515 lines
61 KiB
XML
515 lines
61 KiB
XML
<document id="592167A9586B5F3F5B27F1A43FDD959B" ID-DOI="10.3897/zookeys.1218.135558" ID-ZooBank="8C9034A6-1E8D-4FF6-8721-004826BB7797" ID-publisher-id="135558" URI-arpha="6EB9F9E6-D286-5BF4-AD5D-7B66E7EB7FD9" URI-zoobank="http://zoobank.org/8C9034A6-1E8D-4FF6-8721-004826BB7797" XM.bibliography_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1731693946473" checkinUser="pensoft" docAuthor="Reyes-Hernández, José L. & Solodovnikov, Alexey" docDate="2024" docId="3699DF7013BF5ECB952C7177CE9A72D0" docLanguage="en" docName="ZooKeys 1218: 81-98" docOrigin="ZooKeys 1218" docSource="https://zookeys.pensoft.net/article/135558/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Nitidocolpus Reyes-Hernández & Solodovnikov, 2024, gen. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="98" masterDocId="6EB9F9E6D2865BF4AD5D7B66E7EB7FD9" masterDocTitle="Two new rove beetle genera in Staphylininae that reduce “ Heterothops ” and “ Quedius ” taxonomic wastebaskets (Coleoptera, Staphylinidae)" masterLastPageNumber="98" masterPageNumber="81" pageNumber="81" updateTime="1731698241112" updateUser="admin">
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<mods:titleInfo id="B1F0F1FF87612E42FA962699585E0055">
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<mods:title id="41384A0A3BD446476A8E4DA7A3FE75BC">Two new rove beetle genera in Staphylininae that reduce “ Heterothops ” and “ Quedius ” taxonomic wastebaskets (Coleoptera, Staphylinidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="CCDBE12ABBFB016F7452CF5819A35A6D" type="personal">
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<mods:roleTerm id="B37575CF83E800CB0738716E12CEE824">Author</mods:roleTerm>
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<mods:namePart id="27D7925F98025B3433F147616FA11C84">Reyes-Hernández, José L.</mods:namePart>
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<mods:nameIdentifier id="A782359FAA4496A8973EF6A14D1053B8" type="ORCID">0000-0002-4726-4439</mods:nameIdentifier>
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<mods:affiliation id="26B3043A959C065C66CA699AAF73A58A">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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</mods:role>
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<mods:namePart id="93E5BFD59903CB51C6FE192304190FAA">Solodovnikov, Alexey</mods:namePart>
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<mods:nameIdentifier id="51DB9F240201DA7B5F7E89B159CE7AD7" type="ORCID">0000-0003-2031-849X</mods:nameIdentifier>
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<mods:affiliation id="F1E83947991C252986AB5B9AC3001B0D">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:typeOfResource id="622EB2ACD95A45ABDEE4DC58F4CA54EE">text</mods:typeOfResource>
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<mods:title id="691355B9B25B1A430DABC27CFCF22907">ZooKeys</mods:title>
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<mods:part id="E43C1FD3586D849F22579FF80CE5CDD0">
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<mods:date id="EA6C479B792D6BFF4EF4E1709D876A87">2024</mods:date>
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<mods:number id="14005CD7130FCE91F866BA331199A6D0">2024-11-15</mods:number>
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<mods:detail id="4A4FCFC1696860C4F2434D89D727662A" type="volume">
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<mods:number id="E6B4D0F1BED6BD990D900473D2CD0DC4">1218</mods:number>
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<mods:start id="F575C49E38EE1A79204BEE5D5FB538F9">81</mods:start>
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<mods:classification id="28D47D960EFE95E9F80554C8DD0B157D">journal article</mods:classification>
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<mods:identifier id="D04F6149A2B4435B8EF87C7696F58FDB" type="DOI">10.3897/zookeys.1218.135558</mods:identifier>
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<treatment id="3699DF7013BF5ECB952C7177CE9A72D0" ID-arpha="3699DF70-13BF-5ECB-952C-7177CE9A72D0" ID-zoobank="https://zoobank.org/9BC238C3-B742-4E1E-8B59-48E8C83288DF" LSID="urn:lsid:plazi:treatment:3699DF7013BF5ECB952C7177CE9A72D0" LSID-ZBK="urn:lsid:zoobank.org:act:9BC238C3-B742-4E1E-8B59-48E8C83288DF" httpUri="http://treatment.plazi.org/id/3699DF7013BF5ECB952C7177CE9A72D0">
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<subSubSection id="D052E54EF3025CA5104A3C52D3EAFEE0" type="nomenclature">
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<paragraph id="9E8380C4211ED5159ACD7DFB08C64C39">
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<heading id="1F5EF7A8B9AA734689444C21458A176F" reason="label">Genus</heading>
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||
<taxonomicName id="D9255429E222663203FF494CE8870F11" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
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||
<emphasis id="ED4914F3FEDC836384E44B394730B4B2" italics="true">Nitidocolpus</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="4D1AFA5E06F609E313D72FE8FB1D4A9B" rank="genus">gen. nov.</taxonomicNameLabel>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="07D01FEBC3C978179D633A8EBEEBD11D" type="description">
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<paragraph id="0DD9FF12396B7B0CAE2A48F55D55BB64">
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||
<figureCitation id="A8167CC06431F769553C64DBFC0092A8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Some Nitidocolpus species and characters A N. illatus (Sharp, 1884) B N. laeticulus (Sharp, 1884) C N. germaini (Bernhauer, 1917) D N. aurofasciatus (Bernhauer, 1917) E N. illatus, pterothoraces F N. columbinus (Bernhauer, 1917), metacoxa and metatrochanter, dissected right leg in dorsal view (side faced to the abdomen). Abbreviations: EXPR, external part of prepectus; INPR, internal part of prepectus; MIV, mesobasisternal intercoxal process with V-shaped projection medially; MNPS, mesanapleural sutures; MTMS, metacoxa with dorsomedial spine; MTTAS, metatrochanter with dorsoapical spine. Photo B by J. Jenkins Shaw. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure5" httpUri="https://binary.pensoft.net/fig/1175478">Figs 5</figureCitation>
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,
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<figureCitation id="0B44E44C9DD2ECF0C9C54567AC747A32" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Morphological details of Nitidocolpus and other relevant genera of Staphylininae A – C Nitidocolpus columbinus D – F Cheilocolpus viridulus G – I Cyrtoquedius sp. J – L Quedionuchus nr. impunctus M – O Quedius nr. advena. A, D, G, J, M head B, E, H, K, N pronotum C, F, I, L, O aedeagus in lateral view, the black arrow pointing down indicates the basal projection of the paramere. Abbreviations: ACP, anterolateral clypeal puncture; AFP, anterior frontal; AMRSP, anterior marginal row of setiferous punctures; APP, additional paired punctures adjacent to AMRSP; BP, basal puncture; FCP, frontoclypeal puncture; IASP, internalmost setiferous puncture of the anterior margin; IOP, interocular punctures; IOR, infraorbital ridge; LLSP, large lateral setiferous puncture; LPSP, large posterior setiferous puncture; NR, nuchal ridge; PFP, posterior frontal puncture; POP, parocular punctures; PPDS, paired punctures on dorsal series; SAP, supra-antennal puncture; SAR, supra-antennal ridge; SLSP, sublateral setiferous puncture; TP, temporal puncture. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure6" httpUri="https://binary.pensoft.net/fig/1175479">6 A – C</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="SECID0EDSAG" type="type_taxon">
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||
<paragraph id="8A5A376B8DE48E8B203293F5019E6ED8">
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||
<heading id="6EEDE74377B9B3C46B0FFC9DC532FF91" reason="title">
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||
<typeStatus id="44E3E4D0DC648C59C29CFF0BF53D81EA">Type</typeStatus>
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species.
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</heading>
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</paragraph>
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<paragraph id="4A08E0D54BB44AB42481CC7744A06C2A">
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<taxonomicName id="61960DFA71E68767F97193695465BCEB" authority="Bernhauer, 1917" authorityName="Bernhauer" authorityYear="1917" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="columbinus">
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<emphasis id="A228E39B164435E57026F804AB87CDD6" italics="true">Quedius columbinus</emphasis>
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Bernhauer, 1917
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</taxonomicName>
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, here designated.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="SECID0EUSAG" type="taxon_list">
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||
<paragraph id="78447F6D03E0C83AD7BCC5A8EE46511D">
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||
<heading id="5D82CBB95499F04E5BF5B407673940D9" reason="title">Included species.</heading>
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||
</paragraph>
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||
<paragraph id="BA118F55B1464896E0479E7507DB99E4">
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||
<taxonomicName id="9CCA75FC28AE0D90963807DA9DCC3F70" authority="(Bernhauer, 1917)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Bernhauer" baseAuthorityYear="1917" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="aurofasciatus" status="comb. nov.">
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||
<emphasis id="BB83F97B7B2EF2603578EE82464DF678" italics="true">Nitidocolpus aurofasciatus</emphasis>
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||
(Bernhauer, 1917)
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||
</taxonomicName>
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,
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<taxonomicNameLabel id="889801091C875B4B652EC8E361CCE8CA" rank="species">comb. nov.</taxonomicNameLabel>
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||
ex.
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<taxonomicName id="2A3D0D73C9AA909D1558C062AFE852E5" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
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<emphasis id="804E636B4C8262BB17D354BCE7993FED" italics="true">Quedius</emphasis>
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||
</taxonomicName>
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||
[
|
||
<specimenCount id="B1573A5C23B85711E166CAB7430EA0DF" count="1" type="generic" typeStatus="syntype">1 syntype</specimenCount>
|
||
from
|
||
<named-content id="DD9EA8A659B1E8F8DD323B80D40AEB5C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/field-museum-natural-history" xlink_title="Field Museum of Natural History, Chicago, Illinois, USA">
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||
<collectionCode id="2781117D46E3B7610A83FA9956FA8ED8" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" type="Museum">FMNH</collectionCode>
|
||
</named-content>
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||
examined];
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||
<taxonomicName id="4F1B9CF3F2DEF769BD0FD469432F32A5" authority="(Sharp, 1884)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1884" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="championi" status="comb. nov.">
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||
<emphasis id="8663ABC194A8DF7F80BA15B7B3F057B5" italics="true">Nitidocolpus championi</emphasis>
|
||
(Sharp, 1884)
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||
</taxonomicName>
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||
,
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||
<taxonomicNameLabel id="1D83FC4ABFACBD6255973A88B1BC8CCA" rank="species">comb. nov.</taxonomicNameLabel>
|
||
ex.
|
||
<taxonomicName id="FE64A534D0F1C469B93E3DAD942386A8" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9573DBF8CBDBCF4D0190C055752BABD5" italics="true">Quedius</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<specimenCount id="16D90C432125D3D8D6DFCE666D4E010D" count="1" type="generic" typeStatus="syntype">1 syntype</specimenCount>
|
||
from
|
||
<named-content id="298367C0EF2549D37A8AE44ADAC2FE84" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/field-museum-natural-history" xlink_title="Field Museum of Natural History, Chicago, Illinois, USA">
|
||
<collectionCode id="F90AE8D557C81D1D729B680D087393C8" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" type="Museum">FMNH</collectionCode>
|
||
</named-content>
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||
examined];
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||
<taxonomicName id="53A8318C45DDF1662C0BC64E6154DE28" authority="(Bernhauer, 1917)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Bernhauer" baseAuthorityYear="1917" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="columbinus" status="comb. nov.">
|
||
<emphasis id="FA76C56CB9F9500AC0FE31F06CF7F40E" italics="true">Nitidocolpus columbinus</emphasis>
|
||
(Bernhauer, 1917)
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||
</taxonomicName>
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||
,
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||
<taxonomicNameLabel id="9840ECB5FBE01DF37B3DC39E5DA50520" rank="species">comb. nov.</taxonomicNameLabel>
|
||
ex.
|
||
<taxonomicName id="F7A1E990859F7BE62A32A44AD607AA51" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="65F966380F75020BAC758982F21587AC" italics="true">Quedius</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<specimenCount id="2A38914883AFBEB2874BA8A90E7C9346" count="2" type="generic" typeStatus="syntype">2 syntypes</specimenCount>
|
||
from
|
||
<named-content id="178B5AF9240B17ABCB09A64080A7C35D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/field-museum-natural-history" xlink_title="Field Museum of Natural History, Chicago, Illinois, USA">
|
||
<collectionCode id="6504880A0784E4F488C0B2C8C8036345" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" type="Museum">FMNH</collectionCode>
|
||
</named-content>
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||
examined];
|
||
<taxonomicName id="FB50C1961640551FBD01B5144254FB8D" authority="(Bernhauer, 1917)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Bernhauer" baseAuthorityYear="1917" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="germaini" status="comb. nov.">
|
||
<emphasis id="A5206C04C6E665F76D227525CC46CCB3" italics="true">Nitidocolpus germaini</emphasis>
|
||
(Bernhauer, 1917)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="71A3485825FB1D229A54F0F150AA66B1" rank="species">comb. nov.</taxonomicNameLabel>
|
||
ex.
|
||
<taxonomicName id="3C3C441F1487661EB81791F8EF8D2365" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F14C2FCAEA45D720BDE77F6E0C22A7E9" italics="true">Quedius</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<specimenCount id="80AD57FD3EFAFFAA44C4ED92DBC2442A" count="1" type="generic" typeStatus="syntype">1 syntype</specimenCount>
|
||
from
|
||
<named-content id="89620B5DD8546A7A1B4F4F4192B90E55" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/field-museum-natural-history" xlink_title="Field Museum of Natural History, Chicago, Illinois, USA">
|
||
<collectionCode id="CECFF9D28FA0910E86F0ABFF15D5DCA4" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" type="Museum">FMNH</collectionCode>
|
||
</named-content>
|
||
examined];
|
||
<taxonomicName id="6E7E989B580D37FBD5D2A82CB584F497" authority="(Sharp, 1884)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1884" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="illatus" status="comb. nov.">
|
||
<emphasis id="54C8438D01B087FEB6790DDCF9DD906E" italics="true">Nitidocolpus illatus</emphasis>
|
||
(Sharp, 1884)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="2CA14E7232C00A7F908F1D8C5A1E1BE6" rank="species">comb. nov.</taxonomicNameLabel>
|
||
ex.
|
||
<taxonomicName id="674B78EA095409372D3247149ED86776" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="47EE25405302540ADC1081D207E83594" italics="true">Quedius</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<specimenCount id="1690860E92C3FD4CAD0D90FE71F75E40" count="1" type="generic" typeStatus="syntype">1 syntype</specimenCount>
|
||
from
|
||
<named-content id="C8D2159C5ACCF82BD5E49C40AB55616F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London, United Kingdom">
|
||
<collectionCode id="FA73F48E2859DDD32683C97524203238" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" type="Herbarium">NHM</collectionCode>
|
||
</named-content>
|
||
examined];
|
||
<taxonomicName id="FA1D40CF50DC27F27B3D72EF3F431697" authority="(Sharp, 1884)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1884" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="laeticulus" status="comb. nov.">
|
||
<emphasis id="ABA6FC1CBE379F9B4DAEA63473E633E5" italics="true">Nitidocolpus laeticulus</emphasis>
|
||
(Sharp, 1884)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="4BAF4B4364144819EDA1E7FCC9B6AB85" rank="species">comb. nov.</taxonomicNameLabel>
|
||
ex.
|
||
<taxonomicName id="259C4239A53AADCC5950BE38900C35CA" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="04772AF80D9280510D6FB594DFAB5A74" italics="true">Quedius</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<specimenCount id="2002368E1550B97C94064ABB01CD8B3A" count="1" type="generic" typeStatus="syntype">1 syntype</specimenCount>
|
||
from
|
||
<named-content id="0846CD16F5931F2BDC5F4661A35238AE" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London, United Kingdom">
|
||
<collectionCode id="DE78B435A65CCB7B151738DA8C137E4D" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" type="Herbarium">NHM</collectionCode>
|
||
</named-content>
|
||
examined];
|
||
<taxonomicName id="2E02A7AE6016BF165876F471831A548D" authority="(Fauvel, 1891)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Fauvel" baseAuthorityYear="1891" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="triangulum" status="comb. nov.">
|
||
<emphasis id="9998F6A591086AB009B9F88328263BFF" italics="true">Nitidocolpus triangulum</emphasis>
|
||
(
|
||
<bibRefCitation id="8343FD47D750BB67A2B9D3EEADFBE175" author="Fauvel A" journalOrPublisher="Revue d’Entomologie" pagination="87-127" refId="B7" refString="Fauvel A (1891) Voyage de M. E. Simon au venezuela (Decembre 1887 - Avril 1888). 11 e Memoire. Revue d'Entomologie 10: 87 - 127." title="Voyage de M. E. Simon au vénézuela (Décembre 1887 - Avril 1888). 11 e Mémoire." volume="10" year="1891">Fauvel, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="EA1AE2E5E76A5A3713E2A04725FCB6D7" rank="species">comb. nov.</taxonomicNameLabel>
|
||
ex.
|
||
<taxonomicName id="C1815A963E1C2F3257586C133B07A9D0" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2F8244FDAB81931249FA30BC7EFA4B9C" italics="true">Quedius</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<specimenCount id="9515E08D14F6C09AA441ADA08BB783CF" count="1" type="generic" typeStatus="syntype">1 syntype</specimenCount>
|
||
from
|
||
<named-content id="6E89391C1A86F132F5449638B4595E3F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institut-royal-des-sciences-naturelles-de-belgique" xlink_title="Institut Royal des Sciences Naturelles de Belgique, Brussels, Belgium">
|
||
<collectionCode id="F30FC1D7168AF2CB494A1B7E67C4A5D8" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" type="">IRSNB</collectionCode>
|
||
</named-content>
|
||
examined].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E2XAG" type="diagnosis">
|
||
<paragraph id="181C40653B9C2C323D18EB3777E19A4B">
|
||
<heading id="16A00FFE603C343A32F6A0289801FAB0" reason="title">Diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="5270BDC0AF27BC2E3247E7989B23F8EE">
|
||
Supra-antennal punctures and paraocular punctures absent (Fig.
|
||
<figureCitation id="277E0FBAD225D331E4ED850F3D4ED286" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Morphological details of Nitidocolpus and other relevant genera of Staphylininae A – C Nitidocolpus columbinus D – F Cheilocolpus viridulus G – I Cyrtoquedius sp. J – L Quedionuchus nr. impunctus M – O Quedius nr. advena. A, D, G, J, M head B, E, H, K, N pronotum C, F, I, L, O aedeagus in lateral view, the black arrow pointing down indicates the basal projection of the paramere. Abbreviations: ACP, anterolateral clypeal puncture; AFP, anterior frontal; AMRSP, anterior marginal row of setiferous punctures; APP, additional paired punctures adjacent to AMRSP; BP, basal puncture; FCP, frontoclypeal puncture; IASP, internalmost setiferous puncture of the anterior margin; IOP, interocular punctures; IOR, infraorbital ridge; LLSP, large lateral setiferous puncture; LPSP, large posterior setiferous puncture; NR, nuchal ridge; PFP, posterior frontal puncture; POP, parocular punctures; PPDS, paired punctures on dorsal series; SAP, supra-antennal puncture; SAR, supra-antennal ridge; SLSP, sublateral setiferous puncture; TP, temporal puncture. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure6" httpUri="https://binary.pensoft.net/fig/1175479">6 A</figureCitation>
|
||
); pronotum without paired dorsal series of setiferous macropunctures (Fig.
|
||
<figureCitation id="05C4E4AD73FFFADAA64FBD967C84F7B6" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Morphological details of Nitidocolpus and other relevant genera of Staphylininae A – C Nitidocolpus columbinus D – F Cheilocolpus viridulus G – I Cyrtoquedius sp. J – L Quedionuchus nr. impunctus M – O Quedius nr. advena. A, D, G, J, M head B, E, H, K, N pronotum C, F, I, L, O aedeagus in lateral view, the black arrow pointing down indicates the basal projection of the paramere. Abbreviations: ACP, anterolateral clypeal puncture; AFP, anterior frontal; AMRSP, anterior marginal row of setiferous punctures; APP, additional paired punctures adjacent to AMRSP; BP, basal puncture; FCP, frontoclypeal puncture; IASP, internalmost setiferous puncture of the anterior margin; IOP, interocular punctures; IOR, infraorbital ridge; LLSP, large lateral setiferous puncture; LPSP, large posterior setiferous puncture; NR, nuchal ridge; PFP, posterior frontal puncture; POP, parocular punctures; PPDS, paired punctures on dorsal series; SAP, supra-antennal puncture; SAR, supra-antennal ridge; SLSP, sublateral setiferous puncture; TP, temporal puncture. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure6" httpUri="https://binary.pensoft.net/fig/1175479">6 B</figureCitation>
|
||
); mesanapleural sutures transverse or nearly transverse but reaching the external part of prepectus (Fig.
|
||
<figureCitation id="03AB624FB58BD3432A865BCB61AA490B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Some Nitidocolpus species and characters A N. illatus (Sharp, 1884) B N. laeticulus (Sharp, 1884) C N. germaini (Bernhauer, 1917) D N. aurofasciatus (Bernhauer, 1917) E N. illatus, pterothoraces F N. columbinus (Bernhauer, 1917), metacoxa and metatrochanter, dissected right leg in dorsal view (side faced to the abdomen). Abbreviations: EXPR, external part of prepectus; INPR, internal part of prepectus; MIV, mesobasisternal intercoxal process with V-shaped projection medially; MNPS, mesanapleural sutures; MTMS, metacoxa with dorsomedial spine; MTTAS, metatrochanter with dorsoapical spine. Photo B by J. Jenkins Shaw. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure5" httpUri="https://binary.pensoft.net/fig/1175478">5 E</figureCitation>
|
||
); mesobasisternal intercoxal process with V-shaped projection medially (Fig.
|
||
<figureCitation id="86CA88BF25616C52AF7E292FD3DC356E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Some Nitidocolpus species and characters A N. illatus (Sharp, 1884) B N. laeticulus (Sharp, 1884) C N. germaini (Bernhauer, 1917) D N. aurofasciatus (Bernhauer, 1917) E N. illatus, pterothoraces F N. columbinus (Bernhauer, 1917), metacoxa and metatrochanter, dissected right leg in dorsal view (side faced to the abdomen). Abbreviations: EXPR, external part of prepectus; INPR, internal part of prepectus; MIV, mesobasisternal intercoxal process with V-shaped projection medially; MNPS, mesanapleural sutures; MTMS, metacoxa with dorsomedial spine; MTTAS, metatrochanter with dorsoapical spine. Photo B by J. Jenkins Shaw. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure5" httpUri="https://binary.pensoft.net/fig/1175478">5 E</figureCitation>
|
||
); metatrochanters with apical spine (Fig.
|
||
<figureCitation id="135D4DB81A329791BD47D5981423D4E5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Some Nitidocolpus species and characters A N. illatus (Sharp, 1884) B N. laeticulus (Sharp, 1884) C N. germaini (Bernhauer, 1917) D N. aurofasciatus (Bernhauer, 1917) E N. illatus, pterothoraces F N. columbinus (Bernhauer, 1917), metacoxa and metatrochanter, dissected right leg in dorsal view (side faced to the abdomen). Abbreviations: EXPR, external part of prepectus; INPR, internal part of prepectus; MIV, mesobasisternal intercoxal process with V-shaped projection medially; MNPS, mesanapleural sutures; MTMS, metacoxa with dorsomedial spine; MTTAS, metatrochanter with dorsoapical spine. Photo B by J. Jenkins Shaw. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure5" httpUri="https://binary.pensoft.net/fig/1175478">5 F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="1DA9578AB3A79AF20937A0C3C61634AF" ID-DOI="10.3897/zookeys.1218.135558.figure5" ID-arpha="BA24B181-2D31-5A39-BAE9-7EC7E37A42EF" httpUri="https://binary.pensoft.net/fig/1175478" startId="F5">
|
||
<paragraph id="F68748BAC8DFE51A68B4146241BE9A96">
|
||
<label id="B57CD1A9F2C14765CA9DA13C47E20895">Figure 5.</label>
|
||
</paragraph>
|
||
<paragraph id="E9583DCFFE10CC837729F976FD05D5A2">
|
||
Some
|
||
<taxonomicName id="53497210DAC1326658A25FF975CAFB36" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="undefined">
|
||
<emphasis id="9DAC792E2BCB7DF8601A653430438DC8" italics="true">Nitidocolpus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
and characters
|
||
<emphasis id="9CBA4E1C747135D0514882F5B427BAC7" bold="true">A</emphasis>
|
||
<taxonomicName id="C2B5554F316FCD72263C7ADBFD35E643" authority="(Sharp, 1884)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1884" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="illatus">
|
||
<emphasis id="B90B72EE8858811423104370673D8277" italics="true">N. illatus</emphasis>
|
||
(Sharp, 1884)
|
||
</taxonomicName>
|
||
<emphasis id="8D8A87C8CA68A0C4EAC346D47B19E1DE" bold="true">B</emphasis>
|
||
<taxonomicName id="05CD16CE00F6F43A7409DE3D2711EFB8" authority="(Sharp, 1884)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1884" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="laeticulus">
|
||
<emphasis id="388D7352890CCE0A96C7BAA7AD4F1AEB" italics="true">N. laeticulus</emphasis>
|
||
(Sharp, 1884)
|
||
</taxonomicName>
|
||
<emphasis id="C4411B626BDFA0493831C178B94E6C4E" bold="true">C</emphasis>
|
||
<taxonomicName id="C1C5876AAD2535240DA3426DAFB235C3" authority="(Bernhauer, 1917)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Bernhauer" baseAuthorityYear="1917" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="germaini">
|
||
<emphasis id="E99FDF2876270A6DF8B3E7A18A1B236B" italics="true">N. germaini</emphasis>
|
||
(Bernhauer, 1917)
|
||
</taxonomicName>
|
||
<emphasis id="D0020B0EF3DC55C62B903610736FDD92" bold="true">D</emphasis>
|
||
<taxonomicName id="4B84DEBEAD26C0FF88641B54A55CE67C" authority="(Bernhauer, 1917)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Bernhauer" baseAuthorityYear="1917" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="aurofasciatus">
|
||
<emphasis id="CF4F8035CB4D22E5B7DB34345F305346" italics="true">N. aurofasciatus</emphasis>
|
||
(Bernhauer, 1917)
|
||
</taxonomicName>
|
||
<emphasis id="F6931EDF0C0EE000CC268455F47C3542" bold="true">E</emphasis>
|
||
<taxonomicName id="01DC1F87492F902CAEB3F195C2EB7607" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1884" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="illatus">
|
||
<emphasis id="128C9A195F2CDF2FAAAEC11709EB61E2" italics="true">N. illatus</emphasis>
|
||
</taxonomicName>
|
||
, pterothoraces
|
||
<emphasis id="8D2956D7FAEAD49D16FD913ABA44952E" bold="true">F</emphasis>
|
||
<taxonomicName id="2B437BEE8E0645F0DCB5B8C27A44F992" authority="(Bernhauer, 1917)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Bernhauer" baseAuthorityYear="1917" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="columbinus">
|
||
<emphasis id="1799D272A48ACC70B8D091353868B849" italics="true">N. columbinus</emphasis>
|
||
(Bernhauer, 1917)
|
||
</taxonomicName>
|
||
, metacoxa and metatrochanter, dissected right leg in dorsal view (side faced to the abdomen). Abbreviations: EXPR, external part of prepectus; INPR, internal part of prepectus; MIV, mesobasisternal intercoxal process with V-shaped projection medially;
|
||
<abbrev id="ABBRID0ES2AG" xlink_title="mesanapleural sutures">MNPS</abbrev>
|
||
, mesanapleural sutures; MTMS, metacoxa with dorsomedial spine; MTTAS, metatrochanter with dorsoapical spine. Photo B by J. Jenkins Shaw. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E22AG" type="description">
|
||
<paragraph id="1C3AAC14DB1E1772798FE08703194050">
|
||
<heading id="48C8FDC42D53ED562942330CC53B3128" reason="title">Description.</heading>
|
||
</paragraph>
|
||
<paragraph id="E2546B41D467FA69E694E93CD59E97D8">
|
||
Small to medium-sized
|
||
<taxonomicName id="BAE32A18CCF6BBCF0476C623A9CB4C2D" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subFamily" subFamily="Staphylininae">Staphylininae</taxonomicName>
|
||
(TL =
|
||
<quantity id="F8CC2A19632023D703443DA100C1FFF7" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="10.0" metricValueMin="4.5" unit="mm" value="7.25" valueMax="10.0" valueMin="4.5">4.5–10 mm</quantity>
|
||
) (Fig.
|
||
<figureCitation id="43C88BF7D231FB04022F0309FCD21430" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Some Nitidocolpus species and characters A N. illatus (Sharp, 1884) B N. laeticulus (Sharp, 1884) C N. germaini (Bernhauer, 1917) D N. aurofasciatus (Bernhauer, 1917) E N. illatus, pterothoraces F N. columbinus (Bernhauer, 1917), metacoxa and metatrochanter, dissected right leg in dorsal view (side faced to the abdomen). Abbreviations: EXPR, external part of prepectus; INPR, internal part of prepectus; MIV, mesobasisternal intercoxal process with V-shaped projection medially; MNPS, mesanapleural sutures; MTMS, metacoxa with dorsomedial spine; MTTAS, metatrochanter with dorsoapical spine. Photo B by J. Jenkins Shaw. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure5" httpUri="https://binary.pensoft.net/fig/1175478">5 A – D</figureCitation>
|
||
). Head: Frontoclypeal (epistomal) suture absent or distinct only laterally and only in teneral (or chemically cleared) specimens; head slightly transverse (
|
||
<abbrev id="ABBRID0EL3AG" xlink_title="head width at the widest point, including eyes">
|
||
<collectionCode id="A116B4800F19AA5AFD612F29B0F785FB">HW</collectionCode>
|
||
</abbrev>
|
||
/
|
||
<abbrev id="ABBRID0EP3AG" xlink_title="head length">
|
||
<collectionCode id="0F3DAD02CB67BCF9DE3AEBCFBC5E4A1F" country="USA" lsid="urn:lsid:biocol.org:col:13245" name="Houghton Lake Wildlife Research Station" type="Herbarium">HL</collectionCode>
|
||
</abbrev>
|
||
≥ 1.1) without distinct posterior angles; disc smooth; frons with microsculpture as transverse waves, without distinct concavity between antennal insertions; neck moderately wide (
|
||
<abbrev id="ABBRID0ET3AG" xlink_title="neck width at the widest point">
|
||
<collectionCode id="F65ED6A6D1542C159665166C1CBA91CE">NW</collectionCode>
|
||
</abbrev>
|
||
/
|
||
<abbrev id="ABBRID0EX3AG" xlink_title="head width at the widest point, including eyes">
|
||
<collectionCode id="7F6398D9D5597322D37E48F91E444493">HW</collectionCode>
|
||
</abbrev>
|
||
> 0.75 but <0.90); dorsal macrosetae (Fig.
|
||
<figureCitation id="D7E614E67846F34BD1D3E668E0B4CE35" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Morphological details of Nitidocolpus and other relevant genera of Staphylininae A – C Nitidocolpus columbinus D – F Cheilocolpus viridulus G – I Cyrtoquedius sp. J – L Quedionuchus nr. impunctus M – O Quedius nr. advena. A, D, G, J, M head B, E, H, K, N pronotum C, F, I, L, O aedeagus in lateral view, the black arrow pointing down indicates the basal projection of the paramere. Abbreviations: ACP, anterolateral clypeal puncture; AFP, anterior frontal; AMRSP, anterior marginal row of setiferous punctures; APP, additional paired punctures adjacent to AMRSP; BP, basal puncture; FCP, frontoclypeal puncture; IASP, internalmost setiferous puncture of the anterior margin; IOP, interocular punctures; IOR, infraorbital ridge; LLSP, large lateral setiferous puncture; LPSP, large posterior setiferous puncture; NR, nuchal ridge; PFP, posterior frontal puncture; POP, parocular punctures; PPDS, paired punctures on dorsal series; SAP, supra-antennal puncture; SAR, supra-antennal ridge; SLSP, sublateral setiferous puncture; TP, temporal puncture. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure6" httpUri="https://binary.pensoft.net/fig/1175479">6 A</figureCitation>
|
||
): anterolateral clypeal punctures present; frontoclypeal punctures present; supra-antennal punctures absent; parocular punctures absent; basal punctures present, single; ventral macrosetae: postocular punctures present; infraorbital punctures absent; postmandibular punctures present; submentum with two pairs of macrosetae; mentum with seta alpha only, seta beta absent; gular sutures separated from each other, gula with distinct transverse basal impression; ventral basal ridge present; postgenal ridge present; postmandibular ridge short, extended parallel to margin of eye, without fusion with any other ridge and distinctly separated from eye margin; infraorbital ridge merged with postgenal ridge; nuchal ridge missing dorsally, present laterally, merged with infraorbital ridge; eyes of medium size (
|
||
<abbrev id="ABBRID0E63AG" xlink_title="eye length in lateral view">EYL</abbrev>
|
||
/
|
||
<abbrev id="ABBRID0ED4AG" xlink_title="head length">
|
||
<collectionCode id="5C4D641BD207315543D2FD2848A990DC" country="USA" lsid="urn:lsid:biocol.org:col:13245" name="Houghton Lake Wildlife Research Station" type="Herbarium">HL</collectionCode>
|
||
</abbrev>
|
||
ratio more than 1 / 2 but less than 3 / 4); gena short (
|
||
<abbrev id="ABBRID0EH4AG" xlink_title="gena length">GL</abbrev>
|
||
/
|
||
<collectionCode id="B39285C8C65738F9DF3D47DE67381274" country="USA" name="Lake Forest College" type="Herbarium">ETL</collectionCode>
|
||
<0.5); tomentose pubescence begins from fourth antennomere, its density on fourth antennomere as on following antennomeres; setation of antennomere 3 almost evenly distributed; antennomere 11 with subapical rounded field, without subapical lateral pits; mandibles with dorso-lateral seta, without dorso-lateral groove, external edge with curved base and apex, straight in middle length; right mandible with proximal tooth and bifid distal tooth, apically deflexed ventrad at an angle of> 20 °; left mandible rather straight; labrum with transparent wide apical membrane, anterior margin emarginate at middle; maxillary palps: palpomere 4 (apical) subconical, with evenly narrowed apex, longer than palpomere 2 and weakly dilated compared to palpomere 3; labial palps: palpomere 3 (apical) more or less fusiform or subconical, distinctly longer than, and about as wide as, palpomere 2.
|
||
</paragraph>
|
||
<caption id="C5828026F215B5D6C8C4CDC7A5B9880C" ID-DOI="10.3897/zookeys.1218.135558.figure6" ID-arpha="C66562ED-C5CB-5792-BB98-24153A71FF18" httpUri="https://binary.pensoft.net/fig/1175479" startId="F6">
|
||
<paragraph id="BB38C84CDBE94D87EBDE66A6040EE438">
|
||
<label id="5B730D12F763C43BA3FC838B02A02AD1">Figure 6.</label>
|
||
</paragraph>
|
||
<paragraph id="0DEF715B28C3E18DBE4059B9A7D77894">
|
||
Morphological details of
|
||
<taxonomicName id="6655517B6708332E67F86B055E6B171B" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="59C6B7C187060308272BE21AC2095E71" italics="true">Nitidocolpus</emphasis>
|
||
</taxonomicName>
|
||
and other relevant genera of
|
||
<taxonomicName id="984A670349033C914FBE9879B0D4CA1E" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subFamily" subFamily="Staphylininae">Staphylininae</taxonomicName>
|
||
<emphasis id="F5EA649840F207663CC580A3296D0C50" bold="true">A – C</emphasis>
|
||
<taxonomicName id="84CC23C1FD7DB520BA7517ED3550E2C6" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Bernhauer" baseAuthorityYear="1917" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="columbinus">
|
||
<emphasis id="99CCC469EE1E28E9B0B0B32706625E41" italics="true">Nitidocolpus columbinus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="F9EE836C58A05D696BF307405FD7F797" bold="true">D – F</emphasis>
|
||
<taxonomicName id="A5846F4DDEF234D1EAD54EB598FD708C" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Erichson" baseAuthorityYear="1840" class="Insecta" family="Staphylinidae" genus="Cheilocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="viridulus">
|
||
<emphasis id="661EC6D1EBEA537571D788CCEEFD661F" italics="true">Cheilocolpus viridulus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BB1EF7DE8E56479C49C304AE0D6EDD41" bold="true">G – I</emphasis>
|
||
<taxonomicName id="57B8072B2B222C3B87970E0FA552F0B0" class="Insecta" family="Staphylinidae" genus="Cyrtoquedius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="undefined">
|
||
<emphasis id="B3533C78F7D5D8202C2BE8E32C7A1BEE" italics="true">Cyrtoquedius</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
<emphasis id="D62F0E3DE175D1CE5FE312DC20A4FCA8" bold="true">J – L</emphasis>
|
||
<taxonomicName id="A3DFDE4535B4050D8FA711A37D4F3667" authorityName="Sharp" authorityYear="1884" class="Insecta" family="Staphylinidae" genus="Quedionuchus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="68EE2EFDE2149FD0740C249CB8E19FC4" italics="true">Quedionuchus</emphasis>
|
||
</taxonomicName>
|
||
nr.
|
||
<emphasis id="6B97960604894D8D3A851FFAB099EF77" italics="true">impunctus</emphasis>
|
||
<emphasis id="767E3D41AF85A83B46B9313502EEF2A4" bold="true">M – O</emphasis>
|
||
<taxonomicName id="447AE49C9679F216C76A259000338F26" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F24F436346FCC20431693C3EABB41B27" italics="true">Quedius</emphasis>
|
||
</taxonomicName>
|
||
nr.
|
||
<emphasis id="FE4379700A63D3328E7F76FD17E48A2C" italics="true">advena</emphasis>
|
||
.
|
||
<emphasis id="2522CD56A3603305AFB6BFEFFB542125" bold="true">A, D, G, J, M</emphasis>
|
||
head
|
||
<emphasis id="1FEA6E02F56DBCBC39BA8E342D55C51F" bold="true">B, E, H, K, N</emphasis>
|
||
pronotum
|
||
<emphasis id="92628CB407BA27B2609E25DE95ED7CE4" bold="true">C, F, I, L, O</emphasis>
|
||
aedeagus in lateral view, the black arrow pointing down indicates the basal projection of the paramere. Abbreviations:
|
||
<abbrev id="ABBRID0EHABG" xlink_title="anterolateral clypeal punctures">ACP</abbrev>
|
||
, anterolateral clypeal puncture; AFP, anterior frontal; AMRSP, anterior marginal row of setiferous punctures; APP, additional paired punctures adjacent to AMRSP; BP, basal puncture; FCP, frontoclypeal puncture; IASP, internalmost setiferous puncture of the anterior margin; IOP, interocular punctures; IOR, infraorbital ridge; LLSP, large lateral setiferous puncture; LPSP, large posterior setiferous puncture; NR, nuchal ridge; PFP, posterior frontal puncture; POP, parocular punctures; PPDS, paired punctures on dorsal series; SAP, supra-antennal puncture;
|
||
<abbrev id="ABBRID0ELABG" xlink_title="supra-antennal ridge">SAR</abbrev>
|
||
, supra-antennal ridge; SLSP, sublateral setiferous puncture; TP, temporal puncture. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A6DFB8566092BD33BF4326FAE21B4F6E">
|
||
Thorax: Prothorax with slightly transverse pronotum (
|
||
<abbrev id="ABBRID0EWABG" xlink_title="pronotum width at the widest point">
|
||
<collectionCode id="7D0B7F98F22A501FB9933CEB73D68279" country="Thailand" name="Paleontological Collections">PW</collectionCode>
|
||
</abbrev>
|
||
/
|
||
<abbrev id="ABBRID0E1ABG" xlink_title="pronotum length along the median line">
|
||
<collectionCode id="0D9A0ACFF63020970629BC9F91A06E55" country="Czech Republic" name="Západoceské muzeum v Plzni" type="Museum">PL</collectionCode>
|
||
</abbrev>
|
||
≥ 1.1), without dorsal or sublateral series of setiferous punctures; prosternum without longitudinal keel; antesternal membrane without distinct semisclerotized patch or patches; probasisternum triangular, with narrowed lateral arms and disc protruding medially, with pair of macrosetae on the upper probasiternum; with rounded postcoxal hypomeral process, interrupted by inferior line. Mesothorax: mesoscutellum without posterior scutellar carina, without sub-basal ridge; elytra with humeral spines or spine-like setae, with even setiferous punctation on disc and epipleuron (sometimes with various setose color patterns); with setiferous punctures at apical margin of elytral suture (underside); mesanapleural sutures transverse or nearly transverse but reaching and fusing with external part of prepectus; mesobasisternum with intercoxal process narrowly pointed into sharp angle, without V-shaped projection medially; mesocoxal cavities contiguous; pericoxal ridge present and complete. Metathorax: wings present, with veins CuA and
|
||
<collectionCode id="E745D98176B724A214E705FA79796B88" country="0" name="Mohonk Preserve, Inc.">MP</collectionCode>
|
||
4 fused; metakatepisternal processes divided; metascutellar mid-longitudinal internal suture well developed (only visible in chemically cleared specimens). Legs: apical tarsomere of all legs without dorsal setae, with one empodial seta distinctly shorter than tarsal claws; protarsomeres 1–3 distinctly wider than long in both sexes, with pale adhesive setae ventrally; procoxa with internal ridge running parallel along external ridge; mesotarsi in both sexes without black comb, sometimes in males ventral side of first mesotarsomere with pale adhesive setae; mesotibiae straight; metatarsomere 1 shorter than metatarsomere 5; metatarsi shorter than metatibiae; metatrochanter apically rounded, with strong straight dorsoapical spine; metacoxae with four or fewer spines on ventral posterolateral lobe, spines on dorsomedial disc also present; basal part of metacoxae distinctly wider and more convex than apical part.
|
||
</paragraph>
|
||
<caption id="78D88F8865660CFA11D8B3CDDE718BDF" ID-DOI="10.3897/zookeys.1218.135558.figure7" ID-arpha="87E2A45E-7E9D-53B3-8BC0-134CB60F5BF4" httpUri="https://binary.pensoft.net/fig/1175480" startId="F7">
|
||
<paragraph id="F4189C9F9D210DC2FE5C1F6F5E012257">
|
||
<label id="350163FB59E349A3E470F869BFECEA9E">Figure 7.</label>
|
||
</paragraph>
|
||
<paragraph id="CF94824C59838D759DB4A02C5F9A1FA6">
|
||
Some
|
||
<taxonomicName id="C69A18DF3AC9899F86B4B60AED33874A" class="Insecta" family="Staphylinidae" genus="Cheilocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="undefined-A">
|
||
<emphasis id="3E04A7299A6F73FDAA4F3D429CB696DA" italics="true">Cheilocolpus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
<emphasis id="AB8062F2B72E8D7E96084F4093654C79" bold="true">A</emphasis>
|
||
<taxonomicName id="0D92A0D4598445940DC9DA3CA50DF3B4" authority="(Solier, 1849)" baseAuthorityName="Solier" baseAuthorityYear="1849" class="Insecta" family="Staphylinidae" genus="Cheilocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="pyrostoma">
|
||
<emphasis id="93A213354E1D5FFBD3FC3A0BDF12E292" italics="true">Ch. pyrostoma</emphasis>
|
||
(Solier, 1849)
|
||
</taxonomicName>
|
||
<emphasis id="A87E1BA87786967FBC6332664A139878" bold="true">B</emphasis>
|
||
<taxonomicName id="C230AD3BBA12D2D776352E301D06A7ED" authority="(Bernhauer, 1917)" baseAuthorityName="Bernhauer" baseAuthorityYear="1917" class="Insecta" family="Staphylinidae" genus="Cheilocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="speciosus">
|
||
<emphasis id="F3246B0196A8FE56923164DEE9AE59B7" italics="true">Ch. speciosus</emphasis>
|
||
(Bernhauer, 1917)
|
||
</taxonomicName>
|
||
<emphasis id="3663FB219F6EAD9F07ABC6C5997CFD62" bold="true">C</emphasis>
|
||
<taxonomicName id="7C170A0D39DE8252B9CE4E8E3BB85A0A" authority="(Erichson, 1840)" baseAuthorityName="Erichson" baseAuthorityYear="1840" class="Insecta" family="Staphylinidae" genus="Cheilocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="viridulus">
|
||
<emphasis id="219509DDF16400C6E8692BBB9E0F1026" italics="true">Ch. viridulus</emphasis>
|
||
(Erichson, 1840)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8846A6DC7A8681B640CC48F7D6A4D28D">
|
||
Abdomen: protergal glands well manifested as rounded cuticular acetabula; tergites III –
|
||
<collectionCode id="4CC665E78F2B9AF50387BBD37C9B2A36" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
|
||
in some species with posterior basal transverse carinae, in some species they are only with anterior carinae; on tergite VII anterior transverse basal carina not continuing to paratergites; tergites III –
|
||
<collectionCode id="1D48DA6314897872E6650664AFF77AAD" country="Norway" name="Mykotektet, National Veterinary Institute">VI</collectionCode>
|
||
with different patterns of posteromarginal large macroseta (
|
||
<collectionCode id="A0A98ECB8C236E4FFECED32B9A9A37A9">PMM</collectionCode>
|
||
) on each side, IV –
|
||
<collectionCode id="55AA58E8D57F5B89C6F0AB545687ECC0" country="Norway" name="Mykotektet, National Veterinary Institute">VI</collectionCode>
|
||
with more than one PMM per side; punctation of tergites in form of fine to moderate impressions, some species with wide glabrous areas; lateral tergal sclerites IX short and slightly laterally flattened; male sternite VIII with medial apical emargination. Aedeagus with paramere fused to median lobe only at base and very closely appressed to median lobe along entire length; base of the paramere flat medially, not projecting upwards in the middle; paramere strongly produced over apex of median lobe. Ovipositor with each second gonocoxite with one medial macroseta, without spine-like setae on outer lateral margin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E1CBG" type="distribution">
|
||
<paragraph id="A932052D6B2682886A1BB9E902DE4477">
|
||
<heading id="655317367B1DDB8BC0D94FF8803366AC" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="8B2B7BB6B5A6CDB6622355E6AD23B5DD">Neotropical Realm: Mexican Transition Zone, Central America, and northern South America.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E6CBG" type="ecology">
|
||
<paragraph id="4A9326E3A6645FDED2595A5AD149E238">
|
||
<heading id="BC1E976E7C7F98586A112B80FA557E34" reason="title">Ecology.</heading>
|
||
</paragraph>
|
||
<paragraph id="F3F64C0E96AC31DE2DD29C2CE1A17BEB">
|
||
This genus has been collected from a variety of microhabitats, including mushrooms (e. g.,
|
||
<taxonomicName id="C6772B0FEF7A627F627509FC1131D398" class="Agaricomycetes" family="Pleurotaceae" genus="Pleurotus" kingdom="Fungi" order="Agaricales" phylum="Basidiomycota" rank="species" species="undetermined">
|
||
<emphasis id="501EBBA9D9B30FA6C88FAC65C7EB3D57" italics="true">Pleurotus</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
), under decomposing logs, beneath bark, and in leaf litter found in cloud, oak, and pine forests. They are also frequently captured using flight-intercept and Malaise traps. Additionally, some species are associated with the inflorescences of plants from the families
|
||
<taxonomicName id="8C6AFC74879E488FB6E68B439AFD8891" class="Liliopsida" family="Arecaceae" kingdom="Plantae" order="Arecales" phylum="Tracheophyta" rank="family">Arecaceae</taxonomicName>
|
||
,
|
||
<taxonomicName id="DBE98E0E2C1A8B31BAAFC3B1FB8017DA" class="Magnoliopsida" family="Betulaceae" kingdom="Plantae" order="Fagales" phylum="Tracheophyta" rank="family">Betulaceae</taxonomicName>
|
||
, and
|
||
<taxonomicName id="47EC1FA8FF666CF4B8D89617AFAC9A9D" class="Liliopsida" family="Heliconiaceae" kingdom="Plantae" order="Zingiberales" phylum="Tracheophyta" rank="family">Heliconiaceae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E2DBG" type="comparisons">
|
||
<paragraph id="F72140B93A3C53D3D5B59B092818ED8A">
|
||
<heading id="080DB3688E88F37CA97B0D209790DCCF" reason="title">Comparison.</heading>
|
||
</paragraph>
|
||
<paragraph id="6F3B1860D4959B06D93B1008D3D00E26">
|
||
<taxonomicName id="3A8F0A96A0C4DA07170CDAE8005353D6" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="FCA67D1199B61CC44C308D8D19435E80" italics="true">Nitidocolpus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="97D9B55EDD3663AED455F326B0DF2402" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
can be distinguished from all other genera of
|
||
<taxonomicName id="BD73FE2C777459746E06BB3FACD423EF" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subFamily" subFamily="Staphylininae">Staphylininae</taxonomicName>
|
||
by the combination of characters mentioned in the diagnosis.
|
||
<taxonomicName id="88AB3BD8A9A96606EAC34CF64D3849D7" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="435E9FE939BB8BFF3D77B1EE6E77E7DB" italics="true">Nitidocolpus</emphasis>
|
||
</taxonomicName>
|
||
is further distinguished from the superficially similar
|
||
<taxonomicName id="795C9C6A049629D939D404AC63F2D62C" authorityName="Kraatz" authorityYear="1857" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subTribe" subTribe="Quediina">Quediina</taxonomicName>
|
||
and Cytoquediina (Fig.
|
||
<figureCitation id="74ADC8772D1A72B0FAFD6832459B53A5" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Morphological details of Nitidocolpus and other relevant genera of Staphylininae A – C Nitidocolpus columbinus D – F Cheilocolpus viridulus G – I Cyrtoquedius sp. J – L Quedionuchus nr. impunctus M – O Quedius nr. advena. A, D, G, J, M head B, E, H, K, N pronotum C, F, I, L, O aedeagus in lateral view, the black arrow pointing down indicates the basal projection of the paramere. Abbreviations: ACP, anterolateral clypeal puncture; AFP, anterior frontal; AMRSP, anterior marginal row of setiferous punctures; APP, additional paired punctures adjacent to AMRSP; BP, basal puncture; FCP, frontoclypeal puncture; IASP, internalmost setiferous puncture of the anterior margin; IOP, interocular punctures; IOR, infraorbital ridge; LLSP, large lateral setiferous puncture; LPSP, large posterior setiferous puncture; NR, nuchal ridge; PFP, posterior frontal puncture; POP, parocular punctures; PPDS, paired punctures on dorsal series; SAP, supra-antennal puncture; SAR, supra-antennal ridge; SLSP, sublateral setiferous puncture; TP, temporal puncture. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure6" httpUri="https://binary.pensoft.net/fig/1175479">6</figureCitation>
|
||
) by the lack of supra-antennal punctures, the presence of a single carina on the mesoscutellar shield (posterior carina absent), by hind wings with fused CuA and
|
||
<collectionCode id="B0DB2AB56106E9B011081BCF408FBD8D" country="0" name="Mohonk Preserve, Inc.">MP</collectionCode>
|
||
4 veins, by a single empodial setae, and by the base of the paramere, which has no upward or forward projection in the middle. Compared to other Neotropical genera that resemble
|
||
<taxonomicName id="8454F70F9A4FF7D73C1CDD8E28399AB8" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C6B9B2F16BE73F7FA2D9058CD0BAB895" italics="true">Nitidocolpus</emphasis>
|
||
</taxonomicName>
|
||
, such as
|
||
<taxonomicName id="E2233370A5C5BBBAF8EAADB696E75A9B" class="Insecta" family="Staphylinidae" genus="Cheilocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FF5B0B7D24633D17582BD425B24695A0" italics="true">Cheilocolpus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3BFEB3AF32BAC825042B2E3B607A0742" authority="Sharp, 1884" authorityName="Sharp" authorityYear="1884" class="Insecta" family="Staphylinidae" genus="Mimosticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B179E17168F0597B3654FE2FBFBE0E1E" italics="true">Mimosticus</emphasis>
|
||
Sharp, 1884
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="67B2DF18671FBE77BE7AD27EA1D641EA" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Heterothops" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F20FE31BF51626F4B5662A329B30020B" italics="true">Heterothops</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="80E5830DE8E856B20E49F16B660E7C42" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1ED2149E8B18CD721229212FD05B6AD3" italics="true">Nitidocolpus</emphasis>
|
||
</taxonomicName>
|
||
is distinguished by the absence of paraocular punctures on the head, the lack of setiferous punctures in or near the center of the pronotal disc, and the presence of postcoxal hypomeral processes. In some species of
|
||
<taxonomicName id="850423B305E76A2CDD11C9CD6D5BC463" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="624A0D3B8DA38B98A64D92741B77CF02" italics="true">Nitidocolpus</emphasis>
|
||
</taxonomicName>
|
||
, such as
|
||
<taxonomicName id="7989A8C757B71CFD4EFAB23C2DF20364" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1884" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="laeticulus">
|
||
<emphasis id="5021AA8071EB390A90929C3FCC18FD6A" italics="true">N. laeticulus</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="8CA7E3E8693119F77FE2B0883B2531A4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Some Nitidocolpus species and characters A N. illatus (Sharp, 1884) B N. laeticulus (Sharp, 1884) C N. germaini (Bernhauer, 1917) D N. aurofasciatus (Bernhauer, 1917) E N. illatus, pterothoraces F N. columbinus (Bernhauer, 1917), metacoxa and metatrochanter, dissected right leg in dorsal view (side faced to the abdomen). Abbreviations: EXPR, external part of prepectus; INPR, internal part of prepectus; MIV, mesobasisternal intercoxal process with V-shaped projection medially; MNPS, mesanapleural sutures; MTMS, metacoxa with dorsomedial spine; MTTAS, metatrochanter with dorsoapical spine. Photo B by J. Jenkins Shaw. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure5" httpUri="https://binary.pensoft.net/fig/1175478">5 B</figureCitation>
|
||
) and
|
||
<taxonomicName id="44C01BF14F1ECBCFD243A9DEEB1B1184" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Bernhauer" baseAuthorityYear="1917" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="aurofasciatus">
|
||
<emphasis id="E52ABCC48CDE491C2C9D2DFA568C72EC" italics="true">N. aurofasciatus</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="191E8248A9883B040ADECC399D35DAD2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Some Nitidocolpus species and characters A N. illatus (Sharp, 1884) B N. laeticulus (Sharp, 1884) C N. germaini (Bernhauer, 1917) D N. aurofasciatus (Bernhauer, 1917) E N. illatus, pterothoraces F N. columbinus (Bernhauer, 1917), metacoxa and metatrochanter, dissected right leg in dorsal view (side faced to the abdomen). Abbreviations: EXPR, external part of prepectus; INPR, internal part of prepectus; MIV, mesobasisternal intercoxal process with V-shaped projection medially; MNPS, mesanapleural sutures; MTMS, metacoxa with dorsomedial spine; MTTAS, metatrochanter with dorsoapical spine. Photo B by J. Jenkins Shaw. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure5" httpUri="https://binary.pensoft.net/fig/1175478">5 D</figureCitation>
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), the basal punctures have shifted anteriorly and thus may be erroneously considered paraocular punctures. However, they can be easily distinguished as basal punctures, because they are the only punctures located near the posterior frontal punctures, aside from the temporal punctures. In other species of
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<taxonomicName id="EEBCC3C617091F98905B72B4D0D8C716" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
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||
<emphasis id="7D7C8C91796B5AC0F7880FF929B42D16" italics="true">Nitidocolpus</emphasis>
|
||
</taxonomicName>
|
||
, such as
|
||
<taxonomicName id="973C7890DD05BF463ABB532F5C0A4799" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1884" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="illatus">
|
||
<emphasis id="73147A09965C3B29E0E6BED64E2151A9" italics="true">N. illatus</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="C7830F06D3D3D687D48411EF774D9859" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Some Nitidocolpus species and characters A N. illatus (Sharp, 1884) B N. laeticulus (Sharp, 1884) C N. germaini (Bernhauer, 1917) D N. aurofasciatus (Bernhauer, 1917) E N. illatus, pterothoraces F N. columbinus (Bernhauer, 1917), metacoxa and metatrochanter, dissected right leg in dorsal view (side faced to the abdomen). Abbreviations: EXPR, external part of prepectus; INPR, internal part of prepectus; MIV, mesobasisternal intercoxal process with V-shaped projection medially; MNPS, mesanapleural sutures; MTMS, metacoxa with dorsomedial spine; MTTAS, metatrochanter with dorsoapical spine. Photo B by J. Jenkins Shaw. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure5" httpUri="https://binary.pensoft.net/fig/1175478">5 A</figureCitation>
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||
), the innermost macrosetae on the anterior margin maybe erroneously considered as a puncture of the dorsal row because it is located far from the anterior margin of pronotum.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EBIBG" type="etymology">
|
||
<paragraph id="8BE992AB09E57F707CF44707183F3FF9">
|
||
<heading id="D3E5A76E4072583447FFC6C84BE6FF0B" reason="title">Etymology.</heading>
|
||
</paragraph>
|
||
<paragraph id="97ED032E54B619B1999BB0235033A7C3">The name is derived from the Latin words “nitidus” and “colpus”, which mean “shiny” and “hit”, respectively. The name refers to the great clarity with which the head and pronotum punctures are seen. The gender is masculine.</paragraph>
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||
</subSubSection>
|
||
</treatment>
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||
</document> |