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<document id="AA492552824B109CFB2A6FBF1059BA22" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629612930141" checkinUser="felipe" docAuthor="Zeidler, Wolfgang" docDate="2009" docId="03C89422FF91DD6934C7FBF1FF65FE32" docLanguage="en" docName="zt02000p117.pdf" docOrigin="Zootaxa 2000" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Prolanceolidae Zeidler 2009, new family" docType="treatment" docVersion="3" lastPageNumber="65" masterDocId="FFF1EC5AFFD1DD283450FFE3FFB2FF90" masterDocTitle="A review of the hyperiidean amphipod superfamily Lanceoloidea Bowman &amp; Gruner, 1973 (Crustacea: Amphipoda: Hyperiidea)" masterLastPageNumber="117" masterPageNumber="1" pageNumber="65" updateTime="1699083125402" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="A32C6B736EE8B6C60715AE50D3E6329C">A review of the hyperiidean amphipod superfamily Lanceoloidea Bowman &amp; Gruner, 1973 (Crustacea: Amphipoda: Hyperiidea)</mods:title>
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<mods:namePart id="075DCCB0B63C3F204132FA06084B2B9F">Zeidler, Wolfgang</mods:namePart>
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<mods:date id="7DF909E8AD8A3D2038228E0ED6641B0D">2009</mods:date>
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<treatment id="03C89422FF91DD6934C7FBF1FF65FE32" ID-DOI="http://doi.org/10.5281/zenodo.5328090" ID-Zenodo-Dep="5328090" LSID="urn:lsid:plazi:treatment:03C89422FF91DD6934C7FBF1FF65FE32" httpUri="http://treatment.plazi.org/id/03C89422FF91DD6934C7FBF1FF65FE32" lastPageId="65" lastPageNumber="65" pageId="64" pageNumber="65">
<subSubSection id="C37B76BFFF91DD6834C7FBF1FD14FBBC" box="[151,678,1042,1068]" pageId="64" pageNumber="65" type="nomenclature">
<paragraph id="8BDE2534FF91DD6834C7FBF1FD14FBBC" blockId="64.[151,678,1042,1068]" box="[151,678,1042,1068]" pageId="64" pageNumber="65">
<heading id="D0969258FF91DD6834C7FBF1FD14FBBC" bold="true" box="[151,678,1042,1068]" fontSize="11" level="1" pageId="64" pageNumber="65" reason="1">
<emphasis id="B915F926FF91DD6834C7FBF1FD14FBBC" bold="true" box="[151,678,1042,1068]" pageId="64" pageNumber="65">
Family
<taxonomicName id="4C615EB7FF91DD6834A6FBF1FDA2FBBC" ID-CoL="7NKMZ" authority="Zeidler, 2009" authorityName="Zeidler" authorityYear="2009" box="[246,528,1042,1068]" class="Malacostraca" family="Prolanceolidae" kingdom="Animalia" order="Amphipoda" pageId="64" pageNumber="65" phylum="Arthropoda" rank="family" status="fam. nov.">PROLANCEOLIDAE</taxonomicName>
,
<taxonomicNameLabel id="A226445DFF91DD68364CFBF1FD14FBBC" box="[540,678,1042,1068]" pageId="64" pageNumber="65" rank="family">new family</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37B76BFFF91DD6834C7FBBEFE58F95E" pageId="64" pageNumber="65" type="diagnosis">
<paragraph id="8BDE2534FF91DD6834C7FBBEFEF8F937" blockId="64.[151,1436,1117,2023]" pageId="64" pageNumber="65">
<emphasis id="B915F926FF91DD6834C7FBBEFEA5FBE7" bold="true" box="[151,279,1117,1143]" pageId="64" pageNumber="65">Diagnosis.</emphasis>
Body length up to
<quantity id="4C9988D1FF91DD683659FBBEFDD0FBE7" box="[521,610,1117,1143]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="64" pageNumber="65" unit="mm" value="14.0">14 mm</quantity>
; pereon not inflated. Head without distinct rostrum. Eyes consist of chain of five ocelli on either side of head. Pereonites all separate. Coxae separate from pereonites. Antennae 1 with stout callynophore, peduncle 3-articulate. Antennae 2 longer than A1. Mandibles with strong palp, length almost 1.5x body length of mandible; third article of palp longer than first and second combined; left with broad lacinia mobilis. Maxillae 1 with broad palp, armed with few setae distally; outer lobe relatively slender, with five strong setae distally; inner lobe rounded, armed with numerous fine setae. Maxillae 2 with relatively broad lobes, the inner slightly shorter than outer one; both armed with row of strong setae together with more slender ones on inner and distal margin. Maxilliped with semicircular outer lobes, with row of fine setae along inner margin, slightly overlapping each other; inner lobes rounded, length 0.4x outer lobes, separate to base, not projected laterally from outer lobes as in other lanceolids. Gnathopod 1 subchelate, with distally expanded carpus and propodus. Gnathopod 2 simple, a little longer and more slender than G1. Pereopods 37 simple. Pereopods 6 &amp; 7 with small retractile dactyls, without spoon-shaped process. Urosomite 23 with partial suture ventrally. Uropoda relatively broad (except first), with articulated exopoda and endopoda. Telson slightly longer than peduncle of U3, very elongate with pointed apex. Gills on pereonites 26. Oostegites on pereonites 25.
</paragraph>
<paragraph id="8BDE2534FF91DD683497F956FE58F95E" blockId="64.[151,1436,1117,2023]" box="[199,490,1717,1743]" pageId="64" pageNumber="65">
One genus:
<emphasis id="B915F926FF91DD683503F956FE58F95E" box="[339,490,1717,1742]" italics="true" pageId="64" pageNumber="65">
<taxonomicName id="4C615EB7FF91DD683503F956FE54F95E" authorityName="Woltereck" authorityYear="1907" box="[339,486,1717,1742]" class="Malacostraca" family="Prolanceolidae" genus="Prolanceola" kingdom="Animalia" order="Amphipoda" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">Prolanceola</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C37B76BFFF91DD693497F93EFF65FE32" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65" type="discussion">
<paragraph id="8BDE2534FF91DD693497F93EFF65FE32" blockId="64.[151,1436,1117,2023]" lastBlockId="65.[151,1436,152,418]" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65">
<emphasis id="B915F926FF91DD683497F93EFEF2F967" bold="true" box="[199,320,1757,1783]" pageId="64" pageNumber="65">Remarks.</emphasis>
This new family is proposed here to accommodate
<taxonomicName id="4C615EB7FF91DD6837F2F93EFB80F966" authorityName="Woltereck" authorityYear="1907" box="[930,1074,1757,1782]" class="Malacostraca" family="Prolanceolidae" genus="Prolanceola" kingdom="Animalia" order="Amphipoda" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="B915F926FF91DD6837F2F93EFB80F966" box="[930,1074,1757,1782]" italics="true" pageId="64" pageNumber="65">Prolanceola</emphasis>
</taxonomicName>
which, prior to this study, was placed in the family
<taxonomicName id="4C615EB7FF91DD6835C0F8E6FD91F88F" authorityName="Bovallius" authorityYear="1887" box="[400,547,1797,1823]" class="Malacostraca" family="Lanceolidae" kingdom="Animalia" order="Amphipoda" pageId="64" pageNumber="65" phylum="Arthropoda" rank="family">Lanceolidae</taxonomicName>
.
<bibRefCitation id="EFF058C5FF91DD683660F8E6FD4DF88F" author="Woltereck, R." box="[560,767,1797,1823]" pageId="64" pageNumber="65" pagination="129 - 132" refId="ref76604" refString="Woltereck, R. (1907) Siebente Mitteilung uber die Valdivia-Hyperiden, Prolanceola vibiliformis n. gen. n. sp. Zoologischer Anzeiger 31 (5 / 6), 129 - 132." type="journal article" year="1907">Woltereck (1907)</bibRefCitation>
regarded
<taxonomicName id="4C615EB7FF91DD683729F8E6FBBBF88E" authorityName="Woltereck" authorityYear="1907" box="[889,1033,1797,1822]" class="Malacostraca" family="Prolanceolidae" genus="Prolanceola" kingdom="Animalia" order="Amphipoda" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="B915F926FF91DD683729F8E6FBBBF88E" box="[889,1033,1797,1822]" italics="true" pageId="64" pageNumber="65">Prolanceola</emphasis>
</taxonomicName>
to be a link between the families
<taxonomicName id="4C615EB7FF91DD6834C7F8CEFEB8F8D7" authorityName="Dana" authorityYear="1852" box="[151,266,1837,1863]" class="Malacostraca" family="Vibiliidae" kingdom="Animalia" order="Amphipoda" pageId="64" pageNumber="65" phylum="Arthropoda" rank="family">Vibiliidae</taxonomicName>
and
<taxonomicName id="4C615EB7FF91DD683512F8CEFE63F8D7" authorityName="Bovallius" authorityYear="1887" box="[322,465,1837,1863]" class="Malacostraca" family="Lanceolidae" kingdom="Animalia" order="Amphipoda" pageId="64" pageNumber="65" phylum="Arthropoda" rank="family">Lanceolidae</taxonomicName>
and later (1909) he placed it in the
<taxonomicName id="4C615EB7FF91DD68373EF8CEFBB2F8D7" authorityName="Bovallius" authorityYear="1887" box="[878,1024,1837,1863]" class="Malacostraca" family="Lanceolidae" kingdom="Animalia" order="Amphipoda" pageId="64" pageNumber="65" phylum="Arthropoda" rank="family">Lanceolidae</taxonomicName>
, presumably based on the retractile dactyls of pereopods 6 &amp; 7. However, this placement is not satisfactory because
<taxonomicName id="4C615EB7FF91DD683009F8B6FB5BF8FE" authorityName="Woltereck" authorityYear="1907" box="[1113,1257,1877,1902]" class="Malacostraca" family="Prolanceolidae" genus="Prolanceola" kingdom="Animalia" order="Amphipoda" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="B915F926FF91DD683009F8B6FB5BF8FE" box="[1113,1257,1877,1902]" italics="true" pageId="64" pageNumber="65">Prolanceola</emphasis>
</taxonomicName>
possesses several unique characters that distinguish it from all other families of
<taxonomicName id="4C615EB7FF91DD6837E5F89EFB89F807" box="[949,1083,1917,1943]" pageId="64" pageNumber="116" rank="tribe" tribe="Hyperiidea">Hyperiidea</taxonomicName>
. The chelate first gnathopoda and the structure of the eyes and mouthparts, especially the mandibles and second maxillae are particularly distinctive characters that distinguish
<taxonomicName id="4C615EB7FF91DD683609F82EFD5BF876" authorityName="Woltereck" authorityYear="1907" box="[601,745,1997,2022]" class="Malacostraca" family="Prolanceolidae" genus="Prolanceola" kingdom="Animalia" order="Amphipoda" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="B915F926FF91DD683609F82EFD5BF876" box="[601,745,1997,2022]" italics="true" pageId="64" pageNumber="65">Prolanceola</emphasis>
</taxonomicName>
and the family
<taxonomicName id="4C615EB7FF91DD6837F9F82EFBE9F877" authorityName="Zeidler" authorityYear="2009" box="[937,1115,1997,2023]" class="Malacostraca" family="Prolanceolidae" kingdom="Animalia" order="Amphipoda" pageId="64" pageNumber="65" phylum="Arthropoda" rank="family">Prolanceolidae</taxonomicName>
. It resembles
<taxonomicName id="4C615EB7FF91DD683151F82EFAFDF876" authorityName="H. Milne Edwards" authorityYear="1830" box="[1281,1359,1997,2022]" class="Malacostraca" family="Vibiliidae" genus="Vibilia" kingdom="Animalia" order="Amphipoda" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="B915F926FF91DD683151F82EFAFDF876" box="[1281,1359,1997,2022]" italics="true" pageId="64" pageNumber="65">Vibilia</emphasis>
</taxonomicName>
in the general habitus and also the arrangement of the ocelli of the eyes is similar to that of
<taxonomicName id="4C615EB7FF90DD6930CEFF7BFF63FF4A" authority="Stebbing, 1888" authorityName="Stebbing" authorityYear="1888" class="Malacostraca" family="Lanceolidae" genus="Aurora" kingdom="Animalia" order="Amphipoda" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="B915F926FF90DD6930CEFF7BFA91FF21" box="[1182,1315,152,177]" italics="true" pageId="65" pageNumber="66">V. australis</emphasis>
<bibRefCitation id="EFF058C5FF90DD69317DFF7BFF63FF4A" author="Stebbing, T. R. R." pageId="65" pageNumber="66" pagination="1 - 1737" refId="ref75322" refString="Stebbing, T. R. R. (1888) Report on the Amphipoda collected by H. M. S. ' Challenger' during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. ' Challenger' during the years 1873 - 76. Zoology 29, i - xxiv &amp; 1 - 1737, pls 1 - 210." type="journal article" year="1888">Stebbing, 1888</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C615EB7FF90DD693559FF23FD04FF4A" authority="Shih &amp; Hendrycks, 2003" authorityName="Shih &amp; Hendrycks" authorityYear="2003" box="[265,694,192,218]" class="Malacostraca" family="Vibiliidae" genus="Vibilia" kingdom="Animalia" order="Amphipoda" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="B915F926FF90DD693559FF23FE3EFF49" box="[265,396,192,217]" italics="true" pageId="65" pageNumber="66">V. elongata</emphasis>
Shih &amp; Hendrycks, 2003
</taxonomicName>
, although in
<taxonomicName id="4C615EB7FF90DD69371CFF23FC42FF49" authorityName="Woltereck" authorityYear="1907" box="[844,1008,192,217]" class="Malacostraca" family="Prolanceolidae" genus="Prolanceola" kingdom="Animalia" order="Amphipoda" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="vibiliformis">
<emphasis id="B915F926FF90DD69371CFF23FC42FF49" box="[844,1008,192,217]" italics="true" pageId="65" pageNumber="66">P. vibiliformis</emphasis>
</taxonomicName>
the ocelli appear to be sub-cuticular. In having a mandibular palp with the third article longer than the first and second combined it resembles the family
<taxonomicName id="4C615EB7FF90DD6934BDFEF3FE02FEBA" authorityName="K. H. Barnard" authorityYear="1930" box="[237,432,272,298]" class="Malacostraca" family="Archaeoscinidae" kingdom="Animalia" order="Amphipoda" pageId="65" pageNumber="66" phylum="Arthropoda" rank="family">Archaeoscinidae</taxonomicName>
and
<taxonomicName id="4C615EB7FF90DD6935BEFEF3FDC1FEB9" authorityName="Barnard" authorityYear="1932" box="[494,627,272,297]" class="Malacostraca" family="Megalanceolidae" genus="Megalanceola" kingdom="Animalia" order="Amphipoda" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="remipes">
<emphasis id="B915F926FF90DD6935BEFEF3FDC1FEB9" box="[494,627,272,297]" italics="true" pageId="65" pageNumber="66">M. remipes</emphasis>
</taxonomicName>
(
<taxonomicName id="4C615EB7FF90DD6936D5FEF3FCFDFEBA" authority="Zeidler, 2009" authorityName="Zeidler" authorityYear="2009" box="[645,847,272,298]" class="Malacostraca" family="Megalanceolidae" kingdom="Animalia" order="Amphipoda" pageId="65" pageNumber="66" phylum="Arthropoda" rank="family" status="fam. nov.">Megalanceolidae</taxonomicName>
<emphasis id="B915F926FF90DD693708FEECFC7FFEB9" bold="true" box="[856,973,271,297]" pageId="65" pageNumber="66">
<taxonomicNameLabel id="A226445DFF90DD693708FEECFC7FFEB9" box="[856,973,271,297]" pageId="65" pageNumber="66" rank="family">fam. nov.</taxonomicNameLabel>
</emphasis>
), but the palp is relatively shorter and less robust and the body of the mandible is rectangular and not as narrowed distally. The manner in which the dactyls are inserted on pereopods 6 &amp; 7 is also similar to that found in
<taxonomicName id="4C615EB7FF90DD69379CFE83FBFCFEE9" authorityName="Barnard" authorityYear="1932" box="[972,1102,352,377]" class="Malacostraca" family="Megalanceolidae" genus="Megalanceola" kingdom="Animalia" order="Amphipoda" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="remipes">
<emphasis id="B915F926FF90DD69379CFE83FBFCFEE9" box="[972,1102,352,377]" italics="true" pageId="65" pageNumber="66">M. remipes</emphasis>
</taxonomicName>
but unlike most other lanceolids.
</paragraph>
</subSubSection>
</treatment>
</document>