treatments-xml/data/49/15/2B/49152B56FFDABA1CFC6DFD0EFEF5FCA4.xml

<document id="E79437AD01FC305CF080010F726A6D5E" ID-CLB-Dataset="55667" ID-DOI="10.26107/RBZ-2020-0097" ID-GBIF-Dataset="6cb155ce-8b9f-48ce-8e6c-b0dc00f8b1cc" ID-ISSN="2345-7600" ID-Zenodo-Dep="5351295" ID-ZooBank="815E4670-B063-4FD8-B31E-3AD89B3A7942" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1630205841501" checkinUser="diego" docAuthor="Schubart, Christoph D. &amp; Ng, Peter K. L." docDate="2020" docId="49152B56FFDABA1CFC6DFD0EFEF5FCA4" docLanguage="en" docName="RafflesBZool.68.891-994.pdf" docOrigin="Raffles Bulletin of Zoology 68" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Pseudosesarma brehieri Ng 2018" docType="treatment" docVersion="12" lastPageNumber="952" masterDocId="B52C532EFFE1BA21FFDDFFB3FF8DFFB1" masterDocTitle="Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species" masterLastPageNumber="994" masterPageNumber="891" pageNumber="950" updateTime="1699203722016" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="40B3F20641AF5CA685D69F740E3612C3">Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species</mods:title>
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<mods:namePart id="35BB9F089E2BD5A4F9961BBE6A8DB8F3">Schubart, Christoph D.</mods:namePart>
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<mods:namePart id="49F3CB690EF2604E85545C5F728DFE7A">Ng, Peter K. L.</mods:namePart>
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<taxonomicName id="06BCE1C3FFDABA1AFC6DFD0EFAA5FD64" ID-CoL="6WGDP" authority="Ng, 2018" authorityName="Ng" authorityYear="2018" box="[944,1320,701,725]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="59" pageNumber="950" phylum="Arthropoda" rank="species" species="brehieri">
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<emphasis id="F3C84652FFDABA1AFC6DFD0EFB37FD64" bold="true" box="[944,1210,701,725]" italics="true" pageId="59" pageNumber="950">Pseudosesarma brehieri</emphasis>
<bibRefCitation id="A52DE7B1FFDABA1AFB1EFD0EFAA5FD64" author="Ng" box="[1219,1320,701,725]" firstAuthor="Ng" pageId="59" pageNumber="950" pagination="185 - 197" refId="ref73639" refString="Ng PKL (2018) On two new species of potamid and sesarmid crabs (Decapoda, Brachyura) from caves in Myanmar. Crustaceana, 91 (2): 185 - 197." type="journal article" year="2018">Ng, 2018</bibRefCitation>
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<paragraph id="C1039A40FFDABA1AFC50FD6EFAC6FD44" blockId="59.[909,1355,701,757]" box="[909,1355,733,757]" pageId="59" pageNumber="950">
(
<figureCitation id="598786C5FFDABA1AFC48FD6EFB8EFD44" box="[917,1027,733,757]" captionStart="Fig" captionStartId="41.[113,148,1553,1575]" captionTargetBox="[113,1451,189,1537]" captionTargetId="figure-14@41.[113,1451,189,1537]" captionText="Fig. 22. Overall habitus, Pseudosesarma species. A, P. edwardsii, lectotype male (17.5 × 16.1 mm) (RMNH-D17a), Mergui Archipelago; B, P. edwardsii, male (13.4 × 11.9 mm) (ZRC 1965.8.2.81), Pulau Pawai, Singapore; C, P. edwardsii, male (12.9 × 11.8 mm) (ZRC 1971.9.24.9), Singapore; D, P. crassimanum, male (15.9 × 14.5 mm) (RMNH-D23313), Singapore; E, P. anteactum, holotype male (16.7 × 14.7 mm) (ZRC 2016.0602), Sri Lanka; F, P. glabrum, holotype male (13.8 × 12.0 mm) (CUSAT 2016-1), Kerala, India; G, P. brehieri, holotype male (17.4 × 15.7 mm) (ZRC 2016.0593), Myanmar; H, P. boulengeri, lectotype male (26.9 × 23.5 mm) (NHM 1919.11.14.1), Basra, Iraq." figureDoi="http://doi.org/10.5281/zenodo.6519005" httpUri="https://zenodo.org/record/6519005/files/figure.png" pageId="59" pageNumber="950">Figs. 22G</figureCitation>
, 
<figureCitation id="598786C5FFDABA1AFBCDFD6EFBB1FD44" box="[1040,1084,733,757]" captionStart="Fig" captionStartId="46.[136,171,1913,1935]" captionTargetBox="[147,1463,189,1899]" captionTargetId="figure-8@46.[147,1463,189,1899]" captionText="Fig. 25. Frontal view of cephalothorax of Pseudosesarma. A, P. edwardsii, lectotype male (17.5 × 16.1 mm) (RMNH-D17a), Mergui Archipelago; B, P. edwardsii, male (13.4 × 11.9 mm) (ZRC 1965.8.2.81), Pulau Pawai, Singapore; C, P. crassimanum, male (15.9 × 14.5 mm) (RMNH-D23313), Singapore; D, P. anteactum, holotype male (16.7 × 14.7 mm) (ZRC 2016.0602), Sri Lanka; E, P. glabrum, holotype male (13.8 × 12.0 mm) (CUSAT 2016-1), Kerala, India; F, P. brehieri, holotype male (17.4 × 15.7 mm) (ZRC 2016.0593), Myanmar; G, P. boulengeri, lectotype male (26.9 × 23.5 mm) (NHM 1919.11.14.1), Basra, Iraq; H, Contusarma bocourti, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand; I, Contusarma cheirogonum, neotype male (24.5 × 21.5 mm) (ZRC 1995.225), Bako National Park, Sarawak; J, Miersarma granosimanum, lectotype male (16.9 × 14.8 mm) (NHM 1880.6), Borneo." figureDoi="http://doi.org/10.5281/zenodo.6519007" httpUri="https://zenodo.org/record/6519007/files/figure.png" pageId="59" pageNumber="950">25F</figureCitation>
, 
<figureCitation id="598786C5FFDABA1AFB94FD6EFBF7FD44" box="[1097,1146,733,757]" captionStart="Fig" captionStartId="49.[113,148,1885,1907]" captionTargetBox="[113,1451,246,1869]" captionTargetId="figure-14@49.[113,1451,246,1869]" captionText="Fig. 27. Outer view of chela of Pseudosesarma. A, P. edwardsii, lectotype male (17.5 × 16.1 mm) (RMNH-D17a), Mergui Archipelago; B, P. edwardsii, male (13.4 × 11.9 mm) (ZRC 1965.8.2.81), Pulau Pawai, Singapore; C, P. edwardsii, male (19.4 × 17.3 mm) (ZRC 1971.9.24.8), Singapore; D, P. crassimanum, male (15.9 × 14.5 mm) (RMNH-D23313), Singapore; E, P. anteactum, holotype male (16.7 × 14.7 mm) (ZRC 2016.0602), Sri Lanka; F, P. glabrum, holotype male (13.8 × 12.0 mm) (CUSAT 2016-1), Kerala, India; G, P. brehieri, holotype male (17.4 × 15.7 mm) (ZRC 2016.0593), Myanmar; H, P. boulengeri, lectotype male (26.9 × 23.5 mm) (NHM 1919.11.14.1), Basra, Iraq." figureDoi="http://doi.org/10.5281/zenodo.6519009" httpUri="https://zenodo.org/record/6519009/files/figure.png" pageId="59" pageNumber="950">27G</figureCitation>
, 
<figureCitation id="598786C5FFDABA1AFB5BFD6EFB3BFD44" box="[1158,1206,733,757]" captionStart="Fig" captionStartId="52.[136,171,1939,1961]" captionTargetBox="[173,1438,189,1924]" captionTargetId="figure-7@52.[173,1438,189,1924]" captionText="Fig. 30. Male anterior thoracic sternum and pleon of Pseudosesarma. A, P edwardsii, lectotype male (17.5 × 16.1 mm) (RMNH-D17a), Mergui Archipelago; B, P. edwardsii, male (13.4 × 11.9 mm) (ZRC 1965.8.2.81), Pulau Pawai, Singapore; C, P. crassimanum, male (15.9 × 14.5 mm) (RMNH-D23313), Singapore; D, P. glabrum, holotype male (13.8 × 12.0 mm) (CUSAT 2016-1), Kerala, India; E, P. anteactum, holotype male (16.7 × 14.7 mm) (ZRC 2016.0602), Sri Lanka; F, P. anteactum, paratype male (21.2 × 19.7 mm) (ZRC 2016.0603), Sri Lanka; G, P. brehieri, holotype male (17.4 × 15.7 mm) (ZRC 2016.0593), Myanmar; H, P. boulengeri, lectotype male (26.9 × 23.5 mm) (NHM 1919.11.14.1), Basra, Iraq." figureDoi="http://doi.org/10.5281/zenodo.6519011" httpUri="https://zenodo.org/record/6519011/files/figure.png" pageId="59" pageNumber="950">30G</figureCitation>
, 
<figureCitation id="598786C5FFDABA1AFB19FD6EFA86FD44" box="[1220,1291,733,757]" captionStart="Fig" captionStartId="64.[136,171,1926,1948]" captionTargetBox="[138,1476,190,1907]" captionTargetId="figure-8@64.[124,1486,183,1911]" captionText="Fig. 37. A–G, Pseudosesarma brehieri, paratype female (14.9 × 13.4 mm) (ZRC 2016.0594), Myanmar; F, G, P. brehieri, male (15.0 × 13.6 mm) (ZRC 2013.0209), West Bengal, India; H, I, P. brehieri, male (14.5 × 13.0 mm) (ZRC 2013.0209), West Bengal, India; J–O, P. boulengeri, lectotype male (26.9 × 23.5 mm) (NHM 1919.11.14.1), Basra, Iraq. A, K, male pleon; B, F, L, left G1 (ventral view, denuded); D, G, M, left G1 (dorsal view, denuded); E, left G2; H, O, left distal part of G1 (dorsal view, denuded); I, N, left distal part of G1 (ventral view, denuded); J, anterior thoracic sternites 1–4. Scales: A = 4.0 mm; B–E, N, O = 1.0 mm; F–I = 0.5 mm; J, K = 5.0 mm; L, M = 2.0 mm. A–E, after Ng (2018: fig. 7E–I)." figureDoi="http://doi.org/10.5281/zenodo.6519019" httpUri="https://zenodo.org/record/6519019/files/figure.png" pageId="59" pageNumber="950">37A–I</figureCitation>
, 
<figureCitation id="598786C5FFDABA1AFACAFD6EFABEFD44" box="[1303,1331,733,757]" captionStart="Fig" captionStartId="74.[136,171,1898,1920]" captionTargetBox="[136,1474,211,1884]" captionTargetId="figure-7@74.[136,1474,211,1884]" captionTargetPageId="74" captionText="Fig. 44. Vulva. A, Pseudosesarma edwardsii, female (16.5 × 15.3 mm) (ZRC 2016.0608), Malaysia; B, Pseudosesarma crassimanum, female (15.9 × 13.8 mm) (ZRC 2000.1768), Malaysia; C, Pseudosesarma crassimanum, female (18.0 × 15.7 mm) (ZRC 2017.1045), Thailand; D, Pseudosesarma anteactum, paratype female (17.5 × 15.4 mm) (ZRC 2016.0603), Sri Lanka; E, Pseudosesarma brehieri, paratype female (14.9 × 13.4 mm) (ZRC 2016.0594), Myanmar; F, Pseudosesarma boulengeri, female (24.2 × 20.0 mm) (ZRC 2014.0335), Iran; G, Contusarma bocourti, female (21.1 × 19.0 mm) (ZRC 2019.1114), Thailand; H, Contusarma cheirogonum, female (26.6 × 23.0 mm) (ZRC 1964.9.28.127), Sarawak; I, Miersarma granosimanum, female (20.1 × 16.4 mm) (ZRC 1965.7.29.166), Malaysia; J, Manarma moeschii, female (19.3 × 17.3 mm) (ZRC 2017.0170), Thailand." figureDoi="http://doi.org/10.5281/zenodo.5351375" httpUri="https://zenodo.org/record/5351375/files/figure.png" pageId="59" pageNumber="950">44</figureCitation>
E)
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<emphasis id="F3C84652FFDABA1AFCF0FCAFFB94FC80" box="[813,1049,796,817]" italics="true" pageId="59" pageNumber="950">Pseudosesarma brehieri</emphasis>
<bibRefCitation id="A52DE7B1FFDABA1AFBFFFCAFFB3EFC80" author="Ng" box="[1058,1203,796,817]" firstAuthor="Ng" page="191" pageId="59" pageNumber="950" pagination="185 - 197" refId="ref73639" refString="Ng PKL (2018) On two new species of potamid and sesarmid crabs (Decapoda, Brachyura) from caves in Myanmar. Crustaceana, 91 (2): 185 - 197." type="journal article" year="2018">Ng, 2018: 191</bibRefCitation>
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, figs. 4–6, 7D–I; 
<treatmentCitation id="401DBC51FFDABA1AFAB8FCAFFC50FCFF" author="Trivedi J &amp; Mitra S &amp; Patel P &amp; Gosavi S &amp; Vachhrajani K" page="4" pageId="59" pageNumber="950" year="2020">
<bibRefCitation id="A52DE7B1FFDABA1AFAB8FCAFFC50FCFF" author="Trivedi" etAl="et al." firstAuthor="Trivedi" page="4" pageId="59" pageNumber="950" pagination="1 - 8" refId="ref78280" refString="Trivedi J, Mitra S &amp; Patel P, Gosavi S &amp; Vachhrajani K (2020) Four new records of brachyuran crabs (Crustacea: Decapoda: Brachyura) from India. Thalassas (2020). doi: 10.1007 / s 41208 - 020 - 00252 - 6, pp. 1 - 8." type="journal article" year="2020">Trivedi et al., 2020: 4</bibRefCitation>
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, fig. 3.
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<emphasis id="F3C84652FFDABA1AFCF0FCC4FB9CFC3E" bold="true" box="[813,1041,887,911]" pageId="59" pageNumber="950">Material examined</emphasis>
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<materialsCitation id="71D4901DFFDABA1AFBFEFCC4FBA8FBBE" ID-GBIF-Occurrence="3763237348" collectingDate="2016-11-26" collectionCode="ZRC" collectorName="F. Brehier" country="Myanmar" latitude="16.525972" location="Nathack Gu" longLatPrecision="1" longitude="97.713554" municipality="Mawlamyine" pageId="59" pageNumber="950" specimenCode="ZRC 2016.0593" specimenCount="1" specimenCount-male="1" stateProvince="Mon State" typeStatus="holotype">
<typeStatus id="1E0724E2FFDABA1AFBFEFCC4FB02FC3E" box="[1059,1167,887,911]" pageId="59" pageNumber="950" type="holotype">Holotype</typeStatus>
: 
<specimenCount id="D7BA51C9FFDABA1AFB40FCC4FB5EFC3E" box="[1181,1235,887,911]" pageId="59" pageNumber="950" type="male">male</specimenCount>
(17.4 × 
<quantity id="064437A5FFDABA1AFAE0FCC4FA29FC3E" box="[1341,1444,887,911]" metricMagnitude="-2" metricUnit="m" metricValue="1.5699999999999998" pageId="59" pageNumber="950" unit="mm" value="15.7">15.7 mm</quantity>
) (
<specimenCode id="911A323BFFDABA1AFCE8FC24FC7DFC1E" box="[821,1008,919,943]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="59" pageNumber="950">ZRC 2016.0593</specimenCode>
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<location id="C463CC9BFFDABA1AFBDAFC24FB02FC1E" LSID="urn:lsid:plazi:treatment:49152B56FFDABA1CFC6DFD0EFEF5FCA4:C463CC9BFFDABA1AFBDAFC24FB02FC1E" box="[1031,1167,919,943]" country="Myanmar" county="Mon State" latitude="16.525972" longLatPrecision="1" longitude="97.713554" municipality="Mawlamyine" name="Nathack Gu" pageId="59" pageNumber="950" stateProvince="Mon State">Nathack Gu</location>
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<location id="C463CC9BFFDABA1AFB7CFC24FAEBFC1E" LSID="urn:lsid:plazi:treatment:49152B56FFDABA1CFC6DFD0EFEF5FCA4:C463CC9BFFDABA1AFB7CFC24FAEBFC1E" box="[1185,1382,919,943]" country="Myanmar" county="Mon State" latitude="16.525972" longLatPrecision="1" longitude="97.713554" municipality="Mawlamyine" name="Two Level Cave" pageId="59" pageNumber="950" stateProvince="Mon State">Two Level Cave</location>
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<locationDeviation id="D0B18362FFDABA1AFAA0FC24FC6CFC7E" pageId="59" pageNumber="950">near Saddan Sin Gu</locationDeviation>
, 
<collectingMunicipality id="2167003AFFDABA1AFC2CFC04FB08FC7E" box="[1009,1157,951,975]" pageId="59" pageNumber="950">Mawlamyine</collectingMunicipality>
(
<collectingMunicipality id="2167003AFFDABA1AFB45FC04FA99FC7E" box="[1176,1300,951,975]" pageId="59" pageNumber="950">Moulmein</collectingMunicipality>
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<collectingRegion id="037854A2FFDABA1AFAF1FC04FA2AFC7E" box="[1324,1447,951,975]" pageId="59" pageNumber="950">Mon State</collectingRegion>
, 
<geoCoordinate id="A488FC87FFDABA1AFCF0FC66FC4CFC5F" box="[813,961,981,1006]" degrees="16" direction="north" minutes="31" orientation="latitude" pageId="59" pageNumber="950" precision="1" seconds="33.5" value="16.525972">16°31′33.5″N</geoCoordinate>
<geoCoordinate id="A488FC87FFDABA1AFC14FC66FBE4FC5F" box="[969,1129,981,1006]" degrees="097" direction="east" minutes="42" orientation="longitude" pageId="59" pageNumber="950" precision="1" seconds="48.8" value="97.713554">097°42′48.8″E</geoCoordinate>
, 
<collectingCountry id="B9ABDAD0FFDABA1AFBA8FC64FB52FC5E" box="[1141,1247,983,1007]" pageId="59" pageNumber="950">Myanmar</collectingCountry>
, coll. 
<collectorName id="6C49FF96FFDABA1AFAFEFC64FA1CFC5E" box="[1315,1425,983,1007]" pageId="59" pageNumber="950">F. Bréhier</collectorName>
et al., 
<date id="B502BC80FFDABA1AFC8FFC44FBACFBBE" box="[850,1057,1015,1039]" pageId="59" pageNumber="950" value="2016-11-26">
<collectingDate id="A5464568FFDABA1AFC8FFC44FBACFBBE" box="[850,1057,1015,1039]" pageId="59" pageNumber="950" value="2016-11-26">26 November 2016</collectingDate>
</date>
.
</materialsCitation>
<materialsCitation id="71D4901DFFDABA1AFBF7FC44FB6FFB9E" ID-GBIF-Occurrence="3763237470" collectingDate="2016-11-26" collectionCode="ZRC" collectorName="Myanmar &amp; F. Brehier" country="India" county="Mon State" latitude="16.525972" location="Others" longLatPrecision="1" longitude="97.713554" municipality="Moulmein" pageId="59" pageNumber="950" specimenCode="ZRC 2016.0594" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="1E0724E2FFDABA1AFBF7FC44FB01FBBE" box="[1066,1164,1015,1039]" pageId="59" pageNumber="950" type="paratype">Paratype</typeStatus>
: 
<specimenCount id="D7BA51C9FFDABA1AFB49FC44FB63FBBE" box="[1172,1262,1015,1039]" pageId="59" pageNumber="950" type="female">1 female</specimenCount>
(14.9 × 
<quantity id="064437A5FFDABA1AFA98FC44FA29FBBE" box="[1349,1444,1015,1039]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="59" pageNumber="950" unit="mm" value="13.4">13.4 mm</quantity>
) (
<specimenCode id="911A323BFFDABA1AFCE9FBA4FC68FB9E" box="[820,997,1047,1071]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="59" pageNumber="950">ZRC 2016.0594</specimenCode>
), same data as holotype.
</materialsCitation>
<materialsCitation id="71D4901DFFDABA1AFB3AFBA4FB00FB1E" ID-GBIF-Occurrence="3763237334" collectingDate="2016-11-26" collectionCode="ZRC" collectorName="Myanmar &amp; F. Brehier" country="India" county="Mon State" latitude="16.525972" location="Others" longLatPrecision="1" longitude="97.713554" municipality="Moulmein" pageId="59" pageNumber="950" specimenCode="ZRC 2016.0594" specimenCount="1" specimenCount-female="1" specimenCount-male="5" stateProvince="West Bengal" typeStatus="paratype">
<location id="C463CC9BFFDABA1AFB3AFBA4FABFFB9E" LSID="urn:lsid:plazi:treatment:49152B56FFDABA1CFC6DFD0EFEF5FCA4:C463CC9BFFDABA1AFB3AFBA4FABFFB9E" box="[1255,1330,1047,1071]" country="India" name="Others" pageId="59" pageNumber="950">Others</location>
: 
<collectingCountry id="B9ABDAD0FFDABA1AFAE7FBA4FA08FB9E" box="[1338,1413,1047,1071]" name="India" pageId="59" pageNumber="950">INDIA</collectingCountry>
– 
<specimenCount id="D7BA51C9FFDABA1AFA43FBA4FCE6FBFE" pageId="59" pageNumber="950" type="male">5 males</specimenCount>
(11.9 × 
<quantity id="064437A5FFDABA1AFC17FB84FBA0FBFF" box="[970,1069,1079,1103]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="59" pageNumber="950" unit="mm" value="10.4">10.4 mm</quantity>
, 12.4 × 
<quantity id="064437A5FFDABA1AFB5AFB84FB67FBFF" box="[1159,1258,1079,1103]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="59" pageNumber="950" unit="mm" value="10.9">10.9 mm</quantity>
, 13.6 × 
<quantity id="064437A5FFDABA1AFA99FB84FA2AFBFF" box="[1348,1447,1079,1103]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="59" pageNumber="950" unit="mm" value="11.7">11.7 mm</quantity>
, 14.5 × 13.0 mm, 15.0 × 
<quantity id="064437A5FFDABA1AFBE4FBE4FB16FBDE" box="[1081,1179,1111,1135]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="59" pageNumber="950" unit="mm" value="13.6">13.6 mm</quantity>
) (
<specimenCode id="911A323BFFDABA1AFB6FFBE4FAEBFBDE" box="[1202,1382,1111,1135]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="59" pageNumber="950">ZRC 2013.0209</specimenCode>
), river bank, 
<collectorName id="6C49FF96FFDABA1AFCACFBC4FC50FB3E" box="[881,989,1143,1167]" pageId="59" pageNumber="950">Bhagirath</collectorName>
, 
<collectorName id="6C49FF96FFDABA1AFC37FBC4FBEFFB3E" box="[1002,1122,1143,1167]" pageId="59" pageNumber="950">Nabadiwip</collectorName>
, 
<collectorName id="6C49FF96FFDABA1AFBADFBC4FB3FFB3E" box="[1136,1202,1143,1167]" pageId="59" pageNumber="950">Nadia</collectorName>
, 
<collectingRegion id="037854A2FFDABA1AFB1DFBC4FAC3FB3E" box="[1216,1358,1143,1167]" country="India" name="West Bengal" pageId="59" pageNumber="950">West Bengal</collectingRegion>
, coll. 
<collectorName id="6C49FF96FFDABA1AFA49FBC4FCE0FB1E" pageId="59" pageNumber="950">Z. Jaafar</collectorName>
et al., 
<date id="B502BC80FFDABA1AFC67FB24FB00FB1E" box="[954,1165,1175,1199]" pageId="59" pageNumber="950" value="2004-12-26">
<collectingDate id="A5464568FFDABA1AFC67FB24FB00FB1E" box="[954,1165,1175,1199]" pageId="59" pageNumber="950" value="2004-12-26">26 December 2004</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFDABA1AFCF0FB64FB80F91E" pageId="59" pageNumber="950" type="diagnosis">
<paragraph id="C1039A40FFDABA1AFCF0FB64FB2FF9DE" blockId="59.[812,1452,1239,1647]" pageId="59" pageNumber="950">
<emphasis id="F3C84652FFDABA1AFCF0FB64FC14FB5E" bold="true" box="[813,921,1239,1263]" pageId="59" pageNumber="950">Diagnosis</emphasis>
. Carapace quadrate, slightly wider than long, width to length ratio 1.1; frontal margin wide, median concavity separating lobes relatively shallow; external orbital tooth short, not reaching to level of front, lateral margin with 2 epibranchial teeth; posterolateral margins subparallel; chela short, stout, fingers shorter than palm, outer surface of chela covered with numerous granules, ventral margin of palm almost straight or gently convex; suture between male thoracic sternites 3 and 4 distinct; male pleon triangular, somite 6 wide; male sternopleonal cavity with press-button of pleonal locking mechanism on sternite 5; G1 stout, distal part prominently dilated forming bulbous structure, chitinous tip short, broad, truncate, fan-like.
</paragraph>
<paragraph id="C1039A40FFDABA1AFCF0F924FB80F91E" blockId="59.[813,1037,1687,1711]" box="[813,1037,1687,1711]" pageId="59" pageNumber="950">
<emphasis id="F3C84652FFDABA1AFCF0F924FCF0F91E" bold="true" box="[813,893,1687,1711]" pageId="59" pageNumber="950">Colour</emphasis>
. 
<collectingRegion id="037854A2FFDABA1AFC51F924FC39F91F" box="[908,948,1687,1710]" country="Cameroon" name="North" pageId="59" pageNumber="950">Not</collectingRegion>
known.
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFDABA1AFCF0F964FAADF7BE" pageId="59" pageNumber="950" type="discussion">
<paragraph id="C1039A40FFDABA1AFCF0F964FAADF7BE" blockId="59.[813,1452,1751,2063]" pageId="59" pageNumber="950">
<emphasis id="F3C84652FFDABA1AFCF0F964FC19F95E" bold="true" box="[813,916,1751,1775]" pageId="59" pageNumber="950">Remarks</emphasis>
. The species was described and figured in detail by 
<bibRefCitation id="A52DE7B1FFDABA1AFCF0F944FC13F8BE" author="Ng" box="[813,926,1783,1807]" firstAuthor="Ng" pageId="59" pageNumber="950" pagination="185 - 197" refId="ref73639" refString="Ng PKL (2018) On two new species of potamid and sesarmid crabs (Decapoda, Brachyura) from caves in Myanmar. Crustaceana, 91 (2): 185 - 197." type="journal article" year="2018">Ng (2018)</bibRefCitation>
. 
<taxonomicName id="06BCE1C3FFDABA1AFC74F944FB20F8BE" authorityName="Ng" authorityYear="2018" box="[937,1197,1783,1807]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="59" pageNumber="950" phylum="Arthropoda" rank="species" species="brehieri">
<emphasis id="F3C84652FFDABA1AFC74F944FB20F8BE" box="[937,1197,1783,1807]" italics="true" pageId="59" pageNumber="950">Pseudosesarma brehieri</emphasis>
</taxonomicName>
most closely resembles 
<taxonomicName id="06BCE1C3FFDABA1AFCF0F8ABFC56F89E" baseAuthorityName="De Man" baseAuthorityYear="1887" box="[813,987,1816,1839]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="59" pageNumber="950" phylum="Arthropoda" rank="species" species="crassimanum">
<emphasis id="F3C84652FFDABA1AFCF0F8ABFC56F89E" box="[813,987,1816,1839]" italics="true" pageId="59" pageNumber="950">P. crassimanum</emphasis>
</taxonomicName>
but can easily be separated by the ventral margin of its adult male chela being almost straight (
<figureCitation id="598786C5FFDABA1AFA5DF884FCD1F8DE" captionStart="Fig" captionStartId="49.[113,148,1885,1907]" captionTargetBox="[113,1451,246,1869]" captionTargetId="figure-14@49.[113,1451,246,1869]" captionText="Fig. 27. Outer view of chela of Pseudosesarma. A, P. edwardsii, lectotype male (17.5 × 16.1 mm) (RMNH-D17a), Mergui Archipelago; B, P. edwardsii, male (13.4 × 11.9 mm) (ZRC 1965.8.2.81), Pulau Pawai, Singapore; C, P. edwardsii, male (19.4 × 17.3 mm) (ZRC 1971.9.24.8), Singapore; D, P. crassimanum, male (15.9 × 14.5 mm) (RMNH-D23313), Singapore; E, P. anteactum, holotype male (16.7 × 14.7 mm) (ZRC 2016.0602), Sri Lanka; F, P. glabrum, holotype male (13.8 × 12.0 mm) (CUSAT 2016-1), Kerala, India; G, P. brehieri, holotype male (17.4 × 15.7 mm) (ZRC 2016.0593), Myanmar; H, P. boulengeri, lectotype male (26.9 × 23.5 mm) (NHM 1919.11.14.1), Basra, Iraq." figureDoi="http://doi.org/10.5281/zenodo.6519009" httpUri="https://zenodo.org/record/6519009/files/figure.png" pageId="59" pageNumber="950">Fig. 27G</figureCitation>
) and the distal half of the G1 being proportionately more dilated with the chitinous distal part very wide and fan-like (
<figureCitation id="598786C5FFDABA1AFC4CF824FBC4F81E" box="[913,1097,1943,1967]" captionStart="Fig" captionStartId="64.[136,171,1926,1948]" captionTargetBox="[138,1476,190,1907]" captionTargetId="figure-8@64.[124,1486,183,1911]" captionText="Fig. 37. A–G, Pseudosesarma brehieri, paratype female (14.9 × 13.4 mm) (ZRC 2016.0594), Myanmar; F, G, P. brehieri, male (15.0 × 13.6 mm) (ZRC 2013.0209), West Bengal, India; H, I, P. brehieri, male (14.5 × 13.0 mm) (ZRC 2013.0209), West Bengal, India; J–O, P. boulengeri, lectotype male (26.9 × 23.5 mm) (NHM 1919.11.14.1), Basra, Iraq. A, K, male pleon; B, F, L, left G1 (ventral view, denuded); D, G, M, left G1 (dorsal view, denuded); E, left G2; H, O, left distal part of G1 (dorsal view, denuded); I, N, left distal part of G1 (ventral view, denuded); J, anterior thoracic sternites 1–4. Scales: A = 4.0 mm; B–E, N, O = 1.0 mm; F–I = 0.5 mm; J, K = 5.0 mm; L, M = 2.0 mm. A–E, after Ng (2018: fig. 7E–I)." figureDoi="http://doi.org/10.5281/zenodo.6519019" httpUri="https://zenodo.org/record/6519019/files/figure.png" pageId="59" pageNumber="950">Fig. 37B–D, F–I</figureCitation>
) (versus ventral margin of adult male chela gently concave, and the G1 distal part less dilated with the chitinous distal part narrow and beak-like in 
<taxonomicName id="06BCE1C3FFDABA1AFA49F86BFC36F7BE" baseAuthorityName="De Man" baseAuthorityYear="1887" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="59" pageNumber="950" phylum="Arthropoda" rank="species" species="crassimanum">
<emphasis id="F3C84652FFDABA1AFA49F86BFC36F7BE" italics="true" pageId="59" pageNumber="950">P. crassimanum</emphasis>
</taxonomicName>
; 
<figureCitation id="598786C5FFDABA1AFC16F844FBB5F7BE" box="[971,1080,2039,2063]" captionStart="Fig" captionStartId="49.[113,148,1885,1907]" captionTargetBox="[113,1451,246,1869]" captionTargetId="figure-14@49.[113,1451,246,1869]" captionText="Fig. 27. Outer view of chela of Pseudosesarma. A, P. edwardsii, lectotype male (17.5 × 16.1 mm) (RMNH-D17a), Mergui Archipelago; B, P. edwardsii, male (13.4 × 11.9 mm) (ZRC 1965.8.2.81), Pulau Pawai, Singapore; C, P. edwardsii, male (19.4 × 17.3 mm) (ZRC 1971.9.24.8), Singapore; D, P. crassimanum, male (15.9 × 14.5 mm) (RMNH-D23313), Singapore; E, P. anteactum, holotype male (16.7 × 14.7 mm) (ZRC 2016.0602), Sri Lanka; F, P. glabrum, holotype male (13.8 × 12.0 mm) (CUSAT 2016-1), Kerala, India; G, P. brehieri, holotype male (17.4 × 15.7 mm) (ZRC 2016.0593), Myanmar; H, P. boulengeri, lectotype male (26.9 × 23.5 mm) (NHM 1919.11.14.1), Basra, Iraq." figureDoi="http://doi.org/10.5281/zenodo.6519009" httpUri="https://zenodo.org/record/6519009/files/figure.png" pageId="59" pageNumber="950">Figs. 27D</figureCitation>
, 
<figureCitation id="598786C5FFDABA1AFB98F844FBEDF7BE" box="[1093,1120,2039,2063]" captionStart="Fig" captionStartId="60.[136,171,2006,2028]" captionTargetBox="[204,1406,196,1992]" captionTargetId="figure-8@60.[201,1408,185,1996]" captionTargetPageId="60" captionText="Fig. 35. A–E, Pseudosesarma crassimanum. A–G, lectotype male (16.3 × 14.6 mm) (NHM 1886.52b), Myanmar; H–M, male (15.9 × 14.3 mm) (ZRC 2008.0442), Ranong; N–R, male (15.9 × 14.5 mm) (RMNH-D23313), Singapore. A, N, pleon; B–F, right G1; G, right G2; H–L, O–R, left G1; M, left G2. Scales: A = 2.0 mm; B, G, H, I, M, O, P = 1.0 mm; N = 5.0 mm; C–F, J–L, Q, R = 0.5 mm. After Ng &amp; Schubart (2017: fig. 11)." figureDoi="http://doi.org/10.5281/zenodo.5351361" httpUri="https://zenodo.org/record/5351361/files/figure.png" pageId="59" pageNumber="950">35</figureCitation>
B–F, H–L, O–R).
</paragraph>
</subSubSection>
<caption id="95C3CAC8FFDDBA1DFF55F865FE1EF7F2" ID-DOI="http://doi.org/10.5281/zenodo.5351361" ID-Zenodo-Dep="5351361" httpUri="https://zenodo.org/record/5351361/files/figure.png" pageId="60" pageNumber="951" startId="60.[136,171,2006,2028]" targetBox="[204,1406,196,1992]" targetPageId="60">
<paragraph id="C1039A40FFDDBA1DFF55F865FE1EF7F2" blockId="60.[136,1474,2006,2115]" pageId="60" pageNumber="951">
Fig. 35. A–E, 
<taxonomicName id="06BCE1C3FFDDBA1DFECEF865FDBDF85D" baseAuthorityName="De Man" baseAuthorityYear="1887" box="[275,560,2006,2028]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="60" pageNumber="951" phylum="Arthropoda" rank="species" species="crassimanum">
<emphasis id="F3C84652FFDDBA1DFECEF865FDBDF85D" box="[275,560,2006,2028]" italics="true" pageId="60" pageNumber="951">Pseudosesarma crassimanum</emphasis>
</taxonomicName>
. A–G, lectotype male (16.3 × 14.6 mm) (NHM 1886.52b), Myanmar; H–M, male (15.9 × 14.3 mm) (ZRC 2008.0442), Ranong; N–R, male (15.9 × 14.5 mm) (RMNH-D23313), Singapore. A, N, pleon; B–F, right G1; G, right G2; H–L, O–R, left G1; M, left G2. Scales: A = 2.0 mm; B, G, H, I, M, O, P = 1.0 mm; N = 5.0 mm; C–F, J–L, Q, R = 0.5 mm. After 
<bibRefCitation id="A52DE7B1FFDDBA1DFA78F7A3FEB6F7F3" author="Ng &amp; Schubart" firstAuthor="Ng" pageId="60" pageNumber="951" pagination="655 - 669" refId="ref74064" refString="Ng PKL &amp; Schubart CD (2017) On the taxonomy of Pseudosesarma edwardsii (De Man, 1887) and P. crassimanum (De Man, 1887) (Crustacea: Decapoda: Brachyura: Sesarmidae), with description of a new species from Sri Lanka. Raffles Bulletin of Zoology, 65: 655 - 669." type="journal article" year="2017">Ng &amp; Schubart (2017</bibRefCitation>
: fig. 11).
</paragraph>
</caption>
<subSubSection id="89A6C9CBFFDCBA1CFFACFF0EFEEAFD24" pageId="61" pageNumber="952" type="biology_ecology">
<paragraph id="C1039A40FFDCBA1CFFACFF0EFEEAFD24" blockId="61.[113,753,189,661]" pageId="61" pageNumber="952">
<emphasis id="F3C84652FFDCBA1CFFACFF0EFF40FF64" bold="true" box="[113,205,189,213]" pageId="61" pageNumber="952">Biology</emphasis>
. 
<bibRefCitation id="A52DE7B1FFDCBA1CFF3FFF0DFE19FF64" author="Ng" box="[226,404,189,214]" firstAuthor="Ng" page="193" pageId="61" pageNumber="952" pagination="185 - 197" refId="ref73639" refString="Ng PKL (2018) On two new species of potamid and sesarmid crabs (Decapoda, Brachyura) from caves in Myanmar. Crustaceana, 91 (2): 185 - 197." type="journal article" year="2018">Ng (2018: 193</bibRefCitation>
, 195) observed that the 
<typeStatus id="1E0724E2FFDCBA1CFD62FF0DFD62FF67" box="[703,751,190,214]" pageId="61" pageNumber="952">type</typeStatus>
specimens of 
<taxonomicName id="06BCE1C3FFDCBA1CFECBFF6DFE18FF44" authorityName="Ng" authorityYear="2018" box="[278,405,221,245]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="61" pageNumber="952" phylum="Arthropoda" rank="species" species="brehieri">
<emphasis id="F3C84652FFDCBA1CFECBFF6DFE18FF44" box="[278,405,221,245]" italics="true" pageId="61" pageNumber="952">P. brehieri</emphasis>
</taxonomicName>
were found in a small upper chamber of a cave at the foot of the hill not far from 
<location id="C463CC9BFFDCBA1CFD7FFF4EFF4DFE84" LSID="urn:lsid:plazi:treatment:49152B56FFDABA1CFC6DFD0EFEF5FCA4:C463CC9BFFDCBA1CFD7FFF4EFF4DFE84" country="Myanmar" name="Saddan Sin Gu" pageId="61" pageNumber="952">Saddan Sin Gu</location>
in southern 
<collectingCountry id="B9ABDAD0FFDCBA1CFE92FEADFE36FE87" box="[335,443,286,310]" name="Myanmar" pageId="61" pageNumber="952">Myanmar</collectingCountry>
. The water in the cave was completely fresh. The site is not far from the sea and the species is clearly not an obligate cave species with well developed eyes and normal colouration. 
<bibRefCitation id="A52DE7B1FFDCBA1CFDC3FECEFF05FE04" author="Trivedi" etAl="et al." firstAuthor="Trivedi" page="4" pageId="61" pageNumber="952" pagination="1 - 8" refId="ref78280" refString="Trivedi J, Mitra S &amp; Patel P, Gosavi S &amp; Vachhrajani K (2020) Four new records of brachyuran crabs (Crustacea: Decapoda: Brachyura) from India. Thalassas (2020). doi: 10.1007 / s 41208 - 020 - 00252 - 6, pp. 1 - 8." type="journal article" year="2020">Trivedi et al. (2020: 4)</bibRefCitation>
recorded the species from 
<collectingCountry id="B9ABDAD0FFDCBA1CFE73FE2EFE6AFE04" box="[430,487,413,437]" name="India" pageId="61" pageNumber="952">India</collectingCountry>
, noting that the material was “collected from [the] muddy shore of 
<location id="C463CC9BFFDCBA1CFDEAFE0EFD5BFE64" LSID="urn:lsid:plazi:treatment:49152B56FFDABA1CFC6DFD0EFEF5FCA4:C463CC9BFFDCBA1CFDEAFE0EFD5BFE64" box="[567,726,445,469]" country="India" name="Hooghly River" pageId="61" pageNumber="952" stateProvince="West Bengal">Hooghly River</location>
at Barrackpore city of 
<collectingRegion id="037854A2FFDCBA1CFE90FE6DFD9CFE44" box="[333,529,477,501]" country="India" name="West Bengal" pageId="61" pageNumber="952">West Bengal state</collectingRegion>
which is considered as upper reaches of Hooghly-Matla estuary, where tidal effects usually occur with a low salinity (0.5 to 4 ppm) of interstitial waters”. The present specimens from 
<collectingCountry id="B9ABDAD0FFDCBA1CFDA3FD8EFD38FDE4" box="[638,693,573,597]" name="India" pageId="61" pageNumber="952">India</collectingCountry>
were collected from the side of a riverbank in an estuarine habitat in 
<collectingRegion id="037854A2FFDCBA1CFF52FDCDFE91FD24" box="[143,284,637,661]" country="India" name="West Bengal" pageId="61" pageNumber="952">West Bengal</collectingRegion>
, 
<collectingCountry id="B9ABDAD0FFDCBA1CFEF4FDCEFEEEFD24" box="[297,355,637,661]" name="India" pageId="61" pageNumber="952">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFDCBA1CFFACFD0EFEF5FCA4" pageId="61" pageNumber="952" type="distribution">
<paragraph id="C1039A40FFDCBA1CFFACFD0EFEF5FCA4" blockId="61.[113,753,701,789]" pageId="61" pageNumber="952">
<emphasis id="F3C84652FFDCBA1CFFACFD0EFE8DFD64" bold="true" box="[113,256,701,725]" pageId="61" pageNumber="952">Distribution</emphasis>
. The species was described from 
<collectingCountry id="B9ABDAD0FFDCBA1CFD5AFD0DFD7DFD67" box="[647,752,702,726]" name="Myanmar" pageId="61" pageNumber="952">Myanmar</collectingCountry>
and we now also have specimens from Bengal in eastern 
<collectingCountry id="B9ABDAD0FFDCBA1CFFACFD4EFF25FCA4" box="[113,168,765,789]" name="India" pageId="61" pageNumber="952">India</collectingCountry>
(ZRC 2013.0209).
</paragraph>
</subSubSection>
</treatment>
</document>