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<mods:title id="3C7C42F283AE796EF8571098E015ABEF">Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species</mods:title>
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<emphasis id="F3C84652FFD4BA14FF18FCEEFE6CFCC4" bold="true" box="[197,481,861,885]" italics="true" pageId="53" pageNumber="944">Pseudosesarma edwardsii</emphasis>
(
<bibRefCitation id="A52DE7B1FFD4BA14FE2CFCEEFD1BFCC4" author="De Man" box="[497,662,861,885]" firstAuthor="De Man" pageId="53" pageNumber="944" pagination="639 - 722" refId="ref72089" refString="Man JG De (1887) Ubersicht der Indopacifischen Arten der Gattung Sesarma Say nebst einer Kritik der von W. Hess und E. Nauck in den Jahren 1865 und 1880 beschrieben Decapoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 2: 639 - 722." type="journal article" year="1887">De Man, 1887</bibRefCitation>
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<subSubSection id="89A6C9CBFFD4BA14FF58FCCEFE56FC04" pageId="53" pageNumber="944" type="description">
<paragraph id="C1039A40FFD4BA14FF58FCCEFE56FC04" blockId="53.[133,732,861,949]" pageId="53" pageNumber="944">
(
<figureCitation id="598786C5FFD4BA14FF50FCCEFE96FC24" box="[141,283,893,917]" captionStart="Fig" captionStartId="41.[113,148,1553,1575]" captionTargetBox="[113,1451,189,1537]" captionTargetId="figure-14@41.[113,1451,189,1537]" captionText="Fig. 22. Overall habitus, Pseudosesarma species. A, P. edwardsii, lectotype male (17.5 × 16.1 mm) (RMNH-D17a), Mergui Archipelago; B, P. edwardsii, male (13.4 × 11.9 mm) (ZRC 1965.8.2.81), Pulau Pawai, Singapore; C, P. edwardsii, male (12.9 × 11.8 mm) (ZRC 1971.9.24.9), Singapore; D, P. crassimanum, male (15.9 × 14.5 mm) (RMNH-D23313), Singapore; E, P. anteactum, holotype male (16.7 × 14.7 mm) (ZRC 2016.0602), Sri Lanka; F, P. glabrum, holotype male (13.8 × 12.0 mm) (CUSAT 2016-1), Kerala, India; G, P. brehieri, holotype male (17.4 × 15.7 mm) (ZRC 2016.0593), Myanmar; H, P. boulengeri, lectotype male (26.9 × 23.5 mm) (NHM 1919.11.14.1), Basra, Iraq." figureDoi="http://doi.org/10.5281/zenodo.6519005" httpUri="https://zenodo.org/record/6519005/files/figure.png" pageId="53" pageNumber="944">Figs. 22AC</figureCitation>
,
<figureCitation id="598786C5FFD4BA14FEF5FCCEFEF4FC24" box="[296,377,893,917]" captionStart="Fig" captionStartId="46.[136,171,1913,1935]" captionTargetBox="[147,1463,189,1899]" captionTargetId="figure-8@46.[147,1463,189,1899]" captionText="Fig. 25. Frontal view of cephalothorax of Pseudosesarma. A, P. edwardsii, lectotype male (17.5 × 16.1 mm) (RMNH-D17a), Mergui Archipelago; B, P. edwardsii, male (13.4 × 11.9 mm) (ZRC 1965.8.2.81), Pulau Pawai, Singapore; C, P. crassimanum, male (15.9 × 14.5 mm) (RMNH-D23313), Singapore; D, P. anteactum, holotype male (16.7 × 14.7 mm) (ZRC 2016.0602), Sri Lanka; E, P. glabrum, holotype male (13.8 × 12.0 mm) (CUSAT 2016-1), Kerala, India; F, P. brehieri, holotype male (17.4 × 15.7 mm) (ZRC 2016.0593), Myanmar; G, P. boulengeri, lectotype male (26.9 × 23.5 mm) (NHM 1919.11.14.1), Basra, Iraq; H, Contusarma bocourti, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand; I, Contusarma cheirogonum, neotype male (24.5 × 21.5 mm) (ZRC 1995.225), Bako National Park, Sarawak; J, Miersarma granosimanum, lectotype male (16.9 × 14.8 mm) (NHM 1880.6), Borneo." figureDoi="http://doi.org/10.5281/zenodo.6519007" httpUri="https://zenodo.org/record/6519007/files/figure.png" pageId="53" pageNumber="944">25A, B</figureCitation>
,
<figureCitation id="598786C5FFD4BA14FE5BFCCEFE5BFC24" box="[390,470,893,917]" captionStart="Fig" captionStartId="49.[113,148,1885,1907]" captionTargetBox="[113,1451,246,1869]" captionTargetId="figure-14@49.[113,1451,246,1869]" captionText="Fig. 27. Outer view of chela of Pseudosesarma. A, P. edwardsii, lectotype male (17.5 × 16.1 mm) (RMNH-D17a), Mergui Archipelago; B, P. edwardsii, male (13.4 × 11.9 mm) (ZRC 1965.8.2.81), Pulau Pawai, Singapore; C, P. edwardsii, male (19.4 × 17.3 mm) (ZRC 1971.9.24.8), Singapore; D, P. crassimanum, male (15.9 × 14.5 mm) (RMNH-D23313), Singapore; E, P. anteactum, holotype male (16.7 × 14.7 mm) (ZRC 2016.0602), Sri Lanka; F, P. glabrum, holotype male (13.8 × 12.0 mm) (CUSAT 2016-1), Kerala, India; G, P. brehieri, holotype male (17.4 × 15.7 mm) (ZRC 2016.0593), Myanmar; H, P. boulengeri, lectotype male (26.9 × 23.5 mm) (NHM 1919.11.14.1), Basra, Iraq." figureDoi="http://doi.org/10.5281/zenodo.6519009" httpUri="https://zenodo.org/record/6519009/files/figure.png" pageId="53" pageNumber="944">27AC</figureCitation>
,
<figureCitation id="598786C5FFD4BA14FE3FFCCEFDB9FC24" box="[482,564,893,917]" captionStart="Fig" captionStartId="52.[136,171,1939,1961]" captionTargetBox="[173,1438,189,1924]" captionTargetId="figure-7@52.[173,1438,189,1924]" captionText="Fig. 30. Male anterior thoracic sternum and pleon of Pseudosesarma. A, P edwardsii, lectotype male (17.5 × 16.1 mm) (RMNH-D17a), Mergui Archipelago; B, P. edwardsii, male (13.4 × 11.9 mm) (ZRC 1965.8.2.81), Pulau Pawai, Singapore; C, P. crassimanum, male (15.9 × 14.5 mm) (RMNH-D23313), Singapore; D, P. glabrum, holotype male (13.8 × 12.0 mm) (CUSAT 2016-1), Kerala, India; E, P. anteactum, holotype male (16.7 × 14.7 mm) (ZRC 2016.0602), Sri Lanka; F, P. anteactum, paratype male (21.2 × 19.7 mm) (ZRC 2016.0603), Sri Lanka; G, P. brehieri, holotype male (17.4 × 15.7 mm) (ZRC 2016.0593), Myanmar; H, P. boulengeri, lectotype male (26.9 × 23.5 mm) (NHM 1919.11.14.1), Basra, Iraq." figureDoi="http://doi.org/10.5281/zenodo.6519011" httpUri="https://zenodo.org/record/6519011/files/figure.png" pageId="53" pageNumber="944">30A, B</figureCitation>
,
<figureCitation id="598786C5FFD4BA14FD9CFCCEFDD0FC24" box="[577,605,893,917]" captionStart="Fig" captionStartId="55.[113,148,1344,1366]" captionTargetBox="[113,1451,189,1328]" captionTargetId="figure-15@55.[113,1451,189,1328]" captionTargetPageId="55" captionText="Fig. 32. Male sternopleonal cavity. A, Pseudosesarma edwardsii, male (20.8 × 18.6mm (ZRC 2003.0084), Pulau Tekong, Singapore; B, Pseudosesarma crassimanum, male (15.1 × 13.3 mm) (ZRC 2000.1768), Sungei Benut, Johor, Malaysia; C, Contusarma cheirogonum, male (15.3 × 14.0 mm) (ZRC 2011.0924), Pulau Tioman, Malaysia; D, Miersarma granosimanum, male (22.5 × 19.2 mm) (ZRC 1965.7.29.164), Sedili River, Johor, Malaysia. Arrow indicates shallow depression on somite 5 to accept G1 tip when pleon closed (depression absent in Miersarma, D)." figureDoi="http://doi.org/10.5281/zenodo.5351355" httpUri="https://zenodo.org/record/5351355/files/figure.png" pageId="53" pageNumber="944">32</figureCitation>
A,
<figureCitation id="598786C5FFD4BA14FDA3FCCEFD16FC24" box="[638,667,893,917]" captionStart="Fig" captionStartId="58.[136,171,1390,1412]" captionTargetBox="[136,1475,190,1374]" captionTargetId="figure-8@58.[125,1484,178,1386]" captionTargetPageId="58" captionText="Fig. 34. Pseudosesarma edwardsii. A, B, D, E, lectotype male (17.5 × 16.1 mm) (RMNH-D17a), Mergui Archipelago; C, male (13.3 × 12.9 mm) (ZRC 2000.2019), Pulau Ubin, Singapore; FH, male (20.5 × 19.3 mm) (ZRC 2016.0608), Langkawi; I, J, male (21.3 × 18.7 mm) (ZRC 2003.0084), Singapore; K, L, male (17.8 × 16.3 mm) (ZRC 2016.0608), Langkawi; M, N, male (12.5 × 10.4 mm) (ZRC 2016.0608), Langkawi. A, anterior thoracic sternites 14; B, C, male pleon; D, H, I, K, M, left G1 (dorsal view, denuded); E, G, J, K, N, left G1 (ventral view, denuded); H, left G2. Scales: AC = 2.0 mm; DJ = 1.0 mm; KN = 0.5 mm. (After Ng &amp; Schubart, 2017: figs. 6, 7)." figureDoi="http://doi.org/10.5281/zenodo.5351359" httpUri="https://zenodo.org/record/5351359/files/figure.png" pageId="53" pageNumber="944">34</figureCitation>
,
<figureCitation id="598786C5FFD4BA14FD75FCCEFD49FC24" box="[680,708,893,917]" captionStart="Fig" captionStartId="74.[136,171,1898,1920]" captionTargetBox="[136,1474,211,1884]" captionTargetId="figure-7@74.[136,1474,211,1884]" captionTargetPageId="74" captionText="Fig. 44. Vulva. A, Pseudosesarma edwardsii, female (16.5 × 15.3 mm) (ZRC 2016.0608), Malaysia; B, Pseudosesarma crassimanum, female (15.9 × 13.8 mm) (ZRC 2000.1768), Malaysia; C, Pseudosesarma crassimanum, female (18.0 × 15.7 mm) (ZRC 2017.1045), Thailand; D, Pseudosesarma anteactum, paratype female (17.5 × 15.4 mm) (ZRC 2016.0603), Sri Lanka; E, Pseudosesarma brehieri, paratype female (14.9 × 13.4 mm) (ZRC 2016.0594), Myanmar; F, Pseudosesarma boulengeri, female (24.2 × 20.0 mm) (ZRC 2014.0335), Iran; G, Contusarma bocourti, female (21.1 × 19.0 mm) (ZRC 2019.1114), Thailand; H, Contusarma cheirogonum, female (26.6 × 23.0 mm) (ZRC 1964.9.28.127), Sarawak; I, Miersarma granosimanum, female (20.1 × 16.4 mm) (ZRC 1965.7.29.166), Malaysia; J, Manarma moeschii, female (19.3 × 17.3 mm) (ZRC 2017.0170), Thailand." figureDoi="http://doi.org/10.5281/zenodo.5351375" httpUri="https://zenodo.org/record/5351375/files/figure.png" pageId="53" pageNumber="944">44</figureCitation>
A,
<figureCitation id="598786C5FFD4BA14FE5AFC2EFE2FFC04" box="[391,418,925,949]" captionStart="Fig" captionStartId="88.[136,171,1875,1897]" captionTargetBox="[136,1474,324,1860]" captionTargetId="figure-7@88.[136,1474,324,1860]" captionTargetPageId="88" captionText="Fig. 56. Colour in life. A, B, Pseudosesarma edwardsii, male (20.5 × 19.3 mm) (ZRC 2016.0608), Langkawi [photograph: Paul Ng]; CE, P. edwardsii, male (19.4 × 18.2 mm) (SMF), Langkawi, Peninsular Malaysia [photograph: CD Schubart]; F, G, P. crassimanum, (12.6 × 10.5 mm) (ZRC 2010.0035), Tioman, Peninsular Malaysia [photograph: PKL Ng]; H, P. boulengeri, specimen not preserved [photograph: Reza Naderloo]." figureDoi="http://doi.org/10.5281/zenodo.5351399" httpUri="https://zenodo.org/record/5351399/files/figure.png" pageId="53" pageNumber="944">56</figureCitation>
AE)
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<taxonomicName id="06BCE1C3FFD4BA14FFACFC6FFE78FC43" ID-CoL="9DBPH" authority="De Man, 1887: 649" authorityName="De Man" authorityPageNumber="649" authorityYear="1887" box="[113,501,988,1010]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="944" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="F3C84652FFD4BA14FFACFC6FFEA6FC40" box="[113,299,988,1009]" italics="true" pageId="53" pageNumber="944">Sesarma edwardsii</emphasis>
<bibRefCitation id="A52DE7B1FFD4BA14FEEFFC6FFE78FC43" author="De Man" box="[306,501,988,1010]" firstAuthor="De Man" page="649" pageId="53" pageNumber="944" pagination="639 - 722" refId="ref72089" refString="Man JG De (1887) Ubersicht der Indopacifischen Arten der Gattung Sesarma Say nebst einer Kritik der von W. Hess und E. Nauck in den Jahren 1865 und 1880 beschrieben Decapoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 2: 639 - 722." type="journal article" year="1887">De Man, 1887: 649</bibRefCitation>
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<taxonomicName id="06BCE1C3FFD4BA14FFACFC4AFE90FBBF" ID-CoL="84HBF" authorityName="Ortmann" authorityYear="1894" box="[113,285,1017,1038]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="944" phylum="Arthropoda" rank="species" species="edwardsi">
<emphasis id="F3C84652FFD4BA14FFACFC4AFE90FBBF" box="[113,285,1017,1038]" italics="true" pageId="53" pageNumber="944">Sesarma edwardsi</emphasis>
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<bibRefCitation id="A52DE7B1FFD4BA14FEEFFC4AFE6AFBBE" author="De Man" box="[306,487,1017,1039]" firstAuthor="De Man" page="185" pageId="53" pageNumber="944" pagination="177 - 240" refId="ref72143" refString="Man JG De (1888) Report on the Podophthalmous Crustacea of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta, by Dr. John Anderson, F. R. S., Superintendent of the Museum. - Part IV. Journal of the Linnean Society of London (Zoology), 22 (139): 177 - 240, pls. 13 - 15." type="journal article" year="1888">
De
<treatmentCitation id="401DBC51FFD4BA14FE8FFC4AFE6AFBBE" author="Man JG De" box="[338,487,1017,1039]" page="185" pageId="53" pageNumber="944" year="1888">Man, 1888: 185</treatmentCitation>
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, pl. 13 figs. 14;
<treatmentCitation id="401DBC51FFD4BA14FD5EFC4AFF71FB9A" author="Lanchester WF" page="757" pageId="53" pageNumber="944" year="1900">
<bibRefCitation id="A52DE7B1FFD4BA14FD5EFC4AFF71FB9A" author="Lanchester" firstAuthor="Lanchester" page="757" pageId="53" pageNumber="944" pagination="719 - 770" refId="ref71180" refString="Lanchester WF (1900) On a collection of crustaceans made at Singapore and Malacca. Part I. Crustacea Brachyura. Proceedings of the Zoological Society of London, 1900 (3): 719 - 770, pls. 44 - 47." type="journal article" year="1900">Lanchester, 1900: 757</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFD4BA14FFACFB80FD2FFBF9" blockId="53.[113,753,988,1327]" box="[113,674,1074,1096]" pageId="53" pageNumber="944">
<treatmentCitationGroup id="E1ACBD6EFFD4BA14FFACFB80FD2FFBF9" box="[113,674,1074,1096]" pageId="53" pageNumber="944">
<taxonomicName id="06BCE1C3FFD4BA14FFACFB80FE03FBF6" ID-CoL="84HBF" authorityName="Ortmann" authorityYear="1894" box="[113,398,1074,1096]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="944" phylum="Arthropoda" rank="species" species="edwardsi" subGenus="Sesarma">
<emphasis id="F3C84652FFD4BA14FFACFB80FF49FBF9" box="[113,196,1075,1096]" italics="true" pageId="53" pageNumber="944">Sesarma</emphasis>
(
<emphasis id="F3C84652FFD4BA14FF09FB80FEAAFBF9" box="[212,295,1075,1096]" italics="true" pageId="53" pageNumber="944">Sesarma</emphasis>
)
<emphasis id="F3C84652FFD4BA14FEEBFB81FE03FBF6" box="[310,398,1074,1095]" italics="true" pageId="53" pageNumber="944">edwardsi</emphasis>
</taxonomicName>
<treatmentCitation id="401DBC51FFD4BA14FE77FB81FDDEFBF6" author="Tesch JJ" box="[426,595,1074,1096]" page="147" pageId="53" pageNumber="944" year="1917">
<bibRefCitation id="A52DE7B1FFD4BA14FE77FB81FDDEFBF6" author="Tesch" box="[426,595,1074,1096]" firstAuthor="Tesch" page="147" pageId="53" pageNumber="944" pagination="127 - 260" refId="ref78063" refString="Tesch JJ (1917) Synopsis of the genera Sesarma, Metasesarma, Sarmatium and Clistocoeloma, with a key to the determination of the IndoPacific species. Zoologische Mededelingen, 3: 127 - 260, pls. 15 - 17." type="journal article" year="1917">Tesch, 1917: 147</bibRefCitation>
</treatmentCitation>
(part?).
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFD4BA14FFACFBFCFDF0FBD4" blockId="53.[113,753,988,1327]" box="[113,637,1103,1125]" pageId="53" pageNumber="944">
<treatmentCitationGroup id="E1ACBD6EFFD4BA14FFACFBFCFDF0FBD4" box="[113,637,1103,1125]" pageId="53" pageNumber="944">
?
<taxonomicName id="06BCE1C3FFD4BA14FFA1FBFCFE2DFBD5" ID-CoL="9DBPH" authorityName="De Man" authorityYear="1887" box="[124,416,1103,1125]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="944" phylum="Arthropoda" rank="species" species="edwardsii" subGenus="Sesarma">
<emphasis id="F3C84652FFD4BA14FFA1FBFCFF42FBD5" box="[124,207,1103,1124]" italics="true" pageId="53" pageNumber="944">Sesarma</emphasis>
(
<emphasis id="F3C84652FFD4BA14FF03FBFCFEBCFBD5" box="[222,305,1103,1124]" italics="true" pageId="53" pageNumber="944">Sesarma</emphasis>
)
<emphasis id="F3C84652FFD4BA14FE9CFBFCFE2DFBD5" box="[321,416,1103,1124]" italics="true" pageId="53" pageNumber="944">edwardsii</emphasis>
</taxonomicName>
<treatmentCitation id="401DBC51FFD4BA14FE66FBFCFDF4FBD4" author="Calman WT" box="[443,633,1103,1125]" page="166" pageId="53" pageNumber="944" year="1925">
<bibRefCitation id="A52DE7B1FFD4BA14FE66FBFCFDF4FBD4" author="Calman" box="[443,633,1103,1125]" firstAuthor="Calman" page="166" pageId="53" pageNumber="944" pagination="166 - 167" refId="ref68142" refString="Calman WT (1925) X. Crustacea Decapoda. Collected in Korinchi and on the West Sumatran Coast, by Messrs. H. C. Robinson and C. Boden Kloss. Results of an Expedition to Korinchi Peak, Sumatra. Journal of the Federated Malay States Museums, 8 (3): 166 - 167." type="journal article" year="1925">Calman, 1925: 166</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFD4BA14FFACFBDFFD04FB33" blockId="53.[113,753,988,1327]" box="[113,649,1132,1154]" pageId="53" pageNumber="944">
<treatmentCitationGroup id="E1ACBD6EFFD4BA14FFACFBDFFD04FB33" box="[113,649,1132,1154]" pageId="53" pageNumber="944">
<taxonomicName id="06BCE1C3FFD4BA14FFACFBDFFE0FFB30" authority="(De Man, 1887)" box="[113,386,1132,1153]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="944" phylum="Arthropoda" rank="species" species="edwarsi" subGenus="Sesarma">
<emphasis id="F3C84652FFD4BA14FFACFBDFFF49FB30" box="[113,196,1132,1153]" italics="true" pageId="53" pageNumber="944">Sesarma</emphasis>
(
<emphasis id="F3C84652FFD4BA14FF09FBDFFEAAFB30" box="[212,295,1132,1153]" italics="true" pageId="53" pageNumber="944">Sesarma</emphasis>
)
<emphasis id="F3C84652FFD4BA14FEEBFBDFFE0FFB30" box="[310,386,1132,1153]" italics="true" pageId="53" pageNumber="944">edwarsi</emphasis>
</taxonomicName>
(sic)
<treatmentCitation id="401DBC51FFD4BA14FE0DFBDFFD08FB33" author="Serene R" box="[464,645,1132,1154]" page="105" pageId="53" pageNumber="944" year="1968">
<bibRefCitation id="A52DE7B1FFD4BA14FE0DFBDFFD08FB33" author="Serene" box="[464,645,1132,1154]" firstAuthor="Serene" page="105" pageId="53" pageNumber="944" pagination="33 - 112" refId="ref76415" refString="Serene R (1968) The Brachyura of the Indo Pacific Region. In: Prodromus for a check list of the nonplanctonic marine fauna of South East Asia. Singapore National Academy of Science, Special Publication No. 1: 33 - 112." type="book chapter" year="1968">Serène, 1968: 105</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFD4BA14FFACFB3AFD51FB2E" blockId="53.[113,753,988,1327]" box="[113,732,1161,1183]" pageId="53" pageNumber="944">
<treatmentCitationGroup id="E1ACBD6EFFD4BA14FFACFB3AFD51FB2E" box="[113,732,1161,1183]" pageId="53" pageNumber="944">
<taxonomicName id="06BCE1C3FFD4BA14FFACFB3AFED1FB2F" authority="(De Man, 1887)" box="[113,348,1161,1182]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="944" phylum="Arthropoda" rank="species" species="edwarsi">
<emphasis id="F3C84652FFD4BA14FFACFB3AFED1FB2F" box="[113,348,1161,1182]" italics="true" pageId="53" pageNumber="944">Pseudosesarma edwarsi</emphasis>
</taxonomicName>
(sic)
<treatmentCitation id="401DBC51FFD4BA14FE77FB3AFD28FB2F" author="Serene R &amp; Soh CL" box="[426,677,1161,1183]" page="399" pageId="53" pageNumber="944" year="1970">
<bibRefCitation id="A52DE7B1FFD4BA14FE77FB3AFD28FB2F" author="Serene &amp; Soh" box="[426,677,1161,1183]" firstAuthor="Serene" page="399" pageId="53" pageNumber="944" pagination="387 - 416" refId="ref76461" refString="Serene R &amp; Soh CL (1970) New IndoPacific genera allied to Sesarma Say 1877 (Brachyura, Decapoda, Crustacea). Treubia, 27: 387 - 416." type="journal article" year="1970">Serène &amp; Soh, 1970: 399</bibRefCitation>
</treatmentCitation>
, 406.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFD4BA14FFACFB16FE25FB45" blockId="53.[113,753,988,1327]" pageId="53" pageNumber="944">
<treatmentCitationGroup id="E1ACBD6EFFD4BA14FFACFB16FE25FB45" pageId="53" pageNumber="944">
<taxonomicName id="06BCE1C3FFD4BA14FFACFB16FEE2FB0B" ID-CoL="6WGDS" baseAuthorityName="De Man" baseAuthorityYear="1887" box="[113,367,1189,1210]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="944" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="F3C84652FFD4BA14FFACFB16FEE2FB0B" box="[113,367,1189,1210]" italics="true" pageId="53" pageNumber="944">Pseudosesarma edwardsii</emphasis>
</taxonomicName>
<treatmentCitation id="401DBC51FFD4BA14FE56FB15FDD4FB0A" author="Tan CGS &amp; Ng PKL" box="[395,601,1190,1211]" page="82" pageId="53" pageNumber="944" year="1994">
<bibRefCitation id="A52DE7B1FFD4BA14FE56FB15FDD4FB0A" author="Tan &amp; Ng" box="[395,601,1190,1211]" firstAuthor="Tan" page="82" pageId="53" pageNumber="944" pagination="75 - 84" refId="ref77891" refString="Tan CGS &amp; Ng PKL (1994) An annotated checklist of mangrove brachyuran crabs in Malaysia and Singapore. Hydrobiologia, 285: 75 - 84." type="journal article" year="1994">Tan &amp; Ng, 1994: 82</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFD4BA14FDB8FB15FF71FB66" author="Fransen CHJM &amp; Holthuis LB &amp; Adema JPHM" page="127" pageId="53" pageNumber="944" year="1997">
<bibRefCitation id="A52DE7B1FFD4BA14FDB8FB15FF71FB66" author="Fransen" etAl="et al." firstAuthor="Fransen" page="127" pageId="53" pageNumber="944" pagination="1 - 344" refId="ref69235" refString="Fransen CHJM, Holthuis LB &amp; Adema JPHM (1997) Typecatalogue of the Decapod Crustacea in the collections of the Nationaal Naturhistorisch Museum, with appendices of pre- 1900 collectors and material. Zoologische Verhandelingen, Leiden, 311: i - xvi, 1 - 344." type="journal article" year="1997">Fransen et al., 1997: 127</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFD4BA14FED5FB70FE57FB66" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[264,474,1218,1240]" page="222" pageId="53" pageNumber="944" year="2008">
<bibRefCitation id="A52DE7B1FFD4BA14FED5FB70FE57FB66" author="Ng" box="[264,474,1218,1240]" etAl="et al." firstAuthor="Ng" page="222" pageId="53" pageNumber="944" pagination="1 - 286" refId="ref73754" refString="Ng PKL, Guinot D &amp; Davie PJF (2008 a) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008" yearSuffix="a">Ng et al., 2008a: 222</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFD4BA14FE3BFB70FD61FB69" author="Ng PKL &amp; Schubart CD" box="[486,748,1218,1240]" page="655" pageId="53" pageNumber="944" year="2017">
<bibRefCitation id="A52DE7B1FFD4BA14FE3BFB70FD61FB69" author="Ng &amp; Schubart" box="[486,748,1218,1240]" firstAuthor="Ng" page="655" pageId="53" pageNumber="944" pagination="655 - 669" refId="ref74064" refString="Ng PKL &amp; Schubart CD (2017) On the taxonomy of Pseudosesarma edwardsii (De Man, 1887) and P. crassimanum (De Man, 1887) (Crustacea: Decapoda: Brachyura: Sesarmidae), with description of a new species from Sri Lanka. Raffles Bulletin of Zoology, 65: 655 - 669." type="journal article" year="2017">Ng &amp; Schubart, 2017: 655</bibRefCitation>
</treatmentCitation>
, figs. 1AC, 2A, B, 37, 12.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFD4BA14FFACFB4FFEB1FA9E" blockId="53.[113,753,988,1327]" pageId="53" pageNumber="944">
<treatmentCitationGroup id="E1ACBD6EFFD4BA14FFACFB4FFEB1FA9E" pageId="53" pageNumber="944">
<collectingRegion id="037854A2FFD4BA14FFACFB4FFF18FAA0" box="[113,149,1276,1297]" country="Cameroon" name="North" pageId="53" pageNumber="944">Not</collectingRegion>
<treatmentCitation id="401DBC51FFD4BA14FF41FB4FFD9AFAA0" author="Ortmann AE" box="[156,535,1276,1297]" page="721" pageId="53" pageNumber="944" year="1894">
<taxonomicName id="06BCE1C3FFD4BA14FF41FB4FFD9AFAA0" ID-CoL="84HBF" authority="Ortmann, 1894: 721" authorityName="Ortmann" authorityPageNumber="721" authorityYear="1894" box="[156,535,1276,1297]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="944" phylum="Arthropoda" rank="species" species="edwardsi">
<emphasis id="F3C84652FFD4BA14FF41FB4FFEC0FAA0" box="[156,333,1276,1297]" italics="true" pageId="53" pageNumber="944">Sesarma edwardsi</emphasis>
<bibRefCitation id="A52DE7B1FFD4BA14FE89FB4FFD9AFAA0" author="Ortmann" box="[340,535,1276,1297]" firstAuthor="Ortmann" page="721" pageId="53" pageNumber="944" pagination="683 - 772" refId="ref74488" refString="Ortmann AE (1894) Abtheilung: Brachyura (Brachyura genuina Boas), III. Unterabtheilung: Cancroidea. 2. Section: Cancrinea, 2. Gruppe: Catametopa. Die Decapoden-Krebse des Strassburger Museums mit besonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur Zeit im Strassburger Museum aufbewahrten Formen. Theil VIII [= Part 8]. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 7 (5): 683 - 772, pl. 23." type="journal article" year="1894">Ortmann, 1894: 721</bibRefCitation>
</taxonomicName>
</treatmentCitation>
=
<taxonomicName id="06BCE1C3FFD4BA14FDEFFB4FFEB5FA9E" ID-CoL="N9DB" authority="(De Man, 1889)" baseAuthorityName="De Man" baseAuthorityYear="1889" class="Malacostraca" family="Sesarmidae" genus="Bresedium" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="944" phylum="Arthropoda" rank="species" species="brevipes">
<emphasis id="F3C84652FFD4BA14FDEFFB4FFD7DFAA0" box="[562,752,1276,1297]" italics="true" pageId="53" pageNumber="944">Bresedium brevipes</emphasis>
(
<bibRefCitation id="A52DE7B1FFD4BA14FF43FAAAFEBCFA9E" author="De Man" box="[158,305,1305,1327]" firstAuthor="De Man" pageId="53" pageNumber="944" pagination="409 - 452" refId="ref72218" refString="Man JG De (1889) Ueber einige neue oder seltene indopacifische Brachyuren. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Thiere, Jena, 4 (2): 409 - 452, pls. 9, 10." type="book chapter" year="1889">De Man, 1889</bibRefCitation>
)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFD4BA14FFACFAE4FBBEFDC4" pageId="53" pageNumber="944" type="materials_examined">
<paragraph id="C1039A40FFD4BA14FFACFAE4FBBEFDC4" blockId="53.[113,753,1367,2127]" lastBlockId="53.[813,1452,189,630]" pageId="53" pageNumber="944">
<emphasis id="F3C84652FFD4BA14FFACFAE4FEDBFADE" bold="true" box="[113,342,1367,1391]" pageId="53" pageNumber="944">Material examined</emphasis>
.
<materialsCitation id="71D4901DFFD4BA14FEB5FAE4FE6CFA1E" ID-GBIF-Occurrence="3763237488" collectionCode="RMNH-D" collectorName="J. Anderson" country="Myanmar" location="Myanmar" pageId="53" pageNumber="944" specimenCode="RMNH-D17" specimenCount="1" typeStatus="lectotype">
<typeStatus id="1E0724E2FFD4BA14FEB5FAE4FE5AFADE" box="[360,471,1367,1391]" pageId="53" pageNumber="944" type="lectotype">Lectotype</typeStatus>
(here designated): male (17.5 ×
<quantity id="064437A5FFD4BA14FF0DFAC4FEBAFA3E" box="[208,311,1399,1423]" metricMagnitude="-2" metricUnit="m" metricValue="1.61" pageId="53" pageNumber="944" unit="mm" value="16.1">16.1 mm</quantity>
) (
<specimenCode id="911A323BFFD4BA14FE8CFAC4FE6FFA3E" box="[337,482,1399,1423]" collectionCode="RMNH-D" pageId="53" pageNumber="944">RMNH-D17</specimenCode>
a), Mergui Archipelago,
<collectingCountry id="B9ABDAD0FFD4BA14FFAFFA24FF51FA1E" box="[114,220,1431,1455]" name="Myanmar" pageId="53" pageNumber="944">Myanmar</collectingCountry>
, coll.
<collectorName id="6C49FF96FFD4BA14FEC1FA24FE10FA1E" box="[284,413,1431,1455]" pageId="53" pageNumber="944">J. Anderson</collectorName>
, 1886
</materialsCitation>
.
<materialsCitation id="71D4901DFFD4BA14FE36FA24FF67FA5E" ID-GBIF-Occurrence="3763237369" collectionCode="RMNH-D" collectorName="J. Anderson" country="Myanmar" location="Myanmar" pageId="53" pageNumber="944" specimenCode="RMNH-D17" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<typeStatus id="1E0724E2FFD4BA14FE36FA24FD06FA1E" box="[491,651,1431,1455]" pageId="53" pageNumber="944" type="paralectotype">Paralectotypes</typeStatus>
:
<specimenCount id="D7BA51C9FFD4BA14FD49FA24FD7CFA1E" box="[660,753,1431,1455]" pageId="53" pageNumber="944" type="female">1 female</specimenCount>
(
<specimenCode id="911A323BFFD4BA14FFA4FA04FE88FA7E" box="[121,261,1463,1487]" collectionCode="RMNH-D" pageId="53" pageNumber="944">RMNH-D17</specimenCode>
b), same data as lectotype; 1 young male (7.9 × 7.0 mm)
</materialsCitation>
,
<materialsCitation id="71D4901DFFD4BA14FF25FA64FEE8F9FE" ID-GBIF-Occurrence="3763237480" collectionCode="NHM" collectorName="Others" country="Singapore" location="Myanmar" pageId="53" pageNumber="944" specimenCode="NHM 1886.52" specimenCount="8" specimenCount-female="8" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD4BA14FF25FA64FEEFFA5E" box="[248,354,1495,1519]" pageId="53" pageNumber="944" type="female">8 females</specimenCount>
(20.3 ×
<quantity id="064437A5FFD4BA14FE18FA64FDA4FA5F" box="[453,553,1495,1519]" metricMagnitude="-2" metricUnit="m" metricValue="1.77" pageId="53" pageNumber="944" unit="mm" value="17.7">17.7 mm</quantity>
, 14.9 ×
<quantity id="064437A5FFD4BA14FD55FA64FD61FA5F" box="[648,748,1495,1519]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="53" pageNumber="944" unit="mm" value="13.2">13.2 mm</quantity>
, 13.8 ×
<quantity id="064437A5FFD4BA14FF1DFA44FEAEF9BF" box="[192,291,1527,1551]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="53" pageNumber="944" unit="mm" value="12.7">12.7 mm</quantity>
, 13.7 ×
<quantity id="064437A5FFD4BA14FEA0FA44FE6DF9BF" box="[381,480,1527,1551]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="53" pageNumber="944" unit="mm" value="12.5">12.5 mm</quantity>
, 12.3 ×
<quantity id="064437A5FFD4BA14FDE6FA44FD13F9BF" box="[571,670,1527,1551]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="53" pageNumber="944" unit="mm" value="11.6">11.6 mm</quantity>
, 12.3 ×
<quantity id="064437A5FFD4BA14FFACF9A4FF5FF99F" box="[113,210,1559,1583]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="53" pageNumber="944" unit="mm" value="11.3">11.3 mm</quantity>
, 10.9 ×
<quantity id="064437A5FFD4BA14FEFAF9A4FE05F99F" box="[295,392,1559,1583]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="53" pageNumber="944" unit="mm" value="10.2">10.2 mm</quantity>
, 8.8 ×
<quantity id="064437A5FFD4BA14FE0DF9A4FDAEF99E" box="[464,547,1559,1583]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="53" pageNumber="944" unit="mm" value="7.9">7.9 mm</quantity>
) (
<specimenCode id="911A323BFFD4BA14FDE5F9A4FD54F99E" box="[568,729,1559,1583]" collectionCode="NHM" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="53" pageNumber="944" type="Herbarium">NHM 1886.52</specimenCode>
a), same data as lectotype.
</materialsCitation>
<materialsCitation id="71D4901DFFD4BA14FEB7F984FEC5F93E" ID-GBIF-Occurrence="3763237569" collectingDate="1966-03-29" collectionCode="NHM" collectorName="Others" country="Singapore" location="Myanmar" pageId="53" pageNumber="944" specimenCode="NHM 1886.52" specimenCount="8" specimenCount-female="8" specimenCount-male="1" typeStatus="paralectotype">
<collectorName id="6C49FF96FFD4BA14FEB7F984FE38F9FE" box="[362,437,1591,1615]" pageId="53" pageNumber="944">Others</collectorName>
:
<collectingCountry id="B9ABDAD0FFD4BA14FE63F984FDD9F9FE" box="[446,596,1591,1615]" name="Singapore" pageId="53" pageNumber="944">SINGAPORE</collectingCountry>
<specimenCount id="D7BA51C9FFD4BA14FDB1F984FD3EF9FE" box="[620,691,1591,1615]" pageId="53" pageNumber="944" type="male">1 male</specimenCount>
(19.4 ×
<quantity id="064437A5FFD4BA14FF57F9E4FF60F9DE" box="[138,237,1623,1647]" metricMagnitude="-2" metricUnit="m" metricValue="1.73" pageId="53" pageNumber="944" unit="mm" value="17.3">17.3 mm</quantity>
) (
<collectionCode id="A7AD0285FFD4BA14FED8F9E4FEB6F9DE" box="[261,315,1623,1647]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="944">ZRC</collectionCode>
1971.9.24.8),
<location id="C463CC9BFFD4BA14FE06F9E4FDF1F9DE" LSID="urn:lsid:plazi:treatment:49152B56FFD4BA16FF18FCEEFA21F849:C463CC9BFFD4BA14FE06F9E4FDF1F9DE" box="[475,636,1623,1647]" country="Singapore" name="Sungei Seletar" pageId="53" pageNumber="944">Sungei Seletar</location>
, coll.
<collectorName id="6C49FF96FFD4BA14FD1CF9E4FF10F93E" pageId="53" pageNumber="944">C.L. Soh</collectorName>
,
<date id="B502BC80FFD4BA14FF7BF9C4FEC5F93E" box="[166,328,1655,1679]" pageId="53" pageNumber="944" value="1966-03-29">
<collectingDate id="A5464568FFD4BA14FF7BF9C4FEC5F93E" box="[166,328,1655,1679]" pageId="53" pageNumber="944" value="1966-03-29">29 March 1966</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71D4901DFFD4BA14FE8CF9C4FE46F91E" ID-GBIF-Occurrence="3763237428" collectionCode="ZRC" collectorName="F. N. Chasen" country="Singapore" location="Pulau Aya" municipality="Merban" pageId="53" pageNumber="944" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD4BA14FE8CF9C4FE1AF93E" box="[337,407,1655,1679]" pageId="53" pageNumber="944" type="male">1 male</specimenCount>
(
<collectionCode id="A7AD0285FFD4BA14FE7EF9C4FE55F93E" box="[419,472,1655,1679]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="944">ZRC</collectionCode>
1965.7.29.50),
<location id="C463CC9BFFD4BA14FDA1F9C4FD60F93E" LSID="urn:lsid:plazi:treatment:49152B56FFD4BA16FF18FCEEFA21F849:C463CC9BFFD4BA14FDA1F9C4FD60F93E" box="[636,749,1655,1679]" country="Singapore" municipality="Merban" name="Pulau Aya" pageId="53" pageNumber="944">Pulau Aya</location>
,
<collectingMunicipality id="2167003AFFD4BA14FFACF924FF4AF91E" box="[113,199,1687,1711]" pageId="53" pageNumber="944">Merban</collectingMunicipality>
, coll.
<collectorName id="6C49FF96FFD4BA14FED9F924FE04F91E" box="[260,393,1687,1711]" pageId="53" pageNumber="944">F.N. Chasen</collectorName>
, 1931
</materialsCitation>
;
<materialsCitation id="71D4901DFFD4BA14FE08F924FD1EF97E" ID-GBIF-Occurrence="3763237404" collectingDate="1966-03-29" collectionCode="ZRC" collectorName="C. L. Soh" country="Singapore" location="Sungei Seletar" pageId="53" pageNumber="944" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD4BA14FE08F924FD96F91E" box="[469,539,1687,1711]" pageId="53" pageNumber="944" type="male">1 male</specimenCount>
(
<collectionCode id="A7AD0285FFD4BA14FDFAF924FDD1F91E" box="[551,604,1687,1711]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="944">ZRC</collectionCode>
1971.9.24.9),
<location id="C463CC9BFFD4BA14FFACF904FE94F97E" LSID="urn:lsid:plazi:treatment:49152B56FFD4BA16FF18FCEEFA21F849:C463CC9BFFD4BA14FFACF904FE94F97E" box="[113,281,1719,1743]" country="Singapore" name="Sungei Seletar" pageId="53" pageNumber="944">Sungei Seletar</location>
, coll.
<collectorName id="6C49FF96FFD4BA14FEB8F904FE42F97E" box="[357,463,1719,1743]" pageId="53" pageNumber="944">C.L. Soh</collectorName>
,
<date id="B502BC80FFD4BA14FE03F904FD1EF97E" box="[478,659,1719,1743]" pageId="53" pageNumber="944" value="1966-03-29">
<collectingDate id="A5464568FFD4BA14FE03F904FD1EF97E" box="[478,659,1719,1743]" pageId="53" pageNumber="944" value="1966-03-29">29 March 1966</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71D4901DFFD4BA14FD7FF904FDF5F8BE" ID-GBIF-Occurrence="3763237472" collectingDate="2001-09-20" collectionCode="ZRC" collectorName="R. Teo" country="Singapore" location="Pulau Ubin" municipality="Asam" pageId="53" pageNumber="944" specimenCode="ZRC 2003.0083" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD4BA14FD7FF904FD62F97E" box="[674,751,1719,1743]" pageId="53" pageNumber="944" type="male">1 male</specimenCount>
(
<specimenCode id="911A323BFFD4BA14FFA4F964FEA6F95E" box="[121,299,1751,1775]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="944">ZRC 2003.0083</specimenCode>
),
<location id="C463CC9BFFD4BA14FEE0F964FE35F95E" LSID="urn:lsid:plazi:treatment:49152B56FFD4BA16FF18FCEEFA21F849:C463CC9BFFD4BA14FEE0F964FE35F95E" box="[317,440,1751,1775]" country="Singapore" municipality="Asam" name="Pulau Ubin" pageId="53" pageNumber="944">Pulau Ubin</location>
, vicinity of
<collectingMunicipality id="2167003AFFD4BA14FDEAF964FDF8F95E" box="[567,629,1751,1775]" pageId="53" pageNumber="944">Asam</collectingMunicipality>
mangroves (
<collectingMethod id="18FDE257FFD4BA14FFA4F944FF7CF8BE" box="[121,241,1783,1807]" pageId="53" pageNumber="944">pitfall trap</collectingMethod>
), coll.
<collectorName id="6C49FF96FFD4BA14FE9CF944FE02F8BF" box="[321,399,1783,1806]" pageId="53" pageNumber="944">R. Teo</collectorName>
,
<date id="B502BC80FFD4BA14FE43F944FDF5F8BE" box="[414,632,1783,1807]" pageId="53" pageNumber="944" value="2001-09-20">
<collectingDate id="A5464568FFD4BA14FE43F944FDF5F8BE" box="[414,632,1783,1807]" pageId="53" pageNumber="944" value="2001-09-20">20 September 2001</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71D4901DFFD4BA14FD5BF944FF52F8FE" ID-GBIF-Occurrence="3763237352" collectingDate="2000-07" collectionCode="ZRC" collectorName="C. D. Schubart" country="Singapore" location="Pulau Ubin" pageId="53" pageNumber="944" specimenCode="ZRC 2000.2019" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD4BA14FD5BF944FD58F8BE" box="[646,725,1783,1807]" pageId="53" pageNumber="944" type="male">1 male</specimenCount>
,
<specimenCount id="D7BA51C9FFD4BA14FD3EF944FF37F89E" pageId="53" pageNumber="944" type="female">1 female</specimenCount>
(
<specimenCode id="911A323BFFD4BA14FF14F8A4FEF0F89E" box="[201,381,1815,1839]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="944">ZRC 2000.2019</specimenCode>
),
<location id="C463CC9BFFD4BA14FE4DF8A4FD80F89E" LSID="urn:lsid:plazi:treatment:49152B56FFD4BA16FF18FCEEFA21F849:C463CC9BFFD4BA14FE4DF8A4FD80F89E" box="[400,525,1815,1839]" country="Singapore" name="Pulau Ubin" pageId="53" pageNumber="944">Pulau Ubin</location>
, coll.
<collectorName id="6C49FF96FFD4BA14FD8DF8A4FD60F89E" box="[592,749,1815,1839]" pageId="53" pageNumber="944">C.D. Schubart</collectorName>
,
<date id="B502BC80FFD4BA14FFACF884FF52F8FE" box="[113,223,1847,1871]" pageId="53" pageNumber="944" value="2000-07">
<collectingDate id="A5464568FFD4BA14FFACF884FF52F8FE" box="[113,223,1847,1871]" pageId="53" pageNumber="944" value="2000-07">July 2000</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71D4901DFFD4BA14FF31F884FD61F8DE" ID-GBIF-Occurrence="3763237359" collectingDate="2011-11-25" collectionCode="ZRC" collectorName="C. D. Schubart" country="Singapore" location="Chek Jawa" municipality="Pulau Ubin" pageId="53" pageNumber="944" specimenCount="3" specimenCount-male="3" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD4BA14FF31F884FECDF8FE" box="[236,320,1847,1871]" pageId="53" pageNumber="944" type="male">3 males</specimenCount>
(
<collectionCode id="A7AD0285FFD4BA14FE8DF884FE04F8FE" box="[336,393,1847,1871]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="944">ZRC</collectionCode>
), swamp along dirt road to
<location id="C463CC9BFFD4BA14FD65F884FF24F8DF" LSID="urn:lsid:plazi:treatment:49152B56FFD4BA16FF18FCEEFA21F849:C463CC9BFFD4BA14FD65F884FF24F8DF" country="Singapore" municipality="Pulau Ubin" name="Chek Jawa" pageId="53" pageNumber="944">Chek Jawa</location>
,
<collectingMunicipality id="2167003AFFD4BA14FF68F8E4FEBCF8DE" box="[181,305,1879,1903]" pageId="53" pageNumber="944">Pulau Ubin</collectingMunicipality>
, coll.
<collectorName id="6C49FF96FFD4BA14FEAEF8E4FD83F8DE" box="[371,526,1879,1903]" pageId="53" pageNumber="944">C.D. Schubart</collectorName>
,
<date id="B502BC80FFD4BA14FDC4F8E4FD61F8DE" box="[537,748,1879,1903]" pageId="53" pageNumber="944" value="2011-11-25">
<collectingDate id="A5464568FFD4BA14FDC4F8E4FD61F8DE" box="[537,748,1879,1903]" pageId="53" pageNumber="944" value="2011-11-25">25 November 2011</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71D4901DFFD4BA14FFACF8C4FD58F81E" ID-GBIF-Occurrence="3763237387" collectingDate="2001-11-16" collectionCode="ZRC" collectorName="B. Y. Lee" country="Singapore" location="Pulau Tekong" pageId="53" pageNumber="944" specimenCode="ZRC 2003.0084" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD4BA14FFACF8C4FF30F83E" box="[113,189,1911,1935]" pageId="53" pageNumber="944" type="male">1 male</specimenCount>
(
<specimenCode id="911A323BFFD4BA14FF12F8C4FE0BF83E" box="[207,390,1911,1935]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="944">ZRC 2003.0084</specimenCode>
), culvert beside reservoir,
<location id="C463CC9BFFD4BA14FD6EF8C4FF4BF81E" LSID="urn:lsid:plazi:treatment:49152B56FFD4BA16FF18FCEEFA21F849:C463CC9BFFD4BA14FD6EF8C4FF4BF81E" country="Singapore" name="Pulau Tekong" pageId="53" pageNumber="944">Pulau Tekong</location>
,
<collectingCountry id="B9ABDAD0FFD4BA14FF09F824FECEF81E" box="[212,323,1943,1967]" name="Singapore" pageId="53" pageNumber="944">Singapore</collectingCountry>
, coll.
<collectorName id="6C49FF96FFD4BA14FE54F824FE7DF81F" box="[393,496,1943,1966]" pageId="53" pageNumber="944">B.Y. Lee</collectorName>
,
<date id="B502BC80FFD4BA14FE20F824FD58F81E" box="[509,725,1943,1967]" pageId="53" pageNumber="944" value="2001-11-16">
<collectingDate id="A5464568FFD4BA14FE20F824FD58F81E" box="[509,725,1943,1967]" pageId="53" pageNumber="944" value="2001-11-16">16 November 2001</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71D4901DFFD4BA14FD3EF824FF27F7BE" ID-GBIF-Occurrence="3763237525" collectingDate="2002-01-31" collectionCode="ZRC" collectorName="T. M. Leong" country="Singapore" location="Pulau Tekong" pageId="53" pageNumber="944" specimenCode="ZRC 2013.1116" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD4BA14FD3EF824FF37F87E" pageId="53" pageNumber="944" type="female">1 female</specimenCount>
(11.5 × 10.0 mm) (
<specimenCode id="911A323BFFD4BA14FE40F804FDD9F87E" box="[413,596,1975,1999]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="944">ZRC 2013.1116</specimenCode>
), near stream, northern axis of
<location id="C463CC9BFFD4BA14FEC3F864FE39F85E" LSID="urn:lsid:plazi:treatment:49152B56FFD4BA16FF18FCEEFA21F849:C463CC9BFFD4BA14FEC3F864FE39F85E" box="[286,436,2007,2031]" country="Singapore" name="Pulau Tekong" pageId="53" pageNumber="944">Pulau Tekong</location>
, coll.
<collectorName id="6C49FF96FFD4BA14FE2FF864FDF9F85E" box="[498,628,2007,2031]" pageId="53" pageNumber="944">T.M. Leong</collectorName>
,
<date id="B502BC80FFD4BA14FDA0F864FF27F7BE" pageId="53" pageNumber="944" value="2002-01-31">
<collectingDate id="A5464568FFD4BA14FDA0F864FF27F7BE" pageId="53" pageNumber="944" value="2002-01-31">31 January 2002</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71D4901DFFD4BA14FF65F844FDB2F7FE" ID-GBIF-Occurrence="3763237393" collectingDate="2012-03-24" collectionCode="ZRC" collectorName="M. Chua" country="Singapore" elevation="300" pageId="53" pageNumber="944" specimenCount="1" typeStatus="paralectotype">
1 ex-ovigerous female (19.1 ×
<quantity id="064437A5FFD4BA14FDD2F844FDFEF7BF" box="[527,627,2039,2063]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" pageId="53" pageNumber="944" unit="mm" value="17.8">17.8 mm</quantity>
, first zoeae reared and preserved) (
<collectionCode id="A7AD0285FFD4BA14FEA9F7A4FE20F79E" box="[372,429,2071,2095]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="944">ZRC</collectionCode>
), in secondary forest,
<quantity id="064437A5FFD4BA14FD74F7A4FD7DF79F" box="[681,752,2071,2095]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="53" pageNumber="944" unit="m" value="300.0">
<elevation id="4A917D73FFD4BA14FD74F7A4FD7DF79F" box="[681,752,2071,2095]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="53" pageNumber="944" unit="m" value="300.0">300 m</elevation>
</quantity>
from coast, coll.
<collectorName id="6C49FF96FFD4BA14FEF4F784FE06F7FE" box="[297,395,2103,2127]" pageId="53" pageNumber="944">M. Chua</collectorName>
,
<date id="B502BC80FFD4BA14FE4BF784FDB2F7FE" box="[406,575,2103,2127]" pageId="53" pageNumber="944" value="2012-03-24">
<collectingDate id="A5464568FFD4BA14FE4BF784FDB2F7FE" box="[406,575,2103,2127]" pageId="53" pageNumber="944" value="2012-03-24">24 March 2012</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="71D4901DFFD4BA14FD97F784FB65FF44" ID-GBIF-Occurrence="3763237474" collectingDate="1967-08-13" collectionCode="ZRC" country="Malaysia" location="Pangkor Island" municipality="Straits" pageId="53" pageNumber="944" specimenCode="ZRC 1984.8031" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
PENINSULAR
<collectingCountry id="B9ABDAD0FFD4BA14FCF0FF0EFC37FF64" box="[813,954,189,213]" name="Malaysia" pageId="53" pageNumber="944">MALAYSIA</collectingCountry>
<specimenCount id="D7BA51C9FFD4BA14FC07FF0EFBABFF64" box="[986,1062,189,213]" pageId="53" pageNumber="944" type="male">1 male</specimenCount>
(
<specimenCode id="911A323BFFD4BA14FBEAFF0EFB66FF64" box="[1079,1259,189,213]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="944">ZRC 1984.8031</specimenCode>
),
<location id="C463CC9BFFD4BA14FADCFF0EFA25FF64" LSID="urn:lsid:plazi:treatment:49152B56FFD4BA16FF18FCEEFA21F849:C463CC9BFFD4BA14FADCFF0EFA25FF64" box="[1281,1448,189,213]" country="Malaysia" municipality="Straits" name="Pangkor Island" pageId="53" pageNumber="944">Pangkor Island</location>
,
<collectingMunicipality id="2167003AFFD4BA14FCF0FF6EFCFCFF44" box="[813,881,221,245]" pageId="53" pageNumber="944">Straits</collectingMunicipality>
of
<collectingCountry id="B9ABDAD0FFD4BA14FC4AFF6EFC79FF44" box="[919,1012,221,245]" name="Malaysia" pageId="53" pageNumber="944">Malacca</collectingCountry>
, coll.
<date id="B502BC80FFD4BA14FBE5FF6EFB65FF44" box="[1080,1256,221,246]" pageId="53" pageNumber="944" value="1967-08-13">
<collectingDate id="A5464568FFD4BA14FBE5FF6EFB65FF44" box="[1080,1256,221,246]" pageId="53" pageNumber="944" value="1967-08-13">13 August 1967</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71D4901DFFD4BA14FB28FF6EFBF0FE84" ID-GBIF-Occurrence="3763237463" collectingDate="1998-06-17" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Malaysia" location="Pulau Langkawi" municipality="Temurun" pageId="53" pageNumber="944" specimenCode="ZRC 2010.0035" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD4BA14FB28FF6EFACDFF44" box="[1269,1344,221,245]" pageId="53" pageNumber="944" type="male">1 male</specimenCount>
,
<specimenCount id="D7BA51C9FFD4BA14FA90FF6EFA21FF44" box="[1357,1452,221,245]" pageId="53" pageNumber="944" type="female">1 female</specimenCount>
(
<specimenCode id="911A323BFFD4BA14FCE9FF4EFC7DFEA4" box="[820,1008,253,277]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="944">ZRC 2010.0035</specimenCode>
),
<location id="C463CC9BFFD4BA14FBDAFF4EFB4FFEA4" LSID="urn:lsid:plazi:treatment:49152B56FFD4BA16FF18FCEEFA21F849:C463CC9BFFD4BA14FBDAFF4EFB4FFEA4" box="[1031,1218,253,277]" country="Malaysia" municipality="Temurun" name="Pulau Langkawi" pageId="53" pageNumber="944">Pulau Langkawi</location>
,
<collectingMunicipality id="2167003AFFD4BA14FB0CFF4DFABBFEA4" box="[1233,1334,254,277]" pageId="53" pageNumber="944">Temurun</collectingMunicipality>
waterfall, coll.
<collectorName id="6C49FF96FFD4BA14FCB8FEADFC54FE87" box="[869,985,286,310]" pageId="53" pageNumber="944">P.K.L. Ng</collectorName>
,
<date id="B502BC80FFD4BA14FC3BFEAEFBF0FE84" box="[998,1149,285,309]" pageId="53" pageNumber="944" value="1998-06-17">
<collectingDate id="A5464568FFD4BA14FC3BFEAEFBF0FE84" box="[998,1149,285,309]" pageId="53" pageNumber="944" value="1998-06-17">17 June 1998</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71D4901DFFD4BA14FB51FEAEFC33FE24" ID-GBIF-Occurrence="3763237362" collectingDate="2003-04-01" collectionCode="ZRC" collectorName="Universiti Sains" country="Malaysia" location="Sungei Temurun Datai" municipality="Langkawi" pageId="53" pageNumber="944" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Kedah" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD4BA14FB51FEAEFB5AFE84" box="[1164,1239,285,309]" pageId="53" pageNumber="944" type="male">1 male</specimenCount>
(13.9 ×
<quantity id="064437A5FFD4BA14FAE7FEAEFA2DFE84" box="[1338,1440,285,309]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="53" pageNumber="944" unit="mm" value="12.9">12.9 mm</quantity>
),
<specimenCount id="D7BA51C9FFD4BA14FCF0FE8EFC06FEE4" box="[813,907,317,341]" pageId="53" pageNumber="944" type="female">1 female</specimenCount>
(13.1 ×
<quantity id="064437A5FFD4BA14FC34FE8EFBC6FEE4" box="[1001,1099,317,341]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="53" pageNumber="944" unit="mm" value="11.6">11.6 mm</quantity>
) (
<collectionCode id="A7AD0285FFD4BA14FBBFFE8EFB16FEE4" box="[1122,1179,317,341]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="944">ZRC</collectionCode>
),
<location id="C463CC9BFFD4BA14FB72FE8EFA2AFEE4" LSID="urn:lsid:plazi:treatment:49152B56FFD4BA16FF18FCEEFA21F849:C463CC9BFFD4BA14FB72FE8EFA2AFEE4" box="[1199,1447,317,341]" country="Malaysia" municipality="Langkawi" name="Sungei Temurun Datai" pageId="53" pageNumber="944" stateProvince="Kedah">Sungei Temurun Datai</location>
,
<collectingMunicipality id="2167003AFFD4BA14FCF0FEEEFC17FEC4" box="[813,922,349,373]" pageId="53" pageNumber="944">Langkawi</collectingMunicipality>
,
<collectingRegion id="037854A2FFD4BA14FC7EFEEEFC66FEC4" box="[931,1003,349,373]" country="Malaysia" name="Kedah" pageId="53" pageNumber="944">Kedah</collectingRegion>
,
<collectingCountry id="B9ABDAD0FFD4BA14FC29FEEEFBD5FEC4" box="[1012,1112,349,373]" name="Malaysia" pageId="53" pageNumber="944">Malaysia</collectingCountry>
, coll.
<collectorName id="6C49FF96FFD4BA14FB48FEEEFAB3FEC4" box="[1173,1342,349,373]" pageId="53" pageNumber="944">Universiti Sains</collectorName>
<collectingCountry id="B9ABDAD0FFD4BA14FA9EFEEEFA2AFEC4" box="[1347,1447,349,373]" name="Malaysia" pageId="53" pageNumber="944">Malaysia</collectingCountry>
,
<date id="B502BC80FFD4BA14FCF0FECEFC33FE24" box="[813,958,381,405]" pageId="53" pageNumber="944" value="2003-04-01">
<collectingDate id="A5464568FFD4BA14FCF0FECEFC33FE24" box="[813,958,381,405]" pageId="53" pageNumber="944" value="2003-04-01">1 April 2003</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71D4901DFFD4BA14FC16FECEFCE8FE44" ID-GBIF-Occurrence="3763237507" collectingDate="2006-03-15" collectionCode="NHMW" collectorName="C. D. Schubart" country="Malaysia" location="Pulau Langkawi" municipality="Air Temurun" pageId="53" pageNumber="944" specimenCode="NHMW 26842" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Kedah" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD4BA14FC16FECEFB9BFE24" box="[971,1046,381,405]" pageId="53" pageNumber="944" type="male">1 male</specimenCount>
(19.4 ×
<quantity id="064437A5FFD4BA14FBA5FECEFB50FE24" box="[1144,1245,381,405]" metricMagnitude="-2" metricUnit="m" metricValue="1.8199999999999998" pageId="53" pageNumber="944" unit="mm" value="18.2">18.2 mm</quantity>
),
<specimenCount id="D7BA51C9FFD4BA14FB2EFECEFADFFE24" box="[1267,1362,381,405]" pageId="53" pageNumber="944" type="female">1 female</specimenCount>
(19.3 ×
<quantity id="064437A5FFD4BA14FCF0FE2EFC03FE04" box="[813,910,413,437]" metricMagnitude="-2" metricUnit="m" metricValue="1.77" pageId="53" pageNumber="944" unit="mm" value="17.7">17.7 mm</quantity>
) (
<specimenCode id="911A323BFFD4BA14FC78FE2EFBC0FE04" box="[933,1101,413,437]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="53" pageNumber="944" type="Museum">NHMW 26842</specimenCode>
),
<location id="C463CC9BFFD4BA14FBBDFE2EFA9FFE04" LSID="urn:lsid:plazi:treatment:49152B56FFD4BA16FF18FCEEFA21F849:C463CC9BFFD4BA14FBBDFE2EFA9FFE04" box="[1120,1298,413,437]" country="Malaysia" municipality="Air Temurun" name="Pulau Langkawi" pageId="53" pageNumber="944" stateProvince="Kedah">Pulau Langkawi</location>
,
<collectingMunicipality id="2167003AFFD4BA14FAC3FE2EFA21FE04" box="[1310,1452,413,437]" pageId="53" pageNumber="944">Air Temurun</collectingMunicipality>
waterfall and stream, coll.
<collectorName id="6C49FF96FFD4BA14FB88FE0EFB7DFE64" box="[1109,1264,445,469]" pageId="53" pageNumber="944">C.D. Schubart</collectorName>
et al.,
<date id="B502BC80FFD4BA14FA9CFE0EFCE8FE44" pageId="53" pageNumber="944" value="2006-03-15">
<collectingDate id="A5464568FFD4BA14FA9CFE0EFCE8FE44" pageId="53" pageNumber="944" value="2006-03-15">15 March 2006</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71D4901DFFD4BA14FCB3FE6EFBA2FDC4" ID-GBIF-Occurrence="3763237520" collectingDate="2016-12-18" collectionCode="ZRC" collectorName="P. K. L. Ng &amp; P. Y. C. Ng" country="Malaysia" county="Peninsular" elevation="200" location="Temurun" municipality="Langkawi" pageId="53" pageNumber="944" specimenCode="ZRC 2016.0608" specimenCount="3" specimenCount-female="3" specimenCount-male="8" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD4BA14FCB3FE6EFC32FE44" box="[878,959,477,501]" pageId="53" pageNumber="944" type="male">8 males</specimenCount>
(1 post-molt, badly damaged),
<specimenCount id="D7BA51C9FFD4BA14FAD8FE6EFAE4FE44" box="[1285,1385,477,501]" pageId="53" pageNumber="944" type="female">3 females</specimenCount>
(
<specimenCode id="911A323BFFD4BA14FAABFE6EFC2EFDA4" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="944">ZRC 2016.0608</specimenCode>
),
<location id="C463CC9BFFD4BA14FC69FE4DFB9BFDA4" LSID="urn:lsid:plazi:treatment:49152B56FFD4BA16FF18FCEEFA21F849:C463CC9BFFD4BA14FC69FE4DFB9BFDA4" box="[948,1046,510,533]" country="Malaysia" county="Peninsular" municipality="Langkawi" name="Temurun" pageId="53" pageNumber="944">Temurun</location>
waterfall, at base of waterfalls, under rocks at side of stream, near bank, about
<quantity id="064437A5FFD4BA14FB2BFDAEFAB0FD84" box="[1270,1341,541,565]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="53" pageNumber="944" unit="m" value="200.0">
<elevation id="4A917D73FFD4BA14FB2BFDAEFAB0FD84" box="[1270,1341,541,565]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="53" pageNumber="944" unit="m" value="200.0">200 m</elevation>
</quantity>
from sea,
<collectingMunicipality id="2167003AFFD4BA14FCF0FD8EFC17FDE4" box="[813,922,573,597]" pageId="53" pageNumber="944">Langkawi</collectingMunicipality>
,
<collectingCounty id="2862E2CCFFD4BA14FC7AFD8EFB95FDE4" box="[935,1048,573,597]" pageId="53" pageNumber="944">Peninsular</collectingCounty>
<collectingCountry id="B9ABDAD0FFD4BA14FBFCFD8EFB08FDE4" box="[1057,1157,573,597]" name="Malaysia" pageId="53" pageNumber="944">Malaysia</collectingCountry>
, coll.
<collectorName id="6C49FF96FFD4BA14FB16FD8DFAB1FDE7" box="[1227,1340,574,598]" pageId="53" pageNumber="944">P.K.L. Ng</collectorName>
&amp;
<collectorName id="6C49FF96FFD4BA14FABEFD8EFCDDFDC7" pageId="53" pageNumber="944">P.Y.C. Ng</collectorName>
,
<date id="B502BC80FFD4BA14FC80FDEEFBA2FDC4" box="[861,1071,605,629]" pageId="53" pageNumber="944" value="2016-12-18">
<collectingDate id="A5464568FFD4BA14FC80FDEEFBA2FDC4" box="[861,1071,605,629]" pageId="53" pageNumber="944" value="2016-12-18">18 December 2016</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFD4BA14FCF0FD2EFA21FB44" pageId="53" pageNumber="944" type="diagnosis">
<paragraph id="C1039A40FFD4BA14FCF0FD2EFBB3FBA4" blockId="53.[812,1452,669,1045]" pageId="53" pageNumber="944">
<emphasis id="F3C84652FFD4BA14FCF0FD2EFC17FD04" bold="true" box="[813,922,669,693]" pageId="53" pageNumber="944">Diagnosis</emphasis>
. Carapace transversely rectangular; frontal margin less wide, median concavity separating lobes relatively more shallow; epibranchial tooth distinct, separated from rest of margin by deep notch; posterolateral margins subparallel; outer surface of chela gently convex, covered with small rounded granules, ventral margin of palm sinuous, unarmed; suture between male thoracic sternites 3 and 4 distinct; male pleon broadly triangular; male sternopleonal cavity with press-button of pleonal locking mechanism on sternite 5; distal part of G1 bifurcated or bilobed, with chitinous part very short. In life, chelae red, pigmentation extending to midway of white fingers.
</paragraph>
<paragraph id="C1039A40FFD4BA14FCF0FB8EFA21FB44" blockId="53.[812,1452,1085,1270]" pageId="53" pageNumber="944">
<emphasis id="F3C84652FFD4BA14FCF0FB8EFCF2FBE4" bold="true" box="[813,895,1085,1109]" pageId="53" pageNumber="944">Colour</emphasis>
. The carapace of fresh
<taxonomicName id="06BCE1C3FFD4BA14FB4CFB8DFA92FBE4" baseAuthorityName="De Man" baseAuthorityYear="1887" box="[1169,1311,1085,1109]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="944" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="F3C84652FFD4BA14FB4CFB8DFA92FBE4" box="[1169,1311,1085,1109]" italics="true" pageId="53" pageNumber="944">P. edwardsii</emphasis>
</taxonomicName>
is brown to reddish-brown, with many specimens having the posterior part of the carapace slightly to much darker in colour (
<figureCitation id="598786C5FFD4BA14FA5DFBCEFCF0FB04" captionStart="Fig" captionStartId="88.[136,171,1875,1897]" captionTargetBox="[136,1474,324,1860]" captionTargetId="figure-7@88.[136,1474,324,1860]" captionTargetPageId="88" captionText="Fig. 56. Colour in life. A, B, Pseudosesarma edwardsii, male (20.5 × 19.3 mm) (ZRC 2016.0608), Langkawi [photograph: Paul Ng]; CE, P. edwardsii, male (19.4 × 18.2 mm) (SMF), Langkawi, Peninsular Malaysia [photograph: CD Schubart]; F, G, P. crassimanum, (12.6 × 10.5 mm) (ZRC 2010.0035), Tioman, Peninsular Malaysia [photograph: PKL Ng]; H, P. boulengeri, specimen not preserved [photograph: Reza Naderloo]." figureDoi="http://doi.org/10.5281/zenodo.5351399" httpUri="https://zenodo.org/record/5351399/files/figure.png" pageId="53" pageNumber="944">Fig. 56A, C</figureCitation>
), with the outer surface of the chela red, the colour extending to at least the middle of both fingers, with the distal half white (
<figureCitation id="598786C5FFD4BA14FC7AFB6EFBC8FB44" box="[935,1093,1245,1269]" captionStart="Fig" captionStartId="88.[136,171,1875,1897]" captionTargetBox="[136,1474,324,1860]" captionTargetId="figure-7@88.[136,1474,324,1860]" captionTargetPageId="88" captionText="Fig. 56. Colour in life. A, B, Pseudosesarma edwardsii, male (20.5 × 19.3 mm) (ZRC 2016.0608), Langkawi [photograph: Paul Ng]; CE, P. edwardsii, male (19.4 × 18.2 mm) (SMF), Langkawi, Peninsular Malaysia [photograph: CD Schubart]; F, G, P. crassimanum, (12.6 × 10.5 mm) (ZRC 2010.0035), Tioman, Peninsular Malaysia [photograph: PKL Ng]; H, P. boulengeri, specimen not preserved [photograph: Reza Naderloo]." figureDoi="http://doi.org/10.5281/zenodo.5351399" httpUri="https://zenodo.org/record/5351399/files/figure.png" pageId="53" pageNumber="944">Fig. 56B, D, E</figureCitation>
) (see also
<bibRefCitation id="A52DE7B1FFD4BA14FB6AFB6DFA2DFB44" author="Ng &amp; Schubart" box="[1207,1440,1245,1270]" firstAuthor="Ng" pageId="53" pageNumber="944" pagination="655 - 669" refId="ref74064" refString="Ng PKL &amp; Schubart CD (2017) On the taxonomy of Pseudosesarma edwardsii (De Man, 1887) and P. crassimanum (De Man, 1887) (Crustacea: Decapoda: Brachyura: Sesarmidae), with description of a new species from Sri Lanka. Raffles Bulletin of Zoology, 65: 655 - 669." type="journal article" year="2017">Ng &amp; Schubart, 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFD4BA16FCF0FAAEFB8AF9E9" lastPageId="55" lastPageNumber="946" pageId="53" pageNumber="944" type="discussion">
<paragraph id="C1039A40FFD4BA14FCF0FAAEFB77F9E7" blockId="53.[812,1452,1309,1622]" pageId="53" pageNumber="944">
<emphasis id="F3C84652FFD4BA14FCF0FAAEFC18FA84" bold="true" box="[813,917,1309,1333]" pageId="53" pageNumber="944">Remarks</emphasis>
. The taxonomy of this species has been discussed at length by
<bibRefCitation id="A52DE7B1FFD4BA14FC6BFA8DFB27FAE4" author="Ng &amp; Schubart" box="[950,1194,1341,1366]" firstAuthor="Ng" pageId="53" pageNumber="944" pagination="655 - 669" refId="ref74064" refString="Ng PKL &amp; Schubart CD (2017) On the taxonomy of Pseudosesarma edwardsii (De Man, 1887) and P. crassimanum (De Man, 1887) (Crustacea: Decapoda: Brachyura: Sesarmidae), with description of a new species from Sri Lanka. Raffles Bulletin of Zoology, 65: 655 - 669." type="journal article" year="2017">Ng &amp; Schubart (2017)</bibRefCitation>
and there is no need to further elaborate. The date of publication for this species (and
<taxonomicName id="06BCE1C3FFD4BA14FCF1FACDFC57FA24" baseAuthorityName="De Man" baseAuthorityYear="1887" box="[812,986,1406,1429]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="944" phylum="Arthropoda" rank="species" species="crassimanum">
<emphasis id="F3C84652FFD4BA14FCF1FACDFC57FA24" box="[812,986,1406,1429]" italics="true" pageId="53" pageNumber="944">P. crassimanum</emphasis>
</taxonomicName>
) follows
<bibRefCitation id="A52DE7B1FFD4BA14FB9BFACDFB74FA24" author="Ng" box="[1094,1273,1405,1430]" etAl="et al." firstAuthor="Ng" pageId="53" pageNumber="944" pagination="216 - 225" refId="ref73795" refString="Ng PKL, Lee BY &amp; Tan HH (2015) Observations on the taxonomy and ecology of Labuanium politum (De Man, 1887) (Crustacea: Decapoda: Sesarmidae), an obligate arboreal crab on the nipah palm, Nypa fructicans (Arecales: Arecaceae). Raffles Bulletin of Zoology, Supplement 31: 216 - 225." type="journal article" year="2015">Ng et al. (2015)</bibRefCitation>
.
<bibRefCitation id="A52DE7B1FFD4BA14FAD8FACDFCF8FA04" author="Ng &amp; Schubart" firstAuthor="Ng" pageId="53" pageNumber="944" pagination="655 - 669" refId="ref74064" refString="Ng PKL &amp; Schubart CD (2017) On the taxonomy of Pseudosesarma edwardsii (De Man, 1887) and P. crassimanum (De Man, 1887) (Crustacea: Decapoda: Brachyura: Sesarmidae), with description of a new species from Sri Lanka. Raffles Bulletin of Zoology, 65: 655 - 669." type="journal article" year="2017">Ng &amp; Schubart (2017)</bibRefCitation>
clarified the differences with
<taxonomicName id="06BCE1C3FFD4BA14FB15FA2DFC27FA64" authority="(De Man, 1887)" baseAuthorityName="De Man" baseAuthorityYear="1887" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="944" phylum="Arthropoda" rank="species" species="crassimanum">
<emphasis id="F3C84652FFD4BA14FB15FA2DFAF7FA04" box="[1224,1402,1438,1461]" italics="true" pageId="53" pageNumber="944">P. crassimanum</emphasis>
(
<bibRefCitation id="A52DE7B1FFD4BA14FA56FA2EFC2FFA64" author="De Man" firstAuthor="De Man" pageId="53" pageNumber="944" pagination="639 - 722" refId="ref72089" refString="Man JG De (1887) Ubersicht der Indopacifischen Arten der Gattung Sesarma Say nebst einer Kritik der von W. Hess und E. Nauck in den Jahren 1865 und 1880 beschrieben Decapoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 2: 639 - 722." type="journal article" year="1887">De Man, 1887</bibRefCitation>
)
</taxonomicName>
and further realised that
<taxonomicName id="06BCE1C3FFD4BA14FB68FA0DFAB6FA64" baseAuthorityName="De Man" baseAuthorityYear="1887" box="[1205,1339,1469,1493]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="944" phylum="Arthropoda" rank="species" sensu="lato" species="edwardsii">
<emphasis id="F3C84652FFD4BA14FB68FA0DFAB6FA64" box="[1205,1339,1469,1493]" italics="true" pageId="53" pageNumber="944">P. edwardsii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8FBFB29FFD4BA14FA9FFA0DFAF1FA64" box="[1346,1404,1469,1493]" pageId="53" pageNumber="944" sensu="lato">s. lat.</taxonomicNameLabel>
was harbouring a number of closely related species, including
<taxonomicName id="06BCE1C3FFD4BA14FCF0FA4DFB49F9A4" authority="Ng &amp; Schubart, 2017" authorityName="Ng &amp; Schubart" authorityYear="2017" box="[813,1220,1533,1558]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="944" phylum="Arthropoda" rank="species" species="anteactum">
<emphasis id="F3C84652FFD4BA14FCF0FA4DFC4CF9A4" box="[813,961,1534,1557]" italics="true" pageId="53" pageNumber="944">P. anteactum</emphasis>
<bibRefCitation id="A52DE7B1FFD4BA14FC11FA4DFB49F9A4" author="Ng &amp; Schubart" box="[972,1220,1533,1558]" firstAuthor="Ng" pageId="53" pageNumber="944" pagination="655 - 669" refId="ref74064" refString="Ng PKL &amp; Schubart CD (2017) On the taxonomy of Pseudosesarma edwardsii (De Man, 1887) and P. crassimanum (De Man, 1887) (Crustacea: Decapoda: Brachyura: Sesarmidae), with description of a new species from Sri Lanka. Raffles Bulletin of Zoology, 65: 655 - 669." type="journal article" year="2017">Ng &amp; Schubart, 2017</bibRefCitation>
</taxonomicName>
, from
<collectingCountry id="B9ABDAD0FFD4BA14FACEFA4EFA08F9A4" box="[1299,1413,1533,1557]" name="Sri Lanka" pageId="53" pageNumber="944">Sri Lanka</collectingCountry>
,
<taxonomicName id="06BCE1C3FFD4BA14FA49FA4DFB4DF984" authority="Ng, Rani &amp; Nandan, 2017" authorityName="Ng, Rani &amp; Nandan" authorityYear="2017" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="944" phylum="Arthropoda" rank="species" species="glabrum">
<emphasis id="F3C84652FFD4BA14FA49FA4DFC04F984" italics="true" pageId="53" pageNumber="944">P. glabrum</emphasis>
<bibRefCitation id="A52DE7B1FFD4BA14FC4EF9ADFB4DF984" author="Ng, Rani &amp; Nandan" box="[915,1216,1565,1590]" firstAuthor="Ng" pageId="53" pageNumber="944" pagination="263 - 270" refId="ref73970" refString="Ng PKL, Rani V &amp; Nandan B (2017 a) A new species of Pseudosesarma Serene &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae) from Cochin in southwest India. Zootaxa, 4311 (2), 263 - 270." type="journal article" year="2017">Ng, Rani &amp; Nandan, 2017</bibRefCitation>
</taxonomicName>
, from western
<collectingCountry id="B9ABDAD0FFD4BA14FAB0F9AEFA2AF984" box="[1389,1447,1565,1589]" name="India" pageId="53" pageNumber="944">India</collectingCountry>
, and
<taxonomicName id="06BCE1C3FFD4BA14FC81F98DFBCFF9E4" authority="Ng, 2018" authorityName="Ng" authorityYear="2018" box="[860,1090,1597,1622]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="944" phylum="Arthropoda" rank="species" species="brehieri">
<emphasis id="F3C84652FFD4BA14FC81F98DFC5CF9E4" box="[860,977,1597,1621]" italics="true" pageId="53" pageNumber="944">P. brehieri</emphasis>
<bibRefCitation id="A52DE7B1FFD4BA14FC07F98DFBCFF9E4" author="Ng" box="[986,1090,1597,1622]" firstAuthor="Ng" pageId="53" pageNumber="944" pagination="185 - 197" refId="ref73639" refString="Ng PKL (2018) On two new species of potamid and sesarmid crabs (Decapoda, Brachyura) from caves in Myanmar. Crustaceana, 91 (2): 185 - 197." type="journal article" year="2018">Ng, 2018</bibRefCitation>
</taxonomicName>
, from
<collectingCountry id="B9ABDAD0FFD4BA14FB56F98DFB7BF9E7" box="[1163,1270,1598,1622]" name="Myanmar" pageId="53" pageNumber="944">Myanmar</collectingCountry>
.
</paragraph>
<paragraph id="C1039A40FFD4BA14FCF0F9CDFADFF8C4" blockId="53.[813,1452,1661,1909]" pageId="53" pageNumber="944">
<bibRefCitation id="A52DE7B1FFD4BA14FCF0F9CDFBA2F924" author="Ng &amp; Schubart" box="[813,1071,1661,1686]" firstAuthor="Ng" pageId="53" pageNumber="944" pagination="655 - 669" refId="ref74064" refString="Ng PKL &amp; Schubart CD (2017) On the taxonomy of Pseudosesarma edwardsii (De Man, 1887) and P. crassimanum (De Man, 1887) (Crustacea: Decapoda: Brachyura: Sesarmidae), with description of a new species from Sri Lanka. Raffles Bulletin of Zoology, 65: 655 - 669." type="journal article" year="2017">Ng &amp; Schubart (2017)</bibRefCitation>
observed that the proportions of the male pleon varies with size. In the largest male, the
<typeStatus id="1E0724E2FFD4BA14FCF0F90EFC1FF964" box="[813,914,1725,1749]" pageId="53" pageNumber="944" type="lectotype">lectotype</typeStatus>
, from the Mergui (17.5 ×
<quantity id="064437A5FFD4BA14FB74F90EFA87F964" box="[1193,1290,1725,1749]" metricMagnitude="-2" metricUnit="m" metricValue="1.61" pageId="53" pageNumber="944" unit="mm" value="16.1">16.1 mm</quantity>
, RMNH-D17), the pleon is very wide (
<figureCitation id="598786C5FFD4BA14FBEEF96EFB2FF944" box="[1075,1186,1757,1781]" captionStart="Fig" captionStartId="52.[136,171,1939,1961]" captionTargetBox="[173,1438,189,1924]" captionTargetId="figure-7@52.[173,1438,189,1924]" captionText="Fig. 30. Male anterior thoracic sternum and pleon of Pseudosesarma. A, P edwardsii, lectotype male (17.5 × 16.1 mm) (RMNH-D17a), Mergui Archipelago; B, P. edwardsii, male (13.4 × 11.9 mm) (ZRC 1965.8.2.81), Pulau Pawai, Singapore; C, P. crassimanum, male (15.9 × 14.5 mm) (RMNH-D23313), Singapore; D, P. glabrum, holotype male (13.8 × 12.0 mm) (CUSAT 2016-1), Kerala, India; E, P. anteactum, holotype male (16.7 × 14.7 mm) (ZRC 2016.0602), Sri Lanka; F, P. anteactum, paratype male (21.2 × 19.7 mm) (ZRC 2016.0603), Sri Lanka; G, P. brehieri, holotype male (17.4 × 15.7 mm) (ZRC 2016.0593), Myanmar; H, P. boulengeri, lectotype male (26.9 × 23.5 mm) (NHM 1919.11.14.1), Basra, Iraq." figureDoi="http://doi.org/10.5281/zenodo.6519011" httpUri="https://zenodo.org/record/6519011/files/figure.png" pageId="53" pageNumber="944">Figs. 30A</figureCitation>
, 34B) but it is narrower in smaller males (
<figureCitation id="598786C5FFD4BA14FC2DF94EFBD0F8A4" box="[1008,1117,1789,1813]" captionStart="Fig" captionStartId="52.[136,171,1939,1961]" captionTargetBox="[173,1438,189,1924]" captionTargetId="figure-7@52.[173,1438,189,1924]" captionText="Fig. 30. Male anterior thoracic sternum and pleon of Pseudosesarma. A, P edwardsii, lectotype male (17.5 × 16.1 mm) (RMNH-D17a), Mergui Archipelago; B, P. edwardsii, male (13.4 × 11.9 mm) (ZRC 1965.8.2.81), Pulau Pawai, Singapore; C, P. crassimanum, male (15.9 × 14.5 mm) (RMNH-D23313), Singapore; D, P. glabrum, holotype male (13.8 × 12.0 mm) (CUSAT 2016-1), Kerala, India; E, P. anteactum, holotype male (16.7 × 14.7 mm) (ZRC 2016.0602), Sri Lanka; F, P. anteactum, paratype male (21.2 × 19.7 mm) (ZRC 2016.0603), Sri Lanka; G, P. brehieri, holotype male (17.4 × 15.7 mm) (ZRC 2016.0593), Myanmar; H, P. boulengeri, lectotype male (26.9 × 23.5 mm) (NHM 1919.11.14.1), Basra, Iraq." figureDoi="http://doi.org/10.5281/zenodo.6519011" httpUri="https://zenodo.org/record/6519011/files/figure.png" pageId="53" pageNumber="944">Figs. 30B</figureCitation>
, 34C). Larger males also have proportionately stouter G1s, with the distal part distinctly bilobed (
<figureCitation id="598786C5FFD4BA14FC56F88EFB84F8E4" box="[907,1033,1853,1877]" captionStart="Fig" captionStartId="58.[136,171,1390,1412]" captionTargetBox="[136,1475,190,1374]" captionTargetId="figure-8@58.[125,1484,178,1386]" captionTargetPageId="58" captionText="Fig. 34. Pseudosesarma edwardsii. A, B, D, E, lectotype male (17.5 × 16.1 mm) (RMNH-D17a), Mergui Archipelago; C, male (13.3 × 12.9 mm) (ZRC 2000.2019), Pulau Ubin, Singapore; FH, male (20.5 × 19.3 mm) (ZRC 2016.0608), Langkawi; I, J, male (21.3 × 18.7 mm) (ZRC 2003.0084), Singapore; K, L, male (17.8 × 16.3 mm) (ZRC 2016.0608), Langkawi; M, N, male (12.5 × 10.4 mm) (ZRC 2016.0608), Langkawi. A, anterior thoracic sternites 14; B, C, male pleon; D, H, I, K, M, left G1 (dorsal view, denuded); E, G, J, K, N, left G1 (ventral view, denuded); H, left G2. Scales: AC = 2.0 mm; DJ = 1.0 mm; KN = 0.5 mm. (After Ng &amp; Schubart, 2017: figs. 6, 7)." figureDoi="http://doi.org/10.5281/zenodo.5351359" httpUri="https://zenodo.org/record/5351359/files/figure.png" pageId="53" pageNumber="944">Fig. 34D, E</figureCitation>
) while those of smaller males are more slender with the distal part less obviously bilobed.
</paragraph>
<paragraph id="C1039A40FFD4BA14FCF0F82EFAFAF784" blockId="53.[813,1452,1949,2101]" pageId="53" pageNumber="944">
The identity of “
<taxonomicName id="06BCE1C3FFD4BA14FC3FF82DFAABF804" authorityName="De Man" authorityYear="1887" box="[994,1318,1949,1973]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="944" phylum="Arthropoda" rank="species" species="edwardsii" subGenus="Sesarma">
<emphasis id="F3C84652FFD4BA14FC3FF82DFBB3F804" box="[994,1086,1950,1973]" italics="true" pageId="53" pageNumber="944">Sesarma</emphasis>
(
<emphasis id="F3C84652FFD4BA14FB8DF82DFB21F804" box="[1104,1196,1950,1973]" italics="true" pageId="53" pageNumber="944">Sesarma</emphasis>
)
<emphasis id="F3C84652FFD4BA14FB60F82EFAABF804" box="[1213,1318,1949,1973]" italics="true" pageId="53" pageNumber="944">edwardsii</emphasis>
</taxonomicName>
” from Pasir Ganing in western
<collectingRegion id="037854A2FFD4BA14FC21F80EFBD8F864" box="[1020,1109,1981,2005]" country="Indonesia" name="Sumatera Utara" pageId="53" pageNumber="944">Sumatra</collectingRegion>
by
<bibRefCitation id="A52DE7B1FFD4BA14FB5CF80EFAD1F864" author="Calman" box="[1153,1372,1981,2005]" firstAuthor="Calman" page="166" pageId="53" pageNumber="944" pagination="166 - 167" refId="ref68142" refString="Calman WT (1925) X. Crustacea Decapoda. Collected in Korinchi and on the West Sumatran Coast, by Messrs. H. C. Robinson and C. Boden Kloss. Results of an Expedition to Korinchi Peak, Sumatra. Journal of the Federated Malay States Museums, 8 (3): 166 - 167." type="journal article" year="1925">Calman (1925: 166)</bibRefCitation>
cannot be confirmed. It is within the range of variation of the present species, so it is here retained within its synoymy. The specimens should be re-examined when possible.
</paragraph>
<caption id="95C3CAC8FFD7BA17FF55F82EFA49F7F2" ID-DOI="http://doi.org/10.5281/zenodo.5351351" ID-Zenodo-Dep="5351351" httpUri="https://zenodo.org/record/5351351/files/figure.png" pageId="54" pageNumber="945" startId="54.[136,171,1949,1971]" targetBox="[169,1440,189,1933]" targetPageId="54">
<paragraph id="C1039A40FFD7BA17FF55F82EFA49F7F2" blockId="54.[136,1476,1948,2115]" pageId="54" pageNumber="945">
Fig. 31. Male anterior thoracic sternum and pleon. A,
<taxonomicName id="06BCE1C3FFD7BA17FD52F82EFCD1F803" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1869" box="[655,860,1949,1970]" class="Malacostraca" family="Sesarmidae" genus="Contusarma" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="945" phylum="Arthropoda" rank="species" species="bocourti">
<emphasis id="F3C84652FFD7BA17FD52F82EFCD1F803" box="[655,860,1949,1970]" italics="true" pageId="54" pageNumber="945">Contusarma bocourti</emphasis>
</taxonomicName>
, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand; B,
<taxonomicName id="06BCE1C3FFD7BA17FF78F809FE2FF87F" baseAuthorityName="Targioni Tozzetti" baseAuthorityYear="1877" box="[165,418,1977,1999]" class="Malacostraca" family="Sesarmidae" genus="Contusarma" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="945" phylum="Arthropoda" rank="species" species="cheirogonum">
<emphasis id="F3C84652FFD7BA17FF78F809FE2FF87F" box="[165,418,1977,1999]" italics="true" pageId="54" pageNumber="945">Contusarma cheirogonum</emphasis>
</taxonomicName>
, male (15.3 × 14.0 mm) (ZRC 2011.0924), Pulau Tioman, Malaysia; C,
<taxonomicName id="06BCE1C3FFD7BA17FBB1F809FAE4F87F" baseAuthorityName="Targioni Tozzetti" baseAuthorityYear="1877" box="[1132,1385,1977,1999]" class="Malacostraca" family="Sesarmidae" genus="Contusarma" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="945" phylum="Arthropoda" rank="species" species="cheirogonum">
<emphasis id="F3C84652FFD7BA17FBB1F809FAE4F87F" box="[1132,1385,1977,1999]" italics="true" pageId="54" pageNumber="945">Contusarma cheirogonum</emphasis>
</taxonomicName>
, neotype male (24.5 × 21.5 mm) (ZRC 1995.225), Bako National Park, Sarawak; D,
<taxonomicName id="06BCE1C3FFD7BA17FCB2F864FBFCF85D" baseAuthorityName="Miers" baseAuthorityYear="1880" box="[879,1137,2007,2028]" class="Malacostraca" family="Sesarmidae" genus="Miersarma" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="945" phylum="Arthropoda" rank="species" species="granosimanum">
<emphasis id="F3C84652FFD7BA17FCB2F864FBFCF85D" box="[879,1137,2007,2028]" italics="true" pageId="54" pageNumber="945">Miersarma granosimanum</emphasis>
</taxonomicName>
, lectotype male (16.9 × 14.8 mm) (NHM 1880.6), Borneo; E,
<taxonomicName id="06BCE1C3FFD7BA17FE41F847FD2DF7B8" baseAuthorityName="Miers" baseAuthorityYear="1880" box="[412,672,2036,2057]" class="Malacostraca" family="Sesarmidae" genus="Miersarma" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="945" phylum="Arthropoda" rank="species" species="granosimanum">
<emphasis id="F3C84652FFD7BA17FE41F847FD2DF7B8" box="[412,672,2036,2057]" italics="true" pageId="54" pageNumber="945">Miersarma granosimanum</emphasis>
</taxonomicName>
, male (22.5 × 19.2 mm) (ZRC 1965.7.29.164), Sedili River, Johor, Malaysia; F,
<taxonomicName id="06BCE1C3FFD7BA17FF55F7A3FEE6F794" baseAuthorityName="Zehntner" baseAuthorityYear="1894" box="[136,363,2064,2085]" class="Malacostraca" family="Sesarmidae" genus="Bresedium" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="945" phylum="Arthropoda" rank="species" species="laevimanum">
<emphasis id="F3C84652FFD7BA17FF55F7A3FEE6F794" box="[136,363,2064,2085]" italics="true" pageId="54" pageNumber="945">Bresedium laevimanum</emphasis>
</taxonomicName>
, lectotype male (20.0 × 17.6 mm) (MNHG), Borneo; G,
<taxonomicName id="06BCE1C3FFD7BA17FC43F7A3FB0CF794" baseAuthorityName="Zehntner" baseAuthorityYear="1894" box="[926,1153,2064,2085]" class="Malacostraca" family="Sesarmidae" genus="Bresedium" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="945" phylum="Arthropoda" rank="species" species="laevimanum">
<emphasis id="F3C84652FFD7BA17FC43F7A3FB0CF794" box="[926,1153,2064,2085]" italics="true" pageId="54" pageNumber="945">Bresedium laevimanum</emphasis>
</taxonomicName>
, male (26.3 × 23.7 mm) (NHM) (lectotype of
<taxonomicName id="06BCE1C3FFD7BA17FED9F79EFE30F7F3" baseAuthorityName="Tweedie" baseAuthorityYear="1940" box="[260,445,2093,2114]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="945" phylum="Arthropoda" rank="species" species="sediliensis">
<emphasis id="F3C84652FFD7BA17FED9F79EFE30F7F3" box="[260,445,2093,2114]" italics="true" pageId="54" pageNumber="945">Sesarma sediliensis</emphasis>
</taxonomicName>
), Johor, Malaysia; H,
<taxonomicName id="06BCE1C3FFD7BA17FD51F79EFCE6F7F3" baseAuthorityName="Zehntner" baseAuthorityYear="1894" box="[652,875,2093,2114]" class="Malacostraca" family="Sesarmidae" genus="Bresedium" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="945" phylum="Arthropoda" rank="species" species="laevimanum">
<emphasis id="F3C84652FFD7BA17FD51F79EFCE6F7F3" box="[652,875,2093,2114]" italics="true" pageId="54" pageNumber="945">Bresedium laevimanum</emphasis>
</taxonomicName>
, male (21.2 × 18.2 mm) (ZRC 1965.7.29.121), Johor, Malaysia.
</paragraph>
</caption>
<caption id="95C3CAC8FFD6BA16FFACFAF3FE84FA79" ID-DOI="http://doi.org/10.5281/zenodo.5351355" ID-Zenodo-Dep="5351355" httpUri="https://zenodo.org/record/5351355/files/figure.png" pageId="55" pageNumber="946" startId="55.[113,148,1344,1366]" targetBox="[113,1451,189,1328]" targetPageId="55">
<paragraph id="C1039A40FFD6BA16FFACFAF3FE84FA79" blockId="55.[113,1451,1344,1481]" pageId="55" pageNumber="946">
Fig. 32. Male sternopleonal cavity. A,
<taxonomicName id="06BCE1C3FFD6BA16FE33FAF3FD66FAE4" baseAuthorityName="De Man" baseAuthorityYear="1887" box="[494,747,1344,1365]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="946" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="F3C84652FFD6BA16FE33FAF3FD66FAE4" box="[494,747,1344,1365]" italics="true" pageId="55" pageNumber="946">Pseudosesarma edwardsii</emphasis>
</taxonomicName>
, male (20.8 × 18.6mm (ZRC 2003.0084), Pulau Tekong, Singapore; B,
<taxonomicName id="06BCE1C3FFD6BA16FFACFAEFFE01FAC3" baseAuthorityName="De Man" baseAuthorityYear="1887" box="[113,396,1372,1394]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="946" phylum="Arthropoda" rank="species" species="crassimanum">
<emphasis id="F3C84652FFD6BA16FFACFAEFFE01FAC3" box="[113,396,1372,1394]" italics="true" pageId="55" pageNumber="946">Pseudosesarma crassimanum</emphasis>
</taxonomicName>
, male (15.1 × 13.3 mm) (ZRC 2000.1768), Sungei Benut, Johor, Malaysia; C,
<taxonomicName id="06BCE1C3FFD6BA16FBAAFAEEFAFFFAC0" baseAuthorityName="Targioni Tozzetti" baseAuthorityYear="1877" box="[1143,1394,1372,1394]" class="Malacostraca" family="Sesarmidae" genus="Contusarma" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="946" phylum="Arthropoda" rank="species" species="cheirogonum">
<emphasis id="F3C84652FFD6BA16FBAAFAEEFAFFFAC0" box="[1143,1394,1372,1394]" italics="true" pageId="55" pageNumber="946">Contusarma cheirogonum</emphasis>
</taxonomicName>
, male (15.3 × 14.0 mm) (ZRC 2011.0924), Pulau Tioman, Malaysia; D,
<taxonomicName id="06BCE1C3FFD6BA16FD35FAC9FC64FA3E" baseAuthorityName="Miers" baseAuthorityYear="1880" box="[744,1001,1402,1423]" class="Malacostraca" family="Sesarmidae" genus="Miersarma" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="946" phylum="Arthropoda" rank="species" species="granosimanum">
<emphasis id="F3C84652FFD6BA16FD35FAC9FC64FA3E" box="[744,1001,1402,1423]" italics="true" pageId="55" pageNumber="946">Miersarma granosimanum</emphasis>
</taxonomicName>
, male (22.5 × 19.2 mm) (ZRC 1965.7.29.164), Sedili River, Johor, Malaysia. Arrow indicates shallow depression on somite 5 to accept G1 tip when pleon closed (depression absent in
<taxonomicName id="06BCE1C3FFD6BA16FFACFA07FF51FA78" authorityName="Schubart &amp; Ng" authorityYear="2020" box="[113,220,1460,1481]" class="Malacostraca" family="Sesarmidae" genus="Miersarma" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="946" phylum="Arthropoda" rank="genus">
<emphasis id="F3C84652FFD6BA16FFACFA07FF51FA78" box="[113,220,1460,1481]" italics="true" pageId="55" pageNumber="946">Miersarma</emphasis>
</taxonomicName>
, D).
</paragraph>
</caption>
<paragraph id="C1039A40FFD6BA16FFACF9B3FB8AF9E9" blockId="55.[113,753,1536,2072]" lastBlockId="55.[813,1452,1536,1624]" pageId="55" pageNumber="946">
Specimens reported as “
<emphasis id="F3C84652FFD6BA16FE64F9B3FD2BF9A9" box="[441,678,1536,1560]" italics="true" pageId="55" pageNumber="946">S e s a r m a e d w a r d s i</emphasis>
” and “
<taxonomicName id="06BCE1C3FFD6BA16FFA3F993FE2DF989" baseAuthorityName="De Man" baseAuthorityYear="1887" box="[126,416,1568,1592]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="946" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="F3C84652FFD6BA16FFA3F993FE2DF989" box="[126,416,1568,1592]" italics="true" pageId="55" pageNumber="946">Pseudosesarma edwardsii</emphasis>
</taxonomicName>
” from many parts of
<collectingCountry id="B9ABDAD0FFD6BA16FD42F992FD61F989" box="[671,748,1569,1592]" name="Myanmar" pageId="55" pageNumber="946">Burma</collectingCountry>
,
<collectingCountry id="B9ABDAD0FFD6BA16FFAFF9F3FF26F9E9" box="[114,171,1600,1624]" name="India" pageId="55" pageNumber="946">India</collectingCountry>
, Andamans,
<collectingCountry id="B9ABDAD0FFD6BA16FEEAF9F3FE3BF9E9" box="[311,438,1600,1624]" name="Bangladesh" pageId="55" pageNumber="946">Bangladesh</collectingCountry>
,
<collectingCountry id="B9ABDAD0FFD6BA16FE1FF9F3FDA1F9E9" box="[450,556,1600,1624]" name="Sri Lanka" pageId="55" pageNumber="946">Sri Lanka</collectingCountry>
by
<bibRefCitation id="A52DE7B1FFD6BA16FD8BF9F3FF28F9C9" author="Alcock" firstAuthor="Alcock" page="416" pageId="55" pageNumber="946" pagination="279 - 486" refId="ref67468" refString="Alcock A (1900) Materials for a carcinological fauna of India. No. 6. The Brachyura Catometopa or Grapsoidea. Journal of the Asiatic Society of Bengal, Calcutta, 69 (3): 279 - 486." type="journal article" year="1900">Alcock (1900: 416)</bibRefCitation>
, Mandal &amp; Nandi (1989: 25), Kathirasan (2000: 193), Dev Roy &amp; Nandi (2001: 18), Dev Roy &amp; Bhadra (2007: 143, pl. 4 fig. 5), Dev Roy (2008: 131), Paul et al. (2012: 193), Holmes et al. (2014: 160) and
<bibRefCitation id="A52DE7B1FFD6BA16FE1FF973FD1BF969" author="Shet" box="[450,662,1728,1752]" etAl="et al." firstAuthor="Shet" page="8" pageId="55" pageNumber="946" refId="ref77035" refString="Shet GN, Subash Chandran MD &amp; Ramachandra TV (2016) Brachyuran crabs of Aghanashini Estuary, South Indian West Coast, Karnataka. In: Proceedings of the Lake 2016: Conference on Conservation and Sustainable Management of Ecologically Sensitive Regions in Western Ghats, 2016, 17 pp." type="book" year="2016">Shet et al. (2016: 8</bibRefCitation>
, 12, fig. 2) (see
<bibRefCitation id="A52DE7B1FFD6BA16FF1EF953FE4FF949" author="Trivedi" box="[195,450,1760,1784]" etAl="et al." firstAuthor="Trivedi" page="73" pageId="55" pageNumber="946" pagination="1 - 83" refId="ref78238" refString="Trivedi JN, Trivedi DJ, Vachhrajani KD &amp; Ng PKL (2018) An annotated checklist of marine brachyuran crabs (Crustacea: Decapoda: Brachyura) of India. Zootaxa, 4502 (1): 1 - 83." type="journal article" year="2018">Trivedi et al., 2018: 73</bibRefCitation>
); as well as Java, Sulawesi, and New
<collectingCountry id="B9ABDAD0FFD6BA16FF06F8B3FEA5F8A9" box="[219,296,1792,1816]" name="Guinea-Bissau" pageId="55" pageNumber="946">Guinea</collectingCountry>
by
<bibRefCitation id="A52DE7B1FFD6BA16FE89F8B3FD96F8A9" author="Tesch" box="[340,539,1792,1816]" firstAuthor="Tesch" page="147" pageId="55" pageNumber="946" pagination="127 - 260" refId="ref78063" refString="Tesch JJ (1917) Synopsis of the genera Sesarma, Metasesarma, Sarmatium and Clistocoeloma, with a key to the determination of the IndoPacific species. Zoologische Mededelingen, 3: 127 - 260, pls. 15 - 17." type="journal article" year="1917">Tesch (1917: 147)</bibRefCitation>
probably belong to other species. The material from
<collectingCountry id="B9ABDAD0FFD6BA16FE17F893FDBFF889" box="[458,562,1824,1848]" name="Sri Lanka" pageId="55" pageNumber="946">Sri Lanka</collectingCountry>
by
<bibRefCitation id="A52DE7B1FFD6BA16FD8AF893FD7DF889" author="Alcock" box="[599,752,1824,1848]" firstAuthor="Alcock" pageId="55" pageNumber="946" pagination="279 - 486" refId="ref67468" refString="Alcock A (1900) Materials for a carcinological fauna of India. No. 6. The Brachyura Catometopa or Grapsoidea. Journal of the Asiatic Society of Bengal, Calcutta, 69 (3): 279 - 486." type="journal article" year="1900">Alcock (1900)</bibRefCitation>
was referred to
<taxonomicName id="06BCE1C3FFD6BA16FEC7F8F2FE2AF8E9" authorityName="Ng &amp; Schubart" authorityYear="2017" box="[282,423,1857,1880]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="946" phylum="Arthropoda" rank="species" species="anteactum">
<emphasis id="F3C84652FFD6BA16FEC7F8F2FE2AF8E9" box="[282,423,1857,1880]" italics="true" pageId="55" pageNumber="946">P. anteactum</emphasis>
</taxonomicName>
; while specimens from
<collectingRegion id="037854A2FFD6BA16FD74F8F3FD7DF8E9" box="[681,752,1856,1880]" country="India" name="Kerala" pageId="55" pageNumber="946">Kerala</collectingRegion>
in West
<collectingCountry id="B9ABDAD0FFD6BA16FF11F8D3FE8EF8C9" box="[204,259,1888,1912]" name="India" pageId="55" pageNumber="946">India</collectingCountry>
probably belong to
<taxonomicName id="06BCE1C3FFD6BA16FE02F8D2FDDBF8C9" authorityName="Ng, Rani &amp; Nandan" authorityYear="2017" box="[479,598,1888,1912]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="946" phylum="Arthropoda" rank="species" species="glabrum">
<emphasis id="F3C84652FFD6BA16FE02F8D2FDDBF8C9" box="[479,598,1888,1912]" italics="true" pageId="55" pageNumber="946">P. glabrum</emphasis>
</taxonomicName>
; as does most western Indian material. Specimens from northeastern
<collectingCountry id="B9ABDAD0FFD6BA16FD64F833FD7DF829" box="[697,752,1920,1944]" name="India" pageId="55" pageNumber="946">India</collectingCountry>
belong to
<taxonomicName id="06BCE1C3FFD6BA16FF39F812FED0F809" authorityName="Ng" authorityYear="2018" box="[228,349,1952,1976]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="946" phylum="Arthropoda" rank="species" species="brehieri">
<emphasis id="F3C84652FFD6BA16FF39F812FED0F809" box="[228,349,1952,1976]" italics="true" pageId="55" pageNumber="946">P. brehieri</emphasis>
</taxonomicName>
and we have some specimens from
<collectingRegion id="037854A2FFD6BA16FFAFF872FE8CF869" box="[114,257,1984,2008]" country="India" name="West Bengal" pageId="55" pageNumber="946">West Bengal</collectingRegion>
that confirm this (
<figureCitation id="598786C5FFD6BA16FE07F873FDDAF869" box="[474,599,1984,2008]" captionStart="Fig" captionStartId="64.[136,171,1926,1948]" captionTargetBox="[138,1476,190,1907]" captionTargetId="figure-8@64.[124,1486,183,1911]" captionText="Fig. 37. AG, Pseudosesarma brehieri, paratype female (14.9 × 13.4 mm) (ZRC 2016.0594), Myanmar; F, G, P. brehieri, male (15.0 × 13.6 mm) (ZRC 2013.0209), West Bengal, India; H, I, P. brehieri, male (14.5 × 13.0 mm) (ZRC 2013.0209), West Bengal, India; JO, P. boulengeri, lectotype male (26.9 × 23.5 mm) (NHM 1919.11.14.1), Basra, Iraq. A, K, male pleon; B, F, L, left G1 (ventral view, denuded); D, G, M, left G1 (dorsal view, denuded); E, left G2; H, O, left distal part of G1 (dorsal view, denuded); I, N, left distal part of G1 (ventral view, denuded); J, anterior thoracic sternites 14. Scales: A = 4.0 mm; BE, N, O = 1.0 mm; FI = 0.5 mm; J, K = 5.0 mm; L, M = 2.0 mm. AE, after Ng (2018: fig. 7EI)." figureDoi="http://doi.org/10.5281/zenodo.6519019" httpUri="https://zenodo.org/record/6519019/files/figure.png" pageId="55" pageNumber="946">Fig. 37FI</figureCitation>
). The precise identities of all these records can only be determined after re-examining the specimens.
<bibRefCitation id="A52DE7B1FFD6BA16FE70F7B3FD28F7A9" author="Ortmann's" box="[429,677,2048,2072]" firstAuthor="Ortmann's" page="721" pageId="55" pageNumber="946" pagination="683 - 772" refId="ref74488" refString="Ortmann AE (1894) Abtheilung: Brachyura (Brachyura genuina Boas), III. Unterabtheilung: Cancroidea. 2. Section: Cancrinea, 2. Gruppe: Catametopa. Die Decapoden-Krebse des Strassburger Museums mit besonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur Zeit im Strassburger Museum aufbewahrten Formen. Theil VIII [= Part 8]. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 7 (5): 683 - 772, pl. 23." type="journal article" year="1894">Ortmanns (1894: 721)</bibRefCitation>
record of “
<taxonomicName id="06BCE1C3FFD6BA16FC8EF9B3FB9EF9A9" authorityName="Ortmann" authorityYear="1894" box="[851,1043,1536,1560]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="946" phylum="Arthropoda" rank="species" species="edwardsi">
<emphasis id="F3C84652FFD6BA16FC8EF9B3FB9EF9A9" box="[851,1043,1536,1560]" italics="true" pageId="55" pageNumber="946">Sesarma edwardsi</emphasis>
</taxonomicName>
” from
<collectingCountry id="B9ABDAD0FFD6BA16FB81F9B3FB31F9A9" box="[1116,1212,1536,1560]" name="Australia" pageId="55" pageNumber="946">Australia</collectingCountry>
is probably
<taxonomicName id="06BCE1C3FFD6BA16FAE6F9B3FBCDF989" authority="(De Man, 1889)" baseAuthorityName="De Man" baseAuthorityYear="1889" class="Malacostraca" family="Sesarmidae" genus="Bresedium" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="946" phylum="Arthropoda" rank="species" species="brevipes">
<emphasis id="F3C84652FFD6BA16FAE6F9B3FC0AF989" italics="true" pageId="55" pageNumber="946">Bresedium brevipes</emphasis>
(
<bibRefCitation id="A52DE7B1FFD6BA16FC4AF993FBB4F989" author="De Man" box="[919,1081,1568,1592]" firstAuthor="De Man" pageId="55" pageNumber="946" pagination="409 - 452" refId="ref72218" refString="Man JG De (1889) Ueber einige neue oder seltene indopacifische Brachyuren. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Thiere, Jena, 4 (2): 409 - 452, pls. 9, 10." type="book chapter" year="1889">De Man, 1889</bibRefCitation>
)
</taxonomicName>
(see also
<bibRefCitation id="A52DE7B1FFD6BA16FB6CF993FAB2F989" author="Laurie" box="[1201,1343,1568,1592]" firstAuthor="Laurie" pageId="55" pageNumber="946" pagination="1" refId="ref71318" refString="Laurie RD (1906) Report on the Brachyura collected by Professor Herdman, at Ceylon, in 1902. In: Herdman WA (ed.) Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar with Supplementary Reports Upon the Marine Biology of Ceylon by Other Naturalists, 5 (Supplemental Reports No. 40): 349 - 432, figs. 1 - 12, pls. 1, 2." type="journal article" year="1906">Laurie, 1906</bibRefCitation>
;
<bibRefCitation id="A52DE7B1FFD6BA16FA91F993FCE8F9E9" author="McNeill" firstAuthor="McNeill" pageId="55" pageNumber="946" pagination="1 - 98" refId="ref72612" refString="McNeill FA (1968) Crustacea, Decapoda &amp; Stomatopoda. Great Barrier Reef Expedition 1928 - 29. Scientific Reports, 7 (1): 1 - 98" type="journal article" year="1968">McNeill, 1968</bibRefCitation>
;
<bibRefCitation id="A52DE7B1FFD6BA16FCAFF9F3FC76F9E9" author="Davie" box="[882,1019,1600,1624]" firstAuthor="Davie" pageId="55" pageNumber="946" refId="ref68823" refString="Davie PJF (2002) Crustacea: Malacostraca: Eucarida (Part 2: Anomura, Brachyura). In: Wells A &amp; Houston WWK (eds.) Zoological Catalogue of Australia. Vol. 19.3 b. CSIRO Publishing, Melbourne, 641 pp." type="book" year="2002">Davie, 2002</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFD6BA16FCF0F933FBEAF8C9" pageId="55" pageNumber="946" type="biology_ecology">
<paragraph id="C1039A40FFD6BA16FCF0F933FBEAF8C9" blockId="55.[813,1453,1664,1912]" pageId="55" pageNumber="946">
<emphasis id="F3C84652FFD6BA16FCF0F933FC08F929" bold="true" box="[813,901,1664,1688]" pageId="55" pageNumber="946">Biology</emphasis>
.
<taxonomicName id="06BCE1C3FFD6BA16FC4AF933FB34F929" baseAuthorityName="De Man" baseAuthorityYear="1887" box="[919,1209,1664,1688]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="946" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="F3C84652FFD6BA16FC4AF933FB34F929" box="[919,1209,1664,1688]" italics="true" pageId="55" pageNumber="946">Pseudosesarma edwardsii</emphasis>
</taxonomicName>
is typically found in well-forested coastal freshwater or brackish water habitats. They are semiterrestrial and live on the banks of streams as well as swamps, and are often found under rocks and vegetation. They are sometimes found several hundred metres from the sea, even at the base of waterfalls. They have small eggs and it is clear their larval development remains tied to the open sea.
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFD6BA16FCF0F813FA21F849" pageId="55" pageNumber="946" type="distribution">
<paragraph id="C1039A40FFD6BA16FCF0F813FA21F849" blockId="55.[813,1453,1952,2041]" pageId="55" pageNumber="946">
<emphasis id="F3C84652FFD6BA16FCF0F813FC4BF809" bold="true" box="[813,966,1952,1976]" pageId="55" pageNumber="946">Distribution</emphasis>
. Mergui Archipelago and Andamans to Peninsular
<collectingCountry id="B9ABDAD0FFD6BA16FC77F873FB80F869" box="[938,1037,1984,2008]" name="Malaysia" pageId="55" pageNumber="946">Malaysia</collectingCountry>
and
<collectingCountry id="B9ABDAD0FFD6BA16FB94F873FB34F869" box="[1097,1209,1984,2008]" name="Singapore" pageId="55" pageNumber="946">Singapore</collectingCountry>
, possibly to Java and Sulawesi (
<bibRefCitation id="A52DE7B1FFD6BA16FC4AF853FBA3F849" author="De Man" box="[919,1070,2016,2040]" firstAuthor="De Man" pageId="55" pageNumber="946" pagination="639 - 722" refId="ref72089" refString="Man JG De (1887) Ubersicht der Indopacifischen Arten der Gattung Sesarma Say nebst einer Kritik der von W. Hess und E. Nauck in den Jahren 1865 und 1880 beschrieben Decapoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 2: 639 - 722." type="journal article" year="1887">De Man, 1887</bibRefCitation>
;
<bibRefCitation id="A52DE7B1FFD6BA16FBE5F853FB35F849" author="Tesch" box="[1080,1208,2016,2040]" firstAuthor="Tesch" pageId="55" pageNumber="946" pagination="127 - 260" refId="ref78063" refString="Tesch JJ (1917) Synopsis of the genera Sesarma, Metasesarma, Sarmatium and Clistocoeloma, with a key to the determination of the IndoPacific species. Zoologische Mededelingen, 3: 127 - 260, pls. 15 - 17." type="journal article" year="1917">Tesch, 1917</bibRefCitation>
;
<bibRefCitation id="A52DE7B1FFD6BA16FB1FF852FA2CF849" author="Ng &amp; Schubart" box="[1218,1441,2016,2041]" firstAuthor="Ng" pageId="55" pageNumber="946" pagination="655 - 669" refId="ref74064" refString="Ng PKL &amp; Schubart CD (2017) On the taxonomy of Pseudosesarma edwardsii (De Man, 1887) and P. crassimanum (De Man, 1887) (Crustacea: Decapoda: Brachyura: Sesarmidae), with description of a new species from Sri Lanka. Raffles Bulletin of Zoology, 65: 655 - 669." type="journal article" year="2017">Ng &amp; Schubart, 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>