geoCoordinate>
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<geoCoordinate id="EE3B07A5FFF87E04FCE7FBA9FCC6FB17" box="[778,848,1134,1161]" degrees="0.010" direction="west" orientation="longitude" pageId="23" pageNumber="716" precision="55" value="-0.01">-0.010</geoCoordinate>
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mm
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</quantity>
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thick; in exozone laminated. Acanthostyles abundant, arranged in longitudinal rows between apertures, slightly varying in size, having narrow hyaline cores and wide laminated sheaths. Heterozooecia absent.
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</paragraph>
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<paragraph id="8BB06162FFF87E04FD48FAE9FCA2FADB" blockId="23.[675,1211,1326,1700]" box="[677,820,1326,1352]" pageId="23" pageNumber="716">COMPARISON</paragraph>
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<paragraph id="8BB06162FFF87E04FD48FA8BFC25F93A" blockId="23.[675,1211,1326,1700]" pageId="23" pageNumber="716">
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<taxonomicName id="4C0F1AE1FFF87E04FD48FA8BFCFCFAF8" authority="Ernst & Fernández & Fernández-Martínez & Vera, 2012" authorityName="Ernst & Fernández & Fernández-Martínez & Vera" authorityYear="2012" box="[677,874,1356,1382]" class="Stenolaemata" family="Rhabdomesidae" genus="Orthopora" kingdom="Animalia" order="Cryptostomida" pageId="23" pageNumber="716" phylum="Bryozoa" rank="species" species="spinosa" status="sp. nov.">
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<emphasis id="B97BBD70FFF87E04FD48FA8BFCFCFAF8" box="[677,874,1356,1382]" italics="true" pageId="23" pageNumber="716">Orthopora spinosa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A248000BFFF87E04FC9EFA8AFC24FAF9" box="[883,946,1357,1383]" pageId="23" pageNumber="716" rank="species">n. sp.</taxonomicNameLabel>
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is similar to
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<taxonomicName id="4C0F1AE1FFF87E04FBA6FA8AFCBBFA18" authority="Ernst, 2011" authorityName="Ernst" authorityYear="2011" class="Stenolaemata" family="Rhabdomesidae" genus="Orthopora" kingdom="Animalia" order="Cryptostomida" pageId="23" pageNumber="716" phylum="Bryozoa" rank="species" species="sincera">
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<emphasis id="B97BBD70FFF87E04FBA6FA8AFB2FFAF8" box="[1099,1209,1357,1383]" italics="true" pageId="23" pageNumber="716">O. sincera</emphasis>
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Ernst, 2011
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</taxonomicName>
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from the Lower to Middle Devonian of NW Spain but differs in having thinner branches (average branch width
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<quantity id="4CF7CC87FFF87E04FBC1FA6CFB0FFA58" box="[1068,1177,1451,1478]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="23" pageNumber="716" unit="mm" value="0.62">0.62 mm</quantity>
|
|||
|
|
vs
|
|||
|
|
<quantity id="4CF7CC87FFF87E04FD48FA0CFC9BFA78" box="[677,781,1483,1510]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" pageId="23" pageNumber="716" unit="mm" value="0.74">0.74 mm</quantity>
|
|||
|
|
in
|
|||
|
|
<taxonomicName id="4C0F1AE1FFF87E04FCDBFA0CFC35FA7B" authorityName="Ernst" authorityYear="2011" box="[822,931,1483,1509]" class="Stenolaemata" family="Rhabdomesidae" genus="Orthopora" kingdom="Animalia" order="Cryptostomida" pageId="23" pageNumber="716" phylum="Bryozoa" rank="species" species="sincera">
|
|||
|
|
<emphasis id="B97BBD70FFF87E04FCDBFA0CFC35FA7B" box="[822,931,1483,1509]" italics="true" pageId="23" pageNumber="716">O. sincera</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
), and in the smaller and more widely spaced apertures (average aperture spacing along branch
|
|||
|
|
<quantity id="4CF7CC87FFF87E04FC7EF9CCFC62F9BB" box="[915,1012,1547,1573]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="23" pageNumber="716" unit="mm" value="0.28">0.28mm</quantity>
|
|||
|
|
vs
|
|||
|
|
<quantity id="4CF7CC87FFF87E04FBFBF9CDFBECF9BB" box="[1046,1146,1546,1573]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="23" pageNumber="716" unit="mm" value="0.36">0.36 mm</quantity>
|
|||
|
|
in
|
|||
|
|
<taxonomicName id="4C0F1AE1FFF87E04FB71F9CCFD7BF9DA" authorityName="Ernst" authorityYear="2011" class="Stenolaemata" family="Rhabdomesidae" genus="Orthopora" kingdom="Animalia" order="Cryptostomida" pageId="23" pageNumber="716" phylum="Bryozoa" rank="species" species="sincera">
|
|||
|
|
<emphasis id="B97BBD70FFF87E04FB71F9CCFD7BF9DA" italics="true" pageId="23" pageNumber="716">O. sincera</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
).
|
|||
|
|
<taxonomicName id="4C0F1AE1FFF87E04FCE8F9EDFC51F9DA" authority="Ernst & Fernández & Fernández-Martínez & Vera, 2012" authorityName="Ernst & Fernández & Fernández-Martínez & Vera" authorityYear="2012" box="[773,967,1578,1604]" class="Stenolaemata" family="Rhabdomesidae" genus="Orthopora" kingdom="Animalia" order="Cryptostomida" pageId="23" pageNumber="716" phylum="Bryozoa" rank="species" species="spinosa" status="sp. nov.">
|
|||
|
|
<emphasis id="B97BBD70FFF87E04FCE8F9EDFC51F9DA" box="[773,967,1578,1604]" italics="true" pageId="23" pageNumber="716">Orthopora spinosa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A248000BFFF87E04FC22F9ECFB9DF9DB" box="[975,1035,1579,1605]" pageId="23" pageNumber="716" rank="species">n. sp.</taxonomicNameLabel>
|
|||
|
|
differs from
|
|||
|
|
<taxonomicName id="4C0F1AE1FFF87E04FB73F9EDFCFCF9FA" authority="Ernst, 2008" authorityName="Ernst" authorityYear="2008" class="Stenolaemata" family="Trematoporidae" genus="Neotrematopora" higherTaxonomySource="CoL" kingdom="Animalia" order="Trepostomatida" pageId="23" pageNumber="716" phylum="Bryozoa" rank="species" species="tenuis">
|
|||
|
|
<emphasis id="B97BBD70FFF87E04FB73F9EDFD75F9FA" italics="true" pageId="23" pageNumber="716">O. tenuis</emphasis>
|
|||
|
|
Ernst,
|
|||
|
|
<quantity id="4CF7CC87FFF87E04FCDCF98DFCFCF9FA" box="[817,874,1610,1636]" metricMagnitude="1" metricUnit="m" metricValue="5.10032" pageId="23" pageNumber="716" unit="in" value="2008.0">2008</quantity>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in having smaller autozooecial apertures (average aperture width
|
|||
|
|
<quantity id="4CF7CC87FFF87E04FBCEF9AEFB0AF91A" box="[1059,1180,1641,1668]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="23" pageNumber="716" unit="mm" value="0.05">0.050 mm</quantity>
|
|||
|
|
vs
|
|||
|
|
<quantity id="4CF7CC87FFF87E04FD48F94EFC8FF93A" box="[677,793,1673,1700]" metricMagnitude="-5" metricUnit="m" metricValue="7.6" pageId="23" pageNumber="716" unit="mm" value="0.076">0.076 mm</quantity>
|
|||
|
|
in
|
|||
|
|
<taxonomicName id="4C0F1AE1FFF87E04FCADF94DFC34F93D" authorityName="Ernst" authorityYear="2008" box="[832,930,1674,1700]" class="Stenolaemata" family="Trematoporidae" genus="Neotrematopora" higherTaxonomySource="CoL" kingdom="Animalia" order="Trepostomatida" pageId="23" pageNumber="716" phylum="Bryozoa" rank="species" species="tenuis">
|
|||
|
|
<emphasis id="B97BBD70FFF87E04FCADF94DFC34F93D" box="[832,930,1674,1700]" italics="true" pageId="23" pageNumber="716">O. tenuis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|