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<document id="35402CA09EEF934974B5F3B797F8AEC9" ID-DOI="10.11646/zootaxa.4000.1.1" ID-GBIF-Dataset="507a3784-35f5-46f5-9492-e0884a502f40" ID-ISSN="1175-5326" ID-Zenodo-Dep="289219" ID-ZooBank="00769C76-A2D4-4792-A5DD-5F2B7EA352CB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461184674989" checkinUser="plazi" docAuthor="Allegro, Gianni &amp; Giachino, Pier Mauro" docDate="2015" docId="039087C7FFA7FF8EE28C4E52343A6874" docLanguage="en" docName="zt04000p048.pdf" docOrigin="Zootaxa 4000 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Blennidus (Agraphoderus) fitzcarraldi Giachino &amp; Allegro 2011" docType="treatment" docVersion="6" lastPageNumber="9" masterDocId="FFA9FFBFFFAFFF86E21B4B7535296F7A" masterDocTitle="Annotated checklist of the Blennidus subgenus Agraphoderus species from Peru with description of B. bombonensis n. sp. and synonymic notes (Coleoptera: Carabidae: Pterostichinae)" masterLastPageNumber="48" masterPageNumber="1" pageNumber="9" updateTime="1698599231025" updateUser="plazi">
<mods:mods id="ED0D745FAF5E2C6ED9FE8C531032903B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AD5E82E4EBF283F9DFA79E01CE9F7F5B">
<mods:title id="9F3EBA20A9FD3C3BC52AAF480C523835">Annotated checklist of the Blennidus subgenus Agraphoderus species from Peru with description of B. bombonensis n. sp. and synonymic notes (Coleoptera: Carabidae: Pterostichinae)</mods:title>
</mods:titleInfo>
<mods:name id="A432A027849B194121E81172D30FC2FE" type="personal">
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</mods:role>
<mods:namePart id="769A780F20B29201F8F955D5B3E57933">Allegro, Gianni</mods:namePart>
</mods:name>
<mods:name id="67F9DD06B359DD8E69CB31C4DB32B7F4" type="personal">
<mods:role id="45E1D924EF80BD39FBBA93D14B5624B1">
<mods:roleTerm id="FBD5ADEB1F5C699AA67D946BBBD0365B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="398E782578E130DA616348C9CB3592CF">Giachino, Pier Mauro</mods:namePart>
</mods:name>
<mods:typeOfResource id="94C820E107A083A31778FDC09124BBA2">text</mods:typeOfResource>
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<mods:title id="11E7C8D8F8AFA732856256A68FF27B85">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="234338A8ECC4BFC08543681838B7D8A9">
<mods:date id="A7301DC4B7C90459CD101434B6573E19">2015</mods:date>
<mods:detail id="A77E64C75914F1D48CFA0797E6679B2F" type="volume">
<mods:number id="C1976A07565F22FB82D78A46A4DCD38E">4000</mods:number>
</mods:detail>
<mods:detail id="8666912FFDE77D5E673FE5DDEF69D1E6" type="issue">
<mods:number id="E900395C334BC70943A47B78D216B965">1</mods:number>
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<mods:start id="EA618C5DC10844E1E6D2BE5AAC1B7937">1</mods:start>
<mods:end id="7E7340DAABFB26A5A2CB304178012F06">48</mods:end>
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<mods:classification id="6F8F17600C7BC2F6BDDFB6CAEE3EF33E">journal article</mods:classification>
<mods:identifier id="38FF43139C2A26CFB2587D5755284276" type="DOI">10.11646/zootaxa.4000.1.1</mods:identifier>
<mods:identifier id="728629B1CE53747C72DE19F11698AE31" type="GBIF-Dataset">507a3784-35f5-46f5-9492-e0884a502f40</mods:identifier>
<mods:identifier id="A8D78A3A68B88CDDB3190F5CDB22FA14" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="51281E34D40016139A93EDE7986A0556" type="Zenodo-Dep">289219</mods:identifier>
<mods:identifier id="1BF47770BCFA3944FE9ED10545588343" type="ZooBank">00769C76-A2D4-4792-A5DD-5F2B7EA352CB</mods:identifier>
</mods:mods>
<treatment id="039087C7FFA7FF8EE28C4E52343A6874" ID-DOI="http://doi.org/10.5281/zenodo.6096032" ID-GBIF-Taxon="127692751" ID-Zenodo-Dep="6096032" LSID="urn:lsid:plazi:treatment:039087C7FFA7FF8EE28C4E52343A6874" httpUri="http://treatment.plazi.org/id/039087C7FFA7FF8EE28C4E52343A6874" lastPageNumber="9" pageId="8" pageNumber="9">
<subSubSection id="C323655AFFA7FF8EE28C4E5234716A19" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8B8636D1FFA7FF8EE28C4E5236826A38" blockId="8.[151,939,1319,1379]" box="[151,939,1319,1346]" pageId="8" pageNumber="9">
<heading id="D0CE81BDFFA7FF8EE28C4E5236826A38" bold="true" box="[151,939,1319,1346]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<taxonomicName id="4C394D52FFA7FF8EE28C4E5236826A38" ID-CoL="M54C" authority="Giachino &amp; Allegro, 2011" authorityName="Giachino &amp; Allegro" authorityYear="2011" box="[151,939,1319,1346]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="fitzcarraldi" subGenus="Agraphoderus">
<emphasis id="B94DEAC3FFA7FF8EE28C4E5236826A38" bold="true" box="[151,939,1319,1346]" pageId="8" pageNumber="9">
<emphasis id="B94DEAC3FFA7FF8EE28C4E52343A6A3B" bold="true" box="[151,275,1319,1345]" italics="true" pageId="8" pageNumber="9">Blennidus</emphasis>
(
<emphasis id="B94DEAC3FFA7FF8EE33F4E5234F96A3B" bold="true" box="[292,464,1319,1345]" italics="true" pageId="8" pageNumber="9">Agraphoderus</emphasis>
)
<emphasis id="B94DEAC3FFA7FF8EE3FA4E52374F6A3B" bold="true" box="[481,614,1319,1345]" italics="true" pageId="8" pageNumber="9">fitzcarraldi</emphasis>
Giachino &amp; Allegro, 2011
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8636D1FFA7FF8EE28C4E3F34716A19" blockId="8.[151,939,1319,1379]" box="[151,344,1354,1379]" pageId="8" pageNumber="9">
(
<figureCitation id="13022A54FFA7FF8EE2844E3F35D56A19" box="[159,252,1354,1379]" captionStart="FIGURES 20 – 23" captionStartId="29.[151,264,1953,1976]" captionTargetBox="[255,1328,212,1900]" captionTargetId="figure@29.[238,1348,193,1932]" captionTargetPageId="29" captionText="FIGURES 20 – 23. Habitus of Blennidus: egens male HT (20); etontii male HT (21); euphaenops male HT (22); fitzcarraldi male HT (23)." httpUri="https://zenodo.org/record/289225/files/figure.png" pageId="8" pageNumber="9">Figs. 23</figureCitation>
,
<figureCitation id="13022A54FFA7FF8EE3134E3F340F6A19" box="[264,294,1354,1379]" captionStart="FIGURES 58 – 65" captionStartId="38.[151,264,1931,1954]" captionTargetBox="[169,1417,210,1898]" captionTargetId="figure@38.[151,1436,193,1910]" captionTargetPageId="38" captionText="FIGURES 58 – 65. Pronotum of Blennidus: egens male HT (58); etontii male HT (59); euphaenops male HT (60); fitzcarraldi male HT (61); huascarani male HT (62); inca male LT (63); jelskii male LT (64); languens male LT (65)." httpUri="https://zenodo.org/record/289233/files/figure.png" pageId="8" pageNumber="9">61</figureCitation>
,
<figureCitation id="13022A54FFA7FF8EE3294E3E34666A19" box="[306,335,1355,1379]" captionStart="FIGURES 94 – 99" captionStartId="43.[151,264,1953,1976]" captionTargetBox="[218,1362,198,1914]" captionTargetId="figure@43.[201,1386,193,1932]" captionTargetPageId="43" captionText="FIGURES 94 – 99. Aedeagus (left lateral and dorsal aspects) of Blennidus: curtatus HT (94); etontii HT (95); euphaenops HT (96); fitzcarraldi HT (97); huascarani HT (98); inca LT (99)." httpUri="https://zenodo.org/record/289238/files/figure.png" pageId="8" pageNumber="9">97</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C323655AFFA7FF8EE28C4EE737826AD2" box="[151,683,1426,1448]" pageId="8" pageNumber="9" type="reference_group">
<paragraph id="8B8636D1FFA7FF8EE28C4EE737826AD2" blockId="8.[151,683,1426,1448]" box="[151,683,1426,1448]" pageId="8" pageNumber="9">
<taxonomicName id="4C394D52FFA7FF8EE28C4EE737826AD2" ID-CoL="M54C" authority="Giachino &amp; Allegro, 2011: 23" authorityName="Giachino &amp; Allegro" authorityPageNumber="23" authorityYear="2011" box="[151,683,1426,1448]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="fitzcarraldi">
<emphasis id="B94DEAC3FFA7FF8EE28C4EE7345D6AD2" box="[151,372,1426,1448]" italics="true" pageId="8" pageNumber="9">Blennidus fitzcarraldi</emphasis>
Giachino &amp; Allegro, 2011: 23
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C323655AFFA7FF8EE28C4EA0361969DB" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="8B8636D1FFA7FF8EE28C4EA036F46A94" blockId="8.[151,1436,1493,1806]" box="[151,989,1493,1518]" pageId="8" pageNumber="9">
<emphasis id="B94DEAC3FFA7FF8EE28C4EA0341C6A94" bold="true" box="[151,309,1493,1518]" pageId="8" pageNumber="9">
<typeStatus id="54828873FFA7FF8EE28C4EA035FB6A94" box="[151,210,1493,1518]" pageId="8" pageNumber="9">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32E7641FFA7FF8EE3274EA3345B6A97" box="[316,370,1494,1517]" name="Peru" pageId="8" pageNumber="9">Peru</collectingCountry>
, Dip. Ancash, San Luis, laguna Huachucocha, m 4200.
</paragraph>
<paragraph id="8B8636D1FFA7FF8EE2DC4E8C37A5694C" blockId="8.[151,1436,1493,1806]" pageId="8" pageNumber="9">
<emphasis id="B94DEAC3FFA7FF8EE2DC4E8C34916968" bold="true" box="[199,440,1529,1554]" pageId="8" pageNumber="9">Examined material.</emphasis>
HT ♂ (CAl); 47 PTT (
<specimenCount id="9D3FFD58FFA7FF8EE0C84E8C3639696B" box="[723,784,1529,1554]" pageId="8" pageNumber="9" type="generic">35 ♂</specimenCount>
and 12 ♀) (complete TS) (CAl, CCa, CGi, CMo, CVi,
<collectionCode id="ED28AE14FFA7FF8EE28C4D6B35C7694F" box="[151,238,1566,1589]" pageId="8" pageNumber="9">BMNH</collectionCode>
,
<collectionCode id="ED28AE14FFA7FF8EE2E24D6B3477694F" box="[249,350,1566,1589]" pageId="8" pageNumber="9">MNHNP</collectionCode>
,
<collectionCode id="ED28AE14FFA7FF8EE3724D6B3492694C" box="[361,443,1566,1590]" pageId="8" pageNumber="9">MRSN</collectionCode>
,
<collectionCode id="ED28AE14FFA7FF8EE3DD4D6B3737694C" box="[454,542,1566,1590]" pageId="8" pageNumber="9">MSNM</collectionCode>
,
<collectionCode id="ED28AE14FFA7FF8EE0314D6B37A9694C" LSID="urn:lsid:biocol.org:col:34404" box="[554,640,1566,1590]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34404" name="University of Alberta, E.H. Strickland Entomological Museum" pageId="8" pageNumber="9">UASM</collectionCode>
).
</paragraph>
<paragraph id="8B8636D1FFA7FF8EE2DC4D34361969DB" blockId="8.[151,1436,1493,1806]" pageId="8" pageNumber="9">
<emphasis id="B94DEAC3FFA7FF8EE2DC4D34342F6920" bold="true" box="[199,262,1601,1626]" pageId="8" pageNumber="9">Note.</emphasis>
This species is easily distinguished from the others from Cordillera Blanca (
<taxonomicName id="4C394D52FFA7FF8EE6BA4D37306A6923" box="[1185,1347,1601,1625]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="huascarani">
<emphasis id="B94DEAC3FFA7FF8EE6BA4D37306A6923" box="[1185,1347,1601,1625]" italics="true" pageId="8" pageNumber="9">B. huascarani</emphasis>
</taxonomicName>
and
<taxonomicName id="4C394D52FFA7FF8EE79F4D3735C76907" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="unistria">
<emphasis id="B94DEAC3FFA7FF8EE79F4D3735C76907" italics="true" pageId="8" pageNumber="9">B. unistria</emphasis>
</taxonomicName>
) by the markedly angulate hind angles of pronotum. Because of the similarities in external morphology and in male genitalia, its closest relative is probably
<taxonomicName id="4C394D52FFA7FF8EE0A84DFF360069DB" box="[691,809,1674,1697]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="unistria">
<emphasis id="B94DEAC3FFA7FF8EE0A84DFF360069DB" box="[691,809,1674,1697]" italics="true" pageId="8" pageNumber="9">B. unistria</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C323655AFFA7FF8EE2DC4DD8343A6874" pageId="8" pageNumber="9" type="distribution">
<paragraph id="8B8636D1FFA7FF8EE2DC4DD836C169BC" blockId="8.[151,1436,1493,1806]" box="[199,1000,1709,1734]" pageId="8" pageNumber="9">
<emphasis id="B94DEAC3FFA7FF8EE2DC4DD8344969BC" bold="true" box="[199,352,1709,1734]" pageId="8" pageNumber="9">Distribution.</emphasis>
Restricted to the Dept. Ancash, within Cordillera Blanca.
</paragraph>
<paragraph id="8B8636D1FFA7FF8EE2DC4DA4343A6874" blockId="8.[151,1436,1493,1806]" pageId="8" pageNumber="9">
<emphasis id="B94DEAC3FFA7FF8EE2DC4DA434036990" bold="true" box="[199,298,1745,1770]" pageId="8" pageNumber="9">Habitat.</emphasis>
All the recorded specimens (legit G. Allegro) are from high altitude (
<quantity id="4CC19B34FFA7FF8EE6524DA431DF6993" box="[1097,1270,1745,1770]" metricMagnitude="3" metricUnit="m" metricValue="4.325" metricValueMax="4.45" metricValueMin="4.2" pageId="8" pageNumber="9" unit="m" value="4325.0" valueMax="4450.0" valueMin="4200.0">4,200–4,450 m</quantity>
a.s.l.) Andean grasslands.
</paragraph>
</subSubSection>
</treatment>
</document>
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