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<document id="FB7EC66189EF018DBB58E2C73FD96A0A" ID-CLB-Dataset="55667" ID-DOI="10.26107/RBZ-2020-0097" ID-GBIF-Dataset="6cb155ce-8b9f-48ce-8e6c-b0dc00f8b1cc" ID-ISSN="2345-7600" ID-Zenodo-Dep="5351295" ID-ZooBank="815E4670-B063-4FD8-B31E-3AD89B3A7942" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1630205841501" checkinUser="diego" docAuthor="Schubart, Christoph D. &amp; Ng, Peter K. L." docDate="2020" docId="49152B56FFFCBA3EFC29FA8DFD7DFF44" docLanguage="en" docName="RafflesBZool.68.891-994.pdf" docOrigin="Raffles Bulletin of Zoology 68" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Danarma Schubart &amp; Ng 2020, new genus" docType="treatment" docUuid="8ABA1C2F-54C4-4A70-8646-9AF6EE2EBBF3" docUuidSource="ZooBank" docVersion="10" lastPageNumber="921" masterDocId="B52C532EFFE1BA21FFDDFFB3FF8DFFB1" masterDocTitle="Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species" masterLastPageNumber="994" masterPageNumber="891" pageNumber="920" updateTime="1699203722016" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="829F1B4BF3813A14C2A82919AAF9145E">Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species</mods:title>
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<mods:namePart id="C5C55ED9920074DBEA05EAE74481182D">Schubart, Christoph D.</mods:namePart>
<mods:affiliation id="9E0987F26888536A8591152D7272FCF8">Zoology &amp; Evolution, University of Regensburg, 93040 Regensburg, Germany</mods:affiliation>
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<mods:namePart id="E6EE23BA9A21DB8669B78A47EDF9C26B">Ng, Peter K. L.</mods:namePart>
<mods:affiliation id="27AFB8F554281763BF17C7A2E74BCF68">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore</mods:affiliation>
<mods:nameIdentifier id="C5F21A709FE7F08B6A47704673371751" type="email">peterng@nus.edu.sg</mods:nameIdentifier>
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<treatment id="49152B56FFFCBA3EFC29FA8DFD7DFF44" ID-DOI="http://doi.org/10.5281/zenodo.5449845" ID-GBIF-Taxon="195197152" ID-Zenodo-Dep="5449845" LSID="urn:lsid:zoobank.org:act:8ABA1C2F-54C4-4A70-8646-9AF6EE2EBBF3" httpUri="http://treatment.plazi.org/id/49152B56FFFCBA3EFC29FA8DFD7DFF44" lastPageId="31" lastPageNumber="921" pageId="29" pageNumber="920">
<subSubSection id="89A6C9CBFFFCBA3CFC29FA8DFB69FAE4" box="[1012,1252,1341,1365]" pageId="29" pageNumber="920" type="nomenclature">
<paragraph id="C1039A40FFFCBA3CFC29FA8DFB69FAE4" blockId="29.[1012,1252,1341,1365]" box="[1012,1252,1341,1365]" pageId="29" pageNumber="920">
<heading id="9A4B2D2CFFFCBA3CFC29FA8DFB69FAE4" bold="true" box="[1012,1252,1341,1365]" centered="true" fontSize="10" level="2" pageId="29" pageNumber="920" reason="2">
<emphasis id="F3C84652FFFCBA3CFC29FA8DFB69FAE4" bold="true" box="[1012,1252,1341,1365]" pageId="29" pageNumber="920">
<taxonomicName id="06BCE1C3FFFCBA3CFC29FA8DFBD0FAE4" ID-CoL="84GNK" authority="Schubart &amp; Ng, 2020" authorityName="Schubart &amp; Ng" authorityYear="2020" box="[1012,1117,1342,1365]" class="Malacostraca" family="Sesarmidae" genus="Danarma" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="920" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="F3C84652FFFCBA3CFC29FA8DFBD0FAE4" bold="true" box="[1012,1117,1342,1365]" italics="true" pageId="29" pageNumber="920">Danarma</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E8FBFB29FFFCBA3CFBB1FA8EFB69FAE4" box="[1132,1252,1341,1365]" pageId="29" pageNumber="920" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFFCBA3CFCF0FACEFBD5FA04" pageId="29" pageNumber="920" type="type_taxon">
<paragraph id="C1039A40FFFCBA3CFCF0FACEFBD5FA04" blockId="29.[813,1451,1405,1461]" pageId="29" pageNumber="920">
<emphasis id="F3C84652FFFCBA3CFCF0FACEFC33FA24" bold="true" box="[813,958,1405,1429]" pageId="29" pageNumber="920">
<typeStatus id="1E0724E2FFFCBA3CFCF0FACEFCEAFA24" box="[813,871,1405,1429]" pageId="29" pageNumber="920">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="06BCE1C3FFFCBA3CFC11FACDFAA2FA24" ID-CoL="7Z29S" authority="Dana, 1851" authorityName="Dana" authorityYear="1851" box="[972,1327,1405,1429]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="920" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="F3C84652FFFCBA3CFC11FACDFB25FA24" box="[972,1192,1405,1429]" italics="true" pageId="29" pageNumber="920">Sesarma obtusifrons</emphasis>
<bibRefCitation id="A52DE7B1FFFCBA3CFB6DFACDFAA2FA24" author="Dana" box="[1200,1327,1405,1429]" firstAuthor="Dana" pageId="29" pageNumber="920" pagination="247 - 254" refId="ref68617" refString="Dana JD (1851) Conspectus crustaceorum quae in orbis terrarum circumnavigatione, Carolo Wilkes e classe Reipublicae Foederatae Duce, lexit et descripsit J. D. Dana. Crustacea Grapsoidea, (Cyclometopa, Edwardsii). Proceedings of the Academy of Natural Sciences of Philadelphia, 5: 247 - 254." type="journal article" year="1851">Dana, 1851</bibRefCitation>
</taxonomicName>
, by present designation. Gender neuter.
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFFCBA3FFCF0FA6EFDBAF953" lastPageId="30" lastPageNumber="921" pageId="29" pageNumber="920" type="diagnosis">
<paragraph id="C1039A40FFFCBA3FFCF0FA6EFDBAF953" blockId="29.[812,1453,1501,2101]" lastBlockId="30.[136,775,1547,1763]" lastPageId="30" lastPageNumber="921" pageId="29" pageNumber="920">
<emphasis id="F3C84652FFFCBA3CFCF0FA6EFC28FA44" bold="true" box="[813,933,1501,1525]" pageId="29" pageNumber="920">Diagnosis</emphasis>
. Carapace ovate-trapezoidal, broader than long; frontal margin entire, prominently deflexed, almost completely shielding antennules from frontal view, margin much longer than posterior carapace margin; lateral margins of carapace entire, posterolateral part distinctly converging; regions of carapace poorly demarcated; postfrontal and epigastric crests separated by relatively deep grooves, margin relatively rounded, regions clearly separated; basal articles of antenna and antennules separated by septum; dorsal margin of palm smooth; outer surface of palm smooth, inner surface with prominent transverse submedian swelling, highest point with transverse ridge of granules, outer surface of palm and pollex convex, outer lower surface of palm smooth; dorsal margin of chelipedal dactylus smooth; inner distal margin of merus of cheliped not lamelliform; inner surfaces of first to third ambulatory coxae with numerous long setae between them; male thoracic sternites 24 relatively narrow in adults, suture between sternites 3 and 4 distinct; male sternopleonal cavity reaching two-thirds length of sternite 4 to just before anterior margin of sternite 2; male thoracic sternite 5 smooth, without depression on anterior part; G1 slender or relatively stout, chitinous part short or relatively long. Vulva on anterior part of sternite 6, anterior edge presses against sternite 5; anterior and posterior sternal vulvar covers flat, triangular, tips overlapping; opening not projecting, below sternal vulvar covers.
</paragraph>
</subSubSection>
<caption id="95C3CAC8FFFFBA3FFF55FA37FA4EFA65" ID-DOI="http://doi.org/10.5281/zenodo.5351327" ID-Zenodo-Dep="5351327" httpUri="https://zenodo.org/record/5351327/files/figure.png" pageId="30" pageNumber="921" startId="30.[136,171,1412,1434]" targetBox="[136,1474,189,1397]" targetPageId="30">
<paragraph id="C1039A40FFFFBA3FFF55FA37FA4EFA65" blockId="30.[136,1475,1412,1492]" pageId="30" pageNumber="921">
Fig. 15. Chela. A, B,
<taxonomicName id="06BCE1C3FFFFBA3FFE81FA36FDA9FA2B" baseAuthorityName="Davie &amp; Ng" baseAuthorityYear="2013" box="[348,548,1413,1434]" class="Malacostraca" family="Sesarmidae" genus="Danarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="921" phylum="Arthropoda" rank="species" species="eurymerus">
<emphasis id="F3C84652FFFFBA3FFE81FA36FDA9FA2B" box="[348,548,1413,1434]" italics="true" pageId="30" pageNumber="921">Danarma eurymerus</emphasis>
</taxonomicName>
, holotype male (18.7 × 14.8 mm) (NMNS-7028002), Lanyu Island, Taiwan; C, D,
<taxonomicName id="06BCE1C3FFFFBA3FFABDFA37FF52FA07" baseAuthorityName="De Man, in Weber" baseAuthorityYear="1897" class="Malacostraca" family="Sesarmidae" genus="Cristarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="921" phylum="Arthropoda" rank="species" species="eulimene">
<emphasis id="F3C84652FFFFBA3FFABDFA37FF52FA07" italics="true" pageId="30" pageNumber="921">Cristarma eulimene</emphasis>
</taxonomicName>
, male (22.2 × 17.2 mm) (ZRC 1968.1.22.2), Inhaca Island, Mozambique; E, F,
<taxonomicName id="06BCE1C3FFFFBA3FFC01FA12FB11FA07" baseAuthorityName="Crosnier" baseAuthorityYear="1965" box="[988,1180,1441,1462]" class="Malacostraca" family="Sesarmidae" genus="Cristarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="921" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="F3C84652FFFFBA3FFC01FA12FB11FA07" box="[988,1180,1441,1462]" italics="true" pageId="30" pageNumber="921">Cristarma ortmanni</emphasis>
</taxonomicName>
, male (20.0 × 15.5 mm) (ZRC 1968.1.22.1), Inhaca Island, Mozambique. A, C, E, outer view; B, inner view; D, F, dorso-lateral view showing pectinated ridge on palm.
</paragraph>
</caption>
<subSubSection id="89A6C9CBFFFFBA3FFF55F8B8FF5FF833" pageId="30" pageNumber="921" type="etymology">
<paragraph id="C1039A40FFFFBA3FFF55F8B8FF5FF833" blockId="30.[136,775,1803,1922]" pageId="30" pageNumber="921">
<emphasis id="F3C84652FFFFBA3FFF55F8B8FE8EF892" bold="true" box="[136,259,1803,1827]" pageId="30" pageNumber="921">Etymology</emphasis>
. The genus name is derived from James D. Dana (18131895), who described the
<typeStatus id="1E0724E2FFFFBA3FFE2BF898FDA9F8F2" box="[502,548,1835,1859]" pageId="30" pageNumber="921">type</typeStatus>
species, in arbritary combination with the genus name
<taxonomicName id="06BCE1C3FFFFBA3FFDD9F8F8FDEDF8D3" authorityName="Say" authorityYear="1817" box="[516,608,1867,1890]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="921" phylum="Arthropoda" rank="genus">
<emphasis id="F3C84652FFFFBA3FFDD9F8F8FDEDF8D3" box="[516,608,1867,1890]" italics="true" pageId="30" pageNumber="921">Sesarma</emphasis>
</taxonomicName>
. The gender is neuter.
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFFFBA3FFF55F818FD63F792" pageId="30" pageNumber="921" type="discussion">
<paragraph id="C1039A40FFFFBA3FFF55F818FD63F792" blockId="30.[136,775,1963,2083]" pageId="30" pageNumber="921">
<emphasis id="F3C84652FFFFBA3FFF55F818FED6F872" bold="true" box="[136,347,1963,1987]" pageId="30" pageNumber="921">Included species</emphasis>
.
<taxonomicName id="06BCE1C3FFFFBA3FFEB2F818FC8DF872" authority="Dana, 1851" authorityName="Dana" authorityYear="1851" box="[367,768,1963,1987]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="921" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="F3C84652FFFFBA3FFEB2F818FDEBF872" box="[367,614,1963,1987]" italics="true" pageId="30" pageNumber="921">Sesarma obtusifrons</emphasis>
<bibRefCitation id="A52DE7B1FFFFBA3FFDAEF818FC8DF872" author="Dana" box="[627,768,1963,1987]" firstAuthor="Dana" pageId="30" pageNumber="921" pagination="247 - 254" refId="ref68617" refString="Dana JD (1851) Conspectus crustaceorum quae in orbis terrarum circumnavigatione, Carolo Wilkes e classe Reipublicae Foederatae Duce, lexit et descripsit J. D. Dana. Crustacea Grapsoidea, (Cyclometopa, Edwardsii). Proceedings of the Academy of Natural Sciences of Philadelphia, 5: 247 - 254." type="journal article" year="1851">Dana, 1851</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="06BCE1C3FFFFBA3FFF54F878FDD1F852" authority="Davie &amp; Ng, 2013" authorityName="Davie &amp; Ng" authorityYear="2013" box="[137,604,1995,2019]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="921" phylum="Arthropoda" rank="species" species="silus">
<emphasis id="F3C84652FFFFBA3FFF54F878FEEAF852" box="[137,359,1995,2019]" italics="true" pageId="30" pageNumber="921">Chiromantes silus</emphasis>
<bibRefCitation id="A52DE7B1FFFFBA3FFEAFF878FDD1F852" author="Davie &amp; Ng" box="[370,604,1995,2019]" firstAuthor="Davie" pageId="30" pageNumber="921" pagination="1 - 25" refId="ref68942" refString="Davie PJF &amp; Ng PKL (2013) A review of Chiromantes obtusifrons (Dana, 1851) (Decapoda: Brachyura: Sesarmidae), with descriptions of four new sibling-species from Christmas Island, Guam and Taiwan. Zootaxa, 3609 (1): 1 - 25." type="journal article" year="2013">Davie &amp; Ng, 2013</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="06BCE1C3FFFFBA3FFDB2F878FE4CF7B2" authority="Davie &amp; Ng, 2013" authorityName="Davie &amp; Ng" authorityYear="2013" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="921" phylum="Arthropoda" rank="species" species="leptomerus">
<emphasis id="F3C84652FFFFBA3FFDB2F878FF70F7B2" italics="true" pageId="30" pageNumber="921">Chiromantes leptomerus</emphasis>
<bibRefCitation id="A52DE7B1FFFFBA3FFEDFF858FE4CF7B2" author="Davie &amp; Ng" box="[258,449,2027,2051]" firstAuthor="Davie" pageId="30" pageNumber="921" pagination="1 - 25" refId="ref68942" refString="Davie PJF &amp; Ng PKL (2013) A review of Chiromantes obtusifrons (Dana, 1851) (Decapoda: Brachyura: Sesarmidae), with descriptions of four new sibling-species from Christmas Island, Guam and Taiwan. Zootaxa, 3609 (1): 1 - 25." type="journal article" year="2013">Davie &amp; Ng, 2013</bibRefCitation>
</taxonomicName>
<taxonomicName id="06BCE1C3FFFFBA3FFE1BF858FE83F792" authority="Davie &amp; Ng, 2013" authorityName="Davie &amp; Ng" authorityYear="2013" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="921" phylum="Arthropoda" rank="species" species="eurymerus">
<emphasis id="F3C84652FFFFBA3FFE1BF858FD4FF7B2" box="[454,706,2027,2051]" italics="true" pageId="30" pageNumber="921">Chiromantes eurymerus</emphasis>
<bibRefCitation id="A52DE7B1FFFFBA3FFD15F858FE83F792" author="Davie &amp; Ng" firstAuthor="Davie" pageId="30" pageNumber="921" pagination="1 - 25" refId="ref68942" refString="Davie PJF &amp; Ng PKL (2013) A review of Chiromantes obtusifrons (Dana, 1851) (Decapoda: Brachyura: Sesarmidae), with descriptions of four new sibling-species from Christmas Island, Guam and Taiwan. Zootaxa, 3609 (1): 1 - 25." type="journal article" year="2013">Davie &amp; Ng, 2013</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="06BCE1C3FFFFBA3FFEC6F7B8FD66F792" authority="Davie &amp; Ng, 2013" authorityName="Davie &amp; Ng" authorityYear="2013" box="[283,747,2059,2083]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="921" phylum="Arthropoda" rank="species" species="garfunkel">
<emphasis id="F3C84652FFFFBA3FFEC6F7B8FD9EF792" box="[283,531,2059,2083]" italics="true" pageId="30" pageNumber="921">Chiromantes garfunkel</emphasis>
<bibRefCitation id="A52DE7B1FFFFBA3FFDC1F7B8FD66F792" author="Davie &amp; Ng" box="[540,747,2059,2083]" firstAuthor="Davie" pageId="30" pageNumber="921" pagination="1 - 25" refId="ref68942" refString="Davie PJF &amp; Ng PKL (2013) A review of Chiromantes obtusifrons (Dana, 1851) (Decapoda: Brachyura: Sesarmidae), with descriptions of four new sibling-species from Christmas Island, Guam and Taiwan. Zootaxa, 3609 (1): 1 - 25." type="journal article" year="2013">Davie &amp; Ng, 2013</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFFFBA3EFC9EF9B8FD7DFF44" lastPageId="31" lastPageNumber="922" pageId="30" pageNumber="921" type="discussion">
<paragraph id="C1039A40FFFFBA3EFC9EF9B8FD7DFF44" blockId="30.[835,1475,1547,2083]" lastBlockId="31.[113,752,189,246]" lastPageId="31" lastPageNumber="922" pageId="30" pageNumber="921">
<emphasis id="F3C84652FFFFBA3FFC9EF9B8FC23F992" bold="true" box="[835,942,1547,1571]" pageId="30" pageNumber="921">Remarks</emphasis>
. As discussed earlier, what used to be called
<taxonomicName id="06BCE1C3FFFFBA3FFC99F998FBD7F9F2" baseAuthorityName="Dana" baseAuthorityYear="1851" box="[836,1114,1579,1603]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="921" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="F3C84652FFFFBA3FFC99F998FBD7F9F2" box="[836,1114,1579,1603]" italics="true" pageId="30" pageNumber="921">Chiromantes obtusifrons</emphasis>
</taxonomicName>
is now a group of five species (
<bibRefCitation id="A52DE7B1FFFFBA3FFC96F9F8FB92F9D2" author="Davie &amp; Ng" box="[843,1055,1611,1635]" firstAuthor="Davie" pageId="30" pageNumber="921" pagination="1 - 25" refId="ref68942" refString="Davie PJF &amp; Ng PKL (2013) A review of Chiromantes obtusifrons (Dana, 1851) (Decapoda: Brachyura: Sesarmidae), with descriptions of four new sibling-species from Christmas Island, Guam and Taiwan. Zootaxa, 3609 (1): 1 - 25." type="journal article" year="2013">Davie &amp; Ng, 2013</bibRefCitation>
), all sharing a peculiar frontal margin that is very broad and prominently bent downwards such that much of the antennae and antennules are completely covered in frontal view. The broad front also results in a pecular carapace shape—clearly trapezoidal with the posterolateral margins strongly converging towards the posterior carapace margin, but with the lateral margins arcuate, resulting in a transversely subovate form (
<figureCitation id="598786C5FFFFBA3FFBB4F898FB7EF8F2" box="[1129,1267,1835,1859]" captionStart="Fig" captionStartId="26.[136,171,1910,1932]" captionTargetBox="[136,1474,199,1895]" captionTargetId="figure-8@26.[136,1474,199,1895]" captionTargetPageId="26" captionText="Fig. 13. Overall habitus. A, Danarma obtusifrons, male (17.7 × 13.2 mm) (UF-FLMNH 14837), Oahu, Hawaii; B, Danarma silus, holotype male (16.3 × 12.8 mm) (ZRC 2012.0787), Guam; C, Danarma eurymerus, holotype male (18.7 × 14.8 mm) (NMNS-7028-002), Lanyu Island, Taiwan; D, Danarma garfunkel, holotype male (17.0 × 13.0 mm) (QM), Greta Beach, Christmas Island; E, Cristarma eulimene, male (22.2 × 17.2 mm) (ZRC 1968.1.22.2), Inhaca Island, Mozambique; F, Cristarma ortmanni, male (20.0 × 15.5 mm) (ZRC 1968.1.22.1), Inhaca Island, Mozambique; G, Trapezarma angolense, neotype male (39.3 × 32.7 mm) (SMF-ZMG 635), Benguella, Angola; H, Platychirarma buettikoferi, paralectotype male (10.0 × 8.7 mm) (SMF-ZMG 636), Fisherman Lake, Liberia; I, Platychirarma buettikoferi, lectotype male (14.1 × 11.6 mm) (RMNH D 148), Liberia; J, Platychirarma buettikoferi, male (12.6 × 10.4 mm) (ZRC 2015.0298), Cameroon." figureDoi="http://doi.org/10.5281/zenodo.5351323" httpUri="https://zenodo.org/record/5351323/files/figure.png" pageId="30" pageNumber="921">Figs. 13AD</figureCitation>
,
<figureCitation id="598786C5FFFFBA3FFB21F898FAC3F8F2" box="[1276,1358,1835,1859]" captionStart="Fig" captionStartId="28.[136,171,1850,1872]" captionTargetBox="[136,1474,287,1835]" captionTargetId="figure-7@28.[136,1474,287,1835]" captionTargetPageId="28" captionText="Fig. 14. Frontal view of cephalothorax. A, Danarma obtusifrons, male (17.7 × 13.2 mm) (UF-FLMNH 14837), Hawaii; B, Danarma garfunkel, holotype male (17.0 × 13.0 mm) (QM, ex ZRC 2009.0822), Christmas Island; C, D, Danarma eurymerus, holotype male (18.7 × 14.8 mm) (NMNS-7028-002), Lanyu Island, Taiwan; E, Cristarma eulimene, male (22.2 × 17.2 mm) (ZRC 1968.1.22.2), Inhaca Island, Mozambique; F, Cristarma ortmanni, male (20.0 × 15.5 mm) (ZRC 1968.1.22.1), Inhaca Island, Mozambique; G, Trapezarma angolense, neotype male (39.3 × 32.7 mm) (SMF-ZMG 635), Benguella, Angola; H, Platychirarma buettikoferi, paralectotype male (10.0 × 8.7 mm) (SMF-ZMG 636), Fisherman Lake, Liberia." figureDoi="http://doi.org/10.5281/zenodo.5351325" httpUri="https://zenodo.org/record/5351325/files/figure.png" pageId="30" pageNumber="921">14AD</figureCitation>
). The outer surface of the palm is also smooth, with no longitudinal ridge, the dactylar finger completely unarmed on the dorsal margin (
<figureCitation id="598786C5FFFFBA3FFC42F838FBA9F812" box="[927,1060,1931,1955]" captionStart="Fig" captionStartId="30.[136,171,1412,1434]" captionTargetBox="[136,1474,189,1397]" captionTargetId="figure-8@30.[136,1474,189,1397]" captionTargetPageId="30" captionText="Fig. 15. Chela. A, B, Danarma eurymerus, holotype male (18.7 × 14.8 mm) (NMNS-7028002), Lanyu Island, Taiwan; C, D, Cristarma eulimene, male (22.2 × 17.2 mm) (ZRC 1968.1.22.2), Inhaca Island, Mozambique; E, F, Cristarma ortmanni, male (20.0 × 15.5 mm) (ZRC 1968.1.22.1), Inhaca Island, Mozambique. A, C, E, outer view; B, inner view; D, F, dorso-lateral view showing pectinated ridge on palm." figureDoi="http://doi.org/10.5281/zenodo.5351327" httpUri="https://zenodo.org/record/5351327/files/figure.png" pageId="30" pageNumber="921">Fig. 15A, B</figureCitation>
), and the surfaces between the first to third ambulatory coxae have distinct dense tufts of long setae (
<figureCitation id="598786C5FFFFBA3FFC96F878FC5CF852" box="[843,977,1995,2019]" captionStart="Fig" captionStartId="34.[136,171,1517,1539]" captionTargetBox="[136,1474,189,1501]" captionTargetId="figure-8@34.[136,1474,189,1501]" captionTargetPageId="34" captionText="Fig. 18. Male anterior thoracic sternum and pleon. A, Danarma obtusifrons, male (19.7 × 14.7 mm) (ZRC 2002.0220), Oahu, Hawaii; B, Danarma eurymerus, holotype male (18.7 × 14.8 mm) (NMNS-7028-002), Lanyu Island, Taiwan; C, Cristarma eulimene, male (22.2 × 17.2 mm) (ZRC 1968.1.22.2), Inhaca Island, Mozambique; D, Cristarma ortmanni, male (20.0 × 15.5 mm) (ZRC 1968.1.22.1), Inhaca Island, Mozambique; E, Trapezarma angolense, neotype male (39.3 × 32.7 mm) (SMF-ZMG 635), Benguella, Angola; F, Platychirarma buettikoferi, paralectotype male (10.0 × 8.7 mm) (SMF-ZMG 636), Fisherman Lake, Liberia." figureDoi="http://doi.org/10.5281/zenodo.5351333" httpUri="https://zenodo.org/record/5351333/files/figure.png" pageId="30" pageNumber="921">Fig. 18A, B</figureCitation>
). The presence of strongly setose ambulatory coxae is a feature shared with species of
<taxonomicName id="06BCE1C3FFFFBA3FFACBF858FA08F7B3" authority="Schubart &amp; Ng, 2020" authorityName="Schubart &amp; Ng" authorityYear="2020" box="[1302,1413,2027,2050]" class="Malacostraca" family="Sesarmidae" genus="Cristarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="921" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="F3C84652FFFFBA3FFACBF858FA08F7B3" box="[1302,1413,2027,2050]" italics="true" pageId="30" pageNumber="921">Cristarma</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E8FBFB29FFFFBA3FFA48F858FC09F792" pageId="30" pageNumber="921" rank="genus">new genus</taxonomicNameLabel>
, from East Africa (see later). This suite of distinctive characters shared by five similar species is strong support for the establishment of a new genus, here named
<taxonomicName id="06BCE1C3FFFEBA3EFD5EFF6DFD64FF44" authorityName="Schubart &amp; Ng" authorityYear="2020" box="[643,745,222,245]" class="Malacostraca" family="Sesarmidae" genus="Danarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="922" phylum="Arthropoda" rank="genus">
<emphasis id="F3C84652FFFEBA3EFD5EFF6DFD64FF44" box="[643,745,222,245]" italics="true" pageId="31" pageNumber="922">Danarma</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>