216 lines
28 KiB
XML
216 lines
28 KiB
XML
<document id="3B76CAF4B58FBFE1782583C00F968AB6" ID-CLB-Dataset="7166" ID-DOI="10.1093/jmammal/gyab023" ID-GBIF-Dataset="d1c8718a-3b16-45c7-867d-c8ba4e9ff482" ID-ISSN="1545-1542" ID-Zenodo-Dep="5613298" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619530923730" checkinUser="felipe" docAuthor="Ochoa, Janine, Mijares, Armand S. B., Piper, Philip J., Reyes, Marian C. & Heaney, Lawrence R." docDate="2021" docId="03A41512FFFE4829FEA02ABD460A0024" docLanguage="en" docName="jMammalogy.102.2.1-22.pdf.imf" docOrigin="Journal of Mammalogy 102 (2)" docStyle="DocumentStyle:8488A0FD6434D222CC79ECAD85E66EFE.2:JourMammal.2015-.journal_article" docStyleId="8488A0FD6434D222CC79ECAD85E66EFE" docStyleName="JourMammal.2015-.journal_article" docStyleVersion="2" docTitle="Crateromys Thomas 1895" docType="treatment" docVersion="9" lastPageNumber="13" masterDocId="FF9D6D6AFFF24825FF8C2A7447020664" masterDocTitle="Three new extinct species from the endemic Philippine cloud rat radiation (Rodentia, Muridae, Phloeomyini)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="13" updateTime="1698937702425" updateUser="ExternalLinkService">
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<mods:titleInfo id="1DBE2146954DECF1898D12F5857EEFDB">
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<mods:title id="13D7DB910B36907D337E94F8830F65E0">Three new extinct species from the endemic Philippine cloud rat radiation (Rodentia, Muridae, Phloeomyini)</mods:title>
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<mods:name id="E3A02D4552D77393F65B848599BB0769" type="personal">
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<mods:namePart id="FA67AE55AC202406A033A27037471A69">Ochoa, Janine</mods:namePart>
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<mods:affiliation id="D5C727BABDCABC39E4D140ECB6B1D59E">Department of Anthropology, University of the Philippines, Diliman, Quezon City 1101, Philippines (JO) Archaeological Studies Program, University of the Philippines, Diliman, Quezon City 1101, Philippines (ASBM, PJP) National Museum of the Philippines, Padre Burgos Street, Manila 1000, Philippines (ASBM, PJP, MCR, LRH) Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA (LRH) Present address of PJP: School of Archaeology and Anthropology, Australian National University, Canberra, ACT 2601, Australia</mods:affiliation>
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<mods:nameIdentifier id="3011DEF29122669368A4BBB21E08575C" type="email">jochoa@up.edu.ph</mods:nameIdentifier>
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<mods:namePart id="A2E57B8DE0B041BD7FFCDA6CBBA6750D">Mijares, Armand S. B.</mods:namePart>
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<mods:affiliation id="057BCB930189658FBFE2C88944957CD2">Department of Anthropology, University of the Philippines, Diliman, Quezon City 1101, Philippines (JO) Archaeological Studies Program, University of the Philippines, Diliman, Quezon City 1101, Philippines (ASBM, PJP) National Museum of the Philippines, Padre Burgos Street, Manila 1000, Philippines (ASBM, PJP, MCR, LRH) Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA (LRH) Present address of PJP: School of Archaeology and Anthropology, Australian National University, Canberra, ACT 2601, Australia</mods:affiliation>
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</mods:name>
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<mods:namePart id="A2157B4FD8FC395895FB0B9D1D8DEE5E">Piper, Philip J.</mods:namePart>
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<mods:affiliation id="A9638F94BCF743A005531252B9EE166F">Department of Anthropology, University of the Philippines, Diliman, Quezon City 1101, Philippines (JO) Archaeological Studies Program, University of the Philippines, Diliman, Quezon City 1101, Philippines (ASBM, PJP) National Museum of the Philippines, Padre Burgos Street, Manila 1000, Philippines (ASBM, PJP, MCR, LRH) Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA (LRH) Present address of PJP: School of Archaeology and Anthropology, Australian National University, Canberra, ACT 2601, Australia</mods:affiliation>
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<mods:namePart id="B018DB842D628B51144F479E58188024">Reyes, Marian C.</mods:namePart>
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<mods:affiliation id="93D0601675FC9AFF768388CB24E0C3B1">Department of Anthropology, University of the Philippines, Diliman, Quezon City 1101, Philippines (JO) Archaeological Studies Program, University of the Philippines, Diliman, Quezon City 1101, Philippines (ASBM, PJP) National Museum of the Philippines, Padre Burgos Street, Manila 1000, Philippines (ASBM, PJP, MCR, LRH) Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA (LRH) Present address of PJP: School of Archaeology and Anthropology, Australian National University, Canberra, ACT 2601, Australia</mods:affiliation>
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<mods:namePart id="69A3A6563A4BA61E2EB927CD52AA7F77">Heaney, Lawrence R.</mods:namePart>
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<mods:affiliation id="32052E2E466386CCF4BEDC3E2816F6F0">Department of Anthropology, University of the Philippines, Diliman, Quezon City 1101, Philippines (JO) Archaeological Studies Program, University of the Philippines, Diliman, Quezon City 1101, Philippines (ASBM, PJP) National Museum of the Philippines, Padre Burgos Street, Manila 1000, Philippines (ASBM, PJP, MCR, LRH) Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA (LRH) Present address of PJP: School of Archaeology and Anthropology, Australian National University, Canberra, ACT 2601, Australia</mods:affiliation>
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<mods:typeOfResource id="3F54031CAA243CB6234B287BF6E672AB">text</mods:typeOfResource>
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<mods:title id="DC5689AF3A495C9378ACD92448351FF3">Journal of Mammalogy</mods:title>
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<mods:part id="EB3538D572D31624970A3AF8A25212C3">
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<mods:date id="9C9A750017964601EC476AA38715C1FD">2021</mods:date>
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<mods:number id="6E1F2C2E76C81B2CC5789BAC32AE8137">2021-04-23</mods:number>
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<mods:detail id="52A35ED34BB67973A0B8BE473F513BA5" type="volume">
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<mods:number id="997CBFBD5580894733050DCA1D63040F">102</mods:number>
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</mods:detail>
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<mods:detail id="0F08F1C35346EBB65DB50170458A72BA" type="issue">
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<mods:number id="D1B53F61615AF96CB248958A7015CD16">2</mods:number>
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<mods:start id="B5CBB2979593B60E7B3424ED03C8B930">1</mods:start>
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<mods:end id="5895A744024E519F1445EFA99156B64C">22</mods:end>
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<mods:classification id="0DF8B4E6DD8F96BA2F4B6FB5B968A445">journal article</mods:classification>
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<mods:identifier id="9B5C2B5DD92F75B476FF200C3DC01401" type="CLB-Dataset">7166</mods:identifier>
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<mods:identifier id="19A7E6CA992C4B0970838AA47C95DD76" type="DOI">10.1093/jmammal/gyab023</mods:identifier>
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<treatment id="03A41512FFFE4829FEA02ABD460A0024" ID-DOI="http://doi.org/10.5281/zenodo.4723436" ID-GBIF-Taxon="181653059" ID-Zenodo-Dep="4723436" LSID="urn:lsid:plazi:treatment:03A41512FFFE4829FEA02ABD460A0024" httpUri="http://treatment.plazi.org/id/03A41512FFFE4829FEA02ABD460A0024" lastPageNumber="13" pageId="12" pageNumber="13">
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<subSubSection id="C317F78FFFFE4829FEA02ABD45930684" box="[300,657,201,224]" pageId="12" pageNumber="13" type="nomenclature">
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<paragraph id="8BB2A404FFFE4829FEA02ABD45930684" blockId="12.[144,816,201,1600]" box="[300,657,201,224]" pageId="12" pageNumber="13">
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<heading id="D0FA1368FFFE4829FEA02ABD45930684" box="[300,657,201,224]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="13" reason="4">
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Genus
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<taxonomicName id="4C0DDF87FFFE4829FEFB2ABD45930684" ID-CoL="3VCV" authority="Thomas, 1895" authorityName="Thomas" authorityYear="1895" box="[375,657,201,224]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
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<emphasis id="B9797816FFFE4829FEFB2ABD46F10684" box="[375,499,201,224]" italics="true" pageId="12" pageNumber="13">Crateromys</emphasis>
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Thomas, 1895
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C317F78FFFFE4829FF1C2A9D460A0024" pageId="12" pageNumber="13" type="reference_group">
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<paragraph id="8BB2A404FFFE4829FF1C2A9D460A0024" blockId="12.[144,816,201,1600]" pageId="12" pageNumber="13">
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<taxonomicName id="4C0DDF87FFFE4829FF1C2A9D46120764" ID-CoL="3VCV" authorityName="Thomas" authorityYear="1895" box="[144,272,233,256]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
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<emphasis id="B9797816FFFE4829FF1C2A9D46120764" box="[144,272,233,256]" italics="true" pageId="12" pageNumber="13">Crateromys</emphasis>
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</taxonomicName>
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is represented by one living species on Luzon (
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<taxonomicName id="4C0DDF87FFFE4829FF152B7D46540744" ID-CoL="Z92W" baseAuthorityName="Meyer" baseAuthorityYear="1895" box="[153,342,265,288]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="schadenbergi">
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<emphasis id="B9797816FFFE4829FF152B7D46540744" box="[153,342,265,288]" italics="true" pageId="12" pageNumber="13">Cr. schadenbergi</emphasis>
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</taxonomicName>
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) and two additional species in other parts of the oceanic
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<collectingCountry id="F31AE494FFFE4829FEB02B5D46BF0724" box="[316,445,297,320]" name="Philippines" pageId="12" pageNumber="13">Philippines</collectingCountry>
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:
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<taxonomicName id="4C0DDF87FFFE4829FE472B5D454E0724" ID-CoL="6BCPF" authorityName="Gonzales & Kennedy" authorityYear="1996" box="[459,588,297,320]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="heaneyi">
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<emphasis id="B9797816FFFE4829FE472B5D454E0724" box="[459,588,297,320]" italics="true" pageId="12" pageNumber="13">Cr. heaneyi</emphasis>
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</taxonomicName>
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from Panay Island, and (possibly) extinct
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<taxonomicName id="4C0DDF87FFFE4829FE132B3D451A0704" ID-CoL="Z92V" authorityName="Musser & Gordon" authorityYear="1981" box="[415,536,329,352]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="paulus">
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<emphasis id="B9797816FFFE4829FE132B3D451A0704" box="[415,536,329,352]" italics="true" pageId="12" pageNumber="13">Cr. paulus</emphasis>
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</taxonomicName>
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from Ilin Island (near Mindoro); as noted above, molecular data indicate that
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<taxonomicName id="4C0DDF87FFFE4829FC812B1D47F207C4" ID-CoL="Z92T" authorityName="Musser, Heaney & Rabor" authorityYear="1985" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="australis">
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<emphasis id="B9797816FFFE4829FC812B1D47F207C4" italics="true" pageId="12" pageNumber="13">Cr. australis</emphasis>
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</taxonomicName>
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from
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<collectingRegion id="49C96AE6FFFE4829FEBF2BFD468807C4" box="[307,394,393,416]" country="Philippines" name="Dinagat Islands" pageId="12" pageNumber="13">Dinagat</collectingRegion>
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Island (near Mindanao) should not be placed as a member of this genus (
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<bibRefCitation id="EF9CD9F5FFFE4829FD9B2BDD45EF07A4" author="Rowsey" box="[535,749,425,448]" etAl="et al." firstAuthor="Rowsey" journalOrPublisher="Evolution" pageId="12" pageNumber="13" pagination="1420 - 1435" part="72" refId="ref19146" refString="ROWSEY, D. M., L. R. HEANEY, AND S. A. JANSA. 2018. Diversification rates of the " Old Endemic " murine rodents of Luzon Island, Philippines are inconsistent with incumbency and ecological opportunity. Evolution 72: 1420 - 1435." title="Diversification rates of the " Old Endemic " murine rodents of Luzon Island, Philippines are inconsistent with incumbency and ecological opportunity" type="journal article" year="2018">Rowsey et al. 2018</bibRefCitation>
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), and we do not include it in our treatment that follows. The cranial traits and features of external appearance of
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<taxonomicName id="4C0DDF87FFFE4829FDF52B9D45FB0464" ID-CoL="3VCV" authorityName="Thomas" authorityYear="1895" box="[633,761,489,512]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
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<emphasis id="B9797816FFFE4829FDF52B9D45FB0464" box="[633,761,489,512]" italics="true" pageId="12" pageNumber="13">Crateromys</emphasis>
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</taxonomicName>
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species were described by Musser and Gordon (1981),
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<bibRefCitation id="EF9CD9F5FFFE4829FD51287D46230424" author="Musser" etAl="et al." firstAuthor="Musser" journalOrPublisher="American Museum Novitates" pageId="12" pageNumber="13" pagination="1 - 25" part="2821" refId="ref18546" refString="MUSSER, G. G., L. R. HEANEY, AND D. S. RABOR. 1985. Philippine rats: description of a new species of Crateromys from Dinagat Island. American Museum Novitates 2821: 1 - 25." title="Philippine rats: description of a new species of Crateromys from Dinagat Island" type="journal article" year="1985">Musser et al. (1985)</bibRefCitation>
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, Gonzales and Kennedy (1996), and
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<bibRefCitation id="EF9CD9F5FFFE4829FD56285D46220404" author="Heaney" etAl="et al." firstAuthor="Heaney" journalOrPublisher="Johns Hopkins University Press. Baltimore, Maryland" pageId="12" pageNumber="13" refId="ref17461" refString="HEANEY, L. R., D. S. BALETE, AND E. A. RICKART. 2016 a. The mammals of Luzon Island: biogeography and natural history of a Philippine fauna. Johns Hopkins University Press. Baltimore, Maryland." title="The mammals of Luzon Island: biogeography and natural history of a Philippine fauna" type="book" year="2016" yearSuffix="a">Heaney et al. (2016a)</bibRefCitation>
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.
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<taxonomicName id="4C0DDF87FFFE4829FEA1283D46AF0404" ID-CoL="3VCV" authorityName="Thomas" authorityYear="1895" box="[301,429,585,608]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
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<emphasis id="B9797816FFFE4829FEA1283D46AF0404" box="[301,429,585,608]" italics="true" pageId="12" pageNumber="13">Crateromys</emphasis>
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</taxonomicName>
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possesses high crowns, chevronate lower molar lophs, and strongly arched upper molar cusps that are separated by deep clefts and ridges (
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<figureCitation id="1336B881FFFE4829FDFC28FD45B804C4" box="[624,698,649,672]" captionStart="Fig" captionStartId="9.[101,136,1865,1886]" captionTargetBox="[106,767,201,1850]" captionTargetId="figure-521@9.[105,767,200,1851]" captionTargetPageId="9" captionText="Fig. 4.—Occlusal view of maxillary molars of (A) Carpomys dakal n. sp. (PNM II-1977-J11A-15997), left side; (B) Carpomys phaeurus (FMNH62291),leftside;(C) Musseromysinopinatus(FMNH193840), left side; (D) Crateromys ballik n. sp. (PNM II-1977-J3-10389), right side (mirrored); (E) Crateromys schadenbergi (FMNH 62291), right side (mirrored); (F) Crateromys heaneyi (CMC M626), left side; (G) Batomys granti (FMNH 214412), left side." figureDoi="http://doi.org/10.5281/zenodo.5613310" httpUri="https://zenodo.org/record/5613310/files/figure.png" pageId="12" pageNumber="13">Figs. 4</figureCitation>
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and
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<figureCitation id="1336B881FFFE4829FD6128FD45F804C4" box="[749,762,649,672]" captionStart="Fig" captionStartId="10.[144,178,1865,1886]" captionTargetBox="[148,809,201,1850]" captionTargetId="figure-534@10.[147,809,200,1851]" captionTargetPageId="10" captionText="Fig. 5.—Occlusal view of mandibular molars of (A) Carpomys dakal n. sp. (PNM II-1977-J11A-16301, holotype), leftside; (B) Carpomys phaeurus (FMNH62291),leftside;(C) Musseromysinopinatus (FMNH 193840), left side; (D) Crateromys ballik n. sp. (PNM II-1977-I8-402, holotype), right side (mirrored); (E) Crateromys schadenbergi (FMNH 62291), left side; (F) Crateromys heaneyi (CMC M626), left side; (G) Batomys cagayanensis n. sp. (PNM II-1977-J3-7554, holotype), right side (mirrored); (H) Batomys granti (FMNH 214412), left side." figureDoi="http://doi.org/10.5281/zenodo.5613314" httpUri="https://zenodo.org/record/5613314/files/figure.png" pageId="12" pageNumber="13">5</figureCitation>
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). As detailed above, this contrasts with the medium crown height and less strongly arched cusps of
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<taxonomicName id="4C0DDF87FFFE4829FE7628BD45680484" ID-CoL="62KJX" authority="Thomas, 1895" box="[506,618,713,736]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
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<emphasis id="B9797816FFFE4829FE7628BD45680484" box="[506,618,713,736]" italics="true" pageId="12" pageNumber="13">Carpomys</emphasis>
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</taxonomicName>
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.
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<taxonomicName id="4C0DDF87FFFE4829FDFB28BD45F50484" ID-CoL="3VCV" authorityName="Thomas" authorityYear="1895" box="[631,759,713,736]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
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<emphasis id="B9797816FFFE4829FDFB28BD45F50484" box="[631,759,713,736]" italics="true" pageId="12" pageNumber="13">Crateromys</emphasis>
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</taxonomicName>
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dentition is most similar to that of
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<taxonomicName id="4C0DDF87FFFE4829FE66289D454A0564" ID-CoL="38TM" authorityName="Thomas" authorityYear="1895" box="[490,584,745,768]" class="Mammalia" family="Muridae" genus="Batomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
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<emphasis id="B9797816FFFE4829FE66289D454A0564" box="[490,584,745,768]" italics="true" pageId="12" pageNumber="13">Batomys</emphasis>
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</taxonomicName>
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, but the members of these two genera on Luzon differ dentally in several respects. Both
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<taxonomicName id="4C0DDF87FFFE4829FF5C295D46520524" ID-CoL="3VCV" authorityName="Thomas" authorityYear="1895" box="[208,336,809,832]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
|
<emphasis id="B9797816FFFE4829FF5C295D46520524" box="[208,336,809,832]" italics="true" pageId="12" pageNumber="13">Crateromys</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C0DDF87FFFE4829FE00295D46E80524" ID-CoL="38TM" authorityName="Thomas" authorityYear="1895" box="[396,490,809,832]" class="Mammalia" family="Muridae" genus="Batomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
|
<emphasis id="B9797816FFFE4829FE00295D46E80524" box="[396,490,809,832]" italics="true" pageId="12" pageNumber="13">Batomys</emphasis>
|
|
</taxonomicName>
|
|
have hypsodont molars, but
|
|
<taxonomicName id="4C0DDF87FFFE4829FF1C293D46120504" ID-CoL="3VCV" authorityName="Thomas" authorityYear="1895" box="[144,272,841,864]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
|
<emphasis id="B9797816FFFE4829FF1C293D46120504" box="[144,272,841,864]" italics="true" pageId="12" pageNumber="13">Crateromys</emphasis>
|
|
</taxonomicName>
|
|
has higher crowns than
|
|
<emphasis id="B9797816FFFE4829FDAB293D458F0504" box="[551,653,841,864]" italics="true" pageId="12" pageNumber="13">
|
|
<taxonomicName id="4C0DDF87FFFE4829FDAB293D458B0504" ID-CoL="38TM" authorityName="Thomas" authorityYear="1895" box="[551,649,841,864]" class="Mammalia" family="Muridae" genus="Batomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">Batomys</taxonomicName>
|
|
.
|
|
</emphasis>
|
|
Upper molars of
|
|
<taxonomicName id="4C0DDF87FFFE4829FF22291D462C05E4" ID-CoL="3VCV" authorityName="Thomas" authorityYear="1895" box="[174,302,873,896]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
|
<emphasis id="B9797816FFFE4829FF22291D462C05E4" box="[174,302,873,896]" italics="true" pageId="12" pageNumber="13">Crateromys</emphasis>
|
|
</taxonomicName>
|
|
generally are more angular, whereas
|
|
<taxonomicName id="4C0DDF87FFFE4829FD43291D442F05E4" ID-CoL="38TM" authorityName="Thomas" authorityYear="1895" box="[719,813,873,896]" class="Mammalia" family="Muridae" genus="Batomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
|
<emphasis id="B9797816FFFE4829FD43291D442F05E4" box="[719,813,873,896]" italics="true" pageId="12" pageNumber="13">Batomys</emphasis>
|
|
</taxonomicName>
|
|
cusps are more rounded in occlusal outline. The angular shape of cusps in
|
|
<taxonomicName id="4C0DDF87FFFE4829FEEF29DD46E105A4" ID-CoL="3VCV" authorityName="Thomas" authorityYear="1895" box="[355,483,937,960]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
|
<emphasis id="B9797816FFFE4829FEEF29DD46E105A4" box="[355,483,937,960]" italics="true" pageId="12" pageNumber="13">Crateromys</emphasis>
|
|
</taxonomicName>
|
|
in part is due to deep clefts that separate lingual and medial cusps of upper molars. In anterocone of M1, t1 and t2 are separated by a deep cleft, and this pattern is repeated for t4/t5 and t7/t
|
|
<quantity id="4CF509E1FFFE4829FDC52E7D45700244" box="[585,626,1033,1056]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="12" pageNumber="13" unit="in" value="8.0">8 in</quantity>
|
|
all upper molars (
|
|
<figureCitation id="1336B881FFFE4829FF152E5D47EF0224" box="[153,237,1065,1088]" captionStart="Fig" captionStartId="4.[861,896,1502,1523]" captionTargetBox="[867,1527,202,1488]" captionTargetId="figure-650@4.[865,1527,200,1489]" captionTargetPageId="4" captionText="Fig. 2.—Maxillary and mandibular left molar toothrows of Carpomys dakal n. sp. (A and B, respectively; PNM II-1977-J11A-15997 and PNM II-1976-M-101) and Crateromys schadenbergi (C and D, respectively; FMNH 62295), showing the location of cusps and conids referred to in the text. pc = posterior cingulum." figureDoi="http://doi.org/10.5281/zenodo.5613304" httpUri="https://zenodo.org/record/5613304/files/figure.png" pageId="12" pageNumber="13">Fig. 2C</figureCitation>
|
|
). These deep indentations are absent or less developed in most upper molar lophs in
|
|
<taxonomicName id="4C0DDF87FFFE4829FD9A2E3D45760204" ID-CoL="38TM" authorityName="Thomas" authorityYear="1895" box="[534,628,1097,1120]" class="Mammalia" family="Muridae" genus="Batomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
|
<emphasis id="B9797816FFFE4829FD9A2E3D45760204" box="[534,628,1097,1120]" italics="true" pageId="12" pageNumber="13">Batomys</emphasis>
|
|
</taxonomicName>
|
|
, with the exception of the posterior lophs (t7 and t8) in M1 and M2. In the
|
|
<taxonomicName id="4C0DDF87FFFE4829FF1C2EFD461202C4" ID-CoL="3VCV" authorityName="Thomas" authorityYear="1895" box="[144,272,1161,1184]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
|
<emphasis id="B9797816FFFE4829FF1C2EFD461202C4" box="[144,272,1161,1184]" italics="true" pageId="12" pageNumber="13">Crateromys</emphasis>
|
|
</taxonomicName>
|
|
M1 anterocone, t1 and t3 are typically oriented anteriorly, whereas these cusps are oriented posteriorly in
|
|
<taxonomicName id="4C0DDF87FFFE4829FF1C2EBD47EC0284" ID-CoL="38TM" authorityName="Thomas" authorityYear="1895" box="[144,238,1225,1248]" class="Mammalia" family="Muridae" genus="Batomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
|
<emphasis id="B9797816FFFE4829FF1C2EBD47EC0284" box="[144,238,1225,1248]" italics="true" pageId="12" pageNumber="13">Batomys</emphasis>
|
|
</taxonomicName>
|
|
, producing a concave occlusal outline. Asimilar pattern is observed in the anterior loph of M2, where t4 and t6 are oriented anteriorly in
|
|
<taxonomicName id="4C0DDF87FFFE4829FE5C2F7D45520344" ID-CoL="3VCV" authorityName="Thomas" authorityYear="1895" box="[464,592,1289,1312]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
|
<emphasis id="B9797816FFFE4829FE5C2F7D45520344" box="[464,592,1289,1312]" italics="true" pageId="12" pageNumber="13">Crateromys</emphasis>
|
|
</taxonomicName>
|
|
and transversely in
|
|
<taxonomicName id="4C0DDF87FFFE4829FF1C2F5D47EC0324" ID-CoL="38TM" authorityName="Thomas" authorityYear="1895" box="[144,238,1321,1344]" class="Mammalia" family="Muridae" genus="Batomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
|
<emphasis id="B9797816FFFE4829FF1C2F5D47EC0324" box="[144,238,1321,1344]" italics="true" pageId="12" pageNumber="13">Batomys</emphasis>
|
|
</taxonomicName>
|
|
. The anterior and posterior lophs of M3 have a more complex occlusal outline in
|
|
<taxonomicName id="4C0DDF87FFFE4829FE492F3D45470304" ID-CoL="3VCV" authorityName="Thomas" authorityYear="1895" box="[453,581,1353,1376]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
|
<emphasis id="B9797816FFFE4829FE492F3D45470304" box="[453,581,1353,1376]" italics="true" pageId="12" pageNumber="13">Crateromys</emphasis>
|
|
</taxonomicName>
|
|
, whereas these lophs have a simpler laminar outline in
|
|
<taxonomicName id="4C0DDF87FFFE4829FD992F1D457103E4" ID-CoL="38TM" authorityName="Thomas" authorityYear="1895" box="[533,627,1385,1408]" class="Mammalia" family="Muridae" genus="Batomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
|
<emphasis id="B9797816FFFE4829FD992F1D457103E4" box="[533,627,1385,1408]" italics="true" pageId="12" pageNumber="13">Batomys</emphasis>
|
|
</taxonomicName>
|
|
. In
|
|
<taxonomicName id="4C0DDF87FFFE4829FD2A2F1D442403E4" ID-CoL="3VCV" authorityName="Thomas" authorityYear="1895" box="[678,806,1385,1408]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
|
<emphasis id="B9797816FFFE4829FD2A2F1D442403E4" box="[678,806,1385,1408]" italics="true" pageId="12" pageNumber="13">Crateromys</emphasis>
|
|
</taxonomicName>
|
|
, the anterior and posterior lophs of m3 are chevronate and a posterior cingulum is occasionally observed; in
|
|
<taxonomicName id="4C0DDF87FFFE4829FD2C2FDD45FC03A4" ID-CoL="38TM" authorityName="Thomas" authorityYear="1895" box="[672,766,1449,1472]" class="Mammalia" family="Muridae" genus="Batomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
|
<emphasis id="B9797816FFFE4829FD2C2FDD45FC03A4" box="[672,766,1449,1472]" italics="true" pageId="12" pageNumber="13">Batomys</emphasis>
|
|
</taxonomicName>
|
|
, the anterior loph is not arched and the posterior loph is laminar or oblong in outline. Our assignment of the following new species to
|
|
<taxonomicName id="4C0DDF87FFFE4829FE842C7D468A0044" ID-CoL="3VCV" authorityName="Thomas" authorityYear="1895" box="[264,392,1545,1568]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
|
<emphasis id="B9797816FFFE4829FE842C7D468A0044" box="[264,392,1545,1568]" italics="true" pageId="12" pageNumber="13">Crateromys</emphasis>
|
|
</taxonomicName>
|
|
is based on its possessing this set of characters.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |