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data/22/54/B3/2254B3092EC155C291F2154CDD92BBD3.xml
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<document id="01437B1BD5EC063A47FA8F4D8AC81362" ID-DOI="10.3897/zookeys.1218.135558" ID-ZooBank="8C9034A6-1E8D-4FF6-8721-004826BB7797" ID-publisher-id="135558" URI-arpha="6EB9F9E6-D286-5BF4-AD5D-7B66E7EB7FD9" URI-zoobank="http://zoobank.org/8C9034A6-1E8D-4FF6-8721-004826BB7797" XM.bibliography_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1731693946473" checkinUser="pensoft" docAuthor="Reyes-Hernández, José L. & Solodovnikov, Alexey" docDate="2024" docId="2254B3092EC155C291F2154CDD92BBD3" docLanguage="en" docName="ZooKeys 1218: 81-98" docOrigin="ZooKeys 1218" docSource="https://zookeys.pensoft.net/article/135558/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Chiquiticus Reyes-Hernández & Solodovnikov, 2024, gen. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="98" masterDocId="6EB9F9E6D2865BF4AD5D7B66E7EB7FD9" masterDocTitle="Two new rove beetle genera in Staphylininae that reduce “ Heterothops ” and “ Quedius ” taxonomic wastebaskets (Coleoptera, Staphylinidae)" masterLastPageNumber="98" masterPageNumber="81" pageNumber="81" updateTime="1731698241112" updateUser="admin">
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<mods:titleInfo id="E28B30654C287530B39E1F2FE8E6F946">
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<mods:title id="6611D72DCFFA4B2DC666AEE2E3C8D2A4">Two new rove beetle genera in Staphylininae that reduce “ Heterothops ” and “ Quedius ” taxonomic wastebaskets (Coleoptera, Staphylinidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="2FBDD9B37363AE2828BC0B35F31A4D8B" type="personal">
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<mods:role id="F62DCD76A60D3BAFF2B4E6D124DC4087">
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<mods:roleTerm id="E0639BE048F90DE22CD7ADA1D9192CB2">Author</mods:roleTerm>
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||||
</mods:role>
|
||||
<mods:namePart id="096794AD3190091530E7E55D9F8F683A">Reyes-Hernández, José L.</mods:namePart>
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||||
<mods:nameIdentifier id="89D5CF7AD867980178D8F363BA5A8BAB" type="ORCID">0000-0002-4726-4439</mods:nameIdentifier>
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<mods:affiliation id="825A0839BB7456D3C5041776389EFF18">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:name id="406403BF63D110DEC50F200E48555B69" type="personal">
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<mods:roleTerm id="602FFA6DF6CB6E4CD314C3EBB343FEB3">Author</mods:roleTerm>
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</mods:role>
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||||
<mods:namePart id="373424EB68ED3C4C7A75F68B5C2A32A6">Solodovnikov, Alexey</mods:namePart>
|
||||
<mods:nameIdentifier id="6E9779687C614451F68E91198A63071C" type="ORCID">0000-0003-2031-849X</mods:nameIdentifier>
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<mods:affiliation id="49E02981A8CE4EED6E41EB5ABBB6CAF6">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:typeOfResource id="855A0F2FB16D12D79DE1FCC078C76E37">text</mods:typeOfResource>
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<mods:titleInfo id="F85027308C2999479BBAD95E079CBFFF">
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<mods:title id="0DE4D783141C53A6E053258CAD7BBCF1">ZooKeys</mods:title>
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<mods:part id="26D10AE1F208952432815725CF01EB71">
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<mods:date id="8FA4962638518CD214FF8C79F66242B4">2024</mods:date>
|
||||
<mods:detail id="A9B3ACB9D4D3909222B5176ABE59AE58" type="pubDate">
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<mods:number id="82798EBC64E3008E65D3510A8DB83E7C">2024-11-15</mods:number>
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<mods:detail id="4EFD59FB1008A5D13D24402E6528D152" type="volume">
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<mods:number id="EA49FD67355A00679CD7F67C3600471F">1218</mods:number>
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<mods:classification id="253F513ACFD559B62B15F9B5B4B3EBEF">journal article</mods:classification>
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<mods:identifier id="7E09252C0D1940D1763B9F96AF875905" type="DOI">10.3897/zookeys.1218.135558</mods:identifier>
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<mods:identifier id="FE9247E8B6AA251638489A1245F46A4F" type="ZooBank">8C9034A6-1E8D-4FF6-8721-004826BB7797</mods:identifier>
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<treatment id="2254B3092EC155C291F2154CDD92BBD3" ID-arpha="2254B309-2EC1-55C2-91F2-154CDD92BBD3" ID-zoobank="https://zoobank.org/4BCF12DF-8231-4269-B13B-917E194D43B9" LSID="urn:lsid:plazi:treatment:2254B3092EC155C291F2154CDD92BBD3" LSID-ZBK="urn:lsid:zoobank.org:act:4BCF12DF-8231-4269-B13B-917E194D43B9" httpUri="http://treatment.plazi.org/id/2254B3092EC155C291F2154CDD92BBD3">
|
||||
<subSubSection id="010E8068AC946F2FADE843D90A4B4841" type="nomenclature">
|
||||
<paragraph id="AEEB44897824D85F60008A6B03AD99F1">
|
||||
<heading id="43A44E6C9C866628C878167B3DB41BE9" reason="label">Genus</heading>
|
||||
<taxonomicName id="F0448DC5ECF0D5604AD2E5DED2789BA5" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="85374FDDC21F200E57C38C41CF6D7910" italics="true">Chiquiticus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="B8F8C25E409BC187F3EBE68015ADB592" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="20649FFCE26DC622E3CDC7E73825F50C" type="description">
|
||||
<paragraph id="679BAFE4C49896FF954D8DBA242978DE">
|
||||
<figureCitation id="95DC55A5AAE26DE82CB04F15EEF5BCC6" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Some Chiquiticus species and characters A C. sp. nr. pusio B C. rambouseki (Blackwelder, 1943) C C. pusio (LeConte, J. L., 1863), head ventral view D C. pusio, abdomen E C. pusio, prosternum F C. pusio, mesothorax. Abbreviations: ASP, antesternal plate; EXPR, external part of prepectus; FS, profurcasternum; ICC, intercoxal carina of the probasisternum; INPR, internal part of prepectus; IOR, infraorbital ridge; LBS, lower probasisternum; MMA, medial macroseta; MNPS, mesanapleural sutures; PMA, posteromarginal large macrosetae; PMR, postmandibular ridge; POR, postoccipital ridge; UBS, upper probasisternum; VBR, ventral basal ridge. Scale bars: 1 mm (A, B, D); 0.1 mm (C, E, F)." figureDoi="10.3897/zookeys.1218.135558.figure1" httpUri="https://binary.pensoft.net/fig/1175474">Figs 1</figureCitation>
|
||||
,
|
||||
<figureCitation id="1269D0D368A8166396999A18C42FB828" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Chiquiticus arizonicus (Smetana, 1971) (Holotype # CNC 935711) A dorsal habitus B head C pronotum. Abbreviations: CATA, connection of the anterior tentorial arms (frontoclypeal “ suture ”); PPDS, paired punctures on dorsal series. Photos of # CNC 935711 (J. Buffam, Canadian National Collection of Insects, Arachnids and Nematodes)." figureDoi="10.3897/zookeys.1218.135558.figure2" httpUri="https://binary.pensoft.net/fig/1175475">2</figureCitation>
|
||||
,
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<figureCitation id="C4E583A1BB696BA23F5CF377260EBED4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. † Chiquiticus cornelli (Chatzimanolis & Engel, 2013) (paratype AMNH DR- 10-1214) A dorsal habitus B head C pronotum. Abbreviations: CATA, connection of the anterior tentorial arms (frontoclypeal “ suture ”); PPDS, paired punctures on dorsal series. Photo of AMNH DR- 10-1214 (A. Pierwola, American Museum of Natural History). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1218.135558.figure3" httpUri="https://binary.pensoft.net/fig/1175476">3</figureCitation>
|
||||
,
|
||||
<figureCitation id="08E8065D2F108AFDC78CFD3668D21D8D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Morphological details of Chiquiticus and Heterothops A – C Chiquiticus sp. aff. pusio from Panama D – F Heterothops sp. undescribed from Costa Rica A, D head B, E pronotum C, F aedeagus in lateral view, the scale to the right is equal to 1 mm. Abbreviations: ACP, anterolateral clypeal puncture; AFP, anterior frontal; AMRS, anterior marginal row of setiferous punctures; BP, basal puncture; CATA, connection of the anterior tentorial arms (frontoclypeal “ suture ”); FCP, frontoclypeal puncture; IASP, internalmost setiferous puncture of the anterior margin; IOR, infraorbital ridge; LLSP, large lateral setiferous puncture; LPSP, large posterior setiferous puncture; NR, nuchal ridge; PFP, posterior frontal puncture; POP, parocular punctures; PPDS, paired punctures on dorsal series; SAR, supra-antennal ridge; SLSP, sublateral setiferous puncture; TP, temporal puncture." figureDoi="10.3897/zookeys.1218.135558.figure4" httpUri="https://binary.pensoft.net/fig/1175477">4 A – C</figureCitation>
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</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E4NAE" type="type_taxon">
|
||||
<paragraph id="3216391FA72BE6A5EFD1FB1227C772E7">
|
||||
<heading id="DC8D7546DA8E2A7EBF23BF0665C7467F" reason="title">
|
||||
<typeStatus id="691F13A26B0921BD2CFC3F7C52418273">Type</typeStatus>
|
||||
species.
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="78EAB27B4A65733016F593298439D688">
|
||||
<taxonomicName id="96460478E3EE456B5297DB5EAD24349A" authority="J. L. LeConte, 1863" authorityName="J. L. LeConte" authorityYear="1863" class="Insecta" family="Staphylinidae" genus="Heterothops" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="pusio">
|
||||
<emphasis id="4F5B22297BDA44E4AFFDE425843B6528" italics="true">Heterothops pusio</emphasis>
|
||||
J. L. LeConte, 1863
|
||||
</taxonomicName>
|
||||
, here designated.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EOOAE" type="taxon_list">
|
||||
<paragraph id="CB83B046457B63919EA7A28FFCDE8923">
|
||||
<heading id="5AF29CFC816CFA591B97CEFF5CD4F815" reason="title">Included species.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="EB8589726E12EAA071E2EA7E8FE1FE09">
|
||||
<taxonomicName id="5185353AD070A000291E1D66A67607F3" authority="(Smetana, 1971)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Smetana" baseAuthorityYear="1971" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="arizonicus" status="comb. nov.">
|
||||
<emphasis id="731E6C12A1891344BA2D31114018ADF6" italics="true">Chiquiticus arizonicus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="465A5F1EB62E2450F1241A1556479C89" DOI="10.4039/entm10379fv" author="Smetana A" refId="B19" refString="Smetana A (1971) Revision of the tribe Quediini of America north of Mexico (Coleoptera: Staphylinidae). The Memoirs of the Entomological Society of Canada 103 (S 79): 1 - 303. https: // doi. org / 10.4039 / entm 10379 fv" year="1971">Smetana, 1971</bibRefCitation>
|
||||
)
|
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</taxonomicName>
|
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,
|
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<taxonomicNameLabel id="02BCA78BC5CB28C55EABA5419DD60B4C" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
ex.
|
||||
<taxonomicName id="3F2A35A9B4BA6F045A2B595C637B824E" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Heterothops" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="95E7531B57DD434A360183A13CC91C0B" italics="true">Heterothops</emphasis>
|
||||
</taxonomicName>
|
||||
[
|
||||
<typeStatus id="4C5192FD8887BE131903AEED7BD140E3" type="holotype">holotype</typeStatus>
|
||||
and
|
||||
<specimenCount id="43F14A86A9D7961A2D89BEAD41D9D28A" count="2" type="generic" typeStatus="paratype">2 paratypes</specimenCount>
|
||||
from
|
||||
<named-content id="73B92C7A232CA3C40F9C480E58EF42DE" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/canadian-national-collection-insects-arachnids-and-nematodes" xlink_title="Canadian National Collection, Ontario, Canada">
|
||||
<collectionCode id="EDD4AABC747C168E87E2FF4686178F3A" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" type="">CNC</collectionCode>
|
||||
</named-content>
|
||||
examined];
|
||||
<taxonomicName id="F48D9569BA9B80DD8EF2357DCEF96269" authority="(Smetana, 1971)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Smetana" baseAuthorityYear="1971" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="campbelli" status="comb. nov.">
|
||||
<emphasis id="71FD28EF150530FC9DA0C29A67B85B2F" italics="true">Chiquiticus campbelli</emphasis>
|
||||
(
|
||||
<bibRefCitation id="D9B863AA0203F8DB12682177B47B2486" DOI="10.4039/entm10379fv" author="Smetana A" refId="B19" refString="Smetana A (1971) Revision of the tribe Quediini of America north of Mexico (Coleoptera: Staphylinidae). The Memoirs of the Entomological Society of Canada 103 (S 79): 1 - 303. https: // doi. org / 10.4039 / entm 10379 fv" year="1971">Smetana, 1971</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="BC6808A7EBF1244F385A5B45B1D20ECD" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
ex.
|
||||
<taxonomicName id="D3BE0429957ED4D057F1026A03B5C170" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Heterothops" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="6D633B1716C2EAF000858861E394CAB9" italics="true">Heterothops</emphasis>
|
||||
</taxonomicName>
|
||||
[
|
||||
<typeStatus id="07167D3C113E6DE6D02A7E96F5E06840" type="holotype">holotype</typeStatus>
|
||||
, and
|
||||
<specimenCount id="A4DE9B8B04B51515D7564CEC83C80EA8" count="7" type="generic" typeStatus="paratype">7 paratypes</specimenCount>
|
||||
from
|
||||
<named-content id="939132ABF6B5FAB3FEC3076CF3FC5FD1" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/canadian-national-collection-insects-arachnids-and-nematodes" xlink_title="Canadian National Collection, Ontario, Canada">
|
||||
<collectionCode id="044C9E4C076F48017033630A2E66CE6E" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" type="">CNC</collectionCode>
|
||||
</named-content>
|
||||
examined]; †
|
||||
<taxonomicName id="A7311BCC122C4CD80478EE88EE65FE1A" authority="(Chatzimanolis & Engel, 2013)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Chatzimanolis & Engel" baseAuthorityYear="2013" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="cornelli" status="comb. nov.">
|
||||
<emphasis id="CB90FAA85A19B5F648D3FAEE9502AAED" italics="true">Chiquiticus cornelli</emphasis>
|
||||
(
|
||||
<bibRefCitation id="D2CBACDBFA12B4BC71507549DF479617" DOI="10.2992/007.081.0401" author="Chatzimanolis S & Engel MS" issue="4" journalOrPublisher="Annals of Carnegie Museum" pagination="281-294" refId="B5" refString="Chatzimanolis S, Engel MS (2013) The Fauna of Staphylininae in Dominican Amber (Coleoptera: Staphylinidae). Annals of Carnegie Museum 81 (4): 281 - 294. https: // doi. org / 10.2992 / 007.081. 0401" title="The Fauna of Staphylininae in Dominican Amber (Coleoptera: Staphylinidae)." volume="81" year="2013">Chatzimanolis & Engel, 2013</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="B79B1848577BE31A5E7239B9E6175807" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
ex.
|
||||
<taxonomicName id="C76B225D979422F93A60F4B20518EE61" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Heterothops" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="4676EF18DEAD48891E40168630BBB49B" italics="true">Heterothops</emphasis>
|
||||
</taxonomicName>
|
||||
[photos of the
|
||||
<typeStatus id="D8BAB844A99C07AB65B1DCAFC11C473E" type="holotype">holotype</typeStatus>
|
||||
and
|
||||
<specimenCount id="BF23038157C6DF6C250BA3BE2F6C3BA0" count="1" type="generic" typeStatus="paratype">one paratype</specimenCount>
|
||||
from
|
||||
<named-content id="BAC54FCBAB0EBC084A508B8AC34848B9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/american-museum-natural-history" xlink_title="American Museum of Natural History, New York, USA">
|
||||
<collectionCode id="804D180084D8B96E16F495D469E78969" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" type="">AMNH</collectionCode>
|
||||
</named-content>
|
||||
examined];
|
||||
<taxonomicName id="CFE3FC7A5270C270B44AB07108FCAF50" authority="(Smetana, 1971)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Smetana" baseAuthorityYear="1971" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="gemellus" status="comb. nov.">
|
||||
<emphasis id="5A05329CA26FD07F3C29586FC4113801" italics="true">Chiquiticus gemellus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="B7426F6740EF60D6912E482FB645CD3E" DOI="10.4039/entm10379fv" author="Smetana A" refId="B19" refString="Smetana A (1971) Revision of the tribe Quediini of America north of Mexico (Coleoptera: Staphylinidae). The Memoirs of the Entomological Society of Canada 103 (S 79): 1 - 303. https: // doi. org / 10.4039 / entm 10379 fv" year="1971">Smetana, 1971</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="4ED1481174192BA4986BA59AAAB585E6" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
ex.
|
||||
<taxonomicName id="1505CEF78CA46C1A0287D61FDE3714BC" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Heterothops" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="EA7AFC1D86B0E80E67DE33D5FFD87F58" italics="true">Heterothops</emphasis>
|
||||
</taxonomicName>
|
||||
[
|
||||
<typeStatus id="199013329049504649352640524A4F31" type="holotype">holotype</typeStatus>
|
||||
from
|
||||
<named-content id="75F6461A191FD90A07F4221AD0627138" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-comparative-zoology" xlink_title="Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, USA">
|
||||
<collectionCode id="609367FCF0140F7B03A1B18AF8F998E3" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" type="Museum">MCZ</collectionCode>
|
||||
</named-content>
|
||||
and
|
||||
<specimenCount id="7189BB9578E25AC519E3FFB10FE0C219" count="2" type="generic" typeStatus="paratype">2 paratypes</specimenCount>
|
||||
from
|
||||
<named-content id="61B6DC5E23B8444DA185EF0BC94632D2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/canadian-national-collection-insects-arachnids-and-nematodes" xlink_title="Canadian National Collection, Ontario, Canada">
|
||||
<collectionCode id="BA5462690D09DEF8F259AA8241705029" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" type="">CNC</collectionCode>
|
||||
</named-content>
|
||||
2 examined]; †
|
||||
<taxonomicName id="2FA70216BC2796B911C8633F10BBB66B" authority="(Chatzimanolis & Engel, 2013)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Chatzimanolis & Engel" baseAuthorityYear="2013" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="infernalis" status="comb. nov.">
|
||||
<emphasis id="7ECC9F2AA461F7F3B9902D95081056FE" italics="true">Chiquiticus infernalis</emphasis>
|
||||
(
|
||||
<bibRefCitation id="1C8723983B8493BAC681BFD5909FB066" DOI="10.2992/007.081.0401" author="Chatzimanolis S & Engel MS" issue="4" journalOrPublisher="Annals of Carnegie Museum" pagination="281-294" refId="B5" refString="Chatzimanolis S, Engel MS (2013) The Fauna of Staphylininae in Dominican Amber (Coleoptera: Staphylinidae). Annals of Carnegie Museum 81 (4): 281 - 294. https: // doi. org / 10.2992 / 007.081. 0401" title="The Fauna of Staphylininae in Dominican Amber (Coleoptera: Staphylinidae)." volume="81" year="2013">Chatzimanolis & Engel, 2013</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="1F444364B3C2BBAE4E29A6D9C5823417" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
ex.
|
||||
<taxonomicName id="008EFEF0761B3218849B5CE2F11A1E45" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Heterothops" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="CA0520135DFACEEC2E46947B2F3A087F" italics="true">Heterothops</emphasis>
|
||||
</taxonomicName>
|
||||
[moved to
|
||||
<taxonomicName id="004C6D971976363E4089528AEDC114CE" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="0DF80EB6B71FC30FDAA6BECFBA3ACB2E" italics="true">Chiquiticus</emphasis>
|
||||
</taxonomicName>
|
||||
based on the data in the publication];
|
||||
<taxonomicName id="9999939889B36467D268A921D0D40A68" authority="(Casey, 1886)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Casey" baseAuthorityYear="1886" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="occidentis" status="comb. nov.">
|
||||
<emphasis id="7D6687C4C6BE5A16D2F0B85E373CFFBA" italics="true">Chiquiticus occidentis</emphasis>
|
||||
(Casey, 1886)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="972CEAD8C8B7E062EF303FE3AB5FC716" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
ex.
|
||||
<taxonomicName id="1DBE2D816AA7D4A0B0450E1FDD19ED3E" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Heterothops" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="6F1C0188BC6E6D9CE8156368EFE644FC" italics="true">Heterothops</emphasis>
|
||||
</taxonomicName>
|
||||
[
|
||||
<typeStatus id="DF48ACBBAFB6EF4A015CEA8ADD963C99" type="holotype">holotype</typeStatus>
|
||||
of
|
||||
<taxonomicName id="AB9DBE412B5C7C8537205533AF3427DE" authority="Fall, 1907" authorityName="Fall" authorityYear="1907" class="Insecta" family="Staphylinidae" genus="Heterothops" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="mediocris">
|
||||
<emphasis id="BCC59C07821CB6E1B57716E3DE264CDA" italics="true">Heterothops mediocris</emphasis>
|
||||
Fall, 1907
|
||||
</taxonomicName>
|
||||
, from
|
||||
<named-content id="7E3C7FC305F531BF276D2EB54E88530F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-comparative-zoology" xlink_title="Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, USA">
|
||||
<collectionCode id="16D52C676C03E39E38F1CD51E43C64DD" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" type="Museum">MCZ</collectionCode>
|
||||
</named-content>
|
||||
examined,
|
||||
<taxonomicName id="23B63248DF0C2302803E6028D67058AF" authorityName="Fall" authorityYear="1907" class="Insecta" family="Staphylinidae" genus="Heterothops" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="mediocris">
|
||||
<emphasis id="E298386DD1458882065844061584EDC5" italics="true">H. mediocris</emphasis>
|
||||
</taxonomicName>
|
||||
is a junior synonym of
|
||||
<taxonomicName id="318DABFF3B37B6F7AB62FBBCAB70D687" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Casey" baseAuthorityYear="1886" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="occidentis">
|
||||
<emphasis id="5882719F672756A6D4405B2A9767475D" italics="true">H. occidentis</emphasis>
|
||||
</taxonomicName>
|
||||
according to
|
||||
<bibRefCitation id="FC1F8014E2E9BF8E25B64B40EE72B557" DOI="10.4039/entm10379fv" author="Smetana A" firstAuthor="Smetana" refId="B19" refString="Smetana A (1971) Revision of the tribe Quediini of America north of Mexico (Coleoptera: Staphylinidae). The Memoirs of the Entomological Society of Canada 103 (S 79): 1–303. https://doi.org/10.4039/entm10379fv" year="1971">Smetana (1971)</bibRefCitation>
|
||||
and non-type material identified by A. Smetana in
|
||||
<named-content id="77C4D9A3AD01230A2F4C8878E4E4065F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/canadian-national-collection-insects-arachnids-and-nematodes" xlink_title="Canadian National Collection, Ontario, Canada">
|
||||
<collectionCode id="0534A5D32CE240BF3FEF18498D665C99" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" type="">CNC</collectionCode>
|
||||
</named-content>
|
||||
];
|
||||
<taxonomicName id="987FBEABC3A618E4D85E1587888A3A29" authority="(LeConte, J. L., 1863)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Reyes-Hernández & Solodovnikov" baseAuthorityYear="1863" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="pusio" status="comb. nov.">
|
||||
<emphasis id="EACA2DCB1E9398D58CCD7E035BFC8D6F" italics="true">Chiquiticus pusio</emphasis>
|
||||
(LeConte, J. L., 1863)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="B3A5E6D376E36471D02530D36D9914B7" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
ex.
|
||||
<taxonomicName id="7BDF475913EB3440082168D0E907F067" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Heterothops" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="9BC5768175882C9291A63E3CB1BCCAA3" italics="true">Heterothops</emphasis>
|
||||
</taxonomicName>
|
||||
[1 presumed
|
||||
<typeStatus id="9E804C4EC2BD0DBAB568647A8A30B856" type="syntype">syntype</typeStatus>
|
||||
or at least historical specimen examined, (
|
||||
<named-content id="0DFC6F4829B4DDBC8A036097753C82E0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-comparative-zoology" xlink_title="Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, USA">
|
||||
<collectionCode id="21B5CBA9D49E9AEF0DC800DEC054CD9D" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" type="Museum">MCZ</collectionCode>
|
||||
</named-content>
|
||||
) conspecific with the type material according to
|
||||
<bibRefCitation id="02743D5F5151FB8D025F55C2089E05CD" DOI="10.4039/entm10379fv" author="Smetana A" firstAuthor="Smetana" refId="B19" refString="Smetana A (1971) Revision of the tribe Quediini of America north of Mexico (Coleoptera: Staphylinidae). The Memoirs of the Entomological Society of Canada 103 (S 79): 1–303. https://doi.org/10.4039/entm10379fv" year="1971">Smetana (1971)</bibRefCitation>
|
||||
and non-type material identified by A. Smetana in
|
||||
<named-content id="EA97A7BEB8B41EBA0099AC997C597BEB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/canadian-national-collection-insects-arachnids-and-nematodes" xlink_title="Canadian National Collection, Ontario, Canada">
|
||||
<collectionCode id="E6368CBB289AF04F7CFA4E50C8B3DCED" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" type="">CNC</collectionCode>
|
||||
</named-content>
|
||||
];
|
||||
<taxonomicName id="F4DD79DC6B654CF03C8C283C0A5780E4" authority="(Blackwelder, 1943)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Blackwelder" baseAuthorityYear="1943" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rambouseki" status="comb. nov.">
|
||||
<emphasis id="E4B767A55DA99A9BD37F7DC95135E92F" italics="true">Chiquiticus rambouseki</emphasis>
|
||||
(Blackwelder, 1943)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="057DB6A9BD1AA11B818AE689A37058A9" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
ex.
|
||||
<taxonomicName id="EB50B95ED6F9B92129509CD7A0F4EF20" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Heterothops" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="693A38D7051F1A6C9044DB6CF94A46A9" italics="true">Heterothops</emphasis>
|
||||
</taxonomicName>
|
||||
[
|
||||
<specimenCount id="9CA245D7551DFC84081861EBCB16B34D" count="1" type="generic" typeStatus="syntype">1 syntype</specimenCount>
|
||||
from
|
||||
<named-content id="97685A6024908C3F5268990FB50F8E2E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London, United Kingdom">
|
||||
<collectionCode id="19EAE90E7082D26A37545A3F8A0AE52B" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" type="Herbarium">NHM</collectionCode>
|
||||
</named-content>
|
||||
examined].
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EGWAE" type="diagnosis">
|
||||
<paragraph id="B716F59CB3411DA6610D6E60B802F69C">
|
||||
<heading id="CAC968AF4A6C9C12481447A0A3DE02C1" reason="title">Diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="F52A9FFAB79C47A59D3767118A8A77CB">
|
||||
Small
|
||||
<taxonomicName id="2EEAA84C2E54023A061E72EAA82BC2D6" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subFamily" subFamily="Staphylininae">Staphylininae</taxonomicName>
|
||||
mainly around
|
||||
<quantity id="8411DD8E5A23CA3A17B802BC96FEEE73" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" unit="mm" value="3.0" valueMax="3.5" valueMin="2.5">2.5–3.5 mm</quantity>
|
||||
long (Fig.
|
||||
<figureCitation id="B3F5D5B741A4E1F6436A7F8BD2D2CD53" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Some Chiquiticus species and characters A C. sp. nr. pusio B C. rambouseki (Blackwelder, 1943) C C. pusio (LeConte, J. L., 1863), head ventral view D C. pusio, abdomen E C. pusio, prosternum F C. pusio, mesothorax. Abbreviations: ASP, antesternal plate; EXPR, external part of prepectus; FS, profurcasternum; ICC, intercoxal carina of the probasisternum; INPR, internal part of prepectus; IOR, infraorbital ridge; LBS, lower probasisternum; MMA, medial macroseta; MNPS, mesanapleural sutures; PMA, posteromarginal large macrosetae; PMR, postmandibular ridge; POR, postoccipital ridge; UBS, upper probasisternum; VBR, ventral basal ridge. Scale bars: 1 mm (A, B, D); 0.1 mm (C, E, F)." figureDoi="10.3897/zookeys.1218.135558.figure1" httpUri="https://binary.pensoft.net/fig/1175474">1 A, B</figureCitation>
|
||||
) with subconical head with ventral basal ridge but without postgenal ridge (Fig.
|
||||
<figureCitation id="E40D99E6F17FB981050B6E5B27E3D1E3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Some Chiquiticus species and characters A C. sp. nr. pusio B C. rambouseki (Blackwelder, 1943) C C. pusio (LeConte, J. L., 1863), head ventral view D C. pusio, abdomen E C. pusio, prosternum F C. pusio, mesothorax. Abbreviations: ASP, antesternal plate; EXPR, external part of prepectus; FS, profurcasternum; ICC, intercoxal carina of the probasisternum; INPR, internal part of prepectus; IOR, infraorbital ridge; LBS, lower probasisternum; MMA, medial macroseta; MNPS, mesanapleural sutures; PMA, posteromarginal large macrosetae; PMR, postmandibular ridge; POR, postoccipital ridge; UBS, upper probasisternum; VBR, ventral basal ridge. Scale bars: 1 mm (A, B, D); 0.1 mm (C, E, F)." figureDoi="10.3897/zookeys.1218.135558.figure1" httpUri="https://binary.pensoft.net/fig/1175474">1 C</figureCitation>
|
||||
); without supra-antennal punctures (Fig.
|
||||
<figureCitation id="366BAC6897395CFD6C507BA30FEB4447" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Morphological details of Chiquiticus and Heterothops A – C Chiquiticus sp. aff. pusio from Panama D – F Heterothops sp. undescribed from Costa Rica A, D head B, E pronotum C, F aedeagus in lateral view, the scale to the right is equal to 1 mm. Abbreviations: ACP, anterolateral clypeal puncture; AFP, anterior frontal; AMRS, anterior marginal row of setiferous punctures; BP, basal puncture; CATA, connection of the anterior tentorial arms (frontoclypeal “ suture ”); FCP, frontoclypeal puncture; IASP, internalmost setiferous puncture of the anterior margin; IOR, infraorbital ridge; LLSP, large lateral setiferous puncture; LPSP, large posterior setiferous puncture; NR, nuchal ridge; PFP, posterior frontal puncture; POP, parocular punctures; PPDS, paired punctures on dorsal series; SAR, supra-antennal ridge; SLSP, sublateral setiferous puncture; TP, temporal puncture." figureDoi="10.3897/zookeys.1218.135558.figure4" httpUri="https://binary.pensoft.net/fig/1175477">4 A</figureCitation>
|
||||
); pronotum transverse (
|
||||
<abbrev id="ABBRID0E5WAE" xlink_title="pronotum width at the widest point">
|
||||
<collectionCode id="7EE5D5B517D081D5E0E0138D5F3CDE34" country="Thailand" name="Paleontological Collections">PW</collectionCode>
|
||||
</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0ECXAE" xlink_title="pronotum length along the median line">
|
||||
<collectionCode id="EC3E70FDE8BBF6A1B5BA28C5C9DB6731" country="Czech Republic" name="Západoceské muzeum v Plzni" type="Museum">PL</collectionCode>
|
||||
</abbrev>
|
||||
≥ 1.1), with its maximum width in posterior half, with paired punctures on dorsal series widely separated from each other (Fig.
|
||||
<figureCitation id="C3871C42E995C32E6480234F52D8BD8E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Morphological details of Chiquiticus and Heterothops A – C Chiquiticus sp. aff. pusio from Panama D – F Heterothops sp. undescribed from Costa Rica A, D head B, E pronotum C, F aedeagus in lateral view, the scale to the right is equal to 1 mm. Abbreviations: ACP, anterolateral clypeal puncture; AFP, anterior frontal; AMRS, anterior marginal row of setiferous punctures; BP, basal puncture; CATA, connection of the anterior tentorial arms (frontoclypeal “ suture ”); FCP, frontoclypeal puncture; IASP, internalmost setiferous puncture of the anterior margin; IOR, infraorbital ridge; LLSP, large lateral setiferous puncture; LPSP, large posterior setiferous puncture; NR, nuchal ridge; PFP, posterior frontal puncture; POP, parocular punctures; PPDS, paired punctures on dorsal series; SAR, supra-antennal ridge; SLSP, sublateral setiferous puncture; TP, temporal puncture." figureDoi="10.3897/zookeys.1218.135558.figure4" httpUri="https://binary.pensoft.net/fig/1175477">4 B</figureCitation>
|
||||
); mesanapleural sutures transverse not reaching or fusing with external part of prepectus (Fig.
|
||||
<figureCitation id="A99BB75ED52C94C558B924B4A37BF701" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Some Chiquiticus species and characters A C. sp. nr. pusio B C. rambouseki (Blackwelder, 1943) C C. pusio (LeConte, J. L., 1863), head ventral view D C. pusio, abdomen E C. pusio, prosternum F C. pusio, mesothorax. Abbreviations: ASP, antesternal plate; EXPR, external part of prepectus; FS, profurcasternum; ICC, intercoxal carina of the probasisternum; INPR, internal part of prepectus; IOR, infraorbital ridge; LBS, lower probasisternum; MMA, medial macroseta; MNPS, mesanapleural sutures; PMA, posteromarginal large macrosetae; PMR, postmandibular ridge; POR, postoccipital ridge; UBS, upper probasisternum; VBR, ventral basal ridge. Scale bars: 1 mm (A, B, D); 0.1 mm (C, E, F)." figureDoi="10.3897/zookeys.1218.135558.figure1" httpUri="https://binary.pensoft.net/fig/1175474">1 F</figureCitation>
|
||||
); males with black combs on first mesotarsomeres, but without black combs on mesotrochanters; tergites VII and VIII without broader, foliose setae in addition to usual acuminate, simple setae; tergite X not fused to lateral tergal sclerites in males.
|
||||
</paragraph>
|
||||
<caption id="7CD288897508E0264E93894FBD8337AF" ID-DOI="10.3897/zookeys.1218.135558.figure1" ID-arpha="795CF57B-E1EB-55C8-8581-E365545C615A" httpUri="https://binary.pensoft.net/fig/1175474" startId="F1">
|
||||
<paragraph id="99862F448D1D4E4518F945B2B0439E9C">
|
||||
<label id="90C5106A2D7520CD5E43EA2F5048BF11">Figure 1.</label>
|
||||
</paragraph>
|
||||
<paragraph id="B17297C9E9B75011E60FDF00D5196902">
|
||||
Some
|
||||
<taxonomicName id="398AF4AAB67A7686A21800EFA7B67E9A" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="undefined">
|
||||
<emphasis id="25F7CB88931EF5DF3FC2394D8C2AF448" italics="true">Chiquiticus</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
and characters
|
||||
<emphasis id="120E872699F910BFC8C4119D374C4371" bold="true">A</emphasis>
|
||||
<emphasis id="56485F4213ECD8B7A34374DEBC41F082" italics="true">C.</emphasis>
|
||||
sp. nr.
|
||||
<taxonomicName id="758D635866DC41CD7F3B9CEBEFA5A09D" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Reyes-Hernández & Solodovnikov" baseAuthorityYear="1863" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="pusio">
|
||||
<emphasis id="855FB211B3828C5F1D32E20699111609" italics="true">pusio</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="9DA044883EF4A490F64853FD55F9662B" bold="true">B</emphasis>
|
||||
<taxonomicName id="B21F33DF837D088A945D4012ED169EA1" authority="(Blackwelder, 1943)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Blackwelder" baseAuthorityYear="1943" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rambouseki">
|
||||
<emphasis id="817A7743AB6004F24140444ADA69789E" italics="true">C. rambouseki</emphasis>
|
||||
(Blackwelder, 1943)
|
||||
</taxonomicName>
|
||||
<emphasis id="0D08C98410F3B36D1071FE0EE47C3A52" bold="true">C</emphasis>
|
||||
<taxonomicName id="B39AED243A47C6EDF01C7C69337B577B" authority="(LeConte, J. L., 1863)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Reyes-Hernández & Solodovnikov" baseAuthorityYear="1863" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="pusio">
|
||||
<emphasis id="ED5D2887AA83FCB38DC3F70D4793D98E" italics="true">C. pusio</emphasis>
|
||||
(LeConte, J. L., 1863)
|
||||
</taxonomicName>
|
||||
, head ventral view
|
||||
<emphasis id="18565E2E82467ED292012B6D5D9C71C8" bold="true">D</emphasis>
|
||||
<taxonomicName id="5553F3C95F1D354C9765331B804D2225" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Reyes-Hernández & Solodovnikov" baseAuthorityYear="1863" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="pusio">
|
||||
<emphasis id="66269AF34304E049B20F63328B84C069" italics="true">C. pusio</emphasis>
|
||||
</taxonomicName>
|
||||
, abdomen
|
||||
<emphasis id="CAFD9E85F5E54C7C8B993F0352F49CD6" bold="true">E</emphasis>
|
||||
<taxonomicName id="0FD602755F4ADE525DF8C78ED03124EA" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Reyes-Hernández & Solodovnikov" baseAuthorityYear="1863" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="pusio">
|
||||
<emphasis id="44B95859244A0DCF7062443396320B4C" italics="true">C. pusio</emphasis>
|
||||
</taxonomicName>
|
||||
, prosternum
|
||||
<emphasis id="F21B33F30960CF741C3C145D83554074" bold="true">F</emphasis>
|
||||
<taxonomicName id="56AB8FC5AF12344BBEFC5BDA869A69AA" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Reyes-Hernández & Solodovnikov" baseAuthorityYear="1863" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="pusio">
|
||||
<emphasis id="14D6FC665F17A2450EDBC9A0B02BE341" italics="true">C. pusio</emphasis>
|
||||
</taxonomicName>
|
||||
, mesothorax. Abbreviations: ASP, antesternal plate; EXPR, external part of prepectus; FS, profurcasternum;
|
||||
<abbrev id="ABBRID0EQ1AE" xlink_title="intercoxal carina">ICC</abbrev>
|
||||
, intercoxal carina of the probasisternum; INPR, internal part of prepectus; IOR, infraorbital ridge; LBS, lower probasisternum;
|
||||
<abbrev id="ABBRID0EU1AE" xlink_title="medial macroseta">MMA</abbrev>
|
||||
, medial macroseta;
|
||||
<abbrev id="ABBRID0EY1AE" xlink_title="mesanapleural sutures">MNPS</abbrev>
|
||||
, mesanapleural sutures;
|
||||
<abbrev id="ABBRID0E31AE" xlink_title="posteromarginal large macroseta">PMA</abbrev>
|
||||
, posteromarginal large macrosetae; PMR, postmandibular ridge; POR, postoccipital ridge;
|
||||
<abbrev id="ABBRID0EA2AE" xlink_title="upper probasisternum">UBS</abbrev>
|
||||
, upper probasisternum; VBR, ventral basal ridge. Scale bars: 1 mm (
|
||||
<emphasis id="8E0B292B5203F8BDEAD3A81FC8FE761E" bold="true">A, B, D</emphasis>
|
||||
); 0.1 mm (
|
||||
<emphasis id="5A831953502F83B83083F37389839DD0" bold="true">C, E, F</emphasis>
|
||||
).
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EN2AE" type="description">
|
||||
<paragraph id="E4863406662FAAD06AC1EC341CF0457D">
|
||||
<heading id="FB18E9BB2A5F8050FBF478B416ACDCA6" reason="title">Description.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="96E584F8644EDC6AFA07BB3B44A34793">
|
||||
Body small (TL =
|
||||
<quantity id="1A47B723612CB5C559F25C912F5FBB3F" metricMagnitude="-3" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="3.9" metricValueMin="2.3" unit="mm" value="3.0999999999999996" valueMax="3.9" valueMin="2.3">2.3–3.9 mm</quantity>
|
||||
). Head (Fig.
|
||||
<figureCitation id="2C45D200C025E02631B2178DE3381D84" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Some Chiquiticus species and characters A C. sp. nr. pusio B C. rambouseki (Blackwelder, 1943) C C. pusio (LeConte, J. L., 1863), head ventral view D C. pusio, abdomen E C. pusio, prosternum F C. pusio, mesothorax. Abbreviations: ASP, antesternal plate; EXPR, external part of prepectus; FS, profurcasternum; ICC, intercoxal carina of the probasisternum; INPR, internal part of prepectus; IOR, infraorbital ridge; LBS, lower probasisternum; MMA, medial macroseta; MNPS, mesanapleural sutures; PMA, posteromarginal large macrosetae; PMR, postmandibular ridge; POR, postoccipital ridge; UBS, upper probasisternum; VBR, ventral basal ridge. Scale bars: 1 mm (A, B, D); 0.1 mm (C, E, F)." figureDoi="10.3897/zookeys.1218.135558.figure1" httpUri="https://binary.pensoft.net/fig/1175474">1 C</figureCitation>
|
||||
): frontoclypeal (epistomal) “ suture ” complete; head about as wide as long (
|
||||
<abbrev id="ABBRID0EX2AE" xlink_title="head width at the widest point, including eyes">
|
||||
<collectionCode id="BC13A1AC19F43BBB2E172493C8469BD2">HW</collectionCode>
|
||||
</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0E22AE" xlink_title="head length">HL</abbrev>
|
||||
> 0.9 but <1.1); neck from moderately (
|
||||
<abbrev id="ABBRID0E62AE" xlink_title="neck width at the widest point">
|
||||
<collectionCode id="E1A1A012FAE3C2E26BE8A04439D330B3">NW</collectionCode>
|
||||
</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0ED3AE" xlink_title="head width at the widest point, including eyes">
|
||||
<collectionCode id="0C5642E7082C1ED8057521F4DAB34E3C">HW</collectionCode>
|
||||
</abbrev>
|
||||
> 0.75 but <0.90) to distinctly wide (
|
||||
<abbrev id="ABBRID0EH3AE" xlink_title="neck width at the widest point">
|
||||
<collectionCode id="F7CA9488D47F16FEBD3060B147245082">NW</collectionCode>
|
||||
</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0EL3AE" xlink_title="head width at the widest point, including eyes">
|
||||
<collectionCode id="CC7052623163644D424E7A880609BDC0">HW</collectionCode>
|
||||
</abbrev>
|
||||
≥ 0.90); dorsal macrosetae: anterolateral clypeal punctures present; frontoclypeal punctures present; supra-antennal punctures absent; parocular punctures present as one on each side or absent; basal punctures present, single; posterior frontal punctures located posterior to temporal punctures; gular sutures separated, gula with distinct transverse basal impression; ventral basal ridge (
|
||||
<collectionCode id="BB85D17216E9A0A0F191FFB23A7B8406">VBR</collectionCode>
|
||||
) present, postgenal ridge absent; postmandibular ridge (
|
||||
<collectionCode id="590A19E030A8897A681E683B0DEBC4A6" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" type="">PMR</collectionCode>
|
||||
) extended diagonally towards gula; infraorbital ridge extends to
|
||||
<collectionCode id="17563523B4AB11A715F5185788F5AB9F" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" type="">PMR</collectionCode>
|
||||
; nuchal ridge absent or rudimentary, at most present as a linear impression but not a ridge; eye from medium large (
|
||||
<abbrev id="ABBRID0EP3AE" xlink_title="eye length in lateral view">EYL</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0ET3AE" xlink_title="head length">HL</abbrev>
|
||||
ratio> 1 / 2 but <3 / 4) to very small (
|
||||
<abbrev id="ABBRID0EX3AE" xlink_title="eye length in lateral view">EYL</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0E23AE" xlink_title="head length">HL</abbrev>
|
||||
ratio less than 3 / 10); antennomere 1 distinctly shorter than antennomeres 3 and 2 combined; male and female antennomere 11 at least two times longer than antennomere 10; left mandible with proximal, not bifid tooth; maxillary palps: palpomere 4 (apical) subulate, distinctly shorter than palpomere 2, palpomere 3 markedly dilated compared to palpomere 4; labial palps: palpomere 3 (apical) more or less cylindrical, distinctly narrower and longer than palpomere 2, parallel-sided and needle-shaped, palpomere 2 markedly dilated at apex.
|
||||
</paragraph>
|
||||
<caption id="C3212E7D64A001BB3C157B179128B275" ID-DOI="10.3897/zookeys.1218.135558.figure2" ID-arpha="BD2D51F9-1D10-5C9F-B51E-8AC92ED3D993" httpUri="https://binary.pensoft.net/fig/1175475" startId="F2">
|
||||
<paragraph id="C441835C53F73CCA3833F60B98311EC5">
|
||||
<label id="83F8667299642ED7ADDEB40D4508A728">Figure 2.</label>
|
||||
</paragraph>
|
||||
<paragraph id="F267DFFEF638B0C87950EE3D59D8712D">
|
||||
<taxonomicName id="30647AD9526A388D76070574B482A241" authority="(Smetana, 1971)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Smetana" baseAuthorityYear="1971" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="arizonicus">
|
||||
<emphasis id="575788F69404DE99AED351C2FEFBB7F8" italics="true">Chiquiticus arizonicus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="34DC07122CEDD3F78C980DC6B1412585" DOI="10.4039/entm10379fv" author="Smetana A" refId="B19" refString="Smetana A (1971) Revision of the tribe Quediini of America north of Mexico (Coleoptera: Staphylinidae). The Memoirs of the Entomological Society of Canada 103 (S 79): 1 - 303. https: // doi. org / 10.4039 / entm 10379 fv" year="1971">Smetana, 1971</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
(Holotype # CNC 935711)
|
||||
<emphasis id="454CBCF49810C19896B465719719A481" bold="true">A</emphasis>
|
||||
dorsal habitus
|
||||
<emphasis id="2B7152FBEC2E3F0DDADC2E3A5F499A20" bold="true">B</emphasis>
|
||||
head
|
||||
<emphasis id="819508FA74192A08DB76D85B3C5A4419" bold="true">C</emphasis>
|
||||
pronotum. Abbreviations:
|
||||
<abbrev id="ABBRID0EY4AE" xlink_title="connection of the anterior tentorial arms">CATA</abbrev>
|
||||
, connection of the anterior tentorial arms (frontoclypeal “ suture ”); PPDS, paired punctures on dorsal series. Photos of # CNC 935711 (J. Buffam, Canadian National Collection of Insects, Arachnids and Nematodes).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="EA6B080B9E6F5E21383F7841996E59C3" ID-DOI="10.3897/zookeys.1218.135558.figure3" ID-arpha="9C3789DD-23F3-50D2-B20E-337333F1847D" httpUri="https://binary.pensoft.net/fig/1175476" startId="F3">
|
||||
<paragraph id="75695ED62E6B949C34625EBDAADF2680">
|
||||
<label id="A4602883C93E6884FA42504D3C1E07D4">Figure 3.</label>
|
||||
</paragraph>
|
||||
<paragraph id="AB6C210A7337547730EB1D03C3C75865">
|
||||
†
|
||||
<taxonomicName id="990E491EF6706DDCF7D2F2DE29A1B30B" authority="(Chatzimanolis & Engel, 2013)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Chatzimanolis & Engel" baseAuthorityYear="2013" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="cornelli">
|
||||
<emphasis id="638BE5A350BBE852DD4369DC28C4CF87" italics="true">Chiquiticus cornelli</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EAFC962CED422CB6B3C2315EC6BAF2D7" DOI="10.2992/007.081.0401" author="Chatzimanolis S & Engel MS" issue="4" journalOrPublisher="Annals of Carnegie Museum" pagination="281-294" refId="B5" refString="Chatzimanolis S, Engel MS (2013) The Fauna of Staphylininae in Dominican Amber (Coleoptera: Staphylinidae). Annals of Carnegie Museum 81 (4): 281 - 294. https: // doi. org / 10.2992 / 007.081. 0401" title="The Fauna of Staphylininae in Dominican Amber (Coleoptera: Staphylinidae)." volume="81" year="2013">Chatzimanolis & Engel, 2013</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
(paratype
|
||||
<named-content id="5E69C5D9525194B279EDB139B0726884" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/american-museum-natural-history" xlink_title="American Museum of Natural History, New York, USA">AMNH</named-content>
|
||||
DR- 10-1214)
|
||||
<emphasis id="3D20297B0E33D23D0CF34DAEE3A15DC0" bold="true">A</emphasis>
|
||||
dorsal habitus
|
||||
<emphasis id="15726343706375C7D20B547BFCE6DEB7" bold="true">B</emphasis>
|
||||
head
|
||||
<emphasis id="770A300E35FC1B9713854C0DFE5F30CC" bold="true">C</emphasis>
|
||||
pronotum. Abbreviations:
|
||||
<abbrev id="ABBRID0E65AE" xlink_title="connection of the anterior tentorial arms">CATA</abbrev>
|
||||
, connection of the anterior tentorial arms (frontoclypeal “ suture ”); PPDS, paired punctures on dorsal series. Photo of
|
||||
<named-content id="AEF7D340E219B3BCBADB90F5AD90E447" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/american-museum-natural-history" xlink_title="American Museum of Natural History, New York, USA">AMNH</named-content>
|
||||
DR- 10-1214 (A. Pierwola, American Museum of Natural History). Scale bars: 0.1 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="6C86460AC56D69F2E75CA9A7F9202AE1">
|
||||
Thorax (Fig.
|
||||
<figureCitation id="B6E571059C260CD15F1CB23AD15141AE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Some Chiquiticus species and characters A C. sp. nr. pusio B C. rambouseki (Blackwelder, 1943) C C. pusio (LeConte, J. L., 1863), head ventral view D C. pusio, abdomen E C. pusio, prosternum F C. pusio, mesothorax. Abbreviations: ASP, antesternal plate; EXPR, external part of prepectus; FS, profurcasternum; ICC, intercoxal carina of the probasisternum; INPR, internal part of prepectus; IOR, infraorbital ridge; LBS, lower probasisternum; MMA, medial macroseta; MNPS, mesanapleural sutures; PMA, posteromarginal large macrosetae; PMR, postmandibular ridge; POR, postoccipital ridge; UBS, upper probasisternum; VBR, ventral basal ridge. Scale bars: 1 mm (A, B, D); 0.1 mm (C, E, F)." figureDoi="10.3897/zookeys.1218.135558.figure1" httpUri="https://binary.pensoft.net/fig/1175474">1 E, F</figureCitation>
|
||||
): prothorax with slightly transverse pronotum (
|
||||
<abbrev id="ABBRID0ET6AE" xlink_title="pronotum width at the widest point">
|
||||
<collectionCode id="C67E87884756B31F9DCA9726F688B3D0" country="Thailand" name="Paleontological Collections">PW</collectionCode>
|
||||
</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0EX6AE" xlink_title="pronotum length along the median line">
|
||||
<collectionCode id="A6DE4B8708A39450DA40F69FDCBD9953" country="Czech Republic" name="Západoceské muzeum v Plzni" type="Museum">PL</collectionCode>
|
||||
</abbrev>
|
||||
≥ 1.1) with one pair of setiferous punctures in dorsal series widely separated from each other, the distance between them being about equal to the distance between them and the lateral margin; sublateral setiferous punctures present or absent; sternacostal (transverse carina) ridge medially not protruding; antesternal membrane with distinct semisclerotized plate (Fig.
|
||||
<figureCitation id="7E9EB07A143906D5001EA9E519A0DD82" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Some Chiquiticus species and characters A C. sp. nr. pusio B C. rambouseki (Blackwelder, 1943) C C. pusio (LeConte, J. L., 1863), head ventral view D C. pusio, abdomen E C. pusio, prosternum F C. pusio, mesothorax. Abbreviations: ASP, antesternal plate; EXPR, external part of prepectus; FS, profurcasternum; ICC, intercoxal carina of the probasisternum; INPR, internal part of prepectus; IOR, infraorbital ridge; LBS, lower probasisternum; MMA, medial macroseta; MNPS, mesanapleural sutures; PMA, posteromarginal large macrosetae; PMR, postmandibular ridge; POR, postoccipital ridge; UBS, upper probasisternum; VBR, ventral basal ridge. Scale bars: 1 mm (A, B, D); 0.1 mm (C, E, F)." figureDoi="10.3897/zookeys.1218.135558.figure1" httpUri="https://binary.pensoft.net/fig/1175474">1 E</figureCitation>
|
||||
); pronotum and prosternum not fused in procoxal cavity, pronotosternal suture complete in cavity; pronotal hypomeron not setose, without postcoxal hypomeral process; upper probasisternum with or without pair of macrosetae. Mesothorax: mesoscutellum without posterior scutellar carina, without sub-basal ridge; elytron without humeral spines or spine-like setae, with evenly setiferous punctation on disc and epipleuron; mesanapleural sutures (Fig.
|
||||
<figureCitation id="DE4077A4840A4D9E920B2DB8F929AD90" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Some Chiquiticus species and characters A C. sp. nr. pusio B C. rambouseki (Blackwelder, 1943) C C. pusio (LeConte, J. L., 1863), head ventral view D C. pusio, abdomen E C. pusio, prosternum F C. pusio, mesothorax. Abbreviations: ASP, antesternal plate; EXPR, external part of prepectus; FS, profurcasternum; ICC, intercoxal carina of the probasisternum; INPR, internal part of prepectus; IOR, infraorbital ridge; LBS, lower probasisternum; MMA, medial macroseta; MNPS, mesanapleural sutures; PMA, posteromarginal large macrosetae; PMR, postmandibular ridge; POR, postoccipital ridge; UBS, upper probasisternum; VBR, ventral basal ridge. Scale bars: 1 mm (A, B, D); 0.1 mm (C, E, F)." figureDoi="10.3897/zookeys.1218.135558.figure1" httpUri="https://binary.pensoft.net/fig/1175474">1 F</figureCitation>
|
||||
) transverse not reaching or fusing with external part of prepectus, medially fused or nearly touching each other; apex of mesobasisternal intercoxal process varies from sharply pointed to obtuse angle, without V-shaped projection medially; mesocoxal cavities contiguous; pericoxal ridge absent. Metathorax with wings present, with veins CuA and
|
||||
<collectionCode id="D629C49B37C979C304C24666DD8D2F15" country="0" name="Mohonk Preserve, Inc.">MP</collectionCode>
|
||||
4 fused in one vein; metakatepisternal processes divided. Legs: protarsomeres expanded in both sexes; apical tarsomere of all legs with one empodial seta distinctly shorter than tarsal claws; protibial row of laterodorsal spines always present in females but maybe absent in males in some species; ventral tibial spur resting at base of apical excavation connected to tibial margin by thin membranous region; first segment of mesotarsi in males with black comb lateroventrally, without pale adhesive setae; mesotrochanters in both sexes without black comb.
|
||||
</paragraph>
|
||||
<paragraph id="0ABEF0B3CBF35073F0CAAE2DEED60C92">
|
||||
Abdomen: tergites III –
|
||||
<collectionCode id="D82E0D4B29F9E7A026CA866B301D27C1" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
|
||||
with only anterior transverse basal carina; tergites III –
|
||||
<collectionCode id="C8CB53A0732FE2263851895E91BB13A4" country="Norway" name="Mykotektet, National Veterinary Institute">VI</collectionCode>
|
||||
with a large posteromarginal macroseta on each side (Fig.
|
||||
<figureCitation id="AB4F745D2C3C1315FA21FC014E9A2D7F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Some Chiquiticus species and characters A C. sp. nr. pusio B C. rambouseki (Blackwelder, 1943) C C. pusio (LeConte, J. L., 1863), head ventral view D C. pusio, abdomen E C. pusio, prosternum F C. pusio, mesothorax. Abbreviations: ASP, antesternal plate; EXPR, external part of prepectus; FS, profurcasternum; ICC, intercoxal carina of the probasisternum; INPR, internal part of prepectus; IOR, infraorbital ridge; LBS, lower probasisternum; MMA, medial macroseta; MNPS, mesanapleural sutures; PMA, posteromarginal large macrosetae; PMR, postmandibular ridge; POR, postoccipital ridge; UBS, upper probasisternum; VBR, ventral basal ridge. Scale bars: 1 mm (A, B, D); 0.1 mm (C, E, F)." figureDoi="10.3897/zookeys.1218.135558.figure1" httpUri="https://binary.pensoft.net/fig/1175474">1 D</figureCitation>
|
||||
); tergites VII and VIII without broader, foliose setae in addition to the usual acuminate, simple setae; anterior transverse basal carina of tergite VII not continuing to paratergites; lateral tergal sclerites IX cylindrical; male sternite VIII with medial apical emargination; male sternum IX symmetrical. Aedeagus with rounded apex.
|
||||
</paragraph>
|
||||
<caption id="A13DCC9401313B832BF67499A15B5C20" ID-DOI="10.3897/zookeys.1218.135558.figure4" ID-arpha="C012DB46-8885-59B2-BA39-5D46C1A66267" httpUri="https://binary.pensoft.net/fig/1175477" startId="F4">
|
||||
<paragraph id="0E4CC41860702EF793FC4A002F540A28">
|
||||
<label id="BA8213C0A825FCA32A3E17007EA62B79">Figure 4.</label>
|
||||
</paragraph>
|
||||
<paragraph id="4241E80E697BBCF997CF9A6DD9E5F5BC">
|
||||
Morphological details of
|
||||
<taxonomicName id="52404ED7AC239D76A31145B0CC253B6A" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="86EF72ACEF76C6BE9EB3FA629E684C1C" italics="true">Chiquiticus</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="A80D31BB19BFEC32EA5BF679C76F39AB" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Heterothops" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="FDBA3D02426E2AF29BF0A5A0FB30A838" italics="true">Heterothops</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="C34DA733BAD67766901DAF39BE9C8C5F" bold="true">A – C</emphasis>
|
||||
<taxonomicName id="D58EA3F4CD1D06A79A613D10E8F74863" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="undetermined">
|
||||
<emphasis id="57371531AEBE88723239BF8BB57B95F8" italics="true">Chiquiticus</emphasis>
|
||||
sp.
|
||||
</taxonomicName>
|
||||
aff.
|
||||
<emphasis id="88E3620AE78221BC426FF207C5DDDCC4" italics="true">pusio</emphasis>
|
||||
from Panama
|
||||
<emphasis id="79140E2E26FB4083343B6CF338D0EED3" bold="true">D – F</emphasis>
|
||||
<taxonomicName id="60F26E409D3D47D07E115A77B641B528" class="Insecta" family="Staphylinidae" genus="Heterothops" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="undetermined">
|
||||
<emphasis id="E6063AA74EAD14C256439DF5F3541242" italics="true">Heterothops</emphasis>
|
||||
sp.
|
||||
</taxonomicName>
|
||||
undescribed from Costa Rica
|
||||
<emphasis id="80BA0E8FFA402193DF3E661B6DBC13ED" bold="true">A, D</emphasis>
|
||||
head
|
||||
<emphasis id="D3ABC965B8B3911BF8A37B0379439E33" bold="true">B, E</emphasis>
|
||||
pronotum
|
||||
<emphasis id="6175E34F34DB296D89A128BE417FA9A7" bold="true">C, F</emphasis>
|
||||
aedeagus in lateral view, the scale to the right is equal to 1 mm. Abbreviations:
|
||||
<abbrev id="ABBRID0E5BAG" xlink_title="anterolateral clypeal punctures">ACP</abbrev>
|
||||
, anterolateral clypeal puncture; AFP, anterior frontal; AMRS, anterior marginal row of setiferous punctures; BP, basal puncture;
|
||||
<abbrev id="ABBRID0ECCAG" xlink_title="connection of the anterior tentorial arms">CATA</abbrev>
|
||||
, connection of the anterior tentorial arms (frontoclypeal “ suture ”); FCP, frontoclypeal puncture; IASP, internalmost setiferous puncture of the anterior margin; IOR, infraorbital ridge; LLSP, large lateral setiferous puncture; LPSP, large posterior setiferous puncture; NR, nuchal ridge; PFP, posterior frontal puncture; POP, parocular punctures; PPDS, paired punctures on dorsal series;
|
||||
<abbrev id="ABBRID0EGCAG" xlink_title="supra-antennal ridge">SAR</abbrev>
|
||||
, supra-antennal ridge; SLSP, sublateral setiferous puncture; TP, temporal puncture.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EPCAG" type="distribution">
|
||||
<paragraph id="6FF2F01290EE4BCAA8815BF230D7731B">
|
||||
<heading id="7E939BF0987FEB08BD66CAA26D8E8C77" reason="title">Distribution.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="11E2BF9E29051A5B76954900FC1E51C5">
|
||||
Nearctic region, Central America, the Caribbean. Introduced into Central Europe:
|
||||
<taxonomicName id="80E70C717E7E1B02E15B90AD1B6CF664" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Reyes-Hernández & Solodovnikov" baseAuthorityYear="1863" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="pusio">
|
||||
<emphasis id="A361AEC9C45599236CB82016864822EF" italics="true">C. pusio</emphasis>
|
||||
</taxonomicName>
|
||||
in
|
||||
<collectingCountry id="7FE9058C53298CA70A825D56B3E32466" name="Germany">Germany</collectingCountry>
|
||||
(
|
||||
<bibRefCitation id="B1CC0F0BC366746005A512CCC417238B" author="Schulke and Renner" firstAuthor="Schulke" issue="1" journalOrPublisher="Linzer biologische Beiträge" pagination="501-507" refId="B18" refString="Schülke M, Renner K (2020) Heterothops pusio Leconte, 1863 eine neue Adventivart aus Nordamerika in Deutschland (Coleoptera, Staphylinidae, Staphylininae, Amblyopinini). Linzer biologische Beiträge 52 (1): 501–507." title="Heterothops pusio Leconte, 1863 eine neue Adventivart aus Nordamerika in Deutschland (Coleoptera, Staphylinidae, Staphylininae, Amblyopinini)." volume="52" year="2020">Schülke and Renner 2020</bibRefCitation>
|
||||
) and another species related to
|
||||
<taxonomicName id="8A4E170B37AC455FDCBB22A43E2A0384" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Reyes-Hernández & Solodovnikov" baseAuthorityYear="1863" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="pusio">
|
||||
<emphasis id="98E6F8FB688C6EEF5DE2D75A90F6F49C" italics="true">C. pusio</emphasis>
|
||||
</taxonomicName>
|
||||
collected in
|
||||
<collectingCountry id="629873C2DB722C8B8CC34F8E4E180D00" name="Austria">Austria</collectingCountry>
|
||||
with a car net.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EPDAG" type="ecology">
|
||||
<paragraph id="191AD25EA05BEAF146165B72721DA558">
|
||||
<heading id="56E8AA473271391FC0EED915079B7188" reason="title">Ecology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B996309A392F2541A69C73AC4ABE60B3">
|
||||
The North American species occur in various
|
||||
<typeStatus id="F56D173CE7B13747D89FE618B49AE115">types</typeStatus>
|
||||
of debris like leaf litter and similar substrates, with some species also occurring facultatively in mammal burrows or nests (
|
||||
<taxonomicName id="BD3EE856BC1C12DE6A1C8982D70A4FDC" authority="Say & Ord, 1825" authorityName="Say & Ord" authorityYear="1825" class="Mammalia" family="Cricetidae" genus="Neotoma" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
|
||||
<emphasis id="2BA9551462D2180F07BD3B3569320421" italics="true">Neotoma</emphasis>
|
||||
Say & Ord, 1825
|
||||
</taxonomicName>
|
||||
), especially in drier regions (
|
||||
<bibRefCitation id="BA4BC9F92494B560B97A9AD0D67A9995" DOI="10.4039/entm10379fv" author="Smetana" firstAuthor="Smetana" refId="B19" refString="Smetana A (1971) Revision of the tribe Quediini of America north of Mexico (Coleoptera: Staphylinidae). The Memoirs of the Entomological Society of Canada 103 (S 79): 1–303. https://doi.org/10.4039/entm10379fv" year="1971">Smetana 1971</bibRefCitation>
|
||||
; Adam Brunke pers. comm.). In Europe, where they are introduced, they have been collected using car nets.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EAEAG" type="comparisons">
|
||||
<paragraph id="EC504E5B7295DFF845ACF63CB8932CDB">
|
||||
<heading id="2E514159F7646AD0F3151A1C64383315" reason="title">Comparison.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="876A999B0C5D1543D1A82B581AB4A661">
|
||||
<taxonomicName id="D363571BE03EFCE80D5B7362C70A05D7" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="E6DDA42E222FCE1E1762B35C3607BB4D" italics="true">Chiquiticus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="134C6C54A7B590D689E5312C66C18457" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
can be distinguished from all other genera of
|
||||
<taxonomicName id="8053CDD658945FFA55FCE5317466C43C" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subFamily" subFamily="Staphylininae">Staphylininae</taxonomicName>
|
||||
in broad sense (
|
||||
<bibRefCitation id="1543F1F031347631CA9BB8B2329EB548" DOI="10.48580/dg9ld-3gk" author="Newton" firstAuthor="Newton" refId="B14" refString="Newton A (2022) StaphBase _ 2022-12 - 12 _ 15: 41: 00 _ - 0600. In: Bánki O, Roskov Y, Döring M, Ower G, Vandepitte L, Hobern D, Remsen D, Schalk P, DeWalt RE, Keping M, Miller J, Orrell T, Aalbu R, Abbott J, Adlard R, Adriaenssens EM, Aedo C, Aescht E, Akkari N, et al. (Eds) Catalogue of Life Checklist (Aug 2022). https://doi.org/10.48580/dg9ld-3gk" year="2022">Newton 2022</bibRefCitation>
|
||||
) by the combination of characters mentioned in the diagnosis. From species close to the
|
||||
<typeStatus id="1D9168B6B388E6A6FF6128AC34D684C0">type</typeStatus>
|
||||
of
|
||||
<taxonomicName id="FC0885DFB298F606CBCF94F12F0F8D45" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Heterothops" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="F7B50F5282DF3BDF36341F7C7C2A5B53" italics="true">Heterothops</emphasis>
|
||||
</taxonomicName>
|
||||
, it is distinguished by the paired setiferous macropunctures of the dorsal series being distinctly separated from each other, by a black comb on first mesotarsomere of males, and by presence of only one posteromarginal macrosetae on each side of tergites III –
|
||||
<collectionCode id="9B3F54F92423F2D878A13C9DF5791D7D" country="Norway" name="Mykotektet, National Veterinary Institute">VI</collectionCode>
|
||||
. In species close to the
|
||||
<typeStatus id="A9ED1DF2FF8CEB86FF3A6E1A94809287">type</typeStatus>
|
||||
of
|
||||
<taxonomicName id="6E45E99FB0D5B50A3F6586D8A55F2253" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Heterothops" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="C44A71897B24DFCA81415BEB000C6898" italics="true">Heterothops</emphasis>
|
||||
</taxonomicName>
|
||||
, the paired setiferous macropunctures of the dorsal series are close together, the distance between them being distinctly less than the distance between dorsal row and respective lateral margin of pronotum. Also, in the true
|
||||
<taxonomicName id="A86E0E1D62A72629DF185E7E6B17CA69" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Heterothops" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="0AB08A03EF4575FC5A92A8B83000A37A" italics="true">Heterothops</emphasis>
|
||||
</taxonomicName>
|
||||
, males do not have a black comb on the first mesotarsomere and their pattern of posteromarginal large macrosetae on tergites III –
|
||||
<collectionCode id="C36A6068D4350C693EB7DA6BB6C9B0E9" country="Norway" name="Mykotektet, National Veterinary Institute">VI</collectionCode>
|
||||
is 1, 1, 2 and 2 per side. For more details on the differentiation between
|
||||
<taxonomicName id="799A1EDB78705733C6624BCD87A59D48" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="02200B5D9419D7F02D71158AE405C707" italics="true">Chiquiticus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="4A88C12702257A770DB9FF8ABFEE3433" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
and
|
||||
<taxonomicName id="6C0E476CDEBB509835566240F892242F" class="Insecta" family="Staphylinidae" genus="Heterothops" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="undefined">
|
||||
<emphasis id="4D2B1BDD4375FC086125CA563AAC56E4" italics="true">Heterothops</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
from Central Europe, see
|
||||
<bibRefCitation id="BDA1B9C1307A36DEDB491A2B7B6AC1F7" author="Schülke M & Renner K" firstAuthor="Schulke" issue="1" journalOrPublisher="Linzer biologische Beiträge" pagination="501-507" refId="B18" refString="Schülke M, Renner K (2020) Heterothops pusio Leconte, 1863 eine neue Adventivart aus Nordamerika in Deutschland (Coleoptera, Staphylinidae, Staphylininae, Amblyopinini). Linzer biologische Beiträge 52 (1): 501–507." title="Heterothops pusio Leconte, 1863 eine neue Adventivart aus Nordamerika in Deutschland (Coleoptera, Staphylinidae, Staphylininae, Amblyopinini)." volume="52" year="2020">Schülke and Renner (2020)</bibRefCitation>
|
||||
. Presence of a frontoclypeal “ suture ” is a rare trait in
|
||||
<taxonomicName id="D65F4F0E0D35613284ABC0FF674571B0" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subFamily" subFamily="Staphylininae">Staphylininae</taxonomicName>
|
||||
, which in
|
||||
<taxonomicName id="E7A7777259B2EAA54EFADEDDB6603B02" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="DC7A72FF4C50684BEBA7875361652F10" italics="true">Chiquiticus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="E307101AB20E17F05044C93C25ECE1AA" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
is always visible. In
|
||||
<taxonomicName id="10376242B52004D6A65390F020FDA293" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subFamily" subFamily="Staphylininae">Staphylininae</taxonomicName>
|
||||
this character is found in few basal lineages, like
|
||||
<taxonomicName id="3D747B1191C1B005259ECAAE4314D044" authority="Bernhauer, 1935" authorityName="Bernhauer" authorityYear="1935" class="Insecta" family="Staphylinidae" genus="Arrowinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="C574C283191E1768381F083BBC09B464" italics="true">Arrowinus</emphasis>
|
||||
Bernhauer, 1935
|
||||
</taxonomicName>
|
||||
, several genera of
|
||||
<taxonomicName id="1C5573FD16A05193EDD80030B635BC53" authorityName="Seevers" authorityYear="1944" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subTribe" subTribe="Amblyopinina">Amblyopinina</taxonomicName>
|
||||
including
|
||||
<taxonomicName id="82A9928A76817334643E9D307F0B5B83" authority="Germain, 1903" authorityName="Germain" authorityYear="1903" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="E2BD4288A2637FF0B46FEA8D42E8B820" italics="true">Loncovilius</emphasis>
|
||||
Germain, 1903
|
||||
</taxonomicName>
|
||||
(
|
||||
<bibRefCitation id="D306F7BF5C05F0964566EBB88D5B227B" DOI="10.1093/zoolinnean/zlad143" author="Reyes-Hernandez" etAl="et al." firstAuthor="Reyes-Hernandez" refId="B15" refString="Reyes-Hernández JL, Hansen AK, Jenkins Shaw J, Solodovnikov A (2024) Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society 202: zlad 143. https://doi.org/10.1093/zoolinnean/zlad143" year="2024">Reyes-Hernández et al. 2024</bibRefCitation>
|
||||
), and in the relatives to
|
||||
<taxonomicName id="3109DB3E2F9006E9CB8F2145A3B233BC" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="6845CFF139DA21692B9945983580EA36" italics="true">Chiquiticus</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="54D200442296369C5094E16F9749BDFE" authority="(Jenkins Shaw et al. 2020)" baseAuthorityName="Jenkins Shaw" baseAuthorityYear="2020" class="Insecta" family="Staphylinidae" genus="Amazonothops" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="FE2C32B9ACA8105E2E3E39F95F5DB561" italics="true">Amazonothops</emphasis>
|
||||
(
|
||||
<bibRefCitation id="553C0C5C077579519CB67BDFB3028D8D" DOI="10.3897/zookeys.904.48592" author="Jenkins Shaw" etAl="et al." firstAuthor="Jenkins Shaw" journalOrPublisher="ZooKeys" pagination="103-115" refId="B11" refString="Jenkins Shaw J, Orlov I, Solodovnikov A (2020) A new genus and species of Staphylininae rove beetle from the Peruvian Amazon (Coleoptera, Staphylinidae). ZooKeys 904: 103–115. https://doi.org/10.3897/zookeys.904.48592" title="A new genus and species of Staphylininae rove beetle from the Peruvian Amazon (Coleoptera, Staphylinidae)." volume="904" year="2020">Jenkins Shaw et al. 2020</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="B4E7A8B1BC448F22B9FC2F798B17F7D4" authorityName="Cameron" authorityYear="1926" class="Insecta" family="Staphylinidae" genus="Ctenandropus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="074392E703787A04DCAF3E121F7DB66A" italics="true">Ctenandropus</emphasis>
|
||||
</taxonomicName>
|
||||
. Additionally,
|
||||
<taxonomicName id="FA7BE994943FC2BE2589BDE25803B46D" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="CF9E16B00E555672CA3DC2F51899FC32" italics="true">Chiquiticus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="4CE6EE536F01DB5D2C8AEC717BC1034B" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
displays an antesternal plate, currently known as a unique character of groups such as
|
||||
<taxonomicName id="8D1CF20AE86F85AD2F941C01846CE6AC" authorityName="Bernhauer" authorityYear="1935" class="Insecta" family="Staphylinidae" genus="Arrowinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="07E0006886C3C63CF48C7D406DAF5D78" italics="true">Arrowinus</emphasis>
|
||||
</taxonomicName>
|
||||
and several tribes united in Xatholininae sensu
|
||||
<bibRefCitation id="AE779C1D2225AA0BDDE7169C68484495" DOI="10.1111/syen.12382" author="Żyła D & Solodovnikov A" firstAuthor="Zyla" issue="1" journalOrPublisher="Systematic Entomology" pagination="114-127" refId="B20" refString="Żyła D, Solodovnikov A (2020) Multilocus phylogeny defines a new classification of Staphylininae (Coleoptera, Staphylinidae), a rove beetle group with high lineage diversity. Systematic Entomology 45 (1): 114–127. https://doi.org/10.1111/syen.12382" title="Multilocus phylogeny defines a new classification of Staphylininae (Coleoptera, Staphylinidae), a rove beetle group with high lineage diversity." volume="45" year="2020">Żyła and Solodovnikov (2020)</bibRefCitation>
|
||||
. However, this plate sometimes is also found in
|
||||
<taxonomicName id="9D667D047E1B307D5DE41E7ABE243800" authorityName="Seevers" authorityYear="1944" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subTribe" subTribe="Amblyopinina">Amblyopinina</taxonomicName>
|
||||
among
|
||||
<taxonomicName id="5CF1B0CA6EF42318870FDB47611F54ED" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="tribe" tribe="Staphylinini">Staphylinini</taxonomicName>
|
||||
, for example in species that are close to the
|
||||
<typeStatus id="EE818C25AC179AACECB61F097CB3D32D">types</typeStatus>
|
||||
of the currently non-monophyletic genera
|
||||
<taxonomicName id="687B8E4518EC0EA198DA2D5D91A08795" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Heterothops" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="BACA5CF26FA262DBF5AF69B9C67B971B" italics="true">Heterothops</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="5B3F2965B1A1068A54211E35FBE3B65E" authority="(Reyes-Hernandez et al. 2024)" baseAuthorityName="Reyes-Hernandez" baseAuthorityYear="2024" class="Insecta" family="Staphylinidae" genus="Cheilocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="A80C55D1AEB41BB2C1CFE2EC2A75EB1F" italics="true">Cheilocolpus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="1EE91A6A1DEB2E63BE3CE51142078DA7" DOI="10.1093/zoolinnean/zlad143" author="Reyes-Hernandez" etAl="et al." firstAuthor="Reyes-Hernandez" refId="B15" refString="Reyes-Hernández JL, Hansen AK, Jenkins Shaw J, Solodovnikov A (2024) Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society 202: zlad 143. https://doi.org/10.1093/zoolinnean/zlad143" year="2024">Reyes-Hernández et al. 2024</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
. Furthermore,
|
||||
<taxonomicName id="BE0C7A56DABCC03AC14659DC22FD0883" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="52D34DCA1A360AC927FD1F286376A8D7" italics="true">Chiquiticus</emphasis>
|
||||
</taxonomicName>
|
||||
exhibits unique characters shared only with the related genera
|
||||
<taxonomicName id="282656B0A42580CF7D6F22D7443C882F" authorityName="Cameron" authorityYear="1926" class="Insecta" family="Staphylinidae" genus="Ctenandropus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="1E16C2F9A27E39561CB9BABAA6EB83E8" italics="true">Ctenandropus</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="7D233D4F24B204411B2DE3603364DE61" class="Insecta" family="Staphylinidae" genus="Amazonothops" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="C65DE289514499579A93C842B77F9A52" italics="true">Amazonothops</emphasis>
|
||||
</taxonomicName>
|
||||
, such as the presence of transverse mesanapleural sutures distinctly separated from the external portion of the prepectus (Fig.
|
||||
<figureCitation id="494D3CDFCD55DA97518966676073B8A6" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Some Chiquiticus species and characters A C. sp. nr. pusio B C. rambouseki (Blackwelder, 1943) C C. pusio (LeConte, J. L., 1863), head ventral view D C. pusio, abdomen E C. pusio, prosternum F C. pusio, mesothorax. Abbreviations: ASP, antesternal plate; EXPR, external part of prepectus; FS, profurcasternum; ICC, intercoxal carina of the probasisternum; INPR, internal part of prepectus; IOR, infraorbital ridge; LBS, lower probasisternum; MMA, medial macroseta; MNPS, mesanapleural sutures; PMA, posteromarginal large macrosetae; PMR, postmandibular ridge; POR, postoccipital ridge; UBS, upper probasisternum; VBR, ventral basal ridge. Scale bars: 1 mm (A, B, D); 0.1 mm (C, E, F)." figureDoi="10.3897/zookeys.1218.135558.figure1" httpUri="https://binary.pensoft.net/fig/1175474">1 F</figureCitation>
|
||||
and fig. 3 E in
|
||||
<bibRefCitation id="9F60C675AAA9E4EDBCCECD07AB250DF0" DOI="10.3897/zookeys.904.48592" author="Jenkins Shaw J & Orlov I & Solodovnikov A" etAl="et al." firstAuthor="Jenkins Shaw" journalOrPublisher="ZooKeys" pagination="103-115" refId="B11" refString="Jenkins Shaw J, Orlov I, Solodovnikov A (2020) A new genus and species of Staphylininae rove beetle from the Peruvian Amazon (Coleoptera, Staphylinidae). ZooKeys 904: 103–115. https://doi.org/10.3897/zookeys.904.48592" title="A new genus and species of Staphylininae rove beetle from the Peruvian Amazon (Coleoptera, Staphylinidae)." volume="904" year="2020">Jenkins Shaw et al. (2020)</bibRefCitation>
|
||||
misinterpreted as “ transvere ridge (TR) ”) and distinctly elongated protergal glands (fig. 3 F in
|
||||
<bibRefCitation id="88AD6EE9701171D7A577B6C3534A7B52" DOI="10.3897/zookeys.904.48592" author="Jenkins Shaw J & Orlov I & Solodovnikov A" etAl="et al." firstAuthor="Jenkins Shaw" journalOrPublisher="ZooKeys" pagination="103-115" refId="B11" refString="Jenkins Shaw J, Orlov I, Solodovnikov A (2020) A new genus and species of Staphylininae rove beetle from the Peruvian Amazon (Coleoptera, Staphylinidae). ZooKeys 904: 103–115. https://doi.org/10.3897/zookeys.904.48592" title="A new genus and species of Staphylininae rove beetle from the Peruvian Amazon (Coleoptera, Staphylinidae)." volume="904" year="2020">Jenkins Shaw et al. (2020)</bibRefCitation>
|
||||
).
|
||||
<taxonomicName id="2718CB5E20124091C0B851B5141418E4" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="3AB73EED4816E9DC3208374C5470E2F9" italics="true">Chiquiticus</emphasis>
|
||||
</taxonomicName>
|
||||
is distinguished from
|
||||
<taxonomicName id="4552CEEC006D0ABA28A4363BE0CA5BC5" authorityName="Cameron" authorityYear="1926" class="Insecta" family="Staphylinidae" genus="Ctenandropus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="A3F84926E85150990B6528BFC889D42A" italics="true">Ctenandropus</emphasis>
|
||||
</taxonomicName>
|
||||
by the subconical head with ventral basal ridge; the transverse pronotum (
|
||||
<abbrev id="ABBRID0EDLAG" xlink_title="pronotum width at the widest point">
|
||||
<collectionCode id="632594B9983FE22CD9CCFFFDC9D6B629" country="Thailand" name="Paleontological Collections">PW</collectionCode>
|
||||
</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0EHLAG" xlink_title="pronotum length along the median line">
|
||||
<collectionCode id="C7DA9641149584E79A008D8B824BF2DB" country="Czech Republic" name="Západoceské muzeum v Plzni" type="Museum">PL</collectionCode>
|
||||
</abbrev>
|
||||
≥ 1.1) with its maximum width in the posterior half; usually by presence of intercoxal carina separating the upper probasisternum and lower probasisternum; and by tergites III –
|
||||
<collectionCode id="929BC2ABC9A3ACD5CDA8E124B8F05865" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
|
||||
lacking a posterior transverse carina. In
|
||||
<taxonomicName id="151D89AF26DEADFC9103DCCAD7A623D1" authorityName="Cameron" authorityYear="1926" class="Insecta" family="Staphylinidae" genus="Ctenandropus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="8A365D79A27958018ABA62817690C302" italics="true">Ctenandropus</emphasis>
|
||||
</taxonomicName>
|
||||
, the head is subquadrate, without postgenal ridge or ventral basal ridge; the pronotum is as long as wide (
|
||||
<abbrev id="ABBRID0ESLAG" xlink_title="pronotum width at the widest point">
|
||||
<collectionCode id="FFDCA5E9240597C8A733892008A3CAEC" country="Thailand" name="Paleontological Collections">PW</collectionCode>
|
||||
</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0EWLAG" xlink_title="pronotum length along the median line">
|
||||
<collectionCode id="EE7F2E7575997DF7DA7F4408002C1F82" country="Czech Republic" name="Západoceské muzeum v Plzni" type="Museum">PL</collectionCode>
|
||||
</abbrev>
|
||||
> 0.9 but <1.1), with its maximum width in the anterior half; without intercoxal carina separating the upper probasisternum and lower probasisternum; and tergites III –
|
||||
<collectionCode id="0465933FB99FBAE2718AE520E4A14A69" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
|
||||
have a posterior transverse carina.
|
||||
<taxonomicName id="74A9D1C697A36EF7AF425E040A6281F1" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="89351FBAAFF0C987C554A361E8EEB2CA" italics="true">Chiquiticus</emphasis>
|
||||
</taxonomicName>
|
||||
is distinguished from
|
||||
<taxonomicName id="38DC544CED7FF356AF758EC881239DBE" class="Insecta" family="Staphylinidae" genus="Amazonothops" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="54CF2B44C49C15CD9563A41B337190C9" italics="true">Amazonothops</emphasis>
|
||||
</taxonomicName>
|
||||
by the absence of a black comb on the mesotrochanters of males and foliaceous setae on the tergites in both sexes, by distinctly short empodial setae, by the tergite X not fused to internal face of lateral sclerites and by paramere with rounded apex and short lateroapical setae.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EIMAG" type="etymology">
|
||||
<paragraph id="FCCC6957F5C78F48E2FDDDBBF2DE5087">
|
||||
<heading id="8A7C5AC91F79F9F0AE0A84C816755D52" reason="title">Etymology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="3D3299B946B08766ADE70A86B057BF5B">The name is derived from the Latinization of the word “Chiquitico”, which is a term used in some Hispanophone countries to refer to very small things. The gender is masculine.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ENMAG" type="New combinations notes">
|
||||
<paragraph id="5608980A631B6F9F326D5FDF0F8F7E53">
|
||||
<heading id="1958174E183936D2B2618DC249392A9B" reason="title">New combinations notes.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="152EF44BFC694201060EEC108E772115">
|
||||
The extinct species †
|
||||
<taxonomicName id="9BB28306CEB72AA0522A3D9A53489DCD" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Chatzimanolis & Engel" baseAuthorityYear="2013" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="infernalis">
|
||||
<emphasis id="7D3353C18AFA30F24D2AFA5539DD3D87" italics="true">C. infernalis</emphasis>
|
||||
</taxonomicName>
|
||||
from Dominican Amber is here restudied based on the original descriptions only. Many details important for the taxonomic placement of this species (e. g. pronotum and mesanapleural sutures) are not visible in the photos of this fossil there and probably they would be available only with the mCT examination. However, such visible features as habitus including small body size, as well as antennal bases located close to each other, very elongated last antennomere that is almost as long as the two preceding ones, oval and setose preapical maxillary palpomere that is clearly wider than the apical one, and wide neck allowed us to conclude that †
|
||||
<taxonomicName id="4C31B8B2E561B805806393639FFC3EB0" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Chatzimanolis & Engel" baseAuthorityYear="2013" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="infernalis">
|
||||
<emphasis id="326557E20DA6DB10708E46AA8A4FAB26" italics="true">C. infernalis</emphasis>
|
||||
</taxonomicName>
|
||||
is an extinct member of the new genus
|
||||
<taxonomicName id="49FD0686128A25128242E19B5D1A1F4C" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="1DF8A41E82DF4C35D40E7E15E8F3669C" italics="true">Chiquiticus</emphasis>
|
||||
</taxonomicName>
|
||||
. For †
|
||||
<taxonomicName id="1D4B422EA1D462EA7926664429B38928" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Chatzimanolis & Engel" baseAuthorityYear="2013" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="cornelli">
|
||||
<emphasis id="EE15CE23BB4CD53B82607F6176B4E3BE" italics="true">C. cornelli</emphasis>
|
||||
</taxonomicName>
|
||||
, in addition to the original description, we were able to examine better photos of the
|
||||
<typeStatus id="62A899F5D65D054B52272F7DE5BB4540" type="holotype">holotype</typeStatus>
|
||||
and
|
||||
<typeStatus id="63A955CCEBC57BBFE5A1A12A45010682" type="paratype">paratype</typeStatus>
|
||||
kindly made available at our request (Fig.
|
||||
<figureCitation id="94705B4C0541B6A7E166696767F779D3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. † Chiquiticus cornelli (Chatzimanolis & Engel, 2013) (paratype AMNH DR- 10-1214) A dorsal habitus B head C pronotum. Abbreviations: CATA, connection of the anterior tentorial arms (frontoclypeal “ suture ”); PPDS, paired punctures on dorsal series. Photo of AMNH DR- 10-1214 (A. Pierwola, American Museum of Natural History). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1218.135558.figure3" httpUri="https://binary.pensoft.net/fig/1175476">3</figureCitation>
|
||||
). In these images, in addition to the characters previously described for †
|
||||
<taxonomicName id="EC665DA374F6B4003C2C0A45DBB24C55" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Chatzimanolis & Engel" baseAuthorityYear="2013" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="infernalis">
|
||||
<emphasis id="703B336400DCAE602C1A3CE4C18A7C92" italics="true">C. infernalis</emphasis>
|
||||
</taxonomicName>
|
||||
, the connection of the anterior tentorial arms (Fig.
|
||||
<figureCitation id="2291D2DEA6683EF0084547EE98863564" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. † Chiquiticus cornelli (Chatzimanolis & Engel, 2013) (paratype AMNH DR- 10-1214) A dorsal habitus B head C pronotum. Abbreviations: CATA, connection of the anterior tentorial arms (frontoclypeal “ suture ”); PPDS, paired punctures on dorsal series. Photo of AMNH DR- 10-1214 (A. Pierwola, American Museum of Natural History). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1218.135558.figure3" httpUri="https://binary.pensoft.net/fig/1175476">3 B</figureCitation>
|
||||
) and the widely separated paired punctures in the dorsal series (Fig.
|
||||
<figureCitation id="5C861FC064FF33DDB33CF94EA5CC6049" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. † Chiquiticus cornelli (Chatzimanolis & Engel, 2013) (paratype AMNH DR- 10-1214) A dorsal habitus B head C pronotum. Abbreviations: CATA, connection of the anterior tentorial arms (frontoclypeal “ suture ”); PPDS, paired punctures on dorsal series. Photo of AMNH DR- 10-1214 (A. Pierwola, American Museum of Natural History). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1218.135558.figure3" httpUri="https://binary.pensoft.net/fig/1175476">3 C</figureCitation>
|
||||
) are visible. It should be noted that both fossil species are known only from females. Also,
|
||||
<specimenCount id="9FFB00962E7DB797582C681B59A6C2C1" count="2" type="generic">two specimens</specimenCount>
|
||||
of the extant
|
||||
<taxonomicName id="EA28576F983C4786472D5110E7DA3FE8" authority="(Blackwelder, 1943)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Blackwelder" baseAuthorityYear="1943" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rambouseki">
|
||||
<emphasis id="91813E590D43A1149A7143053296C301" italics="true">C. rambouseki</emphasis>
|
||||
(Blackwelder, 1943)
|
||||
</taxonomicName>
|
||||
that we examined were females and not in the best condition; however, the combination of their visible characters allows the species to be placed in
|
||||
<taxonomicName id="C3AFCE3AB0508C4C2537C8A2F4BEFED7" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="120BB1264917E846E229C8A4B6364480" italics="true">Chiquiticus</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EEPAG" type="Biogeographic note">
|
||||
<paragraph id="90FD0E401EE14A4BD45F9AE67E224C27">
|
||||
<heading id="AED7E11F606DE0A585EA23B45A0CDCCD" reason="title">Biogeographic note.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="65A050827AD7260AD786AA353A4AA7A8">
|
||||
As far as currently known, the Nearctic is the region with the greatest species diversity of
|
||||
<taxonomicName id="4A92841C079D151CEF080BDFC71D29C0" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="F1C4FA194A796B70C171AA30C58C0E50" italics="true">Chiquiticus</emphasis>
|
||||
</taxonomicName>
|
||||
. However, the
|
||||
<taxonomicName id="D0DFBFCA23D3EA0A02B3AD2F0206269E" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Chiquiticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="1FA63D2448FBE6001BB9BA202DC8AE61" italics="true">Chiquiticus</emphasis>
|
||||
</taxonomicName>
|
||||
fossils found in mid-Miocene Dominican amber (
|
||||
<bibRefCitation id="3C3E5E2AA4DE14C7217ED004DFCE4AFE" DOI="10.2992/007.081.0401" author="Chatzimanolis and Engel" firstAuthor="Chatzimanolis" issue="4" journalOrPublisher="Annals of Carnegie Museum" pagination="281-294" refId="B5" refString="Chatzimanolis S, Engel MS (2013) The Fauna of Staphylininae in Dominican Amber (Coleoptera: Staphylinidae). Annals of Carnegie Museum 81 (4): 281–294. https://doi.org/10.2992/007.081.0401" title="The Fauna of Staphylininae in Dominican Amber (Coleoptera: Staphylinidae)." volume="81" year="2013">Chatzimanolis and Engel 2013</bibRefCitation>
|
||||
) could suggest that Nearctic speciation took place after the genus dispersed there from the southern hemisphere where its closely related genera
|
||||
<taxonomicName id="F506F2D05AAF1B691823B21D4AA760BF" class="Insecta" family="Staphylinidae" genus="Amazonothops" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="4BD8369E462077246B3DFCC3D2E8B70C" italics="true">Amazonothops</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4DAF4BECE13DABEF90DD2BADDD41DC7E" authorityName="Cameron" authorityYear="1926" class="Insecta" family="Staphylinidae" genus="Ctenandropus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="00946204132B3F3C66F0A16AE3A1A5C8" italics="true">Ctenandropus</emphasis>
|
||||
</taxonomicName>
|
||||
are also found (Reyes-Hernández et al. in prep.). Such dispersal may have occurred via the GAARlandia land bridge (
|
||||
<bibRefCitation id="5F580B7921FEDE6B007E3F9C8F5833C9" DOI="10.2747/0020-6814.48.9.791" author="Iturralde-Vinent" firstAuthor="Iturralde-Vinent" issue="9" journalOrPublisher="International Geology Review" pagination="791-827" refId="B9" refString="Iturralde-Vinent M (2006) Meso-Cenozoic Caribbean paleogeography: implications for the historical biogeography of the region. International Geology Review 48 (9): 791–827. https://doi.org/10.2747/0020-6814.48.9.791" title="Meso-Cenozoic Caribbean paleogeography: implications for the historical biogeography of the region." volume="48" year="2006">Iturralde-Vinent 2006</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="8CCE7092B086EB73A3D233B0D8B17BBD" DOI="10.1016/j.ympev.2017.06.010" author="Masonick" etAl="et al." firstAuthor="Masonick" journalOrPublisher="Molecular Phylogenetics and Evolution" pagination="225-233" refId="B13" refString="Masonick P, Michael A, Frankenberg S, Rabitsch W, Weirauch C (2017) Molecular phylogenetics and biogeography of the ambush bugs (Hemiptera: Reduviidae: Phymatinae). Molecular Phylogenetics and Evolution 114: 225–233. https://doi.org/10.1016/j.ympev.2017.06.010" title="Molecular phylogenetics and biogeography of the ambush bugs (Hemiptera: Reduviidae: Phymatinae)." volume="114" year="2017">Masonick et al. 2017</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="FC789915C8A523C5AB7DE20CC700491B" DOI="10.1344/GeologicaActa2023.21.5" author="Iturralde-Vinent and MacPhee" firstAuthor="Iturralde-Vinent" refId="B10" refString="Iturralde-Vinent MA, MacPhee RDE (2023) New evidence for late Eocene-early Oligocene uplift of Aves Ridge and paleogeography of GAARlandia. Geologica Acta 21.5: 1–10. https://doi.org/10.1344/GeologicaActa2023.21.5" year="2023">Iturralde-Vinent and MacPhee 2023</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
515
data/36/99/DF/3699DF7013BF5ECB952C7177CE9A72D0.xml
Normal file
515
data/36/99/DF/3699DF7013BF5ECB952C7177CE9A72D0.xml
Normal file
|
|
@ -0,0 +1,515 @@
|
|||
<document id="592167A9586B5F3F5B27F1A43FDD959B" ID-DOI="10.3897/zookeys.1218.135558" ID-ZooBank="8C9034A6-1E8D-4FF6-8721-004826BB7797" ID-publisher-id="135558" URI-arpha="6EB9F9E6-D286-5BF4-AD5D-7B66E7EB7FD9" URI-zoobank="http://zoobank.org/8C9034A6-1E8D-4FF6-8721-004826BB7797" XM.bibliography_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1731693946473" checkinUser="pensoft" docAuthor="Reyes-Hernández, José L. & Solodovnikov, Alexey" docDate="2024" docId="3699DF7013BF5ECB952C7177CE9A72D0" docLanguage="en" docName="ZooKeys 1218: 81-98" docOrigin="ZooKeys 1218" docSource="https://zookeys.pensoft.net/article/135558/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Nitidocolpus Reyes-Hernández & Solodovnikov, 2024, gen. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="98" masterDocId="6EB9F9E6D2865BF4AD5D7B66E7EB7FD9" masterDocTitle="Two new rove beetle genera in Staphylininae that reduce “ Heterothops ” and “ Quedius ” taxonomic wastebaskets (Coleoptera, Staphylinidae)" masterLastPageNumber="98" masterPageNumber="81" pageNumber="81" updateTime="1731698241112" updateUser="admin">
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||||
<mods:mods id="07C83D9FC2796649BFA1FCA866330F57" xmlns:mods="http://www.loc.gov/mods/v3">
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||||
<mods:titleInfo id="B1F0F1FF87612E42FA962699585E0055">
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<mods:title id="41384A0A3BD446476A8E4DA7A3FE75BC">Two new rove beetle genera in Staphylininae that reduce “ Heterothops ” and “ Quedius ” taxonomic wastebaskets (Coleoptera, Staphylinidae)</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name id="CCDBE12ABBFB016F7452CF5819A35A6D" type="personal">
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||||
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||||
<mods:namePart id="27D7925F98025B3433F147616FA11C84">Reyes-Hernández, José L.</mods:namePart>
|
||||
<mods:nameIdentifier id="A782359FAA4496A8973EF6A14D1053B8" type="ORCID">0000-0002-4726-4439</mods:nameIdentifier>
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<mods:affiliation id="26B3043A959C065C66CA699AAF73A58A">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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</mods:name>
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<mods:name id="E29A776A4F5BB44DC7677387989BD150" type="personal">
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</mods:role>
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<mods:namePart id="93E5BFD59903CB51C6FE192304190FAA">Solodovnikov, Alexey</mods:namePart>
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<mods:nameIdentifier id="51DB9F240201DA7B5F7E89B159CE7AD7" type="ORCID">0000-0003-2031-849X</mods:nameIdentifier>
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<mods:affiliation id="F1E83947991C252986AB5B9AC3001B0D">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:title id="691355B9B25B1A430DABC27CFCF22907">ZooKeys</mods:title>
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<mods:part id="E43C1FD3586D849F22579FF80CE5CDD0">
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<mods:date id="EA6C479B792D6BFF4EF4E1709D876A87">2024</mods:date>
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<mods:number id="14005CD7130FCE91F866BA331199A6D0">2024-11-15</mods:number>
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</mods:detail>
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<mods:detail id="4A4FCFC1696860C4F2434D89D727662A" type="volume">
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<mods:number id="E6B4D0F1BED6BD990D900473D2CD0DC4">1218</mods:number>
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<mods:classification id="28D47D960EFE95E9F80554C8DD0B157D">journal article</mods:classification>
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<mods:identifier id="D04F6149A2B4435B8EF87C7696F58FDB" type="DOI">10.3897/zookeys.1218.135558</mods:identifier>
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<mods:identifier id="E1C380BE614FEE05E66FA6FBBCF0F545" type="ZooBank">8C9034A6-1E8D-4FF6-8721-004826BB7797</mods:identifier>
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</mods:mods>
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<treatment id="3699DF7013BF5ECB952C7177CE9A72D0" ID-arpha="3699DF70-13BF-5ECB-952C-7177CE9A72D0" ID-zoobank="https://zoobank.org/9BC238C3-B742-4E1E-8B59-48E8C83288DF" LSID="urn:lsid:plazi:treatment:3699DF7013BF5ECB952C7177CE9A72D0" LSID-ZBK="urn:lsid:zoobank.org:act:9BC238C3-B742-4E1E-8B59-48E8C83288DF" httpUri="http://treatment.plazi.org/id/3699DF7013BF5ECB952C7177CE9A72D0">
|
||||
<subSubSection id="D052E54EF3025CA5104A3C52D3EAFEE0" type="nomenclature">
|
||||
<paragraph id="9E8380C4211ED5159ACD7DFB08C64C39">
|
||||
<heading id="1F5EF7A8B9AA734689444C21458A176F" reason="label">Genus</heading>
|
||||
<taxonomicName id="D9255429E222663203FF494CE8870F11" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="ED4914F3FEDC836384E44B394730B4B2" italics="true">Nitidocolpus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="4D1AFA5E06F609E313D72FE8FB1D4A9B" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="07D01FEBC3C978179D633A8EBEEBD11D" type="description">
|
||||
<paragraph id="0DD9FF12396B7B0CAE2A48F55D55BB64">
|
||||
<figureCitation id="A8167CC06431F769553C64DBFC0092A8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Some Nitidocolpus species and characters A N. illatus (Sharp, 1884) B N. laeticulus (Sharp, 1884) C N. germaini (Bernhauer, 1917) D N. aurofasciatus (Bernhauer, 1917) E N. illatus, pterothoraces F N. columbinus (Bernhauer, 1917), metacoxa and metatrochanter, dissected right leg in dorsal view (side faced to the abdomen). Abbreviations: EXPR, external part of prepectus; INPR, internal part of prepectus; MIV, mesobasisternal intercoxal process with V-shaped projection medially; MNPS, mesanapleural sutures; MTMS, metacoxa with dorsomedial spine; MTTAS, metatrochanter with dorsoapical spine. Photo B by J. Jenkins Shaw. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure5" httpUri="https://binary.pensoft.net/fig/1175478">Figs 5</figureCitation>
|
||||
,
|
||||
<figureCitation id="0B44E44C9DD2ECF0C9C54567AC747A32" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Morphological details of Nitidocolpus and other relevant genera of Staphylininae A – C Nitidocolpus columbinus D – F Cheilocolpus viridulus G – I Cyrtoquedius sp. J – L Quedionuchus nr. impunctus M – O Quedius nr. advena. A, D, G, J, M head B, E, H, K, N pronotum C, F, I, L, O aedeagus in lateral view, the black arrow pointing down indicates the basal projection of the paramere. Abbreviations: ACP, anterolateral clypeal puncture; AFP, anterior frontal; AMRSP, anterior marginal row of setiferous punctures; APP, additional paired punctures adjacent to AMRSP; BP, basal puncture; FCP, frontoclypeal puncture; IASP, internalmost setiferous puncture of the anterior margin; IOP, interocular punctures; IOR, infraorbital ridge; LLSP, large lateral setiferous puncture; LPSP, large posterior setiferous puncture; NR, nuchal ridge; PFP, posterior frontal puncture; POP, parocular punctures; PPDS, paired punctures on dorsal series; SAP, supra-antennal puncture; SAR, supra-antennal ridge; SLSP, sublateral setiferous puncture; TP, temporal puncture. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure6" httpUri="https://binary.pensoft.net/fig/1175479">6 A – C</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EDSAG" type="type_taxon">
|
||||
<paragraph id="8A5A376B8DE48E8B203293F5019E6ED8">
|
||||
<heading id="6EEDE74377B9B3C46B0FFC9DC532FF91" reason="title">
|
||||
<typeStatus id="44E3E4D0DC648C59C29CFF0BF53D81EA">Type</typeStatus>
|
||||
species.
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="4A08E0D54BB44AB42481CC7744A06C2A">
|
||||
<taxonomicName id="61960DFA71E68767F97193695465BCEB" authority="Bernhauer, 1917" authorityName="Bernhauer" authorityYear="1917" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="columbinus">
|
||||
<emphasis id="A228E39B164435E57026F804AB87CDD6" italics="true">Quedius columbinus</emphasis>
|
||||
Bernhauer, 1917
|
||||
</taxonomicName>
|
||||
, here designated.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EUSAG" type="taxon_list">
|
||||
<paragraph id="78447F6D03E0C83AD7BCC5A8EE46511D">
|
||||
<heading id="5D82CBB95499F04E5BF5B407673940D9" reason="title">Included species.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="BA118F55B1464896E0479E7507DB99E4">
|
||||
<taxonomicName id="9CCA75FC28AE0D90963807DA9DCC3F70" authority="(Bernhauer, 1917)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Bernhauer" baseAuthorityYear="1917" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="aurofasciatus" status="comb. nov.">
|
||||
<emphasis id="BB83F97B7B2EF2603578EE82464DF678" italics="true">Nitidocolpus aurofasciatus</emphasis>
|
||||
(Bernhauer, 1917)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="889801091C875B4B652EC8E361CCE8CA" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
ex.
|
||||
<taxonomicName id="2A3D0D73C9AA909D1558C062AFE852E5" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="804E636B4C8262BB17D354BCE7993FED" italics="true">Quedius</emphasis>
|
||||
</taxonomicName>
|
||||
[
|
||||
<specimenCount id="B1573A5C23B85711E166CAB7430EA0DF" count="1" type="generic" typeStatus="syntype">1 syntype</specimenCount>
|
||||
from
|
||||
<named-content id="DD9EA8A659B1E8F8DD323B80D40AEB5C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/field-museum-natural-history" xlink_title="Field Museum of Natural History, Chicago, Illinois, USA">
|
||||
<collectionCode id="2781117D46E3B7610A83FA9956FA8ED8" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" type="Museum">FMNH</collectionCode>
|
||||
</named-content>
|
||||
examined];
|
||||
<taxonomicName id="4F1B9CF3F2DEF769BD0FD469432F32A5" authority="(Sharp, 1884)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1884" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="championi" status="comb. nov.">
|
||||
<emphasis id="8663ABC194A8DF7F80BA15B7B3F057B5" italics="true">Nitidocolpus championi</emphasis>
|
||||
(Sharp, 1884)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="1D83FC4ABFACBD6255973A88B1BC8CCA" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
ex.
|
||||
<taxonomicName id="FE64A534D0F1C469B93E3DAD942386A8" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="9573DBF8CBDBCF4D0190C055752BABD5" italics="true">Quedius</emphasis>
|
||||
</taxonomicName>
|
||||
[
|
||||
<specimenCount id="16D90C432125D3D8D6DFCE666D4E010D" count="1" type="generic" typeStatus="syntype">1 syntype</specimenCount>
|
||||
from
|
||||
<named-content id="298367C0EF2549D37A8AE44ADAC2FE84" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/field-museum-natural-history" xlink_title="Field Museum of Natural History, Chicago, Illinois, USA">
|
||||
<collectionCode id="F90AE8D557C81D1D729B680D087393C8" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" type="Museum">FMNH</collectionCode>
|
||||
</named-content>
|
||||
examined];
|
||||
<taxonomicName id="53A8318C45DDF1662C0BC64E6154DE28" authority="(Bernhauer, 1917)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Bernhauer" baseAuthorityYear="1917" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="columbinus" status="comb. nov.">
|
||||
<emphasis id="FA76C56CB9F9500AC0FE31F06CF7F40E" italics="true">Nitidocolpus columbinus</emphasis>
|
||||
(Bernhauer, 1917)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="9840ECB5FBE01DF37B3DC39E5DA50520" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
ex.
|
||||
<taxonomicName id="F7A1E990859F7BE62A32A44AD607AA51" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="65F966380F75020BAC758982F21587AC" italics="true">Quedius</emphasis>
|
||||
</taxonomicName>
|
||||
[
|
||||
<specimenCount id="2A38914883AFBEB2874BA8A90E7C9346" count="2" type="generic" typeStatus="syntype">2 syntypes</specimenCount>
|
||||
from
|
||||
<named-content id="178B5AF9240B17ABCB09A64080A7C35D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/field-museum-natural-history" xlink_title="Field Museum of Natural History, Chicago, Illinois, USA">
|
||||
<collectionCode id="6504880A0784E4F488C0B2C8C8036345" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" type="Museum">FMNH</collectionCode>
|
||||
</named-content>
|
||||
examined];
|
||||
<taxonomicName id="FB50C1961640551FBD01B5144254FB8D" authority="(Bernhauer, 1917)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Bernhauer" baseAuthorityYear="1917" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="germaini" status="comb. nov.">
|
||||
<emphasis id="A5206C04C6E665F76D227525CC46CCB3" italics="true">Nitidocolpus germaini</emphasis>
|
||||
(Bernhauer, 1917)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="71A3485825FB1D229A54F0F150AA66B1" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
ex.
|
||||
<taxonomicName id="3C3C441F1487661EB81791F8EF8D2365" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="F14C2FCAEA45D720BDE77F6E0C22A7E9" italics="true">Quedius</emphasis>
|
||||
</taxonomicName>
|
||||
[
|
||||
<specimenCount id="80AD57FD3EFAFFAA44C4ED92DBC2442A" count="1" type="generic" typeStatus="syntype">1 syntype</specimenCount>
|
||||
from
|
||||
<named-content id="89620B5DD8546A7A1B4F4F4192B90E55" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/field-museum-natural-history" xlink_title="Field Museum of Natural History, Chicago, Illinois, USA">
|
||||
<collectionCode id="CECFF9D28FA0910E86F0ABFF15D5DCA4" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" type="Museum">FMNH</collectionCode>
|
||||
</named-content>
|
||||
examined];
|
||||
<taxonomicName id="6E7E989B580D37FBD5D2A82CB584F497" authority="(Sharp, 1884)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1884" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="illatus" status="comb. nov.">
|
||||
<emphasis id="54C8438D01B087FEB6790DDCF9DD906E" italics="true">Nitidocolpus illatus</emphasis>
|
||||
(Sharp, 1884)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="2CA14E7232C00A7F908F1D8C5A1E1BE6" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
ex.
|
||||
<taxonomicName id="674B78EA095409372D3247149ED86776" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="47EE25405302540ADC1081D207E83594" italics="true">Quedius</emphasis>
|
||||
</taxonomicName>
|
||||
[
|
||||
<specimenCount id="1690860E92C3FD4CAD0D90FE71F75E40" count="1" type="generic" typeStatus="syntype">1 syntype</specimenCount>
|
||||
from
|
||||
<named-content id="C8D2159C5ACCF82BD5E49C40AB55616F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London, United Kingdom">
|
||||
<collectionCode id="FA73F48E2859DDD32683C97524203238" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" type="Herbarium">NHM</collectionCode>
|
||||
</named-content>
|
||||
examined];
|
||||
<taxonomicName id="FA1D40CF50DC27F27B3D72EF3F431697" authority="(Sharp, 1884)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1884" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="laeticulus" status="comb. nov.">
|
||||
<emphasis id="ABA6FC1CBE379F9B4DAEA63473E633E5" italics="true">Nitidocolpus laeticulus</emphasis>
|
||||
(Sharp, 1884)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="4BAF4B4364144819EDA1E7FCC9B6AB85" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
ex.
|
||||
<taxonomicName id="259C4239A53AADCC5950BE38900C35CA" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="04772AF80D9280510D6FB594DFAB5A74" italics="true">Quedius</emphasis>
|
||||
</taxonomicName>
|
||||
[
|
||||
<specimenCount id="2002368E1550B97C94064ABB01CD8B3A" count="1" type="generic" typeStatus="syntype">1 syntype</specimenCount>
|
||||
from
|
||||
<named-content id="0846CD16F5931F2BDC5F4661A35238AE" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London, United Kingdom">
|
||||
<collectionCode id="DE78B435A65CCB7B151738DA8C137E4D" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" type="Herbarium">NHM</collectionCode>
|
||||
</named-content>
|
||||
examined];
|
||||
<taxonomicName id="2E02A7AE6016BF165876F471831A548D" authority="(Fauvel, 1891)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Fauvel" baseAuthorityYear="1891" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="triangulum" status="comb. nov.">
|
||||
<emphasis id="9998F6A591086AB009B9F88328263BFF" italics="true">Nitidocolpus triangulum</emphasis>
|
||||
(
|
||||
<bibRefCitation id="8343FD47D750BB67A2B9D3EEADFBE175" author="Fauvel A" journalOrPublisher="Revue d’Entomologie" pagination="87-127" refId="B7" refString="Fauvel A (1891) Voyage de M. E. Simon au venezuela (Decembre 1887 - Avril 1888). 11 e Memoire. Revue d'Entomologie 10: 87 - 127." title="Voyage de M. E. Simon au vénézuela (Décembre 1887 - Avril 1888). 11 e Mémoire." volume="10" year="1891">Fauvel, 1891</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="EA1AE2E5E76A5A3713E2A04725FCB6D7" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
ex.
|
||||
<taxonomicName id="C1815A963E1C2F3257586C133B07A9D0" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="2F8244FDAB81931249FA30BC7EFA4B9C" italics="true">Quedius</emphasis>
|
||||
</taxonomicName>
|
||||
[
|
||||
<specimenCount id="9515E08D14F6C09AA441ADA08BB783CF" count="1" type="generic" typeStatus="syntype">1 syntype</specimenCount>
|
||||
from
|
||||
<named-content id="6E89391C1A86F132F5449638B4595E3F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institut-royal-des-sciences-naturelles-de-belgique" xlink_title="Institut Royal des Sciences Naturelles de Belgique, Brussels, Belgium">
|
||||
<collectionCode id="F30FC1D7168AF2CB494A1B7E67C4A5D8" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" type="">IRSNB</collectionCode>
|
||||
</named-content>
|
||||
examined].
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E2XAG" type="diagnosis">
|
||||
<paragraph id="181C40653B9C2C323D18EB3777E19A4B">
|
||||
<heading id="16A00FFE603C343A32F6A0289801FAB0" reason="title">Diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="5270BDC0AF27BC2E3247E7989B23F8EE">
|
||||
Supra-antennal punctures and paraocular punctures absent (Fig.
|
||||
<figureCitation id="277E0FBAD225D331E4ED850F3D4ED286" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Morphological details of Nitidocolpus and other relevant genera of Staphylininae A – C Nitidocolpus columbinus D – F Cheilocolpus viridulus G – I Cyrtoquedius sp. J – L Quedionuchus nr. impunctus M – O Quedius nr. advena. A, D, G, J, M head B, E, H, K, N pronotum C, F, I, L, O aedeagus in lateral view, the black arrow pointing down indicates the basal projection of the paramere. Abbreviations: ACP, anterolateral clypeal puncture; AFP, anterior frontal; AMRSP, anterior marginal row of setiferous punctures; APP, additional paired punctures adjacent to AMRSP; BP, basal puncture; FCP, frontoclypeal puncture; IASP, internalmost setiferous puncture of the anterior margin; IOP, interocular punctures; IOR, infraorbital ridge; LLSP, large lateral setiferous puncture; LPSP, large posterior setiferous puncture; NR, nuchal ridge; PFP, posterior frontal puncture; POP, parocular punctures; PPDS, paired punctures on dorsal series; SAP, supra-antennal puncture; SAR, supra-antennal ridge; SLSP, sublateral setiferous puncture; TP, temporal puncture. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure6" httpUri="https://binary.pensoft.net/fig/1175479">6 A</figureCitation>
|
||||
); pronotum without paired dorsal series of setiferous macropunctures (Fig.
|
||||
<figureCitation id="05C4E4AD73FFFADAA64FBD967C84F7B6" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Morphological details of Nitidocolpus and other relevant genera of Staphylininae A – C Nitidocolpus columbinus D – F Cheilocolpus viridulus G – I Cyrtoquedius sp. J – L Quedionuchus nr. impunctus M – O Quedius nr. advena. A, D, G, J, M head B, E, H, K, N pronotum C, F, I, L, O aedeagus in lateral view, the black arrow pointing down indicates the basal projection of the paramere. Abbreviations: ACP, anterolateral clypeal puncture; AFP, anterior frontal; AMRSP, anterior marginal row of setiferous punctures; APP, additional paired punctures adjacent to AMRSP; BP, basal puncture; FCP, frontoclypeal puncture; IASP, internalmost setiferous puncture of the anterior margin; IOP, interocular punctures; IOR, infraorbital ridge; LLSP, large lateral setiferous puncture; LPSP, large posterior setiferous puncture; NR, nuchal ridge; PFP, posterior frontal puncture; POP, parocular punctures; PPDS, paired punctures on dorsal series; SAP, supra-antennal puncture; SAR, supra-antennal ridge; SLSP, sublateral setiferous puncture; TP, temporal puncture. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure6" httpUri="https://binary.pensoft.net/fig/1175479">6 B</figureCitation>
|
||||
); mesanapleural sutures transverse or nearly transverse but reaching the external part of prepectus (Fig.
|
||||
<figureCitation id="03AB624FB58BD3432A865BCB61AA490B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Some Nitidocolpus species and characters A N. illatus (Sharp, 1884) B N. laeticulus (Sharp, 1884) C N. germaini (Bernhauer, 1917) D N. aurofasciatus (Bernhauer, 1917) E N. illatus, pterothoraces F N. columbinus (Bernhauer, 1917), metacoxa and metatrochanter, dissected right leg in dorsal view (side faced to the abdomen). Abbreviations: EXPR, external part of prepectus; INPR, internal part of prepectus; MIV, mesobasisternal intercoxal process with V-shaped projection medially; MNPS, mesanapleural sutures; MTMS, metacoxa with dorsomedial spine; MTTAS, metatrochanter with dorsoapical spine. Photo B by J. Jenkins Shaw. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure5" httpUri="https://binary.pensoft.net/fig/1175478">5 E</figureCitation>
|
||||
); mesobasisternal intercoxal process with V-shaped projection medially (Fig.
|
||||
<figureCitation id="86CA88BF25616C52AF7E292FD3DC356E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Some Nitidocolpus species and characters A N. illatus (Sharp, 1884) B N. laeticulus (Sharp, 1884) C N. germaini (Bernhauer, 1917) D N. aurofasciatus (Bernhauer, 1917) E N. illatus, pterothoraces F N. columbinus (Bernhauer, 1917), metacoxa and metatrochanter, dissected right leg in dorsal view (side faced to the abdomen). Abbreviations: EXPR, external part of prepectus; INPR, internal part of prepectus; MIV, mesobasisternal intercoxal process with V-shaped projection medially; MNPS, mesanapleural sutures; MTMS, metacoxa with dorsomedial spine; MTTAS, metatrochanter with dorsoapical spine. Photo B by J. Jenkins Shaw. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure5" httpUri="https://binary.pensoft.net/fig/1175478">5 E</figureCitation>
|
||||
); metatrochanters with apical spine (Fig.
|
||||
<figureCitation id="135D4DB81A329791BD47D5981423D4E5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Some Nitidocolpus species and characters A N. illatus (Sharp, 1884) B N. laeticulus (Sharp, 1884) C N. germaini (Bernhauer, 1917) D N. aurofasciatus (Bernhauer, 1917) E N. illatus, pterothoraces F N. columbinus (Bernhauer, 1917), metacoxa and metatrochanter, dissected right leg in dorsal view (side faced to the abdomen). Abbreviations: EXPR, external part of prepectus; INPR, internal part of prepectus; MIV, mesobasisternal intercoxal process with V-shaped projection medially; MNPS, mesanapleural sutures; MTMS, metacoxa with dorsomedial spine; MTTAS, metatrochanter with dorsoapical spine. Photo B by J. Jenkins Shaw. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure5" httpUri="https://binary.pensoft.net/fig/1175478">5 F</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="1DA9578AB3A79AF20937A0C3C61634AF" ID-DOI="10.3897/zookeys.1218.135558.figure5" ID-arpha="BA24B181-2D31-5A39-BAE9-7EC7E37A42EF" httpUri="https://binary.pensoft.net/fig/1175478" startId="F5">
|
||||
<paragraph id="F68748BAC8DFE51A68B4146241BE9A96">
|
||||
<label id="B57CD1A9F2C14765CA9DA13C47E20895">Figure 5.</label>
|
||||
</paragraph>
|
||||
<paragraph id="E9583DCFFE10CC837729F976FD05D5A2">
|
||||
Some
|
||||
<taxonomicName id="53497210DAC1326658A25FF975CAFB36" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="undefined">
|
||||
<emphasis id="9DAC792E2BCB7DF8601A653430438DC8" italics="true">Nitidocolpus</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
and characters
|
||||
<emphasis id="9CBA4E1C747135D0514882F5B427BAC7" bold="true">A</emphasis>
|
||||
<taxonomicName id="C2B5554F316FCD72263C7ADBFD35E643" authority="(Sharp, 1884)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1884" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="illatus">
|
||||
<emphasis id="B90B72EE8858811423104370673D8277" italics="true">N. illatus</emphasis>
|
||||
(Sharp, 1884)
|
||||
</taxonomicName>
|
||||
<emphasis id="8D8A87C8CA68A0C4EAC346D47B19E1DE" bold="true">B</emphasis>
|
||||
<taxonomicName id="05CD16CE00F6F43A7409DE3D2711EFB8" authority="(Sharp, 1884)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1884" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="laeticulus">
|
||||
<emphasis id="388D7352890CCE0A96C7BAA7AD4F1AEB" italics="true">N. laeticulus</emphasis>
|
||||
(Sharp, 1884)
|
||||
</taxonomicName>
|
||||
<emphasis id="C4411B626BDFA0493831C178B94E6C4E" bold="true">C</emphasis>
|
||||
<taxonomicName id="C1C5876AAD2535240DA3426DAFB235C3" authority="(Bernhauer, 1917)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Bernhauer" baseAuthorityYear="1917" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="germaini">
|
||||
<emphasis id="E99FDF2876270A6DF8B3E7A18A1B236B" italics="true">N. germaini</emphasis>
|
||||
(Bernhauer, 1917)
|
||||
</taxonomicName>
|
||||
<emphasis id="D0020B0EF3DC55C62B903610736FDD92" bold="true">D</emphasis>
|
||||
<taxonomicName id="4B84DEBEAD26C0FF88641B54A55CE67C" authority="(Bernhauer, 1917)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Bernhauer" baseAuthorityYear="1917" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="aurofasciatus">
|
||||
<emphasis id="CF4F8035CB4D22E5B7DB34345F305346" italics="true">N. aurofasciatus</emphasis>
|
||||
(Bernhauer, 1917)
|
||||
</taxonomicName>
|
||||
<emphasis id="F6931EDF0C0EE000CC268455F47C3542" bold="true">E</emphasis>
|
||||
<taxonomicName id="01DC1F87492F902CAEB3F195C2EB7607" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1884" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="illatus">
|
||||
<emphasis id="128C9A195F2CDF2FAAAEC11709EB61E2" italics="true">N. illatus</emphasis>
|
||||
</taxonomicName>
|
||||
, pterothoraces
|
||||
<emphasis id="8D2956D7FAEAD49D16FD913ABA44952E" bold="true">F</emphasis>
|
||||
<taxonomicName id="2B437BEE8E0645F0DCB5B8C27A44F992" authority="(Bernhauer, 1917)" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Bernhauer" baseAuthorityYear="1917" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="columbinus">
|
||||
<emphasis id="1799D272A48ACC70B8D091353868B849" italics="true">N. columbinus</emphasis>
|
||||
(Bernhauer, 1917)
|
||||
</taxonomicName>
|
||||
, metacoxa and metatrochanter, dissected right leg in dorsal view (side faced to the abdomen). Abbreviations: EXPR, external part of prepectus; INPR, internal part of prepectus; MIV, mesobasisternal intercoxal process with V-shaped projection medially;
|
||||
<abbrev id="ABBRID0ES2AG" xlink_title="mesanapleural sutures">MNPS</abbrev>
|
||||
, mesanapleural sutures; MTMS, metacoxa with dorsomedial spine; MTTAS, metatrochanter with dorsoapical spine. Photo B by J. Jenkins Shaw. Scale bars: 1 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E22AG" type="description">
|
||||
<paragraph id="1C3AAC14DB1E1772798FE08703194050">
|
||||
<heading id="48C8FDC42D53ED562942330CC53B3128" reason="title">Description.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="E2546B41D467FA69E694E93CD59E97D8">
|
||||
Small to medium-sized
|
||||
<taxonomicName id="BAE32A18CCF6BBCF0476C623A9CB4C2D" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subFamily" subFamily="Staphylininae">Staphylininae</taxonomicName>
|
||||
(TL =
|
||||
<quantity id="F8CC2A19632023D703443DA100C1FFF7" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="10.0" metricValueMin="4.5" unit="mm" value="7.25" valueMax="10.0" valueMin="4.5">4.5–10 mm</quantity>
|
||||
) (Fig.
|
||||
<figureCitation id="43C88BF7D231FB04022F0309FCD21430" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Some Nitidocolpus species and characters A N. illatus (Sharp, 1884) B N. laeticulus (Sharp, 1884) C N. germaini (Bernhauer, 1917) D N. aurofasciatus (Bernhauer, 1917) E N. illatus, pterothoraces F N. columbinus (Bernhauer, 1917), metacoxa and metatrochanter, dissected right leg in dorsal view (side faced to the abdomen). Abbreviations: EXPR, external part of prepectus; INPR, internal part of prepectus; MIV, mesobasisternal intercoxal process with V-shaped projection medially; MNPS, mesanapleural sutures; MTMS, metacoxa with dorsomedial spine; MTTAS, metatrochanter with dorsoapical spine. Photo B by J. Jenkins Shaw. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure5" httpUri="https://binary.pensoft.net/fig/1175478">5 A – D</figureCitation>
|
||||
). Head: Frontoclypeal (epistomal) suture absent or distinct only laterally and only in teneral (or chemically cleared) specimens; head slightly transverse (
|
||||
<abbrev id="ABBRID0EL3AG" xlink_title="head width at the widest point, including eyes">
|
||||
<collectionCode id="A116B4800F19AA5AFD612F29B0F785FB">HW</collectionCode>
|
||||
</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0EP3AG" xlink_title="head length">
|
||||
<collectionCode id="0F3DAD02CB67BCF9DE3AEBCFBC5E4A1F" country="USA" lsid="urn:lsid:biocol.org:col:13245" name="Houghton Lake Wildlife Research Station" type="Herbarium">HL</collectionCode>
|
||||
</abbrev>
|
||||
≥ 1.1) without distinct posterior angles; disc smooth; frons with microsculpture as transverse waves, without distinct concavity between antennal insertions; neck moderately wide (
|
||||
<abbrev id="ABBRID0ET3AG" xlink_title="neck width at the widest point">
|
||||
<collectionCode id="F65ED6A6D1542C159665166C1CBA91CE">NW</collectionCode>
|
||||
</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0EX3AG" xlink_title="head width at the widest point, including eyes">
|
||||
<collectionCode id="7F6398D9D5597322D37E48F91E444493">HW</collectionCode>
|
||||
</abbrev>
|
||||
> 0.75 but <0.90); dorsal macrosetae (Fig.
|
||||
<figureCitation id="D7E614E67846F34BD1D3E668E0B4CE35" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Morphological details of Nitidocolpus and other relevant genera of Staphylininae A – C Nitidocolpus columbinus D – F Cheilocolpus viridulus G – I Cyrtoquedius sp. J – L Quedionuchus nr. impunctus M – O Quedius nr. advena. A, D, G, J, M head B, E, H, K, N pronotum C, F, I, L, O aedeagus in lateral view, the black arrow pointing down indicates the basal projection of the paramere. Abbreviations: ACP, anterolateral clypeal puncture; AFP, anterior frontal; AMRSP, anterior marginal row of setiferous punctures; APP, additional paired punctures adjacent to AMRSP; BP, basal puncture; FCP, frontoclypeal puncture; IASP, internalmost setiferous puncture of the anterior margin; IOP, interocular punctures; IOR, infraorbital ridge; LLSP, large lateral setiferous puncture; LPSP, large posterior setiferous puncture; NR, nuchal ridge; PFP, posterior frontal puncture; POP, parocular punctures; PPDS, paired punctures on dorsal series; SAP, supra-antennal puncture; SAR, supra-antennal ridge; SLSP, sublateral setiferous puncture; TP, temporal puncture. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure6" httpUri="https://binary.pensoft.net/fig/1175479">6 A</figureCitation>
|
||||
): anterolateral clypeal punctures present; frontoclypeal punctures present; supra-antennal punctures absent; parocular punctures absent; basal punctures present, single; ventral macrosetae: postocular punctures present; infraorbital punctures absent; postmandibular punctures present; submentum with two pairs of macrosetae; mentum with seta alpha only, seta beta absent; gular sutures separated from each other, gula with distinct transverse basal impression; ventral basal ridge present; postgenal ridge present; postmandibular ridge short, extended parallel to margin of eye, without fusion with any other ridge and distinctly separated from eye margin; infraorbital ridge merged with postgenal ridge; nuchal ridge missing dorsally, present laterally, merged with infraorbital ridge; eyes of medium size (
|
||||
<abbrev id="ABBRID0E63AG" xlink_title="eye length in lateral view">EYL</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0ED4AG" xlink_title="head length">
|
||||
<collectionCode id="5C4D641BD207315543D2FD2848A990DC" country="USA" lsid="urn:lsid:biocol.org:col:13245" name="Houghton Lake Wildlife Research Station" type="Herbarium">HL</collectionCode>
|
||||
</abbrev>
|
||||
ratio more than 1 / 2 but less than 3 / 4); gena short (
|
||||
<abbrev id="ABBRID0EH4AG" xlink_title="gena length">GL</abbrev>
|
||||
/
|
||||
<collectionCode id="B39285C8C65738F9DF3D47DE67381274" country="USA" name="Lake Forest College" type="Herbarium">ETL</collectionCode>
|
||||
<0.5); tomentose pubescence begins from fourth antennomere, its density on fourth antennomere as on following antennomeres; setation of antennomere 3 almost evenly distributed; antennomere 11 with subapical rounded field, without subapical lateral pits; mandibles with dorso-lateral seta, without dorso-lateral groove, external edge with curved base and apex, straight in middle length; right mandible with proximal tooth and bifid distal tooth, apically deflexed ventrad at an angle of> 20 °; left mandible rather straight; labrum with transparent wide apical membrane, anterior margin emarginate at middle; maxillary palps: palpomere 4 (apical) subconical, with evenly narrowed apex, longer than palpomere 2 and weakly dilated compared to palpomere 3; labial palps: palpomere 3 (apical) more or less fusiform or subconical, distinctly longer than, and about as wide as, palpomere 2.
|
||||
</paragraph>
|
||||
<caption id="C5828026F215B5D6C8C4CDC7A5B9880C" ID-DOI="10.3897/zookeys.1218.135558.figure6" ID-arpha="C66562ED-C5CB-5792-BB98-24153A71FF18" httpUri="https://binary.pensoft.net/fig/1175479" startId="F6">
|
||||
<paragraph id="BB38C84CDBE94D87EBDE66A6040EE438">
|
||||
<label id="5B730D12F763C43BA3FC838B02A02AD1">Figure 6.</label>
|
||||
</paragraph>
|
||||
<paragraph id="0DEF715B28C3E18DBE4059B9A7D77894">
|
||||
Morphological details of
|
||||
<taxonomicName id="6655517B6708332E67F86B055E6B171B" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="59C6B7C187060308272BE21AC2095E71" italics="true">Nitidocolpus</emphasis>
|
||||
</taxonomicName>
|
||||
and other relevant genera of
|
||||
<taxonomicName id="984A670349033C914FBE9879B0D4CA1E" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subFamily" subFamily="Staphylininae">Staphylininae</taxonomicName>
|
||||
<emphasis id="F5EA649840F207663CC580A3296D0C50" bold="true">A – C</emphasis>
|
||||
<taxonomicName id="84CC23C1FD7DB520BA7517ED3550E2C6" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Bernhauer" baseAuthorityYear="1917" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="columbinus">
|
||||
<emphasis id="99CCC469EE1E28E9B0B0B32706625E41" italics="true">Nitidocolpus columbinus</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="F9EE836C58A05D696BF307405FD7F797" bold="true">D – F</emphasis>
|
||||
<taxonomicName id="A5846F4DDEF234D1EAD54EB598FD708C" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Erichson" baseAuthorityYear="1840" class="Insecta" family="Staphylinidae" genus="Cheilocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="viridulus">
|
||||
<emphasis id="661EC6D1EBEA537571D788CCEEFD661F" italics="true">Cheilocolpus viridulus</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="BB1EF7DE8E56479C49C304AE0D6EDD41" bold="true">G – I</emphasis>
|
||||
<taxonomicName id="57B8072B2B222C3B87970E0FA552F0B0" class="Insecta" family="Staphylinidae" genus="Cyrtoquedius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="undefined">
|
||||
<emphasis id="B3533C78F7D5D8202C2BE8E32C7A1BEE" italics="true">Cyrtoquedius</emphasis>
|
||||
sp.
|
||||
</taxonomicName>
|
||||
<emphasis id="D62F0E3DE175D1CE5FE312DC20A4FCA8" bold="true">J – L</emphasis>
|
||||
<taxonomicName id="A3DFDE4535B4050D8FA711A37D4F3667" authorityName="Sharp" authorityYear="1884" class="Insecta" family="Staphylinidae" genus="Quedionuchus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="68EE2EFDE2149FD0740C249CB8E19FC4" italics="true">Quedionuchus</emphasis>
|
||||
</taxonomicName>
|
||||
nr.
|
||||
<emphasis id="6B97960604894D8D3A851FFAB099EF77" italics="true">impunctus</emphasis>
|
||||
<emphasis id="767E3D41AF85A83B46B9313502EEF2A4" bold="true">M – O</emphasis>
|
||||
<taxonomicName id="447AE49C9679F216C76A259000338F26" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="F24F436346FCC20431693C3EABB41B27" italics="true">Quedius</emphasis>
|
||||
</taxonomicName>
|
||||
nr.
|
||||
<emphasis id="FE4379700A63D3328E7F76FD17E48A2C" italics="true">advena</emphasis>
|
||||
.
|
||||
<emphasis id="2522CD56A3603305AFB6BFEFFB542125" bold="true">A, D, G, J, M</emphasis>
|
||||
head
|
||||
<emphasis id="1FEA6E02F56DBCBC39BA8E342D55C51F" bold="true">B, E, H, K, N</emphasis>
|
||||
pronotum
|
||||
<emphasis id="92628CB407BA27B2609E25DE95ED7CE4" bold="true">C, F, I, L, O</emphasis>
|
||||
aedeagus in lateral view, the black arrow pointing down indicates the basal projection of the paramere. Abbreviations:
|
||||
<abbrev id="ABBRID0EHABG" xlink_title="anterolateral clypeal punctures">ACP</abbrev>
|
||||
, anterolateral clypeal puncture; AFP, anterior frontal; AMRSP, anterior marginal row of setiferous punctures; APP, additional paired punctures adjacent to AMRSP; BP, basal puncture; FCP, frontoclypeal puncture; IASP, internalmost setiferous puncture of the anterior margin; IOP, interocular punctures; IOR, infraorbital ridge; LLSP, large lateral setiferous puncture; LPSP, large posterior setiferous puncture; NR, nuchal ridge; PFP, posterior frontal puncture; POP, parocular punctures; PPDS, paired punctures on dorsal series; SAP, supra-antennal puncture;
|
||||
<abbrev id="ABBRID0ELABG" xlink_title="supra-antennal ridge">SAR</abbrev>
|
||||
, supra-antennal ridge; SLSP, sublateral setiferous puncture; TP, temporal puncture. Scale bars: 1 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="A6DFB8566092BD33BF4326FAE21B4F6E">
|
||||
Thorax: Prothorax with slightly transverse pronotum (
|
||||
<abbrev id="ABBRID0EWABG" xlink_title="pronotum width at the widest point">
|
||||
<collectionCode id="7D0B7F98F22A501FB9933CEB73D68279" country="Thailand" name="Paleontological Collections">PW</collectionCode>
|
||||
</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0E1ABG" xlink_title="pronotum length along the median line">
|
||||
<collectionCode id="0D9A0ACFF63020970629BC9F91A06E55" country="Czech Republic" name="Západoceské muzeum v Plzni" type="Museum">PL</collectionCode>
|
||||
</abbrev>
|
||||
≥ 1.1), without dorsal or sublateral series of setiferous punctures; prosternum without longitudinal keel; antesternal membrane without distinct semisclerotized patch or patches; probasisternum triangular, with narrowed lateral arms and disc protruding medially, with pair of macrosetae on the upper probasiternum; with rounded postcoxal hypomeral process, interrupted by inferior line. Mesothorax: mesoscutellum without posterior scutellar carina, without sub-basal ridge; elytra with humeral spines or spine-like setae, with even setiferous punctation on disc and epipleuron (sometimes with various setose color patterns); with setiferous punctures at apical margin of elytral suture (underside); mesanapleural sutures transverse or nearly transverse but reaching and fusing with external part of prepectus; mesobasisternum with intercoxal process narrowly pointed into sharp angle, without V-shaped projection medially; mesocoxal cavities contiguous; pericoxal ridge present and complete. Metathorax: wings present, with veins CuA and
|
||||
<collectionCode id="E745D98176B724A214E705FA79796B88" country="0" name="Mohonk Preserve, Inc.">MP</collectionCode>
|
||||
4 fused; metakatepisternal processes divided; metascutellar mid-longitudinal internal suture well developed (only visible in chemically cleared specimens). Legs: apical tarsomere of all legs without dorsal setae, with one empodial seta distinctly shorter than tarsal claws; protarsomeres 1–3 distinctly wider than long in both sexes, with pale adhesive setae ventrally; procoxa with internal ridge running parallel along external ridge; mesotarsi in both sexes without black comb, sometimes in males ventral side of first mesotarsomere with pale adhesive setae; mesotibiae straight; metatarsomere 1 shorter than metatarsomere 5; metatarsi shorter than metatibiae; metatrochanter apically rounded, with strong straight dorsoapical spine; metacoxae with four or fewer spines on ventral posterolateral lobe, spines on dorsomedial disc also present; basal part of metacoxae distinctly wider and more convex than apical part.
|
||||
</paragraph>
|
||||
<caption id="78D88F8865660CFA11D8B3CDDE718BDF" ID-DOI="10.3897/zookeys.1218.135558.figure7" ID-arpha="87E2A45E-7E9D-53B3-8BC0-134CB60F5BF4" httpUri="https://binary.pensoft.net/fig/1175480" startId="F7">
|
||||
<paragraph id="F4189C9F9D210DC2FE5C1F6F5E012257">
|
||||
<label id="350163FB59E349A3E470F869BFECEA9E">Figure 7.</label>
|
||||
</paragraph>
|
||||
<paragraph id="CF94824C59838D759DB4A02C5F9A1FA6">
|
||||
Some
|
||||
<taxonomicName id="C69A18DF3AC9899F86B4B60AED33874A" class="Insecta" family="Staphylinidae" genus="Cheilocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="undefined-A">
|
||||
<emphasis id="3E04A7299A6F73FDAA4F3D429CB696DA" italics="true">Cheilocolpus</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
<emphasis id="AB8062F2B72E8D7E96084F4093654C79" bold="true">A</emphasis>
|
||||
<taxonomicName id="0D92A0D4598445940DC9DA3CA50DF3B4" authority="(Solier, 1849)" baseAuthorityName="Solier" baseAuthorityYear="1849" class="Insecta" family="Staphylinidae" genus="Cheilocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="pyrostoma">
|
||||
<emphasis id="93A213354E1D5FFBD3FC3A0BDF12E292" italics="true">Ch. pyrostoma</emphasis>
|
||||
(Solier, 1849)
|
||||
</taxonomicName>
|
||||
<emphasis id="A87E1BA87786967FBC6332664A139878" bold="true">B</emphasis>
|
||||
<taxonomicName id="C230AD3BBA12D2D776352E301D06A7ED" authority="(Bernhauer, 1917)" baseAuthorityName="Bernhauer" baseAuthorityYear="1917" class="Insecta" family="Staphylinidae" genus="Cheilocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="speciosus">
|
||||
<emphasis id="F3246B0196A8FE56923164DEE9AE59B7" italics="true">Ch. speciosus</emphasis>
|
||||
(Bernhauer, 1917)
|
||||
</taxonomicName>
|
||||
<emphasis id="3663FB219F6EAD9F07ABC6C5997CFD62" bold="true">C</emphasis>
|
||||
<taxonomicName id="7C170A0D39DE8252B9CE4E8E3BB85A0A" authority="(Erichson, 1840)" baseAuthorityName="Erichson" baseAuthorityYear="1840" class="Insecta" family="Staphylinidae" genus="Cheilocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="viridulus">
|
||||
<emphasis id="219509DDF16400C6E8692BBB9E0F1026" italics="true">Ch. viridulus</emphasis>
|
||||
(Erichson, 1840)
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8846A6DC7A8681B640CC48F7D6A4D28D">
|
||||
Abdomen: protergal glands well manifested as rounded cuticular acetabula; tergites III –
|
||||
<collectionCode id="4CC665E78F2B9AF50387BBD37C9B2A36" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
|
||||
in some species with posterior basal transverse carinae, in some species they are only with anterior carinae; on tergite VII anterior transverse basal carina not continuing to paratergites; tergites III –
|
||||
<collectionCode id="1D48DA6314897872E6650664AFF77AAD" country="Norway" name="Mykotektet, National Veterinary Institute">VI</collectionCode>
|
||||
with different patterns of posteromarginal large macroseta (
|
||||
<collectionCode id="A0A98ECB8C236E4FFECED32B9A9A37A9">PMM</collectionCode>
|
||||
) on each side, IV –
|
||||
<collectionCode id="55AA58E8D57F5B89C6F0AB545687ECC0" country="Norway" name="Mykotektet, National Veterinary Institute">VI</collectionCode>
|
||||
with more than one PMM per side; punctation of tergites in form of fine to moderate impressions, some species with wide glabrous areas; lateral tergal sclerites IX short and slightly laterally flattened; male sternite VIII with medial apical emargination. Aedeagus with paramere fused to median lobe only at base and very closely appressed to median lobe along entire length; base of the paramere flat medially, not projecting upwards in the middle; paramere strongly produced over apex of median lobe. Ovipositor with each second gonocoxite with one medial macroseta, without spine-like setae on outer lateral margin.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E1CBG" type="distribution">
|
||||
<paragraph id="A932052D6B2682886A1BB9E902DE4477">
|
||||
<heading id="655317367B1DDB8BC0D94FF8803366AC" reason="title">Distribution.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8B2B7BB6B5A6CDB6622355E6AD23B5DD">Neotropical Realm: Mexican Transition Zone, Central America, and northern South America.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E6CBG" type="ecology">
|
||||
<paragraph id="4A9326E3A6645FDED2595A5AD149E238">
|
||||
<heading id="BC1E976E7C7F98586A112B80FA557E34" reason="title">Ecology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="F3F64C0E96AC31DE2DD29C2CE1A17BEB">
|
||||
This genus has been collected from a variety of microhabitats, including mushrooms (e. g.,
|
||||
<taxonomicName id="C6772B0FEF7A627F627509FC1131D398" class="Agaricomycetes" family="Pleurotaceae" genus="Pleurotus" kingdom="Fungi" order="Agaricales" phylum="Basidiomycota" rank="species" species="undetermined">
|
||||
<emphasis id="501EBBA9D9B30FA6C88FAC65C7EB3D57" italics="true">Pleurotus</emphasis>
|
||||
spp.
|
||||
</taxonomicName>
|
||||
), under decomposing logs, beneath bark, and in leaf litter found in cloud, oak, and pine forests. They are also frequently captured using flight-intercept and Malaise traps. Additionally, some species are associated with the inflorescences of plants from the families
|
||||
<taxonomicName id="8C6AFC74879E488FB6E68B439AFD8891" class="Liliopsida" family="Arecaceae" kingdom="Plantae" order="Arecales" phylum="Tracheophyta" rank="family">Arecaceae</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="DBE98E0E2C1A8B31BAAFC3B1FB8017DA" class="Magnoliopsida" family="Betulaceae" kingdom="Plantae" order="Fagales" phylum="Tracheophyta" rank="family">Betulaceae</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="47EC1FA8FF666CF4B8D89617AFAC9A9D" class="Liliopsida" family="Heliconiaceae" kingdom="Plantae" order="Zingiberales" phylum="Tracheophyta" rank="family">Heliconiaceae</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E2DBG" type="comparisons">
|
||||
<paragraph id="F72140B93A3C53D3D5B59B092818ED8A">
|
||||
<heading id="080DB3688E88F37CA97B0D209790DCCF" reason="title">Comparison.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="6F3B1860D4959B06D93B1008D3D00E26">
|
||||
<taxonomicName id="3A8F0A96A0C4DA07170CDAE8005353D6" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="FCA67D1199B61CC44C308D8D19435E80" italics="true">Nitidocolpus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="97D9B55EDD3663AED455F326B0DF2402" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
can be distinguished from all other genera of
|
||||
<taxonomicName id="BD73FE2C777459746E06BB3FACD423EF" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subFamily" subFamily="Staphylininae">Staphylininae</taxonomicName>
|
||||
by the combination of characters mentioned in the diagnosis.
|
||||
<taxonomicName id="88AB3BD8A9A96606EAC34CF64D3849D7" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="435E9FE939BB8BFF3D77B1EE6E77E7DB" italics="true">Nitidocolpus</emphasis>
|
||||
</taxonomicName>
|
||||
is further distinguished from the superficially similar
|
||||
<taxonomicName id="795C9C6A049629D939D404AC63F2D62C" authorityName="Kraatz" authorityYear="1857" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subTribe" subTribe="Quediina">Quediina</taxonomicName>
|
||||
and Cytoquediina (Fig.
|
||||
<figureCitation id="74ADC8772D1A72B0FAFD6832459B53A5" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Morphological details of Nitidocolpus and other relevant genera of Staphylininae A – C Nitidocolpus columbinus D – F Cheilocolpus viridulus G – I Cyrtoquedius sp. J – L Quedionuchus nr. impunctus M – O Quedius nr. advena. A, D, G, J, M head B, E, H, K, N pronotum C, F, I, L, O aedeagus in lateral view, the black arrow pointing down indicates the basal projection of the paramere. Abbreviations: ACP, anterolateral clypeal puncture; AFP, anterior frontal; AMRSP, anterior marginal row of setiferous punctures; APP, additional paired punctures adjacent to AMRSP; BP, basal puncture; FCP, frontoclypeal puncture; IASP, internalmost setiferous puncture of the anterior margin; IOP, interocular punctures; IOR, infraorbital ridge; LLSP, large lateral setiferous puncture; LPSP, large posterior setiferous puncture; NR, nuchal ridge; PFP, posterior frontal puncture; POP, parocular punctures; PPDS, paired punctures on dorsal series; SAP, supra-antennal puncture; SAR, supra-antennal ridge; SLSP, sublateral setiferous puncture; TP, temporal puncture. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure6" httpUri="https://binary.pensoft.net/fig/1175479">6</figureCitation>
|
||||
) by the lack of supra-antennal punctures, the presence of a single carina on the mesoscutellar shield (posterior carina absent), by hind wings with fused CuA and
|
||||
<collectionCode id="B0DB2AB56106E9B011081BCF408FBD8D" country="0" name="Mohonk Preserve, Inc.">MP</collectionCode>
|
||||
4 veins, by a single empodial setae, and by the base of the paramere, which has no upward or forward projection in the middle. Compared to other Neotropical genera that resemble
|
||||
<taxonomicName id="8454F70F9A4FF7D73C1CDD8E28399AB8" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="C6B9B2F16BE73F7FA2D9058CD0BAB895" italics="true">Nitidocolpus</emphasis>
|
||||
</taxonomicName>
|
||||
, such as
|
||||
<taxonomicName id="E2233370A5C5BBBAF8EAADB696E75A9B" class="Insecta" family="Staphylinidae" genus="Cheilocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="FF5B0B7D24633D17582BD425B24695A0" italics="true">Cheilocolpus</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="3BFEB3AF32BAC825042B2E3B607A0742" authority="Sharp, 1884" authorityName="Sharp" authorityYear="1884" class="Insecta" family="Staphylinidae" genus="Mimosticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B179E17168F0597B3654FE2FBFBE0E1E" italics="true">Mimosticus</emphasis>
|
||||
Sharp, 1884
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="67B2DF18671FBE77BE7AD27EA1D641EA" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Heterothops" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="F20FE31BF51626F4B5662A329B30020B" italics="true">Heterothops</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="80E5830DE8E856B20E49F16B660E7C42" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="1ED2149E8B18CD721229212FD05B6AD3" italics="true">Nitidocolpus</emphasis>
|
||||
</taxonomicName>
|
||||
is distinguished by the absence of paraocular punctures on the head, the lack of setiferous punctures in or near the center of the pronotal disc, and the presence of postcoxal hypomeral processes. In some species of
|
||||
<taxonomicName id="850423B305E76A2CDD11C9CD6D5BC463" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="624A0D3B8DA38B98A64D92741B77CF02" italics="true">Nitidocolpus</emphasis>
|
||||
</taxonomicName>
|
||||
, such as
|
||||
<taxonomicName id="7989A8C757B71CFD4EFAB23C2DF20364" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1884" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="laeticulus">
|
||||
<emphasis id="5021AA8071EB390A90929C3FCC18FD6A" italics="true">N. laeticulus</emphasis>
|
||||
</taxonomicName>
|
||||
(Fig.
|
||||
<figureCitation id="8CA7E3E8693119F77FE2B0883B2531A4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Some Nitidocolpus species and characters A N. illatus (Sharp, 1884) B N. laeticulus (Sharp, 1884) C N. germaini (Bernhauer, 1917) D N. aurofasciatus (Bernhauer, 1917) E N. illatus, pterothoraces F N. columbinus (Bernhauer, 1917), metacoxa and metatrochanter, dissected right leg in dorsal view (side faced to the abdomen). Abbreviations: EXPR, external part of prepectus; INPR, internal part of prepectus; MIV, mesobasisternal intercoxal process with V-shaped projection medially; MNPS, mesanapleural sutures; MTMS, metacoxa with dorsomedial spine; MTTAS, metatrochanter with dorsoapical spine. Photo B by J. Jenkins Shaw. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure5" httpUri="https://binary.pensoft.net/fig/1175478">5 B</figureCitation>
|
||||
) and
|
||||
<taxonomicName id="44C01BF14F1ECBCFD243A9DEEB1B1184" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Bernhauer" baseAuthorityYear="1917" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="aurofasciatus">
|
||||
<emphasis id="E52ABCC48CDE491C2C9D2DFA568C72EC" italics="true">N. aurofasciatus</emphasis>
|
||||
</taxonomicName>
|
||||
(Fig.
|
||||
<figureCitation id="191E8248A9883B040ADECC399D35DAD2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Some Nitidocolpus species and characters A N. illatus (Sharp, 1884) B N. laeticulus (Sharp, 1884) C N. germaini (Bernhauer, 1917) D N. aurofasciatus (Bernhauer, 1917) E N. illatus, pterothoraces F N. columbinus (Bernhauer, 1917), metacoxa and metatrochanter, dissected right leg in dorsal view (side faced to the abdomen). Abbreviations: EXPR, external part of prepectus; INPR, internal part of prepectus; MIV, mesobasisternal intercoxal process with V-shaped projection medially; MNPS, mesanapleural sutures; MTMS, metacoxa with dorsomedial spine; MTTAS, metatrochanter with dorsoapical spine. Photo B by J. Jenkins Shaw. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure5" httpUri="https://binary.pensoft.net/fig/1175478">5 D</figureCitation>
|
||||
), the basal punctures have shifted anteriorly and thus may be erroneously considered paraocular punctures. However, they can be easily distinguished as basal punctures, because they are the only punctures located near the posterior frontal punctures, aside from the temporal punctures. In other species of
|
||||
<taxonomicName id="EEBCC3C617091F98905B72B4D0D8C716" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="7D7C8C91796B5AC0F7880FF929B42D16" italics="true">Nitidocolpus</emphasis>
|
||||
</taxonomicName>
|
||||
, such as
|
||||
<taxonomicName id="973C7890DD05BF463ABB532F5C0A4799" authorityName="Reyes-Hernández & Solodovnikov" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1884" class="Insecta" family="Staphylinidae" genus="Nitidocolpus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="illatus">
|
||||
<emphasis id="73147A09965C3B29E0E6BED64E2151A9" italics="true">N. illatus</emphasis>
|
||||
</taxonomicName>
|
||||
(Fig.
|
||||
<figureCitation id="C7830F06D3D3D687D48411EF774D9859" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Some Nitidocolpus species and characters A N. illatus (Sharp, 1884) B N. laeticulus (Sharp, 1884) C N. germaini (Bernhauer, 1917) D N. aurofasciatus (Bernhauer, 1917) E N. illatus, pterothoraces F N. columbinus (Bernhauer, 1917), metacoxa and metatrochanter, dissected right leg in dorsal view (side faced to the abdomen). Abbreviations: EXPR, external part of prepectus; INPR, internal part of prepectus; MIV, mesobasisternal intercoxal process with V-shaped projection medially; MNPS, mesanapleural sutures; MTMS, metacoxa with dorsomedial spine; MTTAS, metatrochanter with dorsoapical spine. Photo B by J. Jenkins Shaw. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.135558.figure5" httpUri="https://binary.pensoft.net/fig/1175478">5 A</figureCitation>
|
||||
), the innermost macrosetae on the anterior margin maybe erroneously considered as a puncture of the dorsal row because it is located far from the anterior margin of pronotum.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EBIBG" type="etymology">
|
||||
<paragraph id="8BE992AB09E57F707CF44707183F3FF9">
|
||||
<heading id="D3E5A76E4072583447FFC6C84BE6FF0B" reason="title">Etymology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="97ED032E54B619B1999BB0235033A7C3">The name is derived from the Latin words “nitidus” and “colpus”, which mean “shiny” and “hit”, respectively. The name refers to the great clarity with which the head and pronotum punctures are seen. The gender is masculine.</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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