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<document id="BEB5AEF32F3B5939E7459E7021630A81" ID-DOI="10.11646/zootaxa.5026.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10100344" ID-ZooBank="07E0C922-6B0F-4916-85C2-AD95146A8F1E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629979906524" checkinUser="plazi" docAuthor="Neita-Moreno, Jhon C." docDate="2021" docId="03932D70B629C37DFF7B0C91A29CD923" docLanguage="en" docName="zootaxa.5026.1.1.pdf" docOrigin="Zootaxa 5026 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyclocephala fasciolata Bates 1888" docType="treatment" docVersion="2" lastPageNumber="11" masterDocId="FFAA5508B623C377FFEC0C06A02ADC12" masterDocTitle="A review of the black species of Cyclocephala Dejean (Coleoptera: Scarabaeidae Dynastinae) from Colombia" masterLastPageNumber="58" masterPageNumber="1" pageNumber="11" updateTime="1699603307102" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E34DED293B56BE7A9C494D2093647DA8">A review of the black species of Cyclocephala Dejean (Coleoptera: Scarabaeidae Dynastinae) from Colombia</mods:title>
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<mods:namePart id="25300136046EF85AA353AAFD3ABFD1DD">Neita-Moreno, Jhon C.</mods:namePart>
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<mods:date id="42C34ED527D5B3E6C1A9E3632F878457">2021</mods:date>
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<treatment id="03932D70B629C37DFF7B0C91A29CD923" LSID="urn:lsid:plazi:treatment:03932D70B629C37DFF7B0C91A29CD923" httpUri="http://treatment.plazi.org/id/03932D70B629C37DFF7B0C91A29CD923" lastPageNumber="11" pageId="10" pageNumber="11">
<subSubSection id="C320CFEDB629C37DFF7B0C91A24CDCA3" box="[151,614,151,178]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="8B859C66B629C37DFF7B0C91A24CDCA3" blockId="10.[151,1436,151,250]" box="[151,614,151,178]" pageId="10" pageNumber="11">
<heading id="D0CD2B0AB629C37DFF7B0C91A24CDCA3" bold="true" box="[151,614,151,178]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<emphasis id="B94E4074B629C37DFF7B0C91A24CDCA3" bold="true" box="[151,614,151,178]" pageId="10" pageNumber="11">
<taxonomicName id="4C3AE7E5B629C37DFF7B0C91A19CDCA0" authorityName="Bates" authorityYear="1888" box="[151,438,151,178]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fasciolata">
<emphasis id="B94E4074B629C37DFF7B0C91A19CDCA0" bold="true" box="[151,438,151,178]" italics="true" pageId="10" pageNumber="11">Cyclocephala fasciolata</emphasis>
</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C320CFEDB629C37DFF7B0CBDA1EAD86F" pageId="10" pageNumber="11" type="description">
<paragraph id="8B859C66B629C37DFF7B0CBDA13FDCEB" blockId="10.[151,1436,151,250]" pageId="10" pageNumber="11">
(
<figureCitation id="130180E3B629C37DFF730CBDA10DDCC7" box="[159,295,187,213]" captionStart="FIGURE 2" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[308,1279,181,1919]" captionTargetId="figure-15@13.[308,1279,181,1919]" captionTargetPageId="13" captionText="FIGURE 2AD. Cyclocephala fasciolata species group. A, Cyclocephala brevipennis male; B, Cyclocephala brevipennis female; C, Cyclocephala fasciolata male; D, Cyclocephala santandereana (holotype). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10100348" httpUri="https://zenodo.org/record/10100348/files/figure.png" pageId="10" pageNumber="11">Figs. 2AD</figureCitation>
;
<figureCitation id="130180E3B629C37DFEDE0CBDA158DCC7" box="[306,370,187,213]" captionStart="FIGURE 6" captionStartId="17.[151,250,1811,1835]" captionTargetBox="[163,1436,193,1776]" captionTargetId="figure-15@17.[151,1436,181,1787]" captionTargetPageId="17" captionText="FIGURE 6AH. Holotype males. A, Cyclocephala marqueti Dechambre habitus; B, Cyclocephala marqueti labels; C, Cyclocephala proxima Dechambre habitus; D, Cyclocephala proxima labels; E, Cyclocephala melane Bates habitus; F, Cyclocephala melane labels; G, Cyclocephala rogerpauli Moore, Branham, &amp; Cave, 2018 [= Cyclocephala nigra Dechambre] habitus; H, Cyclocephala rogerpauli labels." figureDoi="http://doi.org/10.5281/zenodo.10100360" httpUri="https://zenodo.org/record/10100360/files/figure.png" pageId="10" pageNumber="11">6EF</figureCitation>
;
<figureCitation id="130180E3B629C37DFE910CBDA1EFDCC7" box="[381,453,187,213]" captionStart="FIGURE 9" captionStartId="20.[151,250,1912,1936]" captionTargetBox="[151,1429,188,1878]" captionTargetId="figure-29@20.[151,1436,181,1888]" captionTargetPageId="20" captionText="FIGURE 9AD. Adult mandibles and maxillae. A, Cyclocephala brevipennis mandible (ventral view); B, Cyclocephala brevipennis maxilla (ventral view); C, Cyclocephala fasciolata mandible (dorsal view); D, Cyclocephala fasciolata maxilla (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.10100366" httpUri="https://zenodo.org/record/10100366/files/figure.png" pageId="10" pageNumber="11">9AD</figureCitation>
;
<figureCitation id="130180E3B629C37DFE3C0CBDA20EDCC7" box="[464,548,187,213]" captionStart="FIGURE 12" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[264,1334,202,1911]" captionTargetId="figure-15@23.[253,1334,181,1919]" captionTargetPageId="23" captionText="FIGURE 12AF. Adult labia. A, Cyclocephala marqueti; B, Cyclocephala proxima; C, Cyclocephala fasciolata; D, Cyclocephala brevipennis; E, Cyclocephala ligyrina; F, Cyclocephala arhuacana." figureDoi="http://doi.org/10.5281/zenodo.10100372" httpUri="https://zenodo.org/record/10100372/files/figure.png" pageId="10" pageNumber="11">12CD</figureCitation>
;
<figureCitation id="130180E3B629C37DFDDC0CBDA254DCC7" box="[560,638,187,213]" captionStart="FIGURE 13" captionStartId="24.[151,250,1821,1845]" captionTargetBox="[153,1423,181,1788]" captionTargetId="figure-29@24.[151,1436,181,1798]" captionTargetPageId="24" captionText="FIGURE 13AF. Adult epipharynxes. A, Cyclocephala marqueti ventral view; B, Cyclocephala marqueti dorsal view; C, Cyclocephala proxima ventral view; D, Cyclocephala proxima dorsal view; E, Cyclocephala brevipennis ventral view; F, Cyclocephala brevipennis dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10100377" httpUri="https://zenodo.org/record/10100377/files/figure.png" pageId="10" pageNumber="11">13EF</figureCitation>
;
<figureCitation id="130180E3B629C37DFD650CBDA2F7DCC7" box="[649,733,187,213]" captionStart="FIGURE 14" captionStartId="25.[151,250,1808,1832]" captionTargetBox="[161,1429,196,1774]" captionTargetId="figure-15@25.[151,1436,181,1785]" captionTargetPageId="25" captionText="FIGURE 14AF. Adult epipharynxes. A, Cyclocephala fasciolata ventral view; B, Cyclocephala fasciolata dorsal view; C, Cyclocephala ligyrina ventral view; D, Cyclocephala ligyrina dorsal view; E, Cyclocephala arhuacana ventral view; F, Cyclocephala arhuacana dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10100380" httpUri="https://zenodo.org/record/10100380/files/figure.png" pageId="10" pageNumber="11">14AB</figureCitation>
;
<figureCitation id="130180E3B629C37DFD040CBDA317DCC4" box="[744,829,187,214]" captionStart="FIGURE 15" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[219,1372,189,1907]" captionTargetId="figure-29@26.[206,1381,181,1919]" captionTargetPageId="26" captionText="FIGURE 15AJ. Adult male large claw (caudal view). A, Cyclcephala marqueti; B, Cyclocephala proxima; C, Cyclocephala brevipennis; D, Cyclocephala fasciolata; E, Cyclocephala santandereana (holotype); F, Cyclocephala dalensi; G, Cyclocephala kaszabi; H, Cyclocephala ligyrina; I, Cyclocephala rufa; J, Cyclocephala arhuacana (holotype). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10100382" httpUri="https://zenodo.org/record/10100382/files/figure.png" pageId="10" pageNumber="11">15CD</figureCitation>
;
<figureCitation id="130180E3B629C37DFCA40CBDA3B3DCC7" box="[840,921,187,213]" captionStart="FIGURE 16" captionStartId="27.[151,250,1676,1700]" captionTargetBox="[167,1426,192,1639]" captionTargetId="figure-15@27.[151,1436,181,1652]" captionTargetPageId="27" captionText="FIGURE 16AJ. Adult male large claw (lateral view). A, Cyclcephala marqueti; B, Cyclocephala proxima; C, Cyclocephala brevipennis; D, Cyclocephala fasciolata; E, Cyclocephala santandereana (holotype); F, Cyclocephala dalensi; G, Cyclocephala kaszabi; H, Cyclocephala ligyrina; I, Cyclocephala rufa; J, Cyclocephala arhuacana (holotype). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10100384" httpUri="https://zenodo.org/record/10100384/files/figure.png" pageId="10" pageNumber="11">16CE</figureCitation>
;
<figureCitation id="130180E3B629C37DFC480CBDA3DCDCC7" box="[932,1014,187,213]" captionStart="FIGURE 17" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[178,1404,181,1911]" captionTargetId="figure-29@28.[169,1418,181,1919]" captionTargetPageId="28" captionText="FIGURE 17AJ. Protarsomere 5 (caudal view). A, Cyclcephala marqueti; B, Cyclocephala proxima; C, Cyclocephala brevipennis; D, Cyclocephala fasciolata; E, Cyclocephala santandereana (holotype); F, Cyclocephala dalensi; G, Cyclocephala kaszabi; H, Cyclocephala ligyrina; I, Cyclocephala rufa; J, Cyclocephala arhuacana (holotype). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10100386" httpUri="https://zenodo.org/record/10100386/files/figure.png" pageId="10" pageNumber="11">17CE</figureCitation>
;
<figureCitation id="130180E3B629C37DFBED0CBDA47DDCC7" box="[1025,1111,187,213]" captionStart="FIGURE 18" captionStartId="29.[151,250,1846,1870]" captionTargetBox="[161,1428,191,1813]" captionTargetId="figure-15@29.[151,1436,181,1822]" captionTargetPageId="29" captionText="FIGURE 18AM. Protibiae (dorsal view). A, Cyclocephala marqueti (male); B, Cyclocephala proxima (male); C, Cyclocephala proxima (female); D, Cyclocephala brevipennis (male); E, Cyclocephala brevipennis (female); F, Cyclocephala fasciolata; G, Cyclocephala santandereana (holotype); H, Cyclocephala dalensi; I, Cyclocephala kaszabi; J, Cyclocephala ligyrina; K, Cyclocephala rufa; L, Cyclocephala arhuacana (holotype); M, Cyclocephala arhuacana (allotype). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10100388" httpUri="https://zenodo.org/record/10100388/files/figure.png" pageId="10" pageNumber="11">18DG</figureCitation>
;
<figureCitation id="130180E3B629C37DFB8E0CBDA4BFDCC7" box="[1122,1173,187,213]" captionStart="FIGURE 20" captionStartId="31.[151,250,1913,1937]" captionTargetBox="[196,1389,161,1875]" captionTargetId="figure-15@31.[185,1402,151,1890]" captionTargetPageId="31" captionText="FIGURE 20AG. Female epipleura (ventral view). A, Cyclocephala marqueti; B, Cyclocephala proxima; C, Cyclocephala rogerpauli; D, Cyclocephala brevipennis; E, Cyclocephala kaszabi; F, Cyclocephala ligyrina; G, Cyclocephala arhuacana." figureDoi="http://doi.org/10.5281/zenodo.10100394" httpUri="https://zenodo.org/record/10100394/files/figure.png" pageId="10" pageNumber="11">20D</figureCitation>
;
<figureCitation id="130180E3B629C37DFB4D0CBDA4D9DCC4" box="[1185,1267,187,214]" captionStart="FIGURE 24" captionStartId="35.[151,250,1840,1864]" captionTargetBox="[152,1423,181,1807]" captionTargetId="figure-15@35.[151,1436,181,1816]" captionTargetPageId="35" captionText="FIGURE 24AH. Pygidia (dorsal view). A, Cyclocephala marqueti; B, Cyclocephala proxima; C, Cyclocephala rogerpauli; D, Cyclocephala brevipennis; E, Cyclocephala santandereana (holotype); F, Cyclocephala dalensi; G, Cyclocephala kaszabi; H, Cyclocephala ligyrina." figureDoi="http://doi.org/10.5281/zenodo.10100402" httpUri="https://zenodo.org/record/10100402/files/figure.png" pageId="10" pageNumber="11">24D, E</figureCitation>
;
<figureCitation id="130180E3B629C37DFB120CBDA579DCC4" box="[1278,1363,187,214]" captionStart="FIGURE 25" captionStartId="39.[151,250,1962,1986]" captionTargetBox="[154,1436,777,1932]" captionTargetId="figure-280@39.[151,1436,777,1939]" captionTargetPageId="39" captionText="FIGURE 25AD. Pygidia and heads (dorsal view). A, Cyclocephala rufa pygidium; B, Cyclocephala arhuacana (holotype) pygidium; C, Cyclocephala brevipennis head (dorsal view); D, Cyclocephala santandereana (holotype) head (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.10100404" httpUri="https://zenodo.org/record/10100404/files/figure.png" pageId="10" pageNumber="11">25CD</figureCitation>
;
<figureCitation id="130180E3B629C37DFAB20CBDA080DCE8" captionStart="FIGURE 27" captionStartId="41.[151,250,1885,1909]" captionTargetBox="[205,1382,190,1855]" captionTargetId="figure-15@41.[193,1395,181,1861]" captionTargetPageId="41" captionText="FIGURE 27AF. Parameres and phallobases, Cyclocephala fasciolata species group. A, Cyclocephala fasciolata parameres, caudal view; B, Cyclocephala fasciolata phallobase, lateral view; C, Cyclocephala brevipennis parameres, caudal view; D, Cyclocephala brevipennis phallobase, lateral view; E, Cyclocephala santandereana parameres, caudal view; F, Cyclocephala santandereana phallobase, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10100409" httpUri="https://zenodo.org/record/10100409/files/figure.png" pageId="10" pageNumber="11">27A F</figureCitation>
; 32F; 39)
</paragraph>
<paragraph id="8B859C66B629C37DFF7B0D21A46FD827" blockId="10.[151,1437,295,1330]" pageId="10" pageNumber="11">
<emphasis id="B94E4074B629C37DFF7B0D21A10BDD53" bold="true" box="[151,289,295,321]" pageId="10" pageNumber="11">Description</emphasis>
. The
<taxonomicName id="4C3AE7E5B629C37DFE8E0D21A25BDD53" authorityName="Bates" authorityYear="1888" box="[354,625,295,321]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fasciolata">
<emphasis id="B94E4074B629C37DFE8E0D21A25BDD53" box="[354,625,295,321]" italics="true" pageId="10" pageNumber="11">Cyclocephala fasciolata</emphasis>
</taxonomicName>
species group consists of four species and is characterized by following combination of characters. Color black except pronotum with 2 transverse, slightly oblique, reddish-orange bands; body broad, subrectangular (
<figureCitation id="130180E3B629C37DFE370D69A27FDD9B" box="[475,597,367,393]" captionStart="FIGURE 2" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[308,1279,181,1919]" captionTargetId="figure-15@13.[308,1279,181,1919]" captionTargetPageId="13" captionText="FIGURE 2AD. Cyclocephala fasciolata species group. A, Cyclocephala brevipennis male; B, Cyclocephala brevipennis female; C, Cyclocephala fasciolata male; D, Cyclocephala santandereana (holotype). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10100348" httpUri="https://zenodo.org/record/10100348/files/figure.png" pageId="10" pageNumber="11">Fig. 2AD</figureCitation>
) (
<taxonomicName id="4C3AE7E5B629C37DFD830D76A30DDD98" authority="Bates" authorityName="Bates" authorityYear="1888" box="[623,807,367,394]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="melane">
<emphasis id="B94E4074B629C37DFD830D76A2A8DD98" box="[623,642,368,394]" italics="true" pageId="10" pageNumber="11">C</emphasis>
.
<emphasis id="B94E4074B629C37DFD7C0D69A2CBDD9B" box="[656,737,367,393]" italics="true" pageId="10" pageNumber="11">melane</emphasis>
Bates
</taxonomicName>
is totally black (
<figureCitation id="130180E3B629C37DFC0A0D69A414DD98" box="[998,1086,367,394]" captionStart="FIGURE 6" captionStartId="17.[151,250,1811,1835]" captionTargetBox="[163,1436,193,1776]" captionTargetId="figure-15@17.[151,1436,181,1787]" captionTargetPageId="17" captionText="FIGURE 6AH. Holotype males. A, Cyclocephala marqueti Dechambre habitus; B, Cyclocephala marqueti labels; C, Cyclocephala proxima Dechambre habitus; D, Cyclocephala proxima labels; E, Cyclocephala melane Bates habitus; F, Cyclocephala melane labels; G, Cyclocephala rogerpauli Moore, Branham, &amp; Cave, 2018 [= Cyclocephala nigra Dechambre] habitus; H, Cyclocephala rogerpauli labels." figureDoi="http://doi.org/10.5281/zenodo.10100360" httpUri="https://zenodo.org/record/10100360/files/figure.png" pageId="10" pageNumber="11">Fig. 6E</figureCitation>
).
<emphasis id="B94E4074B629C37DFBBE0D69A4B1DD9B" bold="true" box="[1106,1179,367,393]" pageId="10" pageNumber="11">Head:</emphasis>
Frons mostly impunctate, frontoclypeal suture slightly indicated (
<figureCitation id="130180E3B629C37DFD6D0D95A2D4DDBF" box="[641,766,403,429]" captionStart="FIGURE 2" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[308,1279,181,1919]" captionTargetId="figure-15@13.[308,1279,181,1919]" captionTargetPageId="13" captionText="FIGURE 2AD. Cyclocephala fasciolata species group. A, Cyclocephala brevipennis male; B, Cyclocephala brevipennis female; C, Cyclocephala fasciolata male; D, Cyclocephala santandereana (holotype). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10100348" httpUri="https://zenodo.org/record/10100348/files/figure.png" pageId="10" pageNumber="11">Fig. 2AD</figureCitation>
). Clypeus with punctures dense to sparse, clypeal apex parabolic, slightly reflexed (
<figureCitation id="130180E3B629C37DFE4C0DB1A236DDC3" box="[416,540,439,465]" captionStart="FIGURE 2" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[308,1279,181,1919]" captionTargetId="figure-15@13.[308,1279,181,1919]" captionTargetPageId="13" captionText="FIGURE 2AD. Cyclocephala fasciolata species group. A, Cyclocephala brevipennis male; B, Cyclocephala brevipennis female; C, Cyclocephala fasciolata male; D, Cyclocephala santandereana (holotype). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10100348" httpUri="https://zenodo.org/record/10100348/files/figure.png" pageId="10" pageNumber="11">Fig. 2AD</figureCitation>
). Mandible without teeth, inner apex slight concave (
<figureCitation id="130180E3B629C37DFB8B0DB1A4F5DDC3" box="[1127,1247,439,465]" captionStart="FIGURE 9" captionStartId="20.[151,250,1912,1936]" captionTargetBox="[151,1429,188,1878]" captionTargetId="figure-29@20.[151,1436,181,1888]" captionTargetPageId="20" captionText="FIGURE 9AD. Adult mandibles and maxillae. A, Cyclocephala brevipennis mandible (ventral view); B, Cyclocephala brevipennis maxilla (ventral view); C, Cyclocephala fasciolata mandible (dorsal view); D, Cyclocephala fasciolata maxilla (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.10100366" httpUri="https://zenodo.org/record/10100366/files/figure.png" pageId="10" pageNumber="11">Fig. 9A, C</figureCitation>
); labium densely setose, apex widely notched, paraglossa slightly developed (
<figureCitation id="130180E3B629C37DFCA20DDDA3F0DDE7" box="[846,986,475,501]" captionStart="FIGURE 12" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[264,1334,202,1911]" captionTargetId="figure-15@23.[253,1334,181,1919]" captionTargetPageId="23" captionText="FIGURE 12AF. Adult labia. A, Cyclocephala marqueti; B, Cyclocephala proxima; C, Cyclocephala fasciolata; D, Cyclocephala brevipennis; E, Cyclocephala ligyrina; F, Cyclocephala arhuacana." figureDoi="http://doi.org/10.5281/zenodo.10100372" httpUri="https://zenodo.org/record/10100372/files/figure.png" pageId="10" pageNumber="11">Fig. 12CD</figureCitation>
); maxilla with galea developed, with 6 teeth, lacinia not developed, apex with slender, long setae (
<figureCitation id="130180E3B629C37DFCF30DF9A3BDDE08" box="[799,919,511,538]" captionStart="FIGURE 9" captionStartId="20.[151,250,1912,1936]" captionTargetBox="[151,1429,188,1878]" captionTargetId="figure-29@20.[151,1436,181,1888]" captionTargetPageId="20" captionText="FIGURE 9AD. Adult mandibles and maxillae. A, Cyclocephala brevipennis mandible (ventral view); B, Cyclocephala brevipennis maxilla (ventral view); C, Cyclocephala fasciolata mandible (dorsal view); D, Cyclocephala fasciolata maxilla (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.10100366" httpUri="https://zenodo.org/record/10100366/files/figure.png" pageId="10" pageNumber="11">Fig. 9B, D</figureCitation>
). Epipharynx rectangular, densely setose, apex projecting at center (ventral view,
<figureCitation id="130180E3B629C37DFDF90E25A28BDE2F" box="[533,673,547,573]" captionStart="FIGURE 13" captionStartId="24.[151,250,1821,1845]" captionTargetBox="[153,1423,181,1788]" captionTargetId="figure-29@24.[151,1436,181,1798]" captionTargetPageId="24" captionText="FIGURE 13AF. Adult epipharynxes. A, Cyclocephala marqueti ventral view; B, Cyclocephala marqueti dorsal view; C, Cyclocephala proxima ventral view; D, Cyclocephala proxima dorsal view; E, Cyclocephala brevipennis ventral view; F, Cyclocephala brevipennis dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10100377" httpUri="https://zenodo.org/record/10100377/files/figure.png" pageId="10" pageNumber="11">Figs. 13EF</figureCitation>
;
<figureCitation id="130180E3B629C37DFD400E25A328DE2F" box="[684,770,547,573]" captionStart="FIGURE 14" captionStartId="25.[151,250,1808,1832]" captionTargetBox="[161,1429,196,1774]" captionTargetId="figure-15@25.[151,1436,181,1785]" captionTargetPageId="25" captionText="FIGURE 14AF. Adult epipharynxes. A, Cyclocephala fasciolata ventral view; B, Cyclocephala fasciolata dorsal view; C, Cyclocephala ligyrina ventral view; D, Cyclocephala ligyrina dorsal view; E, Cyclocephala arhuacana ventral view; F, Cyclocephala arhuacana dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10100380" httpUri="https://zenodo.org/record/10100380/files/figure.png" pageId="10" pageNumber="11">14AB</figureCitation>
). Antenna with 10 antennomeres, club subequal in length to antennomeres 27 (slightly shorter in female)
<emphasis id="B94E4074B629C37DFD4C0E41A308DE73" bold="true" box="[672,802,583,609]" pageId="10" pageNumber="11">Pronotum:</emphasis>
Surface with punctures moderate in density, moderately large to large (especially on sides). Base with marginal bead (Fig. 32A).
<emphasis id="B94E4074B629C37DFC550E6DA424DE97" bold="true" box="[953,1038,619,645]" pageId="10" pageNumber="11">Elytra:</emphasis>
Surface with punctacte rows, punctures of striae and intervals similar to those of pronotum; entire surface often coarsely and transversely wrinkled obscuring rows of punctures. Wing vein AA
<subScript id="17BE9E23B629C37DFD6D0EC0A2A3DEC6" attach="left" box="[641,649,710,724]" fontSize="6" pageId="10" pageNumber="11">3</subScript>
equal length to AA
<subScript id="17BE9E23B629C37DFC8E0EC0A340DEC6" attach="left" box="[866,874,710,724]" fontSize="6" pageId="10" pageNumber="11">4</subScript>
(
<figureCitation id="130180E3B629C37DFC940EB5A42EDEDF" box="[888,1028,691,717]" captionStart="FIGURE 22" captionStartId="33.[151,250,1736,1760]" captionTargetBox="[155,1423,191,1701]" captionTargetId="figure-15@33.[151,1436,181,1712]" captionTargetPageId="33" captionText="FIGURE 22AD. Wing. Cyclocephala fasciolata species group. A, Cyclocephala brevipennis; B, C, D, Cyclocephala fasciolata." figureDoi="http://doi.org/10.5281/zenodo.10100398" httpUri="https://zenodo.org/record/10100398/files/figure.png" pageId="10" pageNumber="11">Fig. 22AD</figureCitation>
). Anterior edge of wing distal of apical hinge glabrous (
<figureCitation id="130180E3B629C37DFE9F0ED1A1F1DEE3" box="[371,475,727,753]" captionStart="FIGURE 22" captionStartId="33.[151,250,1736,1760]" captionTargetBox="[155,1423,191,1701]" captionTargetId="figure-15@33.[151,1436,181,1712]" captionTargetPageId="33" captionText="FIGURE 22AD. Wing. Cyclocephala fasciolata species group. A, Cyclocephala brevipennis; B, C, D, Cyclocephala fasciolata." figureDoi="http://doi.org/10.5281/zenodo.10100398" httpUri="https://zenodo.org/record/10100398/files/figure.png" pageId="10" pageNumber="11">Fig. 22D</figureCitation>
).
<emphasis id="B94E4074B629C37DFE020ED1A240DEE3" bold="true" box="[494,618,727,753]" pageId="10" pageNumber="11">Pygidium:</emphasis>
Surface usually moderately densely punctate; punctures moderate in size to more usually moderately large, ocellate, setigerous; setae short, tawny, moderately dense (
<figureCitation id="130180E3B629C37DFB730EFDA50CDF07" box="[1183,1318,763,789]" captionStart="FIGURE 24" captionStartId="35.[151,250,1840,1864]" captionTargetBox="[152,1423,181,1807]" captionTargetId="figure-15@35.[151,1436,181,1816]" captionTargetPageId="35" captionText="FIGURE 24AH. Pygidia (dorsal view). A, Cyclocephala marqueti; B, Cyclocephala proxima; C, Cyclocephala rogerpauli; D, Cyclocephala brevipennis; E, Cyclocephala santandereana (holotype); F, Cyclocephala dalensi; G, Cyclocephala kaszabi; H, Cyclocephala ligyrina." figureDoi="http://doi.org/10.5281/zenodo.10100402" httpUri="https://zenodo.org/record/10100402/files/figure.png" pageId="10" pageNumber="11">Fig. 24DE</figureCitation>
). In lateral view, surface in males evenly convex, weakly convex to nearly flat in females.
<emphasis id="B94E4074B629C37DFBF10F19A474DF2B" bold="true" box="[1053,1118,799,825]" pageId="10" pageNumber="11">Legs:</emphasis>
Protibia bidentate in males, basal tooth slightly removed from others (
<figureCitation id="130180E3B629C37DFD810F45A33CDF4F" box="[621,790,835,861]" captionStart="FIGURE 18" captionStartId="29.[151,250,1846,1870]" captionTargetBox="[161,1428,191,1813]" captionTargetId="figure-15@29.[151,1436,181,1822]" captionTargetPageId="29" captionText="FIGURE 18AM. Protibiae (dorsal view). A, Cyclocephala marqueti (male); B, Cyclocephala proxima (male); C, Cyclocephala proxima (female); D, Cyclocephala brevipennis (male); E, Cyclocephala brevipennis (female); F, Cyclocephala fasciolata; G, Cyclocephala santandereana (holotype); H, Cyclocephala dalensi; I, Cyclocephala kaszabi; J, Cyclocephala ligyrina; K, Cyclocephala rufa; L, Cyclocephala arhuacana (holotype); M, Cyclocephala arhuacana (allotype). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10100388" httpUri="https://zenodo.org/record/10100388/files/figure.png" pageId="10" pageNumber="11">Fig. 18D, FG</figureCitation>
). Protarsus in male enlarged, tarsomeres 24 successively gradually larger, fourth with small tooth on ventral surface, fifth large, curved, with small angulation at base on ventral side (
<figureCitation id="130180E3B629C37DFECA0F8DA19CDFB7" box="[294,438,907,933]" captionStart="FIGURE 17" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[178,1404,181,1911]" captionTargetId="figure-29@28.[169,1418,181,1919]" captionTargetPageId="28" captionText="FIGURE 17AJ. Protarsomere 5 (caudal view). A, Cyclcephala marqueti; B, Cyclocephala proxima; C, Cyclocephala brevipennis; D, Cyclocephala fasciolata; E, Cyclocephala santandereana (holotype); F, Cyclocephala dalensi; G, Cyclocephala kaszabi; H, Cyclocephala ligyrina; I, Cyclocephala rufa; J, Cyclocephala arhuacana (holotype). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10100386" httpUri="https://zenodo.org/record/10100386/files/figure.png" pageId="10" pageNumber="11">Figs. 17CE</figureCitation>
), all tarsomeres longitudinally striate on dorsal face, median claw large, curved, apex cleft (
<figureCitation id="130180E3B629C37DFF4C0FA9A10FDFD8" box="[160,293,943,970]" captionStart="FIGURE 15" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[219,1372,189,1907]" captionTargetId="figure-29@26.[206,1381,181,1919]" captionTargetPageId="26" captionText="FIGURE 15AJ. Adult male large claw (caudal view). A, Cyclcephala marqueti; B, Cyclocephala proxima; C, Cyclocephala brevipennis; D, Cyclocephala fasciolata; E, Cyclocephala santandereana (holotype); F, Cyclocephala dalensi; G, Cyclocephala kaszabi; H, Cyclocephala ligyrina; I, Cyclocephala rufa; J, Cyclocephala arhuacana (holotype). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10100382" httpUri="https://zenodo.org/record/10100382/files/figure.png" pageId="10" pageNumber="11">Fig. 15CE</figureCitation>
); external edge with some striations (
<figureCitation id="130180E3B629C37DFD2E0FA9A362DFD8" box="[706,840,943,970]" captionStart="FIGURE 16" captionStartId="27.[151,250,1676,1700]" captionTargetBox="[167,1426,192,1639]" captionTargetId="figure-15@27.[151,1436,181,1652]" captionTargetPageId="27" captionText="FIGURE 16AJ. Adult male large claw (lateral view). A, Cyclcephala marqueti; B, Cyclocephala proxima; C, Cyclocephala brevipennis; D, Cyclocephala fasciolata; E, Cyclocephala santandereana (holotype); F, Cyclocephala dalensi; G, Cyclocephala kaszabi; H, Cyclocephala ligyrina; I, Cyclocephala rufa; J, Cyclocephala arhuacana (holotype). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10100384" httpUri="https://zenodo.org/record/10100384/files/figure.png" pageId="10" pageNumber="11">Fig. 16CE</figureCitation>
). Metatarsus subequal in length to metatibia. Females with protibia tridentate (
<figureCitation id="130180E3B629C37DFE4B0FD5A226DFFF" box="[423,524,979,1005]" captionStart="FIGURE 18" captionStartId="29.[151,250,1846,1870]" captionTargetBox="[161,1428,191,1813]" captionTargetId="figure-15@29.[151,1436,181,1822]" captionTargetPageId="29" captionText="FIGURE 18AM. Protibiae (dorsal view). A, Cyclocephala marqueti (male); B, Cyclocephala proxima (male); C, Cyclocephala proxima (female); D, Cyclocephala brevipennis (male); E, Cyclocephala brevipennis (female); F, Cyclocephala fasciolata; G, Cyclocephala santandereana (holotype); H, Cyclocephala dalensi; I, Cyclocephala kaszabi; J, Cyclocephala ligyrina; K, Cyclocephala rufa; L, Cyclocephala arhuacana (holotype); M, Cyclocephala arhuacana (allotype). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10100388" httpUri="https://zenodo.org/record/10100388/files/figure.png" pageId="10" pageNumber="11">Fig. 18E</figureCitation>
).
<emphasis id="B94E4074B629C37DFDF30FD5A253DFFF" bold="true" box="[543,633,979,1005]" pageId="10" pageNumber="11">Venter:</emphasis>
Prosternal process moderately long, columnar, apex obliquely flattened in a transverse oval with anterior 2/3 to 4/5 raised into convex “button”.
<emphasis id="B94E4074B629C37DFC0C0FF1A440D803" bold="true" box="[992,1130,1015,1041]" pageId="10" pageNumber="11">Parameres:</emphasis>
As in
<figureCitation id="130180E3B629C37DFB550FF1A517D803" box="[1209,1341,1015,1041]" captionStart="FIGURE 27" captionStartId="41.[151,250,1885,1909]" captionTargetBox="[205,1382,190,1855]" captionTargetId="figure-15@41.[193,1395,181,1861]" captionTargetPageId="41" captionText="FIGURE 27AF. Parameres and phallobases, Cyclocephala fasciolata species group. A, Cyclocephala fasciolata parameres, caudal view; B, Cyclocephala fasciolata phallobase, lateral view; C, Cyclocephala brevipennis parameres, caudal view; D, Cyclocephala brevipennis phallobase, lateral view; E, Cyclocephala santandereana parameres, caudal view; F, Cyclocephala santandereana phallobase, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10100409" httpUri="https://zenodo.org/record/10100409/files/figure.png" pageId="10" pageNumber="11">Fig. 27AF</figureCitation>
. Female similar to male, but protarsus simple. Gonocoxite and gonosternite fused (Fig. 32F).
</paragraph>
<paragraph id="8B859C66B629C37DFF2B0839A1EAD86F" blockId="10.[151,1437,295,1330]" pageId="10" pageNumber="11">
<emphasis id="B94E4074B629C37DFF2B0839A175D84B" bold="true" box="[199,351,1087,1113]" pageId="10" pageNumber="11">Composition</emphasis>
. This group includes the following five species, of which, two species are distributed in
<collectingCountry id="F32DDCF6B629C37DFAAB0839A096D86F" name="Colombia" pageId="10" pageNumber="11">Colombia</collectingCountry>
, and are treated below:
</paragraph>
</subSubSection>
<subSubSection id="C320CFEDB629C37DFF7B0881A29CD923" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="8B859C66B629C37DFF7B0881A3CCD8B3" blockId="10.[151,1437,295,1330]" box="[151,998,1158,1186]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB2ABB48B629C37DFF7B0881A3CCD8B3" box="[151,998,1158,1186]" pageId="10" pageNumber="11">
<treatmentCitation id="0A9BBA77B629C37DFF7B0881A2F1D8B0" author="Ratcliffe, B. C. &amp; Cave, R. D." box="[151,731,1159,1186]" page="155" pageId="10" pageNumber="11" year="2002">
<taxonomicName id="4C3AE7E5B629C37DFF7B0881A2F1D8B0" authority="Ratcliffe &amp; Cave, 2002: 155" authorityName="Ratcliffe &amp; Cave" authorityPageNumber="155" authorityYear="2002" box="[151,731,1159,1186]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="abrelata">
<emphasis id="B94E4074B629C37DFF7B0881A1BCD8B3" box="[151,406,1159,1185]" italics="true" pageId="10" pageNumber="11">Cyclocephala abrelata</emphasis>
<bibRefCitation id="EFABE197B629C37DFE710881A2F1D8B0" author="Ratcliffe, B. C. &amp; Cave, R. D." box="[413,731,1159,1186]" pageId="10" pageNumber="11" pagination="152 - 157" refId="ref24130" refString="Ratcliffe, B. C. &amp; Cave, R. D. (2002) New species of Cyclocephala from Honduras and El Salvador (Coleoptera: Scarabaeidae: Dynastinae: Cyclocephalini). The Coleopterist Bulletin, 56, 152 - 157. https: // doi. org / 10.1649 / 0010 - 065 X (2002) 056 [0152: NSOCFH] 2.0. CO; 2" type="journal article" year="2002">Ratcliffe &amp; Cave, 2002: 155</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[original combination].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B859C66B629C37DFF7B08ADA3BDD8D7" blockId="10.[151,1437,295,1330]" box="[151,919,1194,1222]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB2ABB48B629C37DFF7B08ADA3BDD8D7" box="[151,919,1194,1222]" pageId="10" pageNumber="11">
<treatmentCitation id="0A9BBA77B629C37DFF7B08ADA2A6D8D7" author="Endrodi, S." box="[151,652,1195,1222]" page="69" pageId="10" pageNumber="11" year="1985">
<taxonomicName id="4C3AE7E5B629C37DFF7B08ADA2A6D8D7" authority="Endrodi, 1985: 69" authorityName="Endrodi" authorityPageNumber="69" authorityYear="1985" box="[151,652,1195,1222]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="B94E4074B629C37DFF7B08ADA190D8D7" box="[151,442,1195,1221]" italics="true" pageId="10" pageNumber="11">Cyclocephala brevipennis</emphasis>
<bibRefCitation id="EFABE197B629C37DFE2D08ADA2A6D8D7" author="Endrodi, S." box="[449,652,1195,1222]" pageId="10" pageNumber="11" refId="ref23141" refString="Endrodi, S. (1985) The Dynastinae of the World. Dr. W. Junk, Dordrecht, 800 pp." type="book" year="1985">Endrődi, 1985: 69</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[original combination].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B859C66B629C37DFF7B08C9A35CD8FB" blockId="10.[151,1437,295,1330]" box="[151,886,1230,1258]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB2ABB48B629C37DFF7B08C9A35CD8FB" box="[151,886,1230,1258]" pageId="10" pageNumber="11">
<treatmentCitation id="0A9BBA77B629C37DFF7B08C9A241D8FB" author="Bates, H. W." box="[151,619,1231,1258]" page="301" pageId="10" pageNumber="11" year="1888">
<taxonomicName id="4C3AE7E5B629C37DFF7B08C9A241D8FB" authority="Bates, 1888: 301" authorityName="Bates" authorityPageNumber="301" authorityYear="1888" box="[151,619,1231,1258]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fasciolata">
<emphasis id="B94E4074B629C37DFF7B08C9A18CD8FB" box="[151,422,1231,1257]" italics="true" pageId="10" pageNumber="11">Cyclocephala fasciolata</emphasis>
<bibRefCitation id="EFABE197B629C37DFE4208D6A241D8FB" author="Bates, H. W." box="[430,619,1231,1258]" pageId="10" pageNumber="11" pagination="296 - 342" refId="ref22786" refString="Bates, H. W. (1888) Pectinicornia and Lamellicornia, family Dynastidae. In: Salvin, O. &amp; Godman, F. D. (editors) Biologia Centrali - Americana. Insecta, Coleoptera. Vol. 2. Part 2. R. H. Porter, London, pp. 296 - 342." type="book chapter" year="1888">Bates, 1888: 301</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[original combination].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B859C66B629C37DFF7B08F5A372D91F" blockId="10.[151,1437,295,1330]" box="[151,856,1266,1294]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB2ABB48B629C37DFF7B08F5A372D91F" box="[151,856,1266,1294]" pageId="10" pageNumber="11">
<treatmentCitation id="0A9BBA77B629C37DFF7B08F5A267D91F" author="Bates, H. W." box="[151,589,1267,1294]" page="310" pageId="10" pageNumber="11" year="1888">
<taxonomicName id="4C3AE7E5B629C37DFF7B08F5A267D91F" authority="Bates, 1888: 310" authorityName="Bates" authorityPageNumber="310" authorityYear="1888" box="[151,589,1267,1294]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="melane">
<emphasis id="B94E4074B629C37DFF7B08F5A1A3D91F" box="[151,393,1267,1293]" italics="true" pageId="10" pageNumber="11">Cyclocephala melane</emphasis>
<bibRefCitation id="EFABE197B629C37DFE7C08F2A267D91F" author="Bates, H. W." box="[400,589,1267,1294]" pageId="10" pageNumber="11" pagination="296 - 342" refId="ref22786" refString="Bates, H. W. (1888) Pectinicornia and Lamellicornia, family Dynastidae. In: Salvin, O. &amp; Godman, F. D. (editors) Biologia Centrali - Americana. Insecta, Coleoptera. Vol. 2. Part 2. R. H. Porter, London, pp. 296 - 342." type="book chapter" year="1888">Bates, 1888: 310</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[original combination].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B859C66B629C37DFF7B0911A29CD923" blockId="10.[151,1437,295,1330]" box="[151,694,1303,1330]" pageId="10" pageNumber="11">
<taxonomicName id="4C3AE7E5B629C37DFF7B0911A20FD923" authority="Neita" authorityName="Neita-Moreno" authorityYear="2021" box="[151,549,1303,1329]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="santandereana" status="sp. nov.">
<emphasis id="B94E4074B629C37DFF7B0911A1F7D923" box="[151,477,1303,1329]" italics="true" pageId="10" pageNumber="11">Cyclocephala santandereana</emphasis>
Neita
</taxonomicName>
,
<taxonomicNameLabel id="A27DFD0FB629C37DFDC3091EA29CD923" box="[559,694,1303,1330]" pageId="10" pageNumber="11" rank="species">new species</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>