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<document id="C596CDD965A9022A928CB1E4B768DF21" ID-DOI="10.1093/isd/ixab003" ID-ISSN="2399-3421" ID-Zenodo-Dep="5384533" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630299435056" checkinUser="valdenar" docAuthor="Means, Jackson C., Hennen, Derek A., Tanabe, Tsutomu &amp; Marek, Paul E." docDate="2021" docId="332087BBFF8FB774471FFC6861EBA761" docLanguage="en" docName="ixab003.pdf" docOrigin="Insect Systematics and Diversity 5 (2), No. 1" docSource="http://dx.doi.org/10.1093/isd/ixab003" docStyle="DocumentStyle:20C34595BA01015AC07A3B955E96DF74.8:InsectSystDivers.2017-.journal_article" docStyleId="20C34595BA01015AC07A3B955E96DF74" docStyleName="InsectSystDivers.2017-.journal_article" docStyleVersion="8" docTitle="Brachoria platana Means, Hennen, Marek 2021, New Species" docType="treatment" docVersion="4" lastPageNumber="14" masterDocId="CF19FFC3FF83B779442DFFF36515A159" masterDocTitle="Phylogenetic Systematics of the Millipede Family Xystodesmidae" masterLastPageNumber="26" masterPageNumber="1" pageNumber="13" updateTime="1699219996674" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="7C55577B4AFE0DF70F21DF315C5A38E6">Phylogenetic Systematics of the Millipede Family Xystodesmidae</mods:title>
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<mods:namePart id="B7C9710D9567D7A39C49ED89E7521F41">Means, Jackson C.</mods:namePart>
<mods:affiliation id="7144E53BB5C7702C42E160466CF623D1">Department of Entomology, Virginia Tech, Blacksburg, VA, USA,</mods:affiliation>
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<mods:namePart id="31B733E0087E85747D9D1DC57E4DCBA9">Hennen, Derek A.</mods:namePart>
<mods:affiliation id="3BF45873F7F69B2DC3B126F1D5D24BCF">Department of Entomology, Virginia Tech, Blacksburg, VA, USA,</mods:affiliation>
</mods:name>
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<mods:namePart id="2B12454E23D3637F4B51E074114E705B">Tanabe, Tsutomu</mods:namePart>
<mods:affiliation id="247E8C6151128E0AEE845928B8DCAB10">Faculty of Advanced Science and Technology, Kumamoto</mods:affiliation>
</mods:name>
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<mods:namePart id="C5B0E32639AAFEAC4F23125888A964E8">Marek, Paul E.</mods:namePart>
<mods:affiliation id="FDA30BC99EB615EE74CBEB7803A53832">Department of Entomology, Virginia Tech, Blacksburg, VA, USA,</mods:affiliation>
<mods:nameIdentifier id="8AFB690EB05AB0C830F7433A93D19CA7" type="email">paulemarek@gmail.com</mods:nameIdentifier>
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<mods:title id="CFF15B6E519DA585172AEC7C2863B867">Insect Systematics and Diversity</mods:title>
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<mods:date id="CBC36921E4C034309D9394823E9F24FB">2021</mods:date>
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<treatment id="332087BBFF8FB774471FFC6861EBA761" ID-DOI="http://doi.org/10.5281/zenodo.5466767" ID-Zenodo-Dep="5466767" LSID="urn:lsid:plazi:treatment:332087BBFF8FB774471FFC6861EBA761" httpUri="http://treatment.plazi.org/id/332087BBFF8FB774471FFC6861EBA761" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<subSubSection id="F3936526FF8FB775471FFC6866E2A578" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="BB3636ADFF8FB775471FFC6866C3A28D" blockId="12.[818,1343,923,980]" pageId="12" pageNumber="13">
<heading id="E07E81C1FF8FB775471FFC6866C3A28D" bold="true" fontSize="10" level="2" pageId="12" pageNumber="13" reason="3">
<emphasis id="89FDEABFFF8FB775471FFC6866C3A28D" bold="true" pageId="12" pageNumber="13">
<taxonomicName id="7C894D2EFF8FB775471FFC68602EA2ED" ID-CoL="86Y53" authority="Means, Hennen, Marek" authorityName="Means, Hennen, Marek" authorityYear="2021" box="[818,1339,923,948]" class="Diplopoda" family="Xystodesmidae" genus="Brachoria" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="platana" status="sp. nov.">
<emphasis id="89FDEABFFF8FB775471FFC686105A2ED" bold="true" box="[818,1040,923,948]" italics="true" pageId="12" pageNumber="13">Brachoria platana</emphasis>
Means, Hennen, Marek
</taxonomicName>
,
<taxonomicNameLabel id="92CE57C4FF8FB775471FFC4866C3A28D" box="[818,982,954,980]" pageId="12" pageNumber="13" rank="species">New Species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="BB3636ADFF8FB775471FFC1466E2A578" blockId="12.[818,1396,999,1057]" pageId="12" pageNumber="13">
<heading id="E07E81C1FF8FB775471FFC1466E2A578" fontSize="9" level="1" pageId="12" pageNumber="13" reason="6">
Vernacular Name: The Carr Creek Mimic Millipede
<figureCitation id="23B22A28FF8FB775471FFBF966B6A578" box="[818,931,1034,1057]" captionStart="Fig" captionStartId="9.[146,173,1782,1799]" captionTargetBox="[284,1303,147,1747]" captionTargetId="figure-2@9.[284,1303,147,1747]" captionTargetPageId="9" captionText="Fig. 5. Color photographs of new species of xystodesmid millipedes and their color morphs. Appalachioria bondi n. sp. (AG): AThree-spotted yellow morph with yellow legs, dorsal view, B Two-spotted pink morph with pink legs, dorsal view, C Three-spotted white morph with white legs, DThree-spotted orange morph with orange legs, ETwo-spotted yellow morph with pink legs, F.Two-spotted white morph with white legs, GThree-spotted yellow morph with pink legs, dorsal view. Appalachioria sierwaldae, n. sp. (HI): H. Three-spotted morph with yellow paranotal spots and orange medial spots and orange legs. I. Threespotted yellow morph with yellow legs. Appalachioria brownae n. sp. (JK): J. Three-spotted yellow morph with yellow legs. K. Two-spotted yellow morph with yellow legs. Brachoria camptera n. sp. (LM): L. Two-spotted orange morph with white legs. M. Two-spotted red morph with red legs. Brachoria cryocybe n. sp. (N): N. Two-spotted morph with red paranotal spots and white stripe and red legs. Brachoria platana n. sp. (OP): O. Striped red morph with red legs. P. Threespotted yellow morph with yellow legs. Rudiloria charityae n. sp. (Q): Q. Three-spotted morph with yellow paranotal spots and orange medial spots and yellow legs. Sigmoria beameri n. sp. (R): R. Two-spotted red morph with light red legs. Nannaria aenigma n. sp. (ST): S. Two-spotted red morph with light red legs. T. Two-spotted white morph with white legs. Nannaria hokie n. sp. (U):Two-spotted red morph with gray legs." figureDoi="http://doi.org/10.5281/zenodo.5384543" httpUri="https://zenodo.org/record/5384543/files/figure.png" pageId="12" pageNumber="13">Figs. 5O,P</figureCitation>
and
<figureCitation id="23B22A28FF8FB77547F7FBF966FAA578" box="[986,1007,1034,1057]" captionStart="Fig" captionStartId="13.[146,173,1182,1199]" captionTargetBox="[314,1274,733,1147]" captionTargetId="figure-347@13.[314,1274,733,1147]" captionTargetPageId="13" captionText="Fig. 11. Brachoria platana n. sp. holotype ♂ left gonopod (VTEC MPE03250). (A) Anterior view; (B) Medial view; (C) Posterior view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5384558" httpUri="https://zenodo.org/record/5384558/files/figure.png" pageId="12" pageNumber="13">11</figureCitation>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F3936526FF8FB775471FFBB766E2A451" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="BB3636ADFF8FB775471FFBB766E2A451" blockId="12.[818,1443,1090,1288]" pageId="12" pageNumber="13">
<emphasis id="89FDEABFFF8FB775471FFBB766FDA50E" bold="true" box="[818,1000,1092,1111]" italics="true" pageId="12" pageNumber="13">Material examined:</emphasis>
Type specimens.
<specimenCount id="AD8FFD24FF8FB77540AEFBB16186A501" box="[1155,1171,1090,1112]" pageId="12" pageNumber="13" type="male"></specimenCount>
<typeStatus id="6432880FFF8FB77540BAFBB761FDA501" box="[1175,1256,1092,1112]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
(
<collectionCode id="DD98AE68FF8FB77540DFFBB76038A501" box="[1266,1325,1092,1112]" pageId="12" pageNumber="13">VTEC</collectionCode>
MPE03250) and
<specimenCount id="AD8FFD24FF8FB7754776FB93667EA52F" box="[859,875,1120,1142]" pageId="12" pageNumber="13" type="female"></specimenCount>
<typeStatus id="6432880FFF8FB775475CFB9066D7A52F" box="[881,962,1123,1142]" pageId="12" pageNumber="13" type="paratype">paratype</typeStatus>
(
<collectionCode id="DD98AE68FF8FB77547E2FB91611FA52F" box="[975,1034,1122,1142]" pageId="12" pageNumber="13">VTEC</collectionCode>
MPE03253) from
<collectingRegion id="794DF84FFF8FB7754091FB916001A52F" box="[1212,1300,1122,1142]" country="United States of America" name="Kentucky" pageId="12" pageNumber="13">Kentucky</collectingRegion>
, Knott County, Carr Creek State Park, woods behind campground and around marina (
<geoCoordinate id="DEBD506AFF8FB77547A9FB6F66EFA5E9" box="[900,1018,1180,1200]" degrees="37.23148" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="1" value="37.23148">37.23148°N</geoCoordinate>
, -
<geoCoordinate id="DEBD506AFF8FB7754023FB6F6190A5E9" box="[1038,1157,1180,1200]" degrees="83.00081" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="1" value="-83.00081">83.00081°W</geoCoordinate>
, Elev.
<quantity id="7C719B48FF8FB77540E5FB6F601FA5E9" box="[1224,1290,1180,1200]" metricMagnitude="2" metricUnit="m" metricValue="3.29" pageId="12" pageNumber="13" unit="m" value="329.0">329 m</quantity>
),
<date id="CF37106DFF8FB7754132FB6F6676A597" pageId="12" pageNumber="13" value="2017-09-27">27 September 2017</date>
, 15:00, dry litter of tulip poplar, sycamore, autumn olive (Coll: J. C. Means and D. A. Hennen). Material examined listed in Supp Table 2 (online only).
</paragraph>
</subSubSection>
<subSubSection id="F3936526FF8FB774471FFAC6679FA7DC" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="BB3636ADFF8FB774471FFAC6679FA7DC" blockId="12.[818,1441,1332,1382]" lastBlockId="13.[146,770,1268,1669]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis id="89FDEABFFF8FB775471FFAC66682A411" bold="true" box="[818,919,1333,1352]" italics="true" pageId="12" pageNumber="13">Diagnosis:</emphasis>
Adult males of
<taxonomicName id="7C894D2EFF8FB7754010FAC761FEA411" authority="Means &amp; Hennen &amp; Tanabe &amp; Marek, 2021" authorityName="Means &amp; Hennen &amp; Tanabe &amp; Marek" authorityYear="2021" box="[1085,1259,1332,1352]" class="Diplopoda" family="Xystodesmidae" genus="Brachoria" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="platana" status="sp. nov.">
<emphasis id="89FDEABFFF8FB7754010FAC761FEA411" box="[1085,1259,1332,1352]" italics="true" pageId="12" pageNumber="13">Brachoria platana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92CE57C4FF8FB77540D8FAC66039A411" box="[1269,1324,1333,1352]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
are distinct from other apheloriine species based on the following combination of characters:
<emphasis id="89FDEABFFF8EB7744516FB0764B1A451" bold="true" box="[315,420,1268,1288]" pageId="13" pageNumber="14">Gonopods.</emphasis>
Gonopodal acropodite curving dorsomedially at apex, with a distal cingulum, separating it from
<taxonomicName id="7C894D2EFF8EB77444BFFADD65E5A41B" authorityName="Chamberlin" authorityYear="1921" box="[146,240,1326,1346]" class="Diplopoda" family="Xystodesmidae" genus="Apheloria" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="89FDEABFFF8EB77444BFFADD65E5A41B" box="[146,240,1326,1346]" italics="true" pageId="13" pageNumber="14">Apheloria</emphasis>
</taxonomicName>
and
<taxonomicName id="7C894D2EFF8EB7744504FADD6496A41B" authorityName="Causey" authorityYear="1955" box="[297,387,1326,1346]" class="Diplopoda" family="Xystodesmidae" genus="Rudiloria" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="89FDEABFFF8EB7744504FADD6496A41B" box="[297,387,1326,1346]" italics="true" pageId="13" pageNumber="14">Rudiloria</emphasis>
</taxonomicName>
(
<figureCitation id="23B22A28FF8EB77445BBFADD64C9A41B" box="[406,476,1326,1346]" captionStart="Fig" captionStartId="13.[146,173,1182,1199]" captionTargetBox="[314,1274,733,1147]" captionTargetId="figure-347@13.[314,1274,733,1147]" captionTargetPageId="13" captionText="Fig. 11. Brachoria platana n. sp. holotype ♂ left gonopod (VTEC MPE03250). (A) Anterior view; (B) Medial view; (C) Posterior view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5384558" httpUri="https://zenodo.org/record/5384558/files/figure.png" pageId="13" pageNumber="14">Fig. 11</figureCitation>
). Prefemoral process reduced to small ridge. Distal zone sinuous, curving first ventrally and then dorsally; in anterior view directed posteriorly at 90° from apex. Acropodite tip with lateral and medial flanges forming an envelope-like structure through which runs an elevated ridge carrying the solenomere (
<figureCitation id="23B22A28FF8EB77445BAFA3264FAA48C" box="[407,495,1473,1493]" captionStart="Fig" captionStartId="13.[146,173,1182,1199]" captionTargetBox="[314,1274,733,1147]" captionTargetId="figure-347@13.[314,1274,733,1147]" captionTargetPageId="13" captionText="Fig. 11. Brachoria platana n. sp. holotype ♂ left gonopod (VTEC MPE03250). (A) Anterior view; (B) Medial view; (C) Posterior view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5384558" httpUri="https://zenodo.org/record/5384558/files/figure.png" pageId="13" pageNumber="14">Fig. 11A</figureCitation>
).
<emphasis id="89FDEABFFF8EB7744624FA326751A48C" bold="true" box="[521,580,1473,1493]" pageId="13" pageNumber="14">Color.</emphasis>
<taxonomicName id="7C894D2EFF8EB774467CFA326617A48C" authority="Means &amp; Hennen &amp; Tanabe &amp; Marek, 2021" authorityName="Means &amp; Hennen &amp; Tanabe &amp; Marek" authorityYear="2021" box="[593,770,1473,1493]" class="Diplopoda" family="Xystodesmidae" genus="Brachoria" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="platana" status="sp. nov.">
<emphasis id="89FDEABFFF8EB774467CFA326617A48C" box="[593,770,1473,1493]" italics="true" pageId="13" pageNumber="14">Brachoria platana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92CE57C4FF8EB77444BFFA2C65DDA4AB" box="[146,200,1503,1522]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
has two color morphs, co-occurring. Tergites always with red or yellow paranotal spots, sometimes also with concolorous metatergal middorsal spots or stripes. Dark-brown to black background. Collum always with concolorous red or yellow anterior and lateral spots, sometimes also with a posterior spot and concolorous marginal lines connecting the spots.
</paragraph>
</subSubSection>
<caption id="EFF66625FF8FB77544BCF8616075A6FA" ID-DOI="http://doi.org/10.5281/zenodo.5384554" ID-Zenodo-Dep="5384554" box="[145,1376,1936,1955]" httpUri="https://zenodo.org/record/5384554/files/figure.png" pageId="12" pageNumber="13" startId="12.[145,172,1938,1955]" targetBox="[314,1274,1453,1903]" targetPageId="12">
<paragraph id="BB3636ADFF8FB77544BCF8616075A6FA" blockId="12.[145,1376,1936,1955]" box="[145,1376,1936,1955]" pageId="12" pageNumber="13">
<emphasis id="89FDEABFFF8FB77544BCF86165D7A6FA" bold="true" box="[145,194,1938,1955]" pageId="12" pageNumber="13">Fig. 9.</emphasis>
<taxonomicName id="7C894D2EFF8FB77544E6F861647EA6FA" authority="Means &amp; Hennen &amp; Tanabe &amp; Marek, 2021" authorityName="Means &amp; Hennen &amp; Tanabe &amp; Marek" authorityYear="2021" box="[203,363,1938,1955]" class="Diplopoda" family="Xystodesmidae" genus="Brachoria" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="camptera" status="sp. nov.">
<emphasis id="89FDEABFFF8FB77544E6F861647EA6FA" box="[203,363,1938,1955]" italics="true" pageId="12" pageNumber="13">Brachoria camptera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92CE57C4FF8FB7754542F8616488A6FA" box="[367,413,1938,1955]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
holotype ♂ left gonopod (VTEC MPE01398). (A) Anterior view; (B) Medial view; (C) Posterior view. Scale bar = 1.0 mm.
</paragraph>
</caption>
<caption id="EFF66625FF8EB77444BFFD9C6072A3D9" ID-DOI="http://doi.org/10.5281/zenodo.5384556" ID-Zenodo-Dep="5384556" box="[146,1383,621,640]" httpUri="https://zenodo.org/record/5384556/files/figure.png" pageId="13" pageNumber="14" startId="13.[146,173,623,640]" targetBox="[314,1274,147,588]" targetPageId="13">
<paragraph id="BB3636ADFF8EB77444BFFD9C6072A3D9" blockId="13.[146,1383,621,640]" box="[146,1383,621,640]" pageId="13" pageNumber="14">
<emphasis id="89FDEABFFF8EB77444BFFD9C65D9A3D9" bold="true" box="[146,204,623,640]" pageId="13" pageNumber="14">Fig. 10.</emphasis>
<taxonomicName id="7C894D2EFF8EB77444FEFD9C6465A3D9" authority="Means &amp; Hennen &amp; Tanabe &amp; Marek, 2021" authorityName="Means &amp; Hennen &amp; Tanabe &amp; Marek" authorityYear="2021" box="[211,368,623,640]" class="Diplopoda" family="Xystodesmidae" genus="Brachoria" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="cryocybe" status="sp. nov.">
<emphasis id="89FDEABFFF8EB77444FEFD9C6465A3D9" box="[211,368,623,640]" italics="true" pageId="13" pageNumber="14">Brachoria cryocybe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92CE57C4FF8EB7744558FD9C64B6A3D9" box="[373,419,623,640]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
holotype ♂ left gonopod (VTEC MPE02644). (A) Anterior view; (B) Medial view; (C) Posterior view. Scale bar = 1.0 mm.
</paragraph>
</caption>
<caption id="EFF66625FF8EB77444BFFB6D604DA5F6" ID-DOI="http://doi.org/10.5281/zenodo.5384558" ID-Zenodo-Dep="5384558" box="[146,1368,1180,1199]" httpUri="https://zenodo.org/record/5384558/files/figure.png" pageId="13" pageNumber="14" startId="13.[146,173,1182,1199]" targetBox="[314,1274,733,1147]" targetPageId="13">
<paragraph id="BB3636ADFF8EB77444BFFB6D604DA5F6" blockId="13.[146,1368,1180,1199]" box="[146,1368,1180,1199]" pageId="13" pageNumber="14">
<emphasis id="89FDEABFFF8EB77444BFFB6D65DFA5F6" bold="true" box="[146,202,1182,1199]" pageId="13" pageNumber="14">Fig. 11.</emphasis>
<taxonomicName id="7C894D2EFF8EB77444FFFB6D6477A5F6" authority="Means &amp; Hennen &amp; Tanabe &amp; Marek, 2021" authorityName="Means &amp; Hennen &amp; Tanabe &amp; Marek" authorityYear="2021" box="[210,354,1182,1199]" class="Diplopoda" family="Xystodesmidae" genus="Brachoria" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="platana" status="sp. nov.">
<emphasis id="89FDEABFFF8EB77444FFFB6D6477A5F6" box="[210,354,1182,1199]" italics="true" pageId="13" pageNumber="14">Brachoria platana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92CE57C4FF8EB774454BFB6D6481A5F6" box="[358,404,1182,1199]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
holotype ♂ left gonopod (VTEC MPE03250). (A) Anterior view; (B) Medial view; (C) Posterior view. Scale bar = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="F3936526FF8EB77444BFF9476464A6D9" pageId="13" pageNumber="14" type="description">
<paragraph id="BB3636ADFF8EB77444BFF94764F2A65B" blockId="13.[146,770,1713,1794]" pageId="13" pageNumber="14">
<emphasis id="89FDEABFFF8EB77444BFF9476411A79E" bold="true" box="[146,260,1716,1735]" italics="true" pageId="13" pageNumber="14">Description:</emphasis>
Supp Table 3 (online only). Based on
<typeStatus id="6432880FFF8EB77446AEF94767CFA791" box="[643,730,1716,1736]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
(
<specimenCount id="AD8FFD24FF8EB77446C6F94267EEA79E" box="[747,763,1713,1735]" pageId="13" pageNumber="14" type="male"></specimenCount>
) MPE03250. Measurements (mm): BL = 41.0, CW = 7.4, IW = 4.7, ISW = 1.6, B11W = 9.4, B11H = 5.4.
</paragraph>
<paragraph id="BB3636ADFF8EB77444BFF8C26464A6D9" blockId="13.[146,770,1841,1920]" pageId="13" pageNumber="14">
<emphasis id="89FDEABFFF8EB77444BFF8C265E4A61D" bold="true" box="[146,241,1841,1860]" italics="true" pageId="13" pageNumber="14">Variation:</emphasis>
Individuals of
<taxonomicName id="7C894D2EFF8EB77445A4F8C26727A61C" authority="Means &amp; Hennen &amp; Tanabe &amp; Marek, 2021" authorityName="Means &amp; Hennen &amp; Tanabe &amp; Marek" authorityYear="2021" box="[393,562,1841,1861]" class="Diplopoda" family="Xystodesmidae" genus="Brachoria" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="platana" status="sp. nov.">
<emphasis id="89FDEABFFF8EB77445A4F8C26727A61C" box="[393,562,1841,1861]" italics="true" pageId="13" pageNumber="14">Brachoria platana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92CE57C4FF8EB7744610F8C16766A61C" box="[573,627,1842,1861]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
are known to exhibit two color morphs, 1) striped red (
<figureCitation id="23B22A28FF8EB7744607F8BD6766A63B" box="[554,627,1870,1890]" captionStart="Fig" captionStartId="9.[146,173,1782,1799]" captionTargetBox="[284,1303,147,1747]" captionTargetId="figure-2@9.[284,1303,147,1747]" captionTargetPageId="9" captionText="Fig. 5. Color photographs of new species of xystodesmid millipedes and their color morphs. Appalachioria bondi n. sp. (AG): AThree-spotted yellow morph with yellow legs, dorsal view, B Two-spotted pink morph with pink legs, dorsal view, C Three-spotted white morph with white legs, DThree-spotted orange morph with orange legs, ETwo-spotted yellow morph with pink legs, F.Two-spotted white morph with white legs, GThree-spotted yellow morph with pink legs, dorsal view. Appalachioria sierwaldae, n. sp. (HI): H. Three-spotted morph with yellow paranotal spots and orange medial spots and orange legs. I. Threespotted yellow morph with yellow legs. Appalachioria brownae n. sp. (JK): J. Three-spotted yellow morph with yellow legs. K. Two-spotted yellow morph with yellow legs. Brachoria camptera n. sp. (LM): L. Two-spotted orange morph with white legs. M. Two-spotted red morph with red legs. Brachoria cryocybe n. sp. (N): N. Two-spotted morph with red paranotal spots and white stripe and red legs. Brachoria platana n. sp. (OP): O. Striped red morph with red legs. P. Threespotted yellow morph with yellow legs. Rudiloria charityae n. sp. (Q): Q. Three-spotted morph with yellow paranotal spots and orange medial spots and yellow legs. Sigmoria beameri n. sp. (R): R. Two-spotted red morph with light red legs. Nannaria aenigma n. sp. (ST): S. Two-spotted red morph with light red legs. T. Two-spotted white morph with white legs. Nannaria hokie n. sp. (U):Two-spotted red morph with gray legs." figureDoi="http://doi.org/10.5281/zenodo.5384543" httpUri="https://zenodo.org/record/5384543/files/figure.png" pageId="13" pageNumber="14">Fig. 5O</figureCitation>
) and 2) threespotted yellow (
<figureCitation id="23B22A28FF8EB7744508F89F6471A6D9" box="[293,356,1900,1920]" captionStart="Fig" captionStartId="9.[146,173,1782,1799]" captionTargetBox="[284,1303,147,1747]" captionTargetId="figure-2@9.[284,1303,147,1747]" captionTargetPageId="9" captionText="Fig. 5. Color photographs of new species of xystodesmid millipedes and their color morphs. Appalachioria bondi n. sp. (AG): AThree-spotted yellow morph with yellow legs, dorsal view, B Two-spotted pink morph with pink legs, dorsal view, C Three-spotted white morph with white legs, DThree-spotted orange morph with orange legs, ETwo-spotted yellow morph with pink legs, F.Two-spotted white morph with white legs, GThree-spotted yellow morph with pink legs, dorsal view. Appalachioria sierwaldae, n. sp. (HI): H. Three-spotted morph with yellow paranotal spots and orange medial spots and orange legs. I. Threespotted yellow morph with yellow legs. Appalachioria brownae n. sp. (JK): J. Three-spotted yellow morph with yellow legs. K. Two-spotted yellow morph with yellow legs. Brachoria camptera n. sp. (LM): L. Two-spotted orange morph with white legs. M. Two-spotted red morph with red legs. Brachoria cryocybe n. sp. (N): N. Two-spotted morph with red paranotal spots and white stripe and red legs. Brachoria platana n. sp. (OP): O. Striped red morph with red legs. P. Threespotted yellow morph with yellow legs. Rudiloria charityae n. sp. (Q): Q. Three-spotted morph with yellow paranotal spots and orange medial spots and yellow legs. Sigmoria beameri n. sp. (R): R. Two-spotted red morph with light red legs. Nannaria aenigma n. sp. (ST): S. Two-spotted red morph with light red legs. T. Two-spotted white morph with white legs. Nannaria hokie n. sp. (U):Two-spotted red morph with gray legs." figureDoi="http://doi.org/10.5281/zenodo.5384543" httpUri="https://zenodo.org/record/5384543/files/figure.png" pageId="13" pageNumber="14">Fig. 5P</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F3936526FF8EB774471FFB0761B4A41B" pageId="13" pageNumber="14" type="biology_ecology">
<paragraph id="BB3636ADFF8EB774471FFB0761B4A41B" blockId="13.[818,1442,1268,1346]" pageId="13" pageNumber="14">
<emphasis id="89FDEABFFF8EB774471FFB076697A45E" bold="true" box="[818,898,1268,1287]" italics="true" pageId="13" pageNumber="14">Ecology:</emphasis>
Individuals of
<taxonomicName id="7C894D2EFF8EB774403AFB0761D5A451" authority="Means &amp; Hennen &amp; Tanabe &amp; Marek, 2021" authorityName="Means &amp; Hennen &amp; Tanabe &amp; Marek" authorityYear="2021" box="[1047,1216,1268,1288]" class="Diplopoda" family="Xystodesmidae" genus="Brachoria" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="platana" status="sp. nov.">
<emphasis id="89FDEABFFF8EB774403AFB0761D5A451" box="[1047,1216,1268,1288]" italics="true" pageId="13" pageNumber="14">Brachoria platana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92CE57C4FF8EB77440E4FB0761EBA45E" box="[1225,1278,1268,1287]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
were found in a mesic deciduous forest of tulip poplar, sycamore and autumn olive by the waters edge of Carr Creek Lake.
</paragraph>
</subSubSection>
<subSubSection id="F3936526FF8EB774471FFA9D666EA4F9" pageId="13" pageNumber="14" type="distribution">
<paragraph id="BB3636ADFF8EB774471FFA9D666EA4F9" blockId="13.[818,1442,1390,1440]" pageId="13" pageNumber="14">
<emphasis id="89FDEABFFF8EB774471FFA9D614DA4DB" box="[818,1112,1390,1410]" italics="true" pageId="13" pageNumber="14">
<emphasis id="89FDEABFFF8EB774471FFA9D66BEA4D8" bold="true" box="[818,939,1390,1409]" italics="true" pageId="13" pageNumber="14">Distribution:</emphasis>
<taxonomicName id="7C894D2EFF8EB774479FFA9D614DA4DB" authority="Means &amp; Hennen &amp; Tanabe &amp; Marek, 2021" authorityName="Means &amp; Hennen &amp; Tanabe &amp; Marek" authorityYear="2021" box="[946,1112,1390,1410]" class="Diplopoda" family="Xystodesmidae" genus="Brachoria" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="platana" status="sp. nov.">Brachoria platana</taxonomicName>
</emphasis>
<taxonomicNameLabel id="92CE57C4FF8EB7744072FA9C6187A4DB" box="[1119,1170,1391,1410]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
is known only from the
<typeStatus id="6432880FFF8EB7744156FA9C60B7A4DB" box="[1403,1442,1391,1410]" pageId="13" pageNumber="14">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="F3936526FF8EB774471FFA3F61EBA761" pageId="13" pageNumber="14" type="etymology">
<paragraph id="BB3636ADFF8EB774471FFA3F61EBA761" blockId="13.[818,1443,1484,1592]" pageId="13" pageNumber="14">
<emphasis id="89FDEABFFF8EB774471FFA3F6688A486" bold="true" box="[818,925,1484,1503]" italics="true" pageId="13" pageNumber="14">Etymology:</emphasis>
This species is named after where the
<typeStatus id="6432880FFF8EB774410EFA3F6061A4B9" box="[1315,1396,1484,1504]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
was discovered, in a grove of American sycamores (
<taxonomicName id="7C894D2EFF8EB77440FBFA1A6083A4A4" box="[1238,1430,1513,1533]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="occidentalis">
<emphasis id="89FDEABFFF8EB77440FBFA1A6083A4A4" box="[1238,1430,1513,1533]" italics="true" pageId="13" pageNumber="14">Platanus occidentalis</emphasis>
</taxonomicName>
). The specific name is a noun in apposition derived from the Latin genus name of the American sycamore,
<taxonomicName id="7C894D2EFF8EB774408EF9D761E7A761" box="[1187,1266,1572,1592]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">
<emphasis id="89FDEABFFF8EB774408EF9D761E7A761" box="[1187,1266,1572,1592]" italics="true" pageId="13" pageNumber="14">Platanus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>