A review of the hyperiidean amphipod superfamily Lanceoloidea Bowman & Gruner, 1973 (Crustacea: Amphipoda: Hyperiidea)
Author
Zeidler, Wolfgang
text
Zootaxa
2009
2009-02-04
2000
1
117
journal article
11755334
Family
PROLANCEOLIDAE
,
new family
Diagnosis.
Body length up to
14 mm
; pereon not inflated. Head without distinct rostrum. Eyes consist of chain of five ocelli on either side of head. Pereonites all separate. Coxae separate from pereonites. Antennae 1 with stout callynophore, peduncle 3-articulate. Antennae 2 longer than A1. Mandibles with strong palp, length almost 1.5x body length of mandible; third article of palp longer than first and second combined; left with broad lacinia mobilis. Maxillae 1 with broad palp, armed with few setae distally; outer lobe relatively slender, with five strong setae distally; inner lobe rounded, armed with numerous fine setae. Maxillae 2 with relatively broad lobes, the inner slightly shorter than outer one; both armed with row of strong setae together with more slender ones on inner and distal margin. Maxilliped with semicircular outer lobes, with row of fine setae along inner margin, slightly overlapping each other; inner lobes rounded, length 0.4x outer lobes, separate to base, not projected laterally from outer lobes as in other lanceolids. Gnathopod 1 subchelate, with distally expanded carpus and propodus. Gnathopod 2 simple, a little longer and more slender than G1. Pereopods 3–7 simple. Pereopods 6 & 7 with small retractile dactyls, without spoon-shaped process. Urosomite 2–3 with partial suture ventrally. Uropoda relatively broad (except first), with articulated exopoda and endopoda. Telson slightly longer than peduncle of U3, very elongate with pointed apex. Gills on pereonites 2–6. Oostegites on pereonites 2–5.
One genus:
Prolanceola
.
Remarks.
This new family is proposed here to accommodate
Prolanceola
which, prior to this study, was placed in the family
Lanceolidae
.
Woltereck (1907)
regarded
Prolanceola
to be a link between the families
Vibiliidae
and
Lanceolidae
and later (1909) he placed it in the
Lanceolidae
, presumably based on the retractile dactyls of pereopods 6 & 7. However, this placement is not satisfactory because
Prolanceola
possesses several unique characters that distinguish it from all other families of
Hyperiidea
. The chelate first gnathopoda and the structure of the eyes and mouthparts, especially the mandibles and second maxillae are particularly distinctive characters that distinguish
Prolanceola
and the family
Prolanceolidae
. It resembles
Vibilia
in the general habitus and also the arrangement of the ocelli of the eyes is similar to that of
V. australis
Stebbing, 1888
and
V. elongata
Shih & Hendrycks, 2003
, although in
P. vibiliformis
the ocelli appear to be sub-cuticular. In having a mandibular palp with the third article longer than the first and second combined it resembles the family
Archaeoscinidae
and
M. remipes
(
Megalanceolidae
fam. nov.
), but the palp is relatively shorter and less robust and the body of the mandible is rectangular and not as narrowed distally. The manner in which the dactyls are inserted on pereopods 6 & 7 is also similar to that found in
M. remipes
but unlike most other lanceolids.