A review of the black species of Cyclocephala Dejean (Coleoptera: Scarabaeidae Dynastinae) from Colombia Author Neita-Moreno, Jhon C. text Zootaxa 2021 2021-08-24 5026 1 1 58 journal article 10.11646/zootaxa.5026.1.1 1175-5326 10100344 07E0C922-6B0F-4916-85C2-AD95146A8F1E Cyclocephala fasciolata species group ( Figs. 2A–D ; 6E–F ; 9A–D ; 12C–D ; 13E–F ; 14A–B ; 15C–D ; 16C–E ; 17C–E ; 18D–G ; 20D ; 24D, E ; 25C–D ; 27A– F ; 32F; 39) Description . The Cyclocephala fasciolata species group consists of four species and is characterized by following combination of characters. Color black except pronotum with 2 transverse, slightly oblique, reddish-orange bands; body broad, subrectangular ( Fig. 2A–D ) ( C . melane Bates is totally black ( Fig. 6E ). Head: Frons mostly impunctate, frontoclypeal suture slightly indicated ( Fig. 2A–D ). Clypeus with punctures dense to sparse, clypeal apex parabolic, slightly reflexed ( Fig. 2A–D ). Mandible without teeth, inner apex slight concave ( Fig. 9A, C ); labium densely setose, apex widely notched, paraglossa slightly developed ( Fig. 12C–D ); maxilla with galea developed, with 6 teeth, lacinia not developed, apex with slender, long setae ( Fig. 9B, D ). Epipharynx rectangular, densely setose, apex projecting at center (ventral view, Figs. 13E–F ; 14A–B ). Antenna with 10 antennomeres, club subequal in length to antennomeres 2–7 (slightly shorter in female) Pronotum: Surface with punctures moderate in density, moderately large to large (especially on sides). Base with marginal bead (Fig. 32A). Elytra: Surface with punctacte rows, punctures of striae and intervals similar to those of pronotum; entire surface often coarsely and transversely wrinkled obscuring rows of punctures. Wing vein AA 3 equal length to AA 4 ( Fig. 22A–D ). Anterior edge of wing distal of apical hinge glabrous ( Fig. 22D ). Pygidium: Surface usually moderately densely punctate; punctures moderate in size to more usually moderately large, ocellate, setigerous; setae short, tawny, moderately dense ( Fig. 24D–E ). In lateral view, surface in males evenly convex, weakly convex to nearly flat in females. Legs: Protibia bidentate in males, basal tooth slightly removed from others ( Fig. 18D, F–G ). Protarsus in male enlarged, tarsomeres 2–4 successively gradually larger, fourth with small tooth on ventral surface, fifth large, curved, with small angulation at base on ventral side ( Figs. 17C–E ), all tarsomeres longitudinally striate on dorsal face, median claw large, curved, apex cleft ( Fig. 15C–E ); external edge with some striations ( Fig. 16C–E ). Metatarsus subequal in length to metatibia. Females with protibia tridentate ( Fig. 18E ). Venter: Prosternal process moderately long, columnar, apex obliquely flattened in a transverse oval with anterior 2/3 to 4/5 raised into convex “button”. Parameres: As in Fig. 27A–F . Female similar to male, but protarsus simple. Gonocoxite and gonosternite fused (Fig. 32F). Composition . This group includes the following five species, of which, two species are distributed in Colombia , and are treated below: Cyclocephala abrelata Ratcliffe & Cave, 2002: 155 [original combination]. Cyclocephala brevipennis Endrődi, 1985: 69 [original combination]. Cyclocephala fasciolata Bates, 1888: 301 [original combination]. Cyclocephala melane Bates, 1888: 310 [original combination]. Cyclocephala santandereana Neita , new species