A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae)
Author
Assing, Volker
text
Linzer biologische Beiträge
2019
2019-07-26
51
1
5
20
journal article
22840
10.5281/zenodo.3763624
31d8ae9c-f528-4f75-a558-65cbf715d338
0253-116X
3763624
Rugilus
(
Rugilus
)
asperrimus
nov.nom.
Rugilus
rugosissimus
ASSING, 2015: 78
ff.; preoccupied.
C o m m e n t: Al Newton kindly drew my attention to the homonymy of
Rugilus
rugossimus
ASSING, 2015
with
Stilicus rugossimus
FAGEL, 1953. The preoccupied name is here replaced with nomen novum
R. asperrimus
.
Rugilus
(
Rugilus
)
gryps
nov.sp.
(
Figs 16-20
,
Map 1
)
T y p e m a t e r i a l:
Holotype
♂: ̎
China
:
Sichuan Prov.
,
Moxi
env.,
Hailuogou valley
,
Gonghe vill.,
1715 m
,
29°37'27''N
,
102°06'28''E
, /
17.-21.VI.2014
, at light, in front of hotel, ruderals and gardens close to margin of mixed forest, J. Hảjek, J. Růžička & M. Tkoč leg. /
Holotypus
♂
Rugilus gryps
sp. n.
, det. V.
Assing 2015
̎ (NMP).
E t y m o l o g y: The specific epithet (Latin: griffin, also griffon or gryphon, a mythical creature with the head and wings of an eagle) is a noun in apposition. It alludes to the shape of the ventral process of the aedeagus, which somewhat resembles the beak of an eagle in lateral view.
D e s c r i p t i o n: Body length
6.2 mm
; length of forebody
3.8 mm
. Coloration: head, pronotum, and abdomen blackish-brown; elytra dark-brown with slight bronze hue; legs dark-yellowish, with the apices of the meso- and metafemora indistinctly and narrowly darker; antennae pale-reddish, with antennomere I slightly darker red.
Head (
Fig. 16
) weakly transverse, 1.04 times as broad as long, broadest across eyes; margins behind eyes smoothly curving towards posterior constriction in dorsal view, posterior angles obsolete; punctation coarse, largely longitudinally confluent. Eyes large and bulging, approximately 0.7 times as long as distance from posterior margin of eyes to posterior constriction. Anterior margin of labrum with two basally fused teeth on either side of the median incision.
Pronotum (
Fig. 16
) 1.15 times as long as broad and 0.75 times as wide as head; midline without impunctate, glossy band; punctation similar to that of head.
Elytra (
Fig. 16
) 0.97 times as long as pronotum and with pronounced humeral angles; punctation dense, moderately coarse, and defined; interstices glossy. Hind wings present. Metatarsomere I slightly longer than the combined length of II and III.
Abdomen approximately 0.9 times as broad as elytra; tergites III-VI with moderately pronounced impressions anteriorly, these impressions with coarse, dense and irregular punctation; punctation of remaining tergal surfaces fine and dense; interstices without distinct microsculpture; posterior margin of tergite VII with palisade fringe.
♂: sternite VII (
Fig. 19
) strongly transverse, approximately 1.4 times as broad as long, with shallow median impression, posterior margin with deep and broad excision, on either side of this excision acutely produced; sternite VIII (
Fig. 20
) approximately 1.15 times as broad as long, posterior excision 0.2 times as deep as length of sternite; aedeagus (
Figs 17-18
) small in relation to body size,
0.6 mm
long, and of compact shape.
C o m p a r a t i v e n o t e s: Based on the similarly derived modifications of the male sternite VII (with large and nearly semi-circular posterior excision), the similarly derived morphology of the aedeagus (small, but robust; ventral process short and forming an angle with the basal portion; internal sac with large black sclerites), as well as the similar external characters,
R. gryps
belongs to the
R. malaisei
group, which previously included five species:
R. asperrimus
ASSING, nov.nom. (Guizhou),
R. wuyicus
ASSING, 2012 (Jiangxi),
R. desectus
ASSING, 2012 (Yunnan),
R. aquilinus
ASSING, 2012 (Sichuan), and
R. malaisei
(SCHEERPELTZ, 1965) (
Burma
). It is distinguished from them as follows:
from
R. asperrimus
and
R. wuyicus
by smaller size, a more slender body, paler antennae, a much less transverse head, much longer elytra, the presence of hind wings, the presence of a palisade fringe at the posterior margin of tergite VII, a deeper posterior excision of the less transverse male sternite VII and more acute projections on either side of this excision, and a slightly smaller aedeagus with a ventral proces of slightly different shape,
from
R. wuyicus
additionally by the more confluent punctation of the head and pronotum;
from
R. malaisei
by smaller size, a more slender head with larger and more bulging eyes, a more slender pronotum, a less transverse male sternite VII with a deeper posterior excision of different shape, and by the slightly different shape of the ventral process of the aedeagus;
from
R. desectus
by a slightly smaller body with a less transverse head and a more slender pronotum, a less deep posterior excision of the male sternite VII, and the completely different shape of the ventral process of the aedeagus;
from
R. aquilinus
by somewhat paler legs and antennae, a less transverse head, relatively longer elytra, a less transverse male sternite VII with a deeper posterior excision of different shape, and by the slightly different shape of the ventral process of the aedeagus.
For illustrations of the external and male sexual characters of
R. asperrimus
,
R. wuyicus
,
R. aquilinus
,
R. desectus
, and
R. malaisei
see
ASSING (2012a
,
2012b
,
2015
). The distributions of the species of the
R. malaisei
group are illustrated in
Map 1.
D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: The type locality is situated to the southeast of the Gongga Shan in
Sichuan
,
China
(
Map 1
). The
holotype
was collected with a light trap near the margin of a mixed forest at an altitude of
1715 m
.