282 lines
36 KiB
XML
282 lines
36 KiB
XML
<document id="92304BCF9EACF58BDC3EB9BBA1FBCBCA" ID-CLB-Dataset="22840" ID-DOI="10.5281/zenodo.3763624" ID-GBIF-Dataset="31d8ae9c-f528-4f75-a558-65cbf715d338" ID-ISSN="0253-116X" ID-Zenodo-Dep="3763624" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587657676076" checkinUser="valdenar" docAuthor="Assing, Volker" docDate="2019" docId="03E9879FDE55FFF670C5FB3AFC1B3A3B" docLanguage="en" docName="Linzer.biol.Beitr.51.1.5-20.pdf.imf" docOrigin="Linzer biologische Beiträge 51 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Rugilus (Rugilus) asperrimus Assing, 2019, nov.nom." docType="treatment" docVersion="5" lastPageNumber="16" masterDocId="FFD0FFE7DE5CFFFD7077FFB2FFB23800" masterDocTitle="A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae)" masterLastPageNumber="20" masterPageNumber="5" pageNumber="14" updateTime="1698835682857" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="712E50A04993262329B58AE05512E0C4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="09BCDECE2945C54AE8F1348EC8BF17BD">
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<mods:title id="5E0FB13E60A95522D837D73398940794">A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae)</mods:title>
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<mods:name id="7E940B32BEF4862B3C69826E3F84A3EE" type="personal">
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<mods:namePart id="B6A6D966540D5D219ACBA9AF58C36A43">Assing, Volker</mods:namePart>
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<mods:title id="B1C3FDE7CA2FBA5FF3A67BF85C375D16">Linzer biologische Beiträge</mods:title>
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</mods:titleInfo>
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<mods:part id="52AA9D887CC3667317152E1D98527C11">
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<mods:date id="A8E48544A007471F7258E8AA95467C08">2019</mods:date>
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<mods:number id="43A8667660756186A37349F2AE094E8A">2019-07-26</mods:number>
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<mods:detail id="CBEF90BAB1783CCC18133DD28AD01879" type="volume">
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<mods:number id="1F8D02B3781095C85EBFF1584F388C33">51</mods:number>
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<mods:detail id="FC733289BB684C884428ECA89B9BE0A1" type="issue">
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<mods:number id="93EEE2D1A9C6071F94140E40CB102769">1</mods:number>
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<mods:start id="B46BDBED0F2AF9B35B66AC6AD506AB4C">5</mods:start>
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<mods:end id="453AC7445E4F967FBD438A131BDDB75C">20</mods:end>
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<mods:identifier id="143BFF3335670963716AC270033F9E7A" type="DOI">10.5281/zenodo.3763624</mods:identifier>
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<mods:identifier id="5C5D8F096AA04C64A872B0068AA05786" type="GBIF-Dataset">31d8ae9c-f528-4f75-a558-65cbf715d338</mods:identifier>
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<mods:identifier id="92542F8B7F975C864EB7BA5159881C32" type="ISSN">0253-116X</mods:identifier>
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<mods:identifier id="81E75F3D7B2E959D4D35029E7730FB36" type="Zenodo-Dep">3763624</mods:identifier>
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</mods:mods>
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<treatment id="03E9879FDE55FFF670C5FB3AFC1B3A3B" ID-GBIF-Taxon="163736951" LSID="urn:lsid:plazi:treatment:03E9879FDE55FFF670C5FB3AFC1B3A3B" httpUri="http://treatment.plazi.org/id/03E9879FDE55FFF670C5FB3AFC1B3A3B" lastPageId="11" lastPageNumber="16" pageId="9" pageNumber="14">
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<subSubSection id="C35A6502DE55FFF470C5FB3AFDE03C9C" box="[178,594,1159,1182]" pageId="9" pageNumber="14" type="nomenclature">
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<paragraph id="8BFF3689DE55FFF470C5FB3AFDE03C9C" blockId="9.[178,594,1159,1182]" box="[178,594,1159,1182]" pageId="9" pageNumber="14">
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<heading id="D0B781E5DE55FFF470C5FB3AFDE03C9C" box="[178,594,1159,1182]" fontSize="8" level="2" pageId="9" pageNumber="14" reason="2">
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<emphasis id="B934EA9BDE55FFF470C5FB3AFDE03C9C" bold="true" box="[178,594,1159,1182]" pageId="9" pageNumber="14">
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<taxonomicName id="4C404D0ADE55FFF470C5FB3AFE5A3C9D" ID-CoL="8Y97G" authorityName="Assing" authorityYear="2019" box="[178,488,1159,1182]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="14" phylum="Arthropoda" rank="species" species="asperrimus" status="nom. nov." subGenus="Rugilus">
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<emphasis id="B934EA9BDE55FFF470C5FB3AFEB13C9E" bold="true" box="[178,259,1160,1182]" italics="true" pageId="9" pageNumber="14">Rugilus</emphasis>
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(
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<emphasis id="B934EA9BDE55FFF47165FB3AFED63C9E" bold="true" box="[274,356,1160,1182]" italics="true" pageId="9" pageNumber="14">Rugilus</emphasis>
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)
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<emphasis id="B934EA9BDE55FFF47104FB3AFE5A3C9D" bold="true" box="[371,488,1160,1181]" italics="true" pageId="9" pageNumber="14">asperrimus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20757E0DE55FFF47199FB35FDE03C9C" box="[494,594,1159,1180]" pageId="9" pageNumber="14">nov.nom.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35A6502DE55FFF470C5FB1CFD043D21" pageId="9" pageNumber="14" type="reference_group">
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<paragraph id="8BFF3689DE55FFF470C5FB1CFD043CC3" blockId="9.[178,694,1197,1219]" box="[178,694,1197,1219]" pageId="9" pageNumber="14">
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<treatmentCitationGroup id="AB5011A7DE55FFF470C5FB1CFD043CC3" box="[178,694,1197,1219]" pageId="9" pageNumber="14">
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<emphasis id="B934EA9BDE55FFF470C5FB1CFEC13CC2" box="[178,371,1198,1218]" italics="true" pageId="9" pageNumber="14">
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<taxonomicName id="4C404D0ADE55FFF470C5FB1CFF453CC2" ID-CoL="8VS2Z" authorityName="Assing" authorityYear="2019" box="[178,247,1198,1218]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="14" phylum="Arthropoda" rank="genus">Rugilus</taxonomicName>
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rugosissimus
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</emphasis>
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<treatmentCitation id="0AE11098DE55FFF4710EFB1FFDA83CC2" author="ASSING V" box="[377,538,1197,1219]" page="78" pageId="9" pageNumber="14" year="2015">
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<bibRefCitation id="EFD14B78DE55FFF4710EFB1FFDA83CC2" author="ASSING V" box="[377,538,1197,1219]" pageId="9" pageNumber="14" pagination="73 - 82" refId="ref8701" refString="ASSING V. (2015): A revision of Palaearctic and Oriental Rugilus. V. Two new species from China and additional records (Coleoptera: Staphylinidae: Paederinae) (Insecta: Coleoptera). Linzer Biologische Beitrage 47 (1): 73 - 82." type="journal article" year="2015">ASSING, 2015: 78</bibRefCitation>
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</treatmentCitation>
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ff.; preoccupied.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BFF3689DE55FFF470C5FB63FD043D21" blockId="9.[178,1085,1233,1315]" pageId="9" pageNumber="14">
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C o m m e n t: Al Newton kindly drew my attention to the homonymy of
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<emphasis id="B934EA9BDE55FFF47398FB60FE973D05" italics="true" pageId="9" pageNumber="14">
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<taxonomicName id="4C404D0ADE55FFF47398FB60FB8F3CE7" ID-CoL="8VS2Z" authorityName="Assing" authorityYear="2019" box="[1007,1085,1234,1255]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="14" phylum="Arthropoda" rank="genus">Rugilus</taxonomicName>
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rugossimus
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</emphasis>
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<bibRefCitation id="EFD14B78DE55FFF4715BFB5DFE0B3D04" author="ASSING V" box="[300,441,1262,1284]" pageId="9" pageNumber="14" pagination="73 - 82" refId="ref8701" refString="ASSING V. (2015): A revision of Palaearctic and Oriental Rugilus. V. Two new species from China and additional records (Coleoptera: Staphylinidae: Paederinae) (Insecta: Coleoptera). Linzer Biologische Beitrage 47 (1): 73 - 82." type="journal article" year="2015">ASSING, 2015</bibRefCitation>
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with
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<emphasis id="B934EA9BDE55FFF47182FB42FD083D05" box="[501,698,1264,1285]" italics="true" pageId="9" pageNumber="14">Stilicus rugossimus</emphasis>
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FAGEL, 1953. The preoccupied name is here replaced with nomen novum
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<taxonomicName id="4C404D0ADE55FFF47255FABCFD023D22" ID-CoL="8Y97G" authorityName="Assing" authorityYear="2019" box="[546,688,1293,1315]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="14" phylum="Arthropoda" rank="species" species="asperrimus">
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<emphasis id="B934EA9BDE55FFF47255FABCFD023D22" box="[546,688,1293,1315]" italics="true" pageId="9" pageNumber="14">R. asperrimus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35A6502DE55FFF670C5FAFFFC1B3A3B" lastPageId="11" lastPageNumber="16" pageId="9" pageNumber="14" type="description">
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<paragraph id="8BFF3689DE55FFF470C5FAFFFB8F3D87" blockId="9.[178,726,1356,1376]" lastBlockId="9.[178,1085,1393,1483]" pageId="9" pageNumber="14">
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<emphasis id="B934EA9BDE55FFF470C5FAFFFD643D62" bold="true" box="[178,726,1356,1379]" pageId="9" pageNumber="14">
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<taxonomicName id="4C404D0ADE55FFF470C5FAFFFE193D63" authority="Assing, 2019" authorityName="Assing" authorityYear="2019" box="[178,427,1356,1379]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="14" phylum="Arthropoda" rank="species" species="gryps" status="sp. nov." subGenus="Rugilus">
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<emphasis id="B934EA9BDE55FFF470C5FAFFFEB13D63" bold="true" box="[178,259,1357,1379]" italics="true" pageId="9" pageNumber="14">Rugilus</emphasis>
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(
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<emphasis id="B934EA9BDE55FFF47165FAFFFED63D63" bold="true" box="[274,356,1357,1379]" italics="true" pageId="9" pageNumber="14">Rugilus</emphasis>
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)
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<emphasis id="B934EA9BDE55FFF47104FAFCFE193D63" bold="true" box="[371,427,1358,1379]" italics="true" pageId="9" pageNumber="14">gryps</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20757E0DE55FFF471C6FAFEFE4C3D61" box="[433,510,1356,1377]" pageId="9" pageNumber="14" rank="species">nov.sp.</taxonomicNameLabel>
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(
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<figureCitation id="137B2A0CDE55FFF4727CFAFEFDC93D61" box="[523,635,1356,1378]" captionStart-0="Figs 13-18" captionStart-1="Figs 19-20" captionStartId-0="5.[178,218,1474,1496]" captionStartId-1="6.[178,218,644,666]" captionTargetBox-0="[182,1038,207,1458]" captionTargetBox-1="[190,1074,225,617]" captionTargetId-0="figure@5.[178,1064,207,1463]" captionTargetId-1="figure@6.[180,1087,216,630]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Figs 13-18: Rugilus reuteri (13-15) and R. gryps (16-18): (13) male habitus; (14) female habitus; (15, 17-18) aedeagus in lateral and in ventral view; (16) forebody. Scale bars: 13-14, 16: 1.0 mm; 15, 17-18: 0.2 mm." captionText-1="Figs 19-20: Rugilus gryps: (19) male sternite VII; (20) male sternite VIII. Scale bar: 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3763628" figureDoi-1="http://doi.org/10.5281/zenodo.3763630" httpUri-0="https://zenodo.org/record/3763628/files/figure.png" httpUri-1="https://zenodo.org/record/3763630/files/figure.png" pageId="9" pageNumber="14">Figs 16-20</figureCitation>
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,
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<figureCitation id="137B2A0CDE55FFF472F0FAFEFD7F3D62" box="[647,717,1356,1378]" captionStart="Map 1" captionStartId="11.[178,223,1307,1328]" captionTargetBox="[178,1077,583,1292]" captionTargetId="figure@11.[178,1077,583,1293]" captionTargetPageId="11" captionText="Map 1: Distributions of the species of the Rugilus malaisei group: R. malaisei (black circle); R. desectus (white circle); R. gryps (black diamond); R. aquilinus (white diamond); R. asperrimus (black star); R. wuyicus (white star)." figureDoi="http://doi.org/10.5281/zenodo.3763636" httpUri="https://zenodo.org/record/3763636/files/figure.png" pageId="9" pageNumber="14">Map 1</figureCitation>
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)
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</emphasis>
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<materialsCitation id="3B283CD4DE55FFF470C5FAC3FB8F3D87" ID-GBIF-Occurrence="2608526785" box="[178,1085,1391,1415]" collectingDate="2014-06-17" collectingDateMax="2014-06-21" collectingDateMin="2014-06-17" country="China" elevation="1715" latitude="29.624166" location="Hailuogou valley" longLatPrecision="20" longitude="102.10777" municipality="Moxi" pageId="9" pageNumber="14" specimenCount="2" stateProvince="Sichuan Prov." typeStatus="holotype">
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T y p e m a t e r i a l:
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<typeStatus id="54FB882BDE55FFF471D9FAC3FDB03D87" box="[430,514,1393,1415]" pageId="9" pageNumber="14" type="holotype">Holotype</typeStatus>
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♂: ̎
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<collectingCountry id="F3577619DE55FFF47240FAC2FDC73D86" box="[567,629,1392,1414]" name="China" pageId="9" pageNumber="14">China</collectingCountry>
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:
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<collectingRegion id="4984F86BDE55FFF472F2FAC3FCBC3D86" box="[645,782,1393,1415]" country="China" name="Sichuan" pageId="9" pageNumber="14">Sichuan Prov.</collectingRegion>
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,
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<collectingMunicipality id="6B9BACF3DE55FFF4736AFAC3FCFC3D87" box="[797,846,1393,1415]" pageId="9" pageNumber="14">Moxi</collectingMunicipality>
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env.,
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<location id="8E9F6052DE55FFF473E3FAC3FB883D87" LSID="urn:lsid:plazi:treatment:03E9879FDE55FFF670C5FB3AFC1B3A3B:8E9F6052DE55FFF473E3FAC3FB883D87" box="[916,1082,1393,1415]" country="China" latitude="29.624166" longLatPrecision="20" longitude="102.10777" municipality="Moxi" name="Hailuogou valley" pageId="9" pageNumber="14" stateProvince="Sichuan Prov.">Hailuogou valley</location>
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,
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</materialsCitation>
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</paragraph>
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<paragraph id="8BFF3689DE55FFF470C5FA3AFD7B3DCB" blockId="9.[178,1085,1393,1483]" pageId="9" pageNumber="14">
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Gonghe vill.,
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<quantity id="4CB89B6CDE55FFF4714DFA3AFE343D9D" box="[314,390,1416,1438]" metricMagnitude="3" metricUnit="m" metricValue="1.7149999999999999" pageId="9" pageNumber="14" unit="m" value="1715.0">
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<elevation id="006DD1BADE55FFF4714DFA3AFE343D9D" box="[314,390,1416,1438]" metricMagnitude="3" metricUnit="m" metricValue="1.7149999999999999" pageId="9" pageNumber="14" unit="m" value="1715.0">1715 m</elevation>
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</quantity>
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,
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<geoCoordinate id="EE74504EDE55FFF471E2FA3AFDB23D9D" box="[405,512,1416,1437]" degrees="29" direction="north" minutes="37" orientation="latitude" pageId="9" pageNumber="14" precision="15" seconds="27" value="29.624166">29°37'27''N</geoCoordinate>
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,
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<geoCoordinate id="EE74504EDE55FFF47267FA3AFD363D9E" box="[528,644,1416,1438]" degrees="102" direction="east" minutes="06" orientation="longitude" pageId="9" pageNumber="14" precision="15" seconds="28" value="102.10777">102°06'28''E</geoCoordinate>
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, /
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<date id="FFFE1049DE55FFF472D2FA3AFC833D9D" box="[677,817,1416,1437]" pageId="9" pageNumber="14" value="2014-06-17" valueMax="2014-06-21" valueMin="2014-06-17">
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<collectingDate id="EFBAE9A1DE55FFF472D2FA3AFC833D9D" box="[677,817,1416,1437]" pageId="9" pageNumber="14" value="2014-06-17" valueMax="2014-06-21" valueMin="2014-06-17">17.-21.VI.2014</collectingDate>
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</date>
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, at light, in front of hotel, ruderals and gardens close to margin of mixed forest, J. Hảjek, J. Růžička & M. Tkoč leg. /
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<typeStatus id="54FB882BDE55FFF4747FFA2CFF513DCA" pageId="9" pageNumber="14" type="holotype">Holotypus</typeStatus>
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♂
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<taxonomicName id="4C404D0ADE55FFF47088FA04FECE3DCA" authority="Assing, 2019" authorityName="Assing" authorityYear="2019" box="[255,380,1462,1482]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="14" phylum="Arthropoda" rank="species" species="gryps" status="sp. nov.">
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<emphasis id="B934EA9BDE55FFF47088FA04FECE3DCA" box="[255,380,1462,1482]" italics="true" pageId="9" pageNumber="14">Rugilus gryps</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20757E0DE55FFF471F5FA07FE063DCA" box="[386,436,1461,1482]" pageId="9" pageNumber="14" rank="species">sp. n.</taxonomicNameLabel>
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, det. V.
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<bibRefCitation id="EFD14B78DE55FFF47276FA07FDC53DCA" author="ASSING V" box="[513,631,1460,1483]" pageId="9" pageNumber="14" pagination="73 - 82" refId="ref8701" refString="ASSING V. (2015): A revision of Palaearctic and Oriental Rugilus. V. Two new species from China and additional records (Coleoptera: Staphylinidae: Paederinae) (Insecta: Coleoptera). Linzer Biologische Beitrage 47 (1): 73 - 82." type="journal article" year="2015">Assing 2015</bibRefCitation>
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̎ (NMP).
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</paragraph>
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<paragraph id="8BFF3689DE55FFF470C5FA6AFE343E42" blockId="9.[178,1085,1496,1602]" pageId="9" pageNumber="14">E t y m o l o g y: The specific epithet (Latin: griffin, also griffon or gryphon, a mythical creature with the head and wings of an eagle) is a noun in apposition. It alludes to the shape of the ventral process of the aedeagus, which somewhat resembles the beak of an eagle in lateral view.</paragraph>
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<paragraph id="8BFF3689DE56FFF770C5FF68FC6A3944" blockId="10.[178,1086,218,324]" pageId="10" pageNumber="15">
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D e s c r i p t i o n: Body length
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<quantity id="4CB89B6CDE56FFF7725EFF68FDCD38EF" box="[553,639,218,240]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="10" pageNumber="15" unit="mm" value="6.2">6.2 mm</quantity>
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; length of forebody
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<quantity id="4CB89B6CDE56FFF77313FF68FC0838EF" box="[868,954,218,239]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="10" pageNumber="15" unit="mm" value="3.8">3.8 mm</quantity>
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. Coloration: head, pronotum, and abdomen blackish-brown; elytra dark-brown with slight bronze hue; legs dark-yellowish, with the apices of the meso- and metafemora indistinctly and narrowly darker; antennae pale-reddish, with antennomere I slightly darker red.
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</paragraph>
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<paragraph id="8BFF3689DE56FFF770C5FEE0FDB839F4" blockId="10.[178,1085,338,500]" pageId="10" pageNumber="15">
|
|
Head (
|
|
<figureCitation id="137B2A0CDE56FFF7708EFEE0FEF53968" box="[249,327,338,360]" captionStart="Figs 13-18" captionStartId="5.[178,218,1474,1496]" captionTargetBox="[182,1038,207,1458]" captionTargetId="figure@5.[178,1064,207,1463]" captionTargetPageId="5" captionText="Figs 13-18: Rugilus reuteri (13-15) and R. gryps (16-18): (13) male habitus; (14) female habitus; (15, 17-18) aedeagus in lateral and in ventral view; (16) forebody. Scale bars: 13-14, 16: 1.0 mm; 15, 17-18: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3763628" httpUri="https://zenodo.org/record/3763628/files/figure.png" pageId="10" pageNumber="15">Fig. 16</figureCitation>
|
|
) weakly transverse, 1.04 times as broad as long, broadest across eyes; margins behind eyes smoothly curving towards posterior constriction in dorsal view, posterior angles obsolete; punctation coarse, largely longitudinally confluent. Eyes large and bulging, approximately 0.7 times as long as distance from posterior margin of eyes to posterior constriction. Anterior margin of labrum with two basally fused teeth on either side of the median incision.
|
|
</paragraph>
|
|
<paragraph id="8BFF3689DE56FFF770C5FDB0FCED3A34" blockId="10.[178,1085,514,564]" pageId="10" pageNumber="15">
|
|
Pronotum (
|
|
<figureCitation id="137B2A0CDE56FFF77152FDB0FEDD3A18" box="[293,367,514,536]" captionStart="Figs 13-18" captionStartId="5.[178,218,1474,1496]" captionTargetBox="[182,1038,207,1458]" captionTargetId="figure@5.[178,1064,207,1463]" captionTargetPageId="5" captionText="Figs 13-18: Rugilus reuteri (13-15) and R. gryps (16-18): (13) male habitus; (14) female habitus; (15, 17-18) aedeagus in lateral and in ventral view; (16) forebody. Scale bars: 13-14, 16: 1.0 mm; 15, 17-18: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3763628" httpUri="https://zenodo.org/record/3763628/files/figure.png" pageId="10" pageNumber="15">Fig. 16</figureCitation>
|
|
) 1.15 times as long as broad and 0.75 times as wide as head; midline without impunctate, glossy band; punctation similar to that of head.
|
|
</paragraph>
|
|
<paragraph id="8BFF3689DE56FFF770C5FDF1FCCF3A93" blockId="10.[178,1086,579,660]" pageId="10" pageNumber="15">
|
|
Elytra (
|
|
<figureCitation id="137B2A0CDE56FFF77177FDF1FEFF3A59" box="[256,333,579,601]" captionStart="Figs 13-18" captionStartId="5.[178,218,1474,1496]" captionTargetBox="[182,1038,207,1458]" captionTargetId="figure@5.[178,1064,207,1463]" captionTargetPageId="5" captionText="Figs 13-18: Rugilus reuteri (13-15) and R. gryps (16-18): (13) male habitus; (14) female habitus; (15, 17-18) aedeagus in lateral and in ventral view; (16) forebody. Scale bars: 13-14, 16: 1.0 mm; 15, 17-18: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3763628" httpUri="https://zenodo.org/record/3763628/files/figure.png" pageId="10" pageNumber="15">Fig. 16</figureCitation>
|
|
) 0.97 times as long as pronotum and with pronounced humeral angles; punctation dense, moderately coarse, and defined; interstices glossy. Hind wings present. Metatarsomere I slightly longer than the combined length of II and III.
|
|
</paragraph>
|
|
<paragraph id="8BFF3689DE56FFF770C5FD11FC013B11" blockId="10.[178,1085,675,785]" pageId="10" pageNumber="15">Abdomen approximately 0.9 times as broad as elytra; tergites III-VI with moderately pronounced impressions anteriorly, these impressions with coarse, dense and irregular punctation; punctation of remaining tergal surfaces fine and dense; interstices without distinct microsculpture; posterior margin of tergite VII with palisade fringe.</paragraph>
|
|
<paragraph id="8BFF3689DE56FFF770C5FCACFB893BAB" blockId="10.[178,1085,798,940]" pageId="10" pageNumber="15">
|
|
♂: sternite VII (
|
|
<figureCitation id="137B2A0CDE56FFF7712CFC92FE153B36" box="[347,423,800,822]" captionStart="Figs 19-20" captionStartId="6.[178,218,644,666]" captionTargetBox="[190,1074,225,617]" captionTargetId="figure@6.[180,1087,216,630]" captionTargetPageId="6" captionText="Figs 19-20: Rugilus gryps: (19) male sternite VII; (20) male sternite VIII. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3763630" httpUri="https://zenodo.org/record/3763630/files/figure.png" pageId="10" pageNumber="15">Fig. 19</figureCitation>
|
|
) strongly transverse, approximately 1.4 times as broad as long, with shallow median impression, posterior margin with deep and broad excision, on either side of this excision acutely produced; sternite VIII (
|
|
<figureCitation id="137B2A0CDE56FFF7736DFCE9FCD53B71" box="[794,871,859,881]" captionStart="Figs 19-20" captionStartId="6.[178,218,644,666]" captionTargetBox="[190,1074,225,617]" captionTargetId="figure@6.[180,1087,216,630]" captionTargetPageId="6" captionText="Figs 19-20: Rugilus gryps: (19) male sternite VII; (20) male sternite VIII. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3763630" httpUri="https://zenodo.org/record/3763630/files/figure.png" pageId="10" pageNumber="15">Fig. 20</figureCitation>
|
|
) approximately 1.15 times as broad as long, posterior excision 0.2 times as deep as length of sternite; aedeagus (
|
|
<figureCitation id="137B2A0CDE56FFF7716BFC27FE393BAC" box="[284,395,917,940]" captionStart="Figs 13-18" captionStartId="5.[178,218,1474,1496]" captionTargetBox="[182,1038,207,1458]" captionTargetId="figure@5.[178,1064,207,1463]" captionTargetPageId="5" captionText="Figs 13-18: Rugilus reuteri (13-15) and R. gryps (16-18): (13) male habitus; (14) female habitus; (15, 17-18) aedeagus in lateral and in ventral view; (16) forebody. Scale bars: 13-14, 16: 1.0 mm; 15, 17-18: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3763628" httpUri="https://zenodo.org/record/3763628/files/figure.png" pageId="10" pageNumber="15">Figs 17-18</figureCitation>
|
|
) small in relation to body size,
|
|
<quantity id="4CB89B6CDE56FFF772BBFC27FCAB3BAB" box="[716,793,917,939]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="15" unit="mm" value="0.6">0.6 mm</quantity>
|
|
long, and of compact shape.
|
|
</paragraph>
|
|
<paragraph id="8BFF3689DE56FFF770C5FC09FE933CBB" blockId="10.[178,1085,955,1211]" pageId="10" pageNumber="15">
|
|
C o m p a r a t i v e n o t e s: Based on the similarly derived modifications of the male sternite VII (with large and nearly semi-circular posterior excision), the similarly derived morphology of the aedeagus (small, but robust; ventral process short and forming an angle with the basal portion; internal sac with large black sclerites), as well as the similar external characters,
|
|
<taxonomicName id="4C404D0ADE56FFF77265FB80FDDA3C47" authorityName="Assing" authorityYear="2019" box="[530,616,1074,1095]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="15" phylum="Arthropoda" rank="species" species="gryps">
|
|
<emphasis id="B934EA9BDE56FFF77265FB80FDDA3C47" box="[530,616,1074,1095]" italics="true" pageId="10" pageNumber="15">R. gryps</emphasis>
|
|
</taxonomicName>
|
|
belongs to the
|
|
<emphasis id="B934EA9BDE56FFF7737EFB80FCCF3C46" box="[777,893,1073,1095]" italics="true" pageId="10" pageNumber="15">R. malaisei</emphasis>
|
|
group, which previously included five species:
|
|
<taxonomicName id="4C404D0ADE56FFF77185FBFDFD363C64" authorityName="Assing" authorityYear="2019" box="[498,644,1103,1124]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="15" phylum="Arthropoda" rank="species" species="asperrimus">
|
|
<emphasis id="B934EA9BDE56FFF77185FBFDFD363C64" box="[498,644,1103,1124]" italics="true" pageId="10" pageNumber="15">R. asperrimus</emphasis>
|
|
</taxonomicName>
|
|
ASSING, nov.nom. (Guizhou),
|
|
<taxonomicName id="4C404D0ADE56FFF773BCFBFDFB8F3C64" authorityName="Assing" authorityYear="2012" box="[971,1085,1103,1124]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="15" phylum="Arthropoda" rank="species" species="wuyicus">
|
|
<emphasis id="B934EA9BDE56FFF773BCFBFDFB8F3C64" box="[971,1085,1103,1124]" italics="true" pageId="10" pageNumber="15">R. wuyicus</emphasis>
|
|
</taxonomicName>
|
|
ASSING, 2012 (Jiangxi),
|
|
<taxonomicName id="4C404D0ADE56FFF771D9FBDFFD903C81" authorityName="Assing" authorityYear="2012" box="[430,546,1132,1154]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="15" phylum="Arthropoda" rank="species" species="desectus">
|
|
<emphasis id="B934EA9BDE56FFF771D9FBDFFD903C81" box="[430,546,1132,1154]" italics="true" pageId="10" pageNumber="15">R. desectus</emphasis>
|
|
</taxonomicName>
|
|
ASSING, 2012 (Yunnan),
|
|
<emphasis id="B934EA9BDE56FFF7735BFBDFFC1A3C81" box="[812,936,1132,1154]" italics="true" pageId="10" pageNumber="15">R. aquilinus</emphasis>
|
|
ASSING, 2012 (Sichuan), and
|
|
<emphasis id="B934EA9BDE56FFF77138FB38FE703C9E" box="[335,450,1161,1183]" italics="true" pageId="10" pageNumber="15">R. malaisei</emphasis>
|
|
(SCHEERPELTZ, 1965) (
|
|
<collectingCountry id="F3577619DE56FFF772C1FB3AFCB33C9E" box="[694,769,1160,1182]" name="Myanmar" pageId="10" pageNumber="15">Burma</collectingCountry>
|
|
). It is distinguished from them as follows:
|
|
</paragraph>
|
|
<paragraph id="8BFF3689DE56FFF770C5FB79FF403D73" blockId="10.[178,1086,1227,1395]" pageId="10" pageNumber="15">
|
|
from
|
|
<taxonomicName id="4C404D0ADE56FFF7709DFB7FFECA3CE2" authorityName="Assing" authorityYear="2019" box="[234,376,1229,1250]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="15" phylum="Arthropoda" rank="species" species="asperrimus">
|
|
<emphasis id="B934EA9BDE56FFF7709DFB7FFECA3CE2" box="[234,376,1229,1250]" italics="true" pageId="10" pageNumber="15">R. asperrimus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C404D0ADE56FFF771DCFB7FFDAB3CE2" authorityName="Assing" authorityYear="2012" box="[427,537,1229,1250]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="15" phylum="Arthropoda" rank="species" species="wuyicus">
|
|
<emphasis id="B934EA9BDE56FFF771DCFB7FFDAB3CE2" box="[427,537,1229,1250]" italics="true" pageId="10" pageNumber="15">R. wuyicus</emphasis>
|
|
</taxonomicName>
|
|
by smaller size, a more slender body, paler antennae, a much less transverse head, much longer elytra, the presence of hind wings, the presence of a palisade fringe at the posterior margin of tergite VII, a deeper posterior excision of the less transverse male sternite VII and more acute projections on either side of this excision, and a slightly smaller aedeagus with a ventral proces of slightly different shape,
|
|
</paragraph>
|
|
<paragraph id="8BFF3689DE56FFF770C5FA31FEAE3DB6" blockId="10.[178,1085,1411,1462]" pageId="10" pageNumber="15">
|
|
from
|
|
<taxonomicName id="4C404D0ADE56FFF77085FA37FED83D9A" authorityName="Assing" authorityYear="2012" box="[242,362,1413,1434]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="15" phylum="Arthropoda" rank="species" species="wuyicus">
|
|
<emphasis id="B934EA9BDE56FFF77085FA37FED83D9A" box="[242,362,1413,1434]" italics="true" pageId="10" pageNumber="15">R. wuyicus</emphasis>
|
|
</taxonomicName>
|
|
additionally by the more confluent punctation of the head and pronotum;
|
|
</paragraph>
|
|
<paragraph id="8BFF3689DE56FFF770C5FA77FE893E33" blockId="10.[178,1085,1477,1587]" pageId="10" pageNumber="15">
|
|
from
|
|
<emphasis id="B934EA9BDE56FFF7709DFA75FEEE3DDC" box="[234,348,1479,1500]" italics="true" pageId="10" pageNumber="15">R. malaisei</emphasis>
|
|
by smaller size, a more slender head with larger and more bulging eyes, a more slender pronotum, a less transverse male sternite VII with a deeper posterior excision of different shape, and by the slightly different shape of the ventral process of the aedeagus;
|
|
</paragraph>
|
|
<paragraph id="8BFF3689DE57FFF670C5FF68FCF93928" blockId="11.[178,1085,218,296]" pageId="11" pageNumber="16">
|
|
from
|
|
<taxonomicName id="4C404D0ADE57FFF67099FF69FED738F0" authorityName="Assing" authorityYear="2012" box="[238,357,219,240]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="16" phylum="Arthropoda" rank="species" species="desectus">
|
|
<emphasis id="B934EA9BDE57FFF67099FF69FED738F0" box="[238,357,219,240]" italics="true" pageId="11" pageNumber="16">R. desectus</emphasis>
|
|
</taxonomicName>
|
|
by a slightly smaller body with a less transverse head and a more slender pronotum, a less deep posterior excision of the male sternite VII, and the completely different shape of the ventral process of the aedeagus;
|
|
</paragraph>
|
|
<paragraph id="8BFF3689DE57FFF670C5FE84FB883984" blockId="11.[178,1085,310,388]" pageId="11" pageNumber="16">
|
|
from
|
|
<emphasis id="B934EA9BDE57FFF6709DFE85FED6394C" box="[234,356,311,332]" italics="true" pageId="11" pageNumber="16">R. aquilinus</emphasis>
|
|
by somewhat paler legs and antennae, a less transverse head, relatively longer elytra, a less transverse male sternite VII with a deeper posterior excision of different shape, and by the slightly different shape of the ventral process of the aedeagus.
|
|
</paragraph>
|
|
<paragraph id="8BFF3689DE57FFF670C5FE20FC3239DF" blockId="11.[178,1085,402,480]" pageId="11" pageNumber="16">
|
|
For illustrations of the external and male sexual characters of
|
|
<taxonomicName id="4C404D0ADE57FFF6735CFE21FC0839A8" authorityName="Assing" authorityYear="2019" box="[811,954,403,424]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="16" phylum="Arthropoda" rank="species" species="asperrimus">
|
|
<emphasis id="B934EA9BDE57FFF6735CFE21FC0839A8" box="[811,954,403,424]" italics="true" pageId="11" pageNumber="16">R. asperrimus</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C404D0ADE57FFF673BFFE21FB8539A8" authorityName="Assing" authorityYear="2012" box="[968,1079,403,424]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="16" phylum="Arthropoda" rank="species" species="wuyicus">
|
|
<emphasis id="B934EA9BDE57FFF673BFFE21FB8539A8" box="[968,1079,403,424]" italics="true" pageId="11" pageNumber="16">R. wuyicus</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<emphasis id="B934EA9BDE57FFF670C5FE1DFE9C39C4" box="[178,302,431,452]" italics="true" pageId="11" pageNumber="16">R. aquilinus</emphasis>
|
|
,
|
|
<taxonomicName id="4C404D0ADE57FFF6714BFE1DFE0339C4" authorityName="Assing" authorityYear="2012" box="[316,433,431,452]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="16" phylum="Arthropoda" rank="species" species="desectus">
|
|
<emphasis id="B934EA9BDE57FFF6714BFE1DFE0339C4" box="[316,433,431,452]" italics="true" pageId="11" pageNumber="16">R. desectus</emphasis>
|
|
</taxonomicName>
|
|
, and
|
|
<emphasis id="B934EA9BDE57FFF6719BFE1DFDD239C4" box="[492,608,431,452]" italics="true" pageId="11" pageNumber="16">R. malaisei</emphasis>
|
|
see
|
|
<bibRefCitation id="EFD14B78DE57FFF672E7FE1CFC9F39C3" author="ASSING V" box="[656,813,430,451]" pageId="11" pageNumber="16" pagination="115 - 190" refId="ref8522" refString="ASSING V. (2012 a): The Rugilus species of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Paederinae). 2 Stuttgarter Beitrage zur Naturkun de A, Neue Serie 5: 115 - 190." type="journal article" year="2012">ASSING (2012a</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="EFD14B78DE57FFF6734EFE1CFCC939C4" author="ASSING V" box="[825,891,430,452]" pageId="11" pageNumber="16" pagination="137 - 149" refId="ref8563" refString="ASSING V. (2012 b): A revision of Palaearctic and Oriental Rugilus LEACH, 1819. II. Three new species from China and additional records (Coleoptera: Staphylinidae: Paederinae). 2 Koleopterologische Rundschau 82: 137 - 149." type="journal article" year="2012">2012b</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="EFD14B78DE57FFF673F0FE1FFC0C39C3" author="ASSING V" box="[903,958,429,451]" pageId="11" pageNumber="16" pagination="73 - 82" refId="ref8701" refString="ASSING V. (2015): A revision of Palaearctic and Oriental Rugilus. V. Two new species from China and additional records (Coleoptera: Staphylinidae: Paederinae) (Insecta: Coleoptera). Linzer Biologische Beitrage 47 (1): 73 - 82." type="journal article" year="2015">2015</bibRefCitation>
|
|
). The distributions of the species of the
|
|
<emphasis id="B934EA9BDE57FFF671A3FE79FDF439E0" box="[468,582,459,480]" italics="true" pageId="11" pageNumber="16">R. malaisei</emphasis>
|
|
group are illustrated in
|
|
<figureCitation id="137B2A0CDE57FFF6734FFE78FC3239DF" box="[824,896,458,479]" captionStart="Map 1" captionStartId="11.[178,223,1307,1328]" captionTargetBox="[178,1077,583,1292]" captionTargetId="figure@11.[178,1077,583,1293]" captionTargetPageId="11" captionText="Map 1: Distributions of the species of the Rugilus malaisei group: R. malaisei (black circle); R. desectus (white circle); R. gryps (black diamond); R. aquilinus (white diamond); R. asperrimus (black star); R. wuyicus (white star)." figureDoi="http://doi.org/10.5281/zenodo.3763636" httpUri="https://zenodo.org/record/3763636/files/figure.png" pageId="11" pageNumber="16">Map 1.</figureCitation>
|
|
</paragraph>
|
|
<paragraph id="8BFF3689DE57FFF670C5FE5CFC1B3A3B" blockId="11.[178,1085,494,572]" pageId="11" pageNumber="16">
|
|
D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: The type locality is situated to the southeast of the Gongga Shan in
|
|
<collectingRegion id="4984F86BDE57FFF67251FDB8FDC83A20" box="[550,634,522,544]" country="China" name="Sichuan" pageId="11" pageNumber="16">Sichuan</collectingRegion>
|
|
,
|
|
<collectingCountry id="F3577619DE57FFF672F2FDB8FD703A20" box="[645,706,522,544]" name="China" pageId="11" pageNumber="16">China</collectingCountry>
|
|
(
|
|
<figureCitation id="137B2A0CDE57FFF672B9FDB8FCA33A20" box="[718,785,522,544]" captionStart="Map 1" captionStartId="11.[178,223,1307,1328]" captionTargetBox="[178,1077,583,1292]" captionTargetId="figure@11.[178,1077,583,1293]" captionTargetPageId="11" captionText="Map 1: Distributions of the species of the Rugilus malaisei group: R. malaisei (black circle); R. desectus (white circle); R. gryps (black diamond); R. aquilinus (white diamond); R. asperrimus (black star); R. wuyicus (white star)." figureDoi="http://doi.org/10.5281/zenodo.3763636" httpUri="https://zenodo.org/record/3763636/files/figure.png" pageId="11" pageNumber="16">Map 1</figureCitation>
|
|
). The
|
|
<typeStatus id="54FB882BDE57FFF67323FDB8FC1F3A20" box="[852,941,522,544]" pageId="11" pageNumber="16" type="holotype">holotype</typeStatus>
|
|
was collected with a light trap near the margin of a mixed forest at an altitude of
|
|
<quantity id="4CB89B6CDE57FFF67321FD97FC173A3B" box="[854,933,549,571]" metricMagnitude="3" metricUnit="m" metricValue="1.7149999999999999" pageId="11" pageNumber="16" unit="m" value="1715.0">1715 m</quantity>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |