563 lines
93 KiB
XML
563 lines
93 KiB
XML
<document id="9B6BCB2D01E200B958BDFA7BF8317169" ID-CLB-Dataset="55667" ID-DOI="10.26107/RBZ-2020-0097" ID-GBIF-Dataset="6cb155ce-8b9f-48ce-8e6c-b0dc00f8b1cc" ID-ISSN="2345-7600" ID-Zenodo-Dep="5351295" ID-ZooBank="815E4670-B063-4FD8-B31E-3AD89B3A7942" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1630205841501" checkinUser="diego" docAuthor="Schubart, Christoph D. & Ng, Peter K. L." docDate="2020" docId="49152B56FFDEBA62FF7CFE2EFE87F84C" docLanguage="en" docName="RafflesBZool.68.891-994.pdf" docOrigin="Raffles Bulletin of Zoology 68" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Contusarma bocourti" docType="treatment" docVersion="11" lastPageNumber="958" masterDocId="B52C532EFFE1BA21FFDDFFB3FF8DFFB1" masterDocTitle="Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species" masterLastPageNumber="994" masterPageNumber="891" pageNumber="954" updateTime="1699203722016" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="71F43B66FEB02F0AE56BC972DB9EFAC8">Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species</mods:title>
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<mods:namePart id="5D485812E61BECE6CA259C8D8A5BF6D7">Schubart, Christoph D.</mods:namePart>
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<mods:affiliation id="A5A49165FCE9E33DD5FD8C75EA520CC7">Zoology & Evolution, University of Regensburg, 93040 Regensburg, Germany</mods:affiliation>
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<mods:namePart id="4B45D78431306940A7283E53BD45C5F0">Ng, Peter K. L.</mods:namePart>
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<mods:affiliation id="DFD14E10EBFBB3575F89D99393F43CF9">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore</mods:affiliation>
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<mods:nameIdentifier id="2762534501E37DC37A8C11272B33DC27" type="email">peterng@nus.edu.sg</mods:nameIdentifier>
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<taxonomicName id="06BCE1C3FFDEBA1EFF7CFE2EFD4DFE04" ID-CoL="84GS6" authority="(A. Milne-Edwards, 1869)" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1869" box="[161,704,413,437]" class="Malacostraca" family="Sesarmidae" genus="Contusarma" kingdom="Animalia" order="Decapoda" pageId="63" pageNumber="954" phylum="Arthropoda" rank="species" species="bocourti">
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<emphasis id="F3C84652FFDEBA1EFF7CFE2EFE03FE04" bold="true" box="[161,398,413,437]" italics="true" pageId="63" pageNumber="954">Contusarma bocourti</emphasis>
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(A.
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<bibRefCitation id="A52DE7B1FFDEBA1EFE1FFE2EFD37FE04" author="Milne-Edwards" box="[450,698,413,437]" firstAuthor="Milne-Edwards" pageId="63" pageNumber="954" pagination="25 - 31" refId="ref72689" refString="Milne-Edwards A (1869) Notes sur quelques nouvelles especes du genre Sesarma (Say). Nouvelle Archives du Museum d'Histoire Naturelle, Paris, 5: 25 - 31." type="journal article" year="1869">Milne-Edwards, 1869</bibRefCitation>
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)
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<subSubSection id="89A6C9CBFFDEBA1EFF5BFE0EFD51FE64" box="[134,732,445,469]" pageId="63" pageNumber="954" type="description">
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<paragraph id="C1039A40FFDEBA1EFF5BFE0EFD51FE64" blockId="63.[134,732,413,469]" box="[134,732,445,469]" pageId="63" pageNumber="954">
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(
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<figureCitation id="598786C5FFDEBA1EFF53FE0EFED4FE64" box="[142,345,445,469]" captionStart="Fig" captionStartId="42.[136,171,1862,1884]" captionTargetBox="[136,1474,233,1847]" captionTargetId="figure-7@42.[136,1474,233,1847]" captionTargetPageId="42" captionText="Fig. 23. A, E, Contusarma bocourti, lectotype male (24.0 × 27.2 mm) (MNHN D-10965), Bangkok, Thailand; B, F, C. bocourti, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand; C, C. bocourti, female (21.1 × 19.0 mm) (ZRC 2019.1114), Bangkok, Thailand; D, H, C. cheirogonum, neotype male (24.5 × 21.5 mm) (ZRC 1995.225), Bako National Park, Sarawak; G, C. cheirogonum, male (23.0 × 20.7 mm) (ZRC 2000.2018), Pulau Ubin, Singapore. A–D, overall habitus; E–H, anterior half of carapace." figureDoi="http://doi.org/10.5281/zenodo.5351341" httpUri="https://zenodo.org/record/5351341/files/figure.png" pageId="63" pageNumber="954">Figs. 23A–C, E, F</figureCitation>
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,
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<figureCitation id="598786C5FFDEBA1EFEBBFE0EFE1AFE64" box="[358,407,445,469]" captionStart="Fig" captionStartId="46.[136,171,1913,1935]" captionTargetBox="[147,1463,189,1899]" captionTargetId="figure-8@46.[147,1463,189,1899]" captionText="Fig. 25. Frontal view of cephalothorax of Pseudosesarma. A, P. edwardsii, lectotype male (17.5 × 16.1 mm) (RMNH-D17a), Mergui Archipelago; B, P. edwardsii, male (13.4 × 11.9 mm) (ZRC 1965.8.2.81), Pulau Pawai, Singapore; C, P. crassimanum, male (15.9 × 14.5 mm) (RMNH-D23313), Singapore; D, P. anteactum, holotype male (16.7 × 14.7 mm) (ZRC 2016.0602), Sri Lanka; E, P. glabrum, holotype male (13.8 × 12.0 mm) (CUSAT 2016-1), Kerala, India; F, P. brehieri, holotype male (17.4 × 15.7 mm) (ZRC 2016.0593), Myanmar; G, P. boulengeri, lectotype male (26.9 × 23.5 mm) (NHM 1919.11.14.1), Basra, Iraq; H, Contusarma bocourti, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand; I, Contusarma cheirogonum, neotype male (24.5 × 21.5 mm) (ZRC 1995.225), Bako National Park, Sarawak; J, Miersarma granosimanum, lectotype male (16.9 × 14.8 mm) (NHM 1880.6), Borneo." figureDoi="http://doi.org/10.5281/zenodo.6519007" httpUri="https://zenodo.org/record/6519007/files/figure.png" pageId="63" pageNumber="954">25H</figureCitation>
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,
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<figureCitation id="598786C5FFDEBA1EFE79FE0EFE32FE64" box="[420,447,445,469]" captionStart="Fig" captionStartId="50.[136,171,1012,1034]" captionTargetBox="[136,1474,189,997]" captionTargetId="figure-499@50.[136,1474,189,997]" captionTargetPageId="50" captionText="Fig. 28. Chela of Contusarma. A, C. bocourti, lectotype male (24.0 × 27.2 mm) (MNHN D-10965), Bangkok, Thailand; B, C, C. bocourti, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand; D, C. cheirogonum, neotype male (24.5 × 21.5 mm) (ZRC 1995.225), Bako National Park, Sarawak. A, B, D, outer view; C, ventro-marginal view." figureDoi="http://doi.org/10.5281/zenodo.5351347" httpUri="https://zenodo.org/record/5351347/files/figure.png" pageId="63" pageNumber="954">28</figureCitation>
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A–C,
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<figureCitation id="598786C5FFDEBA1EFDDDFE0EFD96FE64" box="[512,539,445,469]" captionStart="Fig" captionStartId="54.[136,171,1949,1971]" captionTargetBox="[169,1440,189,1933]" captionTargetId="figure-8@54.[169,1440,189,1933]" captionTargetPageId="54" captionText="Fig. 31. Male anterior thoracic sternum and pleon. A, Contusarma bocourti, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand; B, Contusarma cheirogonum, male (15.3 × 14.0 mm) (ZRC 2011.0924), Pulau Tioman, Malaysia; C, Contusarma cheirogonum, neotype male (24.5 × 21.5 mm) (ZRC 1995.225), Bako National Park, Sarawak; D, Miersarma granosimanum, lectotype male (16.9 × 14.8 mm) (NHM 1880.6), Borneo; E, Miersarma granosimanum, male (22.5 × 19.2 mm) (ZRC 1965.7.29.164), Sedili River, Johor, Malaysia; F, Bresedium laevimanum, lectotype male (20.0 × 17.6 mm) (MNHG), Borneo; G, Bresedium laevimanum, male (26.3 × 23.7 mm) (NHM) (lectotype of Sesarma sediliensis), Johor, Malaysia; H, Bresedium laevimanum, male (21.2 × 18.2 mm) (ZRC 1965.7.29.121), Johor, Malaysia." figureDoi="http://doi.org/10.5281/zenodo.5351351" httpUri="https://zenodo.org/record/5351351/files/figure.png" pageId="63" pageNumber="954">31</figureCitation>
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A,
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<figureCitation id="598786C5FFDEBA1EFDE0FE0EFDD7FE64" box="[573,602,445,469]" captionStart="Fig" captionStartId="65.[113,148,1520,1542]" captionTargetBox="[114,1451,192,1507]" captionTargetId="figure-15@65.[93,1473,173,1513]" captionTargetPageId="65" captionText="Fig. 38. Contusarma bocourti, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand. A, anterior thoracic sternites 1–4; B, left third maxilliped (denuded); C, male pleon; D, left G1 (ventral view, denuded); E, left G1 (dorsa view); F, left distal part of G1 (ventral view, denuded); G, left distal part of G1 (dorsal view, denuded); H, left distal part of G1 (mesial view). Scales: A, C = 5.0 mm; B = 2.5 mm; D, E = 1.0 mm; F–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5351363" httpUri="https://zenodo.org/record/5351363/files/figure.png" pageId="63" pageNumber="954">38</figureCitation>
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,
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<figureCitation id="598786C5FFDEBA1EFDBAFE0EFD0EFE64" box="[615,643,445,469]" captionStart="Fig" captionStartId="74.[136,171,1898,1920]" captionTargetBox="[136,1474,211,1884]" captionTargetId="figure-7@74.[136,1474,211,1884]" captionTargetPageId="74" captionText="Fig. 44. Vulva. A, Pseudosesarma edwardsii, female (16.5 × 15.3 mm) (ZRC 2016.0608), Malaysia; B, Pseudosesarma crassimanum, female (15.9 × 13.8 mm) (ZRC 2000.1768), Malaysia; C, Pseudosesarma crassimanum, female (18.0 × 15.7 mm) (ZRC 2017.1045), Thailand; D, Pseudosesarma anteactum, paratype female (17.5 × 15.4 mm) (ZRC 2016.0603), Sri Lanka; E, Pseudosesarma brehieri, paratype female (14.9 × 13.4 mm) (ZRC 2016.0594), Myanmar; F, Pseudosesarma boulengeri, female (24.2 × 20.0 mm) (ZRC 2014.0335), Iran; G, Contusarma bocourti, female (21.1 × 19.0 mm) (ZRC 2019.1114), Thailand; H, Contusarma cheirogonum, female (26.6 × 23.0 mm) (ZRC 1964.9.28.127), Sarawak; I, Miersarma granosimanum, female (20.1 × 16.4 mm) (ZRC 1965.7.29.166), Malaysia; J, Manarma moeschii, female (19.3 × 17.3 mm) (ZRC 2017.0170), Thailand." figureDoi="http://doi.org/10.5281/zenodo.5351375" httpUri="https://zenodo.org/record/5351375/files/figure.png" pageId="63" pageNumber="954">44</figureCitation>
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G,
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<figureCitation id="598786C5FFDEBA1EFD78FE0EFD4DFE64" box="[677,704,445,469]" captionStart="Fig" captionStartId="89.[113,148,1854,1876]" captionTargetBox="[112,1450,344,1838]" captionTargetId="figure-14@89.[112,1450,344,1838]" captionTargetPageId="89" captionText="Fig. 57. Colour in life. A, Contusarma bocourti, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand [photograph: Phaibul Naiyanetr]; B–D, C. cheirogonum, male (26.3 × 23.0 mm) (ZRC 1995.0226), Bako, Sarawak [photograph: PKL Ng]; E, G, C. cheirogonum, male, Tioman, Peninsular Malaysia, specimen not collected [photograph: Chua Ee Kiam]; F, C. cheirogonum, Tioman, Peninsular Malaysia, female (specimen not collected); H, C. cheirogonum, specimen not collected, Bako, Sarawak [photograph: Jongkar Grinang]." figureDoi="http://doi.org/10.5281/zenodo.5351401" httpUri="https://zenodo.org/record/5351401/files/figure.png" pageId="63" pageNumber="954">57</figureCitation>
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A)
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<treatmentCitation id="401DBC51FFDEBA1EFFACFE4FFDCCFDA0" author="Milne-Edwards A" box="[113,577,508,530]" page="28" pageId="63" pageNumber="954" year="1869">
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<taxonomicName id="06BCE1C3FFDEBA1EFFACFE4FFDCCFDA0" ID-CoL="7Z294" authority="A. Milne-Edwards, 1869: 28" authorityName="A. Milne-Edwards" authorityPageNumber="28" authorityYear="1869" box="[113,577,508,530]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="63" pageNumber="954" phylum="Arthropoda" rank="species" species="bocourti">
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<emphasis id="F3C84652FFDEBA1EFFACFE4FFE90FDA0" box="[113,285,508,529]" italics="true" pageId="63" pageNumber="954">Sesarma bocourti</emphasis>
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A.
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<bibRefCitation id="A52DE7B1FFDEBA1EFE99FE4FFDCCFDA0" author="Milne-Edwards" box="[324,577,508,530]" firstAuthor="Milne-Edwards" page="28" pageId="63" pageNumber="954" pagination="25 - 31" refId="ref72689" refString="Milne-Edwards A (1869) Notes sur quelques nouvelles especes du genre Sesarma (Say). Nouvelle Archives du Museum d'Histoire Naturelle, Paris, 5: 25 - 31." type="journal article" year="1869">Milne-Edwards, 1869: 28</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</paragraph>
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<paragraph id="C1039A40FFDEBA1EFFACFDAAFD1CFD9E" blockId="63.[113,753,508,645]" box="[113,657,537,559]" pageId="63" pageNumber="954">
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<treatmentCitationGroup id="E1ACBD6EFFDEBA1EFFACFDAAFD1CFD9E" box="[113,657,537,559]" pageId="63" pageNumber="954">
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<taxonomicName id="06BCE1C3FFDEBA1EFFACFDAAFE0AFD9F" ID-CoL="7Z294" authorityName="A. Milne-Edwards" authorityYear="1869" box="[113,391,537,558]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="63" pageNumber="954" phylum="Arthropoda" rank="species" species="bocourti" subGenus="Sesarma">
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<emphasis id="F3C84652FFDEBA1EFFACFDAAFF49FD9F" box="[113,196,537,558]" italics="true" pageId="63" pageNumber="954">Sesarma</emphasis>
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(
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<emphasis id="F3C84652FFDEBA1EFF09FDAAFEAAFD9F" box="[212,295,537,558]" italics="true" pageId="63" pageNumber="954">Sesarma</emphasis>
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)
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<emphasis id="F3C84652FFDEBA1EFEEBFDAAFE0AFD9F" box="[310,391,537,558]" italics="true" pageId="63" pageNumber="954">bocourti</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="401DBC51FFDEBA1EFE7EFDAAFDC0FD9E" author="Tesch JJ" box="[419,589,537,559]" page="135" pageId="63" pageNumber="954" year="1917">
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<bibRefCitation id="A52DE7B1FFDEBA1EFE7EFDAAFDC0FD9E" author="Tesch" box="[419,589,537,559]" firstAuthor="Tesch" page="135" pageId="63" pageNumber="954" pagination="127 - 260" refId="ref78063" refString="Tesch JJ (1917) Synopsis of the genera Sesarma, Metasesarma, Sarmatium and Clistocoeloma, with a key to the determination of the IndoPacific species. Zoologische Mededelingen, 3: 127 - 260, pls. 15 - 17." type="journal article" year="1917">Tesch, 1917: 135</bibRefCitation>
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</treatmentCitation>
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(part).
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</treatmentCitationGroup>
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</paragraph>
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|
||
<taxonomicName id="06BCE1C3FFDEBA1EFFACFD86FED2FDFB" ID-CoL="7YLX6" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1869" box="[113,351,565,586]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="63" pageNumber="954" phylum="Arthropoda" rank="species" species="bocourti">
|
||
<emphasis id="F3C84652FFDEBA1EFFACFD86FED2FDFB" box="[113,351,565,586]" italics="true" pageId="63" pageNumber="954">Pseudosesarma bocourti</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="401DBC51FFDEBA1EFEAAFD86FDC9FDFA" author="Naiyanetr P" box="[375,580,565,587]" page="101" pageId="63" pageNumber="954" year="1998">
|
||
<bibRefCitation id="A52DE7B1FFDEBA1EFEAAFD86FDC9FDFA" author="Naiyanetr" box="[375,580,565,587]" firstAuthor="Naiyanetr" page="101" pageId="63" pageNumber="954" refId="ref73221" refString="Naiyanetr P (1998) Checklist of Crustacean Fauna in Thailand (Decapoda, Stomatopoda, Anostraca, Myodocopa and Isopoda). Office of Environmental Policy and Planning, Bangkok, 161 pp." type="book" year="1998">Naiyanetr, 1998: 101</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="401DBC51FFDEBA1EFD92FD86FF22FDD6" author="Naiyanetr P" page="17" pageId="63" pageNumber="954" year="2007">
|
||
<bibRefCitation id="A52DE7B1FFDEBA1EFD92FD86FF22FDD6" author="Naiyanetr" firstAuthor="Naiyanetr" page="17" pageId="63" pageNumber="954" refId="ref73256" refString="Naiyanetr P (2007) Checklist of Crustacean Fauna in Thailand (Decapoda, Stomatopoda, Anostraca, Myodocopa and Isopoda). Office of Natural Resources and Environmental Policy and Planning, Bangkok, 196 pp." type="book" year="2007">Naiyanetr, 2007: 17</bibRefCitation>
|
||
</treatmentCitation>
|
||
(unnumbered fig.), 115;
|
||
<treatmentCitation id="401DBC51FFDEBA1EFE73FDE0FD01FDD6" author="Ng PKL, Guinot D & Davie PJF" box="[430,652,594,616]" page="222" pageId="63" pageNumber="954" year="2008">
|
||
<bibRefCitation id="A52DE7B1FFDEBA1EFE73FDE0FD05FDD6" author="Ng" box="[430,648,594,616]" etAl="et al." firstAuthor="Ng" page="222" pageId="63" pageNumber="954" pagination="1 - 286" refId="ref73754" refString="Ng PKL, Guinot D & Davie PJF (2008 a) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008" yearSuffix="a">Ng et al., 2008a: 222</bibRefCitation>
|
||
;
|
||
</treatmentCitation>
|
||
<treatmentCitation id="401DBC51FFDEBA1EFD48FDE0FEB2FD35" author="Hoang NK, Do VN, Nguyen TT & Nguyen TTH" page="75, 78" pageId="63" pageNumber="954" year="2012">
|
||
<bibRefCitation id="A52DE7B1FFDEBA1EFD48FDE0FE98FD34" author="Hoang" etAl="et al." firstAuthor="Hoang" page="75" pageId="63" pageNumber="954" pagination="72 - 80" refId="ref70272" refString="Hoang NK, Do VN, Nguyen TT & Nguyen TTH (2012) Preliminary data of three zoobenthos groups (Brachyura, Gastropoda and Bivalviva) found in the mangrove forest ecosystems of the Tran De Estuary, Soc Trang Province. Journal of Science of HNUE (Hanoi National University of Education), Chemical and Biological Sciences, 57 (8): 72 - 80." type="journal article" year="2012">Hoang et al., 2012: 75</bibRefCitation>
|
||
, 78.
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="89A6C9CBFFDEBA1EFFACFD1EFD07FC14" pageId="63" pageNumber="954" type="materials_examined">
|
||
<paragraph id="C1039A40FFDEBA1EFFACFD1EFD07FC14" blockId="63.[113,754,685,934]" pageId="63" pageNumber="954">
|
||
<emphasis id="F3C84652FFDEBA1EFFACFD1EFEDBFD74" bold="true" box="[113,342,685,709]" pageId="63" pageNumber="954">Material examined</emphasis>
|
||
.
|
||
<materialsCitation id="71D4901DFFDEBA1EFEB5FD1DFD60FD54" ID-GBIF-Occurrence="3763237464" collectionCode="MNHN" location="Thailand" pageId="63" pageNumber="954" specimenCount="1" stateProvince="Krung Thep Mahanakhon" typeStatus="lectotype">
|
||
<typeStatus id="1E0724E2FFDEBA1EFEB5FD1DFE5AFD77" box="[360,471,686,710]" pageId="63" pageNumber="954" type="lectotype">Lectotype</typeStatus>
|
||
(here designated): male (24.0 ×
|
||
<quantity id="064437A5FFDEBA1EFF16FD7EFEA2FD54" box="[203,303,717,741]" metricMagnitude="-2" metricUnit="m" metricValue="2.7199999999999998" pageId="63" pageNumber="954" unit="mm" value="27.2">27.2 mm</quantity>
|
||
) (
|
||
<collectionCode id="A7AD0285FFDEBA1EFE9BFD7EFE14FD54" box="[326,409,717,741]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="63" pageNumber="954">MNHN</collectionCode>
|
||
D-10965),
|
||
<collectingRegion id="037854A2FFDEBA1EFDC6FD7EFDF3FD54" box="[539,638,717,741]" country="Thailand" name="Krung Thep Mahanakhon" pageId="63" pageNumber="954">Bangkok</collectingRegion>
|
||
,
|
||
<location id="C463CC9BFFDEBA1EFD56FD7EFD60FD54" LSID="urn:lsid:plazi:treatment:49152B56FFDEBA62FF7CFE2EFE87F84C:C463CC9BFFDEBA1EFD56FD7EFD60FD54" box="[651,749,717,741]" name="Thailand" pageId="63" pageNumber="954" stateProvince="Krung Thep Mahanakhon">Thailand</location>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="71D4901DFFDEBA1EFFACFD5EFEFEFC94" ID-GBIF-Occurrence="3763237447" collectingDate="1998-08-01" collectionCode="MNHN" collectorName="Pongsathron" country="Thailand" county="Silom" location="Hotel Mae-Noon" municipality="Chao Phraya" pageId="63" pageNumber="954" specimenCode="ZRC 2000.0952, ZRC 2019.1114" specimenCount="1" specimenCount-female="1" stateProvince="Krung Thep Mahanakhon" typeStatus="paralectotype">
|
||
<typeStatus id="1E0724E2FFDEBA1EFFACFD5EFE8BFCB4" box="[113,262,749,773]" pageId="63" pageNumber="954" type="paralectotype">Paralectotype</typeStatus>
|
||
: male (19.1 ×
|
||
<quantity id="064437A5FFDEBA1EFE77FD5EFD80FCB4" box="[426,525,749,773]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" pageId="63" pageNumber="954" unit="mm" value="21.5">21.5 mm</quantity>
|
||
) (
|
||
<collectionCode id="A7AD0285FFDEBA1EFDF8FD5EFDF5FCB4" box="[549,632,749,773]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="63" pageNumber="954">MNHN</collectionCode>
|
||
D-10965), same data as lectotype.
|
||
</materialsCitation>
|
||
<materialsCitation id="71D4901DFFDEBA1EFEA0FCBDFF23FC34" ID-GBIF-Occurrence="3763237477" collectingDate="1998-08-01" collectionCode="MNHN" collectorName="Pongsathron" country="Thailand" county="Silom" location="Hotel Mae-Noon" municipality="Chao Phraya" pageId="63" pageNumber="954" specimenCode="ZRC 2000.0952, ZRC 2019.1114" specimenCount="1" specimenCount-female="1" stateProvince="Krung Thep Mahanakhon" typeStatus="paralectotype">
|
||
<collectingCountry id="B9ABDAD0FFDEBA1EFEA0FCBDFD8AFC97" box="[381,519,782,806]" name="Thailand" pageId="63" pageNumber="954">THAILAND</collectingCountry>
|
||
– male (25.3 ×
|
||
<quantity id="064437A5FFDEBA1EFD1CFCBEFF11FCF4" metricMagnitude="-2" metricUnit="m" metricValue="2.33" pageId="63" pageNumber="954" unit="mm" value="23.3">23.3 mm</quantity>
|
||
) (
|
||
<specimenCode id="911A323BFFDEBA1EFF68FC9EFEE1FCF4" box="[181,364,813,837]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="63" pageNumber="954">ZRC 2000.0952</specimenCode>
|
||
),
|
||
<collectingCounty id="2862E2CCFFDEBA1EFE5CFC9EFE49FCF4" box="[385,452,813,837]" pageId="63" pageNumber="954">Silom</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="2167003AFFDEBA1EFE0FFC9EFDD2FCF4" box="[466,607,813,837]" pageId="63" pageNumber="954">Chao Phraya</collectingMunicipality>
|
||
at cemetery,
|
||
<location id="C463CC9BFFDEBA1EFFACFCFEFEBFFCD4" LSID="urn:lsid:plazi:treatment:49152B56FFDEBA62FF7CFE2EFE87F84C:C463CC9BFFDEBA1EFFACFCFEFEBFFCD4" box="[113,306,845,869]" country="Thailand" county="Silom" municipality="Chao Phraya" name="Hotel Mae-Noon" pageId="63" pageNumber="954" stateProvince="Krung Thep Mahanakhon">Hotel Mae-Noon</location>
|
||
,
|
||
<collectingRegion id="037854A2FFDEBA1EFE9DFCFEFE28FCD4" box="[320,421,845,869]" country="Thailand" name="Krung Thep Mahanakhon" pageId="63" pageNumber="954">Bangkok</collectingRegion>
|
||
, coll.
|
||
<collectorName id="6C49FF96FFDEBA1EFE32FCFEFDF6FCD4" box="[495,635,845,869]" pageId="63" pageNumber="954">Pongsathron</collectorName>
|
||
,
|
||
<date id="B502BC80FFDEBA1EFD57FCFEFF27FC34" pageId="63" pageNumber="954" value="1998-08-01">
|
||
<collectingDate id="A5464568FFDEBA1EFD57FCFEFF27FC34" pageId="63" pageNumber="954" value="1998-08-01">1 August 1998</collectingDate>
|
||
</date>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="71D4901DFFDEBA1EFF68FCDEFD0BFC14" ID-GBIF-Occurrence="3763237526" collectingDate="1998-08-01" collectionCode="MNHN" collectorName="Pongsathron" country="Thailand" county="Silom" location="Hotel Mae-Noon" municipality="Chao Phraya" pageId="63" pageNumber="954" specimenCode="ZRC 2000.0952, ZRC 2019.1114" specimenCount="1" specimenCount-female="1" stateProvince="Krung Thep Mahanakhon" typeStatus="paralectotype">
|
||
<specimenCount id="D7BA51C9FFDEBA1EFF68FCDEFE9FFC34" box="[181,274,877,901]" pageId="63" pageNumber="954" type="female">1 female</specimenCount>
|
||
(
|
||
<specimenCode id="911A323BFFDEBA1EFEC2FCDEFE5FFC34" box="[287,466,877,901]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="63" pageNumber="954">ZRC 2019.1114</specimenCode>
|
||
),
|
||
<collectingRegion id="037854A2FFDEBA1EFE3EFCDEFDCBFC34" box="[483,582,877,901]" country="Thailand" name="Krung Thep Mahanakhon" pageId="63" pageNumber="954">Bangkok</collectingRegion>
|
||
, Thailand, from aquarium trade, via
|
||
<collectorName id="6C49FF96FFDEBA1EFE90FC3EFE5DFC14" box="[333,464,909,933]" pageId="63" pageNumber="954">C. Lukhaup</collectorName>
|
||
,
|
||
<date id="B502BC80FFDEBA1EFE01FC3EFD0BFC14" box="[476,646,909,934]" pageId="63" pageNumber="954" value="2007-01-09">
|
||
<collectingDate id="A5464568FFDEBA1EFE01FC3EFD0BFC14" box="[476,646,909,934]" pageId="63" pageNumber="954" value="2007-01-09">9 January 2007</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="89A6C9CBFFDEBA1EFFACFC7EFE2FFAF4" pageId="63" pageNumber="954" type="diagnosis">
|
||
<paragraph id="C1039A40FFDEBA1EFFACFC7EFF3BFB14" blockId="63.[113,753,973,1189]" pageId="63" pageNumber="954">
|
||
<emphasis id="F3C84652FFDEBA1EFFACFC7EFF52FC54" bold="true" box="[113,223,973,997]" pageId="63" pageNumber="954">Diagnosis</emphasis>
|
||
. Anterior dorsal carapace regions with relatively dense short black setae; outer surface of adult chela covered with large flattened granules, those on margins large, sharp; margins of ambulatory segments with dense short setae; male pleon relatively transversely more narrow; G1 relatively stouter, chitinous distal part gently curved, proportionately wider.
|
||
</paragraph>
|
||
<paragraph id="C1039A40FFDEBA1EFFACFB7EFE2FFAF4" blockId="63.[113,753,1229,1349]" pageId="63" pageNumber="954">
|
||
<emphasis id="F3C84652FFDEBA1EFFACFB7EFF4EFB54" bold="true" box="[113,195,1229,1253]" pageId="63" pageNumber="954">Colour</emphasis>
|
||
. Carapace relatively dark, gastric regions almost black; distal margins of ambulatory merus, carpus, and propodus with darker bands (
|
||
<figureCitation id="598786C5FFDEBA1EFE71FABEFD80FA94" box="[428,525,1293,1317]" captionStart="Fig" captionStartId="89.[113,148,1854,1876]" captionTargetBox="[112,1450,344,1838]" captionTargetId="figure-14@89.[112,1450,344,1838]" captionTargetPageId="89" captionText="Fig. 57. Colour in life. A, Contusarma bocourti, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand [photograph: Phaibul Naiyanetr]; B–D, C. cheirogonum, male (26.3 × 23.0 mm) (ZRC 1995.0226), Bako, Sarawak [photograph: PKL Ng]; E, G, C. cheirogonum, male, Tioman, Peninsular Malaysia, specimen not collected [photograph: Chua Ee Kiam]; F, C. cheirogonum, Tioman, Peninsular Malaysia, female (specimen not collected); H, C. cheirogonum, specimen not collected, Bako, Sarawak [photograph: Jongkar Grinang]." figureDoi="http://doi.org/10.5281/zenodo.5351401" httpUri="https://zenodo.org/record/5351401/files/figure.png" pageId="63" pageNumber="954">Fig. 57A</figureCitation>
|
||
) (see also
|
||
<bibRefCitation id="A52DE7B1FFDEBA1EFD5CFABEFF59FAF4" author="Naiyanetr" firstAuthor="Naiyanetr" page="17" pageId="63" pageNumber="954" refId="ref73256" refString="Naiyanetr P (2007) Checklist of Crustacean Fauna in Thailand (Decapoda, Stomatopoda, Anostraca, Myodocopa and Isopoda). Office of Natural Resources and Environmental Policy and Planning, Bangkok, 196 pp." type="book" year="2007">Naiyanetr, 2007: 17</bibRefCitation>
|
||
, unnumbered fig.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="89A6C9CBFFDEBA60FFACFADEFB4BF7BA" lastPageId="65" lastPageNumber="956" pageId="63" pageNumber="954" type="discussion">
|
||
<paragraph id="C1039A40FFDEBA1EFFACFADEFBA8FD04" blockId="63.[113,753,1389,2117]" lastBlockId="63.[813,1452,189,693]" pageId="63" pageNumber="954">
|
||
<emphasis id="F3C84652FFDEBA1EFFACFADEFF57FA34" bold="true" box="[113,218,1389,1413]" pageId="63" pageNumber="954">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="06BCE1C3FFDEBA1EFF31FADDFE22FA34" authorityName="A. Milne-Edwards" authorityYear="1869" box="[236,431,1389,1413]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="63" pageNumber="954" phylum="Arthropoda" rank="species" species="bocourti">
|
||
<emphasis id="F3C84652FFDEBA1EFF31FADDFE22FA34" box="[236,431,1389,1413]" italics="true" pageId="63" pageNumber="954">Sesarma bocourti</emphasis>
|
||
</taxonomicName>
|
||
was described briefly by A.
|
||
<bibRefCitation id="A52DE7B1FFDEBA1EFFAFFA3EFEFAFA14" author="Milne-Edwards" box="[114,375,1421,1445]" firstAuthor="Milne-Edwards" pageId="63" pageNumber="954" pagination="25 - 31" refId="ref72689" refString="Milne-Edwards A (1869) Notes sur quelques nouvelles especes du genre Sesarma (Say). Nouvelle Archives du Museum d'Histoire Naturelle, Paris, 5: 25 - 31." type="journal article" year="1869">Milne-Edwards (1869)</bibRefCitation>
|
||
from an unspecified number of specimens from
|
||
<collectingRegion id="037854A2FFDEBA1EFEC6FA1EFEF1FA74" box="[283,380,1453,1477]" country="Thailand" name="Krung Thep Mahanakhon" pageId="63" pageNumber="954">Bangkok</collectingRegion>
|
||
,
|
||
<collectingCountry id="B9ABDAD0FFDEBA1EFE59FA1EFE69FA74" box="[388,484,1453,1477]" name="Thailand" pageId="63" pageNumber="954">Thailand</collectingCountry>
|
||
. The extant specimens (all dried) are therefore
|
||
<typeStatus id="1E0724E2FFDEBA1EFE9AFA7DFE25FA57" box="[327,424,1486,1510]" pageId="63" pageNumber="954" type="syntype">syntypes</typeStatus>
|
||
. The largest male (24.0 ×
|
||
<quantity id="064437A5FFDEBA1EFD1CFA7EFF10F9B4" metricMagnitude="-2" metricUnit="m" metricValue="2.7199999999999998" pageId="63" pageNumber="954" unit="mm" value="27.2">27.2 mm</quantity>
|
||
, MNHN D-10965) is here selected as the
|
||
<typeStatus id="1E0724E2FFDEBA1EFD93FA5EFD22F9B4" box="[590,687,1517,1541]" pageId="63" pageNumber="954" type="lectotype">lectotype</typeStatus>
|
||
of the species (
|
||
<figureCitation id="598786C5FFDEBA1EFF12F9BEFEB9F994" box="[207,308,1549,1573]" captionStart="Fig" captionStartId="42.[136,171,1862,1884]" captionTargetBox="[136,1474,233,1847]" captionTargetId="figure-7@42.[136,1474,233,1847]" captionTargetPageId="42" captionText="Fig. 23. A, E, Contusarma bocourti, lectotype male (24.0 × 27.2 mm) (MNHN D-10965), Bangkok, Thailand; B, F, C. bocourti, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand; C, C. bocourti, female (21.1 × 19.0 mm) (ZRC 2019.1114), Bangkok, Thailand; D, H, C. cheirogonum, neotype male (24.5 × 21.5 mm) (ZRC 1995.225), Bako National Park, Sarawak; G, C. cheirogonum, male (23.0 × 20.7 mm) (ZRC 2000.2018), Pulau Ubin, Singapore. A–D, overall habitus; E–H, anterior half of carapace." figureDoi="http://doi.org/10.5281/zenodo.5351341" httpUri="https://zenodo.org/record/5351341/files/figure.png" pageId="63" pageNumber="954">Fig. 23A</figureCitation>
|
||
).
|
||
<taxonomicName id="06BCE1C3FFDEBA1EFE95F9BDFF25F9F4" authority="Targioni Tozzetti, 1877" authorityName="Targioni Tozzetti" authorityYear="1877" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="63" pageNumber="954" phylum="Arthropoda" rank="species" species="cheirogona">
|
||
<emphasis id="F3C84652FFDEBA1EFE95F9BDFDA8F994" box="[328,549,1549,1573]" italics="true" pageId="63" pageNumber="954">Sesarma cheirogona</emphasis>
|
||
<bibRefCitation id="A52DE7B1FFDEBA1EFDF3F9BEFF25F9F4" author="Targioni Tozzetti" firstAuthor="Targioni Tozzetti" pageId="63" pageNumber="954" pagination="1 - 257" refId="ref77991" refString="Targioni Tozzetti A (1877) Crostacei brachyuri e anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R Istituto di Studi Superiori Pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1: i - xxix + 1 - 257, pls. 1 - 12." type="journal article" year="1877">Targioni Tozzetti, 1877</bibRefCitation>
|
||
</taxonomicName>
|
||
, was described on the basis of a single male (22.0 × 20.0 mm) supposedly collected from Yokohama in
|
||
<collectingCountry id="B9ABDAD0FFDEBA1EFDBFF9FDFD2FF9D4" box="[610,674,1614,1637]" name="Japan" pageId="63" pageNumber="954">Japan</collectingCountry>
|
||
.
|
||
<bibRefCitation id="A52DE7B1FFDEBA1EFD71F9FEFF7CF934" author="Nobili" firstAuthor="Nobili" page="507" pageId="63" pageNumber="954" pagination="473 - 523" refId="ref74323" refString="Nobili G (1900) Decapodi e stomatopodi Indo-Malesi. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 20 (40): 473 - 523, figs. 1 - 4." type="journal article" year="1900">Nobili (1900: 507)</bibRefCitation>
|
||
apparently examined the
|
||
<typeStatus id="1E0724E2FFDEBA1EFDD3F9DEFDE6F934" box="[526,619,1645,1669]" pageId="63" pageNumber="954" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="06BCE1C3FFDEBA1EFD49F9DDFF66F914" authorityName="Targioni Tozzetti" authorityYear="1877" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="63" pageNumber="954" phylum="Arthropoda" rank="species" species="cheirogona">
|
||
<emphasis id="F3C84652FFDEBA1EFD49F9DDFF66F914" italics="true" pageId="63" pageNumber="954">Sesarma cheirogona</emphasis>
|
||
</taxonomicName>
|
||
and commented that there were no significant differences from his specimens of
|
||
<taxonomicName id="06BCE1C3FFDEBA1EFE2FF91DFD3FF974" authorityName="A. Milne-Edwards" authorityYear="1869" box="[498,690,1709,1733]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="63" pageNumber="954" phylum="Arthropoda" rank="species" species="bocourti">
|
||
<emphasis id="F3C84652FFDEBA1EFE2FF91DFD3FF974" box="[498,690,1709,1733]" italics="true" pageId="63" pageNumber="954">Sesarma bocourti</emphasis>
|
||
</taxonomicName>
|
||
from Sumatra collected by the Siboga Expedition and synonymised both species.
|
||
<bibRefCitation id="A52DE7B1FFDEBA1EFEDAF95EFE15F8B4" author="Tesch" box="[263,408,1773,1797]" firstAuthor="Tesch" pageId="63" pageNumber="954" pagination="127 - 260" refId="ref78063" refString="Tesch JJ (1917) Synopsis of the genera Sesarma, Metasesarma, Sarmatium and Clistocoeloma, with a key to the determination of the IndoPacific species. Zoologische Mededelingen, 3: 127 - 260, pls. 15 - 17." type="journal article" year="1917">Tesch (1917)</bibRefCitation>
|
||
agreed with the synonymy but noted there were
|
||
<specimenCount id="D7BA51C9FFDEBA1EFEE5F8BDFE29F897" box="[312,420,1806,1830]" pageId="63" pageNumber="954" type="generic">two types</specimenCount>
|
||
of male pleons for what had been called
|
||
<taxonomicName id="06BCE1C3FFDEBA1EFF27F89DFEF8F8F4" authorityName="A. Milne-Edwards" authorityYear="1869" box="[250,373,1837,1861]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="63" pageNumber="954" phylum="Arthropoda" rank="species" species="bocourti">
|
||
<emphasis id="F3C84652FFDEBA1EFF27F89DFEF8F8F4" box="[250,373,1837,1861]" italics="true" pageId="63" pageNumber="954">S. bocourti</emphasis>
|
||
</taxonomicName>
|
||
in Southeast Asia; one relatively more narrow (
|
||
<bibRefCitation id="A52DE7B1FFDEBA1EFEC8F8FEFE28F8D4" author="Tesch" box="[277,421,1869,1893]" firstAuthor="Tesch" pageId="63" pageNumber="954" pagination="127 - 260" refId="ref78063" refString="Tesch JJ (1917) Synopsis of the genera Sesarma, Metasesarma, Sarmatium and Clistocoeloma, with a key to the determination of the IndoPacific species. Zoologische Mededelingen, 3: 127 - 260, pls. 15 - 17." type="journal article" year="1917">Tesch, 1917</bibRefCitation>
|
||
: fig. 2a; from an unspecified location in Borneo and Deli, Sumatra) and one relatively wider (
|
||
<bibRefCitation id="A52DE7B1FFDEBA1EFF60F83EFEC8F814" author="Tesch" box="[189,325,1933,1957]" firstAuthor="Tesch" pageId="63" pageNumber="954" pagination="127 - 260" refId="ref78063" refString="Tesch JJ (1917) Synopsis of the genera Sesarma, Metasesarma, Sarmatium and Clistocoeloma, with a key to the determination of the IndoPacific species. Zoologische Mededelingen, 3: 127 - 260, pls. 15 - 17." type="journal article" year="1917">Tesch, 1917</bibRefCitation>
|
||
: fig. 2b; from Balikpapan, Borneo). He, however, recognised just one species as he commented that they were otherwise similar externally.
|
||
<bibRefCitation id="A52DE7B1FFDEBA1EFDEAF87EFD7DF854" author="Urita" box="[567,752,1997,2021]" firstAuthor="Urita" page="20" pageId="63" pageNumber="954" refId="ref78486" refString="Urita T (1926) A check list of Brachyura found in Kagoshima Pref., Japan. The Tsingtao Times, Tsingtao, 40 pp." type="book" year="1926">Urita (1926: 20)</bibRefCitation>
|
||
recorded
|
||
<taxonomicName id="06BCE1C3FFDEBA1EFF09F85DFECAF7B4" authorityName="A. Milne-Edwards" authorityYear="1869" box="[212,327,2029,2053]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="63" pageNumber="954" phylum="Arthropoda" rank="species" species="bocourti">
|
||
<emphasis id="F3C84652FFDEBA1EFF09F85DFECAF7B4" box="[212,327,2029,2053]" italics="true" pageId="63" pageNumber="954">S. bocourti</emphasis>
|
||
</taxonomicName>
|
||
from “near coast of
|
||
<collectingRegion id="037854A2FFDEBA1EFDC1F85EFD06F7B4" box="[540,651,2029,2053]" country="Japan" name="Kagoshima" pageId="63" pageNumber="954">Kagosima</collectingRegion>
|
||
, Satuma” in
|
||
<collectingCountry id="B9ABDAD0FFDEBA1EFF52F7BDFF43F794" box="[143,206,2062,2085]" name="Japan" pageId="63" pageNumber="954">Japan</collectingCountry>
|
||
, but no material was indicated, and no figure was provided so its identity cannot be confirmed.
|
||
<bibRefCitation id="A52DE7B1FFDEBA1EFDBBF79EFCD5FF64" author="Sakai" box="[614,856,189,2117]" firstAuthor="Sakai" page="685" pageId="63" pageNumber="954" pagination="1 - 6" refId="ref75618" refString="Sakai T (1939) Studies on the Crabs of Japan. IV. Brachygnatha, Brachyrhyncha. Yokendo Co., Ltd., Tokyo. Pp. 365 - 741 + 1 - 23 [= Index], figs. 1 - 129, pls. XLII - CXI, tables 1 - 6." type="book chapter" year="1939">Sakai (1939: 685</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="A52DE7B1FFDEBA1EFCBFFF0EFC43FF64" author="Sakai" box="[866,974,189,213]" firstAuthor="Sakai" page="661" pageId="63" pageNumber="954" pagination="1 - 251" refId="ref75757" refString="Sakai T (1976) Crabs of Japan and the Adjacent Seas. Kodansha Ltd., Tokyo. In three volumes. xxix + 773 pp. [English Text], 461 pp. [Japanese Text]. Pp. 1 - 16, pls. 1 - 251 [Plates volume]." type="book chapter" year="1976">1976: 661</bibRefCitation>
|
||
) listed the species for the Japanese fauna but noted that it was based on
|
||
<bibRefCitation id="A52DE7B1FFDEBA1EFB9BFF6EFAECFF44" author="Targioni Tozzetti's" box="[1094,1377,221,245]" firstAuthor="Targioni Tozzetti's" pageId="63" pageNumber="954" pagination="1 - 257" refId="ref77991" refString="Targioni Tozzetti A (1877) Crostacei brachyuri e anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R Istituto di Studi Superiori Pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1: i - xxix + 1 - 257, pls. 1 - 12." type="journal article" year="1877">Targioni Tozzetti’s (1877)</bibRefCitation>
|
||
record and he had no specimens.
|
||
<bibRefCitation id="A52DE7B1FFDEBA1EFB84FF4EFA85FEA4" author="Tweedie" box="[1113,1288,253,277]" firstAuthor="Tweedie" pageId="63" pageNumber="954" pagination="88 - 113" refId="ref78385" refString="Tweedie MWF (1940) New and interesting Malaysian species of Sesarma and Utica (Crustacea, Brachyura). Bulletin of the Raffles Museum, 16: 88 - 113, figs. 1 - 12, pl. 24." type="journal article" year="1940">Tweedie (1940)</bibRefCitation>
|
||
discussed this and commented that broader pleons are generally associated with smaller specimens and the differences were probably due to variation.
|
||
<bibRefCitation id="A52DE7B1FFDEBA1EFC34FEEEFB4DFEC4" author="Tweedie" box="[1001,1216,349,373]" firstAuthor="Tweedie" page="91" pageId="63" pageNumber="954" pagination="88 - 113" refId="ref78385" refString="Tweedie MWF (1940) New and interesting Malaysian species of Sesarma and Utica (Crustacea, Brachyura). Bulletin of the Raffles Museum, 16: 88 - 113, figs. 1 - 12, pl. 24." type="journal article" year="1940">Tweedie (1940: 91)</bibRefCitation>
|
||
also commented that “If
|
||
<taxonomicName id="06BCE1C3FFDEBA1EFC8FFECDFB7FFE24" authority="Targioni Tozzetti" authorityName="Targioni Tozzetti" authorityYear="1877" box="[850,1266,381,405]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="63" pageNumber="954" phylum="Arthropoda" rank="species" species="cheirogona">
|
||
<emphasis id="F3C84652FFDEBA1EFC8FFECDFBA2FE24" box="[850,1071,381,405]" italics="true" pageId="63" pageNumber="954">Sesarma cheirogona</emphasis>
|
||
Targioni Tozzetti
|
||
</taxonomicName>
|
||
from Yokohama (1877, p. 141) is really identical with
|
||
<taxonomicName id="06BCE1C3FFDEBA1EFADDFE2EFAEFFE04" authorityName="A. Milne-Edwards" authorityYear="1869" box="[1280,1378,413,437]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="63" pageNumber="954" phylum="Arthropoda" rank="species" species="bocourti">
|
||
<emphasis id="F3C84652FFDEBA1EFADDFE2EFAEFFE04" box="[1280,1378,413,437]" italics="true" pageId="63" pageNumber="954">bocourti</emphasis>
|
||
</taxonomicName>
|
||
, I am inclined to think that the specimen was wrongly localised. It was collected during the world cruise of the ‘Magenta’, which included visits to Borneo and Sumatra”. There have thus been no reliable confirmed records of this species north of
|
||
<collectingCountry id="B9ABDAD0FFDEBA1EFC96FD8EFC20FDE4" box="[843,941,573,597]" name="Thailand" pageId="63" pageNumber="954">Thailand</collectingCountry>
|
||
, despite intensive collecting and study of crabs in
|
||
<collectingCountry id="B9ABDAD0FFDEBA1EFC97FDEDFC07FDC4" box="[842,906,606,629]" name="Japan" pageId="63" pageNumber="954">Japan</collectingCountry>
|
||
. This supports Tweedie’s suggestion that the type locality is likely to have been from the Indo-Malayan region rather than Yokahama.
|
||
</paragraph>
|
||
<paragraph id="C1039A40FFDEBA1EFCF0FD6EFBCEFA44" blockId="63.[812,1452,733,1525]" pageId="63" pageNumber="954">
|
||
Examination of the present extensive collection has revealed differences between the specimens from
|
||
<collectingCountry id="B9ABDAD0FFDEBA1EFACEFD4EFAF5FCA4" box="[1299,1400,765,789]" name="Thailand" pageId="63" pageNumber="954">Thailand</collectingCountry>
|
||
and the rest of Southeast Asia (southern Peninsular
|
||
<collectingCountry id="B9ABDAD0FFDEBA1EFA9EFCAEFA25FC84" box="[1347,1448,797,821]" name="Malaysia" pageId="63" pageNumber="954">Malaysia</collectingCountry>
|
||
,
|
||
<collectingCountry id="B9ABDAD0FFDEBA1EFCF0FC8EFC11FCE4" box="[813,924,829,853]" name="Singapore" pageId="63" pageNumber="954">Singapore</collectingCountry>
|
||
, and
|
||
<collectingCountry id="B9ABDAD0FFDEBA1EFC0AFC8EFBB5FCE4" box="[983,1080,829,853]" name="Malaysia" pageId="63" pageNumber="954">Sarawak</collectingCountry>
|
||
). Most significantly, their G1s are slightly different, with those from
|
||
<collectingCountry id="B9ABDAD0FFDEBA1EFB43FCEEFB70FCC4" box="[1182,1277,861,885]" name="Thailand" pageId="63" pageNumber="954">Thailand</collectingCountry>
|
||
being relatively stouter, with the chitinous distal part gently curved and proportionately wider (
|
||
<figureCitation id="598786C5FFDEBA1EFBF8FC2EFB27FC04" box="[1061,1194,925,949]" captionStart="Fig" captionStartId="65.[113,148,1520,1542]" captionTargetBox="[114,1451,192,1507]" captionTargetId="figure-15@65.[93,1473,173,1513]" captionTargetPageId="65" captionText="Fig. 38. Contusarma bocourti, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand. A, anterior thoracic sternites 1–4; B, left third maxilliped (denuded); C, male pleon; D, left G1 (ventral view, denuded); E, left G1 (dorsa view); F, left distal part of G1 (ventral view, denuded); G, left distal part of G1 (dorsal view, denuded); H, left distal part of G1 (mesial view). Scales: A, C = 5.0 mm; B = 2.5 mm; D, E = 1.0 mm; F–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5351363" httpUri="https://zenodo.org/record/5351363/files/figure.png" pageId="63" pageNumber="954">Fig. 38D–H</figureCitation>
|
||
). Those from the rest of Southeast Asia are relatively more slender, and the chitinous distal part is straight and proportionately narrower (
|
||
<figureCitation id="598786C5FFDEBA1EFAA2FC6EFC39FBA4" captionStart="Fig" captionStartId="66.[138,173,1852,1874]" captionTargetBox="[135,1473,257,1836]" captionTargetId="figure-8@66.[131,1478,252,1841]" captionTargetPageId="66" captionText="Fig. 39. Contusarma cheirogonum. A–C, holotype male of Sesarma cheirogona (after Targioni Tozzetti, 1877: pl. 9 fig. 2); D, E, F–J, K, male (26.3 × 23.0 mm) (ZRC 1995.226), Bako National Park, Sarawak; L–N, neotype male (24.5 × 21.5 mm) (ZRC 1995.225), Bako National Park, Sarawak; O, male (24.5 × 21.3 mm) (ZRC 1964.9.28.140), Johor, Malaysia; P, male (22.3 × 20.5 mm) (ZRC 1964.9.28.127), Saribas, Sarawak; Q, male (28.8 × 25.5 mm) (ZRC 1964.9.28.126), Saribas, Sarawak. A, D, carapace; B, outer view of left chela; C, thoracic sternites and pleon; E, O–Q, pleon; F, L, left G1 (dorsal view, denuded); M, G, H, left distal part of G1 (dorsal view, denuded); N, I, left distal part of G1 (mesial view); J, left distal part of G1 (ventral view, denuded); K, left G2 (denuded). A–C, after Targioni Tozzetti (1877: pl. 9 fig. 2); D, H, J, K, after Ng (1995: fig. 14). Scales: D, E, K, O–Q = 5.0 mm; F, L = 2.0 mm; G–K, M, N = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5351365" httpUri="https://zenodo.org/record/5351365/files/figure.png" pageId="63" pageNumber="954">Fig. 39F–J, L–N</figureCitation>
|
||
). In addition, while the male pleon does have a degree of natural variability in proportions, as discussed by
|
||
<bibRefCitation id="A52DE7B1FFDEBA1EFC93FB8EFC77FBE4" author="Tweedie" box="[846,1018,1085,1109]" firstAuthor="Tweedie" pageId="63" pageNumber="954" pagination="88 - 113" refId="ref78385" refString="Tweedie MWF (1940) New and interesting Malaysian species of Sesarma and Utica (Crustacea, Brachyura). Bulletin of the Raffles Museum, 16: 88 - 113, figs. 1 - 12, pl. 24." type="journal article" year="1940">Tweedie (1940)</bibRefCitation>
|
||
, that of Thai specimens is still relatively transversely narrower (notably somite 6) (
|
||
<figureCitation id="598786C5FFDEBA1EFB06FBEEFABAFBC4" box="[1243,1335,1117,1141]" captionStart="Fig" captionStartId="65.[113,148,1520,1542]" captionTargetBox="[114,1451,192,1507]" captionTargetId="figure-15@65.[93,1473,173,1513]" captionTargetPageId="65" captionText="Fig. 38. Contusarma bocourti, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand. A, anterior thoracic sternites 1–4; B, left third maxilliped (denuded); C, male pleon; D, left G1 (ventral view, denuded); E, left G1 (dorsa view); F, left distal part of G1 (ventral view, denuded); G, left distal part of G1 (dorsal view, denuded); H, left distal part of G1 (mesial view). Scales: A, C = 5.0 mm; B = 2.5 mm; D, E = 1.0 mm; F–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5351363" httpUri="https://zenodo.org/record/5351363/files/figure.png" pageId="63" pageNumber="954">Fig. 38C</figureCitation>
|
||
) compared to those from the rest of Southeast Asia (
|
||
<figureCitation id="598786C5FFDEBA1EFB0FFBCEFAFCFB24" box="[1234,1393,1149,1173]" captionStart="Fig" captionStartId="66.[138,173,1852,1874]" captionTargetBox="[135,1473,257,1836]" captionTargetId="figure-8@66.[131,1478,252,1841]" captionTargetPageId="66" captionText="Fig. 39. Contusarma cheirogonum. A–C, holotype male of Sesarma cheirogona (after Targioni Tozzetti, 1877: pl. 9 fig. 2); D, E, F–J, K, male (26.3 × 23.0 mm) (ZRC 1995.226), Bako National Park, Sarawak; L–N, neotype male (24.5 × 21.5 mm) (ZRC 1995.225), Bako National Park, Sarawak; O, male (24.5 × 21.3 mm) (ZRC 1964.9.28.140), Johor, Malaysia; P, male (22.3 × 20.5 mm) (ZRC 1964.9.28.127), Saribas, Sarawak; Q, male (28.8 × 25.5 mm) (ZRC 1964.9.28.126), Saribas, Sarawak. A, D, carapace; B, outer view of left chela; C, thoracic sternites and pleon; E, O–Q, pleon; F, L, left G1 (dorsal view, denuded); M, G, H, left distal part of G1 (dorsal view, denuded); N, I, left distal part of G1 (mesial view); J, left distal part of G1 (ventral view, denuded); K, left G2 (denuded). A–C, after Targioni Tozzetti (1877: pl. 9 fig. 2); D, H, J, K, after Ng (1995: fig. 14). Scales: D, E, K, O–Q = 5.0 mm; F, L = 2.0 mm; G–K, M, N = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5351365" httpUri="https://zenodo.org/record/5351365/files/figure.png" pageId="63" pageNumber="954">Fig. 39E, O–Q</figureCitation>
|
||
). The adult male from
|
||
<collectingCountry id="B9ABDAD0FFDEBA1EFC00FB2EFBB1FB04" box="[989,1084,1181,1205]" name="Thailand" pageId="63" pageNumber="954">Thailand</collectingCountry>
|
||
is also distinctly more setose over the whole carapace and ambulatory legs (
|
||
<figureCitation id="598786C5FFDEBA1EFB37FB0EFAEAFB64" box="[1258,1383,1213,1237]" captionStart="Fig" captionStartId="42.[136,171,1862,1884]" captionTargetBox="[136,1474,233,1847]" captionTargetId="figure-7@42.[136,1474,233,1847]" captionTargetPageId="42" captionText="Fig. 23. A, E, Contusarma bocourti, lectotype male (24.0 × 27.2 mm) (MNHN D-10965), Bangkok, Thailand; B, F, C. bocourti, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand; C, C. bocourti, female (21.1 × 19.0 mm) (ZRC 2019.1114), Bangkok, Thailand; D, H, C. cheirogonum, neotype male (24.5 × 21.5 mm) (ZRC 1995.225), Bako National Park, Sarawak; G, C. cheirogonum, male (23.0 × 20.7 mm) (ZRC 2000.2018), Pulau Ubin, Singapore. A–D, overall habitus; E–H, anterior half of carapace." figureDoi="http://doi.org/10.5281/zenodo.5351341" httpUri="https://zenodo.org/record/5351341/files/figure.png" pageId="63" pageNumber="954">Fig. 23B, F</figureCitation>
|
||
), with the granules on the chelae proportionately stouter and larger (
|
||
<figureCitation id="598786C5FFDEBA1EFCE9FB4EFC37FAA4" box="[820,954,1277,1301]" captionStart="Fig" captionStartId="50.[136,171,1012,1034]" captionTargetBox="[136,1474,189,997]" captionTargetId="figure-499@50.[136,1474,189,997]" captionTargetPageId="50" captionText="Fig. 28. Chela of Contusarma. A, C. bocourti, lectotype male (24.0 × 27.2 mm) (MNHN D-10965), Bangkok, Thailand; B, C, C. bocourti, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand; D, C. cheirogonum, neotype male (24.5 × 21.5 mm) (ZRC 1995.225), Bako National Park, Sarawak. A, B, D, outer view; C, ventro-marginal view." figureDoi="http://doi.org/10.5281/zenodo.5351347" httpUri="https://zenodo.org/record/5351347/files/figure.png" pageId="63" pageNumber="954">Fig. 28A–C</figureCitation>
|
||
). Those from the rest of Southeast Asia have sparsely setose to almost glabrous carapaces and legs (
|
||
<figureCitation id="598786C5FFDEBA1EFA5DFAAEFC0CFAE4" captionStart="Fig" captionStartId="42.[136,171,1862,1884]" captionTargetBox="[136,1474,233,1847]" captionTargetId="figure-7@42.[136,1474,233,1847]" captionTargetPageId="42" captionText="Fig. 23. A, E, Contusarma bocourti, lectotype male (24.0 × 27.2 mm) (MNHN D-10965), Bangkok, Thailand; B, F, C. bocourti, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand; C, C. bocourti, female (21.1 × 19.0 mm) (ZRC 2019.1114), Bangkok, Thailand; D, H, C. cheirogonum, neotype male (24.5 × 21.5 mm) (ZRC 1995.225), Bako National Park, Sarawak; G, C. cheirogonum, male (23.0 × 20.7 mm) (ZRC 2000.2018), Pulau Ubin, Singapore. A–D, overall habitus; E–H, anterior half of carapace." figureDoi="http://doi.org/10.5281/zenodo.5351341" httpUri="https://zenodo.org/record/5351341/files/figure.png" pageId="63" pageNumber="954">Fig. 23G, H</figureCitation>
|
||
), and the granules on the chelae are smaller (
|
||
<figureCitation id="598786C5FFDEBA1EFA5DFA8EFCD3FAC4" captionStart="Fig" captionStartId="50.[136,171,1012,1034]" captionTargetBox="[136,1474,189,997]" captionTargetId="figure-499@50.[136,1474,189,997]" captionTargetPageId="50" captionText="Fig. 28. Chela of Contusarma. A, C. bocourti, lectotype male (24.0 × 27.2 mm) (MNHN D-10965), Bangkok, Thailand; B, C, C. bocourti, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand; D, C. cheirogonum, neotype male (24.5 × 21.5 mm) (ZRC 1995.225), Bako National Park, Sarawak. A, B, D, outer view; C, ventro-marginal view." figureDoi="http://doi.org/10.5281/zenodo.5351347" httpUri="https://zenodo.org/record/5351347/files/figure.png" pageId="63" pageNumber="954">Fig. 28D</figureCitation>
|
||
). The degree of setation is not discernible on the
|
||
<typeStatus id="1E0724E2FFDEBA1EFAAEFAEDFA26FAC7" box="[1395,1451,1374,1398]" pageId="63" pageNumber="954">types</typeStatus>
|
||
of
|
||
<taxonomicName id="06BCE1C3FFDEBA1EFC96FACDFC48FA24" authorityName="A. Milne-Edwards" authorityYear="1869" box="[843,965,1405,1429]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="63" pageNumber="954" phylum="Arthropoda" rank="species" species="bocourti">
|
||
<emphasis id="F3C84652FFDEBA1EFC96FACDFC48FA24" box="[843,965,1405,1429]" italics="true" pageId="63" pageNumber="954">C. bocourti</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="598786C5FFDEBA1EFC0BFACEFBD4FA24" box="[982,1113,1405,1429]" captionStart="Fig" captionStartId="42.[136,171,1862,1884]" captionTargetBox="[136,1474,233,1847]" captionTargetId="figure-7@42.[136,1474,233,1847]" captionTargetPageId="42" captionText="Fig. 23. A, E, Contusarma bocourti, lectotype male (24.0 × 27.2 mm) (MNHN D-10965), Bangkok, Thailand; B, F, C. bocourti, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand; C, C. bocourti, female (21.1 × 19.0 mm) (ZRC 2019.1114), Bangkok, Thailand; D, H, C. cheirogonum, neotype male (24.5 × 21.5 mm) (ZRC 1995.225), Bako National Park, Sarawak; G, C. cheirogonum, male (23.0 × 20.7 mm) (ZRC 2000.2018), Pulau Ubin, Singapore. A–D, overall habitus; E–H, anterior half of carapace." figureDoi="http://doi.org/10.5281/zenodo.5351341" httpUri="https://zenodo.org/record/5351341/files/figure.png" pageId="63" pageNumber="954">Fig. 23A, E</figureCitation>
|
||
), but the setae may have fallen off due to their age and dried condition. The phylogenetic tree supports this, with a sister-group relationship between the two species (
|
||
<figureCitation id="598786C5FFDEBA1EFC39FA6EFBBBFA44" box="[996,1078,1501,1525]" captionStart="Fig" captionStartId="91.[113,149,1757,1779]" captionTargetBox="[116,1448,352,1737]" captionTargetId="figure-15@91.[86,1475,347,1749]" captionTargetPageId="91" captionText="Fig. 59. Phylogenetic relationships among selected taxa of Sesarmidae (91 OTUs), including all representatives of Chiromantes and Pseudosesarma previous to current revision of these genera. In addition, several species of Bresedium and Sesarmops are considered, as the revision also involves some of their former members. Topology and confidence values were determined by means of a phylogenetic reconstruction based on 1,096 basepairs of mtDNA (16S-tRNALeu-ND1), conducted with MrBayes, GTR+I+G as substitution model, and Parasesarma plicatum as outgroup (midpoint rooting). Posterior probabilities were determined after 4 million generations with a burn-in of 25%. All species so far classified within Chiromantes (without obvious anterolateral teeth) are shaded in blue, whereas those species that used to be classified in Pseudosesarma (carrying at least one pair of anterolateral teeth) are shaded in yellow." figureDoi="http://doi.org/10.5281/zenodo.5351405" httpUri="https://zenodo.org/record/5351405/files/figure.png" pageId="63" pageNumber="954">Fig. 59</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C1039A40FFDEBA60FCF0F9AEFDE7F7BA" blockId="63.[812,1453,1565,2101]" lastBlockId="65.[113,753,1683,2059]" lastPageId="65" lastPageNumber="956" pageId="63" pageNumber="954">
|
||
While we have no doubt that the Thai material is
|
||
<taxonomicName id="06BCE1C3FFDEBA1EFCF3F98DFB99F9E4" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1869" box="[814,1044,1597,1621]" class="Malacostraca" family="Sesarmidae" genus="Contusarma" kingdom="Animalia" order="Decapoda" pageId="63" pageNumber="954" phylum="Arthropoda" rank="species" sensu="stricto" species="bocourti">
|
||
<emphasis id="F3C84652FFDEBA1EFCF3F98DFB99F9E4" box="[814,1044,1597,1621]" italics="true" pageId="63" pageNumber="954">Contusarma bocourti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E8FBFB29FFDEBA1EFBC6F98DFBD5F9E4" box="[1051,1112,1598,1621]" pageId="63" pageNumber="954" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
, Hoang et al.’s (2012) material of “
|
||
<taxonomicName id="06BCE1C3FFDEBA1EFC80F9EEFBF2F9C4" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1869" box="[861,1151,1629,1653]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="63" pageNumber="954" phylum="Arthropoda" rank="species" species="bocourti">
|
||
<emphasis id="F3C84652FFDEBA1EFC80F9EEFBF2F9C4" box="[861,1151,1629,1653]" italics="true" pageId="63" pageNumber="954">Pseudosesarma bocourti</emphasis>
|
||
</taxonomicName>
|
||
” from southern
|
||
<collectingCountry id="B9ABDAD0FFDEBA1EFA94F9EEFA27F9C4" box="[1353,1450,1629,1653]" name="Vietnam" pageId="63" pageNumber="954">Vietnam</collectingCountry>
|
||
should be re-examined if possible. It is provisionally identified as
|
||
<taxonomicName id="06BCE1C3FFDEBA1EFC1DF92DFBCCF904" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1869" box="[960,1089,1693,1717]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="63" pageNumber="954" phylum="Arthropoda" rank="species" species="bocourti">
|
||
<emphasis id="F3C84652FFDEBA1EFC1DF92DFBCCF904" box="[960,1089,1693,1717]" italics="true" pageId="63" pageNumber="954">C. bocourti</emphasis>
|
||
</taxonomicName>
|
||
. The identity of the Malaysian,
|
||
<collectingCountry id="B9ABDAD0FFDEBA1EFCF0F90EFC12F964" box="[813,927,1725,1749]" name="Singapore" pageId="63" pageNumber="954">Singapore</collectingCountry>
|
||
, and Bornean specimens is more problematic. This is because the real origin of
|
||
<taxonomicName id="06BCE1C3FFDEBA1EFB70F96DFA02F944" authorityName="Targioni Tozzetti" authorityYear="1877" box="[1197,1423,1757,1781]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="63" pageNumber="954" phylum="Arthropoda" rank="species" species="cheirogona">
|
||
<emphasis id="F3C84652FFDEBA1EFB70F96DFA02F944" box="[1197,1423,1757,1781]" italics="true" pageId="63" pageNumber="954">Sesarma cheirogona</emphasis>
|
||
</taxonomicName>
|
||
is unclear, as discussed above.
|
||
<bibRefCitation id="A52DE7B1FFDEBA1EFBAEF94EFAAEF8A4" author="Tweedie" box="[1139,1315,1789,1813]" firstAuthor="Tweedie" pageId="63" pageNumber="954" pagination="88 - 113" refId="ref78385" refString="Tweedie MWF (1940) New and interesting Malaysian species of Sesarma and Utica (Crustacea, Brachyura). Bulletin of the Raffles Museum, 16: 88 - 113, figs. 1 - 12, pl. 24." type="journal article" year="1940">Tweedie (1940)</bibRefCitation>
|
||
commented that
|
||
<bibRefCitation id="A52DE7B1FFDEBA1EFCBDF8AEFB08F884" author="Targioni Tozzetti's" box="[864,1157,1821,1845]" firstAuthor="Targioni Tozzetti's" pageId="63" pageNumber="954" pagination="1 - 257" refId="ref77991" refString="Targioni Tozzetti A (1877) Crostacei brachyuri e anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R Istituto di Studi Superiori Pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1: i - xxix + 1 - 257, pls. 1 - 12." type="journal article" year="1877">Targioni Tozzetti’s (1877)</bibRefCitation>
|
||
material of
|
||
<taxonomicName id="06BCE1C3FFDEBA1EFACFF8ADFA21F884" authorityName="Targioni Tozzetti" authorityYear="1877" box="[1298,1452,1821,1845]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="63" pageNumber="954" phylum="Arthropoda" rank="species" species="cheirogona">
|
||
<emphasis id="F3C84652FFDEBA1EFACFF8ADFA21F884" box="[1298,1452,1821,1845]" italics="true" pageId="63" pageNumber="954">S. cheirogona</emphasis>
|
||
</taxonomicName>
|
||
could have originated from Borneo or Sumatra, but that expedition also visited
|
||
<collectingCountry id="B9ABDAD0FFDEBA1EFBFDF8EEFB02F8C4" box="[1056,1167,1885,1909]" name="Singapore" pageId="63" pageNumber="954">Singapore</collectingCountry>
|
||
, Java, and
|
||
<collectingCountry id="B9ABDAD0FFDEBA1EFB22F8EEFAD3F8C4" box="[1279,1374,1885,1909]" name="Vietnam" pageId="63" pageNumber="954">Vietnam</collectingCountry>
|
||
; before the vessel reached
|
||
<collectingCountry id="B9ABDAD0FFDEBA1EFBDCF8CDFBCFF824" box="[1025,1090,1918,1941]" name="Japan" pageId="63" pageNumber="954">Japan</collectingCountry>
|
||
. Comparing the figures given in
|
||
<bibRefCitation id="A52DE7B1FFDEBA1EFCF0F82EFBA0F804" author="Targioni Tozzetti" box="[813,1069,1949,1973]" firstAuthor="Targioni Tozzetti" pageId="63" pageNumber="954" pagination="1 - 257" refId="ref77991" refString="Targioni Tozzetti A (1877) Crostacei brachyuri e anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R Istituto di Studi Superiori Pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1: i - xxix + 1 - 257, pls. 1 - 12." type="journal article" year="1877">Targioni Tozzetti (1877)</bibRefCitation>
|
||
, however, it seems likely his material came from
|
||
<collectingCountry id="B9ABDAD0FFDEBA1EFC76F80EFB9AF864" box="[939,1047,1981,2005]" name="Singapore" pageId="63" pageNumber="954">Singapore</collectingCountry>
|
||
or Borneo (or adjacent areas) as his figure (
|
||
<figureCitation id="598786C5FFDEBA1EFCA6F86EFC52F844" box="[891,991,2013,2037]" captionStart="Fig" captionStartId="66.[138,173,1852,1874]" captionTargetBox="[135,1473,257,1836]" captionTargetId="figure-8@66.[131,1478,252,1841]" captionTargetPageId="66" captionText="Fig. 39. Contusarma cheirogonum. A–C, holotype male of Sesarma cheirogona (after Targioni Tozzetti, 1877: pl. 9 fig. 2); D, E, F–J, K, male (26.3 × 23.0 mm) (ZRC 1995.226), Bako National Park, Sarawak; L–N, neotype male (24.5 × 21.5 mm) (ZRC 1995.225), Bako National Park, Sarawak; O, male (24.5 × 21.3 mm) (ZRC 1964.9.28.140), Johor, Malaysia; P, male (22.3 × 20.5 mm) (ZRC 1964.9.28.127), Saribas, Sarawak; Q, male (28.8 × 25.5 mm) (ZRC 1964.9.28.126), Saribas, Sarawak. A, D, carapace; B, outer view of left chela; C, thoracic sternites and pleon; E, O–Q, pleon; F, L, left G1 (dorsal view, denuded); M, G, H, left distal part of G1 (dorsal view, denuded); N, I, left distal part of G1 (mesial view); J, left distal part of G1 (ventral view, denuded); K, left G2 (denuded). A–C, after Targioni Tozzetti (1877: pl. 9 fig. 2); D, H, J, K, after Ng (1995: fig. 14). Scales: D, E, K, O–Q = 5.0 mm; F, L = 2.0 mm; G–K, M, N = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5351365" httpUri="https://zenodo.org/record/5351365/files/figure.png" pageId="63" pageNumber="954">Fig. 39C</figureCitation>
|
||
;
|
||
<bibRefCitation id="A52DE7B1FFDEBA1EFC31F86EFB7DF844" author="Targioni Tozzetti" box="[1004,1264,2013,2037]" firstAuthor="Targioni Tozzetti" pageId="63" pageNumber="954" pagination="1 - 257" refId="ref77991" refString="Targioni Tozzetti A (1877) Crostacei brachyuri e anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R Istituto di Studi Superiori Pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1: i - xxix + 1 - 257, pls. 1 - 12." type="journal article" year="1877">Targioni Tozzetti, 1877</bibRefCitation>
|
||
: fig. 2d) shows a proportionately wider pleon similar to those from this area (
|
||
<figureCitation id="598786C5FFDEBA1EFCE8F7AEFC53F784" box="[821,990,2077,2101]" captionStart="Fig" captionStartId="66.[138,173,1852,1874]" captionTargetBox="[135,1473,257,1836]" captionTargetId="figure-8@66.[131,1478,252,1841]" captionTargetPageId="66" captionText="Fig. 39. Contusarma cheirogonum. A–C, holotype male of Sesarma cheirogona (after Targioni Tozzetti, 1877: pl. 9 fig. 2); D, E, F–J, K, male (26.3 × 23.0 mm) (ZRC 1995.226), Bako National Park, Sarawak; L–N, neotype male (24.5 × 21.5 mm) (ZRC 1995.225), Bako National Park, Sarawak; O, male (24.5 × 21.3 mm) (ZRC 1964.9.28.140), Johor, Malaysia; P, male (22.3 × 20.5 mm) (ZRC 1964.9.28.127), Saribas, Sarawak; Q, male (28.8 × 25.5 mm) (ZRC 1964.9.28.126), Saribas, Sarawak. A, D, carapace; B, outer view of left chela; C, thoracic sternites and pleon; E, O–Q, pleon; F, L, left G1 (dorsal view, denuded); M, G, H, left distal part of G1 (dorsal view, denuded); N, I, left distal part of G1 (mesial view); J, left distal part of G1 (ventral view, denuded); K, left G2 (denuded). A–C, after Targioni Tozzetti (1877: pl. 9 fig. 2); D, H, J, K, after Ng (1995: fig. 14). Scales: D, E, K, O–Q = 5.0 mm; F, L = 2.0 mm; G–K, M, N = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5351365" httpUri="https://zenodo.org/record/5351365/files/figure.png" pageId="63" pageNumber="954">Fig. 39E, O–Q</figureCitation>
|
||
). For this reason, we refer all the present material from Peninsular
|
||
<collectingCountry id="B9ABDAD0FFA0BA60FE5DF920FE69F91A" box="[384,484,1683,1707]" name="Malaysia" pageId="65" pageNumber="956">Malaysia</collectingCountry>
|
||
,
|
||
<collectingCountry id="B9ABDAD0FFA0BA60FE32F920FDD3F91A" box="[495,606,1683,1707]" name="Singapore" pageId="65" pageNumber="956">Singapore</collectingCountry>
|
||
, and
|
||
<collectingCountry id="B9ABDAD0FFA0BA60FD49F920FD7DF91A" box="[660,752,1683,1707]" name="Malaysia" pageId="65" pageNumber="956">Sarawak</collectingCountry>
|
||
to
|
||
<taxonomicName id="06BCE1C3FFA0BA60FF4CF907FECEF97A" baseAuthorityName="Targioni Tozzetti" baseAuthorityYear="1877" box="[145,323,1715,1739]" class="Malacostraca" family="Sesarmidae" genus="Contusarma" kingdom="Animalia" order="Decapoda" pageId="65" pageNumber="956" phylum="Arthropoda" rank="species" species="cheirogonum">
|
||
<emphasis id="F3C84652FFA0BA60FF4CF907FECEF97A" box="[145,323,1715,1739]" italics="true" pageId="65" pageNumber="956">C. cheirogonum</emphasis>
|
||
</taxonomicName>
|
||
. The identity of the specimen figured by
|
||
<bibRefCitation id="A52DE7B1FFA0BA60FF48F960FED8F95A" author="Tesch" box="[149,341,1747,1771]" firstAuthor="Tesch" page="138" pageId="65" pageNumber="956" pagination="127 - 260" refId="ref78063" refString="Tesch JJ (1917) Synopsis of the genera Sesarma, Metasesarma, Sarmatium and Clistocoeloma, with a key to the determination of the IndoPacific species. Zoologische Mededelingen, 3: 127 - 260, pls. 15 - 17." type="journal article" year="1917">Tesch (1917: 138</bibRefCitation>
|
||
, fig. 2b) with a very wide male pleon from Balikpapan in eastern Kalimantan in Borneo cannot be ascertained without a re-examination at a future date. This may simply be intraspecific variation because
|
||
<specimenCount id="D7BA51C9FFA0BA60FD14F887FF5AF8DA" pageId="65" pageNumber="956" type="generic">one specimen</specimenCount>
|
||
from
|
||
<collectingCountry id="B9ABDAD0FFA0BA60FEC1F8E0FEF5F8DA" box="[284,376,1875,1899]" name="Malaysia" pageId="65" pageNumber="956">Sarawak</collectingCountry>
|
||
has a similarly proportioned male pleon (
|
||
<figureCitation id="598786C5FFA0BA60FF60F8C0FEACF83A" box="[189,289,1907,1931]" captionStart="Fig" captionStartId="54.[136,171,1949,1971]" captionTargetBox="[169,1440,189,1933]" captionTargetId="figure-8@54.[169,1440,189,1933]" captionTargetPageId="54" captionText="Fig. 31. Male anterior thoracic sternum and pleon. A, Contusarma bocourti, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand; B, Contusarma cheirogonum, male (15.3 × 14.0 mm) (ZRC 2011.0924), Pulau Tioman, Malaysia; C, Contusarma cheirogonum, neotype male (24.5 × 21.5 mm) (ZRC 1995.225), Bako National Park, Sarawak; D, Miersarma granosimanum, lectotype male (16.9 × 14.8 mm) (NHM 1880.6), Borneo; E, Miersarma granosimanum, male (22.5 × 19.2 mm) (ZRC 1965.7.29.164), Sedili River, Johor, Malaysia; F, Bresedium laevimanum, lectotype male (20.0 × 17.6 mm) (MNHG), Borneo; G, Bresedium laevimanum, male (26.3 × 23.7 mm) (NHM) (lectotype of Sesarma sediliensis), Johor, Malaysia; H, Bresedium laevimanum, male (21.2 × 18.2 mm) (ZRC 1965.7.29.121), Johor, Malaysia." figureDoi="http://doi.org/10.5281/zenodo.5351351" httpUri="https://zenodo.org/record/5351351/files/figure.png" pageId="65" pageNumber="956">Fig. 31C</figureCitation>
|
||
), with the other male specimens from this location having narrower pleons. The records from the rest of Borneo and Sumatra by
|
||
<bibRefCitation id="A52DE7B1FFA0BA60FE74F807FDDCF87A" author="De Man" box="[425,593,1971,1995]" firstAuthor="De Man" pageId="65" pageNumber="956" pagination="21 - 36" refId="ref72026" refString="Man JG De (1880) On some species of the genus Sesarma Say and Cardisoma Latr. Notes Leyden Museum, 2 (5): 21 - 36." type="journal article" year="1880">De Man (1880</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="A52DE7B1FFA0BA60FDBDF800FD17F87A" author="De Man" box="[608,666,1971,1995]" firstAuthor="De Man" journalOrPublisher="Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere" pageId="65" pageNumber="956" pagination="75 - 218" part="9" refId="ref72333" refString="Man JG De (1895) Bericht uber die von Herrn Schiffscaptan Storm zu Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der JavaSee gesammelten Decapoden und Stomatopoden. Zweiter Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 9: 75 - 218." title="Bericht uber die von Herrn Schiffscaptan Storm zu Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der JavaSee gesammelten Decapoden und Stomatopoden. Zweiter Theil" type="journal article" year="1895">1895</bibRefCitation>
|
||
),
|
||
<bibRefCitation id="A52DE7B1FFA0BA60FD6CF800FF36F85A" author="Miers" firstAuthor="Miers" pageId="65" pageNumber="956" pagination="304 - 317" refId="ref72642" refString="Miers EJ (1880) On a collection of Crustacea from the Malayan Region. Part II. Telphusidea, Catometopa and Oxystomata. Annals and Magazine of Natural History, Series 5, 5 (28): 304 - 317, pl. 14." type="journal article" year="1880">Miers (1880)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="A52DE7B1FFA0BA60FF14F860FEEDF85A" author="Nobili" box="[201,352,2003,2027]" firstAuthor="Nobili" pageId="65" pageNumber="956" pagination="473 - 523" refId="ref74323" refString="Nobili G (1900) Decapodi e stomatopodi Indo-Malesi. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 20 (40): 473 - 523, figs. 1 - 4." type="journal article" year="1900">Nobili (1900)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="A52DE7B1FFA0BA60FEB0F860FE72F85A" author="Tesch" box="[365,511,2003,2027]" firstAuthor="Tesch" pageId="65" pageNumber="956" pagination="127 - 260" refId="ref78063" refString="Tesch JJ (1917) Synopsis of the genera Sesarma, Metasesarma, Sarmatium and Clistocoeloma, with a key to the determination of the IndoPacific species. Zoologische Mededelingen, 3: 127 - 260, pls. 15 - 17." type="journal article" year="1917">Tesch (1917)</bibRefCitation>
|
||
, and
|
||
<bibRefCitation id="A52DE7B1FFA0BA60FDE0F867FD4AF85A" author="Roux" box="[573,711,2003,2027]" firstAuthor="Roux" pageId="65" pageNumber="956" pagination="1" refId="ref75491" refString="Roux J (1933) Crustaces Decapodes d'eau douce. In: Resultats Scientifiques du Voyage aux Indes Orientales Neerlandaises de LL. AA. RR. le Prince et la Princesse Leopold de Belgique. Memoires du Musee Royal d'Histoire Naturelle de Belgique, Hors Serie [= Supplement] 3, No. 14 [= 3 (14)]: 1 - 18, figs. 1, 2." type="journal article" year="1933">Roux (1933)</bibRefCitation>
|
||
are also provisionally referred to
|
||
<taxonomicName id="06BCE1C3FFA0BA60FE68F847FDEEF7BA" baseAuthorityName="Targioni Tozzetti" baseAuthorityYear="1877" box="[437,611,2035,2059]" class="Malacostraca" family="Sesarmidae" genus="Contusarma" kingdom="Animalia" order="Decapoda" pageId="65" pageNumber="956" phylum="Arthropoda" rank="species" species="cheirogonum">
|
||
<emphasis id="F3C84652FFA0BA60FE68F847FDEEF7BA" box="[437,611,2035,2059]" italics="true" pageId="65" pageNumber="956">C. cheirogonum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="95C3CAC8FFA1BA61FF55F835FD2CF79A" ID-DOI="http://doi.org/10.5281/zenodo.6519019" ID-Zenodo-Dep="6519019" httpUri="https://zenodo.org/record/6519019/files/figure.png" pageId="64" pageNumber="955" startId="64.[136,171,1926,1948]" subCaptionStartIDs="64.[573,601,2070,2092]" subCaptionStarts="fig. 7E" targetBox="[138,1476,190,1907]" targetPageId="64">
|
||
<paragraph id="C1039A40FFA1BA61FF55F835FD2CF79A" blockId="64.[136,1475,1925,2092]" pageId="64" pageNumber="955">
|
||
Fig. 37. A–G,
|
||
<taxonomicName id="06BCE1C3FFA1BA61FEC5F835FD8EF82A" authorityName="Ng" authorityYear="2018" box="[280,515,1926,1947]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="64" pageNumber="955" phylum="Arthropoda" rank="species" species="brehieri">
|
||
<emphasis id="F3C84652FFA1BA61FEC5F835FD8EF82A" box="[280,515,1926,1947]" italics="true" pageId="64" pageNumber="955">Pseudosesarma brehieri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="1E0724E2FFA1BA61FDCCF835FDE9F82A" box="[529,612,1926,1947]" pageId="64" pageNumber="955" type="paratype">paratype</typeStatus>
|
||
female (14.9 ×
|
||
<quantity id="064437A5FFA1BA61FCDBF835FCD2F82A" box="[774,863,1926,1947]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="64" pageNumber="955" unit="mm" value="13.4">13.4 mm</quantity>
|
||
) (ZRC 2016.0594),
|
||
<collectingCountry id="B9ABDAD0FFA1BA61FBFAF835FB0AF82A" box="[1063,1159,1926,1947]" name="Myanmar" pageId="64" pageNumber="955">Myanmar</collectingCountry>
|
||
; F, G,
|
||
<taxonomicName id="06BCE1C3FFA1BA61FB10F834FABBF82A" authorityName="Ng" authorityYear="2018" box="[1229,1334,1926,1948]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="64" pageNumber="955" phylum="Arthropoda" rank="species" species="brehieri">
|
||
<emphasis id="F3C84652FFA1BA61FB10F834FABBF82A" box="[1229,1334,1926,1948]" italics="true" pageId="64" pageNumber="955">P. brehieri</emphasis>
|
||
</taxonomicName>
|
||
, male (15.0 ×
|
||
<quantity id="064437A5FFA1BA61FF55F810FF6CF809" box="[136,225,1955,1977]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="64" pageNumber="955" unit="mm" value="13.6">13.6 mm</quantity>
|
||
) (ZRC 2013.0209),
|
||
<collectingRegion id="037854A2FFA1BA61FE77F810FDA4F808" box="[426,553,1955,1977]" country="India" name="West Bengal" pageId="64" pageNumber="955">West Bengal</collectingRegion>
|
||
,
|
||
<collectingCountry id="B9ABDAD0FFA1BA61FDE8F810FDE4F808" box="[565,617,1955,1977]" name="India" pageId="64" pageNumber="955">India</collectingCountry>
|
||
; H, I,
|
||
<taxonomicName id="06BCE1C3FFA1BA61FD77F810FC99F809" authorityName="Ng" authorityYear="2018" box="[682,788,1955,1976]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="64" pageNumber="955" phylum="Arthropoda" rank="species" species="brehieri">
|
||
<emphasis id="F3C84652FFA1BA61FD77F810FC99F809" box="[682,788,1955,1976]" italics="true" pageId="64" pageNumber="955">P. brehieri</emphasis>
|
||
</taxonomicName>
|
||
, male (14.5 × 13.0 mm) (ZRC 2013.0209),
|
||
<collectingRegion id="037854A2FFA1BA61FB17F810FAC7F808" box="[1226,1354,1955,1977]" country="India" name="West Bengal" pageId="64" pageNumber="955">West Bengal</collectingRegion>
|
||
,
|
||
<collectingCountry id="B9ABDAD0FFA1BA61FA8BF810FA07F808" box="[1366,1418,1955,1977]" name="India" pageId="64" pageNumber="955">India</collectingCountry>
|
||
; J–O,
|
||
<taxonomicName id="06BCE1C3FFA1BA61FF55F873FE82F865" authorityName="Calman" authorityYear="1920" box="[136,271,1983,2005]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="64" pageNumber="955" phylum="Arthropoda" rank="species" species="boulengeri">
|
||
<emphasis id="F3C84652FFA1BA61FF55F873FE82F865" box="[136,271,1983,2005]" italics="true" pageId="64" pageNumber="955">P. boulengeri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="1E0724E2FFA1BA61FEC3F80CFEFAF864" box="[286,375,1983,2005]" pageId="64" pageNumber="955" type="lectotype">lectotype</typeStatus>
|
||
male (26.9 ×
|
||
<quantity id="064437A5FFA1BA61FDD7F80CFDE9F864" box="[522,612,1983,2005]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="64" pageNumber="955" unit="mm" value="23.5">23.5 mm</quantity>
|
||
) (NHM 1919.11.14.1),
|
||
<collectingRegion id="037854A2FFA1BA61FC89F873FC00F864" box="[852,909,1984,2005]" country="Iraq" name="Al Basrah" pageId="64" pageNumber="955">Basra</collectingRegion>
|
||
,
|
||
<collectingCountry id="B9ABDAD0FFA1BA61FC47F873FC49F864" box="[922,964,1984,2005]" name="Iraq" pageId="64" pageNumber="955">Iraq</collectingCountry>
|
||
. A, K, male pleon; B, F, L, left G1 (ventral view, denuded); D, G, M, left G1 (dorsal view, denuded); E, left G2; H, O, left distal part of G1 (dorsal view, denuded); I, N, left distal part of G1 (ventral view, denuded); J, anterior thoracic sternites 1–4. Scales: A = 4.0 mm; B–E, N, O = 1.0 mm; F–I =
|
||
<quantity id="064437A5FFA1BA61FADEF84AFADCF7BE" box="[1283,1361,2041,2063]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="64" pageNumber="955" unit="mm" value="0.5">0.5 mm</quantity>
|
||
; J, K = 5.0 mm; L, M = 2.0 mm. A–E, after
|
||
<bibRefCitation id="A52DE7B1FFA1BA61FE0CF7A5FDBFF79A" author="Ng" box="[465,562,2070,2091]" firstAuthor="Ng" pageId="64" pageNumber="955" pagination="185 - 197" refId="ref73639" refString="Ng PKL (2018) On two new species of potamid and sesarmid crabs (Decapoda, Brachyura) from caves in Myanmar. Crustaceana, 91 (2): 185 - 197." type="journal article" year="2018">Ng (2018</bibRefCitation>
|
||
: fig. 7E–I).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="95C3CAC8FFA0BA60FFACFA43FE69F9ED" ID-DOI="http://doi.org/10.5281/zenodo.5351363" ID-Zenodo-Dep="5351363" httpUri="https://zenodo.org/record/5351363/files/figure.png" pageId="65" pageNumber="956" startId="65.[113,148,1520,1542]" targetBox="[114,1451,192,1507]" targetPageId="65">
|
||
<paragraph id="C1039A40FFA0BA60FFACFA43FE69F9ED" blockId="65.[113,1452,1520,1628]" pageId="65" pageNumber="956">
|
||
Fig. 38.
|
||
<taxonomicName id="06BCE1C3FFA0BA60FF1BFA43FE1BF9B4" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1869" box="[198,406,1520,1541]" class="Malacostraca" family="Sesarmidae" genus="Contusarma" kingdom="Animalia" order="Decapoda" pageId="65" pageNumber="956" phylum="Arthropoda" rank="species" species="bocourti">
|
||
<emphasis id="F3C84652FFA0BA60FF1BFA43FE1BF9B4" box="[198,406,1520,1541]" italics="true" pageId="65" pageNumber="956">Contusarma bocourti</emphasis>
|
||
</taxonomicName>
|
||
, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand. A, anterior thoracic sternites 1–4; B, left third maxilliped (denuded); C, male pleon; D, left G1 (ventral view, denuded); E, left G1 (dorsa view); F, left distal part of G1 (ventral view, denuded); G, left distal part of G1 (dorsal view, denuded); H, left distal part of G1 (mesial view). Scales: A, C = 5.0 mm; B = 2.5 mm; D, E = 1.0 mm; F–H = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C1039A40FFA0BA60FCF0F920FB4BF7BA" blockId="65.[812,1452,1683,2060]" pageId="65" pageNumber="956">
|
||
The types of
|
||
<bibRefCitation id="A52DE7B1FFA0BA60FC65F920FB32F91A" author="Targioni Tozzetti" box="[952,1215,1683,1707]" firstAuthor="Targioni Tozzetti" pageId="65" pageNumber="956" pagination="1 - 257" refId="ref77991" refString="Targioni Tozzetti A (1877) Crostacei brachyuri e anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R Istituto di Studi Superiori Pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1: i - xxix + 1 - 257, pls. 1 - 12." type="journal article" year="1877">Targioni Tozzetti (1877)</bibRefCitation>
|
||
are lost.
|
||
<bibRefCitation id="A52DE7B1FFA0BA60FAFDF927FCD0F97A" author="Lucas" firstAuthor="Lucas" page="200" pageId="65" pageNumber="956" pagination="148 - 247" refId="ref71976" refString="Lucas JS (1981) Spider crabs of the family Hymenosomatidae (Crustacea: Brachyura) with particular reference to Australian species: Systematics and biology. Records of the Australian Museum, 1980, 33 (4): 148 - 247, figs. 1 - 10." type="journal article" year="1981">Lucas (1981: 200)</bibRefCitation>
|
||
notes that “The type specimens which were kept in the Museo Zoologico de “La Specola”, Firenze,
|
||
<collectingCountry id="B9ABDAD0FFA0BA60FAC0F960FADDF95A" box="[1309,1360,1747,1771]" name="Italy" pageId="65" pageNumber="956">Italy</collectingCountry>
|
||
, are not extant; they were lost during World War II (M. Poggesi, pers. comm.)”. Gianna Innocenti (Universitá di Firenze) conducted a fresh search in 2019 but also came to the conclusion the type of
|
||
<taxonomicName id="06BCE1C3FFA0BA60FCA0F8E7FBD5F8DA" authorityName="Targioni Tozzetti" authorityYear="1877" box="[893,1112,1875,1899]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="65" pageNumber="956" phylum="Arthropoda" rank="species" species="cheirogona">
|
||
<emphasis id="F3C84652FFA0BA60FCA0F8E7FBD5F8DA" box="[893,1112,1875,1899]" italics="true" pageId="65" pageNumber="956">Sesarma cheirogona</emphasis>
|
||
</taxonomicName>
|
||
is no longer extant. To stabilise the taxonomy of the two species in
|
||
<taxonomicName id="06BCE1C3FFA0BA60FB62F8C7FACEF83A" authorityName="Schubart & Ng" authorityYear="2020" box="[1215,1347,1908,1931]" class="Malacostraca" family="Sesarmidae" genus="Contusarma" kingdom="Animalia" order="Decapoda" pageId="65" pageNumber="956" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3C84652FFA0BA60FB62F8C7FACEF83A" box="[1215,1347,1908,1931]" italics="true" pageId="65" pageNumber="956">Contusarma</emphasis>
|
||
</taxonomicName>
|
||
, we here select a male (24.5 ×
|
||
<quantity id="064437A5FFA0BA60FBECF820FB17F81A" box="[1073,1178,1939,1963]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" pageId="65" pageNumber="956" unit="mm" value="21.5">21.5 mm</quantity>
|
||
) (ZRC 1995.225) from Bako National Park,
|
||
<collectingCountry id="B9ABDAD0FFA0BA60FBC5F800FBFAF87A" box="[1048,1143,1971,1995]" name="Malaysia" pageId="65" pageNumber="956">Sarawak</collectingCountry>
|
||
, as the
|
||
<typeStatus id="1E0724E2FFA0BA60FB0DF807FAA8F87D" box="[1232,1317,1972,1996]" pageId="65" pageNumber="956" type="neotype">neotype</typeStatus>
|
||
of
|
||
<taxonomicName id="06BCE1C3FFA0BA60FA92F807FB30F85A" authority="Targioni Tozzetti, 1877" authorityName="Targioni Tozzetti" authorityYear="1877" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="65" pageNumber="956" phylum="Arthropoda" rank="species" species="cheirogona">
|
||
<emphasis id="F3C84652FFA0BA60FA92F807FC25F85A" italics="true" pageId="65" pageNumber="956">Sesarma cheirogona</emphasis>
|
||
<bibRefCitation id="A52DE7B1FFA0BA60FC6FF860FB30F85A" author="Targioni Tozzetti" box="[946,1213,2003,2027]" firstAuthor="Targioni Tozzetti" pageId="65" pageNumber="956" pagination="1 - 257" refId="ref77991" refString="Targioni Tozzetti A (1877) Crostacei brachyuri e anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R Istituto di Studi Superiori Pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1: i - xxix + 1 - 257, pls. 1 - 12." type="journal article" year="1877">Targioni Tozzetti, 1877</bibRefCitation>
|
||
</taxonomicName>
|
||
. This is necessary to stabilise the taxonomy of the species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="95C3CAC8FFA3BA63FF57F88FFA48F84F" ID-DOI="http://doi.org/10.5281/zenodo.5351365" ID-Zenodo-Dep="5351365" httpUri="https://zenodo.org/record/5351365/files/figure.png" pageId="66" pageNumber="957" startId="66.[138,173,1852,1874]" targetBox="[135,1473,257,1836]" targetPageId="66">
|
||
<paragraph id="C1039A40FFA3BA63FF57F88FFA48F84F" blockId="66.[138,1477,1852,2047]" pageId="66" pageNumber="957">
|
||
Fig. 39.
|
||
<taxonomicName id="06BCE1C3FFA3BA63FF02F88FFE50F8E0" baseAuthorityName="Targioni Tozzetti" baseAuthorityYear="1877" box="[223,477,1852,1873]" class="Malacostraca" family="Sesarmidae" genus="Contusarma" kingdom="Animalia" order="Decapoda" pageId="66" pageNumber="957" phylum="Arthropoda" rank="species" species="cheirogonum">
|
||
<emphasis id="F3C84652FFA3BA63FF02F88FFE50F8E0" box="[223,477,1852,1873]" italics="true" pageId="66" pageNumber="957">Contusarma cheirogonum</emphasis>
|
||
</taxonomicName>
|
||
. A–C, holotype male of
|
||
<taxonomicName id="06BCE1C3FFA3BA63FD05F88FFC2DF8E0" authorityName="Targioni Tozzetti" authorityYear="1877" box="[728,928,1852,1873]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="66" pageNumber="957" phylum="Arthropoda" rank="species" species="cheirogona">
|
||
<emphasis id="F3C84652FFA3BA63FD05F88FFC2DF8E0" box="[728,928,1852,1873]" italics="true" pageId="66" pageNumber="957">Sesarma cheirogona</emphasis>
|
||
</taxonomicName>
|
||
(after
|
||
<bibRefCitation id="A52DE7B1FFA3BA63FC3BF88FFB5FF8E0" author="Targioni Tozzetti" box="[998,1234,1852,1874]" firstAuthor="Targioni Tozzetti" pageId="66" pageNumber="957" pagination="1 - 257" refId="ref77991" refString="Targioni Tozzetti A (1877) Crostacei brachyuri e anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R Istituto di Studi Superiori Pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1: i - xxix + 1 - 257, pls. 1 - 12." type="journal article" year="1877">Targioni Tozzetti, 1877</bibRefCitation>
|
||
: pl. 9 fig. 2); D, E, F–J, K, male (26.3 × 23.0 mm) (ZRC 1995.226), Bako National Park, Sarawak; L–N, neotype male (24.5 × 21.5 mm) (ZRC 1995.225), Bako National Park, Sarawak; O, male (24.5 × 21.3 mm) (ZRC 1964.9.28.140), Johor, Malaysia; P, male (22.3 × 20.5 mm) (ZRC 1964.9.28.127), Saribas, Sarawak; Q, male (28.8 × 25.5 mm) (ZRC 1964.9.28.126), Saribas, Sarawak. A, D, carapace; B, outer view of left chela; C, thoracic sternites and pleon; E, O–Q, pleon; F, L, left G1 (dorsal view, denuded); M, G, H, left distal part of G1 (dorsal view, denuded); N, I, left distal part of G1 (mesial view); J, left distal part of G1 (ventral view, denuded); K, left G2 (denuded). A–C, after
|
||
<bibRefCitation id="A52DE7B1FFA3BA63FAADF87FFE96F84F" author="Targioni Tozzetti" firstAuthor="Targioni Tozzetti" pageId="66" pageNumber="957" pagination="1 - 257" refId="ref77991" refString="Targioni Tozzetti A (1877) Crostacei brachyuri e anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R Istituto di Studi Superiori Pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1: i - xxix + 1 - 257, pls. 1 - 12." type="journal article" year="1877">Targioni Tozzetti (1877</bibRefCitation>
|
||
: pl. 9 fig. 2); D, H, J, K, after
|
||
<bibRefCitation id="A52DE7B1FFA3BA63FD9BF85AFD2BF84E" author="Ng" box="[582,678,2025,2047]" firstAuthor="Ng" pageId="66" pageNumber="957" pagination="181 - 205" refId="ref73590" refString="Ng PKL (1995) The freshwater crabs and prawns (Crustacea: Decapoda) of Bako National Park, Sarawak, Malaysia, with descriptions of one new genus and three new species. Raffles Bulletin of Zoology, 43 (1): 181 - 205." type="journal article" year="1995">Ng (1995</bibRefCitation>
|
||
: fig. 14). Scales: D, E, K, O–Q = 5.0 mm; F, L = 2.0 mm; G–K, M, N = 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="95C3CAC8FFA2BA62FFACF9F2FB72F91C" ID-DOI="http://doi.org/10.5281/zenodo.5351367" ID-Zenodo-Dep="5351367" httpUri="https://zenodo.org/record/5351367/files/figure.png" pageId="67" pageNumber="958" startId="67.[113,148,1601,1623]" targetBox="[118,1453,193,1587]" targetPageId="67">
|
||
<paragraph id="C1039A40FFA2BA62FFACF9F2FB72F91C" blockId="67.[113,1452,1601,1709]" pageId="67" pageNumber="958">
|
||
Fig. 40.
|
||
<taxonomicName id="06BCE1C3FFA2BA62FF1BF9F1FE44F9E6" baseAuthorityName="Miers" baseAuthorityYear="1880" box="[198,457,1602,1623]" class="Malacostraca" family="Sesarmidae" genus="Miersarma" kingdom="Animalia" order="Decapoda" pageId="67" pageNumber="958" phylum="Arthropoda" rank="species" species="granosimanum">
|
||
<emphasis id="F3C84652FFA2BA62FF1BF9F1FE44F9E6" box="[198,457,1602,1623]" italics="true" pageId="67" pageNumber="958">Miersarma granosimanum</emphasis>
|
||
</taxonomicName>
|
||
. A–E, lectotype male (16.9 × 14.8 mm) (NHM 1880.6), Borneo; F–I, male (22.5 × 19.2 mm) (ZRC 1965.7.29.164), Sedili River, Johor, Malaysia; J, male (19.5 × 16.7 mm) (ZRC 1965.7.29.165), Sedili River, Johor, Malaysia. A, male pleon; B, F, left G1 (ventral view, denuded); C, G, J, left G1 (dorsal view, denuded); D, H, left distal part of G1 (ventral view, denuded); E, I, left distal part of G1 (dorsal view, denuded). Scales: A = 5.0 mm; B, C, F, G, J = 1.0 mm; D, E, H, I = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="89A6C9CBFFA2BA62FFACF957FD17F82D" pageId="67" pageNumber="958" type="biology_ecology">
|
||
<paragraph id="C1039A40FFA2BA62FFACF957FD17F82D" blockId="67.[113,752,1764,1949]" pageId="67" pageNumber="958">
|
||
<emphasis id="F3C84652FFA2BA62FFACF957FF4AF94D" bold="true" box="[113,199,1764,1788]" pageId="67" pageNumber="958">Biology</emphasis>
|
||
. Probably very similar to
|
||
<taxonomicName id="06BCE1C3FFA2BA62FE07F956FD7DF94D" baseAuthorityName="Targioni Tozzetti" baseAuthorityYear="1877" box="[474,752,1764,1788]" class="Malacostraca" family="Sesarmidae" genus="Contusarma" kingdom="Animalia" order="Decapoda" pageId="67" pageNumber="958" phylum="Arthropoda" rank="species" species="cheirogonum">
|
||
<emphasis id="F3C84652FFA2BA62FE07F956FD7DF94D" box="[474,752,1764,1788]" italics="true" pageId="67" pageNumber="958">Contusarma cheirogonum</emphasis>
|
||
</taxonomicName>
|
||
(see below), but detailed data or field observations are not available. One recent specimen (ZRC 2000.0952) was obtained from along the Chao Phraya River, from Silom, a site about
|
||
<quantity id="064437A5FFA2BA62FF3CF8D6FEA8F8CD" box="[225,293,1892,1917]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="67" pageNumber="958" unit="km" value="20.0">20 km</quantity>
|
||
from the sea. This is in the main city area of
|
||
<collectingRegion id="037854A2FFA2BA62FF4DF837FF7DF82D" box="[144,240,1924,1948]" country="Thailand" name="Krung Thep Mahanakhon" pageId="67" pageNumber="958">Bangkok</collectingRegion>
|
||
but there is still strong tidal influence.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="89A6C9CBFFA2BA62FFACF877FE87F84C" pageId="67" pageNumber="958" type="distribution">
|
||
<paragraph id="C1039A40FFA2BA62FFACF877FE87F84C" blockId="67.[113,752,1988,2045]" pageId="67" pageNumber="958">
|
||
<emphasis id="F3C84652FFA2BA62FFACF877FF71F86D" bold="true" box="[113,252,1988,2012]" pageId="67" pageNumber="958">Distribution</emphasis>
|
||
. Known only from western coast of
|
||
<collectingCountry id="B9ABDAD0FFA2BA62FD56F877FD60F86D" box="[651,749,1988,2012]" name="Thailand" pageId="67" pageNumber="958">Thailand</collectingCountry>
|
||
, Indian Ocean.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |