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<document id="83FA5686AD1554B8AC2557D5125EB93B" ID-CLB-Dataset="8298" ID-DOI="10.11646/zootaxa.4926.2.5" ID-GBIF-Dataset="0b136600-8384-4130-8c2e-94c6678f3dd2" ID-ISSN="1175-5326" ID-Zenodo-Dep="4506132" ID-ZooBank="FE8A891C-293F-483F-B892-E58CA2D7387C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612508629043" checkinUser="plazi" docAuthor="Razanajaonarivalona, Elysé Hugo, Moravec, Jiří &amp; Rakotomanana, Hajanirina" docDate="2021" docId="03F75D7ECD39FFF0FF5CFF7DFE001F9E" docLanguage="en" docName="zootaxa.4926.2.5.pdf" docOrigin="Zootaxa 4926 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pogonostoma (Bathypogonum) levigatum subsp. lucens Rivalier 1970" docType="treatment" docVersion="7" lastPageNumber="258" masterDocId="FFCE2506CD34FFFDFFCBFFEAFFE01949" masterDocTitle="New or rare Madagascar tiger beetles- 20. Pogonostoma (Bathypogonum) horimichioi sp. nov. and supplemented characters of P. (B) levigatum levigatum (W. Horn) and P. (B.) levigatum lucens Rivalier (Coleoptera: Cicindelidae)" masterLastPageNumber="262" masterPageNumber="245" pageNumber="258" updateTime="1698927655442" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="B6B2F4E002D7AA1AF9832EA00974B30F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EF374C6DFBCA30CA47B38A5BABC2CF5C">
<mods:title id="970EC31BE6C2C012E4703EB4168C34B8">New or rare Madagascar tiger beetles- 20. Pogonostoma (Bathypogonum) horimichioi sp. nov. and supplemented characters of P. (B) levigatum levigatum (W. Horn) and P. (B.) levigatum lucens Rivalier (Coleoptera: Cicindelidae)</mods:title>
</mods:titleInfo>
<mods:name id="AFC522329AD606CB1F99FE759F8F0727" type="personal">
<mods:role id="8777FB959C8FDEEEAD9C0D6674C488DE">
<mods:roleTerm id="6CA152740E829B23FD5233FB49830180">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4E4E70AA0A818E55F4F0F02C25B15AFF">Razanajaonarivalona, Elysé Hugo</mods:namePart>
<mods:nameIdentifier id="0CE4F858C9A93DA694F5291688FB57A8" type="ORCID">0000-0002-9430-6793</mods:nameIdentifier>
<mods:nameIdentifier id="BF1FEF69B2FCA5543344FD4E995E80F3" type="email">relysehugo@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name id="5606B914014FB95D5BD29DFD0166731C" type="personal">
<mods:role id="D9DDAF9132ACD2F1E7938B3177EEF5B1">
<mods:roleTerm id="F1615690DC8F0C17858AF006A853657F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="83DFED26D17CA2F7C52E4F7B5752509F">Moravec, Jiří</mods:namePart>
<mods:affiliation id="0DE17C39BDE45EBBAC8F2D282E11C986">Mendel University in Brno, Department of Forest Ecology, Zemědělská 3, 613 00 Brno, Czech Republic</mods:affiliation>
</mods:name>
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<mods:role id="12EB96E8A2B9F75EEB2840DE0DE3F5E3">
<mods:roleTerm id="07B96B21F60933FE52E0907B7421E000">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1976C901D6561E1D69DA5B6452C50B3F">Rakotomanana, Hajanirina</mods:namePart>
<mods:nameIdentifier id="73C17A558834077FC026FFD512EC461E" type="ORCID">0000-0002-9430-6793</mods:nameIdentifier>
<mods:affiliation id="205C4BA375212DBEA84CF46A5DCEA62F">Département de Zoologie et Biodiversité Animale, Faculté des Sciences, Université d’Antananarivo, BP 906, Antananarivo 101, Madagascar. rakotomh @ yahoo. com; https: // orcid. org / 0000 - 0002 - 0547 - 7161</mods:affiliation>
<mods:nameIdentifier id="094A8B41481721E2041DB870A53D5BEE" type="email">relysehugo@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="30DCCE057D2E490E05A20A193B183E2C">text</mods:typeOfResource>
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<mods:title id="819095F9446B2B3BB94D142C5F7E28EE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1C0BD4939285DFD34D21AD917FAB06C0">
<mods:date id="5AFA3B945E8198EDA57B310B39773D93">2021</mods:date>
<mods:detail id="8C9559E4D8262046BA5EB108E2D5ADDF" type="pubDate">
<mods:number id="FEBA1E52DDF8DC9508EA6757EB9F3724">2021-02-05</mods:number>
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<mods:detail id="DCDF22F64623F285E983D15CEFECC832" type="volume">
<mods:number id="D24BB67D815B0C70B40F7DAB84F8ACC7">4926</mods:number>
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<mods:classification id="060ADFEB7B84BB24BB22B08C1DAC308E">journal article</mods:classification>
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<mods:identifier id="53AA3AD75784B959D6C713897BD30762" type="DOI">10.11646/zootaxa.4926.2.5</mods:identifier>
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<treatment id="03F75D7ECD39FFF0FF5CFF7DFE001F9E" ID-DOI="http://doi.org/10.5281/zenodo.4534305" ID-GBIF-Taxon="177318856" ID-Zenodo-Dep="4534305" LSID="urn:lsid:plazi:treatment:03F75D7ECD39FFF0FF5CFF7DFE001F9E" httpUri="http://treatment.plazi.org/id/03F75D7ECD39FFF0FF5CFF7DFE001F9E" lastPageNumber="258" pageId="13" pageNumber="258">
<subSubSection id="C344BFE3CD39FFF0FF5CFF7DFC7819F8" box="[151,920,151,178]" pageId="13" pageNumber="258" type="nomenclature">
<paragraph id="8BE1EC68CD39FFF0FF5CFF7DFC7819F8" blockId="13.[151,920,151,213]" box="[151,920,151,178]" pageId="13" pageNumber="258">
<heading id="D0A95B04CD39FFF0FF5CFF7DFC7819F8" bold="true" box="[151,920,151,178]" fontSize="11" level="1" pageId="13" pageNumber="258" reason="1">
<taxonomicName id="4C5E97EBCD39FFF0FF5CFF7DFC7819F8" ID-CoL="8MC53" authority="Rivalier, 1970" authorityName="Rivalier" authorityYear="1970" box="[151,920,151,178]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="258" phylum="Arthropoda" rank="subSpecies" species="levigatum" subGenus="Bathypogonum" subSpecies="lucens">
<emphasis id="B92A307ACD39FFF0FF5CFF7DFC7819F8" bold="true" box="[151,920,151,178]" pageId="13" pageNumber="258">
<emphasis id="B92A307ACD39FFF0FF5CFF7DFEDC19F8" bold="true" box="[151,316,151,177]" italics="true" pageId="13" pageNumber="258">Pogonostoma</emphasis>
(
<emphasis id="B92A307ACD39FFF0FE86FF7DFDE719FB" bold="true" box="[333,519,151,178]" italics="true" pageId="13" pageNumber="258">Bathypogonum</emphasis>
)
<emphasis id="B92A307ACD39FFF0FDD3FF7DFD0419FB" bold="true" box="[536,740,151,178]" italics="true" pageId="13" pageNumber="258">levigatum lucens</emphasis>
<bibRefCitation id="EFCF9199CD39FFF0FD27FF7DFC7819F8" author="Rivalier, E." box="[748,920,151,177]" pageId="13" pageNumber="258" pagination="269 - 338" refId="ref9106" refString="Rivalier, E. (1970) Le genre Pogonostoma (Col. Cicindelidae). Revision avec description d'especes nouvelles. Annales de la Societe entomologique de France, New Series, 6, 269 - 338." type="journal article" year="1970">Rivalier, 1970</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C344BFE3CD39FFF0FF5CFF51FE9B199C" box="[151,379,187,213]" pageId="13" pageNumber="258" type="description">
<paragraph id="8BE1EC68CD39FFF0FF5CFF51FE9B199C" blockId="13.[151,920,151,213]" box="[151,379,187,213]" pageId="13" pageNumber="258">
(
<figureCitation id="1365F0EDCD39FFF0FF54FF51FF16199C" box="[159,246,187,213]" captionStart="FIGURES 26–28" captionStartId="10.[151,264,1841,1865]" captionTargetBox="[159,1423,189,1805]" captionTargetId="figure-23@10.[151,1435,181,1817]" captionTargetPageId="10" captionText="FIGURES 26–28. Pogonostoma (Bathypogonum) levigatum, habitat. 26–27—P. (B.) l. levigatum W. Horn. 26—♁, 15.6 mm, Amboaniho, ST (SDEI); 27—♁, 15.3 mm, Marotaonala (CCJM); 28—P. (B.) l. lucens Rivalier, ♀, 15.5 mm, Île de Beraphia, LT (MNHN). Bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4506142" httpUri="https://zenodo.org/record/4506142/files/figure.png" pageId="13" pageNumber="258">Figs 28</figureCitation>
,
<figureCitation id="1365F0EDCD39FFF0FEC9FF51FEAA199C" box="[258,330,187,213]" captionStart="FIGURES 39–44" captionStartId="14.[151,264,1867,1891]" captionTargetBox="[158,1429,187,1840]" captionTargetId="figure-23@14.[151,1435,181,1844]" captionTargetPageId="14" captionText="FIGURES 39–44. Pogonostoma (Bathypogonum) levigatum. 39–42— P. (B.) l. levigatum W. Horn. 39–40—pronotum: 39—♁, Marotaonala (CCJM); 40—♀, Vohemar (CCJM ex APCA); 41–44—male aedeagus or its apex: 41—showing internal sac, Ma- rotaonala; 42—ditto, dorsal view; 43–44—P. (B.) l. lucens Rivalier. 43—Ramena River (SDEI); 44—ditto, dorsal view. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4506146" httpUri="https://zenodo.org/record/4506146/files/figure.png" pageId="13" pageNumber="258">43–44</figureCitation>
,
<figureCitation id="1365F0EDCD39FFF0FE9DFF51FE93199C" box="[342,371,187,213]" captionStart="FIGURES 45–47" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[215,1373,193,1907]" captionTargetId="figure-17@15.[209,1379,181,1919]" captionTargetPageId="15" captionText="FIGURES 45–47. Distribution of Pogonostoma (Bathypogonum). 45– P. (B.) horimichioi sp. nov.; 46—P. (B.) l. levigatum W. Horn; 47—P. (B.) l. lucens Rivalier." figureDoi="http://doi.org/10.5281/zenodo.4506148" httpUri="https://zenodo.org/record/4506148/files/figure.png" pageId="13" pageNumber="258">47</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C344BFE3CD39FFF0FF5CFEEAFB83187F" pageId="13" pageNumber="258" type="reference_group">
<paragraph id="8BE1EC68CD39FFF0FF5CFEEAFB3C1851" blockId="13.[151,1244,256,310]" box="[151,1244,256,280]" pageId="13" pageNumber="258">
<treatmentCitationGroup id="AB4ECB46CD39FFF0FF5CFEEAFB3C1851" box="[151,1244,256,280]" pageId="13" pageNumber="258">
<taxonomicName id="4C5E97EBCD39FFF0FF5CFEEAFDAA1851" authority="Rivalier, 1970" box="[151,586,256,280]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="258" phylum="Arthropoda" rank="species" species="laevigatum" subGenus="Bathypogonum">
<emphasis id="B92A307ACD39FFF0FF5CFEEAFEC2185E" box="[151,290,256,279]" italics="true" pageId="13" pageNumber="258">Pogonostoma</emphasis>
(
<emphasis id="B92A307ACD39FFF0FEFAFEEAFE2A1851" box="[305,458,256,280]" italics="true" pageId="13" pageNumber="258">Bathypogonum</emphasis>
)
<emphasis id="B92A307ACD39FFF0FE12FEEAFDAA1851" box="[473,586,256,280]" italics="true" pageId="13" pageNumber="258">laevigatum</emphasis>
</taxonomicName>
(sic!)
<emphasis id="B92A307ACD39FFF0FD47FEEAFD2D1851" box="[652,717,256,280]" italics="true" pageId="13" pageNumber="258">lucens</emphasis>
<treatmentCitation id="0AFFCA79CD39FFF0FD18FEEAFC761851" author="Rivalier, E." box="[723,918,256,280]" page="312" pageId="13" pageNumber="258" year="1970">
<bibRefCitation id="EFCF9199CD39FFF0FD18FEEAFC761851" author="Rivalier, E." box="[723,918,256,280]" pageId="13" pageNumber="258" pagination="269 - 338" refId="ref9106" refString="Rivalier, E. (1970) Le genre Pogonostoma (Col. Cicindelidae). Revision avec description d'especes nouvelles. Annales de la Societe entomologique de France, New Series, 6, 269 - 338." type="journal article" year="1970">Rivalier, 1970: 312</bibRefCitation>
</treatmentCitation>
(incorrect subsequent spelling).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE1EC68CD39FFF0FF5CFEF5FB83187F" blockId="13.[151,1244,256,310]" box="[151,1123,286,310]" pageId="13" pageNumber="258">
<treatmentCitationGroup id="AB4ECB46CD39FFF0FF5CFEF5FB83187F" box="[151,1123,286,310]" pageId="13" pageNumber="258">
<treatmentCitation id="0AFFCA79CD39FFF0FF5CFEF5FCAD187F" author="Moravec, J." box="[151,845,286,310]" page="84" pageId="13" pageNumber="258" year="2007">
<taxonomicName id="4C5E97EBCD39FFF0FF5CFEF5FCAD187F" ID-CoL="8MC53" authority=": Moravec 2007: 84" authorityName="Moravec" authorityPageNumber="84" authorityYear="2007" box="[151,845,286,310]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="258" phylum="Arthropoda" rank="subSpecies" species="levigatum" subGenus="Bathypogonum" subSpecies="lucens">
<emphasis id="B92A307ACD39FFF0FF5CFEF5FEC2187F" box="[151,290,287,310]" italics="true" pageId="13" pageNumber="258">Pogonostoma</emphasis>
(
<emphasis id="B92A307ACD39FFF0FEFAFEF4FE2A187F" box="[305,458,286,310]" italics="true" pageId="13" pageNumber="258">Bathypogonum</emphasis>
)
<emphasis id="B92A307ACD39FFF0FE12FEF4FD65187F" box="[473,645,286,310]" italics="true" pageId="13" pageNumber="258">levigatum lucens</emphasis>
:
<bibRefCitation id="EFCF9199CD39FFF0FD59FEF5FCAD187F" author="Moravec, J." box="[658,845,286,310]" pageId="13" pageNumber="258" refId="ref8440" refString="Moravec, J. (2007) Tiger beetles of Madagascar 1. A monograph of the genus Pogonostoma (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, 499 pp." type="book" year="2007">Moravec 2007: 84</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 272–285, 1662–1664.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C344BFE3CD39FFF0FF5CFE8FFAB31BAE" pageId="13" pageNumber="258" type="materials_examined">
<paragraph id="8BE1EC68CD39FFF0FF5CFE8FFF1318EA" blockId="13.[151,1437,356,1751]" pageId="13" pageNumber="258">
<materialsCitation id="3B36E635CD39FFF0FF5CFE8FFF0F18EA" ID-GBIF-Occurrence="3033947312" collectorName="de Beraphia" country="Madagascar" location="western Sambirano" municipality="Ile de Beraphia" pageId="13" pageNumber="258" specimenCount="1" typeStatus="holotype">
<emphasis id="B92A307ACD39FFF0FF5CFE8FFED41837" bold="true" box="[151,308,356,383]" pageId="13" pageNumber="258">
<typeStatus id="54E552CACD39FFF0FF5CFE8FFF321836" box="[151,210,357,383]" pageId="13" pageNumber="258">Type</typeStatus>
locality
</emphasis>
.
<collectingCountry id="F349ACF8CD39FFF0FE8DFE8EFE2F1837" box="[326,463,356,382]" name="Madagascar" pageId="13" pageNumber="258">Madagascar</collectingCountry>
,
<location id="8E81BAB3CD39FFF0FE16FE8FFD5D1837" LSID="urn:lsid:plazi:treatment:03F75D7ECD39FFF0FF5CFF7DFE001F9E:8E81BAB3CD39FFF0FE16FE8FFD5D1837" box="[477,701,356,383]" country="Madagascar" municipality="Ile de Beraphia" name="western Sambirano" pageId="13" pageNumber="258">western Sambirano</location>
:
<collectingMunicipality id="6B857612CD39FFF0FD00FE8CFC9E1837" box="[715,894,356,385]" pageId="13" pageNumber="258">
Île
<collectorName id="26AB89BECD39FFF0FD3FFE8EFC9E1837" box="[756,894,356,382]" pageId="13" pageNumber="258">de Beraphia</collectorName>
</collectingMunicipality>
(island of Nosy Berafia of the Radama archipelago)
</materialsCitation>
.
</paragraph>
<paragraph id="8BE1EC68CD39FFF0FF0CFE47FDFE1B1E" blockId="13.[151,1437,356,1751]" pageId="13" pageNumber="258">
<emphasis id="B92A307ACD39FFF0FF0CFE47FE8C188F" bold="true" box="[199,364,428,455]" pageId="13" pageNumber="258">Type material</emphasis>
.
<materialsCitation id="3B36E635CD39FFF0FEB3FE47FB121B46" ID-GBIF-Occurrence="3033947301" collectionCode="MNHN" collectorName="Coll. R. Oberthur" country="Madagascar" location="Ile de Beraphia" pageId="13" pageNumber="258" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
<typeStatus id="54E552CACD39FFF0FEB3FE47FE08188E" box="[376,488,429,455]" pageId="13" pageNumber="258" type="lectotype">Lectotype</typeStatus>
<specimenCount id="9D5827E1CD39FFF0FE25FE47FDE3188E" box="[494,515,429,455]" pageId="13" pageNumber="258" type="female">♀</specimenCount>
in
<collectionCode id="ED4F74ADCD39FFF0FDE8FE47FD99188E" box="[547,633,429,455]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="258">MNHN</collectionCode>
(designated by Moravec 2070), labelled: “
<collectingCountry id="F349ACF8CD39FFF0FB80FE46FB35188F" box="[1099,1237,428,454]" name="Madagascar" pageId="13" pageNumber="258">Madagascar</collectingCountry>
/
<location id="8E81BAB3CD39FFF0FB23FE44FA73188F" LSID="urn:lsid:plazi:treatment:03F75D7ECD39FFF0FF5CFF7DFE001F9E:8E81BAB3CD39FFF0FB23FE44FA73188F" box="[1256,1427,428,457]" country="Madagascar" name="Ile de Beraphia" pageId="13" pageNumber="258">Île de Beraphia</location>
” / Cote Ouest, 1934”; [printed]; “Muséum Paris, 1952 /
<collectorName id="26AB89BECD39FFF0FCC9FE3AFC2A18A3" box="[770,970,464,491]" pageId="13" pageNumber="258">Coll. R. Oberthür</collectorName>
” [bluish, printed]; “
<typeStatus id="54E552CACD39FFF0FB62FE3BFAFC18A2" box="[1193,1308,465,491]" pageId="13" pageNumber="258" type="lectotype">Lectotype</typeStatus>
/
<taxonomicName id="4C5E97EBCD39FFF0FAF9FE3BFE761B46" authority="" authorityName="Jeannel" authorityYear="1946" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="258" phylum="Arthropoda" rank="subGenus" subGenus="Bathypogonum">Pogonostoma (Bathypogonum)</taxonomicName>
/ levigatum lucens
<bibRefCitation id="EFCF9199CD39FFF0FDBBFE1EFCEF1B46" author="Rivalier, E." box="[624,783,500,527]" pageId="13" pageNumber="258" pagination="269 - 338" refId="ref9106" refString="Rivalier, E. (1970) Le genre Pogonostoma (Col. Cicindelidae). Revision avec description d'especes nouvelles. Annales de la Societe entomologique de France, New Series, 6, 269 - 338." type="journal article" year="1970">Rivalier, 1970</bibRefCitation>
/ design. Jiří Moravec 2003” [red, printed]
</materialsCitation>
.
<materialsCitation id="3B36E635CD39FFF0FB36FE1EFDF91B1E" ID-GBIF-Occurrence="3033947306" collectionCode="MNHN" country="Madagascar" pageId="13" pageNumber="258" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<typeStatus id="54E552CACD39FFF0FB36FE1EFA791B47" box="[1277,1433,500,526]" pageId="13" pageNumber="258" type="paralectotype">Paralectotype</typeStatus>
.
<specimenCount id="9D5827E1CD39FFF0FF5CFDF3FF221B7A" box="[151,194,537,563]" pageId="13" pageNumber="258" type="female">1 ♀</specimenCount>
in
<collectionCode id="ED4F74ADCD39FFF0FF2CFDF3FEDD1B7A" box="[231,317,537,563]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="258">MNHN</collectionCode>
with same labels and: “Revision Jiří Moravec 2003: /
<typeStatus id="54E552CACD39FFF0FC6DFDF2FBDE1B7B" box="[934,1086,536,562]" pageId="13" pageNumber="258" type="paralectotype">Paralectotype</typeStatus>
/
<taxonomicName id="4C5E97EBCD39FFF0FB9DFDF3FB0E1B7A" authorityName="Klug" authorityYear="1835" box="[1110,1262,537,563]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="258" phylum="Arthropoda" rank="genus">Pogonostoma</taxonomicName>
/ levigatum lu- cens
<bibRefCitation id="EFCF9199CD39FFF0FF1BFDD6FE911B1E" author="Rivalier, E." box="[208,369,572,599]" pageId="13" pageNumber="258" pagination="269 - 338" refId="ref9106" refString="Rivalier, E. (1970) Le genre Pogonostoma (Col. Cicindelidae). Revision avec description d'especes nouvelles. Annales de la Societe entomologique de France, New Series, 6, 269 - 338." type="journal article" year="1970">Rivalier, 1970</bibRefCitation>
” [red, printed]
</materialsCitation>
.
</paragraph>
<paragraph id="8BE1EC68CD39FFF0FF0CFD8AFAB31BAE" blockId="13.[151,1437,356,1751]" pageId="13" pageNumber="258">
<emphasis id="B92A307ACD39FFF0FF0CFD8AFE1A1B33" bold="true" box="[199,506,608,634]" pageId="13" pageNumber="258">Other material examined</emphasis>
. 5 ♁♁,
<materialsCitation id="3B36E635CD39FFF0FD96FD8BFD291BA0" ID-GBIF-Occurrence="3033947304" collectionCode="SDEI, MFNB" collectorName="W. Horn " determinerName="W. Horn" location="Antananarivo" pageId="13" pageNumber="258" specimenCount="2" specimenCount-female="2" stateProvince="Antananarivo">
<specimenCount id="9D5827E1CD39FFF0FD96FD8BFD7F1B32" box="[605,671,609,635]" pageId="13" pageNumber="258" type="female">2 ♀♀</specimenCount>
in
<collectionCode id="ED4F74ADCD39FFF0FD02FD8BFCE71B32" box="[713,775,609,635]" pageId="13" pageNumber="258">SDEI</collectionCode>
[filed as “
<emphasis id="B92A307ACD39FFF0FC4EFD8AFBEE1B33" box="[901,1038,608,634]" italics="true" pageId="13" pageNumber="258">levipunctum</emphasis>
<collectorName id="26AB89BECD39FFF0FBDCFD8BFB9A1B32" box="[1047,1146,609,635]" pageId="13" pageNumber="258">W. Horn</collectorName>
i. l.”]: “
<collectorName id="26AB89BECD39FFF0FB16FD8BFA801B32" box="[1245,1376,609,635]" pageId="13" pageNumber="258">R. Ramena</collectorName>
, NW Madag. /
<collectorName id="26AB89BECD39FFF0FF34FD6FFE811BD7" box="[255,353,644,671]" pageId="13" pageNumber="258">J. Mellis</collectorName>
” // “
<collectorName id="26AB89BECD39FFF0FE5BFD6EFDCD1BD6" box="[400,557,644,671]" pageId="13" pageNumber="258">Coll. W. Horn</collectorName>
/ DEI Eberswalde”. 1 ♁ in
<collectionCode id="ED4F74ADCD39FFF0FC92FD6FFC7C1BD6" box="[857,924,645,671]" pageId="13" pageNumber="258">SDEI</collectionCode>
: “Bealanana, NW Madag.”; “levipunctum m.” // “
<collectorName id="26AB89BECD39FFF0FF73FD42FEB71B8A" box="[184,343,680,707]" pageId="13" pageNumber="258">Coll. W. Horn</collectorName>
/ DEI Eberswalde”. 1 ♁ in
<collectionCode id="ED4F74ADCD39FFF0FD40FD43FD2E1B8A" box="[651,718,681,707]" pageId="13" pageNumber="258">SDEI</collectionCode>
: “Tananarive / Winter 24”. 1 ♁ in
<collectionCode id="ED4F74ADCD39FFF0FB80FD43FB7F1B8A" box="[1099,1183,681,707]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="13" pageNumber="258">MFNB</collectionCode>
: “
<collectingRegion id="499A228ACD39FFF0FB78FD42FAB11B8B" box="[1203,1361,680,706]" country="Madagascar" name="Antananarivo" pageId="13" pageNumber="258">Antananarivo</collectingRegion>
” (sic!) / “
<taxonomicName id="4C5E97EBCD39FFF0FF64FD27FEAB1BAE" authorityName="Klug" authorityYear="1835" box="[175,331,717,743]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="258" phylum="Arthropoda" rank="genus">Pogonostoma</taxonomicName>
/ levigatum /
<determinerName id="7E3D00ACCD39FFF0FE2DFD27FDA61BAE" box="[486,582,717,743]" pageId="13" pageNumber="258">W. Horn</determinerName>
det”. 2 ♁♁
</materialsCitation>
,
<materialsCitation id="3B36E635CD39FFF0FD1DFD27FAAE1BAE" ID-GBIF-Occurrence="3033947308" box="[726,1358,716,743]" collectionCode="MFNB" country="Madagascar" location="Madagascar" pageId="13" pageNumber="258" specimenCount="4" specimenCount-female="4">
<specimenCount id="9D5827E1CD39FFF0FD1DFD27FCF51BAE" box="[726,789,717,743]" pageId="13" pageNumber="258" type="female">4 ♀♀</specimenCount>
in
<collectionCode id="ED4F74ADCD39FFF0FCF2FD27FC6D1BAE" box="[825,909,717,743]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="13" pageNumber="258">MFNB</collectionCode>
: “
<collectingCountry id="F349ACF8CD39FFF0FC6BFD26FBCD1BAF" box="[928,1069,716,742]" name="Madagascar" pageId="13" pageNumber="258">Madagascar</collectingCountry>
” [filed as “
<taxonomicName id="4C5E97EBCD39FFF0FB67FD27FAD61BAF" box="[1196,1334,716,742]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="258" phylum="Arthropoda" rank="species" species="levigatum">
<emphasis id="B92A307ACD39FFF0FB67FD27FAD61BAF" box="[1196,1334,716,742]" italics="true" pageId="13" pageNumber="258">P. levigatum</emphasis>
</taxonomicName>
”]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C344BFE3CD39FFF0FF0CFD1AFC461DF2" pageId="13" pageNumber="258" type="diagnosis">
<paragraph id="8BE1EC68CD39FFF0FF0CFD1AFC621AD2" blockId="13.[151,1437,356,1751]" pageId="13" pageNumber="258">
<emphasis id="B92A307ACD39FFF0FF0CFD1AFE271A43" bold="true" box="[199,455,752,778]" pageId="13" pageNumber="258">Differential diagnosis</emphasis>
. Distinguished from the nominotypical subspecies by its even brighter body coloration (
<figureCitation id="1365F0EDCD39FFF0FF6BFCFEFF111A66" box="[160,241,788,815]" captionStart="FIGURES 26–28" captionStartId="10.[151,264,1841,1865]" captionTargetBox="[159,1423,189,1805]" captionTargetId="figure-23@10.[151,1435,181,1817]" captionTargetPageId="10" captionText="FIGURES 26–28. Pogonostoma (Bathypogonum) levigatum, habitat. 26–27—P. (B.) l. levigatum W. Horn. 26—♁, 15.6 mm, Amboaniho, ST (SDEI); 27—♁, 15.3 mm, Marotaonala (CCJM); 28—P. (B.) l. lucens Rivalier, ♀, 15.5 mm, Île de Beraphia, LT (MNHN). Bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4506142" httpUri="https://zenodo.org/record/4506142/files/figure.png" pageId="13" pageNumber="258">Fig. 28</figureCitation>
) and following features: elytra metallic blue or green-blue, often with golden lustre on elytral disc; elytral punctures shallower and with wider and flatter, notably shiny intervals; setal vesture predominantly whitish; lacinia paler with ochre-testaceous outer area; antennomeres 6–11 brownish. Aedeagus (
<figureCitation id="1365F0EDCD39FFF0FBD2FCB6FB791A3E" box="[1049,1177,860,887]" captionStart="FIGURES 39–44" captionStartId="14.[151,264,1867,1891]" captionTargetBox="[158,1429,187,1840]" captionTargetId="figure-23@14.[151,1435,181,1844]" captionTargetPageId="14" captionText="FIGURES 39–44. Pogonostoma (Bathypogonum) levigatum. 39–42— P. (B.) l. levigatum W. Horn. 39–40—pronotum: 39—♁, Marotaonala (CCJM); 40—♀, Vohemar (CCJM ex APCA); 41–44—male aedeagus or its apex: 41—showing internal sac, Ma- rotaonala; 42—ditto, dorsal view; 43–44—P. (B.) l. lucens Rivalier. 43—Ramena River (SDEI); 44—ditto, dorsal view. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4506146" httpUri="https://zenodo.org/record/4506146/files/figure.png" pageId="13" pageNumber="258">Figs 44–43</figureCitation>
) shaped generally as in the nominotypical subspecies with variably longer or shorter apex.
</paragraph>
<paragraph id="8BE1EC68CD39FFF0FF0CFC4FFE041AAA" blockId="13.[151,1437,356,1751]" pageId="13" pageNumber="258">
For the detailed redescription and illustrations of other characters, including of the
<typeStatus id="54E552CACD39FFF0FB93FC4EFB221AF7" box="[1112,1218,932,958]" pageId="13" pageNumber="258" type="lectotype">lectotype</typeStatus>
, see the monograph of the genus (
<bibRefCitation id="EFCF9199CD39FFF0FEE4FC23FE381AAA" author="Moravec, J." box="[303,472,969,995]" pageId="13" pageNumber="258" refId="ref8440" refString="Moravec, J. (2007) Tiger beetles of Madagascar 1. A monograph of the genus Pogonostoma (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, 499 pp." type="book" year="2007">Moravec 2007</bibRefCitation>
).
</paragraph>
<paragraph id="8BE1EC68CD39FFF0FF0CFC06FC461DF2" blockId="13.[151,1437,356,1751]" pageId="13" pageNumber="258">
<emphasis id="B92A307ACD39FFF0FF0CFC06FEA71D4F" bold="true" box="[199,327,1004,1030]" pageId="13" pageNumber="258">Variability</emphasis>
. Besides the rather variable coloration, also the length of the aedeagus apex considerably varies. As discussed and illustrated previously (
<bibRefCitation id="EFCF9199CD39FFF0FDA2FBFBFCF31D62" author="Moravec, J." box="[617,787,1041,1067]" pageId="13" pageNumber="258" refId="ref8440" refString="Moravec, J. (2007) Tiger beetles of Madagascar 1. A monograph of the genus Pogonostoma (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, 499 pp." type="book" year="2007">Moravec 2007</bibRefCitation>
, figs 278–284), the specimens from Ramena, Bealanana and “Tananarive”, arranged in SDEI under the unpublished name “
<emphasis id="B92A307ACD39FFF0FC4FFBDEFBED1D07" box="[900,1037,1076,1102]" italics="true" pageId="13" pageNumber="258">levipunctum</emphasis>
“ as well as the seven specimens in MFNB, show a considerable variability in the shape of the aedeagus apex. Nevertheless, as intermediate shapes of the aedeagi were found within the examined adults (also from the same locality), the differences in the length of the aedeagus apex were considered as a mere variability (
<bibRefCitation id="EFCF9199CD39FFF0FD3AFB4BFC7A1DF2" author="Moravec, J." box="[753,922,1185,1211]" pageId="13" pageNumber="258" refId="ref8440" refString="Moravec, J. (2007) Tiger beetles of Madagascar 1. A monograph of the genus Pogonostoma (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, 499 pp." type="book" year="2007">Moravec 2007</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C344BFE3CD39FFF0FF0CFB2EFBEF1F6B" pageId="13" pageNumber="258" type="distribution">
<paragraph id="8BE1EC68CD39FFF0FF0CFB2EFBEF1F6B" blockId="13.[151,1437,356,1751]" pageId="13" pageNumber="258">
<emphasis id="B92A307ACD39FFF0FF0CFB2EFE091D97" bold="true" box="[199,489,1220,1247]" pageId="13" pageNumber="258">Distribution and biology</emphasis>
.
<taxonomicName id="4C5E97EBCD39FFF0FE3DFB2FFCB21D96" authority="" authorityName="Jeannel" authorityYear="1946" box="[502,850,1220,1247]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="258" phylum="Arthropoda" rank="subGenus" subGenus="Bathypogonum">
<emphasis id="B92A307ACD39FFF0FE3DFB2FFD701D97" box="[502,656,1221,1246]" italics="true" pageId="13" pageNumber="258">Pogonostoma</emphasis>
(
<emphasis id="B92A307ACD39FFF0FD6BFB2EFCAA1D97" box="[672,842,1220,1246]" italics="true" pageId="13" pageNumber="258">Bathypogonum</emphasis>
)
</taxonomicName>
<emphasis id="B92A307ACD39FFF0FC93FB2EFC551D97" box="[856,949,1220,1246]" italics="true" pageId="13" pageNumber="258">l. lucens</emphasis>
is considered a geographical subspecies described by
<bibRefCitation id="EFCF9199CD39FFF0FEDBFB02FE5B1C4A" author="Rivalier, E." box="[272,443,1256,1283]" pageId="13" pageNumber="258" pagination="269 - 338" refId="ref9106" refString="Rivalier, E. (1970) Le genre Pogonostoma (Col. Cicindelidae). Revision avec description d'especes nouvelles. Annales de la Societe entomologique de France, New Series, 6, 269 - 338." type="journal article" year="1970">Rivalier (1970)</bibRefCitation>
from Sambirano. Besides the
<typeStatus id="54E552CACD39FFF0FCC0FB03FCDB1C4A" box="[779,827,1257,1283]" pageId="13" pageNumber="258">type</typeStatus>
locality, the western near-shore island of Nosy Berafia (= Île de Beraphia) of the Radama archipelago, it occurs also in the area of the Ramena river south-east of Ambanja, both localities belong to western Sambirano. One examined specimen is labelled “Bealanana”, which is a district in an inland northern
<collectingCountry id="F349ACF8CD39FFF0FEA2FABEFE121C27" box="[361,498,1364,1390]" name="Madagascar" pageId="13" pageNumber="258">Madagascar</collectingCountry>
, but its northern border is adjacent to the district of Ambanja; thus the specimen may come from a forest in northern areas of the district, in the past probably connected by a forest strip running from the area of Ambondrona (southern part of the large Tsaratanana massif, Sambirano). The labels “
<collectingRegion id="499A228ACD39FFF0FB3FFA76FA721CFF" box="[1268,1426,1436,1462]" country="Madagascar" name="Antananarivo" pageId="13" pageNumber="258">Antananarivo</collectingRegion>
” and “Tananarive” are either erroneous or more probably meaning “
<collectingCountry id="F349ACF8CD39FFF0FC4EFA2AFBF41C93" box="[901,1044,1472,1498]" name="Madagascar" pageId="13" pageNumber="258">Madagascar</collectingCountry>
”, as commonly labelled as such by insect dealers. The numerous specimens in MFNB, labelled “
<collectingCountry id="F349ACF8CD39FFF0FC9BFA0EFC3F1CB7" box="[848,991,1508,1534]" name="Madagascar" pageId="13" pageNumber="258">Madagascar</collectingCountry>
”, possess quite homogenous characters and belong probably to one population, although their exact origin is unknown.
</paragraph>
</subSubSection>
<subSubSection id="C344BFE3CD39FFF0FF0CF9C7FE001F9E" pageId="13" pageNumber="258" type="discussion">
<paragraph id="8BE1EC68CD39FFF0FF0CF9C7FE001F9E" blockId="13.[151,1437,356,1751]" pageId="13" pageNumber="258">
<emphasis id="B92A307ACD39FFF0FF0CF9C7FED41F0E" bold="true" box="[199,308,1581,1607]" pageId="13" pageNumber="258">Remarks</emphasis>
.
<bibRefCitation id="EFCF9199CD39FFF0FE89F9C6FE0D1F0E" author="Rivalier, E." box="[322,493,1580,1607]" pageId="13" pageNumber="258" pagination="269 - 338" refId="ref9106" refString="Rivalier, E. (1970) Le genre Pogonostoma (Col. Cicindelidae). Revision avec description d'especes nouvelles. Annales de la Societe entomologique de France, New Series, 6, 269 - 338." type="journal article" year="1970">Rivalier (1970)</bibRefCitation>
mentioned that he described this subspecies according to one male and two females. However, only the two females coming from the type locality were found in MNHN by the second author of the present paper (see
<bibRefCitation id="EFCF9199CD39FFF0FEA0F99FFDF31FC6" author="Moravec, J." box="[363,531,1653,1679]" pageId="13" pageNumber="258" refId="ref8440" refString="Moravec, J. (2007) Tiger beetles of Madagascar 1. A monograph of the genus Pogonostoma (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, 499 pp." type="book" year="2007">Moravec 2007</bibRefCitation>
); fortunately,
<bibRefCitation id="EFCF9199CD39FFF0FD64F99EFCBB1FC6" author="Rivalier, E." box="[687,859,1652,1679]" pageId="13" pageNumber="258" pagination="269 - 338" refId="ref9106" refString="Rivalier, E. (1970) Le genre Pogonostoma (Col. Cicindelidae). Revision avec description d'especes nouvelles. Annales de la Societe entomologique de France, New Series, 6, 269 - 338." type="journal article" year="1970">Rivalier (1970)</bibRefCitation>
mentioned that the aedeagus (of the missing male) was identical with that of the nominotypical subspecies. Consequently, one of the existing females was designated as
<typeStatus id="54E552CACD39FFF0FF7DF956FEFD1F9F" box="[182,285,1724,1750]" pageId="13" pageNumber="258" type="lectotype">lectotype</typeStatus>
(
<bibRefCitation id="EFCF9199CD39FFF0FEE0F957FE341F9E" author="Moravec, J." box="[299,468,1725,1751]" pageId="13" pageNumber="258" refId="ref8440" refString="Moravec, J. (2007) Tiger beetles of Madagascar 1. A monograph of the genus Pogonostoma (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, 499 pp." type="book" year="2007">Moravec 2007</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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