197 lines
20 KiB
XML
197 lines
20 KiB
XML
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<mods:title id="A97868F7D5E0C7CD768B97F6EED5CBBB">A redescription of TraCheloSaUrUS fiSCheri from the Buntsandstein (Middle Triassic) of Bernburg, Germany: the first European DinoCephaloSaUrUS-like marine reptile and its systematic implications for long-necked early</mods:title>
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<mods:namePart id="1AE9D0EAE9C18E5CDC4901D372FC28CF">Spiekman, Stephan N. F.</mods:namePart>
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<mods:namePart id="302344608584311AC2E4FBCFDD7EAA25">Ezcurra, Martín D.</mods:namePart>
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<mods:namePart id="B8382230B8CAB41404EEDBFDF5B2E1B0">Wang, Wei</mods:namePart>
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<mods:namePart id="9D403065A1AD09019F5A4D8C68E56833">Mujal, Eudald</mods:namePart>
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<mods:namePart id="E62EAB11052D4B686AD08DF4CE9015B3">Buchwitz, Michael</mods:namePart>
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<treatment id="FF5D3B314D36C57840AB92D9FB78FEA7" LSID="urn:lsid:plazi:treatment:FF5D3B314D36C57840AB92D9FB78FEA7" httpUri="http://treatment.plazi.org/id/FF5D3B314D36C57840AB92D9FB78FEA7" lastPageNumber="8" pageId="7" pageNumber="8">
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<paragraph id="774B8A274D36C57840AB92D9FDCBFE2B" blockId="7.[151,774,1224,1285]" pageId="7" pageNumber="8">
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<taxonomicName id="B0F4F1A44D36C57840AB92D9FE13FFCB" authority="Abel, 1919" authorityName="Abel" authorityYear="1919" box="[151,487,1228,1252]" family="Trachelosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Chordata" rank="family">
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<emphasis id="458056354D36C57840AB92D9FE13FFCB" bold="true" box="[151,487,1228,1252]" pageId="7" pageNumber="8">
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Trachelosauridae
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<bibRefCitation id="1365F7D64D36C578415592D9FE13FFCB" author="Abel, O." box="[361,487,1228,1252]" pageId="7" pageNumber="8" refId="ref22127" refString="Abel, O. (1919). Die Stamme der Wirbeltiere. W de Gruyter." type="book" year="1919">Abel, 1919</bibRefCitation>
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</emphasis>
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</taxonomicName>
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<superScript id="8081276F4D36C57841DB92DDFE04FFF6" attach="none" box="[487,496,1224,1241]" fontSize="7" pageId="7" pageNumber="8">1</superScript>
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<emphasis id="458056354D36C57841CC92D9FDCBFE2B" bold="true" pageId="7" pageNumber="8">
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(= Dinocephalosauridae
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<bibRefCitation id="1365F7D64D36C57840AB92F9FDC2FE2B" author="Spiekman, S. N. F. & Fraser, N. C. & Scheyer, T. M." box="[151,566,1260,1285]" pageId="7" pageNumber="8" refId="ref27999" refString="Spiekman, S. N. F., Fraser, N. C., & Scheyer, T. M. (2021 b). A new phylogenetic hypothesis of Tanystropheidae (Diapsida, Archosauromorpha) and other " protorosaurs ", and its implications for the early evolution of stem archosaurs. PeerJ, 9, e 11143. https: // doi. org / 10.7717 / peerj. 11143" type="journal volume" year="2021">Spiekman, Fraser and Scheyer, 2021</bibRefCitation>
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)
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="3FEED9AC4D36C578437E9745FBBBF888" pageId="7" pageNumber="8" type="description">
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<paragraph id="774B8A274D36C578437E9745FBBDFA87" blockId="7.[813,1437,336,1416]" pageId="7" pageNumber="8">
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<emphasis id="458056354D36C578437E9745FB4EFA47" bold="true" box="[834,1210,336,360]" pageId="7" pageNumber="8">Phylocode registration number.</emphasis>
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<taxonomicName id="B0F4F1A44D36C57844F99745FA74FA47" box="[1221,1408,336,360]" family="Trachelosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Chordata" rank="family">Trachelosauridae</taxonomicName>
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is identified in the international clade names repository as registration number 1028.
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</paragraph>
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<paragraph id="774B8A274D36C578437E97A5FA69F987" blockId="7.[813,1437,336,1416]" pageId="7" pageNumber="8">
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||
<emphasis id="458056354D36C578437E97A5FBBEFAE7" bold="true" box="[834,1098,432,456]" pageId="7" pageNumber="8">Phylogenetic definition.</emphasis>
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The definition of
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||
<taxonomicName id="B0F4F1A44D36C578453A97A5FCA6FAC7" family="Trachelosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Chordata" rank="family">Trachelosauridae</taxonomicName>
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is modified from the definition for Dinocephalosauridae (
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<bibRefCitation id="1365F7D64D36C578430F97E5FBD4F927" author="Spiekman, S. N. F. & Fraser, N. C. & Scheyer, T. M." box="[819,1056,496,520]" pageId="7" pageNumber="8" refId="ref27999" refString="Spiekman, S. N. F., Fraser, N. C., & Scheyer, T. M. (2021 b). A new phylogenetic hypothesis of Tanystropheidae (Diapsida, Archosauromorpha) and other " protorosaurs ", and its implications for the early evolution of stem archosaurs. PeerJ, 9, e 11143. https: // doi. org / 10.7717 / peerj. 11143" type="journal volume" year="2021">Spiekman et al., 2021b</bibRefCitation>
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) as follows: the most inclusive clade containing
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<taxonomicName id="B0F4F1A44D36C578439B9405FAD6F907" authority="Broili, 1918" authorityName="Broili & Fischer" authorityYear="1918" box="[935,1314,528,552]" family="Trachelosauridae" genus="Trachelosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="fischeri">
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<emphasis id="458056354D36C578439B9405FB6DF907" box="[935,1177,528,552]" italics="true" pageId="7" pageNumber="8">Trachelosaurus fischeri</emphasis>
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||
<bibRefCitation id="1365F7D64D36C57844999405FAD6F907" author="Broili, F. & Fischer, E." box="[1189,1314,528,552]" pageId="7" pageNumber="8" pagination="359 - 414" refId="ref22657" refString="Broili, F., & Fischer, E. (1918). Trachelosaurus fischeri nov. gen. nov. sp. Ein neuer Saurier aus dem Buntsandstein von Bernburg. Jahrbuch Der Koniglich Preussischen Geologischen Landesanstalt Zu Berlin, 37 (1), 359 - 414." type="journal article" year="1918">Broili, 1918</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="B0F4F1A44D36C578455C9405FB74F967" authority="Li, 2003" authorityName=", Gracilicollum" authorityYear="2003" family="Tanystropheidae" genus="Dinocephalosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="orientalis">
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<emphasis id="458056354D36C578455C9405FBD3F967" italics="true" pageId="7" pageNumber="8">Dinocephalosaurus orientalis</emphasis>
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<bibRefCitation id="1365F7D64D36C57844109425FB74F967" author="Li, C." box="[1068,1152,560,584]" pageId="7" pageNumber="8" pagination="419 - 423" refId="ref25564" refString="Li, C. (2003). First record of protorosaurid reptile (Order Protorosauria) from the Middle Triassic of China. Acta Geologica Sinica, 77 (4), 419 - 423. https: // doi. org / 10.1111 / j. 1755 - 6724.2003. tb 00122. x" type="journal article" year="2003">Li, 2003</bibRefCitation>
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</taxonomicName>
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but not
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<taxonomicName id="B0F4F1A44D36C57844E49425FBB7F947" authority="(Bassani, 1886)" baseAuthorityName="Bassani" baseAuthorityYear="1886" family="Tanystropheidae" genus="Tanystropheus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="longobardicus">
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<emphasis id="458056354D36C57844E49425FC68F947" italics="true" pageId="7" pageNumber="8">Tanystropheus longobardicus</emphasis>
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(
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<bibRefCitation id="1365F7D64D36C57843979445FBCEF947" author="Bassani, F." box="[939,1082,592,616]" pageId="7" pageNumber="8" pagination="15 - 72" refId="ref22337" refString="Bassani, F. (1886). Sui fossili e sull'eta degli schisti bituminosi triasici di Besano in Lombardia. Atti Societa Italiana di scienze naturali, 29, 15 - 72." type="journal article" year="1886">Bassani, 1886</bibRefCitation>
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)
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</taxonomicName>
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,
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<taxonomicName id="B0F4F1A44D36C578446C9444FCABF9A7" authority="Nopcsa, 1931" authorityName="Nopcsa" authorityYear="1931" class="Insecta" family="Cerambycidae" genus="Macrocnemus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bassanii">
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<emphasis id="458056354D36C578446C9444FACBF947" box="[1104,1343,592,616]" italics="true" pageId="7" pageNumber="8">Macrocnemus bassanii</emphasis>
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<bibRefCitation id="1365F7D64D36C578457B9445FCABF9A7" author="Nopcsa, F. B." pageId="7" pageNumber="8" pagination="252 - 255" refId="ref26348" refString="Nopcsa, F. B. (1931). Notizen Uber Macrochemus Bassanii nov. gen. et spec. Centralblatt fur Mineralogie, Geologie und Palaontologie B, 7, 252 - 255." type="journal article" year="1931">Nopcsa, 1931</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="B0F4F1A44D36C57843509464FAF0F9A7" authority="von Meyer, 1832" authorityName="von Meyer" authorityYear="1832" box="[876,1284,624,648]" family="Protorosauridae" genus="Protorosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="speneri">
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<emphasis id="458056354D36C57843509464FBBAF9A7" box="[876,1102,625,648]" italics="true" pageId="7" pageNumber="8">Protorosaurus speneri</emphasis>
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von
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<bibRefCitation id="1365F7D64D36C57844B89465FAF0F9A7" author="Meyer, H." box="[1156,1284,624,648]" pageId="7" pageNumber="8" refId="ref25895" refString="Meyer, H. v. (1832). Palaeologica zur Geschichte der Erde und ihrer Geschopfe. S. Schmerber." type="book" year="1832">Meyer, 1832</bibRefCitation>
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</taxonomicName>
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||
, or
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||
<taxonomicName id="B0F4F1A44D36C578450C9465FBD3F987" authority="Parrington, 1935" authorityName="Parrington" authorityYear="1935" family="Protorosauridae" genus="Prolacerta" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="broomi">
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<emphasis id="458056354D36C578450C9465FC83F987" italics="true" pageId="7" pageNumber="8">Prolacerta broomi</emphasis>
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||
<bibRefCitation id="1365F7D64D36C57843479485FBD3F987" author="Parrington, F. R." box="[891,1063,656,680]" pageId="7" pageNumber="8" pagination="197 - 205" refId="ref26619" refString="Parrington, F. R. (1935). On Prolacerta broomi, gen. et sp. n. and the origin of lizards. Annals and Magazine of Natural History Series 10, 16 (92), 197 - 205. https: // doi. org / 10.1080 / 00222933508655037" type="journal article" year="1935">Parrington, 1935</bibRefCitation>
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||
</taxonomicName>
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. This is a maximum clade definition.
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||
</paragraph>
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<paragraph id="774B8A274D36C578437E94A5FC04F9C8" blockId="7.[813,1437,336,1416]" pageId="7" pageNumber="8">
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||
<emphasis id="458056354D36C578437E94A5FBCEF9E7" bold="true" box="[834,1082,688,712]" pageId="7" pageNumber="8">Reference phylogeny.</emphasis>
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Phylogenetic hypotheses recovered in this paper.
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</paragraph>
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||
<paragraph id="774B8A274D36C578437E94E5FBBBF888" blockId="7.[813,1437,336,1416]" pageId="7" pageNumber="8">
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||
<emphasis id="458056354D36C578437E94E5FC14F827" bold="true" box="[834,992,752,776]" pageId="7" pageNumber="8">Composition.</emphasis>
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The composition is based on our reference phylogenies.
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<taxonomicName id="B0F4F1A44D36C578442A9505FB25F807" box="[1046,1233,784,808]" family="Trachelosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Chordata" rank="family">Trachelosauridae</taxonomicName>
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includes the following nominal species:
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<taxonomicName id="B0F4F1A44D36C578446C9525FA63F867" authorityName=", Gracilicollum" authorityYear="2003" box="[1104,1431,816,840]" family="Tanystropheidae" genus="Dinocephalosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="orientalis">
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<emphasis id="458056354D36C578446C9525FA63F867" box="[1104,1431,816,840]" italics="true" pageId="7" pageNumber="8">Dinocephalosaurus orientalis</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="B0F4F1A44D36C57843119545FB34F848" authority=", Austronaga" authorityName="Austronaga" box="[813,1216,848,872]" family="Trachelosauridae" genus="Trachelosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="fischeri">
|
||
<emphasis id="458056354D36C57843119545FBDAF847" box="[813,1070,848,872]" italics="true" pageId="7" pageNumber="8">Trachelosaurus fischeri</emphasis>
|
||
,
|
||
<emphasis id="458056354D36C578447E9545FB34F848" box="[1090,1216,848,871]" italics="true" pageId="7" pageNumber="8">Austronaga</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="458056354D36C57844F19544FAE9F847" box="[1229,1309,849,872]" italics="true" pageId="7" pageNumber="8">minuta</emphasis>
|
||
,
|
||
<taxonomicName id="B0F4F1A44D36C578450D9545FC5AF8A7" genus="Pectodens" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="zhenyuensis">
|
||
<emphasis id="458056354D36C578450D9545FC5AF8A7" italics="true" pageId="7" pageNumber="8">Pectodens zhenyuensis</emphasis>
|
||
</taxonomicName>
|
||
, and ambiguously
|
||
<emphasis id="458056354D36C57844B89565FA90F8A7" box="[1156,1380,880,904]" italics="true" pageId="7" pageNumber="8">Gracilicollum latens</emphasis>
|
||
, and
|
||
<taxonomicName id="B0F4F1A44D36C57843119584FBBDF888" box="[813,1097,912,936]" genus="Fuyuansaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="acutirostris">
|
||
<emphasis id="458056354D36C57843119584FBBDF888" box="[813,1097,912,936]" italics="true" pageId="7" pageNumber="8">Fuyuansaurus acutirostris</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3FEED9AC4D36C578437E95A5FB78FEA7" pageId="7" pageNumber="8" type="diagnosis">
|
||
<paragraph id="774B8A274D36C578437E95A5FB78FEA7" blockId="7.[813,1437,336,1416]" pageId="7" pageNumber="8">
|
||
<emphasis id="458056354D36C578437E95A5FC4EF8E7" bold="true" box="[834,954,944,968]" pageId="7" pageNumber="8">Diagnosis.</emphasis>
|
||
<taxonomicName id="B0F4F1A44D36C57843F895A5FB8FF8E7" box="[964,1147,944,968]" family="Trachelosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Chordata" rank="family">Trachelosauridae</taxonomicName>
|
||
is a clade that is defined among other archosauromorphs by possessing the following unique combination of character states: external naris positioned far from the anterior end of the premaxilla, resulting in an anteroposteriorly deep base of the prenarial process; jugal without posterior process; cervical column composed of ten or more vertebrae (also present in some deeply nested tanystropheids); axis with postzygapophysis confluent with and not protruding posteriorly from the base of the neural spine in lateral view (also present in
|
||
<taxonomicName id="B0F4F1A44D36C578431192E5FC0AFE28" box="[813,1022,1264,1288]" family="Tanystropheidae" genus="Tanystropheus" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="458056354D36C578431192E5FC33FE27" box="[813,967,1264,1288]" italics="true" pageId="7" pageNumber="8">Tanystropheus</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B0F4F1A44D36C578440192E5FABAFE27" authorityName="Fraser & Rieppel" authorityYear="2006" box="[1085,1358,1264,1288]" family="Tanystropheidae" genus="Amotosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="rotfeldensis">
|
||
<emphasis id="458056354D36C578440192E5FABAFE27" box="[1085,1358,1264,1288]" italics="true" pageId="7" pageNumber="8">Amotosaurus rotfeldensis</emphasis>
|
||
</taxonomicName>
|
||
); mid-dorsal vertebrae with very wide and ‘wing-like’ transverse processes; presence of holocephalous anterior dorsal ribs (also present in some tanystropheids); metatarsal V without a hook-shaped proximal end.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |