263 lines
43 KiB
XML
263 lines
43 KiB
XML
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<mods:title id="33EB7B1AFBD2AB1EA790DFE2B5CBC12E">Revision of Madagascan Cleridae (Coleoptera: Cleroidea). Part I. - The genus Biflabelotillus Pic, 1949 (Tillinae)</mods:title>
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<subSubSection id="C321B684FFFDFFF913E9FE3EFDDFABA6" box="[151,530,474,501]" pageId="1" pageNumber="385" type="nomenclature">
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<paragraph id="8B84E50FFFFDFFF913E9FE3EFDDFABA6" blockId="1.[151,530,474,501]" box="[151,530,474,501]" pageId="1" pageNumber="385">
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<heading id="D0CC5263FFFDFFF913E9FE3EFDDFABA6" bold="true" box="[151,530,474,501]" fontSize="11" level="1" pageId="1" pageNumber="385" reason="1">
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<emphasis id="B94F391DFFFDFFF913E9FE3EFDDFABA6" bold="true" box="[151,530,474,501]" pageId="1" pageNumber="385">
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Genus
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<taxonomicName id="4C3B9E8CFFFDFFF91390FE3EFDDFABA6" authority="Pic, 1949" authorityName="Pic" authorityYear="1949" box="[238,530,474,501]" class="Insecta" family="Cleridae" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
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<emphasis id="B94F391DFFFDFFF91390FE3EFE54ABA7" bold="true" box="[238,409,474,501]" italics="true" pageId="1" pageNumber="385">Biflabelotillus</emphasis>
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<bibRefCitation id="EFAA98FEFFFDFFF912DFFE3EFDDFABA6" author="Pic, M." box="[417,530,474,500]" pageId="1" pageNumber="385" pagination="1 - 16" refId="ref5365" refString="Pic, M. (1949) Clerides de Madagascar. Diversites Entomologiques V., 1949, 1 - 16." type="journal article" year="1949">Pic, 1949</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C321B684FFFDFFF913E9FDFEFBC5A860" box="[151,1032,538,562]" pageId="1" pageNumber="385" type="reference_group">
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<paragraph id="8B84E50FFFFDFFF913E9FDFEFBC5A860" blockId="1.[151,1436,538,685]" box="[151,1032,538,562]" pageId="1" pageNumber="385">
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<treatmentCitationGroup id="AB2BC221FFFDFFF913E9FDFEFBC5A860" box="[151,1032,538,562]" pageId="1" pageNumber="385">
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<treatmentCitation id="0A9AC31EFFFDFFF913E9FDFEFE65A860" author="Pic, M." box="[151,424,538,562]" page="3" pageId="1" pageNumber="385" year="1949">
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<taxonomicName id="4C3B9E8CFFFDFFF913E9FDFEFE65A860" authority="Pic, 1949: 3" authorityName="Pic" authorityPageNumber="3" authorityYear="1949" box="[151,424,538,562]" class="Insecta" family="Cleridae" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
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<emphasis id="B94F391DFFFDFFF913E9FDFEFEEAA860" box="[151,295,538,562]" italics="true" pageId="1" pageNumber="385">Biflabelotillus</emphasis>
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<bibRefCitation id="EFAA98FEFFFDFFF91250FDFEFE65A860" author="Pic, M." box="[302,424,538,562]" pageId="1" pageNumber="385" pagination="1 - 16" refId="ref5365" refString="Pic, M. (1949) Clerides de Madagascar. Diversites Entomologiques V., 1949, 1 - 16." type="journal article" year="1949">Pic, 1949: 3</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(original description). –
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<treatmentCitation id="0A9AC31EFFFDFFF911D8FDFEFBC9A860" author="Edwards, M. A. & Hopwood, A. T." box="[678,1028,538,562]" page="36" pageId="1" pageNumber="385" year="1966">
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<bibRefCitation id="EFAA98FEFFFDFFF911D8FDFEFC13A860" author="Edwards, M. A. & Hopwood, A. T." box="[678,990,538,562]" pageId="1" pageNumber="385" refId="ref5094" refString="Edwards, M. A. & Hopwood, A. T. (1966) Nomenclator Zoologicus. Vol. VI. 1946 - 1955. Zoological Society of London, London, viii + 329 pp." type="book" year="1966">EDWARDS & HOPWOOD (1966)</bibRefCitation>
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: 36
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C321B684FFFDFFF91770FDFEFDE8A802" pageId="1" pageNumber="385" type="type_taxon">
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<paragraph id="8B84E50FFFFDFFF91770FDFEFDE8A802" blockId="1.[151,1436,538,685]" pageId="1" pageNumber="385">
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<typeStatus id="54805BADFFFDFFF91770FDFEFB8DA860" box="[1038,1088,538,562]" pageId="1" pageNumber="385">Type</typeStatus>
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species:
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<taxonomicName id="4C3B9E8CFFFDFFF917E0FDFEFA51A860" authorityYear="1949" box="[1182,1436,538,562]" pageId="1" pageNumber="385">
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<emphasis id="B94F391DFFFDFFF917E0FDFEFA51A860" box="[1182,1436,538,562]" italics="true" pageId="1" pageNumber="385">Biflabelotillus foveolatus</emphasis>
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</taxonomicName>
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<bibRefCitation id="EFAA98FEFFFDFFF913B9FDDCFE8CA803" author="Pic, M." box="[199,321,568,593]" pageId="1" pageNumber="385" pagination="1 - 16" refId="ref5365" refString="Pic, M. (1949) Clerides de Madagascar. Diversites Entomologiques V., 1949, 1 - 16." type="journal article" year="1949">Pic, 1949: 3</bibRefCitation>
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(original description).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C321B684FFFDFFF913E9FDB3FB90A8FF" pageId="1" pageNumber="385" type="reference_group">
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||
<paragraph id="8B84E50FFFFDFFF913E9FDB3FC1EA83D" blockId="1.[151,1436,538,685]" box="[151,979,599,623]" pageId="1" pageNumber="385">
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<emphasis id="B94F391DFFFDFFF913E9FDB3FEEAA83D" box="[151,295,599,623]" italics="true" pageId="1" pageNumber="385">Biflabellotilus</emphasis>
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[sic!] –
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<bibRefCitation id="EFAA98FEFFFDFFF91202FDB3FE27A83D" author="Pic, M." box="[380,490,599,623]" pageId="1" pageNumber="385" pagination="1 - 16" refId="ref5365" refString="Pic, M. (1949) Clerides de Madagascar. Diversites Entomologiques V., 1949, 1 - 16." type="journal article" year="1949">Pic (1949)</bibRefCitation>
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||
: 3 (used as
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<emphasis id="B94F391DFFFDFFF91121FDB3FD22A83D" box="[607,751,599,623]" italics="true" pageId="1" pageNumber="385">Biflabellotilus</emphasis>
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||
<emphasis id="B94F391DFFFDFFF9118BFDB3FC90A83D" box="[757,861,599,623]" italics="true" pageId="1" pageNumber="385">foveolatus</emphasis>
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||
<bibRefCitation id="EFAA98FEFFFDFFF9101DFDB3FC0AA83D" author="Pic, M." box="[867,967,599,623]" pageId="1" pageNumber="385" pagination="1 - 16" refId="ref5365" refString="Pic, M. (1949) Clerides de Madagascar. Diversites Entomologiques V., 1949, 1 - 16." type="journal article" year="1949">Pic, 1949</bibRefCitation>
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||
).
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</paragraph>
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||
<paragraph id="8B84E50FFFFDFFF913E9FD92FB90A8FF" blockId="1.[151,1436,538,685]" pageId="1" pageNumber="385">
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||
<taxonomicName id="4C3B9E8CFFFDFFF913E9FD92FEE3A8DC" authorityName="Pic" authorityYear="1949" box="[151,302,630,654]" class="Insecta" family="Cleridae" genus="Biflabellotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
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<emphasis id="B94F391DFFFDFFF913E9FD92FEE3A8DC" box="[151,302,630,654]" italics="true" pageId="1" pageNumber="385">Biflabellotillus</emphasis>
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</taxonomicName>
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||
[sic!] –
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||
<bibRefCitation id="EFAA98FEFFFDFFF912F7FD92FD9EA8DC" author="Corporaal, J. B." box="[393,595,630,654]" pageId="1" pageNumber="385" refId="ref5062" refString="Corporaal, J. B. (1950) Cleridae. Coleopterorum Catalogus. Supplementa Pars 23. Uitgeverij. Dr. W. Junk, ' s-Gravenhage, 373 pp." type="book" year="1950">CORPORAAL (1950)</bibRefCitation>
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: 17 (catalogue);
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<bibRefCitation id="EFAA98FEFFFDFFF9107FFD92FC7CA8DC" author="Winkler, J. R." box="[769,945,630,654]" pageId="1" pageNumber="385" refId="ref5415" refString="Winkler, J. R. (1961) Die Buntkafer (Cleridae). A. Ziemsen Verlag, Wittenberg, Lutherstadt, 108 pp." type="book" year="1961">WINKLER (1961)</bibRefCitation>
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: 20, 32 (list of genera);
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<bibRefCitation id="EFAA98FEFFFDFFF917CEFD92FA82A8DC" author="Menier, J. J." box="[1200,1359,630,654]" pageId="1" pageNumber="385" pagination="75 - 77" refId="ref5316" refString="Menier, J. J. (1979) Une espece malgache inedite du genere Biflabellotillus: B. peyrierasi, n. sp. (Col. Cleridae). Revue franc ¸ aise d'Entomologie, New Series, 1 (8), 75 - 77." type="journal article" year="1979">MENIER (1979)</bibRefCitation>
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: 75, 76 (redefinition, key to species);
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<bibRefCitation id="EFAA98FEFFFDFFF91288FD71FD7BA8FF" author="Mawdsley, J." box="[502,694,661,685]" pageId="1" pageNumber="385" pagination="19 - 29" refId="ref5261" refString="Mawdsley, J. (1993) A key to the species of the genus Myrmecomaea Fairmaire (Coleoptera: Cleridae) with checklist of species, notes on mimicry, and discussion of endemism in Madagascan Cleridae. Y. E. S. Quarterly, 10 (4), 19 - 29" type="journal article" year="1993">MAWDSLEY (1993)</bibRefCitation>
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: 26 (list of endemic Madagascan genera).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C321B684FFFDFFF913E9FD3FFE67A96F" pageId="1" pageNumber="385" type="multiple">
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||
<paragraph id="8B84E50FFFFDFFF913E9FD3FFE67A96F" blockId="1.[151,1437,731,1981]" pageId="1" pageNumber="385">
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<emphasis id="B94F391DFFFDFFF913E9FD3FFF1BA8A7" bold="true" box="[151,214,731,757]" pageId="1" pageNumber="385">Note.</emphasis>
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<bibRefCitation id="EFAA98FEFFFDFFF913A3FD3FFE9EA8A7" author="Pic, M." box="[221,339,731,757]" pageId="1" pageNumber="385" pagination="1 - 16" refId="ref5365" refString="Pic, M. (1949) Clerides de Madagascar. Diversites Entomologiques V., 1949, 1 - 16." type="journal article" year="1949">Pic (1949)</bibRefCitation>
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used two different spellings in his original paper,
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||
<taxonomicName id="4C3B9E8CFFFDFFF910FBFD3FFBE9A8A7" authorityName="Pic" authorityYear="1949" box="[901,1060,731,757]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
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<emphasis id="B94F391DFFFDFFF910FBFD3FFBE9A8A7" box="[901,1060,731,757]" italics="true" pageId="1" pageNumber="385">Biflabelotillus</emphasis>
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</taxonomicName>
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and
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<emphasis id="B94F391DFFFDFFF91724FD3FFB34A8A7" box="[1114,1273,731,757]" italics="true" pageId="1" pageNumber="385">Biflabellotilus</emphasis>
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(pp. 3). As far as we know, no one has acted as First Reviser (
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<bibRefCitation id="EFAA98FEFFFDFFF911DDFD1BFCE5A94B" author="ICZN" box="[675,808,767,793]" pageId="1" pageNumber="385" refId="ref5225" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. Adopted by the International Union of Biological Sciences. International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
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||
, Article 24.2.3) and
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<emphasis id="B94F391DFFFDFFF91772FD1BFB62A94B" bold="true" box="[1036,1199,767,793]" pageId="1" pageNumber="385">we here select</emphasis>
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<taxonomicName id="4C3B9E8CFFFDFFF917C8FD1BFA98A94B" authorityName="Pic" authorityYear="1949" box="[1206,1365,767,793]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
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<emphasis id="B94F391DFFFDFFF917C8FD1BFA98A94B" box="[1206,1365,767,793]" italics="true" pageId="1" pageNumber="385">Biflabelotillus</emphasis>
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</taxonomicName>
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as the correct original spelling.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C321B684FFFDFFF913B9FCA2FC10AE9B" pageId="1" pageNumber="385" type="diagnosis">
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||
<paragraph id="8B84E50FFFFDFFF913B9FCA2FC10AE9B" blockId="1.[151,1437,731,1981]" pageId="1" pageNumber="385">
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||
<emphasis id="B94F391DFFFDFFF913B9FCA2FE8FA932" bold="true" box="[199,322,838,864]" pageId="1" pageNumber="385">Diagnosis.</emphasis>
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Although a review of Madagascan tilline genera is needed to determine the generic status for most of the genera (
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<bibRefCitation id="EFAA98FEFFFDFFF91247FC8FFE02A9D7" author="Menier, J. J." box="[313,463,875,901]" pageId="1" pageNumber="385" pagination="75 - 77" refId="ref5316" refString="Menier, J. J. (1979) Une espece malgache inedite du genere Biflabellotillus: B. peyrierasi, n. sp. (Col. Cleridae). Revue franc ¸ aise d'Entomologie, New Series, 1 (8), 75 - 77." type="journal article" year="1979">Menier 1979</bibRefCitation>
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||
, Gerstmeier & Weis 2009),
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<taxonomicName id="4C3B9E8CFFFDFFF91074FC8FFC64A9D7" authorityName="Pic" authorityYear="1949" box="[778,937,875,901]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
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<emphasis id="B94F391DFFFDFFF91074FC8FFC64A9D7" box="[778,937,875,901]" italics="true" pageId="1" pageNumber="385">Biflabelotillus</emphasis>
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||
</taxonomicName>
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seems to be a relatively well defined genus. According to
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<bibRefCitation id="EFAA98FEFFFDFFF91251FC6BFE1FA9FB" author="Menier, J. J." box="[303,466,911,937]" pageId="1" pageNumber="385" pagination="75 - 77" refId="ref5316" refString="Menier, J. J. (1979) Une espece malgache inedite du genere Biflabellotillus: B. peyrierasi, n. sp. (Col. Cleridae). Revue franc ¸ aise d'Entomologie, New Series, 1 (8), 75 - 77." type="journal article" year="1979">Menier (1979)</bibRefCitation>
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, this genus belongs to the group of several genera (besides
|
||
<taxonomicName id="4C3B9E8CFFFDFFF91720FC6BFB30A9FB" authorityName="Pic" authorityYear="1949" box="[1118,1277,911,937]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94F391DFFFDFFF91720FC6BFB30A9FB" box="[1118,1277,911,937]" italics="true" pageId="1" pageNumber="385">Biflabelotillus</emphasis>
|
||
</taxonomicName>
|
||
e.g.
|
||
<taxonomicName id="4C3B9E8CFFFDFFF9164EFC6BFE81A99F" authority="Fairmaire, 1903" authorityName="Fairmaire" authorityYear="1903" class="Insecta" family="Cleridae" genus="Archalius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94F391DFFFDFFF9164EFC6BFA51A9FB" box="[1328,1436,911,937]" italics="true" pageId="1" pageNumber="385">Archalius</emphasis>
|
||
Fairmaire, 1903
|
||
</taxonomicName>
|
||
) within Madagascan
|
||
<taxonomicName id="4C3B9E8CFFFDFFF9113FFC57FD6EA99F" authorityName="Latreille" authorityYear="1802" box="[577,675,947,973]" class="Insecta" family="Cleridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="family">Cleridae</taxonomicName>
|
||
, whose elytral surface is covered with bituberculate foveae - a pair of tubercles situated opposite each other on top of a large round depression (see also
|
||
<figureCitation id="1300F98AFFFDFFF91719FC33FB72A9A3" box="[1127,1215,983,1009]" captionStart="FIGURE 13–20" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 13–20. Biflabelotillus foveolatus Pic, 1949—13–15 (lectotype), 16–20 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="1" pageNumber="385">Figs 20</figureCitation>
|
||
,
|
||
<figureCitation id="1300F98AFFFDFFF917B2FC33FB27A9A3" box="[1228,1258,983,1009]" captionStart="FIGURE 21–27" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[225,1371,187,1919]" captionTargetId="figure-17@6.[215,1371,181,1919]" captionTargetPageId="6" captionText="FIGURE 21–27. Biflabelotillus peyrierasi Menier, 1979—21–23 (holotype), 24–27 (non-type, male). Fig. 21—labels of the holotype; 21, 24—habitus, dorsal view; 23, 25—habitus, lateral view; 26—scutellum; 27—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915941" httpUri="https://zenodo.org/record/13915941/files/figure.png" pageId="1" pageNumber="385">27</figureCitation>
|
||
,
|
||
<figureCitation id="1300F98AFFFDFFF91786FC33FADBA9A3" box="[1272,1302,983,1009]" captionStart="FIGURE 28–32" captionStartId="7.[151,250,1586,1610]" captionTargetBox="[170,1418,184,1562]" captionTargetId="figure-26@7.[170,1418,181,1562]" captionTargetPageId="7" captionText="FIGURE 28–32. Biflabelotillus peyrierasi Menier, 1979, female. Fig. 28—habitus, dorsal view; 29—habitus, ventral view; 30—antenna (red arrow indicates last antennomere); 31—scutellum; 32—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915943" httpUri="https://zenodo.org/record/13915943/files/figure.png" pageId="1" pageNumber="385">32</figureCitation>
|
||
,
|
||
<figureCitation id="1300F98AFFFDFFF9165DFC33FA8CA9A3" box="[1315,1345,983,1009]" captionStart="FIGURE 33–37" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 33–37. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="1" pageNumber="385">37</figureCitation>
|
||
, 41). It, however, can reliably be distinguished by biflabeloid antennae, with two flabelae situated at the beginning of each antennomere III–X. Moreover, the flabelae have the same diameter along their whole length (see also
|
||
<figureCitation id="1300F98AFFFDFFF91673FBFBFABDAE6B" box="[1293,1392,1055,1081]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="1" pageNumber="385">Figs 1–3</figureCitation>
|
||
). In other Madagascan genera which possess flabeloid or biflabeloid antennae, the flabelae are situated in the middle or at the end of antennomeres III–X and their diameter is not equal along their length but usually gradually widened toward their apex. Moreover, in some genera (e.g. in
|
||
<taxonomicName id="4C3B9E8CFFFDFFF91186FB6FFB8CAEF7" authority="Fairmaire, 1885" authorityName="Fairmaire" authorityYear="1885" box="[760,1089,1162,1189]" class="Insecta" family="Cleridae" genus="Diplocladus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94F391DFFFDFFF91186FB6FFC4CAEF7" box="[760,897,1163,1189]" italics="true" pageId="1" pageNumber="385">Diplocladus</emphasis>
|
||
Fairmaire, 1885
|
||
</taxonomicName>
|
||
; Gerstmeier & Weis 2009) the base of flabelae is so close that it seems to be in fact one bifurcate flabella.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C321B684FFFDFFFA13B9FB36FB0EAAAB" lastPageId="2" lastPageNumber="386" pageId="1" pageNumber="385" type="description">
|
||
<paragraph id="8B84E50FFFFDFFF913B9FB36FF07AF43" blockId="1.[151,1437,731,1981]" pageId="1" pageNumber="385">
|
||
<emphasis id="B94F391DFFFDFFF913B9FB36FEB8AEBE" bold="true" box="[199,373,1234,1260]" pageId="1" pageNumber="385">Redescription.</emphasis>
|
||
Body cylindrical, shiny, sparsely pubescent. Body length
|
||
<quantity id="4CC348EAFFFDFFF9176DFB36FB61AEBF" box="[1043,1196,1234,1261]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" metricValueMax="11.9" metricValueMin="7.5" pageId="1" pageNumber="385" unit="mm" value="9.7" valueMax="11.9" valueMin="7.5">7.5–11.9 mm</quantity>
|
||
. Body width
|
||
<quantity id="4CC348EAFFFDFFF91636FB37FF0BAF43" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.6" metricValueMin="2.1" pageId="1" pageNumber="385" unit="mm" value="2.85" valueMax="3.6" valueMin="2.1">2.1–3.6 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B84E50FFFFDFFF913B9FAFFFDEAAC2B" blockId="1.[151,1437,731,1981]" pageId="1" pageNumber="385">
|
||
<emphasis id="B94F391DFFFDFFF913B9FAFFFEC3AF67" bold="true" box="[199,270,1307,1333]" pageId="1" pageNumber="385">Head.</emphasis>
|
||
Head including eyes broader than pronotum. Eyes large, with sparse erect setae, protruding laterally, anteriorly kidney-shaped (
|
||
<figureCitation id="1300F98AFFFDFFF912BDFADBFDF6AF0B" box="[451,571,1343,1369]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="1" pageNumber="385">Figs 7–12</figureCitation>
|
||
). Frons and vertex wrinkled. Labrum bilobed, anterior margin with yellow setae and emarginate in middle (
|
||
<figureCitation id="1300F98AFFFDFFF9117EFA87FDB8AF2F" box="[512,629,1379,1405]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="1" pageNumber="385">Figs 7–12</figureCitation>
|
||
). Terminal maxillary palpomeres cylindrical; terminal labial palpomeres securiform. Clypeus shiny, straight anteriorly, slightly dilated posteriad, sparsely pubescent.Antennae 11-segmented, biflabeloid, with flabelae situated at base of each antennomere III-X (
|
||
<figureCitation id="1300F98AFFFDFFF910C4FA4FFBEFAF97" box="[954,1058,1451,1477]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="1" pageNumber="385">Figs 1–3</figureCitation>
|
||
). Flabelae 2–3 times longer than length of respective antennomere. Flabelae equal in width along their whole length, divided, unjointed.Antennomere I thickened, approx. 2.5 longer than antennomere II, which is strongly shortened. Antennomere II with a small pronounced bump on inner side. Antennomeres III-X approx. equal in length. Dorsal side of antennomeres sparsely covered with long yellow recumbent hairs whereas inner side of antennomeres and flabelae is densely covered with shorter white erect hairs (
|
||
<figureCitation id="1300F98AFFFDFFF912CAF9BBFDD4AC2B" box="[436,537,1631,1657]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="1" pageNumber="385">Figs 1–3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B84E50FFFFDFFF913B9F967FCC4AC93" blockId="1.[151,1437,731,1981]" pageId="1" pageNumber="385">
|
||
<emphasis id="B94F391DFFFDFFF913B9F967FE8AACCF" bold="true" box="[199,327,1667,1693]" pageId="1" pageNumber="385">Pronotum.</emphasis>
|
||
Shiny, longer than wide, cylindrical, surface roughly and irregularly wrinkled, whole surface covered with long erect and semi-erect hairs (
|
||
<figureCitation id="1300F98AFFFDFFF911E8F943FD30AC93" box="[662,765,1703,1729]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="1" pageNumber="385">Figs 7–9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B84E50FFFFDFFF913B9F92FFD09AD27" blockId="1.[151,1437,731,1981]" pageId="1" pageNumber="385">
|
||
<emphasis id="B94F391DFFFDFFF913B9F92FFED7ACB7" bold="true" box="[199,282,1739,1765]" pageId="1" pageNumber="385">Elytra.</emphasis>
|
||
Elytra shiny, parallel or subparallel, distinctly longer than their width, with posterior half parallel or slightly widening posteriad, whole surface sparsely covered with semi-erect yellow hairs, more densely in lateral part of elytra. Humeri well developed, distinct, almost rectangular. Elytral surface with bituberculate foveae - a pair of tubercles situated opposite each other on the top of the large round depressions (
|
||
<figureCitation id="1300F98AFFFDFFF91759F8D3FBB0AD03" box="[1063,1149,1847,1873]" captionStart="FIGURE 13–20" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 13–20. Biflabelotillus foveolatus Pic, 1949—13–15 (lectotype), 16–20 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="1" pageNumber="385">Figs 20</figureCitation>
|
||
,
|
||
<figureCitation id="1300F98AFFFDFFF917F9F8D3FB68AD03" box="[1159,1189,1847,1873]" captionStart="FIGURE 21–27" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[225,1371,187,1919]" captionTargetId="figure-17@6.[215,1371,181,1919]" captionTargetPageId="6" captionText="FIGURE 21–27. Biflabelotillus peyrierasi Menier, 1979—21–23 (holotype), 24–27 (non-type, male). Fig. 21—labels of the holotype; 21, 24—habitus, dorsal view; 23, 25—habitus, lateral view; 26—scutellum; 27—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915941" httpUri="https://zenodo.org/record/13915941/files/figure.png" pageId="1" pageNumber="385">27</figureCitation>
|
||
,
|
||
<figureCitation id="1300F98AFFFDFFF917CEF8D3FB01AD03" box="[1200,1228,1847,1873]" captionStart="FIGURE 28–32" captionStartId="7.[151,250,1586,1610]" captionTargetBox="[170,1418,184,1562]" captionTargetId="figure-26@7.[170,1418,181,1562]" captionTargetPageId="7" captionText="FIGURE 28–32. Biflabelotillus peyrierasi Menier, 1979, female. Fig. 28—habitus, dorsal view; 29—habitus, ventral view; 30—antenna (red arrow indicates last antennomere); 31—scutellum; 32—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915943" httpUri="https://zenodo.org/record/13915943/files/figure.png" pageId="1" pageNumber="385">32</figureCitation>
|
||
and
|
||
<figureCitation id="1300F98AFFFDFFF9167EF8D3FAEDAD03" box="[1280,1312,1847,1873]" captionStart="FIGURE 33–37" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 33–37. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="1" pageNumber="385">37</figureCitation>
|
||
). The latter arranged in regular rows (
|
||
<figureCitation id="1300F98AFFFDFFF912C6F8BFFDC2AD27" box="[440,527,1883,1909]" captionStart="FIGURE 13–20" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 13–20. Biflabelotillus foveolatus Pic, 1949—13–15 (lectotype), 16–20 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="1" pageNumber="385">Figs 17</figureCitation>
|
||
,
|
||
<figureCitation id="1300F98AFFFDFFF91165F8BFFDF4AD27" box="[539,569,1883,1909]" captionStart="FIGURE 21–27" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[225,1371,187,1919]" captionTargetId="figure-17@6.[215,1371,181,1919]" captionTargetPageId="6" captionText="FIGURE 21–27. Biflabelotillus peyrierasi Menier, 1979—21–23 (holotype), 24–27 (non-type, male). Fig. 21—labels of the holotype; 21, 24—habitus, dorsal view; 23, 25—habitus, lateral view; 26—scutellum; 27—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915941" httpUri="https://zenodo.org/record/13915941/files/figure.png" pageId="1" pageNumber="385">24</figureCitation>
|
||
,
|
||
<figureCitation id="1300F98AFFFDFFF9113BF8BFFDACAD27" box="[581,609,1883,1909]" captionStart="FIGURE 28–32" captionStartId="7.[151,250,1586,1610]" captionTargetBox="[170,1418,184,1562]" captionTargetId="figure-26@7.[170,1418,181,1562]" captionTargetPageId="7" captionText="FIGURE 28–32. Biflabelotillus peyrierasi Menier, 1979, female. Fig. 28—habitus, dorsal view; 29—habitus, ventral view; 30—antenna (red arrow indicates last antennomere); 31—scutellum; 32—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915943" httpUri="https://zenodo.org/record/13915943/files/figure.png" pageId="1" pageNumber="385">28</figureCitation>
|
||
and
|
||
<figureCitation id="1300F98AFFFDFFF911E6F8BFFD75AD27" box="[664,696,1883,1909]" captionStart="FIGURE 33–37" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 33–37. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="1" pageNumber="385">34</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B84E50FFFFDFFF913B9F89AFC0FADCB" blockId="1.[151,1437,731,1981]" box="[199,962,1918,1945]" pageId="1" pageNumber="385">
|
||
<emphasis id="B94F391DFFFDFFF913B9F89AFE8BADCA" bold="true" box="[199,326,1918,1944]" pageId="1" pageNumber="385">Scutellum.</emphasis>
|
||
Almost square or subtriangular (
|
||
<figureCitation id="1300F98AFFFDFFF911C8F89BFCC0ADCB" box="[694,781,1919,1945]" captionStart="FIGURE 13–20" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 13–20. Biflabelotillus foveolatus Pic, 1949—13–15 (lectotype), 16–20 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="1" pageNumber="385">Figs 19</figureCitation>
|
||
,
|
||
<figureCitation id="1300F98AFFFDFFF91067F89BFCFAADCB" box="[793,823,1919,1945]" captionStart="FIGURE 21–27" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[225,1371,187,1919]" captionTargetId="figure-17@6.[215,1371,181,1919]" captionTargetPageId="6" captionText="FIGURE 21–27. Biflabelotillus peyrierasi Menier, 1979—21–23 (holotype), 24–27 (non-type, male). Fig. 21—labels of the holotype; 21, 24—habitus, dorsal view; 23, 25—habitus, lateral view; 26—scutellum; 27—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915941" httpUri="https://zenodo.org/record/13915941/files/figure.png" pageId="1" pageNumber="385">26</figureCitation>
|
||
,
|
||
<figureCitation id="1300F98AFFFDFFF9103DF89BFC92ADCB" box="[835,863,1919,1945]" captionStart="FIGURE 28–32" captionStartId="7.[151,250,1586,1610]" captionTargetBox="[170,1418,184,1562]" captionTargetId="figure-26@7.[170,1418,181,1562]" captionTargetPageId="7" captionText="FIGURE 28–32. Biflabelotillus peyrierasi Menier, 1979, female. Fig. 28—habitus, dorsal view; 29—habitus, ventral view; 30—antenna (red arrow indicates last antennomere); 31—scutellum; 32—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915943" httpUri="https://zenodo.org/record/13915943/files/figure.png" pageId="1" pageNumber="385">31</figureCitation>
|
||
and
|
||
<figureCitation id="1300F98AFFFDFFF910EBF89BFC78ADCB" box="[917,949,1919,1945]" captionStart="FIGURE 33–37" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 33–37. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="1" pageNumber="385">36</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B84E50FFFFDFFF913B9F847FCFDADEF" blockId="1.[151,1437,731,1981]" box="[199,816,1955,1981]" pageId="1" pageNumber="385">
|
||
<emphasis id="B94F391DFFFDFFF913B9F847FECBADEF" bold="true" box="[199,262,1955,1981]" pageId="1" pageNumber="385">Legs.</emphasis>
|
||
Relatively densely covered with long erect hairs.
|
||
</paragraph>
|
||
<paragraph id="8B84E50FFFFEFFFA13B9FF73FB0EAAAB" blockId="2.[151,1436,150,285]" pageId="2" pageNumber="386">
|
||
<emphasis id="B94F391DFFFEFFFA13B9FF73FEB1AAE2" bold="true" box="[199,380,150,177]" pageId="2" pageNumber="386">Lower surface.</emphasis>
|
||
Shiny, covered with long semi-erect hairs; pro-intercoxal process narrow, slightly expanded distally; anterior margin of mesoventrite straight, lateral margins divergent posteriad, meso-intercoxal process narrow, pointed; metaventrite subquadrate; abdomen with six visible ventrites (
|
||
<figureCitation id="1300F98AFFFEFFFA1770FF3BFBADAAAB" box="[1038,1120,223,249]" captionStart="FIGURE 13–20" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 13–20. Biflabelotillus foveolatus Pic, 1949—13–15 (lectotype), 16–20 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="2" pageNumber="386">Fig. 18</figureCitation>
|
||
,
|
||
<figureCitation id="1300F98AFFFEFFFA1712FF3BFB47AAAB" box="[1132,1162,223,249]" captionStart="FIGURE 28–32" captionStartId="7.[151,250,1586,1610]" captionTargetBox="[170,1418,184,1562]" captionTargetId="figure-26@7.[170,1418,181,1562]" captionTargetPageId="7" captionText="FIGURE 28–32. Biflabelotillus peyrierasi Menier, 1979, female. Fig. 28—habitus, dorsal view; 29—habitus, ventral view; 30—antenna (red arrow indicates last antennomere); 31—scutellum; 32—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915943" httpUri="https://zenodo.org/record/13915943/files/figure.png" pageId="2" pageNumber="386">29</figureCitation>
|
||
,
|
||
<figureCitation id="1300F98AFFFEFFFA17E8FF3AFB78AAAB" box="[1174,1205,222,249]" captionStart="FIGURE 33–37" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 33–37. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="2" pageNumber="386">35</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C321B684FFFEFFFA13B9FEE6FD14AB4F" box="[199,729,258,285]" pageId="2" pageNumber="386" type="distribution">
|
||
<paragraph id="8B84E50FFFFEFFFA13B9FEE6FD14AB4F" blockId="2.[151,1436,150,285]" box="[199,729,258,285]" pageId="2" pageNumber="386">
|
||
<emphasis id="B94F391DFFFEFFFA13B9FEE6FEACAB4E" bold="true" box="[199,353,258,284]" pageId="2" pageNumber="386">Distribution.</emphasis>
|
||
Central and Eastern
|
||
<collectingCountry id="F32CA59FFFFEFFFA1135FEE7FD19AB4F" box="[587,724,259,285]" name="Madagascar" pageId="2" pageNumber="386">Madagascar</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |