487 lines
62 KiB
XML
487 lines
62 KiB
XML
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<mods:title id="7EE6765F178146CCD22B867BAD17DE81">Taxonomic revision of the Phanaeus endymion species group (Coleoptera: Scarabaeidae), with the descriptions of five new species</mods:title>
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<mods:namePart id="96D298A1342CE454987192CA82BF0421">Moctezuma, Victor</mods:namePart>
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<mods:affiliation id="FAFF833D7D086F8B24A9DF9A79A9AB26">Red de Ecoetología, Instituto de Ecología, A. C. Carretera antigua a Coatepec 351, El Haya, Xalapa, Veracruz 91073, Mexico.</mods:affiliation>
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<mods:affiliation id="4EF7D80342A61F6AF74585094824F600">Red de Ecoetología, Instituto de Ecología, A. C. Carretera antigua a Coatepec 351, El Haya, Xalapa, Veracruz 91073, Mexico.</mods:affiliation>
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<mods:nameIdentifier id="AC27658B560A8895A3CCD967E3A9B8DB" type="email">gonzalohalffter@inecol.mx</mods:nameIdentifier>
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<mods:title id="5BB211EF7FD6C2C61A987845966C206F">European Journal of Taxonomy</mods:title>
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<treatment id="03CA87F2FFEAFFD4FD844748BCE73C8D" ID-DOI="http://doi.org/10.5281/zenodo.4726223" ID-GBIF-Taxon="181753768" ID-Zenodo-Dep="4726223" LSID="urn:lsid:plazi:treatment:03CA87F2FFEAFFD4FD844748BCE73C8D" httpUri="http://treatment.plazi.org/id/03CA87F2FFEAFFD4FD844748BCE73C8D" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
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<subSubSection id="C379656FFFEAFFD5FD844748BB7A3F67" box="[556,1032,1347,1374]" pageId="20" pageNumber="21" type="nomenclature">
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<paragraph id="8BDC36E4FFEAFFD5FD844748BB7A3F67" blockId="20.[556,1032,1347,1411]" box="[556,1032,1347,1374]" pageId="20" pageNumber="21">
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<heading id="D0948188FFEAFFD5FD844748BB7A3F67" box="[556,1032,1347,1374]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="21" reason="2">
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<taxonomicName id="4C634D67FFEAFFD5FD844748BB7A3F67" ID-CoL="4FSJM" authority="Edmonds, 1979" authorityName="Edmonds" authorityYear="1979" box="[556,1032,1347,1374]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="halffterorum">
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<emphasis id="B917EAF6FFEAFFD5FD844748BC3A3F64" bold="true" box="[556,840,1347,1374]" italics="true" pageId="20" pageNumber="21">Phanaeus halffterorum</emphasis>
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<bibRefCitation id="EFF24B15FFEAFFD5FCE74748BB7A3F67" author="Edmonds W. D." box="[847,1032,1347,1373]" pageId="20" pageNumber="21" pagination="99 - 105" refId="ref37459" refString="Edmonds W. D. 1979. A new species of Phanaeus from Mexico (Coleoptera: Scarabaeidae). Pan-Pacific Entomologist 55: 99 - 105." type="journal article" year="1979">Edmonds, 1979</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BDC36E4FFEAFFD5FD3F4763BCEE3FB8" blockId="20.[556,1032,1347,1411]" box="[663,924,1384,1411]" pageId="20" pageNumber="21">
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<figureCitation id="13582A61FFEAFFD5FD3F4763BD823FB8" box="[663,752,1384,1410]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta & Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma & Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="20" pageNumber="21">Figs 1F</figureCitation>
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,
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<figureCitation id="13582A61FFEAFFD5FD544763BC693FB9" box="[764,795,1384,1411]" captionStart="Fig" captionStartId="50.[189,232,1890,1916]" captionTargetBox="[193,1392,927,1854]" captionTargetId="figure-235@50.[189,1398,924,1857]" captionTargetPageId="50" captionText="Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017;P. edmondsi Moctezuma, Deloya & Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 by Jose Luis Sánchez-Huerta." figureDoi="http://doi.org/10.5281/zenodo.4836796" httpUri="https://zenodo.org/record/4836796/files/figure.png" pageId="20" pageNumber="21">15</figureCitation>
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,
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<figureCitation id="13582A61FFEAFFD5FC804763BC2C3FB8" box="[808,862,1384,1410]" captionStart="Fig" captionStartId="55.[189,232,1681,1707]" captionTargetBox="[258,1329,264,1646]" captionTargetId="figure-157@55.[258,1329,264,1646]" captionTargetPageId="55" captionText="Fig. 18. Pronotum of females within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta).C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta).H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002. K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836802" httpUri="https://zenodo.org/record/4836802/files/figure.png" pageId="20" pageNumber="21">18G</figureCitation>
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,
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||
<figureCitation id="13582A61FFEAFFD5FCC24763BCEE3FB8" box="[874,924,1384,1410]" captionStart="Fig" captionStartId="56.[189,231,1678,1704]" captionTargetBox="[306,1282,265,1646]" captionTargetId="figure-158@56.[306,1282,265,1646]" captionTargetPageId="56" captionText="Fig. 19. Elytral integument of species within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez- Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta). H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002.K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836804" httpUri="https://zenodo.org/record/4836804/files/figure.png" pageId="20" pageNumber="21">19G</figureCitation>
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</paragraph>
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<paragraph id="8BDC36E4FFEAFFD5FF1547A0BCE03FFC" blockId="20.[189,1090,1451,1515]" box="[189,914,1451,1478]" pageId="20" pageNumber="21">
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<treatmentCitationGroup id="AB7311CAFFEAFFD5FF1547A0BCE03FFC" box="[189,914,1451,1478]" pageId="20" pageNumber="21">
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<treatmentCitation id="0AC210F5FFEAFFD5FF1547A0BDB43FFF" author="Edmonds W. D. & Halffter G." box="[189,710,1451,1477]" page="321" pageId="20" pageNumber="21" year="1978">
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<taxonomicName id="4C634D67FFEAFFD5FF1547A0BDB43FFF" ID-CoL="4FSJM" authority="Edmonds, 1978: 321" authorityName="Edmonds" authorityPageNumber="321" authorityYear="1978" box="[189,710,1451,1477]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="halffterorum">
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||
<emphasis id="B917EAF6FFEAFFD5FF1547A0BEB93FFF" box="[189,459,1451,1477]" italics="true" pageId="20" pageNumber="21">Phanaeus halffterorum</emphasis>
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<bibRefCitation id="EFF24B15FFEAFFD5FE7B47A0BDB43FFF" author="Edmonds W. D. & Halffter G." box="[467,710,1451,1477]" pageId="20" pageNumber="21" pagination="307 - 331" refId="ref37704" refString="Edmonds W. D. & Halffter G. 1978. Taxonomic review of immature dung beetles of the dung beetles of the subfamily Scarabaeinae (Coleoptera: Scarabaeidae). Systematic Entomology 3: 307 - 331. https: // doi. org / 10.1111 / j. 1365 - 3113.1978. tb 00002. x" type="journal article" year="1978">Edmonds, 1978: 321</bibRefCitation>
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</taxonomicName>
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(nomen nudum).
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</paragraph>
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<paragraph id="8BDC36E4FFEAFFD5FF1547DBBB303FD1" blockId="20.[189,1090,1451,1515]" box="[189,1090,1488,1515]" pageId="20" pageNumber="21">
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<treatmentCitationGroup id="AB7311CAFFEAFFD5FF1547DBBB303FD1" box="[189,1090,1488,1515]" pageId="20" pageNumber="21">
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<treatmentCitation id="0AC210F5FFEAFFD5FF1547DBBDC83FD0" author="Edmonds W. D." box="[189,698,1488,1514]" page="99" pageId="20" pageNumber="21" year="1979">
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<taxonomicName id="4C634D67FFEAFFD5FF1547DBBDC83FD0" ID-CoL="4FSJM" authority="Edmonds, 1979: 99" authorityName="Edmonds" authorityPageNumber="99" authorityYear="1979" box="[189,698,1488,1514]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="halffterorum">
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<emphasis id="B917EAF6FFEAFFD5FF1547DBBEB93FD0" box="[189,459,1488,1514]" italics="true" pageId="20" pageNumber="21">Phanaeus halffterorum</emphasis>
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<bibRefCitation id="EFF24B15FFEAFFD5FE7B47DBBDC83FD0" author="Edmonds W. D." box="[467,698,1488,1514]" pageId="20" pageNumber="21" pagination="99 - 105" refId="ref37459" refString="Edmonds W. D. 1979. A new species of Phanaeus from Mexico (Coleoptera: Scarabaeidae). Pan-Pacific Entomologist 55: 99 - 105." type="journal article" year="1979">Edmonds, 1979: 99</bibRefCitation>
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</taxonomicName>
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, 102–105, figs 1–3, 6–8 (in part).
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</paragraph>
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<paragraph id="8BDC36E4FFEAFFD5FF154418BEB13D6F" blockId="20.[189,1399,1555,2025]" pageId="20" pageNumber="21">
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<treatmentCitationGroup id="AB7311CAFFEAFFD5FF154418BEB13D6F" pageId="20" pageNumber="21">
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<taxonomicName id="4C634D67FFEAFFD5FF154418BEBD3C17" ID-CoL="4FSJM" authorityName="Edmonds" authorityYear="1979" box="[189,463,1555,1581]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="halffterorum">
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<emphasis id="B917EAF6FFEAFFD5FF154418BEBD3C17" box="[189,463,1555,1581]" italics="true" pageId="20" pageNumber="21">Phanaeus halffterorum</emphasis>
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</taxonomicName>
|
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–
|
||
<treatmentCitation id="0AC210F5FFEAFFD5FE5C4418BCFC3C17" author="Halffter G. & Edmonds W. D." box="[500,910,1555,1581]" page="88" pageId="20" pageNumber="21" year="1982">
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<bibRefCitation id="EFF24B15FFEAFFD5FE5C4418BCFC3C17" author="Halffter G. & Edmonds W. D." box="[500,910,1555,1581]" pageId="20" pageNumber="21" refId="ref38583" refString="Halffter G. & Edmonds W. D. 1982. The nesting Behavior of Dung Beetles (Scarabaeinae). An ecological and evolutive Approach. Instituto de Ecologia, Mexico City." type="book" year="1982">Halffter & Edmonds 1982: 88–89</bibRefCitation>
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</treatmentCitation>
|
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(in part). —
|
||
<treatmentCitation id="0AC210F5FFEAFFD5FB9D4418BE643C69" author="Anduaga S. & Halffter G." page="157" pageId="20" pageNumber="21" year="1991">
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<bibRefCitation id="EFF24B15FFEAFFD5FB9D4418BE643C69" author="Anduaga S. & Halffter G." pageId="20" pageNumber="21" pagination="151 - 169" refId="ref35569" refString="Anduaga S. & Halffter G. 1991. Micofagia en Scarabaeidae. In: Navarrete-Heredia J. L. & Quiroz- Rocha G. A. (eds) Memorias del I Simposio Nacional sobre la Interaccion Insecto-Hongo: 151 - 169. S. M. E. Facultad de Ciencias, IGU, Veracruz." type="book chapter" year="1991">Anduaga & Halffter 1991: 157</bibRefCitation>
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</treatmentCitation>
|
||
(in part). —
|
||
<treatmentCitation id="0AC210F5FFEAFFD5FE194433BC483C69" author="Delgado-Castillo L. & Navarrete-Heredia J. L. & Blackaller-Bages J." box="[433,826,1592,1619]" page="125" pageId="20" pageNumber="21" year="1993">
|
||
<bibRefCitation id="EFF24B15FFEAFFD5FE194433BC483C69" author="Delgado-Castillo L. & Navarrete-Heredia J. L. & Blackaller-Bages J." box="[433,826,1592,1619]" pageId="20" pageNumber="21" pagination="121 - 126" refId="ref36993" refString="Delgado-Castillo L., Navarrete-Heredia J. L. & Blackaller-Bages J. 1993. A new Mexican species of Onthophagus with mycophagous habits (Coleoptera: Scarabaeidae: Scarabaeinae). The Coleopterists Bulletin 47: 121 - 126. Available from https: // www. jstor. org / stable / 4008857 [accessed 2 Mar. 2021]." type="journal article" year="1993">
|
||
Delgado-Castillo
|
||
<emphasis id="B917EAF6FFEAFFD5FD294432BDC73C68" box="[641,693,1592,1618]" italics="true" pageId="20" pageNumber="21">et al</emphasis>
|
||
. 1993: 125
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</bibRefCitation>
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||
</treatmentCitation>
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||
(in part). —
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<treatmentCitation id="0AC210F5FFEAFFD5FC7D4433BBAC3C68" author="Deloya C. & Burgos A. & Blackaller J. & Lobo J. M." box="[981,1246,1592,1619]" page="21" pageId="20" pageNumber="21" year="1993">
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<bibRefCitation id="EFF24B15FFEAFFD5FC7D4433BBAC3C68" author="Deloya C. & Burgos A. & Blackaller J. & Lobo J. M." box="[981,1246,1592,1619]" pageId="20" pageNumber="21" pagination="15 - 55" refId="ref37208" refString="Deloya C., Burgos A., Blackaller J. & Lobo J. M. 1993. Los coleopteros lamelicornios de Cuernavaca, Morelos, Mexico (Passalidae, Trogidae, Scarabaeidae y Melolonthidae). Boletin de la Sociedad veracruzana de Zoologia 3: 15 - 55." type="journal article" year="1993">
|
||
Deloya
|
||
<emphasis id="B917EAF6FFEAFFD5FB994432BB173C68" box="[1073,1125,1592,1618]" italics="true" pageId="20" pageNumber="21">et al</emphasis>
|
||
. 1993: 21
|
||
</bibRefCitation>
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||
</treatmentCitation>
|
||
, 39 (in part); 2014: 77 (in part). —
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||
<treatmentCitation id="0AC210F5FFEAFFD5FE584456BDAE3C42" author="Anduaga S." box="[496,732,1629,1656]" page="125" pageId="20" pageNumber="21" year="2000">
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||
<bibRefCitation id="EFF24B15FFEAFFD5FE584456BDAE3C42" author="Anduaga S." box="[496,732,1629,1656]" pageId="20" pageNumber="21" pagination="119 - 130" refId="ref35521" refString="Anduaga S. 2000. Escarabajos coprofagos (Coleoptera: Scarabaeoidea) asociados a hongos en la Sierra Madre Occidental, Durango, Mexico: con una compilacion de las especies micetofagas. Acta Zoologica Mexicana (n. s.) 80: 119 - 130." type="journal article" year="2000">Anduaga 2000: 125</bibRefCitation>
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||
</treatmentCitation>
|
||
, 130 (in part). —
|
||
<treatmentCitation id="0AC210F5FFEAFFD5FC1D4456BA043C4D" author="Lopez-Guerrero Y. & Halffter G." box="[949,1398,1629,1655]" page="241" pageId="20" pageNumber="21" year="2000">
|
||
<bibRefCitation id="EFF24B15FFEAFFD5FC1D4456BA043C4D" author="Lopez-Guerrero Y. & Halffter G." box="[949,1398,1629,1655]" pageId="20" pageNumber="21" pagination="225 - 285" refId="ref39888" refString="Lopez-Guerrero Y. & Halffter G. 2000. Evolution of the spermatheca in the Scarabaeoidea (Coleoptera). Fragmenta Entomologica, Roma 32: 225 - 285." type="journal article" year="2000">López-Guerrero & Halffter 2000: 241</bibRefCitation>
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||
</treatmentCitation>
|
||
(in part). —
|
||
<treatmentCitation id="0AC210F5FFEAFFD5FE2E4489BD103CA6" author="Arnaud P." box="[390,610,1666,1692]" page="96" pageId="20" pageNumber="21" year="2002">
|
||
<bibRefCitation id="EFF24B15FFEAFFD5FE2E4489BD103CA6" author="Arnaud P." box="[390,610,1666,1692]" pageId="20" pageNumber="21" refId="ref35859" refString="Arnaud P. 2002 b. Phanaeini. Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus. Hillside Books, Canterbury." type="book" year="2002">Arnaud 2002b: 96</bibRefCitation>
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||
</treatmentCitation>
|
||
(in part). —
|
||
<treatmentCitation id="0AC210F5FFEAFFD5FCA24489BCB93CA6" author="Price D. L." box="[778,971,1666,1693]" page="197" pageId="20" pageNumber="21" year="2005">
|
||
<bibRefCitation id="EFF24B15FFEAFFD5FCA24489BCB93CA6" author="Price D. L." box="[778,971,1666,1693]" pageId="20" pageNumber="21" pagination="197 - 203" refId="ref41704" refString="Price D. L. 2005. Descriptions of the male and female genitalia of Phanaeus (Macleay) (Scarabaeidae: Scarabaeinae): the vindex species group. The Coleopterists Bulletin 59: 197 - 203. https: // doi. org / 10.1649 / 743" type="journal article" year="2005">Price 2005: 197</bibRefCitation>
|
||
</treatmentCitation>
|
||
(in part); 2007: 17, figs 52–54. —
|
||
<treatmentCitation id="0AC210F5FFEAFFD5FF4244ACBE873CFB" author="Edmonds W. D." box="[234,501,1703,1729]" page="31" pageId="20" pageNumber="21" year="2006">
|
||
<bibRefCitation id="EFF24B15FFEAFFD5FF4244ACBE873CFB" author="Edmonds W. D." box="[234,501,1703,1729]" pageId="20" pageNumber="21" pagination="31 - 37" refId="ref37655" refString="Edmonds W. D. 2006. A new species of Phanaeus Macleay (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini) from Oaxaca, Mexico. Zootaxa 1171 (3): 31 - 37. https: // doi. org / 10.11646 / zootaxa. 1171.1.3" type="journal article" year="2006">Edmonds 2006: 31–32</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 36, fig. 7 (in part). —
|
||
<treatmentCitation id="0AC210F5FFEAFFD5FCA844ACBB553CFB" author="Ceballos G. & List R. & Garduno G. & Lopez-Cano R. & Munozcano-Quintanar M. J. & Collado E. & San-Roman J. E." box="[768,1063,1703,1729]" page="397" pageId="20" pageNumber="21" year="2009">
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||
<bibRefCitation id="EFF24B15FFEAFFD5FCA844ACBB553CFB" author="Ceballos G. & List R. & Garduno G. & Lopez-Cano R. & Munozcano-Quintanar M. J. & Collado E. & San-Roman J. E." box="[768,1063,1703,1729]" pageId="20" pageNumber="21" refId="ref36502" refString="Ceballos G., List R., Garduno G., Lopez-Cano R., Munozcano-Quintanar M. J., Collado E. & San-Roman J. E. 2009. La diversidad biologica del Estado de Mexico. Estudio de estado. Gobierno del Estado de Mexico, Toluca de Lerdo." type="book" year="2009">
|
||
Ceballos
|
||
<emphasis id="B917EAF6FFEAFFD5FCC544A3BCD23CFB" box="[877,928,1703,1729]" italics="true" pageId="20" pageNumber="21">et al</emphasis>
|
||
. 2009: 397
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AC210F5FFEAFFD5FBFD44ACBF8B3CDD" author="Edmonds W. D. & Zidek J." page="5" pageId="20" pageNumber="21" year="2012">
|
||
<bibRefCitation id="EFF24B15FFEAFFD5FBFD44ACBF8B3CDD" author="Edmonds W. D. & Zidek J." pageId="20" pageNumber="21" pagination="1 - 108" refId="ref37762" refString="Edmonds W. D. & Zidek J. 2012. Taxonomy of Phanaeus revisited: Revised keys to and comments on species of the New World dung beetle genus Phanaeus Macleay, 1819 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini). Insecta Mundi 274: 1 - 108." type="journal article" year="2012">Edmonds & Zídek 2012: 5</bibRefCitation>
|
||
</treatmentCitation>
|
||
(in part). —
|
||
<treatmentCitation id="0AC210F5FFEAFFD5FE3144C7BD073CDD" author="Krajcik M." box="[409,629,1740,1767]" page="150" pageId="20" pageNumber="21" year="2006">
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||
<bibRefCitation id="EFF24B15FFEAFFD5FE3144C7BD073CDD" author="Krajcik M." box="[409,629,1740,1767]" pageId="20" pageNumber="21" pagination="1 - 189" refId="ref39668" refString="Krajcik M. 2006. Checklist of Scarabaeoidea of the World. 1. Scarabaeinae (Coleoptera: Scarabaeidae: Scarabaeinae). Animma. X. Suplement 3: 1 - 189." type="journal article" year="2006">Krajcik 2006: 150</bibRefCitation>
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||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AC210F5FFEAFFD5FD0044C7BB5E3CDD" author="Moctezuma V. & Halffter G." box="[680,1068,1740,1767]" httpUri="http://treatment.plazi.org/id/DD6BB5FB8EDC36DE0A5A792CDAD4EFE1" page="52" pageId="20" pageNumber="21" year="2017">
|
||
<bibRefCitation id="EFF24B15FFEAFFD5FD0044C7BB5E3CDD" author="Moctezuma V. & Halffter G." box="[680,1068,1740,1767]" pageId="20" pageNumber="21" pagination="47 - 56" refId="ref40548" refString="Moctezuma V. & Halffter G. 2017. A new species of Phanaeus Macleay (Coleoptera: Scarabaeidae: Scarabaeinae) from Los Chimalapas, Oaxaca, Mexico. The Coleopterists Bulletin 71: 47 - 56. https: // doi. org / 10.1649 / 0010 - 065 X- 71.1.47" type="journal article" year="2017">Moctezuma & Halffter 2017: 52</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 54–55, fig. 23 (in part). — Moctezuma
|
||
<emphasis id="B917EAF6FFEAFFD5FED644F9BEC13D31" box="[382,435,1777,1803]" italics="true" pageId="20" pageNumber="21">et al</emphasis>
|
||
. 2017: 113–115, 118–119, 122, 130–132, figs 1–5; 2019: 253, fig. 5. —
|
||
<treatmentCitation id="0AC210F5FFEAFFD5FAB344FABED93D0A" author="Lizardo V. & Escobar F. & Rojas-Soto O." page="273" pageId="20" pageNumber="21" year="2017">
|
||
<bibRefCitation id="EFF24B15FFEAFFD5FAB344FABED93D0A" author="Lizardo V. & Escobar F. & Rojas-Soto O." pageId="20" pageNumber="21" pagination="271 - 294" refId="ref39838" refString="Lizardo V., Escobar F. & Rojas-Soto O. 2017. Diversity and distribution of Phanaeini (Coleoptera: Scarabaeidae: Scarabaeinae) in Mexico. Zootaxa 4358 (3): 271 - 294. https: // doi. org / 10.11646 / zootaxa. 4358.2.3" type="journal article" year="2017">
|
||
Lizardo
|
||
<emphasis id="B917EAF6FFEAFFD5FF42451DBE523D0A" box="[234,288,1814,1840]" italics="true" pageId="20" pageNumber="21">et al</emphasis>
|
||
. 2017: 273
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 275, 292, fig.13 (in part). —
|
||
<treatmentCitation id="0AC210F5FFEAFFD5FCB0451DBB3D3D0A" author="Kohlmann B. & Arriaga-Jimenez A. & Ros M." box="[792,1103,1814,1840]" httpUri="http://treatment.plazi.org/id/7E2DEA7201CB96E41326FCCC778FC035" page="69" pageId="20" pageNumber="21" year="2018">
|
||
<bibRefCitation id="EFF24B15FFEAFFD5FCB0451DBB3D3D0A" author="Kohlmann B. & Arriaga-Jimenez A. & Ros M." box="[792,1103,1814,1840]" pageId="20" pageNumber="21" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. & Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">
|
||
Kohlmann
|
||
<emphasis id="B917EAF6FFEAFFD5FC35451DBCA03D0A" box="[925,978,1814,1840]" italics="true" pageId="20" pageNumber="21">et al</emphasis>
|
||
. 2018: 69
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||
</bibRefCitation>
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||
</treatmentCitation>
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||
, 81, 88–89. —
|
||
<treatmentCitation id="0AC210F5FFEAFFD5FAA3451DBEB23D6F" author="Gillett C. P. D. T. & Toussaint E. F. A." page="2" pageId="20" pageNumber="21" year="2020">
|
||
<bibRefCitation id="EFF24B15FFEAFFD5FAA3451DBEB23D6F" author="Gillett C. P. D. T. & Toussaint E. F. A." pageId="20" pageNumber="21" pagination="661 - 682" refId="ref38342" refString="Gillett C. P. D. T. & Toussaint E. F. A. 2020. Macroevolution and shifts in the feeding biology of the New World scarab beetle tribe Phanaeini (Coleoptera: Scarabaeidae: Scarabaeinae). Biological Journal of the Linnean Society 130: 661 - 682. https: // doi. org / 10.1093 / biolinnean / blaa 058" type="journal article" year="2020">Gillett & Toussaint 2020: 2</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFEAFFD5FF154554BC1A3DD2" blockId="20.[189,1399,1555,2025]" pageId="20" pageNumber="21">
|
||
<treatmentCitationGroup id="AB7311CAFFEAFFD5FF154554BC1A3DD2" pageId="20" pageNumber="21">
|
||
<taxonomicName id="4C634D67FFEAFFD5FF154554BDE43D43" ID-CoL="4FSJM" authorityName="Edmonds" authorityYear="1979" box="[189,662,1887,1914]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="halffterorum" subGenus="Notiophanaeus">
|
||
<emphasis id="B917EAF6FFEAFFD5FF154554BE433D43" box="[189,305,1887,1913]" italics="true" pageId="20" pageNumber="21">Phanaeus</emphasis>
|
||
(
|
||
<emphasis id="B917EAF6FFEAFFD5FEE94554BE813D43" box="[321,499,1887,1913]" italics="true" pageId="20" pageNumber="21">Notiophanaeus</emphasis>
|
||
)
|
||
<emphasis id="B917EAF6FFEAFFD5FDAB4554BDE43D43" box="[515,662,1887,1913]" italics="true" pageId="20" pageNumber="21">halffterorum</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AC210F5FFEAFFD5FD1A4554BCF33D40" author="Edmonds W. D." box="[690,897,1887,1914]" page="2" pageId="20" pageNumber="21" year="1994">
|
||
<bibRefCitation id="EFF24B15FFEAFFD5FD1A4554BCF33D40" author="Edmonds W. D." box="[690,897,1887,1914]" pageId="20" pageNumber="21" pagination="1 - 105" refId="ref37487" refString="Edmonds W. D. 1994. Revision of Phanaeus Macleay, a New World genus of Scarabaeinae dung beetles (Coleoptera: Scarabaeidae, Scarabaeinae). Contributions in Science, Natural History Museum of Los Angeles County 443: 1 - 105." type="journal article" year="1994">Edmonds 1994: 2</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 8–9, 19, 39, 41, 43–44, 101, figs 211, 213, 217–218, 221 (in part); 2003: 61, 65 (in part). —
|
||
<treatmentCitation id="0AC210F5FFEAFFD5FC86458FBB713DA5" author="Arnaud P." box="[814,1027,1924,1951]" page="95" pageId="20" pageNumber="21" year="2002">
|
||
<bibRefCitation id="EFF24B15FFEAFFD5FC86458FBB713DA5" author="Arnaud P." box="[814,1027,1924,1951]" pageId="20" pageNumber="21" refId="ref35859" refString="Arnaud P. 2002 b. Phanaeini. Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus. Hillside Books, Canterbury." type="book" year="2002">Arnaud 2002b: 95</bibRefCitation>
|
||
</treatmentCitation>
|
||
(in part). —
|
||
<treatmentCitation id="0AC210F5FFEAFFD5FB36458FBE313DFE" author="Edmonds W. D. & Zidek J." page="3" pageId="20" pageNumber="21" year="2012">
|
||
<bibRefCitation id="EFF24B15FFEAFFD5FB36458FBE313DFE" author="Edmonds W. D. & Zidek J." pageId="20" pageNumber="21" pagination="1 - 108" refId="ref37762" refString="Edmonds W. D. & Zidek J. 2012. Taxonomy of Phanaeus revisited: Revised keys to and comments on species of the New World dung beetle genus Phanaeus Macleay, 1819 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini). Insecta Mundi 274: 1 - 108." type="journal article" year="2012">Edmonds & Zídek 2012: 3</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 12, figs 134–135, 137, 143–147 (in part). —
|
||
<treatmentCitation id="0AC210F5FFEAFFD5FCFA45A2BB143DFE" author="Lizardo V. & Escobar F. & Rojas-Soto O." box="[850,1126,1961,1988]" page="272" pageId="20" pageNumber="21" year="2017">
|
||
<bibRefCitation id="EFF24B15FFEAFFD5FCFA45A2BB143DFE" author="Lizardo V. & Escobar F. & Rojas-Soto O." box="[850,1126,1961,1988]" pageId="20" pageNumber="21" pagination="271 - 294" refId="ref39838" refString="Lizardo V., Escobar F. & Rojas-Soto O. 2017. Diversity and distribution of Phanaeini (Coleoptera: Scarabaeidae: Scarabaeinae) in Mexico. Zootaxa 4358 (3): 271 - 294. https: // doi. org / 10.11646 / zootaxa. 4358.2.3" type="journal article" year="2017">
|
||
Lizardo
|
||
<emphasis id="B917EAF6FFEAFFD5FC1B45A1BC973DF9" box="[947,997,1961,1987]" italics="true" pageId="20" pageNumber="21">et al</emphasis>
|
||
. 2017: 272
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
(in part). —
|
||
<treatmentCitation id="0AC210F5FFEAFFD5FB5345A2BEE53DD2" author="Kohlmann B. & Arriaga-Jimenez A. & Ros M." httpUri="http://treatment.plazi.org/id/7E2DEA7201CB96E41326FCCC778FC035" page="80" pageId="20" pageNumber="21" year="2018">
|
||
<bibRefCitation id="EFF24B15FFEAFFD5FB5345A2BEE53DD2" author="Kohlmann B. & Arriaga-Jimenez A. & Ros M." pageId="20" pageNumber="21" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. & Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">
|
||
Kohlmann
|
||
<emphasis id="B917EAF6FFEAFFD5FF4145C4BE6E3DD2" box="[233,284,1998,2024]" italics="true" pageId="20" pageNumber="21">et al</emphasis>
|
||
. 2018: 80
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||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AC210F5FFEAFFD5FE6F45C5BC163DD2" author="Zaragoza-Caballero S. & Rodriguez-Miron G. M. & Vega-Badillo V. & Gonzalez-Ramirez M. & Zurita-Garcia M. L. & Dominguez-Leon D. E. & Lopez-Perez S. & Gutierrez-Carranza I. G. & Cifuentes-Ruiz P. & Hernandez C. X. & Ramirez-del-Valle E. & Gutierrez N." box="[455,868,1998,2025]" page="43" pageId="20" pageNumber="21" year="2019">
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||
<bibRefCitation id="EFF24B15FFEAFFD5FE6F45C5BC163DD2" author="Zaragoza-Caballero S. & Rodriguez-Miron G. M. & Vega-Badillo V. & Gonzalez-Ramirez M. & Zurita-Garcia M. L. & Dominguez-Leon D. E. & Lopez-Perez S. & Gutierrez-Carranza I. G. & Cifuentes-Ruiz P. & Hernandez C. X. & Ramirez-del-Valle E. & Gutierrez N." box="[455,868,1998,2025]" pageId="20" pageNumber="21" pagination="1 - 122" refId="ref43055" refString="Zaragoza-Caballero S., Rodriguez-Miron G. M., Vega-Badillo V., Gonzalez-Ramirez M., Zurita-Garcia M. L., Dominguez-Leon D. E., Lopez-Perez S., Gutierrez-Carranza I. G., Cifuentes-Ruiz P., Perez- Hernandez C. X., Ramirez-del-Valle E. & Gutierrez N. 2019. A checklist of the Coleoptera (Insecta) from Morelos, Mexico. Zootaxa 4580: 1 - 122. https: // doi. org / 10.11646 / zootaxa. 4580.1.1" type="journal article" year="2019">
|
||
Zaragoza-Caballero
|
||
<emphasis id="B917EAF6FFEAFFD5FD1F45C4BD993DD2" box="[695,747,1998,2024]" italics="true" pageId="20" pageNumber="21">et al</emphasis>
|
||
. 2019: 43
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C379656FFFEBFFD4FF154302BDD13927" pageId="21" pageNumber="22" type="materials_examined">
|
||
<paragraph id="8BDC36E4FFEBFFD4FF154302BE1E3B18" blockId="21.[189,364,264,291]" box="[189,364,264,291]" pageId="21" pageNumber="22">
|
||
<heading id="D0948188FFEBFFD4FF154302BE1E3B18" bold="true" box="[189,364,264,291]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
|
||
<emphasis id="B917EAF6FFEBFFD4FF154302BE1E3B18" bold="true" box="[189,364,264,291]" pageId="21" pageNumber="22">Type material</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFEBFFD4FF424331BE8B3B6F" blockId="21.[189,1057,314,379]" box="[234,505,314,341]" pageId="21" pageNumber="22">
|
||
<emphasis id="B917EAF6FFEBFFD4FF424331BE2E3B6E" bold="true" box="[234,348,314,340]" pageId="21" pageNumber="22">
|
||
<typeStatus id="54D88846FFEBFFD4FF424331BE2E3B6E" box="[234,348,314,340]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
(not studied)
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFEBFFD4FF15436ABB533B41" blockId="21.[189,1057,314,379]" box="[189,1057,353,379]" pageId="21" pageNumber="22">
|
||
<materialsCitation id="3B0B3CB9FFEBFFD4FF15436ABB533B41" ID-GBIF-Occurrence="3093980440" box="[189,1057,353,379]" collectionCode="CAS" country="Mexico" location="Temascaltepec" municipality="State" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" stateProvince="State of Mexico" typeStatus="holotype">
|
||
<collectingCountry id="F3747674FFEBFFD4FF15436ABE433B41" box="[189,305,353,379]" name="Mexico" pageId="21" pageNumber="22">MEXICO</collectingCountry>
|
||
•
|
||
<specimenCount id="9D65FD6DFFEBFFD4FEE2436ABE103B41" box="[330,354,353,379]" pageId="21" pageNumber="22" type="male">♂</specimenCount>
|
||
,
|
||
<bibRefCitation id="EFF24B15FFEBFFD4FEC6436ABD223B41" author="Edmonds W. D." box="[366,592,353,379]" pageId="21" pageNumber="22" pagination="99 - 105" refId="ref37459" refString="Edmonds W. D. 1979. A new species of Phanaeus from Mexico (Coleoptera: Scarabaeidae). Pan-Pacific Entomologist 55: 99 - 105." type="journal article" year="1979">Edmonds 1979: 99</bibRefCitation>
|
||
;
|
||
<collectingRegion id="49A7F806FFEBFFD4FDF3436ABC683B41" box="[603,794,353,379]" pageId="21" pageNumber="22">State of Mexico</collectingRegion>
|
||
,
|
||
<location id="8EBC603FFFEBFFD4FC8D436ABCA73B41" LSID="urn:lsid:plazi:treatment:03CA87F2FFEAFFD4FD844748BCE73C8D:8EBC603FFFEBFFD4FC8D436ABCA73B41" box="[805,981,353,379]" country="Mexico" municipality="State" name="Temascaltepec" pageId="21" pageNumber="22" stateProvince="State of Mexico">Temascaltepec</location>
|
||
;
|
||
<collectionCode id="ED72AE21FFEBFFD4FC49436ABB6C3B41" box="[993,1054,353,379]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="21" pageNumber="22" type="Museum">CAS</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFEBFFD4FF4243ADBD043BFA" blockId="21.[189,1399,421,679]" box="[234,630,421,448]" pageId="21" pageNumber="22">
|
||
<emphasis id="B917EAF6FFEBFFD4FF4243ADBEBA3B85" bold="true" box="[234,456,421,448]" pageId="21" pageNumber="22">
|
||
<typeStatus id="54D88846FFEBFFD4FF4243ADBE153BFA" box="[234,359,422,448]" pageId="21" pageNumber="22" type="paratype">Paratypes</typeStatus>
|
||
revised
|
||
</emphasis>
|
||
(
|
||
<specimenCount id="9D65FD6DFFEBFFD4FE7143AEBD6F3BFA" box="[473,541,421,448]" pageId="21" pageNumber="22" type="male">6 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D65FD6DFFEBFFD4FD8243ADBD183BFA" box="[554,618,422,448]" pageId="21" pageNumber="22" type="female">2 ♀♀</specimenCount>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFEBFFD4FF1543C7BCF7389D" blockId="21.[189,1399,421,679]" pageId="21" pageNumber="22">
|
||
<materialsCitation id="3B0B3CB9FFEBFFD4FF1543C7BD643BDC" ID-GBIF-Occurrence="3093980449" box="[189,534,460,487]" country="Mexico" location="State of Mexico" pageId="21" pageNumber="22" specimenCount="1" stateProvince="State of Mexico">
|
||
<collectingCountry id="F3747674FFEBFFD4FF1543C7BE433BDC" box="[189,305,460,486]" name="Mexico" pageId="21" pageNumber="22">MEXICO</collectingCountry>
|
||
–
|
||
<emphasis id="B917EAF6FFEBFFD4FEF943C7BD643BDC" bold="true" box="[337,534,460,486]" pageId="21" pageNumber="22">
|
||
<collectingRegion id="49A7F806FFEBFFD4FEF943C7BD643BDC" box="[337,534,460,486]" pageId="21" pageNumber="22">State of Mexico</collectingRegion>
|
||
</emphasis>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0B3CB9FFEBFFD4FD9B43C7BB173837" ID-GBIF-Occurrence="3093980469" collectingDate="1976-07-10" collectionCode="IEXA" collectorName="W. D. Edmonds & P. Reyes & B. Kohlmann" country="Mexico" elevation="2200" location="Real de Arriba" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" stateProvince="State">
|
||
<specimenCount id="9D65FD6DFFEBFFD4FD9B43C7BD113BDD" box="[563,611,460,487]" pageId="21" pageNumber="22" type="male">1 ♂</specimenCount>
|
||
; “
|
||
<locationDeviation id="9A6E2FC6FFEBFFD4FDD643C7BCFC3BDC" box="[638,910,460,487]" location="Temascaltepec" pageId="21" pageNumber="22">
|
||
<quantity id="4C9B9B01FFEBFFD4FDD643C7BDC93BDC" box="[638,699,460,487]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="21" pageNumber="22" unit="km" value="5.0">5 km</quantity>
|
||
E Temascaltepec
|
||
</locationDeviation>
|
||
,
|
||
<location id="8EBC603FFFEBFFD4FC3343C7BB383BDC" LSID="urn:lsid:plazi:treatment:03CA87F2FFEAFFD4FD844748BCE73C8D:8EBC603FFFEBFFD4FC3343C7BB383BDC" box="[923,1098,460,486]" country="Mexico" name="Real de Arriba" pageId="21" pageNumber="22" stateProvince="State">Real de Arriba</location>
|
||
(
|
||
<quantity id="4C9B9B01FFEBFFD4FBF343C7BBC83BDD" box="[1115,1210,460,487]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="21" pageNumber="22" unit="m" value="2200.0">
|
||
<elevation id="004ED1D7FFEBFFD4FBF343C7BBC83BDD" box="[1115,1210,460,487]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="21" pageNumber="22" unit="m" value="2200.0">2200 m</elevation>
|
||
</quantity>
|
||
),
|
||
<date id="FFDD1024FFEBFFD4FB7943C7BA003BDC" box="[1233,1394,460,486]" pageId="21" pageNumber="22" value="1976-07-10">
|
||
<collectingDate id="EF99E9CCFFEBFFD4FB7943C7BA003BDC" box="[1233,1394,460,486]" pageId="21" pageNumber="22" value="1976-07-10">10–VII–1976</collectingDate>
|
||
</date>
|
||
, fungus, oak-pine forest,
|
||
<collectorName id="26965332FFEBFFD4FE7743F8BDED3836" box="[479,671,498,525]" pageId="21" pageNumber="22">W. D. Edmonds</collectorName>
|
||
,
|
||
<collectorName id="26965332FFEBFFD4FD0543F8BC673837" box="[685,789,499,525]" pageId="21" pageNumber="22">P. Reyes</collectorName>
|
||
,
|
||
<collectorName id="26965332FFEBFFD4FC8A43F8BCB33836" box="[802,961,498,525]" pageId="21" pageNumber="22">B. Kohlmann</collectorName>
|
||
cols.”;
|
||
<collectionCode id="ED72AE21FFEBFFD4FBB743F8BB173837" box="[1055,1125,499,525]" pageId="21" pageNumber="22">IEXA</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0B3CB9FFEBFFD4FBD743F8BD663809" ID-GBIF-Occurrence="3093980344" collectingDate="1976-07-10" collectionCode="TAMU" collectorName="W. D. Edmonds & P. Reyes & B. Kohlmann" country="Mexico" elevation="2200" location="Real de Arriba" pageId="21" pageNumber="22" specimenCount="1" specimenCount-female="1" stateProvince="State">
|
||
<specimenCount id="9D65FD6DFFEBFFD4FBD743F8BBDC3837" box="[1151,1198,499,525]" pageId="21" pageNumber="22" type="female">1 ♀</specimenCount>
|
||
; same collection data as for preceding;
|
||
<collectionCode id="ED72AE21FFEBFFD4FE684012BD663809" box="[448,532,537,563]" collectionName="USA, Texas, College Station, Texas A & M University" pageId="21" pageNumber="22">TAMU</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0B3CB9FFEBFFD4FD844012BBD63809" ID-GBIF-Occurrence="3093980398" box="[556,1188,537,563]" collectingDate="1976-07-10" collectionCode="VMC" collectorName="W. D. Edmonds & P. Reyes & B. Kohlmann" country="Mexico" elevation="2200" location="Real de Arriba" pageId="21" pageNumber="22" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="State">
|
||
<specimenCount id="9D65FD6DFFEBFFD4FD844012BD2B3809" box="[556,601,537,563]" pageId="21" pageNumber="22" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D65FD6DFFEBFFD4FDCE4012BDE03809" box="[614,658,537,563]" pageId="21" pageNumber="22" type="female">1 ♀</specimenCount>
|
||
; same collection data as for preceding;
|
||
<collectionCode id="ED72AE21FFEBFFD4FBC94012BBD63809" box="[1121,1188,537,563]" pageId="21" pageNumber="22">VMC</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0B3CB9FFEBFFD4FB144012BE1738BA" ID-GBIF-Occurrence="3093980504" collectedFrom="fungus in pine-oak forest" collectingDate="1976-07-11" collectionCode="TAMU" collectorName="W. D. Edmonds & P. Reyes & B. Kohlmann" country="Mexico" elevation="2360" location="8 km W Temascaltepec" pageId="21" pageNumber="22" specimenCount="2" specimenCount-male="2" stateProvince="State">
|
||
<specimenCount id="9D65FD6DFFEBFFD4FB144012BA733809" box="[1212,1281,537,563]" pageId="21" pageNumber="22" type="male">2 ♂♂</specimenCount>
|
||
; “
|
||
<locationDeviation id="9A6E2FC6FFEBFFD4FAB14012BE1E3863" pageId="21" pageNumber="22">
|
||
<quantity id="4C9B9B01FFEBFFD4FAB14012BA263809" box="[1305,1364,537,563]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="21" pageNumber="22" unit="km" value="8.0">8 km</quantity>
|
||
W Temascaltepec
|
||
</locationDeviation>
|
||
,
|
||
<quantity id="4C9B9B01FFEBFFD4FED04034BEA03860" box="[376,466,575,602]" metricMagnitude="3" metricUnit="m" metricValue="2.36" pageId="21" pageNumber="22" unit="m" value="2360.0">
|
||
<elevation id="004ED1D7FFEBFFD4FED04034BEA03860" box="[376,466,575,602]" metricMagnitude="3" metricUnit="m" metricValue="2.36" pageId="21" pageNumber="22" unit="m" value="2360.0">2360 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFDD1024FFEBFFD4FE764034BD123863" box="[478,608,575,601]" pageId="21" pageNumber="22" value="1976-07-11">
|
||
<collectingDate id="EF99E9CCFFEBFFD4FE764034BD123863" box="[478,608,575,601]" pageId="21" pageNumber="22" value="1976-07-11">11–VII–76</collectingDate>
|
||
</date>
|
||
,
|
||
<collectedFrom id="387F7519FFEBFFD4FDC44034BCE73863" box="[620,917,575,601]" pageId="21" pageNumber="22">fungus in pine-oak forest</collectedFrom>
|
||
,
|
||
<collectorName id="26965332FFEBFFD4FC37404BBB2E3863" box="[927,1116,575,602]" pageId="21" pageNumber="22">W. D. Edmonds</collectorName>
|
||
,
|
||
<collectorName id="26965332FFEBFFD4FBCF404BBBBC3860" box="[1127,1230,576,602]" pageId="21" pageNumber="22">P. Reyes</collectorName>
|
||
,
|
||
<collectorName id="26965332FFEBFFD4FB71404BBA053863" box="[1241,1399,575,602]" pageId="21" pageNumber="22">B. Kohlmann</collectorName>
|
||
cols.”;
|
||
<collectionCode id="ED72AE21FFEBFFD4FEB9406DBE1738BA" box="[273,357,614,640]" collectionName="USA, Texas, College Station, Texas A & M University" pageId="21" pageNumber="22">TAMU</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0B3CB9FFEBFFD4FE2B406DBCA638BA" ID-GBIF-Occurrence="3093980364" box="[387,980,614,640]" collectedFrom="fungus in pine-oak forest" collectingDate="1976-07-11" collectionCode="VMC" collectorName="W. D. Edmonds & P. Reyes" country="Mexico" elevation="2360" location="8 km W Temascaltepec" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" stateProvince="State">
|
||
<specimenCount id="9D65FD6DFFEBFFD4FE2B406DBEC638BA" box="[387,436,614,640]" pageId="21" pageNumber="22" type="male">1 ♂</specimenCount>
|
||
; same collection data as for preceding;
|
||
<collectionCode id="ED72AE21FFEBFFD4FC39406DBCA638BA" box="[913,980,614,640]" pageId="21" pageNumber="22">VMC</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0B3CB9FFEBFFD4FC59406DBCF3389D" ID-GBIF-Occurrence="3093980518" collectingDate="1932-07" collectionCode="TAMU" collectorName="Mexico D. F. & Hinton" country="Mexico" elevation="1920" location="Real de Arriba" pageId="21" pageNumber="22" specimenCode="BM 1939-583" specimenCount="1" specimenCount-male="1" stateProvince="State">
|
||
<specimenCount id="9D65FD6DFFEBFFD4FC59406DBB5038BA" box="[1009,1058,614,640]" pageId="21" pageNumber="22" type="male">1 ♂</specimenCount>
|
||
; “
|
||
<location id="8EBC603FFFEBFFD4FB93406DBB8238BA" LSID="urn:lsid:plazi:treatment:03CA87F2FFEAFFD4FD844748BCE73C8D:8EBC603FFFEBFFD4FB93406DBB8238BA" box="[1083,1264,614,640]" country="Mexico" name="Real de Arriba" pageId="21" pageNumber="22" stateProvince="State">Real de Arriba</location>
|
||
,
|
||
<date id="FFDD1024FFEBFFD4FB55406DBA0038BA" box="[1277,1394,614,640]" bridgedPair="-" pageId="21" pageNumber="22" value="1932-07">
|
||
<collectingDate id="EF99E9CCFFEBFFD4FB55406DBA0038BA" box="[1277,1394,614,640]" bridgedPair="-" pageId="21" pageNumber="22" value="1932-07">VII–1932</collectingDate>
|
||
</date>
|
||
,
|
||
<quantity id="4C9B9B01FFEBFFD4FF154087BE66389C" box="[189,276,652,678]" metricMagnitude="3" metricUnit="m" metricValue="1.92024" pageId="21" pageNumber="22" unit="ft" value="6300.0">
|
||
<elevation id="004ED1D7FFEBFFD4FF154087BE66389C" box="[189,276,652,678]" metricMagnitude="3" metricUnit="m" metricValue="1.92024" pageId="21" pageNumber="22" unit="ft" value="6300.0">6300 ft</elevation>
|
||
</quantity>
|
||
,
|
||
<collectorName id="26965332FFEBFFD4FE884087BECE389D" box="[288,444,652,679]" pageId="21" pageNumber="22">México D. F.</collectorName>
|
||
,
|
||
<collectorName id="26965332FFEBFFD4FE604087BD6B389C" box="[456,537,652,678]" pageId="21" pageNumber="22">Hinton</collectorName>
|
||
coll.,
|
||
<specimenCode id="DBC59E9FFFEBFFD4FDCB4086BC61389D" box="[611,787,652,679]" pageId="21" pageNumber="22">BM 1939–583</specimenCode>
|
||
”;
|
||
<collectionCode id="ED72AE21FFEBFFD4FC814086BCF3389D" box="[809,897,653,679]" collectionName="USA, Texas, College Station, Texas A & M University" pageId="21" pageNumber="22">TAMU</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFEBFFD4FF1540DABE2F38D1" blockId="21.[189,349,721,747]" box="[189,349,721,747]" pageId="21" pageNumber="22">
|
||
<heading id="D0948188FFEBFFD4FF1540DABE2F38D1" bold="true" box="[189,349,721,747]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
|
||
<emphasis id="B917EAF6FFEBFFD4FF1540DABE2F38D1" bold="true" box="[189,349,721,747]" pageId="21" pageNumber="22">Type locality</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFEBFFD4FF154108BDD13927" blockId="21.[189,675,771,797]" box="[189,675,771,797]" pageId="21" pageNumber="22">
|
||
<collectingCountry id="F3747674FFEBFFD4FF154108BE683927" box="[189,282,771,797]" name="Mexico" pageId="21" pageNumber="22">Mexico</collectingCountry>
|
||
, State of
|
||
<collectingCountry id="F3747674FFEBFFD4FE2F4108BE963927" box="[391,484,771,797]" name="Mexico" pageId="21" pageNumber="22">Mexico</collectingCountry>
|
||
, Temascaltepec.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C379656FFFEBFFD4FF15414CBB3939AE" box="[189,1099,839,916]" pageId="21" pageNumber="22" type="distribution">
|
||
<paragraph id="8BDC36E4FFEBFFD4FF15414CBE24395B" blockId="21.[189,342,839,865]" box="[189,342,839,865]" pageId="21" pageNumber="22">
|
||
<heading id="D0948188FFEBFFD4FF15414CBE24395B" bold="true" box="[189,342,839,865]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
|
||
<emphasis id="B917EAF6FFEBFFD4FF15414CBE24395B" bold="true" box="[189,342,839,865]" pageId="21" pageNumber="22">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFEBFFD4FF154172BB3939AE" blockId="21.[189,1099,889,916]" box="[189,1099,889,916]" pageId="21" pageNumber="22">
|
||
Central Trans-Mexican Volcanic Belt, State of
|
||
<collectingRegion id="49A7F806FFEBFFD4FD4A4172BC4E39A9" box="[738,828,889,915]" country="Mexico" name="Mexico" pageId="21" pageNumber="22">Mexico</collectingRegion>
|
||
and
|
||
<collectingRegion id="49A7F806FFEBFFD4FCDC4172BCA439A9" box="[884,982,889,915]" country="Mexico" name="Morelos" pageId="21" pageNumber="22">Morelos</collectingRegion>
|
||
(
|
||
<figureCitation id="13582A61FFEBFFD4FC4E4172BB4F39AE" box="[998,1085,889,916]" captionStart="Fig" captionStartId="50.[189,232,1890,1916]" captionTargetBox="[193,1392,927,1854]" captionTargetId="figure-235@50.[189,1398,924,1857]" captionTargetPageId="50" captionText="Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017;P. edmondsi Moctezuma, Deloya & Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 by Jose Luis Sánchez-Huerta." figureDoi="http://doi.org/10.5281/zenodo.4836796" httpUri="https://zenodo.org/record/4836796/files/figure.png" pageId="21" pageNumber="22">Fig. 15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C379656FFFEBFFD4FF1541B5BCE73C8D" pageId="21" pageNumber="22" type="discussion">
|
||
<paragraph id="8BDC36E4FFEBFFD4FF1541B5BE5D39E2" blockId="21.[189,303,958,984]" box="[189,303,958,984]" pageId="21" pageNumber="22">
|
||
<heading id="D0948188FFEBFFD4FF1541B5BE5D39E2" bold="true" box="[189,303,958,984]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
|
||
<emphasis id="B917EAF6FFEBFFD4FF1541B5BE5D39E2" bold="true" box="[189,303,958,984]" pageId="21" pageNumber="22">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFEBFFD4FF1541FABB9B3F5E" blockId="21.[188,1401,1008,1381]" pageId="21" pageNumber="22">
|
||
Mean length
|
||
<quantity id="4C9B9B01FFEBFFD4FEF541FBBEBB3E31" box="[349,457,1008,1035]" metricMagnitude="-2" metricUnit="m" metricValue="1.7399999999999998" pageId="21" pageNumber="22" unit="mm" value="17.4">17.4 mm</quantity>
|
||
(
|
||
<quantity id="4C9B9B01FFEBFFD4FE7341FBBDFD3E31" box="[475,655,1008,1035]" metricMagnitude="-2" metricUnit="m" metricValue="1.6649999999999998" metricValueMax="1.9899999999999998" metricValueMin="1.34" pageId="21" pageNumber="22" unit="mm" value="16.65" valueMax="19.9" valueMin="13.4">13.4–19.9 mm</quantity>
|
||
). The specimens from
|
||
<collectingRegion id="49A7F806FFEBFFD4FC0B41FBBB773E30" box="[931,1029,1008,1034]" country="Mexico" name="Morelos" pageId="21" pageNumber="22">Morelos</collectingRegion>
|
||
(
|
||
<bibRefCitation id="EFF24B15FFEBFFD4FBBF41FBBB893E30" author="Deloya C. & Burgos A. & Blackaller J. & Lobo J. M." box="[1047,1275,1008,1035]" pageId="21" pageNumber="22" pagination="15 - 55" refId="ref37208" refString="Deloya C., Burgos A., Blackaller J. & Lobo J. M. 1993. Los coleopteros lamelicornios de Cuernavaca, Morelos, Mexico (Passalidae, Trogidae, Scarabaeidae y Melolonthidae). Boletin de la Sociedad veracruzana de Zoologia 3: 15 - 55." type="journal article" year="1993">
|
||
Deloya
|
||
<emphasis id="B917EAF6FFEBFFD4FBD041FABBDC3E30" box="[1144,1198,1008,1034]" italics="true" pageId="21" pageNumber="22">et al</emphasis>
|
||
. 1993
|
||
</bibRefCitation>
|
||
) were not studied by us. This species was erroneously reported from
|
||
<collectingRegion id="49A7F806FFEBFFD4FCCD461DBC8A3E0A" box="[869,1016,1046,1072]" country="Mexico" name="Distrito Federal" pageId="21" pageNumber="22">Mexico City</collectingRegion>
|
||
(
|
||
<bibRefCitation id="EFF24B15FFEBFFD4FBAE461DBBC73E0B" author="Arnaud P." box="[1030,1205,1046,1073]" pageId="21" pageNumber="22" refId="ref35859" refString="Arnaud P. 2002 b. Phanaeini. Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus. Hillside Books, Canterbury." type="book" year="2002">Arnaud 2002b</bibRefCitation>
|
||
). This mistake is attributed to
|
||
<bibRefCitation id="EFF24B15FFEBFFD4FEF84636BE8B3E62" author="Hinton H. E." box="[336,505,1085,1112]" pageId="21" pageNumber="22" pagination="129 - 130" refId="ref39055" refString="Hinton H. E. 1935. Anotaciones acerca de las costumbres micetofagicas de las especies de Phanaeus. Anales del Instituto de Biologia, UNAM (Serie Zoologia) 6: 129 - 130." type="journal article" year="1935">Hinton (1935)</bibRefCitation>
|
||
, who recorded it from Real de Arriba,
|
||
<collectingCountry id="F3747674FFEBFFD4FC1F4636BB633E6D" box="[951,1041,1085,1111]" name="Mexico" pageId="21" pageNumber="22">Mexico</collectingCountry>
|
||
D.F. Real de Arriba is actually located in the State of
|
||
<collectingCountry id="F3747674FFEBFFD4FE614668BD543E47" box="[457,550,1123,1149]" name="Mexico" pageId="21" pageNumber="22">Mexico</collectingCountry>
|
||
. Despite the fact that Moctezuma
|
||
<emphasis id="B917EAF6FFEBFFD4FC16466FBC833E47" box="[958,1009,1123,1149]" italics="true" pageId="21" pageNumber="22">et al</emphasis>
|
||
. (2017) split
|
||
<taxonomicName id="4C634D67FFEBFFD4FB3C466FBA373E47" authorityName="Edmonds" authorityYear="1979" box="[1172,1349,1123,1149]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="halffterorum">
|
||
<emphasis id="B917EAF6FFEBFFD4FB3C466FBA373E47" box="[1172,1349,1123,1149]" italics="true" pageId="21" pageNumber="22">P. halffterorum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C634D67FFEBFFD4FF164680BE2B3E9E" authorityName="Moctezuma, Sanchez-Huerta & Halffter" authorityYear="2017" box="[190,345,1162,1188]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="bravoensis">
|
||
<emphasis id="B917EAF6FFEBFFD4FF164680BE2B3E9E" box="[190,345,1162,1188]" italics="true" pageId="21" pageNumber="22">P. bravoensis</emphasis>
|
||
</taxonomicName>
|
||
, the colouration pattern of
|
||
<taxonomicName id="4C634D67FFEBFFD4FD324680BC383E9E" authorityName="Edmonds" authorityYear="1979" box="[666,842,1162,1188]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="halffterorum">
|
||
<emphasis id="B917EAF6FFEBFFD4FD324680BC383E9E" box="[666,842,1162,1188]" italics="true" pageId="21" pageNumber="22">P. halffterorum</emphasis>
|
||
</taxonomicName>
|
||
remains as indicated by
|
||
<bibRefCitation id="EFF24B15FFEBFFD4FBC74681BA443E9E" author="Edmonds W. D." box="[1135,1334,1162,1188]" pageId="21" pageNumber="22" pagination="99 - 105" refId="ref37459" refString="Edmonds W. D. 1979. A new species of Phanaeus from Mexico (Coleoptera: Scarabaeidae). Pan-Pacific Entomologist 55: 99 - 105." type="journal article" year="1979">Edmonds (1979)</bibRefCitation>
|
||
, with bright metallic green or dark metallic blue specimens. In a review of the immature dung beetles of
|
||
<taxonomicName id="4C634D67FFEBFFD4FF1546DCBDC43ECB" authority="(Edmonds & Halffter 1978)" baseAuthorityName="Edmonds & Halffter" baseAuthorityYear="1978" box="[189,694,1239,1265]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subFamily" subFamily="Scarabaeinae">
|
||
Scarabaeinae (
|
||
<bibRefCitation id="EFF24B15FFEBFFD4FEC246DCBDDF3ECB" author="Edmonds W. D. & Halffter G." box="[362,685,1239,1265]" pageId="21" pageNumber="22" pagination="307 - 331" refId="ref37704" refString="Edmonds W. D. & Halffter G. 1978. Taxonomic review of immature dung beetles of the dung beetles of the subfamily Scarabaeinae (Coleoptera: Scarabaeidae). Systematic Entomology 3: 307 - 331. https: // doi. org / 10.1111 / j. 1365 - 3113.1978. tb 00002. x" type="journal article" year="1978">Edmonds & Halffter 1978</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, the name
|
||
<taxonomicName id="4C634D67FFEBFFD4FC9746D3BBC83ECB" authority="Edmonds, 1978" authorityName="Edmonds" authorityYear="1978" box="[831,1210,1239,1265]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="halffterorum">
|
||
<emphasis id="B917EAF6FFEBFFD4FC9746D3BC813ECB" box="[831,1011,1239,1265]" italics="true" pageId="21" pageNumber="22">P. halffterorum</emphasis>
|
||
<bibRefCitation id="EFF24B15FFEBFFD4FC5646DCBBC83ECB" author="Edmonds W. D. & Halffter G." box="[1022,1210,1239,1265]" pageId="21" pageNumber="22" pagination="307 - 331" refId="ref37704" refString="Edmonds W. D. & Halffter G. 1978. Taxonomic review of immature dung beetles of the dung beetles of the subfamily Scarabaeinae (Coleoptera: Scarabaeidae). Systematic Entomology 3: 307 - 331. https: // doi. org / 10.1111 / j. 1365 - 3113.1978. tb 00002. x" type="journal article" year="1978">Edmonds, 1978</bibRefCitation>
|
||
</taxonomicName>
|
||
was published. Nevertheless, this may be considered as a nomen nudum under Article 13 of the Code (
|
||
<bibRefCitation id="EFF24B15FFEBFFD4FB7346F5BA183F22" author="ICZN" box="[1243,1386,1278,1304]" pageId="21" pageNumber="22" refId="ref39375" refString="ICZN. 1999. International Code of Zoological Nomenclature. Fourth Edition. International Commission on Zoological Nomenclature, the International Trust for Zoological Nomenclature, London." type="book" year="1999">ICZN 1999</bibRefCitation>
|
||
). Consequently, the same name was available later for the same or a different concept under Arts 21, 50; while
|
||
<taxonomicName id="4C634D67FFEBFFD4FEAD4740BD1C3F5F" authority="Edmonds 1979" authorityName="Edmonds" authorityYear="1979" box="[261,622,1354,1381]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="halffterorum">
|
||
<emphasis id="B917EAF6FFEBFFD4FEAD4740BEC73F5E" box="[261,437,1354,1380]" italics="true" pageId="21" pageNumber="22">P. halffterorum</emphasis>
|
||
<bibRefCitation id="EFF24B15FFEBFFD4FE154741BD1C3F5F" author="Edmonds W. D." box="[445,622,1354,1381]" pageId="21" pageNumber="22" pagination="99 - 105" refId="ref37459" refString="Edmonds W. D. 1979. A new species of Phanaeus from Mexico (Coleoptera: Scarabaeidae). Pan-Pacific Entomologist 55: 99 - 105." type="journal article" year="1979">Edmonds 1979</bibRefCitation>
|
||
</taxonomicName>
|
||
must be considered the available authorship and date.
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFEBFFD4FF15479BBCE73C8D" blockId="21.[189,1400,1423,1719]" pageId="21" pageNumber="22">
|
||
To the original description of the male we add that the right lobe of the endophallite copulatrix is more developed than the left lobe; the right lobe ois btusely triangular; the left lobe is obtusely rectangular; the central ridge lis ess developed than the central column (
|
||
<figureCitation id="13582A61FFEBFFD4FC2A47D7BCA93FCC" box="[898,987,1500,1526]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta & Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma & Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="21" pageNumber="22">Fig. 1F</figureCitation>
|
||
). For the female can be added that the trituberculate cephalic carina has conical, nearly aligned tubercles; the middle tubercle is more raised than the lateral tubercles; the pronotal process is trituberculate, with a posterior concavity; all the tubercles are rounded; the middle tubercle is slightly more developed and projected frontally than the lateral tubercles; the pronotal midline is distinctly impressed, with superficially impressed punctures; the pronotal surface is smooth, with almost effaced punctures.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |