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<document id="27AD87551DFC73C2758A8009FA411891" ID-CLB-Dataset="55667" ID-DOI="10.26107/RBZ-2020-0097" ID-GBIF-Dataset="6cb155ce-8b9f-48ce-8e6c-b0dc00f8b1cc" ID-ISSN="2345-7600" ID-Zenodo-Dep="5351295" ID-ZooBank="815E4670-B063-4FD8-B31E-3AD89B3A7942" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1630205841501" checkinUser="diego" docAuthor="Schubart, Christoph D. &amp; Ng, Peter K. L." docDate="2020" docId="49152B56FFF4BA3AFC57FECEFF73FAA4" docLanguage="en" docName="RafflesBZool.68.891-994.pdf" docOrigin="Raffles Bulletin of Zoology 68" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Orisarma intermedium" docType="treatment" docVersion="12" lastPageNumber="918" masterDocId="B52C532EFFE1BA21FFDDFFB3FF8DFFB1" masterDocTitle="Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species" masterLastPageNumber="994" masterPageNumber="891" pageNumber="912" updateTime="1699203722016" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="E9F69E2B85E34EA04C5C23BC3E9EE74B">Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species</mods:title>
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<mods:namePart id="50558BC88DEE2E7750C5528AADDF5213">Schubart, Christoph D.</mods:namePart>
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<mods:namePart id="E7331BD37A6EBA38E7FB3FA9F55ED435">Ng, Peter K. L.</mods:namePart>
<mods:affiliation id="DB3638F7DBA1A5F36D798F7530B422D7">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore</mods:affiliation>
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<taxonomicName id="06BCE1C3FFF4BA34FC57FECEFAC0FE24" ID-CoL="84H2K" authority="(De Haan, 1835)" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[906,1357,381,405]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="912" phylum="Arthropoda" rank="species" species="intermedium">
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<emphasis id="F3C84652FFF4BA34FC57FECEFB04FE24" bold="true" box="[906,1161,381,405]" italics="true" pageId="21" pageNumber="912">Orisarma intermedium</emphasis>
(De Haan, 1835)
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<subSubSection id="89A6C9CBFFF4BA34FCE4FE2EFB49FE64" pageId="21" pageNumber="912" type="description">
<paragraph id="C1039A40FFF4BA34FCE4FE2EFB49FE64" blockId="21.[825,1439,381,469]" pageId="21" pageNumber="912">
(
<figureCitation id="598786C5FFF4BA34FC9CFE2EFC32FE04" box="[833,959,413,437]" captionStart="Fig" captionStartId="8.[136,171,1926,1948]" captionTargetBox="[136,1476,212,1910]" captionTargetId="figure-7@8.[136,1476,212,1910]" captionTargetPageId="8" captionText="Fig. 2. Overall habitus. A, Orisarma intermedium, lectotype male (23.0 × 19.9 mm) (RMNH-D165), Japan; B, O. intermedium, male (27.1 × 25.2 mm) (ZRC 1970.2.23.6), Japan; C, O. intermedium, male (27.0 × 25.0 mm) (ZRC 2013.0140), Kyushu, Japan; D, O. intermedium, male (29.5 × 26.5 mm) (ZRC 2001.0034), Pingtung, Taiwan; E, O. intermedium, male (34.2 × 30.7 mm) (ZRC 2001.0034), Pingtung, Taiwan; F, O. intermedium, male (37.5 × 35.8 mm) (ZRC 2014.0265), Kumejima Island, Japan; G, O. sinense, lectotype male (19.0 × 16.6 mm) (MNHN-BP3635a), China; H, O. sinense, paralectotype female (18.8 × 16.9 mm) (MNHN-BP3635b), China; I, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China; J, Orisarma sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China." figureDoi="http://doi.org/10.5281/zenodo.5351299" httpUri="https://zenodo.org/record/5351299/files/figure.png" pageId="21" pageNumber="912">Figs. 2AF</figureCitation>
,
<figureCitation id="598786C5FFF4BA34FC16FE2EFC63FE04" box="[971,1006,413,437]" captionStart="Fig" captionStartId="10.[136,171,1435,1457]" captionTargetBox="[136,1476,189,1419]" captionTargetId="figure-7@10.[136,1476,189,1419]" captionTargetPageId="10" captionText="Fig. 3. Frontal view of cephalothorax. A, Chiromantes haematocheir, lectotype male (33.7 × 29.7 mm) (RMNH-D160), Japan; B, Orisarma dehaani, lectotype male (39.5 × 35.7 mm) (RMNH-D157), Japan; C, O. neglectum, neotype male (35.6 × 31.8 mm) (ZRC 1998.310), Shanghai, China; D, O. intermedium, male (27.1 × 25.2 mm) (ZRC 1970.2.23.6), Japan; E, O. sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China; F, O. patshuni, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong." figureDoi="http://doi.org/10.5281/zenodo.5351301" httpUri="https://zenodo.org/record/5351301/files/figure.png" pageId="21" pageNumber="912">3D</figureCitation>
,
<figureCitation id="598786C5FFF4BA34FC26FE2EFBBAFE04" box="[1019,1079,413,437]" captionStart="Fig" captionStartId="12.[135,170,1954,1976]" captionTargetBox="[136,1476,241,1939]" captionTargetId="figure-7@12.[136,1476,241,1939]" captionTargetPageId="12" captionText="Fig. 4. A, F, Orisarma dehaani, male (36.4 × 34.4 mm) (ZRC 2011.1027), Kumejima Island, Japan; B, D, O. dehaani, male (27.6 × 29.6 mm) (ZRC 2002.0223), Hong Kong; C, E, G, O. neglectum, male (31.7 × 28.0 mm) (ZRC 1998.309), Shanghai, China; H, J, O. intermedium, male (29.5 × 26.5 mm) (ZRC 2001.0034), Pingtung, Taiwan; I, K, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China. AC, dorsal view of carapace; D, E, H, I, frontal and anterolateral margins; F, G, J, K, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.5351303" httpUri="https://zenodo.org/record/5351303/files/figure.png" pageId="21" pageNumber="912">4H, J</figureCitation>
,
<figureCitation id="598786C5FFF4BA34FB98FE2EFB04FE04" box="[1093,1161,413,437]" captionStart="Fig" captionStartId="16.[135,170,1734,1756]" captionTargetBox="[136,1476,189,1718]" captionTargetId="figure-7@16.[136,1476,189,1718]" captionTargetPageId="16" captionText="Fig. 6. Chela. A, Orisarma intermedium, lectotype male (23.0 × 19.9 mm) (RMNH-D165), Japan; B, O. intermedium, male (27.1 × 25.2 mm) (ZRC 1970.2.23.6), Japan; C, Orisarma intermedium, male (34.7 × 31.6 mm) (ZRC 2001.0034), Pingtung, Taiwan; D, O. intermedium, male (37.5 × 35.8 mm) (ZRC 2014.0265), Kumejima Island, Japan; E, O. sinense, lectotype male (19.0 × 16.6 mm) (MNHN-BP3635a), China; F, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China; G, H, O. sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China. AG, outer view; H, inner view." figureDoi="http://doi.org/10.5281/zenodo.5351307" httpUri="https://zenodo.org/record/5351307/files/figure.png" pageId="21" pageNumber="912">6AD</figureCitation>
,
<figureCitation id="598786C5FFF4BA34FB48FE2EFB5AFE04" box="[1173,1239,413,437]" captionStart="Fig" captionStartId="18.[135,170,1505,1527]" captionTargetBox="[136,1476,189,1489]" captionTargetId="figure-7@18.[136,1476,189,1489]" captionTargetPageId="18" captionText="Fig. 8. Right third and fourth ambulatory legs. A, Orisarma intermedium, male (27.0 × 25.0 mm) (ZRC 2013.0140), Kyushu, Japan; B, O. intermedium, male (29.5 × 26.5 mm) (ZRC 2001.0034), Pingtung, Taiwan; C, O. intermedium, male (37.5 × 35.8 mm) (ZRC 2014.0265), Kumejima Island, Japan; D, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China." figureDoi="http://doi.org/10.5281/zenodo.5351311" httpUri="https://zenodo.org/record/5351311/files/figure.png" pageId="21" pageNumber="912">8AC</figureCitation>
,
<figureCitation id="598786C5FFF4BA34FB39FE2EFA89FE04" box="[1252,1284,413,437]" captionStart="Fig" captionStartId="20.[135,170,1977,1999]" captionTargetBox="[195,1417,189,1962]" captionTargetId="figure-7@20.[195,1417,189,1962]" captionTargetPageId="20" captionText="Fig. 9. Male anterior thoracic sternum and pleon. A, Chiromantes haematocheir, lectotype male (33.7 × 29.7 mm) (RMNH-D160), Japan; B, C. ryukyuanus, holotype male (33.1 × 29.6 mm) (RUMF-ZC-539), Okinawa Island, Japan; C, Orisarma dehaani, lectotype male (39.5 × 35.7 mm) (RMNH-D157), Japan; D, O. neglectum, neotype male (35.6 × 31.8 mm) (ZRC 1998.310), Shanghai, China; E, O. intermedium, lectotype male (23.0 × 19.9 mm) (RMNH-D165), Japan; F, O. sinense, lectotype male (19.0 × 16.6 mm) (MNHN-BP3635a), China; G, O. sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China; H, O. patshuni, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong." figureDoi="http://doi.org/10.5281/zenodo.5351313" httpUri="https://zenodo.org/record/5351313/files/figure.png" pageId="21" pageNumber="912">9E</figureCitation>
,
<figureCitation id="598786C5FFF4BA34FACCFE2EFAD2FE04" box="[1297,1375,413,437]" captionStart="Fig" captionStartId="25.[113,148,1906,1928]" captionTargetBox="[116,1451,292,1890]" captionTargetId="figure-15@25.[99,1466,286,1902]" captionTargetPageId="25" captionText="Fig. 12. AE, Orisarma intermedium, male (27.1 × 25.2 mm) (ZRC 1970.2.23.6), Japan; FJ, O. sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China; KO, O. patshuni, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong. A, F, K, male pleons; B, G, L, left G1 (ventral view, denuded); C, H, M, left G1 (dorsal view, denuded); D, I, N, left distal part of G1 (ventral view, denuded); E, J, O, left distal part of G1 (dorsal view, denuded). Scales: A, F = 5.0 mm; BE, GJ, KM = 1.0 mm; N, O = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5351321" httpUri="https://zenodo.org/record/5351321/files/figure.png" pageId="21" pageNumber="912">12AE</figureCitation>
,
<figureCitation id="598786C5FFF4BA34FAB1FE2EFBA8FE64" captionStart="Fig" captionStartId="56.[136,171,1595,1617]" captionTargetBox="[136,1474,189,1580]" captionTargetId="figure-7@56.[136,1474,189,1580]" captionTargetPageId="56" captionText="Fig. 33. AE, male anterior thoracic sternum and pleon; FH, left ambulatory leg. A, D, H, Manarma moeschii, male (18.2 × 16.3 mm) (ZRC 2000.1926), Thailand; B, E, M. johorense, male (14.6 × 12.6 mm) (ZRC 1971.9.24.14), Seletar River, Singapore; C, F, Orisarma intermedium, male (34.7 × 31.6 mm) (ZRC 2001.0034), Pingtung, Taiwan; G, O. patshuni (, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong." figureDoi="http://doi.org/10.5281/zenodo.5351357" httpUri="https://zenodo.org/record/5351357/files/figure.png" pageId="21" pageNumber="912">33C, F</figureCitation>
,
<figureCitation id="598786C5FFF4BA34FBEFFE0EFBECFE64" box="[1074,1121,445,469]" captionStart="Fig" captionStartId="72.[136,171,1898,1920]" captionTargetBox="[136,1474,211,1882]" captionTargetId="figure-8@72.[136,1474,211,1882]" captionTargetPageId="72" captionText="Fig. 43. Vulva. A, Chiromantes haematocheir, female (27.0 × 23.7 mm) (ZRC 2002.0225), Japan; B, Orisarma dehaani, female (20.4 × 17.8 mm) (ZRC 2012.0057), Japan; C, Orisarma intermedium, female (31.8 × 28.4 mm) (ZRC 2013.0140), Japan; D, Orisarma patshuni, female (20.5 × 18.1 mm) (ZRC 2014.0817), Hong Kong; E, Danarma obtusifrons, female (15.7 × 11.6 mm) (UF-FLMNH 14837), Hawaii; F, Danarma eurymerus, paratype ovigerous female (18.0 × 13.4 mm) (ZRC 2012.0956), Taiwan; G, Cristarma eulimene, female (15.6 × 12.8 mm) (ZRC 1968.1.22.3), Mozambique; H, Cristarma ortmanni, female (13.1 × 10.2 mm) (ZRC 2000.1783), Kenya; I, Trapezarma angolense, female (18.2 × 14.3 mm) (ZRC 2015.0297), Cameroon; J, Platychirarma buettikoferi, ovigerous female (11.3 × 9.1 mm) (ZRC 2015.0298), Cameroon." figureDoi="http://doi.org/10.5281/zenodo.5351373" httpUri="https://zenodo.org/record/5351373/files/figure.png" pageId="21" pageNumber="912">43C</figureCitation>
,
<figureCitation id="598786C5FFF4BA34FBB3FE0EFB31FE64" box="[1134,1212,445,469]" captionStart="Fig" captionStartId="86.[136,171,1434,1456]" captionTargetBox="[136,1474,189,1419]" captionTargetId="figure-7@86.[136,1474,189,1419]" captionTargetPageId="86" captionText="Fig. 54. Colour in life. A, Orisarma intermedium, Pingtung, Taiwan, specimen not collected [photograph: PKL Ng]; B, O. intermedium, female (37.5 × 35.8 mm) (ZRC 2014.0265), Kumejima Island, Japan [photograph: PKL Ng]; C, O. intermedium, male (23.2 × 20.0 mm) (ZRC 2019.0540), Hong Kong [photograph: Paul Ng]; D, O. sinense, Shanghai, China, specimen not collected [photograph: Liu Wenliang]." figureDoi="http://doi.org/10.5281/zenodo.5351395" httpUri="https://zenodo.org/record/5351395/files/figure.png" pageId="21" pageNumber="912">54AC</figureCitation>
)
</paragraph>
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<treatmentCitationGroup id="E1ACBD6EFFF4BA34FCF0FE4FFA26FDA3" box="[813,1451,508,530]" pageId="21" pageNumber="912">
<taxonomicName id="06BCE1C3FFF4BA34FCF0FE4FFABCFDA3" ID-CoL="6L4VW" authority="De Haan, 1835: 61" authorityName="De Haan" authorityPageNumber="61" authorityYear="1835" box="[813,1329,508,530]" class="Malacostraca" family="Grapsidae" genus="Grapsus" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="912" phylum="Arthropoda" rank="species" species="intermedius" subGenus="Pachysoma">
<emphasis id="F3C84652FFF4BA34FCF0FE4FFCF3FDA0" box="[813,894,508,529]" italics="true" pageId="21" pageNumber="912">Grapsus</emphasis>
(
<emphasis id="F3C84652FFF4BA34FC56FE4FFC77FDA0" box="[907,1018,508,529]" italics="true" pageId="21" pageNumber="912">Pachysoma</emphasis>
)
<emphasis id="F3C84652FFF4BA34FBDAFE4FFBF5FDA0" box="[1031,1144,508,529]" italics="true" pageId="21" pageNumber="912">intermedius</emphasis>
<treatmentCitation id="401DBC51FFF4BA34FBA0FE4FFABCFDA3" author="Haan W De" box="[1149,1329,508,530]" page="61" pageId="21" pageNumber="912" year="1835">De Haan, 1835: 61</treatmentCitation>
</taxonomicName>
, pl. 16 fig. 5.
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</paragraph>
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<treatmentCitationGroup id="E1ACBD6EFFF4BA34FCF0FDAAFBDAFD35" pageId="21" pageNumber="912">
<taxonomicName id="06BCE1C3FFF4BA34FCF0FDAAFC7FFD9F" ID-CoL="993LQ" authority="(De Haan, 1835)" box="[813,1010,537,558]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="912" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="F3C84652FFF4BA34FCF0FDAAFC7FFD9F" box="[813,1010,537,558]" italics="true" pageId="21" pageNumber="912">Sesarma intermedia</emphasis>
</taxonomicName>
H.
<treatmentCitation id="401DBC51FFF4BA34FBEDFDAAFAB0FD9E" author="Milne Edwards H" box="[1072,1341,537,559]" page="186" pageId="21" pageNumber="912" year="1853">
<bibRefCitation id="A52DE7B1FFF4BA34FBEDFDAAFAB0FD9E" author="Milne Edwards" box="[1072,1341,537,559]" firstAuthor="Milne Edwards" page="186" pageId="21" pageNumber="912" pagination="163 - 228" refId="ref72839" refString="Milne Edwards H (1853) Memoires sur la famille des Ocypodiens, suite. Annales de Sciences Naturelle, Zoology, Paris, Serie 3 (Zoologie) 20: 163 - 228, pls. 6 - 11. [A continuation of H. Milne Edwards, 1852, and reprinted with it in the undated Melanges Carcinologiques pp. 129 - 196]" type="journal article" year="1853">Milne Edwards, 1853: 186</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFF4BA34FA97FDAAFC38FDFA" author="Stimpson W" page="105" pageId="21" pageNumber="912" year="1858">
<bibRefCitation id="A52DE7B1FFF4BA34FA97FDAAFC38FDFA" author="Stimpson" firstAuthor="Stimpson" page="105" pageId="21" pageNumber="912" pagination="93 - 110" refId="ref77553" refString="Stimpson W (1858) Crustacea Ocypodoidea. Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem, a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers Ducibus, observavit et descripsit. Pars V. Proceedings of the Academy of Natural Sciences of Philadelphia, 10: 93 - 110." type="journal article" year="1858">Stimpson, 1858: 105</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFF4BA34FC63FD86FBFCFDFA" author="Herklots JA" box="[958,1137,565,587]" page="17" pageId="21" pageNumber="912" year="1861">
<bibRefCitation id="A52DE7B1FFF4BA34FC63FD86FBFCFDFA" author="Herklots" box="[958,1137,565,587]" firstAuthor="Herklots" page="17" pageId="21" pageNumber="912" pagination="1 - 43" refId="ref70050" refString="Herklots JA (1861) Symbolae Carcinologicae. I. Catalogue des Crustaces qui ont servi de base au systeme carcinologique de M. W. de Haan, redige d'apres la collection du Musee des Pays- Bas et les Crustaces de la Faune du Japan. Tijdschrift voor Entomologie uitgegeven door de Nederlandische Entomologische Vereeniging, Leyden, 1861: 1 - 43." type="journal article" year="1861">Herklots, 1861: 17</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="A52DE7B1FFF4BA34FBA6FD85FAA4FDFA" author="De Man" box="[1147,1321,565,587]" firstAuthor="De Man" page="25" pageId="21" pageNumber="912" pagination="21 - 36" refId="ref72026" refString="Man JG De (1880) On some species of the genus Sesarma Say and Cardisoma Latr. Notes Leyden Museum, 2 (5): 21 - 36." type="journal article" year="1880">
De
<treatmentCitation id="401DBC51FFF4BA34FB41FD85FAA4FDFA" author="Man JG De" box="[1180,1321,565,587]" page="25" pageId="21" pageNumber="912" year="1880">Man, 1880: 25</treatmentCitation>
</bibRefCitation>
;
<treatmentCitation id="401DBC51FFF4BA34FAEFFD86FCF4FDD6" author="Miers EJ" page="314" pageId="21" pageNumber="912" year="1880">
<bibRefCitation id="A52DE7B1FFF4BA34FAEFFD86FCF4FDD6" author="Miers" firstAuthor="Miers" page="314" pageId="21" pageNumber="912" pagination="304 - 317" refId="ref72642" refString="Miers EJ (1880) On a collection of Crustacea from the Malayan Region. Part II. Telphusidea, Catometopa and Oxystomata. Annals and Magazine of Natural History, Series 5, 5 (28): 304 - 317, pl. 14." type="journal article" year="1880">Miers, 1880: 314</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="A52DE7B1FFF4BA34FC5EFDE0FBCCFDD9" author="De Man" box="[899,1089,594,616]" firstAuthor="De Man" page="649" pageId="21" pageNumber="912" pagination="639 - 722" refId="ref72089" refString="Man JG De (1887) Ubersicht der Indopacifischen Arten der Gattung Sesarma Say nebst einer Kritik der von W. Hess und E. Nauck in den Jahren 1865 und 1880 beschrieben Decapoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 2: 639 - 722." type="journal article" year="1887">
De
<treatmentCitation id="401DBC51FFF4BA34FC78FDE0FBCCFDD9" author="Man JG De" box="[933,1089,594,616]" page="649" pageId="21" pageNumber="912" year="1887">Man, 1887: 649</treatmentCitation>
</bibRefCitation>
;
<bibRefCitation id="A52DE7B1FFF4BA34FB96FDE0FA84FDD6" author="De Man" box="[1099,1289,594,616]" firstAuthor="De Man" page="337" pageId="21" pageNumber="912" pagination="225 - 264" refId="ref72297" refString="Man JG De (1892) Carcinological studies in the Leyden Museum, No. 6. Notes Leyden Museum, 14 (36): 225 - 264, pls. 7 - 10." type="journal article" year="1892">
De
<treatmentCitation id="401DBC51FFF4BA34FBB0FDE0FA84FDD6" author="Man JG De" box="[1133,1289,594,616]" page="337" pageId="21" pageNumber="912" year="1892">Man, 1892: 337</treatmentCitation>
</bibRefCitation>
;
<treatmentCitation id="401DBC51FFF4BA34FACEFDE1FCF4FD35" author="Ortmann AE" page="721" pageId="21" pageNumber="912" year="1894">
<bibRefCitation id="A52DE7B1FFF4BA34FACEFDE1FCF4FD35" author="Ortmann" firstAuthor="Ortmann" page="721" pageId="21" pageNumber="912" pagination="683 - 772" refId="ref74488" refString="Ortmann AE (1894) Abtheilung: Brachyura (Brachyura genuina Boas), III. Unterabtheilung: Cancroidea. 2. Section: Cancrinea, 2. Gruppe: Catametopa. Die Decapoden-Krebse des Strassburger Museums mit besonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur Zeit im Strassburger Museum aufbewahrten Formen. Theil VIII [= Part 8]. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 7 (5): 683 - 772, pl. 23." type="journal article" year="1894">Ortmann, 1894: 721</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFF4BA34FC58FDDCFBDEFD35" author="Stimpson W" box="[901,1107,623,645]" page="133" pageId="21" pageNumber="912" year="1907">
<bibRefCitation id="A52DE7B1FFF4BA34FC58FDDCFBDEFD35" author="Stimpson" box="[901,1107,623,645]" firstAuthor="Stimpson" page="133" pageId="21" pageNumber="912" pagination="1 - 240" refId="ref77609" refString="Stimpson W (1907) Report on the Crustacea (Brachyura and Anomura) collected by the North Pacific Exploring Expedition, 1853 - 1856. Smithsonian Miscellaneous Collections, 49 (1717): 1 - 240, pls. 1 - 26." type="journal article" year="1907">Stimpson, 1907: 133</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFF4BA34FCF0FD3FFA82FD10" blockId="21.[813,1452,508,1307]" box="[813,1295,652,674]" pageId="21" pageNumber="912">
<treatmentCitationGroup id="E1ACBD6EFFF4BA34FCF0FD3FFA82FD10" box="[813,1295,652,674]" pageId="21" pageNumber="912">
<taxonomicName id="06BCE1C3FFF4BA34FCF0FD3FFC75FD10" authorityName="De Haan" authorityYear="1835" box="[813,1016,652,673]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="912" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="F3C84652FFF4BA34FCF0FD3FFC75FD10" box="[813,1016,652,673]" italics="true" pageId="21" pageNumber="912">Sesarma intermedius</emphasis>
</taxonomicName>
<treatmentCitation id="401DBC51FFF4BA34FBC9FD3FFA86FD10" author="Fransen CHJM &amp; Holthuis LB &amp; Adema JPHM" box="[1044,1291,652,674]" page="130" pageId="21" pageNumber="912" year="1997">
<bibRefCitation id="A52DE7B1FFF4BA34FBC9FD3FFA86FD10" author="Fransen" box="[1044,1291,652,674]" etAl="et al." firstAuthor="Fransen" page="130" pageId="21" pageNumber="912" pagination="1 - 344" refId="ref69235" refString="Fransen CHJM, Holthuis LB &amp; Adema JPHM (1997) Typecatalogue of the Decapod Crustacea in the collections of the Nationaal Naturhistorisch Museum, with appendices of pre- 1900 collectors and material. Zoologische Verhandelingen, Leiden, 311: i - xvi, 1 - 344." type="journal article" year="1997">Fransen et al., 1997: 130</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="C1039A40FFF4BA34FCF0FD1AFBE7FC83" blockId="21.[813,1452,508,1307]" pageId="21" pageNumber="912">
<treatmentCitationGroup id="E1ACBD6EFFF4BA34FCF0FD1AFBE7FC83" pageId="21" pageNumber="912">
<taxonomicName id="06BCE1C3FFF4BA34FCF0FD1AFBD6FD0F" ID-CoL="993LQ" authority="(De Haan, 1835)" box="[813,1115,681,702]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="912" phylum="Arthropoda" rank="species" species="intermedia" subGenus="Sesarma">
<emphasis id="F3C84652FFF4BA34FCF0FD1AFC0DFD0F" box="[813,896,681,702]" italics="true" pageId="21" pageNumber="912">Sesarma</emphasis>
(
<emphasis id="F3C84652FFF4BA34FC4DFD1AFC6EFD0F" box="[912,995,681,702]" italics="true" pageId="21" pageNumber="912">Sesarma</emphasis>
)
<emphasis id="F3C84652FFF4BA34FC2EFD1AFBD6FD0F" box="[1011,1115,681,702]" italics="true" pageId="21" pageNumber="912">intermedia</emphasis>
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<treatmentCitation id="401DBC51FFF4BA34FBA5FD1AFAABFD0E" author="Tesch JJ" box="[1144,1318,681,703]" page="162" pageId="21" pageNumber="912" year="1917">
<bibRefCitation id="A52DE7B1FFF4BA34FBA5FD1AFAABFD0E" author="Tesch" box="[1144,1318,681,703]" firstAuthor="Tesch" page="162" pageId="21" pageNumber="912" pagination="127 - 260" refId="ref78063" refString="Tesch JJ (1917) Synopsis of the genera Sesarma, Metasesarma, Sarmatium and Clistocoeloma, with a key to the determination of the IndoPacific species. Zoologische Mededelingen, 3: 127 - 260, pls. 15 - 17." type="journal article" year="1917">Tesch, 1917: 162</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFF4BA34FAEFFD1AFCFBFD6A" author="Balss H" page="155" pageId="21" pageNumber="912" year="1922">
<bibRefCitation id="A52DE7B1FFF4BA34FAEFFD1AFCFBFD6A" author="Balss" firstAuthor="Balss" page="155" pageId="21" pageNumber="912" pagination="1" refId="ref67843" refString="Balss H (1922) Ostasiatische Decapoden, IV. Die Brachyrhynchen (Cancridea). Archiv fur Naturgeschichte, 88 A (11): 94 - 165, figs. 1, 2, pls. 1, 2." type="journal article" year="1922">Balss, 1922: 155</bibRefCitation>
</treatmentCitation>
(part);
<treatmentCitation id="401DBC51FFF4BA34FC63FD76FBEFFD6A" author="Sakai T" box="[958,1122,709,731]" page="235" pageId="21" pageNumber="912" year="1936">
<bibRefCitation id="A52DE7B1FFF4BA34FC63FD76FBEFFD6A" author="Sakai" box="[958,1122,709,731]" firstAuthor="Sakai" page="235" pageId="21" pageNumber="912" refId="ref75565" refString="Sakai T (1936) Crabs of Japan. 66 plates in life colours with descriptions. Sanseido, Tokyo, 239 pp., 12 pp. [literature cited], 27 pp. [bibliography and index], 66 pls. [dated 1935 but published in 1936]" type="book" year="1936">Sakai, 1936: 235</bibRefCitation>
</treatmentCitation>
, pl. 65 fig. 3;
<treatmentCitation id="401DBC51FFF4BA34FB38FD76FA04FD6A" author="Sakai T" box="[1253,1417,709,731]" page="684" pageId="21" pageNumber="912" year="1939">
<bibRefCitation id="A52DE7B1FFF4BA34FB38FD76FA04FD6A" author="Sakai" box="[1253,1417,709,731]" firstAuthor="Sakai" page="684" pageId="21" pageNumber="912" pagination="1 - 6" refId="ref75618" refString="Sakai T (1939) Studies on the Crabs of Japan. IV. Brachygnatha, Brachyrhyncha. Yokendo Co., Ltd., Tokyo. Pp. 365 - 741 + 1 - 23 [= Index], figs. 1 - 129, pls. XLII - CXI, tables 1 - 6." type="book chapter" year="1939">Sakai, 1939: 684</bibRefCitation>
</treatmentCitation>
, pl. 78 fig. 3;
<treatmentCitation id="401DBC51FFF4BA34FC68FD51FBD5FD49" author="Shen CJ" box="[949,1112,738,760]" page="95" pageId="21" pageNumber="912" year="1940">
<bibRefCitation id="A52DE7B1FFF4BA34FC68FD51FBD5FD49" author="Shen" box="[949,1112,738,760]" firstAuthor="Shen" page="95" pageId="21" pageNumber="912" pagination="211 - 242" refId="ref76939" refString="Shen CJ (1940 a) The brachyuran fauna of Hong Kong. Journal of the Hong Kong Fisheries Research Station, 1: 211 - 242." type="journal article" year="1940" yearSuffix="a">Shen, 1940a: 95</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFF4BA34FBB9FD51FA99FD49" author="Shen C" box="[1124,1300,738,760]" page="236" pageId="21" pageNumber="912" year="1940">
<bibRefCitation id="A52DE7B1FFF4BA34FBB9FD51FA99FD49" author="Shen" box="[1124,1300,738,760]" firstAuthor="Shen" page="236" pageId="21" pageNumber="912" pagination="69 - 104" refId="ref76967" refString="Shen C (1940 b) On the collection of crabs of south China. Bulletin of the Fan Memorial Institute of Biology, Zoology Series, Peiping, 10 (2): 69 - 104." type="journal article" year="1940" yearSuffix="b">Shen, 1940b: 236</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFF4BA34FAFDFD51FCF5FCA5" author="Kamita T" page="224" pageId="21" pageNumber="912" year="1941">
<bibRefCitation id="A52DE7B1FFF4BA34FAFDFD51FCF5FCA5" author="Kamita" firstAuthor="Kamita" page="224" pageId="21" pageNumber="912" refId="ref70869" refString="Kamita T (1941) Studies of the decapod crustaceans of Chosen. Pt. 1. Crabs. The Fisheries Society of Chosen, Keijo, 289 pp., 146 text-figs., 1 map." type="book" year="1941">Kamita, 1941: 224</bibRefCitation>
</treatmentCitation>
, fig. 124;
<treatmentCitation id="401DBC51FFF4BA34FC01FD4CFB0FFCA5" author="Sakai T" box="[988,1154,767,789]" page="202" pageId="21" pageNumber="912" year="1965">
<bibRefCitation id="A52DE7B1FFF4BA34FC01FD4CFB0FFCA5" author="Sakai" box="[988,1154,767,789]" firstAuthor="Sakai" page="202" pageId="21" pageNumber="912" pagination="1 - 92" refId="ref75676" refString="Sakai T (1965) The Crabs of the Sagami Bay collected by His Majesty the Emperor of Japan. Maruzen Co., Tokyo. xvi + 206 pp. [English section], figs. 1 - 27, pls. 1 - 100, pp. 1 - 92 [in Japanese] + 1 - 26 [Bibliography and Index in English] + 27 - 32 [Index in Japanese], 1 map." type="book chapter" year="1965">Sakai, 1965: 202</bibRefCitation>
</treatmentCitation>
, pl. 97 fig. 3;
<treatmentCitation id="401DBC51FFF4BA34FAD1FD4CFA2AFCA5" author="Kim HS" box="[1292,1447,767,789]" page="489" pageId="21" pageNumber="912" year="1973">
<bibRefCitation id="A52DE7B1FFF4BA34FAD1FD4CFA2AFCA5" author="Kim" box="[1292,1447,767,789]" firstAuthor="Kim" page="489" pageId="21" pageNumber="912" refId="ref70907" refString="Kim HS (1973) Anomura and Brachyura. In: Illustrated Encyclopaedia of Fauna and Flora of Korea. Vol. 14. Ministry of Education, Sam Wha Publishing Co., Republic of Korea, 694 pp., 112 pls." type="book" year="1973">Kim, 1973: 489</bibRefCitation>
</treatmentCitation>
, text-fig. 218, pl. 94 fig. 168.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFF4BA34FCF0FC8AFAB2FCFE" blockId="21.[813,1452,508,1307]" box="[813,1343,825,847]" pageId="21" pageNumber="912">
<treatmentCitationGroup id="E1ACBD6EFFF4BA34FCF0FC8AFAB2FCFE" box="[813,1343,825,847]" pageId="21" pageNumber="912">
<taxonomicName id="06BCE1C3FFF4BA34FCF0FC8AFBE6FCFF" authority="(De Haan, 1835)" box="[813,1131,825,846]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="912" phylum="Arthropoda" rank="species" species="intermedium" subGenus="Sesarma">
<emphasis id="F3C84652FFF4BA34FCF0FC8AFC0DFCFF" box="[813,896,825,846]" italics="true" pageId="21" pageNumber="912">Sesarma</emphasis>
(
<emphasis id="F3C84652FFF4BA34FC52FC8AFC6FFCFF" box="[911,994,825,846]" italics="true" pageId="21" pageNumber="912">Sesarma</emphasis>
)
<emphasis id="F3C84652FFF4BA34FC2FFC8AFBE6FCFF" box="[1010,1131,825,846]" italics="true" pageId="21" pageNumber="912">intermedium</emphasis>
</taxonomicName>
<treatmentCitation id="401DBC51FFF4BA34FB5BFC8AFAB6FCFE" author="Serene R" box="[1158,1339,825,847]" page="105" pageId="21" pageNumber="912" year="1968">
<bibRefCitation id="A52DE7B1FFF4BA34FB5BFC8AFAB6FCFE" author="Serene" box="[1158,1339,825,847]" firstAuthor="Serene" page="105" pageId="21" pageNumber="912" pagination="33 - 112" refId="ref76415" refString="Serene R (1968) The Brachyura of the Indo Pacific Region. In: Prodromus for a check list of the nonplanctonic marine fauna of South East Asia. Singapore National Academy of Science, Special Publication No. 1: 33 - 112." type="book chapter" year="1968">Serène, 1968: 105</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFF4BA34FCF0FCE5FA24FB75" blockId="21.[813,1452,508,1307]" pageId="21" pageNumber="912">
<treatmentCitationGroup id="E1ACBD6EFFF4BA34FCF0FCE5FA24FB75" pageId="21" pageNumber="912">
<taxonomicName id="06BCE1C3FFF4BA34FCF0FCE5FB98FCDB" baseAuthorityName="W.de Haan" baseAuthorityYear="1835" box="[813,1045,853,875]" class="Malacostraca" family="Sesarmidae" genus="Sesarmops" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="912" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFF4BA34FCF0FCE5FB98FCDB" box="[813,1045,853,875]" italics="true" pageId="21" pageNumber="912">Sesarmops intermedium</emphasis>
</taxonomicName>
<treatmentCitation id="401DBC51FFF4BA34FBF2FCE5FAA5FCDA" author="Serene R &amp; Soh CL" box="[1071,1320,853,876]" page="401" pageId="21" pageNumber="912" year="1970">
<bibRefCitation id="A52DE7B1FFF4BA34FBF2FCE5FAA5FCDA" author="Serene &amp; Soh" box="[1071,1320,853,876]" firstAuthor="Serene" page="401" pageId="21" pageNumber="912" pagination="387 - 416" refId="ref76461" refString="Serene R &amp; Soh CL (1970) New IndoPacific genera allied to Sesarma Say 1877 (Brachyura, Decapoda, Crustacea). Treubia, 27: 387 - 416." type="journal article" year="1970">Serène &amp; Soh, 1970: 401</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFF4BA34FAEFFCE6FCF5FC39" author="Sakai T" page="659" pageId="21" pageNumber="912" year="1976">
<bibRefCitation id="A52DE7B1FFF4BA34FAEFFCE6FCF5FC39" author="Sakai" firstAuthor="Sakai" page="659" pageId="21" pageNumber="912" pagination="1 - 251" refId="ref75757" refString="Sakai T (1976) Crabs of Japan and the Adjacent Seas. Kodansha Ltd., Tokyo. In three volumes. xxix + 773 pp. [English Text], 461 pp. [Japanese Text]. Pp. 1 - 16, pls. 1 - 251 [Plates volume]." type="book chapter" year="1976">Sakai, 1976: 659</bibRefCitation>
</treatmentCitation>
, pl. 224 fig. 3;
<bibRefCitation id="A52DE7B1FFF4BA34FBD5FCC0FB38FC36" author="Matsuzawa" box="[1032,1205,882,904]" firstAuthor="Matsuzawa" pageId="21" pageNumber="912" refId="ref72590" refString="Matsuzawa K (1977) Sea Shore Animals of Muroto. Kansai-Insatsu- Kogyi, Japan, unumbered pages, 126 pls." type="book" year="1977">Matsuzawa, 1977</bibRefCitation>
: pl. 111 fig. 1;
<treatmentCitation id="401DBC51FFF4BA34FA99FCC1FCE1FC15" author="Soh CL" page="11" pageId="21" pageNumber="912" year="1978">
<bibRefCitation id="A52DE7B1FFF4BA34FA99FCC1FCE1FC15" author="Soh" firstAuthor="Soh" page="11" pageId="21" pageNumber="912" pagination="9 - 22" refId="ref77463" refString="Soh CL (1978) On a collection of sesarmine crabs (Decapoda, Brachyura, Grapsidae) from Hong Kong. Memoirs of the Hong Kong Natural History Society, 13: 9 - 22." type="journal article" year="1978">Soh, 1978: 11</bibRefCitation>
</treatmentCitation>
, pl. 4b;
<treatmentCitation id="401DBC51FFF4BA34FC1EFC3CFBF2FC15" author="Takeda M" box="[963,1151,911,933]" page="221" pageId="21" pageNumber="912" year="1982">
<bibRefCitation id="A52DE7B1FFF4BA34FC1EFC3CFBF2FC15" author="Takeda" box="[963,1151,911,933]" firstAuthor="Takeda" page="221" pageId="21" pageNumber="912" refId="ref77752" refString="Takeda M (1982) Keys to the Japanese and foreign crustaceans fully illustrated in colors. Hokuryukan, Tokyo, vi + 285 pp., 58 pp. (keys)." type="book" year="1982">Takeda, 1982: 221</bibRefCitation>
</treatmentCitation>
, fig. 655;
<treatmentCitation id="401DBC51FFF4BA34FB34FC3CFA25FC15" author="Miyake S" box="[1257,1448,911,933]" page="179" pageId="21" pageNumber="912" year="1983">
<bibRefCitation id="A52DE7B1FFF4BA34FB34FC3CFA25FC15" author="Miyake" box="[1257,1448,911,933]" firstAuthor="Miyake" page="179" pageId="21" pageNumber="912" refId="ref72931" refString="Miyake S (1983) Japanese Crustacean Decapods and Stomatopods in Color, vol. II. Brachyura (Crabs). First Edition. Hoikusha, Osaka, viii + 277 pp., 64 pls., unnumbered figs." type="book" year="1983">Miyake, 1983: 179</bibRefCitation>
</treatmentCitation>
, pl. 60-3;
<treatmentCitation id="401DBC51FFF4BA34FC69FC1FFB5FFC70" author="Nagai S &amp; Nomura K" box="[948,1234,940,962]" page="44" pageId="21" pageNumber="912" year="1988">
<bibRefCitation id="A52DE7B1FFF4BA34FC69FC1FFB5FFC70" author="Nagai &amp; Nomura" box="[948,1234,940,962]" firstAuthor="Nagai" page="44" pageId="21" pageNumber="912" refId="ref73185" refString="Nagai S &amp; Nomura K (1988) Crustacea (Brachyura), [The guide book of marine organisms of Okinawa]. No. 7. Shinsei Tosho, Okinawa, 250 pp." type="book" year="1988">Nagai &amp; Nomura, 1988: 44</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFF4BA34FB3CFC1FFA2AFC73" author="Muraoka K" box="[1249,1447,940,962]" page="54" pageId="21" pageNumber="912" year="1998">
<bibRefCitation id="A52DE7B1FFF4BA34FB3CFC1FFA2AFC73" author="Muraoka" box="[1249,1447,940,962]" firstAuthor="Muraoka" page="54" pageId="21" pageNumber="912" pagination="1 - 67" refId="ref72999" refString="Muraoka K (1998) Catalogue of the Brachyuran and Anomuran Crabs donated by Prof. Dr. Tune Sakai to the Kanagawa Prefectural Museum. Catalogue of the Collection in the Kanagawa Prefectural Museum of Natural History, Odawara, Kanagawa, Japan, No. 11: 1 - 67." type="book chapter" year="1998">Muraoka, 1998: 54</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFF4BA34FC8EFC7AFBA4FC6F" author="Minemizu R" box="[851,1065,969,991]" page="298" pageId="21" pageNumber="912" year="2000">
<bibRefCitation id="A52DE7B1FFF4BA34FC8EFC7AFBA4FC6F" author="Minemizu" box="[851,1065,969,991]" firstAuthor="Minemizu" page="298" pageId="21" pageNumber="912" refId="ref72906" refString="Minemizu R (2000) [Marine decapod and stomatopod crustaceans mainly from Japan]. Bun-Ichi Sogo Shuppan, Tokyo, 344 pp." type="book" year="2000">Minemizu, 2000: 298</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFF4BA34FBE8FC7AFA82FC6F" author="Kobayashi S" box="[1077,1295,969,991]" page="122" pageId="21" pageNumber="912" year="2000">
<bibRefCitation id="A52DE7B1FFF4BA34FBE8FC7AFA82FC6F" author="Kobayashi" box="[1077,1295,969,991]" firstAuthor="Kobayashi" page="122" pageId="21" pageNumber="912" pagination="113 - 130" refId="ref70994" refString="Kobayashi S (2000) Distribution pattern and ecology of brachyuran crabs in the riverine environment: their significance in the ecosystem and present condition. Ecological Civil Engineering, 3 (1): 113 - 130." type="journal article" year="2000">Kobayashi, 2000: 122</bibRefCitation>
</treatmentCitation>
, fig. 2o;
<treatmentCitation id="401DBC51FFF4BA34FAB4FC7AFC42FC4A" author="Tsai M-L &amp; Li J-J &amp; Dai C-F" page="61" pageId="21" pageNumber="912" year="2000">
<bibRefCitation id="A52DE7B1FFF4BA34FAB4FC7AFC42FC4A" author="Tsai" etAl="et al." firstAuthor="Tsai" page="61" pageId="21" pageNumber="912" pagination="61 - 78" refId="ref78337" refString="Tsai M-L, Li J-J &amp; Dai C-F (2000) Is large body size always advantageous for terrestrial adaptation? A study of water balance in a semi-terrestrial crab, Sesarmops intermedium (Crustacea: Grapsidae). Evolutionary Ecology, 14: 61 - 78." type="journal article" year="2000">Tsai et al., 2000: 61</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFF4BA34FC01FC55FB19FC4A" author="Ng PKL &amp; Wang C-H &amp; Ho P-H &amp; Shih H-T" box="[988,1172,997,1019]" page="43" pageId="21" pageNumber="912" year="2001">
<bibRefCitation id="A52DE7B1FFF4BA34FC01FC55FB19FC4A" author="Ng" box="[988,1172,997,1019]" etAl="et al." firstAuthor="Ng" page="43" pageId="21" pageNumber="912" pagination="1 - 86" refId="ref74237" refString="Ng PKL, Wang C-H, Ho P-H &amp; Shih H-T (2001) An annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). National Taiwan Museum Special Publication Series, 11: 1 - 86, 8 colour pls." type="journal article" year="2001">Ng et al., 2001: 43</bibRefCitation>
</treatmentCitation>
[includes references for this species from
<collectingCountry id="B9ABDAD0FFF4BA34FC04FBB1FBACFBA9" box="[985,1057,1026,1048]" name="Taiwan" pageId="21" pageNumber="912">Taiwan</collectingCountry>
up to 2001];
<treatmentCitation id="401DBC51FFF4BA34FB72FBB0FAB8FBA6" author="Ho P-H" box="[1199,1333,1026,1048]" page="32" pageId="21" pageNumber="912" year="2003">
<bibRefCitation id="A52DE7B1FFF4BA34FB72FBB0FAB8FBA6" author="Ho" box="[1199,1333,1026,1048]" firstAuthor="Ho" page="32" pageId="21" pageNumber="912" refId="ref70247" refString="Ho P-H (2003) Land crabs in the coastal forest of Kenting National Park. Kenting National Park, Pingtung, 47 pp." type="book" year="2003">Ho, 2003: 32</bibRefCitation>
</treatmentCitation>
, 33;
<treatmentCitation id="401DBC51FFF4BA34FAB7FBB1FC7AFB84" author="Cuesta JA &amp; Guerao G &amp; Liu H-C &amp; Schubart CD" page="162" pageId="21" pageNumber="912" year="2006">
<bibRefCitation id="A52DE7B1FFF4BA34FAB7FBB1FC7AFB84" author="Cuesta" etAl="et al." firstAuthor="Cuesta" page="162" pageId="21" pageNumber="912" pagination="151 - 173" refId="ref68440" refString="Cuesta JA, Guerao G, Liu H-C &amp; Schubart CD (2006) Morphology of the first zoeal stages of eleven Sesarmidae (Crustacea, Brachyura, Thoracotremata) from the Indo-West Pacific, with a summary of familial larval characters. Invertebrate Reproduction &amp; Development, 49 (3): 151 - 173." type="journal article" year="2006">Cuesta et al., 2006: 162</bibRefCitation>
</treatmentCitation>
, fig. 13AE;
<treatmentCitation id="401DBC51FFF4BA34FBA2FBACFA25FB85" author="Takeda M &amp; Ueshima R" box="[1151,1448,1055,1077]" page="99" pageId="21" pageNumber="912" year="2006">
<bibRefCitation id="A52DE7B1FFF4BA34FBA2FBACFA25FB85" author="Takeda &amp; Ueshima" box="[1151,1448,1055,1077]" firstAuthor="Takeda" page="99" pageId="21" pageNumber="912" pagination="61 - 105" refId="ref77817" refString="Takeda M &amp; Ueshima R (2006) Crabs preserved in the Department of Zoology, the University Museum, the University of Tokyo. In: Ueshima R (ed.) Catalogue of Invertebrate Collection Deposited in the Department of Zoology, The University Museum, the University of Tokyo. The University Museum, the University of Tokyo Materials Reports, 62: 61 - 105. [In Japanese]" type="journal article" year="2006">Takeda &amp; Ueshima, 2006: 99</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFF4BA34FC8FFB8FFBA0FBE0" author="Yang S &amp; Chen H &amp; Jiang W" box="[850,1069,1084,1106]" page="802" pageId="21" pageNumber="912" year="2008">
<bibRefCitation id="A52DE7B1FFF4BA34FC8FFB8FFBA0FBE0" author="Yang" box="[850,1069,1084,1106]" etAl="et al." firstAuthor="Yang" page="802" pageId="21" pageNumber="912" pagination="761 - 810" refId="ref78808" refString="Yang S, Chen H &amp; Jiang W (2008) Crustacea: Decapoda Brachyura. In: Liu R (ed.) Checklist of Marine Biota of China Seas. Institute of Oceanology, Chinese Academy of Sciences. Pp. 761 - 810." type="book chapter" year="2008">Yang et al., 2008: 802</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFF4BA34FBE5FB8FFB41FBE0" author="Lee J-H" box="[1080,1228,1084,1105]" page="133" pageId="21" pageNumber="912" year="2008">
<bibRefCitation id="A52DE7B1FFF4BA34FBE5FB8FFB41FBE0" author="Lee" box="[1080,1228,1084,1105]" firstAuthor="Lee" page="133" pageId="21" pageNumber="912" refId="ref71440" refString="Lee J-H (2008) A Field Guide to Crabs in Taiwan. Bigtrees Company, Taipei, 174 pp." type="book" year="2008">Lee, 2008: 133</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFF4BA34FB0AFB8FFAD1FBE0" author="Liu H-C" box="[1239,1372,1084,1106]" page="49" pageId="21" pageNumber="912" year="2009">
<bibRefCitation id="A52DE7B1FFF4BA34FB0AFB8FFAD1FBE0" author="Liu" box="[1239,1372,1084,1106]" firstAuthor="Liu" page="49" pageId="21" pageNumber="912" refId="ref71856" refString="Liu H-C (2009) Diversity, population biology and managements of land crabs in the Kenting National Park. The National Kenting Headquarters, Kenting, 77 pp." type="book" year="2009">Liu, 2009: 49</bibRefCitation>
</treatmentCitation>
, fig. 19;
<treatmentCitation id="401DBC51FFF4BA34FC8FFBEAFB00FBDE" author="Naderloo R &amp; Schubart CD" box="[850,1165,1113,1135]" page="62" pageId="21" pageNumber="912" year="2009">
<bibRefCitation id="A52DE7B1FFF4BA34FC8FFBEAFB00FBDE" author="Naderloo &amp; Schubart" box="[850,1165,1113,1135]" firstAuthor="Naderloo" page="62" pageId="21" pageNumber="912" pagination="61 - 68" refId="ref73102" refString="Naderloo R &amp; Schubart CD (2009) Redescription and mitochondrial identification of Chiromantes boulengeri (Calman, 1920) (Decapoda: Brachyura: Sesarmidae) based on fresh material from the Persian Gulf, Iran. Zootaxa, 2128: 61 - 68." type="journal article" year="2009">Naderloo &amp; Schubart, 2009: 62</bibRefCitation>
</treatmentCitation>
, 63, 67;
<treatmentCitation id="401DBC51FFF4BA34FB3AFBEAFCE1FB3A" author="Liu H-C &amp; Wang C-H" page="62" pageId="21" pageNumber="912" year="2010">
<bibRefCitation id="A52DE7B1FFF4BA34FB3AFBEAFCE1FB3A" author="Liu &amp; Wang" firstAuthor="Liu" page="62" pageId="21" pageNumber="912" refId="ref71886" refString="Liu H-C &amp; Wang C-H (2010) Taiwan Coastal Wetland Crabs. Wild Bird Society of Taipei, Taipei, 79 pp." type="book" year="2010">Liu &amp; Wang, 2010: 62</bibRefCitation>
</treatmentCitation>
, 63;
<treatmentCitation id="401DBC51FFF4BA34FC7CFBC6FA9FFB3A" author="Rademacher M &amp; Mengedoht O" box="[929,1298,1141,1163]" page="31" pageId="21" pageNumber="912" year="2011">
<bibRefCitation id="A52DE7B1FFF4BA34FC7CFBC6FA9FFB3A" author="Rademacher &amp; Mengedoht" box="[929,1298,1141,1163]" firstAuthor="Rademacher" page="31" pageId="21" pageNumber="912" refId="ref74843" refString="Rademacher M &amp; Mengedoht O (2011) Krabben-Fibel. Die schonsten Krabben fur das Aquaterrarium. Dahne Verlag, Ettlingen, Germany, 95 pp." type="book" year="2011">Rademacher &amp; Mengedoht, 2011: 31</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFF4BA34FAFDFBC5FC34FB19" author="Huang H-D &amp; Liu H-C" page="267" pageId="21" pageNumber="912" year="2012">
<bibRefCitation id="A52DE7B1FFF4BA34FAFDFBC5FC34FB19" author="Huang &amp; Liu" firstAuthor="Huang" page="267" pageId="21" pageNumber="912" pagination="267 - 267" refId="ref70431" refString="Huang H-D &amp; Liu H-C (2012) Comparison of visual orientation between two land crabs Sesarmops intermedium and Scandarma lintou in Kenting National Park, southern Taiwan. Integrative and Comparative Biology, 52: 267 - 267." type="journal article" year="2012">Huang &amp; Liu, 2012: 267</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFF4BA34FC18FB21FB18FB19" author="Li J-J &amp; Chiu Y-W" box="[965,1173,1170,1192]" page="57" pageId="21" pageNumber="912" year="2013">
<bibRefCitation id="A52DE7B1FFF4BA34FC18FB21FB18FB19" author="Li &amp; Chiu" box="[965,1173,1170,1192]" firstAuthor="Li" page="57" pageId="21" pageNumber="912" refId="ref71539" refString="Li J-J &amp; Chiu Y-W (2013) Land crabs of Hengchun Peninsula. National Museum of Marine Biology &amp; Aquarium, Pingtung, 95 pp." type="book" year="2013">Li &amp; Chiu, 2013: 57</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFF4BA34FB7CFB20FA2AFB16" author="Toyota K &amp; Seki S" box="[1185,1447,1170,1192]" page="192" pageId="21" pageNumber="912" year="2014">
<bibRefCitation id="A52DE7B1FFF4BA34FB7CFB20FA2AFB16" author="Toyota &amp; Seki" box="[1185,1447,1170,1192]" firstAuthor="Toyota" page="192" pageId="21" pageNumber="912" refId="ref78208" refString="Toyota K &amp; Seki S (2014) Shrimps and Crabs. Nature Watching Guidebook, Freshwater Shrimps and Crabs in Japan. Seibundo Shinkosha Publishing, 256 pp." type="book" year="2014">Toyota &amp; Seki, 2014: 192</bibRefCitation>
</treatmentCitation>
, 193;
<treatmentCitation id="401DBC51FFF4BA34FC58FB03FBDAFB74" author="Ng PKL &amp; Shih H-T &amp; Ho P-H &amp; Wang C-H" box="[901,1111,1199,1221]" page="106" pageId="21" pageNumber="912" year="2017">
<bibRefCitation id="A52DE7B1FFF4BA34FC58FB03FBDAFB74" author="Ng" box="[901,1111,1199,1221]" etAl="et al." firstAuthor="Ng" page="106" pageId="21" pageNumber="912" pagination="1 - 185" refId="ref74189" refString="Ng PKL, Shih H-T, Ho P-H &amp; Wang C-H (2017 b) An updated annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). Journal of the National Taiwan Museum, 70 (3 - 4): 1 - 185." type="journal article" year="2017" yearSuffix="b">Ng et al., 2017b: 106</bibRefCitation>
</treatmentCitation>
[for other references for
<collectingCountry id="B9ABDAD0FFF4BA34FA8FFB1DFA12FB75" box="[1362,1439,1198,1220]" name="Taiwan" pageId="21" pageNumber="912">Taiwan</collectingCountry>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFF4BA34FCF0FB7FFA9BFB50" blockId="21.[813,1452,508,1307]" box="[813,1302,1228,1250]" pageId="21" pageNumber="912">
<treatmentCitationGroup id="E1ACBD6EFFF4BA34FCF0FB7FFA9BFB50" box="[813,1302,1228,1250]" pageId="21" pageNumber="912">
<taxonomicName id="06BCE1C3FFF4BA34FCF0FB7FFBAEFB50" authorityName="De Haan" authorityYear="1835" box="[813,1059,1228,1249]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="912" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="F3C84652FFF4BA34FCEAFB7FFC12FB50" box="[823,927,1228,1249]" italics="true" pageId="21" pageNumber="912">Sesarmops</emphasis>
<emphasis id="F3C84652FFF4BA34FC6FFB7FFBAEFB50" box="[946,1059,1228,1249]" italics="true" pageId="21" pageNumber="912">intermedius</emphasis>
</taxonomicName>
<treatmentCitation id="401DBC51FFF4BA34FBE2FB7FFA9FFB50" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[1087,1298,1228,1250]" page="220" pageId="21" pageNumber="912" year="2008">
<bibRefCitation id="A52DE7B1FFF4BA34FBE2FB7FFA9FFB50" author="Ng" box="[1087,1298,1228,1250]" etAl="et al." firstAuthor="Ng" page="220" pageId="21" pageNumber="912" pagination="1 - 286" refId="ref73754" refString="Ng PKL, Guinot D &amp; Davie PJF (2008 a) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008" yearSuffix="a">Ng et al., 2008a: 220</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFF4BA34FCF0FB5AFB0EFAAA" blockId="21.[813,1452,508,1307]" pageId="21" pageNumber="912">
<treatmentCitationGroup id="E1ACBD6EFFF4BA34FCF0FB5AFB0EFAAA" pageId="21" pageNumber="912">
<taxonomicName id="06BCE1C3FFF4BA34FCF0FB5AFB83FB4F" ID-CoL="7Z2CM" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[813,1038,1257,1278]" class="Malacostraca" family="Sesarmidae" genus="Sesarmops" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="912" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="F3C84652FFF4BA34FCF0FB5AFB83FB4F" box="[813,1038,1257,1278]" italics="true" pageId="21" pageNumber="912">Sesarmops intermedius</emphasis>
</taxonomicName>
<treatmentCitation id="401DBC51FFF4BA34FBF0FB5AFA9CFB4E" author="Hong J-S &amp; Choi J-W &amp; Tsutsumi H" box="[1069,1297,1257,1279]" page="259" pageId="21" pageNumber="912" year="2010">
<bibRefCitation id="A52DE7B1FFF4BA34FBF0FB5AFA9CFB4E" author="Hong" box="[1069,1297,1257,1279]" etAl="et al." firstAuthor="Hong" page="259" pageId="21" pageNumber="912" pagination="255 - 263" refId="ref70376" refString="Hong J-S, Choi J-W &amp; Tsutsumi H (2010) Concluding remarks on the joint survey of macrobenthic fauna on Suncheon Tidal Flats by the participants of &quot; Korea and Japan Joint Symposium on Biology of Tidal Flats 2009 &quot;. Plankton Benthos Research, 5 (Supplement): 255 - 263." type="journal article" year="2010">Hong et al., 2010: 259</bibRefCitation>
</treatmentCitation>
, fig. 3b;
<treatmentCitation id="401DBC51FFF4BA34FAB0FB5AFC50FAAA" author="Lee S-K &amp; Jung J &amp; Kang S &amp; Kim W" page="183" pageId="21" pageNumber="912" year="2012">
<bibRefCitation id="A52DE7B1FFF4BA34FAB0FB5AFC50FAAA" author="Lee" etAl="et al." firstAuthor="Lee" page="183" pageId="21" pageNumber="912" pagination="183 - 185" refId="ref71461" refString="Lee S-K, Jung J, Kang S &amp; Kim W (2012) Isolation and characterization of microsatellite markers for the endangered crab species Sesarmops intermedius. Conservation Genetics and Resources, 4: 183 - 185." type="journal article" year="2012">Lee et al., 2012: 183</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFF4BA34FC35FAB5FB0DFAAA" author="Lee HJ" box="[1000,1152,1285,1307]" page="321" pageId="21" pageNumber="912" year="2015">
<bibRefCitation id="A52DE7B1FFF4BA34FC35FAB5FB0DFAAA" author="Lee" box="[1000,1152,1285,1307]" firstAuthor="Lee" page="321" pageId="21" pageNumber="912" pagination="321 - 324" refId="ref71400" refString="Lee HJ (2015) First zoea of Sesarmops intermedius (De Haan, 1835) (Crustacea: Decapoda: Brachyura: Sesarmidae). Fisheries &amp; Aquatic Sciences, 18 (3): 321 - 324." type="journal article" year="2015">Lee, 2015: 321</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFF4BA39FCF0FAF0FDF0FEA4" lastPageId="24" lastPageNumber="915" pageId="21" pageNumber="912" type="materials_examined">
<paragraph id="C1039A40FFF4BA39FCF0FAF0FDF0FEA4" blockId="21.[812,1452,1347,2108]" lastBlockId="24.[136,775,189,278]" lastPageId="24" lastPageNumber="915" pageId="21" pageNumber="912">
<emphasis id="F3C84652FFF4BA34FCF0FAF0FB9CFAEA" bold="true" box="[813,1041,1347,1371]" pageId="21" pageNumber="912">Material examined</emphasis>
.
<materialsCitation id="71D4901DFFF4BA34FBFEFAF7FB7DFACD" ID-GBIF-Occurrence="3763237368" collectionCode="RMNH" collectorName="Japan &amp; H. Burger." pageId="21" pageNumber="912" specimenCode="D 165" specimenCount="1" typeStatus="lectotype">
<typeStatus id="1E0724E2FFF4BA34FBFEFAF7FB1FFAED" box="[1059,1170,1348,1372]" pageId="21" pageNumber="912" type="lectotype">Lectotype</typeStatus>
male (23.0 ×
<quantity id="064437A5FFF4BA34FAE0FAF7FA29FAED" box="[1341,1444,1348,1372]" metricMagnitude="-2" metricUnit="m" metricValue="1.9899999999999998" pageId="21" pageNumber="912" unit="mm" value="19.9">19.9 mm</quantity>
) (
<collectionCode id="A7AD0285FFF4BA34FCE9FAD7FC0BFACD" box="[820,902,1380,1404]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="912" type="Museum">RMNH</collectionCode>
<specimenCode id="911A323BFFF4BA34FC4CFAD7FC57FACD" box="[913,986,1380,1404]" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="912" type="Museum">D 165</specimenCode>
),
<collectorName id="6C49FF96FFF4BA34FC2CFAD7FBBCFACA" box="[1009,1073,1380,1403]" pageId="21" pageNumber="912">Japan</collectorName>
, coll.
<collectorName id="6C49FF96FFF4BA34FBA4FAD7FB7DFACD" box="[1145,1264,1380,1404]" pageId="21" pageNumber="912">H. Bürger.</collectorName>
</materialsCitation>
<materialsCitation id="71D4901DFFF4BA34FB27FAD7FADCFA2D" ID-GBIF-Occurrence="3763237435" collectionCode="RMNH" collectorName="Others" country="Japan" pageId="21" pageNumber="912" specimenCode="D 165" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<typeStatus id="1E0724E2FFF4BA34FB27FAD7FA1FFACD" box="[1274,1426,1380,1404]" pageId="21" pageNumber="912" type="paralectotype">Paralectotype</typeStatus>
:
<specimenCount id="D7BA51C9FFF4BA34FA42FAD7FCFAFA2D" pageId="21" pageNumber="912" type="female">1 female</specimenCount>
(
<collectionCode id="A7AD0285FFF4BA34FC54FA37FC51FA2D" box="[905,988,1412,1436]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="912" type="Museum">RMNH</collectionCode>
<specimenCode id="911A323BFFF4BA34FC3AFA37FBBCFA2D" box="[999,1073,1412,1436]" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="912" type="Museum">D 165</specimenCode>
), same data as lectotype.
</materialsCitation>
<materialsCitation id="71D4901DFFF4BA34FA86FA37FB9BFA6D" ID-GBIF-Occurrence="3763237530" collectionCode="RMNH" collectorName="Others" country="Japan" pageId="21" pageNumber="912" specimenCode="D 165" specimenCount="1" specimenCount-female="1" specimenCount-male="1" typeStatus="paralectotype">
<collectorName id="6C49FF96FFF4BA34FA86FA37FA25FA2D" box="[1371,1448,1412,1436]" pageId="21" pageNumber="912">Others</collectorName>
:
<collectingCountry id="B9ABDAD0FFF4BA34FCF0FA17FC0CFA0D" box="[813,897,1444,1468]" name="Japan" pageId="21" pageNumber="912">JAPAN</collectingCountry>
<specimenCount id="D7BA51C9FFF4BA34FC79FA17FC7CFA0D" box="[932,1009,1444,1468]" pageId="21" pageNumber="912" type="male">1 male</specimenCount>
(27.1 ×
<quantity id="064437A5FFF4BA34FB84FA17FB32FA0D" box="[1113,1215,1444,1468]" metricMagnitude="-2" metricUnit="m" metricValue="2.52" pageId="21" pageNumber="912" unit="mm" value="25.2">25.2 mm</quantity>
) (
<collectionCode id="A7AD0285FFF4BA34FB05FA17FA83FA0D" box="[1240,1294,1444,1468]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="912">ZRC</collectionCode>
1970.2.23.6), coll.
<collectorName id="6C49FF96FFF4BA34FCB4FA77FC40FA6D" box="[873,973,1476,1500]" pageId="21" pageNumber="912">T. Sakai</collectorName>
, 1968
</materialsCitation>
;
<materialsCitation id="71D4901DFFF4BA34FBFBFA77FA2AFA4D" ID-GBIF-Occurrence="3763237425" collectionCode="ZRC" collectorName="T. Sakai" country="Japan" location="Sagami Bay" municipality="Manazuru" pageId="21" pageNumber="912" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFF4BA34FBFBFA77FB04FA6D" box="[1062,1161,1476,1500]" pageId="21" pageNumber="912" type="female">1 female</specimenCount>
(22.7 × 20.0 mm) (
<collectionCode id="A7AD0285FFF4BA34FAA8FA77FA21FA6D" box="[1397,1452,1476,1500]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="912">ZRC</collectionCode>
1970.8.27.3),
<collectingMunicipality id="2167003AFFF4BA34FC1EFA57FBBFFA4A" box="[963,1074,1508,1531]" pageId="21" pageNumber="912">Manazuru</collectingMunicipality>
,
<location id="C463CC9BFFF4BA34FBE3FA57FB4EFA4D" LSID="urn:lsid:plazi:treatment:49152B56FFF4BA3AFC57FECEFF73FAA4:C463CC9BFFF4BA34FBE3FA57FB4EFA4D" box="[1086,1219,1508,1532]" country="Japan" municipality="Manazuru" name="Sagami Bay" pageId="21" pageNumber="912">Sagami Bay</location>
, coll.
<collectorName id="6C49FF96FFF4BA34FADBFA57FAEEFA4D" box="[1286,1379,1508,1532]" pageId="21" pageNumber="912">T. Sakai</collectorName>
, 1968
</materialsCitation>
;
<materialsCitation id="71D4901DFFF4BA34FCF0F9B7FC55F9ED" ID-GBIF-Occurrence="3763237479" collectingDate="2000-08-30" collectionCode="ZRC" collectorName="H. Suzuki" country="Japan" location="Satsuma Peninsula" municipality="Izaku River" pageId="21" pageNumber="912" specimenCount="1" specimenCount-female="1" stateProvince="Kagoshima" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFF4BA34FCF0F9B7FC04F9AD" box="[813,905,1540,1564]" pageId="21" pageNumber="912" type="female">1 female</specimenCount>
(23.0 ×
<quantity id="064437A5FFF4BA34FC3CF9B7FBCDF9AD" box="[993,1088,1540,1564]" metricMagnitude="-2" metricUnit="m" metricValue="2.04" pageId="21" pageNumber="912" unit="mm" value="20.4">20.4 mm</quantity>
) (
<collectionCode id="A7AD0285FFF4BA34FB88F9B7FB03F9AD" box="[1109,1166,1540,1564]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="912">ZRC</collectionCode>
), river mouth,
<collectingMunicipality id="2167003AFFF4BA34FAF5F9B7FA2AF9AD" box="[1320,1447,1540,1564]" pageId="21" pageNumber="912">Izaku River</collectingMunicipality>
,
<location id="C463CC9BFFF4BA34FCF0F997FC74F98D" LSID="urn:lsid:plazi:treatment:49152B56FFF4BA3AFC57FECEFF73FAA4:C463CC9BFFF4BA34FCF0F997FC74F98D" box="[813,1017,1572,1596]" country="Japan" municipality="Izaku River" name="Satsuma Peninsula" pageId="21" pageNumber="912" stateProvince="Kagoshima">Satsuma Peninsula</location>
,
<collectingRegion id="037854A2FFF4BA34FBD8F997FB7BF98D" box="[1029,1270,1572,1596]" country="Japan" name="Kagoshima" pageId="21" pageNumber="912">Kagoshima Prefecture</collectingRegion>
, coll.
<collectorName id="6C49FF96FFF4BA34FAE4F997FA25F98D" box="[1337,1448,1572,1596]" pageId="21" pageNumber="912">H. Suzuki</collectorName>
,
<date id="B502BC80FFF4BA34FCF0F9F7FC55F9ED" box="[813,984,1604,1628]" pageId="21" pageNumber="912" value="2000-08-30">
<collectingDate id="A5464568FFF4BA34FCF0F9F7FC55F9ED" box="[813,984,1604,1628]" pageId="21" pageNumber="912" value="2000-08-30">30 August 2000</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71D4901DFFF4BA34FC3CF9F7FC56F90D" ID-GBIF-Occurrence="3763237312" collectingDate="2002-09-02" collectionCode="ZRC" collectorName="Japan &amp; J. Lai" country="Japan" county="Tomioka" location="Shikizuki" municipality="Amakusa" pageId="21" pageNumber="912" specimenCode="ZRC 2013.0140" specimenCount="4" specimenCount-female="4" stateProvince="Kagoshima" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFF4BA34FC3CF9F7FBBCF9ED" box="[993,1073,1604,1628]" pageId="21" pageNumber="912" type="male">4 males</specimenCount>
(largest 27.0 × 25.0 mm),
<specimenCount id="D7BA51C9FFF4BA34FA9BF9F7FA21F9ED" box="[1350,1452,1604,1628]" pageId="21" pageNumber="912" type="female">4 females</specimenCount>
(largest 32.1 ×
<quantity id="064437A5FFF4BA34FC15F9D7FBABF9CD" box="[968,1062,1636,1660]" metricMagnitude="-2" metricUnit="m" metricValue="2.93" pageId="21" pageNumber="912" unit="mm" value="29.3">29.3 mm</quantity>
) (
<specimenCode id="911A323BFFF4BA34FBE7F9D7FB64F9CD" box="[1082,1257,1636,1660]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="912">ZRC 2013.0140</specimenCode>
), in brackish water stream,
<collectingCounty id="2862E2CCFFF4BA34FC5CF937FC69F92D" box="[897,996,1668,1692]" pageId="21" pageNumber="912">Tomioka</collectingCounty>
,
<collectingMunicipality id="2167003AFFF4BA34FC32F937FBDAF92D" box="[1007,1111,1668,1692]" pageId="21" pageNumber="912">Amakusa</collectingMunicipality>
,
<location id="C463CC9BFFF4BA34FBBCF937FB47F92D" LSID="urn:lsid:plazi:treatment:49152B56FFF4BA3AFC57FECEFF73FAA4:C463CC9BFFF4BA34FBBCF937FB47F92D" box="[1121,1226,1668,1692]" country="Japan" county="Tomioka" municipality="Amakusa" name="Shikizuki" pageId="21" pageNumber="912" stateProvince="Kagoshima">Shikizuki</location>
,
<collectorName id="6C49FF96FFF4BA34FB08F937FA99F92A" box="[1237,1300,1668,1691]" pageId="21" pageNumber="912">Japan</collectorName>
, coll.
<collectorName id="6C49FF96FFF4BA34FA88F937FA1EF92D" box="[1365,1427,1668,1692]" pageId="21" pageNumber="912">J. Lai</collectorName>
,
<date id="B502BC80FFF4BA34FA43F937FC56F90D" pageId="21" pageNumber="912" value="2002-09-02">
<collectingDate id="A5464568FFF4BA34FA43F937FC56F90D" pageId="21" pageNumber="912" value="2002-09-02">2 September 2002</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71D4901DFFF4BA34FC38F917FC4EF8AD" ID-GBIF-Occurrence="3763237572" collectingDate="2000-06-15" collectionCode="ZRC" collectorName="Y. Cai &amp; N. K. Ng" country="Japan" location="Ryukyu Islands" municipality="Iriomote Island" pageId="21" pageNumber="912" specimenCode="ZRC 2000.2262" specimenCount="2" specimenCount-female="2" stateProvince="Kagoshima" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFF4BA34FC38F917FBA6F90D" box="[997,1067,1700,1724]" pageId="21" pageNumber="912" type="male">1 male</specimenCount>
(36.4 ×
<quantity id="064437A5FFF4BA34FB5CF917FB6CF90D" box="[1153,1249,1700,1724]" metricMagnitude="-2" metricUnit="m" metricValue="3.2399999999999998" pageId="21" pageNumber="912" unit="mm" value="32.4">32.4 mm</quantity>
),
<specimenCount id="D7BA51C9FFF4BA34FB2FF917FADAF90D" box="[1266,1367,1700,1724]" pageId="21" pageNumber="912" type="female">2 females</specimenCount>
(largest 29.0 ×
<quantity id="064437A5FFF4BA34FCA9F977FC5EF96A" box="[884,979,1732,1756]" metricMagnitude="-2" metricUnit="m" metricValue="2.69" pageId="21" pageNumber="912" unit="mm" value="26.9">26.9 mm</quantity>
, smallest 33.5 ×
<quantity id="064437A5FFF4BA34FBA2F977FB53F96D" box="[1151,1246,1732,1756]" metricMagnitude="-2" metricUnit="m" metricValue="3.04" pageId="21" pageNumber="912" unit="mm" value="30.4">30.4 mm</quantity>
) (
<specimenCode id="911A323BFFF4BA34FB2CF977FA2DF96D" box="[1265,1440,1732,1756]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="912">ZRC 2000.2262</specimenCode>
),
<collectingMunicipality id="2167003AFFF4BA34FCF1F957FC5BF94D" box="[812,982,1764,1788]" pageId="21" pageNumber="912">Iriomote Island</collectingMunicipality>
,
<location id="C463CC9BFFF4BA34FC3EF957FB1DF94D" LSID="urn:lsid:plazi:treatment:49152B56FFF4BA3AFC57FECEFF73FAA4:C463CC9BFFF4BA34FC3EF957FB1DF94D" box="[995,1168,1764,1788]" country="Japan" municipality="Iriomote Island" name="Ryukyu Islands" pageId="21" pageNumber="912" stateProvince="Kagoshima">Ryukyu Islands</location>
, coll.
<collectorName id="6C49FF96FFF4BA34FB0BF957FA93F94D" box="[1238,1310,1764,1788]" pageId="21" pageNumber="912">Y. Cai</collectorName>
&amp;
<collectorName id="6C49FF96FFF4BA34FA9BF957FA2AF94D" box="[1350,1447,1764,1788]" pageId="21" pageNumber="912">N.K. Ng</collectorName>
,
<date id="B502BC80FFF4BA34FCF0F8B7FC4EF8AD" box="[813,963,1796,1820]" pageId="21" pageNumber="912" value="2000-06-15">
<collectingDate id="A5464568FFF4BA34FCF0F8B7FC4EF8AD" box="[813,963,1796,1820]" pageId="21" pageNumber="912" value="2000-06-15">15 June 2000</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71D4901DFFF4BA34FC12F8B7FBF8F8ED" ID-GBIF-Occurrence="3763237431" collectingDate="2000-06-15" collectionCode="ZRC" collectorName="Y. X. Cai &amp; N. K. Ng" country="Japan" location="Nagama river" municipality="Japan" pageId="21" pageNumber="912" specimenCode="ZRC 2000.2265" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFF4BA34FC12F8B7FBA2F8AD" box="[975,1071,1796,1820]" pageId="21" pageNumber="912" type="female">1 female</specimenCount>
(
<specimenCode id="911A323BFFF4BA34FBE3F8B7FB7EF8AD" box="[1086,1267,1796,1820]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="912">ZRC 2000.2265</specimenCode>
),
<collectingMunicipality id="2167003AFFF4BA34FADBF8B7FAC8F8AA" box="[1286,1349,1796,1819]" pageId="21" pageNumber="912">Japan</collectingMunicipality>
,
<location id="C463CC9BFFF4BA34FA8FF8B7FCECF88D" LSID="urn:lsid:plazi:treatment:49152B56FFF4BA3AFC57FECEFF73FAA4:C463CC9BFFF4BA34FA8FF8B7FCECF88D" country="Japan" municipality="Japan" name="Nagama river" pageId="21" pageNumber="912" stateProvince="Okinawa">Nagama river</location>
second site from headwaters,
<collectingRegion id="037854A2FFF4BA34FB19F897FAA7F88D" box="[1220,1322,1828,1852]" country="Japan" name="Okinawa" pageId="21" pageNumber="912">Okinawa</collectingRegion>
, coll.
<collectorName id="6C49FF96FFF4BA34FAABF897FCDFF8ED" pageId="21" pageNumber="912">Y.X. Cai</collectorName>
&amp;
<collectorName id="6C49FF96FFF4BA34FCABF8F7FC58F8ED" box="[886,981,1860,1884]" pageId="21" pageNumber="912">N.K. Ng</collectorName>
,
<date id="B502BC80FFF4BA34FC3CF8F7FBF8F8ED" box="[993,1141,1860,1884]" pageId="21" pageNumber="912" value="2000-06-15">
<collectingDate id="A5464568FFF4BA34FC3CF8F7FBF8F8ED" box="[993,1141,1860,1884]" pageId="21" pageNumber="912" value="2000-06-15">15 June 2000</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71D4901DFFF4BA34FB5CF8F7FBBCF80D" ID-GBIF-Occurrence="3763237420" collectingDate="2000-06-15" collectionCode="ZRC" collectorName="Y. X. Cai &amp; N. K. Ng" country="Japan" location="Nagama River" municipality="Japan" pageId="21" pageNumber="912" specimenCode="ZRC 2000.2266" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFF4BA34FB5CF8F7FB52F8ED" box="[1153,1247,1860,1884]" pageId="21" pageNumber="912" type="female">1 female</specimenCount>
(
<specimenCode id="911A323BFFF4BA34FB33F8F7FA2CF8ED" box="[1262,1441,1860,1884]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="912">ZRC 2000.2266</specimenCode>
),
<collectingMunicipality id="2167003AFFF4BA34FCF0F8D7FCE1F8CA" box="[813,876,1892,1915]" pageId="21" pageNumber="912">Japan</collectingMunicipality>
, along the track to the headwaters of
<location id="C463CC9BFFF4BA34FAD5F8D7FA2AF8CD" LSID="urn:lsid:plazi:treatment:49152B56FFF4BA3AFC57FECEFF73FAA4:C463CC9BFFF4BA34FAD5F8D7FA2AF8CD" box="[1288,1447,1892,1916]" country="Japan" municipality="Japan" name="Nagama River" pageId="21" pageNumber="912" stateProvince="Okinawa">Nagama River</location>
, found in dead fallen tree trunk,
<collectingRegion id="037854A2FFF4BA34FB5AF837FB67F82D" box="[1159,1258,1924,1948]" country="Japan" name="Okinawa" pageId="21" pageNumber="912">Okinawa</collectingRegion>
, coll.
<collectorName id="6C49FF96FFF4BA34FAF0F837FA03F82D" box="[1325,1422,1924,1948]" pageId="21" pageNumber="912">Y.X. Cai</collectorName>
&amp;
<collectorName id="6C49FF96FFF4BA34FCF0F817FC00F80D" box="[813,909,1956,1980]" pageId="21" pageNumber="912">N.K. Ng</collectorName>
,
<date id="B502BC80FFF4BA34FC47F817FBBCF80D" box="[922,1073,1956,1980]" pageId="21" pageNumber="912" value="2000-06-15">
<collectingDate id="A5464568FFF4BA34FC47F817FBBCF80D" box="[922,1073,1956,1980]" pageId="21" pageNumber="912" value="2000-06-15">15 June 2000</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71D4901DFFF4BA34FBE3F817FA1DF86D" ID-GBIF-Occurrence="3763237550" collectingDate="2012-09-18" collectionCode="ZRC" collectorName="B. Y. Lee" country="Japan" location="Yenbaru" pageId="21" pageNumber="912" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFF4BA34FBE3F817FB04F80D" box="[1086,1161,1956,1980]" pageId="21" pageNumber="912" type="male">1 male</specimenCount>
(29.3 ×
<quantity id="064437A5FFF4BA34FB36F817FAC3F80D" box="[1259,1358,1956,1980]" metricMagnitude="-2" metricUnit="m" metricValue="2.68" pageId="21" pageNumber="912" unit="mm" value="26.8">26.8 mm</quantity>
) (
<collectionCode id="A7AD0285FFF4BA34FABAF817FA2DF80D" box="[1383,1440,1956,1980]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="912">ZRC</collectionCode>
),
<location id="C463CC9BFFF4BA34FCF0F877FC01F86D" LSID="urn:lsid:plazi:treatment:49152B56FFF4BA3AFC57FECEFF73FAA4:C463CC9BFFF4BA34FCF0F877FC01F86D" box="[813,908,1988,2012]" country="Japan" name="Yenbaru" pageId="21" pageNumber="912" stateProvince="Okinawa">Yenbaru</location>
,
<collectingRegion id="037854A2FFF4BA34FC44F877FC71F86D" box="[921,1020,1988,2012]" country="Japan" name="Okinawa" pageId="21" pageNumber="912">Okinawa</collectingRegion>
, coll.
<collectorName id="6C49FF96FFF4BA34FB9FF877FB24F86A" box="[1090,1193,1988,2011]" pageId="21" pageNumber="912">B.Y. Lee</collectorName>
,
<date id="B502BC80FFF4BA34FB6AF877FA1DF86D" box="[1207,1424,1988,2012]" pageId="21" pageNumber="912" value="2012-09-18">
<collectingDate id="A5464568FFF4BA34FB6AF877FA1DF86D" box="[1207,1424,1988,2012]" pageId="21" pageNumber="912" value="2012-09-18">18 September 2012</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71D4901DFFF4BA34FA43F877FC76F78D" ID-GBIF-Occurrence="3763237341" collectingDate="2006-11-17" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Japan" county="Zenda Forest" location="Ryukyu Islands" municipality="Kume Island" pageId="21" pageNumber="912" specimenCode="ZRC 2014.0265" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFF4BA34FA43F877FCEFF84D" pageId="21" pageNumber="912" type="male">1 male</specimenCount>
(37.5 ×
<quantity id="064437A5FFF4BA34FC14F857FBA2F84D" box="[969,1071,2020,2044]" metricMagnitude="-2" metricUnit="m" metricValue="3.5799999999999996" pageId="21" pageNumber="912" unit="mm" value="35.8">35.8 mm</quantity>
) (
<specimenCode id="911A323BFFF4BA34FB95F857FA8FF84D" box="[1096,1282,2020,2044]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="912">ZRC 2014.0265</specimenCode>
),
<collectingCounty id="2862E2CCFFF4BA34FAC5F857FA26F84A" box="[1304,1451,2020,2044]" pageId="21" pageNumber="912">Zenda Forest</collectingCounty>
Park,
<collectingMunicipality id="2167003AFFF4BA34FCB0F7B7FC76F7AD" box="[877,1019,2052,2076]" pageId="21" pageNumber="912">Kume Island</collectingMunicipality>
,
<location id="C463CC9BFFF4BA34FBDAF7B7FB3EF7AD" LSID="urn:lsid:plazi:treatment:49152B56FFF4BA3AFC57FECEFF73FAA4:C463CC9BFFF4BA34FBDAF7B7FB3EF7AD" box="[1031,1203,2052,2076]" country="Japan" county="Zenda Forest" municipality="Kume Island" name="Ryukyu Islands" pageId="21" pageNumber="912" stateProvince="Okinawa">Ryukyu Islands</location>
, coll.
<collectorName id="6C49FF96FFF4BA34FB2AF7B7FAEAF7AD" box="[1271,1383,2052,2076]" pageId="21" pageNumber="912">P.K.L. Ng</collectorName>
et al.,
<date id="B502BC80FFF4BA34FCF0F797FC76F78D" box="[813,1019,2084,2108]" pageId="21" pageNumber="912" value="2006-11-17">
<collectingDate id="A5464568FFF4BA34FCF0F797FC76F78D" box="[813,1019,2084,2108]" pageId="21" pageNumber="912" value="2006-11-17">17 November 2006</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="71D4901DFFF4BA37FBD9F797FF48F889" ID-GBIF-Occurrence="3763237483" collectingDate="2002-06-22" collectionCode="ZRC" collectorName="K. X. Lee" country="Taiwan" county="northeastern Taiwan" lastPageId="22" lastPageNumber="913" latitude="23.981667" location="Tahsi" longLatPrecision="21" longitude="121.610275" municipality="Ilan county" pageId="21" pageNumber="912" specimenCode="ZRC 2001.0033" specimenCount="1" specimenCount-female="1" specimenCount-juvenile="1" stateProvince="Hualien" typeStatus="paralectotype">
<collectingCountry id="B9ABDAD0FFF4BA34FBD9F797FBFDF78D" box="[1028,1136,2084,2108]" name="Taiwan" pageId="21" pageNumber="912">TAIWAN</collectingCountry>
<specimenCount id="D7BA51C9FFF4BA34FB55F797FB69F78D" box="[1160,1252,2084,2108]" pageId="21" pageNumber="912" type="female">1 female</specimenCount>
(
<specimenCode id="911A323BFFF4BA34FB2DF797FA2CF78D" box="[1264,1441,2084,2108]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="912">ZRC 2001.0033</specimenCode>
),
<collectingCounty id="2862E2CCFFF7BA37FF55F8B3FEE2F8A9" box="[136,367,1792,1816]" pageId="22" pageNumber="913">northeastern Taiwan</collectingCounty>
,
<collectingMunicipality id="2167003AFFF7BA37FEA3F8B3FD8DF8A9" box="[382,512,1792,1816]" pageId="22" pageNumber="913">Ilan county</collectingMunicipality>
,
<location id="C463CC9BFFF7BA37FDD2F8B3FDC3F8A9" LSID="urn:lsid:plazi:treatment:49152B56FFF4BA3AFC57FECEFF73FAA4:C463CC9BFFF7BA37FDD2F8B3FDC3F8A9" box="[527,590,1792,1816]" country="Taiwan" county="northeastern Taiwan" municipality="Ilan county" name="Tahsi" pageId="22" pageNumber="913">Tahsi</location>
, coll.
<collectorName id="6C49FF96FFF7BA37FD45F8B3FC8FF8A6" box="[664,770,1792,1815]" pageId="22" pageNumber="913">K.X. Lee</collectorName>
, 2000;
</materialsCitation>
<materialsCitation id="71D4901DFFF7BA37FF16F893FE02F829" ID-GBIF-Occurrence="3763237531" collectingDate="2002-06-22" collectionCode="ZRC" collectorName="K. X. Lee" country="Taiwan" county="northeastern Taiwan" latitude="23.981667" location="Tahsi" longLatPrecision="21" longitude="121.610275" municipality="Ilan county" pageId="22" pageNumber="913" specimenCode="ZRC 2001.0033" specimenCount="1" specimenCount-female="1" specimenCount-juvenile="1" stateProvince="Hualien" typeStatus="paralectotype">
1 ovigerous female,
<specimenCount id="D7BA51C9FFF7BA37FE7CF893FD87F889" box="[417,522,1824,1848]" pageId="22" pageNumber="913" type="juvenile">1 juvenile</specimenCount>
male,
<specimenCount id="D7BA51C9FFF7BA37FD8DF893FD34F889" box="[592,697,1824,1848]" pageId="22" pageNumber="913" type="juvenile">1 juvenile</specimenCount>
female (
<collectionCode id="A7AD0285FFF7BA37FF4DF8F3FF45F8E9" box="[144,200,1856,1880]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="22" pageNumber="913">ZRC</collectionCode>
),
<collectingRegion id="037854A2FFF7BA37FF0AF8F3FEFAF8E9" box="[215,375,1856,1880]" country="Taiwan" name="Hualien" pageId="22" pageNumber="913">Hualien county</collectingRegion>
,
<collectingRegion id="037854A2FFF7BA37FE5DF8F3FE5FF8E9" box="[384,466,1856,1880]" country="Taiwan" name="Hualien" pageId="22" pageNumber="913">Hualien</collectingRegion>
city, mouth of
<collectingMunicipality id="2167003AFFF7BA37FDB6F8F3FD3BF8E9" box="[619,694,1856,1880]" pageId="22" pageNumber="913">Meilun</collectingMunicipality>
stream,
<geoCoordinate id="A488FC87FFF7BA37FF55F8EDFE8BF8C6" box="[136,262,1886,1911]" degrees="23" direction="north" minutes="58" orientation="latitude" pageId="22" pageNumber="913" precision="15" seconds="54" value="23.981667">23°5854″N</geoCoordinate>
<geoCoordinate id="A488FC87FFF7BA37FED1F8EDFE1AF8C6" box="[268,407,1886,1911]" degrees="121" direction="east" minutes="36" orientation="longitude" pageId="22" pageNumber="913" precision="15" seconds="37" value="121.610275">121°3637″E</geoCoordinate>
,
<location id="C463CC9BFFF7BA37FE7DF8D3FDC9F8C9" LSID="urn:lsid:plazi:treatment:49152B56FFF4BA3AFC57FECEFF73FAA4:C463CC9BFFF7BA37FE7DF8D3FDC9F8C9" box="[416,580,1888,1912]" country="Taiwan" latitude="23.981667" longLatPrecision="21" longitude="121.610275" municipality="Meilun" name="eastern Taiwan" pageId="22" pageNumber="913" stateProvince="Hualien">eastern Taiwan</location>
, coll.
<collectorName id="6C49FF96FFF7BA37FD5CF8D3FD60F8C9" box="[641,749,1888,1912]" pageId="22" pageNumber="913">P.K.L. Ng</collectorName>
&amp;
<collectorName id="6C49FF96FFF7BA37FF55F833FF7FF829" box="[136,242,1920,1944]" pageId="22" pageNumber="913">H.-C. Liu</collectorName>
,
<date id="B502BC80FFF7BA37FF20F833FE02F829" box="[253,399,1920,1944]" pageId="22" pageNumber="913" value="2002-06-22">
<collectingDate id="A5464568FFF7BA37FF20F833FE02F829" box="[253,399,1920,1944]" pageId="22" pageNumber="913" value="2002-06-22">22 June 2002</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71D4901DFFF7BA37FE44F833FE47F869" ID-GBIF-Occurrence="3763237426" collectingDate="1999-09-14" collectionCode="ZRC" collectorName="H. - C. Liu &amp; C. D. Schubart" country="Taiwan" location="Rangkon river" pageId="22" pageNumber="913" specimenCode="ZRC 2000.1834" specimenCount="3" specimenCount-female="1" specimenCount-juvenile="1" specimenCount-male="1" stateProvince="Hualien" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFF7BA37FE44F833FE69F829" box="[409,484,1920,1944]" pageId="22" pageNumber="913" type="male">1 male</specimenCount>
,
<specimenCount id="D7BA51C9FFF7BA37FE32F833FDC3F829" box="[495,590,1920,1944]" pageId="22" pageNumber="913" type="female">1 female</specimenCount>
,
<specimenCount id="D7BA51C9FFF7BA37FD84F833FD4EF829" box="[601,707,1920,1944]" pageId="22" pageNumber="913" type="juvenile">1 juvenile</specimenCount>
(
<specimenCode id="911A323BFFF7BA37FD0CF833FF73F809" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="22" pageNumber="913">ZRC 2000.1834</specimenCode>
),
<location id="C463CC9BFFF7BA37FECFF813FE23F809" LSID="urn:lsid:plazi:treatment:49152B56FFF4BA3AFC57FECEFF73FAA4:C463CC9BFFF7BA37FECFF813FE23F809" box="[274,430,1952,1976]" country="Taiwan" name="Rangkon river" pageId="22" pageNumber="913" stateProvince="Hualien">Rangkon river</location>
mouth, coll.
<collectorName id="6C49FF96FFF7BA37FD99F813FD20F809" box="[580,685,1952,1976]" pageId="22" pageNumber="913">H.-C. Liu</collectorName>
&amp;
<collectorName id="6C49FF96FFF7BA37FD09F813FF67F869" pageId="22" pageNumber="913">C.D. Schubart</collectorName>
,
<date id="B502BC80FFF7BA37FF2BF873FE47F869" box="[246,458,1984,2008]" pageId="22" pageNumber="913" value="1999-09-14">
<collectingDate id="A5464568FFF7BA37FF2BF873FE47F869" box="[246,458,1984,2008]" pageId="22" pageNumber="913" value="1999-09-14">14 September 1999</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71D4901DFFF7BA37FE0AF873FB62F8A9" ID-GBIF-Occurrence="3763237344" collectionCode="ZRC" collectorName="Hengchun Peninsula &amp; P. K. L. Ng" country="Taiwan" location="Kenting National Park" municipality="Tung Kou" pageId="22" pageNumber="913" specimenCode="ZRC 1999.527" specimenCount="6" specimenCount-female="6" stateProvince="Pingtung" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFF7BA37FE0AF873FDA7F869" box="[471,554,1984,2008]" pageId="22" pageNumber="913" type="male">4 males</specimenCount>
(largest 23.5 ×
<quantity id="064437A5FFF7BA37FD05F873FF39F846" metricMagnitude="-2" metricUnit="m" metricValue="2.12" pageId="22" pageNumber="913" unit="mm" value="21.2">21.2 mm</quantity>
, smallest 29.0 ×
<quantity id="064437A5FFF7BA37FEA8F853FE56F849" box="[373,475,2016,2040]" metricMagnitude="-2" metricUnit="m" metricValue="2.58" pageId="22" pageNumber="913" unit="mm" value="25.8">25.8 mm</quantity>
),
<specimenCount id="D7BA51C9FFF7BA37FE2CF853FDD6F849" box="[497,603,2016,2040]" pageId="22" pageNumber="913" type="female">6 females</specimenCount>
(largest 23.9 ×
<quantity id="064437A5FFF7BA37FF55F7B3FF67F7A6" box="[136,234,2048,2072]" metricMagnitude="-2" metricUnit="m" metricValue="2.07" pageId="22" pageNumber="913" unit="mm" value="20.7">20.7 mm</quantity>
, smallest 28.0 ×
<quantity id="064437A5FFF7BA37FE7DF7B3FD8CF7A9" box="[416,513,2048,2072]" metricMagnitude="-2" metricUnit="m" metricValue="2.42" pageId="22" pageNumber="913" unit="mm" value="24.2">24.2 mm</quantity>
) (
<specimenCode id="911A323BFFF7BA37FDCAF7B3FD31F7A9" box="[535,700,2048,2072]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="22" pageNumber="913">ZRC 1999.527</specimenCode>
),
<collectingMunicipality id="2167003AFFF7BA37FD12F7B3FF3BF786" pageId="22" pageNumber="913">Tung Kou</collectingMunicipality>
stream,
<location id="C463CC9BFFF7BA37FEC7F793FD9AF789" LSID="urn:lsid:plazi:treatment:49152B56FFF4BA3AFC57FECEFF73FAA4:C463CC9BFFF7BA37FEC7F793FD9AF789" box="[282,535,2080,2104]" country="Taiwan" municipality="Tung Kou" name="Kenting National Park" pageId="22" pageNumber="913" stateProvince="Pingtung">Kenting National Park</location>
,
<collectingRegion id="037854A2FFF7BA37FDF8F793FD04F789" box="[549,649,2080,2104]" country="Taiwan" name="Pingtung" pageId="22" pageNumber="913">Pingtung</collectingRegion>
,
<collectorName id="6C49FF96FFF7BA37FD45F793FC20F8A9" box="[664,941,1792,2104]" pageId="22" pageNumber="913">Hengchun Peninsula</collectorName>
, coll.
<collectorName id="6C49FF96FFF7BA37FC29F8B3FBE8F8A9" box="[1012,1125,1792,1816]" pageId="22" pageNumber="913">P.K.L. Ng</collectorName>
et al., 2000
</materialsCitation>
;
<materialsCitation id="71D4901DFFF7BA37FB20F8B3FCF1F8C9" ID-GBIF-Occurrence="3763237498" collectingDate="2002-08-05" collectionCode="ZRC" collectorName="P. K. L. Ng &amp; C. - H. Wang" country="Taiwan" county="Kang Kou Hsi" location="southern Taiwan" municipality="Kentin Park" pageId="22" pageNumber="913" specimenCode="ZRC 2002.0477" specimenCount="6" specimenCount-female="5" specimenCount-male="1" stateProvince="Pingtung" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFF7BA37FB20F8B3FAC7F8A9" box="[1277,1354,1792,1816]" pageId="22" pageNumber="913" type="male">1 male</specimenCount>
,
<specimenCount id="D7BA51C9FFF7BA37FA85F8B3FA4FF8A9" box="[1368,1474,1792,1816]" pageId="22" pageNumber="913" type="female">5 females</specimenCount>
(
<specimenCode id="911A323BFFF7BA37FC97F893FB8DF889" box="[842,1024,1824,1848]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="22" pageNumber="913">ZRC 2002.0477</specimenCode>
),
<collectingCounty id="2862E2CCFFF7BA37FBC8F893FB34F889" box="[1045,1209,1824,1848]" pageId="22" pageNumber="913">Kang Kou Hsi</collectingCounty>
,
<collectingMunicipality id="2167003AFFF7BA37FB1BF893FAC0F889" box="[1222,1357,1824,1848]" pageId="22" pageNumber="913">Kentin Park</collectingMunicipality>
,
<collectingRegion id="037854A2FFF7BA37FA86F893FA33F889" box="[1371,1470,1824,1848]" country="Taiwan" name="Pingtung" pageId="22" pageNumber="913">Pingtung</collectingRegion>
,
<location id="C463CC9BFFF7BA37FC9EF8F3FC77F8E9" LSID="urn:lsid:plazi:treatment:49152B56FFF4BA3AFC57FECEFF73FAA4:C463CC9BFFF7BA37FC9EF8F3FC77F8E9" box="[835,1018,1856,1880]" country="Taiwan" county="Kang Kou Hsi" municipality="Kentin Park" name="southern Taiwan" pageId="22" pageNumber="913" stateProvince="Pingtung">southern Taiwan</location>
, coll.
<collectorName id="6C49FF96FFF7BA37FBE1F8F3FB26F8E9" box="[1084,1195,1856,1880]" pageId="22" pageNumber="913">P.K.L. Ng</collectorName>
&amp;
<collectorName id="6C49FF96FFF7BA37FB12F8F3FAD9F8E9" box="[1231,1364,1856,1880]" pageId="22" pageNumber="913">C.-H. Wang</collectorName>
,
<date id="B502BC80FFF7BA37FABDF8F3FCF1F8C9" pageId="22" pageNumber="913" value="2002-08-05">
<collectingDate id="A5464568FFF7BA37FABDF8F3FCF1F8C9" pageId="22" pageNumber="913" value="2002-08-05">5 August 2002</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71D4901DFFF7BA37FC56F8D3FBBAF809" ID-GBIF-Occurrence="3763237401" collectingDate="2000-11-07" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Taiwan" location="Chang-Wai" municipality="Hengchun" pageId="22" pageNumber="913" specimenCode="ZRC 2001.0034" specimenCount="2" specimenCount-male="2" stateProvince="Pingtung" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFF7BA37FC56F8D3FC6FF8C9" box="[907,994,1888,1912]" pageId="22" pageNumber="913" type="male">2 males</specimenCount>
(29.5 ×
<quantity id="064437A5FFF7BA37FB95F8D3FB20F8C6" box="[1096,1197,1888,1912]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" pageId="22" pageNumber="913" unit="mm" value="26.5">26.5 mm</quantity>
, 34.2 ×
<quantity id="064437A5FFF7BA37FAD2F8D3FAFEF8C9" box="[1295,1395,1888,1912]" metricMagnitude="-2" metricUnit="m" metricValue="3.07" pageId="22" pageNumber="913" unit="mm" value="30.7">30.7 mm</quantity>
) (
<specimenCode id="911A323BFFF7BA37FA51F8D3FC34F829" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="22" pageNumber="913">ZRC 2001.0034</specimenCode>
),
<collectingRegion id="037854A2FFF7BA37FC10F833FBBDF829" box="[973,1072,1920,1944]" country="Taiwan" name="Pingtung" pageId="22" pageNumber="913">Pingtung</collectingRegion>
,
<collectingMunicipality id="2167003AFFF7BA37FBE0F833FB20F829" box="[1085,1197,1920,1944]" pageId="22" pageNumber="913">Hengchun</collectingMunicipality>
,
<location id="C463CC9BFFF7BA37FB67F833FAB5F829" LSID="urn:lsid:plazi:treatment:49152B56FFF4BA3AFC57FECEFF73FAA4:C463CC9BFFF7BA37FB67F833FAB5F829" box="[1210,1336,1920,1944]" country="Taiwan" municipality="Hengchun" name="Chang-Wai" pageId="22" pageNumber="913" stateProvince="Pingtung">Chang-Wai</location>
, coll.
<collectorName id="6C49FF96FFF7BA37FAA1F833FCEBF809" pageId="22" pageNumber="913">P.K.L. Ng</collectorName>
,
<date id="B502BC80FFF7BA37FCAFF813FBBAF809" box="[882,1079,1952,1976]" pageId="22" pageNumber="913" value="2000-11-07">
<collectingDate id="A5464568FFF7BA37FCAFF813FBBAF809" box="[882,1079,1952,1976]" pageId="22" pageNumber="913" value="2000-11-07">7 November 2000</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71D4901DFFF7BA37FB9FF813FBE9F849" ID-GBIF-Occurrence="3763237357" collectingDate="2007-05-07" collectionCode="ZRC" collectorName="S. H. Tan" country="Taiwan" location="Taiwan" pageId="22" pageNumber="913" specimenCode="ZRC 2007.0208" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Pingtung" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFF7BA37FB9FF813FB00F809" box="[1090,1165,1952,1976]" pageId="22" pageNumber="913" type="male">1 male</specimenCount>
,
<specimenCount id="D7BA51C9FFF7BA37FB44F813FB7BF809" box="[1177,1270,1952,1976]" pageId="22" pageNumber="913" type="female">1 female</specimenCount>
(
<specimenCode id="911A323BFFF7BA37FAD9F813FA3AF809" box="[1284,1463,1952,1976]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="22" pageNumber="913">ZRC 2007.0208</specimenCode>
), purchased from aquarium trade, supposedly from
<location id="C463CC9BFFF7BA37FAB6F873FA33F869" LSID="urn:lsid:plazi:treatment:49152B56FFF4BA3AFC57FECEFF73FAA4:C463CC9BFFF7BA37FAB6F873FA33F869" box="[1387,1470,1984,2008]" country="Taiwan" name="Taiwan" pageId="22" pageNumber="913" stateProvince="Pingtung">Taiwan</location>
, coll.
<collectorName id="6C49FF96FFF7BA37FCAAF853FC55F846" box="[887,984,2016,2040]" pageId="22" pageNumber="913">S.H. Tan</collectorName>
,
<date id="B502BC80FFF7BA37FC3FF853FBE9F849" box="[994,1124,2016,2040]" pageId="22" pageNumber="913" value="2007-05-07">
<collectingDate id="A5464568FFF7BA37FC3FF853FBE9F849" box="[994,1124,2016,2040]" pageId="22" pageNumber="913" value="2007-05-07">7 May 2007</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71D4901DFFF7BA37FBB0F853FA32F789" ID-GBIF-Occurrence="3763237559" collectingDate="1999-09-15" collectorName="H. - C. Liu &amp; C. D. Schubart" country="Taiwan" location="southern Taiwan" municipality="Checheng-Pingtung" pageId="22" pageNumber="913" specimenCount="2" specimenCount-female="2" stateProvince="Pingtung" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFF7BA37FBB0F853FB5EF849" box="[1133,1235,2016,2040]" pageId="22" pageNumber="913" type="female">2 females</specimenCount>
(17.8 ×
<quantity id="064437A5FFF7BA37FAF7F853FA07F846" box="[1322,1418,2016,2040]" metricMagnitude="-2" metricUnit="m" metricValue="1.58" pageId="22" pageNumber="913" unit="mm" value="15.8">15.8 mm</quantity>
, 15.5 ×
<quantity id="064437A5FFF7BA37FC84F7B3FC31F7A9" box="[857,956,2048,2072]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="22" pageNumber="913" unit="mm" value="13.4">13.4 mm</quantity>
), rice field mouth,
<collectingMunicipality id="2167003AFFF7BA37FB58F7B3FAD1F7A9" box="[1157,1372,2048,2072]" pageId="22" pageNumber="913">Checheng-Pingtung</collectingMunicipality>
,
<location id="C463CC9BFFF7BA37FABBF7B3FC19F789" LSID="urn:lsid:plazi:treatment:49152B56FFF4BA3AFC57FECEFF73FAA4:C463CC9BFFF7BA37FABBF7B3FC19F789" country="Taiwan" municipality="Checheng-Pingtung" name="southern Taiwan" pageId="22" pageNumber="913" stateProvince="Pingtung">southern Taiwan</location>
, coll.
<collectorName id="6C49FF96FFF7BA37FC0DF793FBB8F789" box="[976,1077,2080,2104]" pageId="22" pageNumber="913">H.-C. Liu</collectorName>
&amp;
<collectorName id="6C49FF96FFF7BA37FB8EF793FB64F789" box="[1107,1257,2080,2104]" pageId="22" pageNumber="913">C.D. Schubart</collectorName>
,
<date id="B502BC80FFF7BA37FB2FF793FA32F789" box="[1266,1471,2080,2104]" pageId="22" pageNumber="913" value="1999-09-15">
<collectingDate id="A5464568FFF7BA37FB2FF793FA32F789" box="[1266,1471,2080,2104]" pageId="22" pageNumber="913" value="1999-09-15">15 September 1999</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="71D4901DFFF9BA39FF55FF0EFDF4FEA4" ID-GBIF-Occurrence="3763237330" collectingDate="2019-05-24" collectionCode="ZRC" collectorName="S. Cannici" country="Hong Kong" latitude="22.495722" location="Mai Po Nature Reserve" longLatPrecision="1" longitude="114.04714" pageId="24" pageNumber="915" specimenCode="ZRC 2019.0540" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paralectotype">
<collectingCountry id="B9ABDAD0FFF9BA39FF55FF0EFEA4FF64" box="[136,297,189,213]" name="Hong Kong" pageId="24" pageNumber="915">HONG KONG</collectingCountry>
<specimenCount id="D7BA51C9FFF9BA39FE98FF0EFE02FF64" box="[325,399,189,213]" pageId="24" pageNumber="915" type="male">1 male</specimenCount>
,
<specimenCount id="D7BA51C9FFF9BA39FE47FF0EFD8FFF64" box="[410,514,189,213]" pageId="24" pageNumber="915" type="female">2 females</specimenCount>
(
<specimenCode id="911A323BFFF9BA39FDCDFF0EFD4EFF64" box="[528,707,189,213]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="24" pageNumber="915">ZRC 2019.0540</specimenCode>
), back mangroves, ca.
<geoCoordinate id="A488FC87FFF9BA39FEF1FF6FFE32FF44" box="[300,447,220,245]" degrees="22" direction="north" minutes="29" orientation="latitude" pageId="24" pageNumber="915" precision="1" seconds="44.6" value="22.495722">22°2944.6″N</geoCoordinate>
<geoCoordinate id="A488FC87FFF9BA39FE19FF6FFDE9FF44" box="[452,612,220,245]" degrees="114" direction="east" minutes="02" orientation="longitude" pageId="24" pageNumber="915" precision="1" seconds="49.7" value="114.04714">114°0249.7″E</geoCoordinate>
,
<location id="C463CC9BFFF9BA39FDB0FF6EFF6CFEA4" LSID="urn:lsid:plazi:treatment:49152B56FFF4BA3AFC57FECEFF73FAA4:C463CC9BFFF9BA39FDB0FF6EFF6CFEA4" country="Hong Kong" latitude="22.495722" longLatPrecision="1" longitude="114.04714" name="Mai Po Nature Reserve" pageId="24" pageNumber="915">Mai Po Nature Reserve</location>
, coll.
<collectorName id="6C49FF96FFF9BA39FEF8FF4EFE1BFEA4" box="[293,406,253,277]" pageId="24" pageNumber="915">S. Cannici</collectorName>
et al.,
<date id="B502BC80FFF9BA39FE39FF4EFDF4FEA4" box="[484,633,253,278]" pageId="24" pageNumber="915" value="2019-05-24">
<collectingDate id="A5464568FFF9BA39FE39FF4EFDF4FEA4" box="[484,633,253,278]" pageId="24" pageNumber="915" value="2019-05-24">24 May 2019</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="95C3CAC8FFF7BA37FF5AF990FA37F978" ID-DOI="http://doi.org/10.5281/zenodo.5351315" ID-Zenodo-Dep="5351315" httpUri="https://zenodo.org/record/5351315/files/figure.png" pageId="22" pageNumber="913" startId="22.[135,170,1571,1593]" targetBox="[136,1477,191,1554]" targetPageId="22">
<paragraph id="C1039A40FFF7BA37FF5AF990FA37F978" blockId="22.[135,1474,1571,1737]" pageId="22" pageNumber="913">
Fig. 10. A, D, EI,
<taxonomicName id="06BCE1C3FFF7BA37FE98F990FDC1F989" baseAuthorityName="De Haan" baseAuthorityYear="1833" box="[325,588,1571,1592]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="913" phylum="Arthropoda" rank="species" species="haematocheir">
<emphasis id="F3C84652FFF7BA37FE98F990FDC1F989" box="[325,588,1571,1592]" italics="true" pageId="22" pageNumber="913">Chiromantes haematocheir</emphasis>
</taxonomicName>
, lectotype male (33.7 × 29.7 mm) (RMNH-D160), Japan; B, male (26.4 × 22.9 mm) (ZRC 1970.8.27.7), Sagami Bay, Japan; C,
<taxonomicName id="06BCE1C3FFF7BA37FE22F9F3FD29F9E5" baseAuthorityName="De Haan" baseAuthorityYear="1833" box="[511,676,1599,1621]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="913" phylum="Arthropoda" rank="species" species="haematocheir">
<emphasis id="F3C84652FFF7BA37FE22F9F3FD29F9E5" box="[511,676,1599,1621]" italics="true" pageId="22" pageNumber="913">C. haematocheir</emphasis>
</taxonomicName>
, male (lectotype of
<taxonomicName id="06BCE1C3FFF7BA37FCAFF98CFBCFF9E4" authorityName="Soh" authorityYear="1978" box="[882,1090,1599,1621]" class="Malacostraca" family="Sesarmidae" genus="Holometopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="913" phylum="Arthropoda" rank="species" species="serenei">
<emphasis id="F3C84652FFF7BA37FCAFF98CFBCFF9E4" box="[882,1090,1599,1621]" italics="true" pageId="22" pageNumber="913">Holometopus serenei</emphasis>
</taxonomicName>
) (18.4 × 16.6 mm) (NHM 1978.107), Hong Kong; JL,
<taxonomicName id="06BCE1C3FFF7BA37FEE6F9EEFE45F9C0" authorityName="Naruse &amp; Ng" authorityYear="2008" box="[315,456,1628,1650]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="913" phylum="Arthropoda" rank="species" species="ryukyuanus">
<emphasis id="F3C84652FFF7BA37FEE6F9EEFE45F9C0" box="[315,456,1628,1650]" italics="true" pageId="22" pageNumber="913">C. ryukyuanus</emphasis>
</taxonomicName>
, holotype male (33.1 × 29.6 mm) (RUMF-ZC-539), Okinawa Island, Japan. A, J, male pleon; B, male pleonal somite 6 and telson; male pleonal somites 46, telson; D, anterior thoracic sternites 14; E, left G1 (ventral view, denuded); F, left G1 (dorsal view, denuded); G, left distal part of G1 (ventral view, denuded); H, left distal part of G1 (dorsal view, denuded); I, L, left G2 (denuded); K, left G1. Scales: A, B, D = 5.0 mm; C, E, F, I = 2.0 mm; G, H = 1.0 mm. JL, after
<bibRefCitation id="A52DE7B1FFF7BA37FB7AF900FAE3F979" author="Naruse &amp; Ng" box="[1191,1390,1715,1736]" firstAuthor="Naruse" pageId="22" pageNumber="913" pagination="1 - 13" refId="ref73402" refString="Naruse T &amp; Ng PKL (2008) A new species of Chiromantes s. str. (Decapoda: Brachyura: Sesarmidae) from the Ryukyu Islands, Japan, with a note on the identity of Holometopus serenei Soh, 1978. Crustacean Research, 37: 1 - 13." type="journal article" year="2008">Naruse &amp; Ng (2008</bibRefCitation>
: fig. 8).
</paragraph>
</caption>
<caption id="95C3CAC8FFF6BA36FFACF82EFECBF7F3" ID-DOI="http://doi.org/10.5281/zenodo.5351319" ID-Zenodo-Dep="5351319" httpUri="https://zenodo.org/record/5351319/files/figure.png" pageId="23" pageNumber="914" startId="23.[113,148,1949,1971]" targetBox="[183,1384,190,1934]" targetPageId="23">
<paragraph id="C1039A40FFF6BA36FFACF82EFECBF7F3" blockId="23.[113,1452,1948,2114]" pageId="23" pageNumber="914">
Fig. 11. AG,
<taxonomicName id="06BCE1C3FFF6BA36FED8F82EFE34F803" authorityName=", De Man" authorityYear="1887" box="[261,441,1949,1970]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="914" phylum="Arthropoda" rank="species" species="dehaani">
<emphasis id="F3C84652FFF6BA36FED8F82EFE34F803" box="[261,441,1949,1970]" italics="true" pageId="23" pageNumber="914">Orisarma dehaani</emphasis>
</taxonomicName>
, lectotype male (39.5 × 35.7 mm) (RMNH-D157), Japan; HL,
<taxonomicName id="06BCE1C3FFF6BA36FB9EF82EFB4AF803" baseAuthorityName="De Man" baseAuthorityYear="1887" box="[1091,1223,1949,1970]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="914" phylum="Arthropoda" rank="species" species="neglectum">
<emphasis id="F3C84652FFF6BA36FB9EF82EFB4AF803" box="[1091,1223,1949,1970]" italics="true" pageId="23" pageNumber="914">O. neglectum</emphasis>
</taxonomicName>
, neotype male (35.6 × 31.8 mm) (ZRC 1998.310), Shanghai, China; MP,
<taxonomicName id="06BCE1C3FFF6BA36FDAFF809FD69F87E" baseAuthorityName="Komai &amp; Ng" baseAuthorityYear="2013" box="[626,740,1978,1999]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="914" phylum="Arthropoda" rank="species" species="magnum">
<emphasis id="F3C84652FFF6BA36FDAFF809FD69F87E" box="[626,740,1978,1999]" italics="true" pageId="23" pageNumber="914">O. magnum</emphasis>
</taxonomicName>
, holotype male (47.2 × 52.2 mm) (CBM-ZC 11452), Ogasawara Island, Japan. A, H, male pleon; B, anterior thoracic sternites 14; C, I, N, left G1 (ventral view, denuded); D, J, left G1 (dorsal view, denuded); E, K, O, left distal part of G1 (ventral view, denuded); F, L, P, left distal part of G1 (dorsal view, denuded); G, left G2 (denuded); M, male pleonal somite 6 and telson; MP, after
<bibRefCitation id="A52DE7B1FFF6BA36FDEFF7A3FD7AF794" author="Komai &amp; Ng" box="[562,759,2064,2086]" firstAuthor="Komai" pageId="23" pageNumber="914" pagination="539 - 551" refId="ref71033" refString="Komai T &amp; Ng PKL (2013) A new species of sesarmid crab of the genus Chiromantes (Crustacea: Decapoda: Brachyura) from the Ogasawara Islands, Japan. Zootaxa, 3681 (5): 539 - 551." type="journal article" year="2013">Komai &amp; Ng (2013</bibRefCitation>
: fig. 5A, C, D, E). Scales: A, B, H = 5.0 mm; C, D, G, I, J = 2.0 mm; E, F, K, L = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="89A6C9CBFFF9BA39FF55FE8EFC8AFBA4" pageId="24" pageNumber="915" type="diagnosis">
<paragraph id="C1039A40FFF9BA39FF55FE8EFDEBFDC4" blockId="24.[136,775,317,629]" pageId="24" pageNumber="915">
<emphasis id="F3C84652FFF9BA39FF55FE8EFF78FEE4" bold="true" box="[136,245,317,341]" pageId="24" pageNumber="915">Diagnosis</emphasis>
. Dorsal surface of carapace relatively less convex, surfaces usually slightly more rugose, with scattered stiff setae on anterior and branchial regions; frontal lobes separated by deep median concavity; epibranchial tooth distinct even in small specimens, always separated by deep notch; posterolateral margins subparallel; granules on outer surface of palm (especially those below longitudinal ridge) rounded, prominent; merus of ambulatory legs relatively longer; chitinous distal part of G1 short, tip truncate. In life, carapace and chelae red, sometimes yellow.
</paragraph>
<paragraph id="C1039A40FFF9BA39FF55FD2EFC8AFBA4" blockId="24.[136,775,669,1045]" pageId="24" pageNumber="915">
<emphasis id="F3C84652FFF9BA39FF55FD2EFF5AFD04" bold="true" box="[136,215,669,693]" pageId="24" pageNumber="915">Colour</emphasis>
. In life, the dorsal surfaces of the carapace, chelipeds, and ambulatory legs are bright red with the fingers white (
<figureCitation id="598786C5FFF9BA39FF4DFD6EFF62FD44" box="[144,239,733,757]" captionStart="Fig" captionStartId="86.[136,171,1434,1456]" captionTargetBox="[136,1474,189,1419]" captionTargetId="figure-7@86.[136,1474,189,1419]" captionTargetPageId="86" captionText="Fig. 54. Colour in life. A, Orisarma intermedium, Pingtung, Taiwan, specimen not collected [photograph: PKL Ng]; B, O. intermedium, female (37.5 × 35.8 mm) (ZRC 2014.0265), Kumejima Island, Japan [photograph: PKL Ng]; C, O. intermedium, male (23.2 × 20.0 mm) (ZRC 2019.0540), Hong Kong [photograph: Paul Ng]; D, O. sinense, Shanghai, China, specimen not collected [photograph: Liu Wenliang]." figureDoi="http://doi.org/10.5281/zenodo.5351395" httpUri="https://zenodo.org/record/5351395/files/figure.png" pageId="24" pageNumber="915">Fig. 54A</figureCitation>
) (see also
<bibRefCitation id="A52DE7B1FFF9BA39FE81FD6EFE7CFD44" author="Miyake" box="[348,497,733,757]" firstAuthor="Miyake" pageId="24" pageNumber="915" refId="ref72931" refString="Miyake S (1983) Japanese Crustacean Decapods and Stomatopods in Color, vol. II. Brachyura (Crabs). First Edition. Hoikusha, Osaka, viii + 277 pp., 64 pls., unnumbered figs." type="book" year="1983">Miyake, 1983</bibRefCitation>
;
<bibRefCitation id="A52DE7B1FFF9BA39FE27FD6DFDEBFD44" author="Lee" box="[506,614,733,757]" firstAuthor="Lee" pageId="24" pageNumber="915" refId="ref71440" refString="Lee J-H (2008) A Field Guide to Crabs in Taiwan. Bigtrees Company, Taipei, 174 pp." type="book" year="2008">Lee, 2008</bibRefCitation>
;
<bibRefCitation id="A52DE7B1FFF9BA39FDB2FD6EFD55FD44" author="Liu" box="[623,728,733,757]" firstAuthor="Liu" pageId="24" pageNumber="915" refId="ref71856" refString="Liu H-C (2009) Diversity, population biology and managements of land crabs in the Kenting National Park. The National Kenting Headquarters, Kenting, 77 pp." type="book" year="2009">Liu, 2009</bibRefCitation>
;
<bibRefCitation id="A52DE7B1FFF9BA39FD3FFD6EFEA5FCA4" author="Liu &amp; Wang" firstAuthor="Liu" pageId="24" pageNumber="915" refId="ref71886" refString="Liu H-C &amp; Wang C-H (2010) Taiwan Coastal Wetland Crabs. Wild Bird Society of Taipei, Taipei, 79 pp." type="book" year="2010">Liu &amp; Wang, 2010</bibRefCitation>
;
<bibRefCitation id="A52DE7B1FFF9BA39FEEFFD4EFD1AFCA4" author="Rademacher &amp; Mengedoht" box="[306,663,765,789]" firstAuthor="Rademacher" pageId="24" pageNumber="915" refId="ref74843" refString="Rademacher M &amp; Mengedoht O (2011) Krabben-Fibel. Die schonsten Krabben fur das Aquaterrarium. Dahne Verlag, Ettlingen, Germany, 95 pp." type="book" year="2011">Rademacher &amp; Mengedoht, 2011</bibRefCitation>
), although some orangish-red specimens are sometimes observed in southern
<collectingCountry id="B9ABDAD0FFF9BA39FF34FC8EFEB1FCE4" box="[233,316,829,853]" name="Taiwan" pageId="24" pageNumber="915">Taiwan</collectingCountry>
. There is an interesting large male collected from the Ryukyus which is bright yellow (
<figureCitation id="598786C5FFF9BA39FD9BFCEEFD2BFCC4" box="[582,678,861,885]" captionStart="Fig" captionStartId="86.[136,171,1434,1456]" captionTargetBox="[136,1474,189,1419]" captionTargetId="figure-7@86.[136,1474,189,1419]" captionTargetPageId="86" captionText="Fig. 54. Colour in life. A, Orisarma intermedium, Pingtung, Taiwan, specimen not collected [photograph: PKL Ng]; B, O. intermedium, female (37.5 × 35.8 mm) (ZRC 2014.0265), Kumejima Island, Japan [photograph: PKL Ng]; C, O. intermedium, male (23.2 × 20.0 mm) (ZRC 2019.0540), Hong Kong [photograph: Paul Ng]; D, O. sinense, Shanghai, China, specimen not collected [photograph: Liu Wenliang]." figureDoi="http://doi.org/10.5281/zenodo.5351395" httpUri="https://zenodo.org/record/5351395/files/figure.png" pageId="24" pageNumber="915">Fig. 54B</figureCitation>
), but this specimen was obtained over a kilometre from the sea on a muddy patch up on a hill, and its colour may be a result of fading. Its morphology is identical to more typically coloured specimens. Smaller specimens and some females may have a dark tranverse band across their frontal margin (
<figureCitation id="598786C5FFF9BA39FD44FC4EFD76FBA4" box="[665,763,1021,1045]" captionStart="Fig" captionStartId="86.[136,171,1434,1456]" captionTargetBox="[136,1474,189,1419]" captionTargetId="figure-7@86.[136,1474,189,1419]" captionTargetPageId="86" captionText="Fig. 54. Colour in life. A, Orisarma intermedium, Pingtung, Taiwan, specimen not collected [photograph: PKL Ng]; B, O. intermedium, female (37.5 × 35.8 mm) (ZRC 2014.0265), Kumejima Island, Japan [photograph: PKL Ng]; C, O. intermedium, male (23.2 × 20.0 mm) (ZRC 2019.0540), Hong Kong [photograph: Paul Ng]; D, O. sinense, Shanghai, China, specimen not collected [photograph: Liu Wenliang]." figureDoi="http://doi.org/10.5281/zenodo.5351395" httpUri="https://zenodo.org/record/5351395/files/figure.png" pageId="24" pageNumber="915">Fig. 54C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFF9BA39FF55FB8EFC7FF824" pageId="24" pageNumber="915" type="discussion">
<paragraph id="C1039A40FFF9BA39FF55FB8EFEABF904" blockId="24.[136,776,1085,1717]" pageId="24" pageNumber="915">
<emphasis id="F3C84652FFF9BA39FF55FB8EFF7DFBE4" bold="true" box="[136,240,1085,1109]" pageId="24" pageNumber="915">Remarks</emphasis>
.
<taxonomicName id="06BCE1C3FFF9BA39FF21FB8DFE7DFBE4" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[252,496,1085,1109]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFF9BA39FF21FB8DFE7DFBE4" box="[252,496,1085,1109]" italics="true" pageId="24" pageNumber="915">Orisarma intermedium</emphasis>
</taxonomicName>
is very close to
<taxonomicName id="06BCE1C3FFF9BA39FD44FB8DFC8AFBE4" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[665,775,1086,1109]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="sinense">
<emphasis id="F3C84652FFF9BA39FD44FB8DFC8AFBE4" box="[665,775,1086,1109]" italics="true" pageId="24" pageNumber="915">O. sinense</emphasis>
</taxonomicName>
in general carapace form and smaller specimens are not easy to separate. Many of the old records of these two species in the literature (especially those on mainland
<collectingCountry id="B9ABDAD0FFF9BA39FD8AFB2EFD15FB04" box="[599,664,1181,1205]" name="China" pageId="24" pageNumber="915">China</collectingCountry>
) will need to be rechecked. In an unpublished key, which was the basis of the more recent identifications of
<bibRefCitation id="A52DE7B1FFF9BA39FDCAFB6EFD3CFB44" author="Serene" box="[535,689,1245,1269]" firstAuthor="Serene" pageId="24" pageNumber="915" pagination="33 - 112" refId="ref76415" refString="Serene R (1968) The Brachyura of the Indo Pacific Region. In: Prodromus for a check list of the nonplanctonic marine fauna of South East Asia. Singapore National Academy of Science, Special Publication No. 1: 33 - 112." type="book chapter" year="1968">Serène (1968)</bibRefCitation>
,
<bibRefCitation id="A52DE7B1FFF9BA39FD63FB6EFE93FAA4" author="Serene &amp; Soh" firstAuthor="Serene" pageId="24" pageNumber="915" pagination="387 - 416" refId="ref76461" refString="Serene R &amp; Soh CL (1970) New IndoPacific genera allied to Sesarma Say 1877 (Brachyura, Decapoda, Crustacea). Treubia, 27: 387 - 416." type="journal article" year="1970">Serène &amp; Soh (1970)</bibRefCitation>
, and
<bibRefCitation id="A52DE7B1FFF9BA39FE8AFB4EFE5CFAA4" author="Soh" box="[343,465,1277,1301]" firstAuthor="Soh" pageId="24" pageNumber="915" pagination="9 - 22" refId="ref77463" refString="Soh CL (1978) On a collection of sesarmine crabs (Decapoda, Brachyura, Grapsidae) from Hong Kong. Memoirs of the Hong Kong Natural History Society, 13: 9 - 22." type="journal article" year="1978">Soh (1978)</bibRefCitation>
, the primary characters were whether the epibranchial tooth is clearly demarcated (present in
<taxonomicName id="06BCE1C3FFF9BA39FF75FA8DFED8FAE4" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[168,341,1341,1365]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFF9BA39FF75FA8DFED8FAE4" box="[168,341,1341,1365]" italics="true" pageId="24" pageNumber="915">O. intermedium</emphasis>
</taxonomicName>
versus absent in
<taxonomicName id="06BCE1C3FFF9BA39FDFDFA8DFD18FAE4" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[544,661,1342,1365]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="sinense">
<emphasis id="F3C84652FFF9BA39FDFDFA8DFD18FAE4" box="[544,661,1342,1365]" italics="true" pageId="24" pageNumber="915">O. sinense</emphasis>
</taxonomicName>
), whether the outer surface of the chela was strongly granulated, and the presence of a low longitudinal median ridge (strongly granulated with a ridge in
<taxonomicName id="06BCE1C3FFF9BA39FE4CFA2DFDB9FA04" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[401,564,1437,1461]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFF9BA39FE4CFA2DFDB9FA04" box="[401,564,1437,1461]" italics="true" pageId="24" pageNumber="915">O. intermedium</emphasis>
</taxonomicName>
versus low granules and no ridge in
<taxonomicName id="06BCE1C3FFF9BA39FEE7FA0DFE24FA64" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[314,425,1470,1493]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="sinense">
<emphasis id="F3C84652FFF9BA39FEE7FA0DFE24FA64" box="[314,425,1470,1493]" italics="true" pageId="24" pageNumber="915">O. sinense</emphasis>
</taxonomicName>
), the relative proportions of the fingers of the chela (dactylar finger longer than height of palm in
<taxonomicName id="06BCE1C3FFF9BA39FF3CFA4DFE04F9A4" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[225,393,1533,1557]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFF9BA39FF3CFA4DFE04F9A4" box="[225,393,1533,1557]" italics="true" pageId="24" pageNumber="915">O. intermedium</emphasis>
</taxonomicName>
versus dactylar finger shorter in
<taxonomicName id="06BCE1C3FFF9BA39FD30FA4DFF5AF984" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="sinense">
<emphasis id="F3C84652FFF9BA39FD30FA4DFF5AF984" italics="true" pageId="24" pageNumber="915">O. sinense</emphasis>
</taxonomicName>
) and ambulatory merus (merus of third ambulatory leg 2.5 times longer than broad in
<taxonomicName id="06BCE1C3FFF9BA39FDD3F98DFD34F9E4" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[526,697,1597,1621]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFF9BA39FDD3F98DFD34F9E4" box="[526,697,1597,1621]" italics="true" pageId="24" pageNumber="915">O. intermedium</emphasis>
</taxonomicName>
versus 3 times in
<taxonomicName id="06BCE1C3FFF9BA39FEDAF9EDFEF0F9C4" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[263,381,1630,1653]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="sinense">
<emphasis id="F3C84652FFF9BA39FEDAF9EDFEF0F9C4" box="[263,381,1630,1653]" italics="true" pageId="24" pageNumber="915">O. sinense</emphasis>
</taxonomicName>
), and proportions of the chitinous distal part of G1 (shorter in
<taxonomicName id="06BCE1C3FFF9BA39FE1FF9CDFDE1F924" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[450,620,1661,1685]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFF9BA39FE1FF9CDFDE1F924" box="[450,620,1661,1685]" italics="true" pageId="24" pageNumber="915">O. intermedium</emphasis>
</taxonomicName>
versus longer in
<taxonomicName id="06BCE1C3FFF9BA39FF78F92DFE9AF904" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[165,279,1694,1717]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="sinense">
<emphasis id="F3C84652FFF9BA39FF78F92DFE9AF904" box="[165,279,1694,1717]" italics="true" pageId="24" pageNumber="915">O. sinense</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="C1039A40FFF9BA39FF55F96EFB90FC44" blockId="24.[136,776,1757,2101]" lastBlockId="24.[835,1475,189,1013]" pageId="24" pageNumber="915">
Examination of the present material reveals that smaller specimens of
<taxonomicName id="06BCE1C3FFF9BA39FEF5F94DFE13F8A4" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[296,414,1790,1813]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="sinense">
<emphasis id="F3C84652FFF9BA39FEF5F94DFE13F8A4" box="[296,414,1790,1813]" italics="true" pageId="24" pageNumber="915">O. sinense</emphasis>
</taxonomicName>
do have the epibranchial tooth relatively low and while it may appear to be absent and/or almost confluent with the rest of the margin, it is nevertheless always visible (as in the case of small specimen from
<collectingCountry id="B9ABDAD0FFF9BA39FD16F8EDFF4BF827" name="Hong Kong" pageId="24" pageNumber="915">Hong Kong</collectingCountry>
, ZRC 1975.6.30.10). In medium-sized specimens, the epibranchial tooth is distinct but demarcated from the rest of the margin by a weak notch (
<figureCitation id="598786C5FFF9BA39FE3DF80EFDA6F864" box="[480,555,1981,2005]" captionStart="Fig" captionStartId="8.[136,171,1926,1948]" captionTargetBox="[136,1476,212,1910]" captionTargetId="figure-7@8.[136,1476,212,1910]" captionTargetPageId="8" captionText="Fig. 2. Overall habitus. A, Orisarma intermedium, lectotype male (23.0 × 19.9 mm) (RMNH-D165), Japan; B, O. intermedium, male (27.1 × 25.2 mm) (ZRC 1970.2.23.6), Japan; C, O. intermedium, male (27.0 × 25.0 mm) (ZRC 2013.0140), Kyushu, Japan; D, O. intermedium, male (29.5 × 26.5 mm) (ZRC 2001.0034), Pingtung, Taiwan; E, O. intermedium, male (34.2 × 30.7 mm) (ZRC 2001.0034), Pingtung, Taiwan; F, O. intermedium, male (37.5 × 35.8 mm) (ZRC 2014.0265), Kumejima Island, Japan; G, O. sinense, lectotype male (19.0 × 16.6 mm) (MNHN-BP3635a), China; H, O. sinense, paralectotype female (18.8 × 16.9 mm) (MNHN-BP3635b), China; I, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China; J, Orisarma sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China." figureDoi="http://doi.org/10.5281/zenodo.5351299" httpUri="https://zenodo.org/record/5351299/files/figure.png" pageId="24" pageNumber="915">Fig. 2I</figureCitation>
). Only in the largest specimen is the epibranchial tooth really prominent, and with a deeper notch separating it from the rest of the margin (
<figureCitation id="598786C5FFF9BA39FF4DF7AEFF64F784" box="[144,233,2077,2101]" captionStart="Fig" captionStartId="8.[136,171,1926,1948]" captionTargetBox="[136,1476,212,1910]" captionTargetId="figure-7@8.[136,1476,212,1910]" captionTargetPageId="8" captionText="Fig. 2. Overall habitus. A, Orisarma intermedium, lectotype male (23.0 × 19.9 mm) (RMNH-D165), Japan; B, O. intermedium, male (27.1 × 25.2 mm) (ZRC 1970.2.23.6), Japan; C, O. intermedium, male (27.0 × 25.0 mm) (ZRC 2013.0140), Kyushu, Japan; D, O. intermedium, male (29.5 × 26.5 mm) (ZRC 2001.0034), Pingtung, Taiwan; E, O. intermedium, male (34.2 × 30.7 mm) (ZRC 2001.0034), Pingtung, Taiwan; F, O. intermedium, male (37.5 × 35.8 mm) (ZRC 2014.0265), Kumejima Island, Japan; G, O. sinense, lectotype male (19.0 × 16.6 mm) (MNHN-BP3635a), China; H, O. sinense, paralectotype female (18.8 × 16.9 mm) (MNHN-BP3635b), China; I, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China; J, Orisarma sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China." figureDoi="http://doi.org/10.5281/zenodo.5351299" httpUri="https://zenodo.org/record/5351299/files/figure.png" pageId="24" pageNumber="915">Figs. 2J</figureCitation>
,
<figureCitation id="598786C5FFF9BA39FF2BF7AEFE9DF784" box="[246,272,2077,2101]" captionStart="Fig" captionStartId="12.[135,170,1954,1976]" captionTargetBox="[136,1476,241,1939]" captionTargetId="figure-7@12.[136,1476,241,1939]" captionTargetPageId="12" captionText="Fig. 4. A, F, Orisarma dehaani, male (36.4 × 34.4 mm) (ZRC 2011.1027), Kumejima Island, Japan; B, D, O. dehaani, male (27.6 × 29.6 mm) (ZRC 2002.0223), Hong Kong; C, E, G, O. neglectum, male (31.7 × 28.0 mm) (ZRC 1998.309), Shanghai, China; H, J, O. intermedium, male (29.5 × 26.5 mm) (ZRC 2001.0034), Pingtung, Taiwan; I, K, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China. AC, dorsal view of carapace; D, E, H, I, frontal and anterolateral margins; F, G, J, K, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.5351303" httpUri="https://zenodo.org/record/5351303/files/figure.png" pageId="24" pageNumber="915">4I</figureCitation>
). In
<taxonomicName id="06BCE1C3FFF9BA39FE98F7ADFE62F784" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[325,495,2077,2101]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFF9BA39FE98F7ADFE62F784" box="[325,495,2077,2101]" italics="true" pageId="24" pageNumber="915">O. intermedium</emphasis>
</taxonomicName>
, the epibranchial tooth is distinct even in small specimens and always separated by a deep notch (e.g.,
<figureCitation id="598786C5FFF9BA39FBC8FF6EFB1FFF44" box="[1045,1170,221,245]" captionStart="Fig" captionStartId="8.[136,171,1926,1948]" captionTargetBox="[136,1476,212,1910]" captionTargetId="figure-7@8.[136,1476,212,1910]" captionTargetPageId="8" captionText="Fig. 2. Overall habitus. A, Orisarma intermedium, lectotype male (23.0 × 19.9 mm) (RMNH-D165), Japan; B, O. intermedium, male (27.1 × 25.2 mm) (ZRC 1970.2.23.6), Japan; C, O. intermedium, male (27.0 × 25.0 mm) (ZRC 2013.0140), Kyushu, Japan; D, O. intermedium, male (29.5 × 26.5 mm) (ZRC 2001.0034), Pingtung, Taiwan; E, O. intermedium, male (34.2 × 30.7 mm) (ZRC 2001.0034), Pingtung, Taiwan; F, O. intermedium, male (37.5 × 35.8 mm) (ZRC 2014.0265), Kumejima Island, Japan; G, O. sinense, lectotype male (19.0 × 16.6 mm) (MNHN-BP3635a), China; H, O. sinense, paralectotype female (18.8 × 16.9 mm) (MNHN-BP3635b), China; I, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China; J, Orisarma sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China." figureDoi="http://doi.org/10.5281/zenodo.5351299" httpUri="https://zenodo.org/record/5351299/files/figure.png" pageId="24" pageNumber="915">Figs. 2AF</figureCitation>
,
<figureCitation id="598786C5FFF9BA39FB43FF6EFB4EFF44" box="[1182,1219,221,245]" captionStart="Fig" captionStartId="12.[135,170,1954,1976]" captionTargetBox="[136,1476,241,1939]" captionTargetId="figure-7@12.[136,1476,241,1939]" captionTargetPageId="12" captionText="Fig. 4. A, F, Orisarma dehaani, male (36.4 × 34.4 mm) (ZRC 2011.1027), Kumejima Island, Japan; B, D, O. dehaani, male (27.6 × 29.6 mm) (ZRC 2002.0223), Hong Kong; C, E, G, O. neglectum, male (31.7 × 28.0 mm) (ZRC 1998.309), Shanghai, China; H, J, O. intermedium, male (29.5 × 26.5 mm) (ZRC 2001.0034), Pingtung, Taiwan; I, K, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China. AC, dorsal view of carapace; D, E, H, I, frontal and anterolateral margins; F, G, J, K, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.5351303" httpUri="https://zenodo.org/record/5351303/files/figure.png" pageId="24" pageNumber="915">4H</figureCitation>
).
<bibRefCitation id="A52DE7B1FFF9BA39FB0AFF6EFADDFF44" author="Soh" box="[1239,1360,221,245]" firstAuthor="Soh" pageId="24" pageNumber="915" pagination="9 - 22" refId="ref77463" refString="Soh CL (1978) On a collection of sesarmine crabs (Decapoda, Brachyura, Grapsidae) from Hong Kong. Memoirs of the Hong Kong Natural History Society, 13: 9 - 22." type="journal article" year="1978">Soh (1978)</bibRefCitation>
suggested that the cheliped of
<taxonomicName id="06BCE1C3FFF9BA39FBC6FF4DFB06FEA4" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[1051,1163,254,277]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="sinense">
<emphasis id="F3C84652FFF9BA39FBC6FF4DFB06FEA4" box="[1051,1163,254,277]" italics="true" pageId="24" pageNumber="915">O. sinense</emphasis>
</taxonomicName>
has relatively shorter fingers but this is not a useful character and was probably due to the small size of the specimens he examined. Larger specimens of
<taxonomicName id="06BCE1C3FFF9BA39FC0AFEEDFBCAFEC4" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[983,1095,350,373]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="sinense">
<emphasis id="F3C84652FFF9BA39FC0AFEEDFBCAFEC4" box="[983,1095,350,373]" italics="true" pageId="24" pageNumber="915">O. sinense</emphasis>
</taxonomicName>
(
<figureCitation id="598786C5FFF9BA39FB8BFEEEFB45FEC4" box="[1110,1224,349,373]" captionStart="Fig" captionStartId="16.[135,170,1734,1756]" captionTargetBox="[136,1476,189,1718]" captionTargetId="figure-7@16.[136,1476,189,1718]" captionTargetPageId="16" captionText="Fig. 6. Chela. A, Orisarma intermedium, lectotype male (23.0 × 19.9 mm) (RMNH-D165), Japan; B, O. intermedium, male (27.1 × 25.2 mm) (ZRC 1970.2.23.6), Japan; C, Orisarma intermedium, male (34.7 × 31.6 mm) (ZRC 2001.0034), Pingtung, Taiwan; D, O. intermedium, male (37.5 × 35.8 mm) (ZRC 2014.0265), Kumejima Island, Japan; E, O. sinense, lectotype male (19.0 × 16.6 mm) (MNHN-BP3635a), China; F, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China; G, H, O. sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China. AG, outer view; H, inner view." figureDoi="http://doi.org/10.5281/zenodo.5351307" httpUri="https://zenodo.org/record/5351307/files/figure.png" pageId="24" pageNumber="915">Fig. 6EH</figureCitation>
) have the same kind of fingers as those of
<taxonomicName id="06BCE1C3FFF9BA39FBCEFECDFB31FE24" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[1043,1212,381,405]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFF9BA39FBCEFECDFB31FE24" box="[1043,1212,381,405]" italics="true" pageId="24" pageNumber="915">O. intermedium</emphasis>
</taxonomicName>
(
<figureCitation id="598786C5FFF9BA39FB11FECEFAC8FE24" box="[1228,1349,381,405]" captionStart="Fig" captionStartId="16.[135,170,1734,1756]" captionTargetBox="[136,1476,189,1718]" captionTargetId="figure-7@16.[136,1476,189,1718]" captionTargetPageId="16" captionText="Fig. 6. Chela. A, Orisarma intermedium, lectotype male (23.0 × 19.9 mm) (RMNH-D165), Japan; B, O. intermedium, male (27.1 × 25.2 mm) (ZRC 1970.2.23.6), Japan; C, Orisarma intermedium, male (34.7 × 31.6 mm) (ZRC 2001.0034), Pingtung, Taiwan; D, O. intermedium, male (37.5 × 35.8 mm) (ZRC 2014.0265), Kumejima Island, Japan; E, O. sinense, lectotype male (19.0 × 16.6 mm) (MNHN-BP3635a), China; F, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China; G, H, O. sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China. AG, outer view; H, inner view." figureDoi="http://doi.org/10.5281/zenodo.5351307" httpUri="https://zenodo.org/record/5351307/files/figure.png" pageId="24" pageNumber="915">Fig. 6AD</figureCitation>
). The outer surface of the adult male chela of the two species is slightly different, but the low longitudinal median ridge is present on both species (
<figureCitation id="598786C5FFF9BA39FC20FE6EFBFBFE44" box="[1021,1142,477,501]" captionStart="Fig" captionStartId="16.[135,170,1734,1756]" captionTargetBox="[136,1476,189,1718]" captionTargetId="figure-7@16.[136,1476,189,1718]" captionTargetPageId="16" captionText="Fig. 6. Chela. A, Orisarma intermedium, lectotype male (23.0 × 19.9 mm) (RMNH-D165), Japan; B, O. intermedium, male (27.1 × 25.2 mm) (ZRC 1970.2.23.6), Japan; C, Orisarma intermedium, male (34.7 × 31.6 mm) (ZRC 2001.0034), Pingtung, Taiwan; D, O. intermedium, male (37.5 × 35.8 mm) (ZRC 2014.0265), Kumejima Island, Japan; E, O. sinense, lectotype male (19.0 × 16.6 mm) (MNHN-BP3635a), China; F, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China; G, H, O. sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China. AG, outer view; H, inner view." figureDoi="http://doi.org/10.5281/zenodo.5351307" httpUri="https://zenodo.org/record/5351307/files/figure.png" pageId="24" pageNumber="915">Fig. 6AG</figureCitation>
). In adult
<taxonomicName id="06BCE1C3FFF9BA39FB35FE6DFA1FFE44" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[1256,1426,477,501]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFF9BA39FB35FE6DFA1FFE44" box="[1256,1426,477,501]" italics="true" pageId="24" pageNumber="915">O. intermedium</emphasis>
</taxonomicName>
, the granules on the outer surface of the palm, especially those below the longitudinal ridge, are rounded and prominent (
<figureCitation id="598786C5FFF9BA39FC91FD8EFC47FDE4" box="[844,970,573,597]" captionStart="Fig" captionStartId="16.[135,170,1734,1756]" captionTargetBox="[136,1476,189,1718]" captionTargetId="figure-7@16.[136,1476,189,1718]" captionTargetPageId="16" captionText="Fig. 6. Chela. A, Orisarma intermedium, lectotype male (23.0 × 19.9 mm) (RMNH-D165), Japan; B, O. intermedium, male (27.1 × 25.2 mm) (ZRC 1970.2.23.6), Japan; C, Orisarma intermedium, male (34.7 × 31.6 mm) (ZRC 2001.0034), Pingtung, Taiwan; D, O. intermedium, male (37.5 × 35.8 mm) (ZRC 2014.0265), Kumejima Island, Japan; E, O. sinense, lectotype male (19.0 × 16.6 mm) (MNHN-BP3635a), China; F, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China; G, H, O. sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China. AG, outer view; H, inner view." figureDoi="http://doi.org/10.5281/zenodo.5351307" httpUri="https://zenodo.org/record/5351307/files/figure.png" pageId="24" pageNumber="915">Fig. 6AD</figureCitation>
). In the case of
<taxonomicName id="06BCE1C3FFF9BA39FB5AFD8DFB71FDE4" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[1159,1276,574,597]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="sinense">
<emphasis id="F3C84652FFF9BA39FB5AFD8DFB71FDE4" box="[1159,1276,574,597]" italics="true" pageId="24" pageNumber="915">O. sinense</emphasis>
</taxonomicName>
, the granules are lower and more flattened (
<figureCitation id="598786C5FFF9BA39FBB5FDEEFB6DFDC4" box="[1128,1248,605,629]" captionStart="Fig" captionStartId="16.[135,170,1734,1756]" captionTargetBox="[136,1476,189,1718]" captionTargetId="figure-7@16.[136,1476,189,1718]" captionTargetPageId="16" captionText="Fig. 6. Chela. A, Orisarma intermedium, lectotype male (23.0 × 19.9 mm) (RMNH-D165), Japan; B, O. intermedium, male (27.1 × 25.2 mm) (ZRC 1970.2.23.6), Japan; C, Orisarma intermedium, male (34.7 × 31.6 mm) (ZRC 2001.0034), Pingtung, Taiwan; D, O. intermedium, male (37.5 × 35.8 mm) (ZRC 2014.0265), Kumejima Island, Japan; E, O. sinense, lectotype male (19.0 × 16.6 mm) (MNHN-BP3635a), China; F, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China; G, H, O. sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China. AG, outer view; H, inner view." figureDoi="http://doi.org/10.5281/zenodo.5351307" httpUri="https://zenodo.org/record/5351307/files/figure.png" pageId="24" pageNumber="915">Fig. 6EG</figureCitation>
). The proportions of the ambulatory legs are not useful, and in
<taxonomicName id="06BCE1C3FFF9BA39FAC5FDCDFA4FFD24" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[1304,1474,637,661]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFF9BA39FAC5FDCDFA4FFD24" box="[1304,1474,637,661]" italics="true" pageId="24" pageNumber="915">O. intermedium</emphasis>
</taxonomicName>
(which supposedly has longer legs), this character is too variable to be reliable. There seems to be a tendency (but not always) for the merus of the ambulatory legs to be relatively wider and shorter in smaller specimens (e.g.,
<figureCitation id="598786C5FFF9BA39FAE1FD4EFA38FCA4" box="[1340,1461,765,789]" captionStart="Fig" captionStartId="18.[135,170,1505,1527]" captionTargetBox="[136,1476,189,1489]" captionTargetId="figure-7@18.[136,1476,189,1489]" captionTargetPageId="18" captionText="Fig. 8. Right third and fourth ambulatory legs. A, Orisarma intermedium, male (27.0 × 25.0 mm) (ZRC 2013.0140), Kyushu, Japan; B, O. intermedium, male (29.5 × 26.5 mm) (ZRC 2001.0034), Pingtung, Taiwan; C, O. intermedium, male (37.5 × 35.8 mm) (ZRC 2014.0265), Kumejima Island, Japan; D, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China." figureDoi="http://doi.org/10.5281/zenodo.5351311" httpUri="https://zenodo.org/record/5351311/files/figure.png" pageId="24" pageNumber="915">Fig. 8A, B</figureCitation>
), with those of large specimens becoming longer and more slender (
<figureCitation id="598786C5FFF9BA39FC7CFC8EFC7AFCE4" box="[929,1015,829,853]" captionStart="Fig" captionStartId="18.[135,170,1505,1527]" captionTargetBox="[136,1476,189,1489]" captionTargetId="figure-7@18.[136,1476,189,1489]" captionTargetPageId="18" captionText="Fig. 8. Right third and fourth ambulatory legs. A, Orisarma intermedium, male (27.0 × 25.0 mm) (ZRC 2013.0140), Kyushu, Japan; B, O. intermedium, male (29.5 × 26.5 mm) (ZRC 2001.0034), Pingtung, Taiwan; C, O. intermedium, male (37.5 × 35.8 mm) (ZRC 2014.0265), Kumejima Island, Japan; D, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China." figureDoi="http://doi.org/10.5281/zenodo.5351311" httpUri="https://zenodo.org/record/5351311/files/figure.png" pageId="24" pageNumber="915">Fig. 8C</figureCitation>
). In fact, on the basis of the specimens of both species available, the ambulatory meri of
<taxonomicName id="06BCE1C3FFF9BA39FA92FCEDFA4FFCC4" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[1359,1474,862,885]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="sinense">
<emphasis id="F3C84652FFF9BA39FA92FCEDFA4FFCC4" box="[1359,1474,862,885]" italics="true" pageId="24" pageNumber="915">O. sinense</emphasis>
</taxonomicName>
actually appear proportionately shorter (
<figureCitation id="598786C5FFF9BA39FB2FFCCEFACBFC24" box="[1266,1350,893,917]" captionStart="Fig" captionStartId="18.[135,170,1505,1527]" captionTargetBox="[136,1476,189,1489]" captionTargetId="figure-7@18.[136,1476,189,1489]" captionTargetPageId="18" captionText="Fig. 8. Right third and fourth ambulatory legs. A, Orisarma intermedium, male (27.0 × 25.0 mm) (ZRC 2013.0140), Kyushu, Japan; B, O. intermedium, male (29.5 × 26.5 mm) (ZRC 2001.0034), Pingtung, Taiwan; C, O. intermedium, male (37.5 × 35.8 mm) (ZRC 2014.0265), Kumejima Island, Japan; D, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China." figureDoi="http://doi.org/10.5281/zenodo.5351311" httpUri="https://zenodo.org/record/5351311/files/figure.png" pageId="24" pageNumber="915">Fig. 8D</figureCitation>
versus
<figureCitation id="598786C5FFF9BA39FA45FCCEFC09FC04" captionStart="Fig" captionStartId="18.[135,170,1505,1527]" captionTargetBox="[136,1476,189,1489]" captionTargetId="figure-7@18.[136,1476,189,1489]" captionTargetPageId="18" captionText="Fig. 8. Right third and fourth ambulatory legs. A, Orisarma intermedium, male (27.0 × 25.0 mm) (ZRC 2013.0140), Kyushu, Japan; B, O. intermedium, male (29.5 × 26.5 mm) (ZRC 2001.0034), Pingtung, Taiwan; C, O. intermedium, male (37.5 × 35.8 mm) (ZRC 2014.0265), Kumejima Island, Japan; D, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China." figureDoi="http://doi.org/10.5281/zenodo.5351311" httpUri="https://zenodo.org/record/5351311/files/figure.png" pageId="24" pageNumber="915">Fig. 8AC</figureCitation>
) when specimens of similar size are compared. The G1 differences, however, are valid and reliable (
<figureCitation id="598786C5FFF9BA39FA97FC0EFA4FFC64" box="[1354,1474,957,981]" captionStart="Fig" captionStartId="25.[113,148,1906,1928]" captionTargetBox="[116,1451,292,1890]" captionTargetId="figure-15@25.[99,1466,286,1902]" captionTargetPageId="25" captionText="Fig. 12. AE, Orisarma intermedium, male (27.1 × 25.2 mm) (ZRC 1970.2.23.6), Japan; FJ, O. sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China; KO, O. patshuni, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong. A, F, K, male pleons; B, G, L, left G1 (ventral view, denuded); C, H, M, left G1 (dorsal view, denuded); D, I, N, left distal part of G1 (ventral view, denuded); E, J, O, left distal part of G1 (dorsal view, denuded). Scales: A, F = 5.0 mm; BE, GJ, KM = 1.0 mm; N, O = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5351321" httpUri="https://zenodo.org/record/5351321/files/figure.png" pageId="24" pageNumber="915">Fig. 12GJ</figureCitation>
versus
<figureCitation id="598786C5FFF9BA39FC4DFC6EFB9CFC44" box="[912,1041,989,1013]" captionStart="Fig" captionStartId="25.[113,148,1906,1928]" captionTargetBox="[116,1451,292,1890]" captionTargetId="figure-15@25.[99,1466,286,1902]" captionTargetPageId="25" captionText="Fig. 12. AE, Orisarma intermedium, male (27.1 × 25.2 mm) (ZRC 1970.2.23.6), Japan; FJ, O. sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China; KO, O. patshuni, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong. A, F, K, male pleons; B, G, L, left G1 (ventral view, denuded); C, H, M, left G1 (dorsal view, denuded); D, I, N, left distal part of G1 (ventral view, denuded); E, J, O, left distal part of G1 (dorsal view, denuded). Scales: A, F = 5.0 mm; BE, GJ, KM = 1.0 mm; N, O = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5351321" httpUri="https://zenodo.org/record/5351321/files/figure.png" pageId="24" pageNumber="915">Fig. 12BE</figureCitation>
).
</paragraph>
<paragraph id="C1039A40FFF9BA39FC9EFBADFA28FA64" blockId="24.[835,1475,1053,1493]" pageId="24" pageNumber="915">
In addition to the epibranchial tooth, chela, and G1 differences noted above, adults of the two species can be separated by the form of the frontal margin and carapace physiognomy. In
<taxonomicName id="06BCE1C3FFF9BA39FCBEFBCDFB83FB24" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[867,1038,1149,1173]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFF9BA39FCBEFBCDFB83FB24" box="[867,1038,1149,1173]" italics="true" pageId="24" pageNumber="915">O. intermedium</emphasis>
</taxonomicName>
, the two frontal lobes are separated by a deep median concavity (
<figureCitation id="598786C5FFF9BA39FBBAFB2EFB30FB04" box="[1127,1213,1181,1205]" captionStart="Fig" captionStartId="12.[135,170,1954,1976]" captionTargetBox="[136,1476,241,1939]" captionTargetId="figure-7@12.[136,1476,241,1939]" captionTargetPageId="12" captionText="Fig. 4. A, F, Orisarma dehaani, male (36.4 × 34.4 mm) (ZRC 2011.1027), Kumejima Island, Japan; B, D, O. dehaani, male (27.6 × 29.6 mm) (ZRC 2002.0223), Hong Kong; C, E, G, O. neglectum, male (31.7 × 28.0 mm) (ZRC 1998.309), Shanghai, China; H, J, O. intermedium, male (29.5 × 26.5 mm) (ZRC 2001.0034), Pingtung, Taiwan; I, K, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China. AC, dorsal view of carapace; D, E, H, I, frontal and anterolateral margins; F, G, J, K, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.5351303" httpUri="https://zenodo.org/record/5351303/files/figure.png" pageId="24" pageNumber="915">Fig. 4H</figureCitation>
) but in
<taxonomicName id="06BCE1C3FFF9BA39FAC4FB2DFA01FB04" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[1305,1420,1182,1205]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="sinense">
<emphasis id="F3C84652FFF9BA39FAC4FB2DFA01FB04" box="[1305,1420,1182,1205]" italics="true" pageId="24" pageNumber="915">O. sinense</emphasis>
</taxonomicName>
, this concavity is distinctly shallower (
<figureCitation id="598786C5FFF9BA39FB78FB0EFB63FB64" box="[1189,1262,1213,1237]" captionStart="Fig" captionStartId="12.[135,170,1954,1976]" captionTargetBox="[136,1476,241,1939]" captionTargetId="figure-7@12.[136,1476,241,1939]" captionTargetPageId="12" captionText="Fig. 4. A, F, Orisarma dehaani, male (36.4 × 34.4 mm) (ZRC 2011.1027), Kumejima Island, Japan; B, D, O. dehaani, male (27.6 × 29.6 mm) (ZRC 2002.0223), Hong Kong; C, E, G, O. neglectum, male (31.7 × 28.0 mm) (ZRC 1998.309), Shanghai, China; H, J, O. intermedium, male (29.5 × 26.5 mm) (ZRC 2001.0034), Pingtung, Taiwan; I, K, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China. AC, dorsal view of carapace; D, E, H, I, frontal and anterolateral margins; F, G, J, K, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.5351303" httpUri="https://zenodo.org/record/5351303/files/figure.png" pageId="24" pageNumber="915">Fig. 4I</figureCitation>
). The dorsal surface of the carapace of
<taxonomicName id="06BCE1C3FFF9BA39FBD6FB6DFB39FB44" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[1035,1204,1245,1269]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFF9BA39FBD6FB6DFB39FB44" box="[1035,1204,1245,1269]" italics="true" pageId="24" pageNumber="915">O. intermedium</emphasis>
</taxonomicName>
is relatively less convex, with the surfaces usually slightly more rugose and with scattered stiff setae all over the anterior and branchial regions (
<figureCitation id="598786C5FFF9BA39FC96FA8EFC45FAE4" box="[843,968,1341,1365]" captionStart="Fig" captionStartId="8.[136,171,1926,1948]" captionTargetBox="[136,1476,212,1910]" captionTargetId="figure-7@8.[136,1476,212,1910]" captionTargetPageId="8" captionText="Fig. 2. Overall habitus. A, Orisarma intermedium, lectotype male (23.0 × 19.9 mm) (RMNH-D165), Japan; B, O. intermedium, male (27.1 × 25.2 mm) (ZRC 1970.2.23.6), Japan; C, O. intermedium, male (27.0 × 25.0 mm) (ZRC 2013.0140), Kyushu, Japan; D, O. intermedium, male (29.5 × 26.5 mm) (ZRC 2001.0034), Pingtung, Taiwan; E, O. intermedium, male (34.2 × 30.7 mm) (ZRC 2001.0034), Pingtung, Taiwan; F, O. intermedium, male (37.5 × 35.8 mm) (ZRC 2014.0265), Kumejima Island, Japan; G, O. sinense, lectotype male (19.0 × 16.6 mm) (MNHN-BP3635a), China; H, O. sinense, paralectotype female (18.8 × 16.9 mm) (MNHN-BP3635b), China; I, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China; J, Orisarma sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China." figureDoi="http://doi.org/10.5281/zenodo.5351299" httpUri="https://zenodo.org/record/5351299/files/figure.png" pageId="24" pageNumber="915">Figs. 2AF</figureCitation>
,
<figureCitation id="598786C5FFF9BA39FC0EFA8EFB82FAE4" box="[979,1039,1341,1365]" captionStart="Fig" captionStartId="12.[135,170,1954,1976]" captionTargetBox="[136,1476,241,1939]" captionTargetId="figure-7@12.[136,1476,241,1939]" captionTargetPageId="12" captionText="Fig. 4. A, F, Orisarma dehaani, male (36.4 × 34.4 mm) (ZRC 2011.1027), Kumejima Island, Japan; B, D, O. dehaani, male (27.6 × 29.6 mm) (ZRC 2002.0223), Hong Kong; C, E, G, O. neglectum, male (31.7 × 28.0 mm) (ZRC 1998.309), Shanghai, China; H, J, O. intermedium, male (29.5 × 26.5 mm) (ZRC 2001.0034), Pingtung, Taiwan; I, K, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China. AC, dorsal view of carapace; D, E, H, I, frontal and anterolateral margins; F, G, J, K, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.5351303" httpUri="https://zenodo.org/record/5351303/files/figure.png" pageId="24" pageNumber="915">4H, J</figureCitation>
). In
<taxonomicName id="06BCE1C3FFF9BA39FB9CFA8DFB3CFAE4" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[1089,1201,1342,1365]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="sinense">
<emphasis id="F3C84652FFF9BA39FB9CFA8DFB3CFAE4" box="[1089,1201,1342,1365]" italics="true" pageId="24" pageNumber="915">O. sinense</emphasis>
</taxonomicName>
, the dorsal surface of the carapace is more convex (notably the branchial), smoother, and the setae are more scattered and sparser on the branchial regions (
<figureCitation id="598786C5FFF9BA39FC79FA2EFB9BFA04" box="[932,1046,1437,1461]" captionStart="Fig" captionStartId="8.[136,171,1926,1948]" captionTargetBox="[136,1476,212,1910]" captionTargetId="figure-7@8.[136,1476,212,1910]" captionTargetPageId="8" captionText="Fig. 2. Overall habitus. A, Orisarma intermedium, lectotype male (23.0 × 19.9 mm) (RMNH-D165), Japan; B, O. intermedium, male (27.1 × 25.2 mm) (ZRC 1970.2.23.6), Japan; C, O. intermedium, male (27.0 × 25.0 mm) (ZRC 2013.0140), Kyushu, Japan; D, O. intermedium, male (29.5 × 26.5 mm) (ZRC 2001.0034), Pingtung, Taiwan; E, O. intermedium, male (34.2 × 30.7 mm) (ZRC 2001.0034), Pingtung, Taiwan; F, O. intermedium, male (37.5 × 35.8 mm) (ZRC 2014.0265), Kumejima Island, Japan; G, O. sinense, lectotype male (19.0 × 16.6 mm) (MNHN-BP3635a), China; H, O. sinense, paralectotype female (18.8 × 16.9 mm) (MNHN-BP3635b), China; I, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China; J, Orisarma sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China." figureDoi="http://doi.org/10.5281/zenodo.5351299" httpUri="https://zenodo.org/record/5351299/files/figure.png" pageId="24" pageNumber="915">Figs. 2I, J</figureCitation>
,
<figureCitation id="598786C5FFF9BA39FBF8FA2EFBEEFA04" box="[1061,1123,1437,1461]" captionStart="Fig" captionStartId="12.[135,170,1954,1976]" captionTargetBox="[136,1476,241,1939]" captionTargetId="figure-7@12.[136,1476,241,1939]" captionTargetPageId="12" captionText="Fig. 4. A, F, Orisarma dehaani, male (36.4 × 34.4 mm) (ZRC 2011.1027), Kumejima Island, Japan; B, D, O. dehaani, male (27.6 × 29.6 mm) (ZRC 2002.0223), Hong Kong; C, E, G, O. neglectum, male (31.7 × 28.0 mm) (ZRC 1998.309), Shanghai, China; H, J, O. intermedium, male (29.5 × 26.5 mm) (ZRC 2001.0034), Pingtung, Taiwan; I, K, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China. AC, dorsal view of carapace; D, E, H, I, frontal and anterolateral margins; F, G, J, K, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.5351303" httpUri="https://zenodo.org/record/5351303/files/figure.png" pageId="24" pageNumber="915">4I, K</figureCitation>
). These characters are valid for both sexes as well as for small and subadult specimens.
</paragraph>
<paragraph id="C1039A40FFF9BA39FC9EFA4DFC47F967" blockId="24.[835,1475,1533,1750]" pageId="24" pageNumber="915">
<bibRefCitation id="A52DE7B1FFF9BA39FC9EFA4DFBBAF9A4" author="De Man's" box="[835,1079,1533,1557]" firstAuthor="De Man's" page="182" pageId="24" pageNumber="915" pagination="177 - 240" refId="ref72143" refString="Man JG De (1888) Report on the Podophthalmous Crustacea of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta, by Dr. John Anderson, F. R. S., Superintendent of the Museum. - Part IV. Journal of the Linnean Society of London (Zoology), 22 (139): 177 - 240, pls. 13 - 15." type="journal article" year="1888">De Mans (1888: 182)</bibRefCitation>
<taxonomicName id="06BCE1C3FFF9BA39FB96FA4DFAAEF9A4" box="[1099,1315,1533,1557]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="F3C84652FFF9BA39FB96FA4DFAAEF9A4" box="[1099,1315,1533,1557]" italics="true" pageId="24" pageNumber="915">Sesarma intermedia</emphasis>
</taxonomicName>
” from Mergui is a misidentification of
<taxonomicName id="06BCE1C3FFF9BA39FBB7F9ADFC0BF9E4" authority="(De Man, 1892)" baseAuthorityName="De Man" baseAuthorityYear="1892" class="Malacostraca" family="Sesarmidae" genus="Manarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="moeschii">
<emphasis id="F3C84652FFF9BA39FBB7F9ADFAC4F984" box="[1130,1353,1565,1589]" italics="true" pageId="24" pageNumber="915">Manarma moeschii</emphasis>
(
<bibRefCitation id="A52DE7B1FFF9BA39FA86F9AEFCF3F9E4" author="De Man" firstAuthor="De Man" pageId="24" pageNumber="915" pagination="225 - 264" refId="ref72297" refString="Man JG De (1892) Carcinological studies in the Leyden Museum, No. 6. Notes Leyden Museum, 14 (36): 225 - 264, pls. 7 - 10." type="journal article" year="1892">De Man, 1892</bibRefCitation>
)
</taxonomicName>
. The specimen figured by
<bibRefCitation id="A52DE7B1FFF9BA39FB40F98EFA83F9E4" author="Soh" box="[1181,1294,1597,1621]" firstAuthor="Soh" pageId="24" pageNumber="915" pagination="9 - 22" refId="ref77463" refString="Soh CL (1978) On a collection of sesarmine crabs (Decapoda, Brachyura, Grapsidae) from Hong Kong. Memoirs of the Hong Kong Natural History Society, 13: 9 - 22." type="journal article" year="1978">Soh (1978</bibRefCitation>
: pl. 4b) is almost certainly
<taxonomicName id="06BCE1C3FFF9BA39FC7BF9EDFBC0F9C4" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[934,1101,1629,1653]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFF9BA39FC7BF9EDFBC0F9C4" box="[934,1101,1629,1653]" italics="true" pageId="24" pageNumber="915">O. intermedium</emphasis>
</taxonomicName>
on the basis of his photograph, and we have also examined material from
<collectingCountry id="B9ABDAD0FFF9BA39FB2EF9CDFAF5F927" box="[1267,1400,1662,1686]" name="Hong Kong" pageId="24" pageNumber="915">Hong Kong</collectingCountry>
. Thus, both
<taxonomicName id="06BCE1C3FFF9BA39FCA6F92DFBAEF904" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[891,1059,1693,1717]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFF9BA39FCA6F92DFBAEF904" box="[891,1059,1693,1717]" italics="true" pageId="24" pageNumber="915">O. intermedium</emphasis>
</taxonomicName>
and
<taxonomicName id="06BCE1C3FFF9BA39FB87F92DFB46F904" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[1114,1227,1694,1717]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="sinense">
<emphasis id="F3C84652FFF9BA39FB87F92DFB46F904" box="[1114,1227,1694,1717]" italics="true" pageId="24" pageNumber="915">O. sinense</emphasis>
</taxonomicName>
occur sympatrically in
<collectingCountry id="B9ABDAD0FFF9BA39FC9EF90DFC4BF967" box="[835,966,1726,1750]" name="Hong Kong" pageId="24" pageNumber="915">Hong Kong</collectingCountry>
.
</paragraph>
<paragraph id="C1039A40FFF9BA39FC9EF94DFC7FF824" blockId="24.[835,1474,1789,1941]" pageId="24" pageNumber="915">
<bibRefCitation id="A52DE7B1FFF9BA39FC9EF94DFBA6F8A4" author="Hoang" box="[835,1067,1789,1814]" etAl="et al." firstAuthor="Hoang" page="75" pageId="24" pageNumber="915" pagination="72 - 80" refId="ref70272" refString="Hoang NK, Do VN, Nguyen TT &amp; Nguyen TTH (2012) Preliminary data of three zoobenthos groups (Brachyura, Gastropoda and Bivalviva) found in the mangrove forest ecosystems of the Tran De Estuary, Soc Trang Province. Journal of Science of HNUE (Hanoi National University of Education), Chemical and Biological Sciences, 57 (8): 72 - 80." type="journal article" year="2012">Hoang et al. (2012: 75</bibRefCitation>
, 78) reported “
<taxonomicName id="06BCE1C3FFF9BA39FB19F94DFA3AF8A4" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[1220,1463,1789,1813]" class="Malacostraca" family="Sesarmidae" genus="Sesarmops" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="F3C84652FFF9BA39FB19F94DFA3AF8A4" box="[1220,1463,1789,1813]" italics="true" pageId="24" pageNumber="915">Sesarmops intermedius</emphasis>
</taxonomicName>
” from southern
<collectingCountry id="B9ABDAD0FFF9BA39FC35F8AEFBC9F884" box="[1000,1092,1821,1845]" name="Vietnam" pageId="24" pageNumber="915">Vietnam</collectingCountry>
but their record is dubious as the species is not known to be present that far south; and it is more likely their material was confused with a species of
<taxonomicName id="06BCE1C3FFF9BA39FC9EF8CEFC67F824" authorityName=", Serene &amp; Soh" authorityYear="1970" box="[835,1002,1917,1941]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="915" phylum="Arthropoda" rank="genus">
<emphasis id="F3C84652FFF9BA39FC9EF8CEFC67F824" box="[835,1002,1917,1941]" italics="true" pageId="24" pageNumber="915">Pseudosesarma</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFF9BA3AFC9EF80EFDF6FD24" lastPageId="27" lastPageNumber="918" pageId="24" pageNumber="915" type="biology_ecology">
<paragraph id="C1039A40FFF9BA3AFC9EF80EFDF6FD24" blockId="24.[835,1475,1981,2102]" lastBlockId="27.[113,753,189,662]" lastPageId="27" lastPageNumber="918" pageId="24" pageNumber="915">
<emphasis id="F3C84652FFF9BA39FC9EF80EFC14F864" bold="true" box="[835,921,1981,2005]" pageId="24" pageNumber="915">Biology</emphasis>
. This species is common in freshwater streams and waters near the sea, which may still be subject to some tidal influence, although they can also be found several hundreds of metres away on hilly areas. They normally occur among dense vegetation along these bodies of water, coming out at night to forage on the sand- and mudflats. They are primarily herbivores and scavengers.
<bibRefCitation id="A52DE7B1FFFABA3AFDABFF4DFF35FE84" author="Hong" etAl="et al." firstAuthor="Hong" pageId="27" pageNumber="918" pagination="255 - 263" refId="ref70376" refString="Hong J-S, Choi J-W &amp; Tsutsumi H (2010) Concluding remarks on the joint survey of macrobenthic fauna on Suncheon Tidal Flats by the participants of &quot; Korea and Japan Joint Symposium on Biology of Tidal Flats 2009 &quot;. Plankton Benthos Research, 5 (Supplement): 255 - 263." type="journal article" year="2010">Hong et al. (2010)</bibRefCitation>
discussed its ecology in
<collectingCountry id="B9ABDAD0FFFABA3AFE09FEADFD94FE84" box="[468,537,286,309]" name="South Korea" pageId="27" pageNumber="918">Korea</collectingCountry>
. The physiology of the species in
<collectingCountry id="B9ABDAD0FFFABA3AFEC5FE8EFEE7FEE4" box="[280,362,317,341]" name="Taiwan" pageId="27" pageNumber="918">Taiwan</collectingCountry>
was studied by
<bibRefCitation id="A52DE7B1FFFABA3AFDF4FE8EFD7DFEE4" author="Tsai" box="[553,752,317,341]" etAl="et al." firstAuthor="Tsai" pageId="27" pageNumber="918" pagination="61 - 78" refId="ref78337" refString="Tsai M-L, Li J-J &amp; Dai C-F (2000) Is large body size always advantageous for terrestrial adaptation? A study of water balance in a semi-terrestrial crab, Sesarmops intermedium (Crustacea: Grapsidae). Evolutionary Ecology, 14: 61 - 78." type="journal article" year="2000">Tsai et al. (2000)</bibRefCitation>
while the visual orientation and habits was done by
<bibRefCitation id="A52DE7B1FFFABA3AFD74FEEDFE87FE24" author="Huang &amp; Liu" firstAuthor="Huang" pageId="27" pageNumber="918" pagination="267 - 267" refId="ref70431" refString="Huang H-D &amp; Liu H-C (2012) Comparison of visual orientation between two land crabs Sesarmops intermedium and Scandarma lintou in Kenting National Park, southern Taiwan. Integrative and Comparative Biology, 52: 267 - 267." type="journal article" year="2012">Huang &amp; Liu (2012)</bibRefCitation>
. The reproductive behaviour of this species in
<collectingCountry id="B9ABDAD0FFFABA3AFF53FE2DFF46FE04" box="[142,203,414,437]" name="Japan" pageId="27" pageNumber="918">Japan</collectingCountry>
was discussed by
<bibRefCitation id="A52DE7B1FFFABA3AFE48FE2DFDB5FE04" author="Kyomo" box="[405,568,413,438]" firstAuthor="Kyomo" pageId="27" pageNumber="918" pagination="319 - 329" refId="ref71147" refString="Kyomo J (1986) Reproductive activities in the sesarmid crab Sesarma intermedia in the coastal and estuarine habitats of Hakata, Japan. Marine Biology, 91: 319 - 329." type="journal article" year="1986">Kyomo (1986)</bibRefCitation>
. The larvae have been reported on by
<bibRefCitation id="A52DE7B1FFFABA3AFE85FE0EFDD7FE64" author="Fukuda &amp; Baba" box="[344,602,445,469]" firstAuthor="Fukuda" pageId="27" pageNumber="918" pagination="61 - 75" refId="ref69321" refString="Fukuda Y &amp; Baba K (1976) Complete larval development of the sesarminid crabs, Chiromantes bidens, Holometopus haematocheir, Parasesarma plicatum, and Sesarmops intermedius, reared in the laboratory. Memoirs of the Faculty of Education, Kumamoto University, Natural Science, 25: 61 - 75." type="journal article" year="1976">Fukuda &amp; Baba (1976)</bibRefCitation>
,
<bibRefCitation id="A52DE7B1FFFABA3AFDBAFE0EFF36FE44" author="Cuesta" etAl="et al." firstAuthor="Cuesta" pageId="27" pageNumber="918" pagination="151 - 173" refId="ref68440" refString="Cuesta JA, Guerao G, Liu H-C &amp; Schubart CD (2006) Morphology of the first zoeal stages of eleven Sesarmidae (Crustacea, Brachyura, Thoracotremata) from the Indo-West Pacific, with a summary of familial larval characters. Invertebrate Reproduction &amp; Development, 49 (3): 151 - 173." type="journal article" year="2006">Cuesta et al. (2006)</bibRefCitation>
, and
<bibRefCitation id="A52DE7B1FFFABA3AFF24FE6DFEF8FE44" author="Lee" box="[249,373,477,501]" firstAuthor="Lee" pageId="27" pageNumber="918" pagination="321 - 324" refId="ref71400" refString="Lee HJ (2015) First zoea of Sesarmops intermedius (De Haan, 1835) (Crustacea: Decapoda: Brachyura: Sesarmidae). Fisheries &amp; Aquatic Sciences, 18 (3): 321 - 324." type="journal article" year="2015">Lee (2015)</bibRefCitation>
. The microsatellite markers of the species were studied by
<bibRefCitation id="A52DE7B1FFFABA3AFEB1FE4DFD97FDA4" author="Lee" box="[364,538,509,533]" etAl="et al." firstAuthor="Lee" pageId="27" pageNumber="918" pagination="183 - 185" refId="ref71461" refString="Lee S-K, Jung J, Kang S &amp; Kim W (2012) Isolation and characterization of microsatellite markers for the endangered crab species Sesarmops intermedius. Conservation Genetics and Resources, 4: 183 - 185." type="journal article" year="2012">Lee et al. (2012)</bibRefCitation>
. Nematode parasites have been reported on by
<bibRefCitation id="A52DE7B1FFFABA3AFE55FDAEFDDDFD84" author="Yoshimura" box="[392,592,541,565]" firstAuthor="Yoshimura" pageId="27" pageNumber="918" pagination="755 - 761" refId="ref78855" refString="Yoshimura K (1990) Two new species of the genus Monhystrium Cobb, 1920 (Monhysteridae: Nematoda) from terrestrial crabs of subfamily Sesarminae (Brachyura) in Japan. Zoological Science, 7 (4): 755 - 761." type="journal article" year="1990">Yoshimura (1990)</bibRefCitation>
. In
<collectingCountry id="B9ABDAD0FFFABA3AFDA5FDAEFD46FD84" box="[632,715,541,565]" name="Taiwan" pageId="27" pageNumber="918">Taiwan</collectingCountry>
,
<taxonomicName id="06BCE1C3FFFABA3AFD0BFDADFF77FDE4" baseAuthorityName="De Haan" baseAuthorityYear="1835" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="918" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFFABA3AFD0BFDADFF77FDE4" italics="true" pageId="27" pageNumber="918">O. intermedium</emphasis>
</taxonomicName>
is collected in large numbers from southern
<collectingCountry id="B9ABDAD0FFFABA3AFFAFFDEEFF4FFDC4" box="[114,194,605,629]" name="Taiwan" pageId="27" pageNumber="918">Taiwan</collectingCountry>
for the aquarium trade (as a freshwater crab), and many populations have been seriously depleted.
</paragraph>
</subSubSection>
<caption id="95C3CAC8FFF8BA38FFACF8C1FB5BF86F" ID-DOI="http://doi.org/10.5281/zenodo.5351321" ID-Zenodo-Dep="5351321" httpUri="https://zenodo.org/record/5351321/files/figure.png" pageId="25" pageNumber="916" startId="25.[113,148,1906,1928]" targetBox="[116,1451,292,1890]" targetPageId="25">
<paragraph id="C1039A40FFF8BA38FFACF8C1FB5BF86F" blockId="25.[113,1452,1906,2015]" pageId="25" pageNumber="916">
Fig. 12. AE,
<taxonomicName id="06BCE1C3FFF8BA38FF22F8C0FE50F836" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[255,477,1906,1928]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="916" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFF8BA38FF22F8C0FE50F836" box="[255,477,1906,1928]" italics="true" pageId="25" pageNumber="916">Orisarma intermedium</emphasis>
</taxonomicName>
, male (27.1 × 25.2 mm) (ZRC 1970.2.23.6), Japan; FJ,
<taxonomicName id="06BCE1C3FFF8BA38FBC9F8C0FBF7F839" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[1044,1146,1907,1928]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="916" phylum="Arthropoda" rank="species" species="sinense">
<emphasis id="F3C84652FFF8BA38FBC9F8C0FBF7F839" box="[1044,1146,1907,1928]" italics="true" pageId="25" pageNumber="916">O. sinense</emphasis>
</taxonomicName>
, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China; KO,
<taxonomicName id="06BCE1C3FFF8BA38FE07F83CFDC0F815" authorityName="Soh" authorityYear="1978" box="[474,589,1935,1956]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="916" phylum="Arthropoda" rank="species" species="patshuni">
<emphasis id="F3C84652FFF8BA38FE07F83CFDC0F815" box="[474,589,1935,1956]" italics="true" pageId="25" pageNumber="916">O. patshuni</emphasis>
</taxonomicName>
, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong. A, F, K, male pleons; B, G, L, left G1 (ventral view, denuded); C, H, M, left G1 (dorsal view, denuded); D, I, N, left distal part of G1 (ventral view, denuded); E, J, O, left distal part of G1 (dorsal view, denuded). Scales: A, F = 5.0 mm; BE, GJ, KM = 1.0 mm; N, O = 0.5 mm.
</paragraph>
</caption>
<caption id="95C3CAC8FFFBBA3BFF55F8C5FA21F789" ID-DOI="http://doi.org/10.5281/zenodo.5351323" ID-Zenodo-Dep="5351323" httpUri="https://zenodo.org/record/5351323/files/figure.png" pageId="26" pageNumber="917" startId="26.[136,171,1910,1932]" targetBox="[136,1474,199,1895]" targetPageId="26">
<paragraph id="C1039A40FFFBBA3BFF55F8C5FA21F789" blockId="26.[136,1475,1910,2105]" pageId="26" pageNumber="917">
Fig. 13. Overall habitus. A,
<taxonomicName id="06BCE1C3FFFBBA3BFE4EF8C4FDECF83A" baseAuthorityName="Dana" baseAuthorityYear="1851" box="[403,609,1910,1932]" class="Malacostraca" family="Sesarmidae" genus="Danarma" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="917" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="F3C84652FFFBBA3BFE4EF8C4FDECF83A" box="[403,609,1910,1932]" italics="true" pageId="26" pageNumber="917">Danarma obtusifrons</emphasis>
</taxonomicName>
, male (17.7 × 13.2 mm) (UF-FLMNH 14837), Oahu, Hawaii; B,
<taxonomicName id="06BCE1C3FFFBBA3BFB08F8C4FAEFF83A" baseAuthorityName="Davie &amp; Ng" baseAuthorityYear="2013" box="[1237,1378,1910,1932]" class="Malacostraca" family="Sesarmidae" genus="Danarma" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="917" phylum="Arthropoda" rank="species" species="silus">
<emphasis id="F3C84652FFFBBA3BFB08F8C4FAEFF83A" box="[1237,1378,1910,1932]" italics="true" pageId="26" pageNumber="917">Danarma silus</emphasis>
</taxonomicName>
, holotype male (16.3 × 12.8 mm) (ZRC 2012.0787), Guam; C,
<taxonomicName id="06BCE1C3FFFBBA3BFDA5F827FCB0F818" baseAuthorityName="Davie &amp; Ng" baseAuthorityYear="2013" box="[632,829,1940,1961]" class="Malacostraca" family="Sesarmidae" genus="Danarma" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="917" phylum="Arthropoda" rank="species" species="eurymerus">
<emphasis id="F3C84652FFFBBA3BFDA5F827FCB0F818" box="[632,829,1940,1961]" italics="true" pageId="26" pageNumber="917">Danarma eurymerus</emphasis>
</taxonomicName>
, holotype male (18.7 × 14.8 mm) (NMNS-7028-002), Lanyu Island, Taiwan; D,
<taxonomicName id="06BCE1C3FFFBBA3BFF27F802FE35F874" baseAuthorityName="Davie &amp; Ng" baseAuthorityYear="2013" box="[250,440,1968,1990]" class="Malacostraca" family="Sesarmidae" genus="Danarma" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="917" phylum="Arthropoda" rank="species" species="garfunkel">
<emphasis id="F3C84652FFFBBA3BFF27F802FE35F874" box="[250,440,1968,1990]" italics="true" pageId="26" pageNumber="917">Danarma garfunkel</emphasis>
</taxonomicName>
, holotype male (17.0 × 13.0 mm) (QM), Greta Beach, Christmas Island; E,
<taxonomicName id="06BCE1C3FFFBBA3BFB4DF803FADDF874" baseAuthorityName="De Man, in Weber" baseAuthorityYear="1897" box="[1168,1360,1968,1989]" class="Malacostraca" family="Sesarmidae" genus="Cristarma" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="917" phylum="Arthropoda" rank="species" species="eulimene">
<emphasis id="F3C84652FFFBBA3BFB4DF803FADDF874" box="[1168,1360,1968,1989]" italics="true" pageId="26" pageNumber="917">Cristarma eulimene</emphasis>
</taxonomicName>
, male (22.2 × 17.2 mm) (ZRC 1968.1.22.2), Inhaca Island, Mozambique; F,
<taxonomicName id="06BCE1C3FFFBBA3BFCDCF87EFC4EF853" baseAuthorityName="Crosnier" baseAuthorityYear="1965" box="[769,963,1997,2018]" class="Malacostraca" family="Sesarmidae" genus="Cristarma" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="917" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="F3C84652FFFBBA3BFCDCF87EFC4EF853" box="[769,963,1997,2018]" italics="true" pageId="26" pageNumber="917">Cristarma ortmanni</emphasis>
</taxonomicName>
, male (20.0 × 15.5 mm) (ZRC 1968.1.22.1), Inhaca Island, Mozambique; G,
<taxonomicName id="06BCE1C3FFFBBA3BFEA7F859FDDAF84F" baseAuthorityName="Brito Capello" baseAuthorityYear="1864" box="[378,599,2025,2047]" class="Malacostraca" family="Sesarmidae" genus="Trapezarma" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="917" phylum="Arthropoda" rank="species" species="angolense">
<emphasis id="F3C84652FFFBBA3BFEA7F859FDDAF84F" box="[378,599,2025,2047]" italics="true" pageId="26" pageNumber="917">Trapezarma angolense</emphasis>
</taxonomicName>
, neotype male (39.3 × 32.7 mm) (SMF-ZMG 635), Benguella, Angola; H,
<taxonomicName id="06BCE1C3FFFBBA3BFAEBF85AFF75F7AA" baseAuthorityName="De Man" baseAuthorityYear="1883" class="Malacostraca" family="Sesarmidae" genus="Platychirarma" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="917" phylum="Arthropoda" rank="species" species="buettikoferi">
<emphasis id="F3C84652FFFBBA3BFAEBF85AFF75F7AA" italics="true" pageId="26" pageNumber="917">Platychirarma buettikoferi</emphasis>
</taxonomicName>
, paralectotype male (10.0 × 8.7 mm) (SMF-ZMG 636), Fisherman Lake, Liberia; I,
<taxonomicName id="06BCE1C3FFFBBA3BFB89F7B5FAD7F7AA" baseAuthorityName="De Man" baseAuthorityYear="1883" box="[1108,1370,2054,2075]" class="Malacostraca" family="Sesarmidae" genus="Platychirarma" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="917" phylum="Arthropoda" rank="species" species="buettikoferi">
<emphasis id="F3C84652FFFBBA3BFB89F7B5FAD7F7AA" box="[1108,1370,2054,2075]" italics="true" pageId="26" pageNumber="917">Platychirarma buettikoferi</emphasis>
</taxonomicName>
, lectotype male (14.1 × 11.6 mm) (RMNH D 148), Liberia; J,
<taxonomicName id="06BCE1C3FFFBBA3BFD57F790FC00F789" baseAuthorityName="De Man" baseAuthorityYear="1883" box="[650,909,2083,2104]" class="Malacostraca" family="Sesarmidae" genus="Platychirarma" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="917" phylum="Arthropoda" rank="species" species="buettikoferi">
<emphasis id="F3C84652FFFBBA3BFD57F790FC00F789" box="[650,909,2083,2104]" italics="true" pageId="26" pageNumber="917">Platychirarma buettikoferi</emphasis>
</taxonomicName>
, male (12.6 × 10.4 mm) (ZRC 2015.0298), Cameroon.
</paragraph>
</caption>
<subSubSection id="89A6C9CBFFFABA3AFFACFD0EFF73FAA4" pageId="27" pageNumber="918" type="distribution">
<paragraph id="C1039A40FFFABA3AFFACFD0EFF73FAA4" blockId="27.[113,754,701,1301]" pageId="27" pageNumber="918">
<emphasis id="F3C84652FFFABA3AFFACFD0EFF71FD64" bold="true" box="[113,252,701,725]" pageId="27" pageNumber="918">Distribution</emphasis>
. Despite the lack of genetic differentiation in the current genetic dataset, we are confident
<taxonomicName id="06BCE1C3FFFABA3AFD94FD6DFD7DFD44" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[585,752,733,757]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="918" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFFABA3AFD94FD6DFD7DFD44" box="[585,752,733,757]" italics="true" pageId="27" pageNumber="918">O. intermedium</emphasis>
</taxonomicName>
and
<taxonomicName id="06BCE1C3FFFABA3AFF7EFD4DFE95FCA4" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[163,280,766,789]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="918" phylum="Arthropoda" rank="species" species="sinense">
<emphasis id="F3C84652FFFABA3AFF7EFD4DFE95FCA4" box="[163,280,766,789]" italics="true" pageId="27" pageNumber="918">O. sinense</emphasis>
</taxonomicName>
are separate species on the basis of their morphology. However, we are less sure about their distribution. In
<collectingCountry id="B9ABDAD0FFFABA3AFEC5FC8DFEDAFCE4" box="[280,343,830,853]" name="Japan" pageId="27" pageNumber="918">Japan</collectingCountry>
,
<collectingCountry id="B9ABDAD0FFFABA3AFEBFFC8DFE2BFCE4" box="[354,422,830,853]" name="South Korea" pageId="27" pageNumber="918">Korea</collectingCountry>
, and
<collectingCountry id="B9ABDAD0FFFABA3AFE03FC8EFDBCFCE4" box="[478,561,829,853]" name="Taiwan" pageId="27" pageNumber="918">Taiwan</collectingCountry>
, only one species,
<taxonomicName id="06BCE1C3FFFABA3AFFACFCEDFE96FCC4" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[113,283,861,885]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="918" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFFABA3AFFACFCEDFE96FCC4" box="[113,283,861,885]" italics="true" pageId="27" pageNumber="918">O. intermedium</emphasis>
</taxonomicName>
, is known for certain. On mainland
<collectingCountry id="B9ABDAD0FFFABA3AFD77FCEEFD61FCC4" box="[682,748,861,885]" name="China" pageId="27" pageNumber="918">China</collectingCountry>
, notably southern
<collectingCountry id="B9ABDAD0FFFABA3AFEE8FCCEFEF4FC24" box="[309,377,893,917]" name="China" pageId="27" pageNumber="918">China</collectingCountry>
, both species have been reported (
<bibRefCitation id="A52DE7B1FFFABA3AFFA7FC2EFE81FC04" author="Shen" box="[122,268,925,949]" firstAuthor="Shen" pageId="27" pageNumber="918" pagination="211 - 242" refId="ref76939" refString="Shen CJ (1940 a) The brachyuran fauna of Hong Kong. Journal of the Hong Kong Fisheries Research Station, 1: 211 - 242." type="journal article" year="1940" yearSuffix="a">Shen, 1940a</bibRefCitation>
, b;
<bibRefCitation id="A52DE7B1FFFABA3AFEE1FC2EFE7AFC04" author="Dai" box="[316,503,925,949]" etAl="et al." firstAuthor="Dai" pageId="27" pageNumber="918" pagination="1 - 295" refId="ref68573" refString="Dai AY, Yang SL, Song YZ &amp; Chen GX (1986) Crabs of the China Seas. China Ocean Press, Beijing, 17 + 642 pp., figs. 1 - 295, pls. 1 - 74." type="book chapter" year="1986">Dai et al., 1986</bibRefCitation>
;
<bibRefCitation id="A52DE7B1FFFABA3AFDDAFC2EFD6EFC04" author="Dai &amp; Yang" box="[519,739,925,950]" firstAuthor="Dai" pageId="27" pageNumber="918" pagination="1 - 74" refId="ref68539" refString="Dai AY &amp; Yang SL (1991) Crabs of the China Seas. China Ocean Press, Beijing and Berlin, xxi + 681 pp., pls. 1 - 74." type="book chapter" year="1991">Dai &amp; Yang, 1991</bibRefCitation>
). Certainly, both species appear to be present in
<collectingCountry id="B9ABDAD0FFFABA3AFDAFFC0DFD7CFC67" box="[626,753,958,982]" name="Hong Kong" pageId="27" pageNumber="918">Hong Kong</collectingCountry>
(see
<bibRefCitation id="A52DE7B1FFFABA3AFF74FC6EFEACFC44" author="Soh" box="[169,289,989,1013]" firstAuthor="Soh" pageId="27" pageNumber="918" pagination="9 - 22" refId="ref77463" refString="Soh CL (1978) On a collection of sesarmine crabs (Decapoda, Brachyura, Grapsidae) from Hong Kong. Memoirs of the Hong Kong Natural History Society, 13: 9 - 22." type="journal article" year="1978">Soh, 1978</bibRefCitation>
). The older material will need to be reexamined and fresh collections made to determine their actual distributional limits. In the most recent treatments of the Chinese fauna (
<bibRefCitation id="A52DE7B1FFFABA3AFE9CFB8EFE61FBE4" author="Dai" box="[321,492,1085,1109]" etAl="et al." firstAuthor="Dai" pageId="27" pageNumber="918" pagination="1 - 295" refId="ref68573" refString="Dai AY, Yang SL, Song YZ &amp; Chen GX (1986) Crabs of the China Seas. China Ocean Press, Beijing, 17 + 642 pp., figs. 1 - 295, pls. 1 - 74." type="book chapter" year="1986">Dai et al., 1986</bibRefCitation>
;
<bibRefCitation id="A52DE7B1FFFABA3AFE25FB8EFD4FFBE4" author="Dai &amp; Yang" box="[504,706,1085,1110]" firstAuthor="Dai" pageId="27" pageNumber="918" pagination="1 - 74" refId="ref68539" refString="Dai AY &amp; Yang SL (1991) Crabs of the China Seas. China Ocean Press, Beijing and Berlin, xxi + 681 pp., pls. 1 - 74." type="book chapter" year="1991">Dai &amp; Yang, 1991</bibRefCitation>
),
<taxonomicName id="06BCE1C3FFFABA3AFD0BFB8DFF75FBC4" baseAuthorityName="De Haan" baseAuthorityYear="1835" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="918" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFFABA3AFD0BFB8DFF75FBC4" italics="true" pageId="27" pageNumber="918">O. intermedium</emphasis>
</taxonomicName>
is not treated at all, even though this species is certainly present in
<collectingCountry id="B9ABDAD0FFFABA3AFEAEFBCEFE48FB24" box="[371,453,1149,1173]" name="Taiwan" pageId="27" pageNumber="918">Taiwan</collectingCountry>
and
<collectingCountry id="B9ABDAD0FFFABA3AFDDFFBCDFD05FB27" box="[514,648,1150,1174]" name="Hong Kong" pageId="27" pageNumber="918">Hong Kong</collectingCountry>
, whereas
<taxonomicName id="06BCE1C3FFFABA3AFFAFFB2DFF6EFB04" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[114,227,1182,1205]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="918" phylum="Arthropoda" rank="species" species="sinense">
<emphasis id="F3C84652FFFABA3AFFAFFB2DFF6EFB04" box="[114,227,1182,1205]" italics="true" pageId="27" pageNumber="918">O. sinense</emphasis>
</taxonomicName>
is treated as a species of
<taxonomicName id="06BCE1C3FFFABA3AFE22FB2DFDFEFB04" authorityName="Serene &amp; Soh" authorityYear="1970" box="[511,627,1182,1205]" class="Malacostraca" family="Sesarmidae" genus="Sesarmops" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="918" phylum="Arthropoda" rank="genus">
<emphasis id="F3C84652FFFABA3AFE22FB2DFDFEFB04" box="[511,627,1182,1205]" italics="true" pageId="27" pageNumber="918">Sesarmops</emphasis>
</taxonomicName>
(see
<bibRefCitation id="A52DE7B1FFFABA3AFD71FB2EFEAFFB64" author="Dai &amp; Yang" firstAuthor="Dai" page="539" pageId="27" pageNumber="918" pagination="1 - 74" refId="ref68539" refString="Dai AY &amp; Yang SL (1991) Crabs of the China Seas. China Ocean Press, Beijing and Berlin, xxi + 681 pp., pls. 1 - 74." type="book chapter" year="1991">Dai &amp; Yang, 1991: 539</bibRefCitation>
, text-fig. 277(2), pl. 69(6); with their figures of the carapace and G1 leaving no doubt of the identity of this species).
</paragraph>
</subSubSection>
</treatment>
</document>