144 lines
14 KiB
XML
144 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.53.4955" ID-PMC="PMC4547023" ID-Pensoft-Pub="1314-2003-53-39" ID-Pensoft-UUID="824BD167D1535A15FC27FF98FFD3A404" ID-PubMed="26312040" ID-Zenodo-Dep="576284" ModsDocID="1314-2003-53-39" checkinTime="1451251414861" checkinUser="pensoft" docAuthor="Luong, Thien Tam, Hovenkamp, Peter H. & Sosef, Marc S. M." docDate="2015" docId="049874F136265294995E4C187AA60870" docLanguage="en" docName="PhytoKeys 53: 39-71" docOrigin="PhytoKeys 53" docPubDate="2015-07-21" docSource="http://dx.doi.org/10.3897/phytokeys.53.4955" docTitle="Orthiopteris samoensis P. H. Hovenkamp & T. T. Luong 2015, spec. nov." docType="treatment" docVersion="3" id="824BD167D1535A15FC27FF98FFD3A404" lastPageNumber="57" masterDocId="824BD167D1535A15FC27FF98FFD3A404" masterDocTitle="Revision of the fern genus Orthiopteris (Saccolomataceae) in Malesia and adjacent regions" masterLastPageNumber="71" masterPageNumber="39" pageNumber="56" updateTime="1668141422086" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the fern genus Orthiopteris (Saccolomataceae) in Malesia and adjacent regions</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Luong, Thien Tam</mods:namePart>
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<mods:affiliation>Department of Ecology - Evolutionary Biology, Viet Nam National University Ho Chi Minh city (VNUHCM) - University of Science. 227 Nguyen Van Cu, Ho Chi Minh City, Vietnam & Naturalis Biodiversity Center, section Botany. PO Box 9517, 2300 RA Leiden, The Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hovenkamp, Peter H.</mods:namePart>
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<mods:affiliation>Naturalis Biodiversity Center, section Botany. PO Box 9517, 2300 RA Leiden, The Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sosef, Marc S. M.</mods:namePart>
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<mods:affiliation>Botanic Garden Meise, Nieuwelaan 38, 1860 Meise, Belgium</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2015-07-21</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>53</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>39</mods:start>
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<mods:end>71</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.53.4955</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.53.4955</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-53-39</mods:identifier>
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<mods:identifier type="Pensoft-UUID">824BD167D1535A15FC27FF98FFD3A404</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576284</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182227826" LSID="urn:lsid:plazi:treatment:049874F136265294995E4C187AA60870" httpUri="http://treatment.plazi.org/id/049874F136265294995E4C187AA60870" lastPageId="18" lastPageNumber="57" pageId="17" pageNumber="56">
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<subSubSection pageId="17" pageNumber="56" type="nomenclature">
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<paragraph pageId="17" pageNumber="56">
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6.
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<taxonomicName LSID="049874F1-3626-5294-995E-4C187AA60870" authority="P. H. Hovenkamp & T. T. Luong" authorityName="P. H. Hovenkamp & T. T. Luong" authorityYear="2015" class="Polypodiopsida" family="Saccolomataceae" genus="Orthiopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Orthiopteris samoensis" order="Polypodiales" pageId="17" pageNumber="56" phylum="Tracheophyta" rank="species" species="samoensis" status="spec. nov.">Orthiopteris samoensis P.H.Hovenkamp & T.T.Luong</taxonomicName>
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<taxonomicNameLabel pageId="17" pageNumber="56">spec. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ultimate segments. Arrows show distance from base of the shallowest sinuses to nearest costule / vein. a Orthiopteris ferulacea, Smith 8932 (K) b Orthiopteris cicutarioides, Carr 13257 (K) c Orthiopteris samoensis, Reinecke 97 a (L) d Orthiopteris trichophylla, Brass 12239 (L) e Orthiopteris campylura var. cautada, Carr 12518 (L) f Orthiopteris campylura var. laxa Mueller s. n. (L) g Orthiopteris campylura var. kingii, Brass 12941 (MICH) h Orthiopteris henriettae, Lam & Meeuse 5892 (L) i Orthiopteris campylura var. insularis Braithwaite 4110 (L). All scale bars 1 mm." figureDoi="10.3897/phytokeys.53.4955.figure2" httpUri="https://binary.pensoft.net/fig/47604" pageId="17" pageNumber="56">Figs 2c</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Sori of Orthiopteris seen from abaxial side. a, b Orthiopteris campylura var. campylura (Cuming 119, MICH) c Orthiopteris campylura var. kingii (Brass 12941, MICH) d Orthiopteris trichophylla (Brass 12239, L) e Orthiopteris tenuis (Smith 9154, L) f Orthiopteris tenuis (Smith 7274, L) g Orthiopteris campylura var. caudata (Carr 12518, L) h Orthiopteris cicutarioides (Carr 13257, K) i, j Orthiopteris campylura var. insularis (Braithwaite 4110, L) k Orthiopteris tenuis (small segment) (Degener 14648, L) l Orthiopteris ferulacea (Smith 8932, K) m, n Orthiopteris firma (Schlechter 14890, L) o, p Orthiopteris campylura var. laxa (Mueller s. n., L) q, r Orthiopteris henriettae (Lam & Meeuse 5892, L) s Orthiopteris henriettae. t Orthiopteris samoensis (Reinecke 97 a, L) u, v Orthiopteris samoensis (Vaupel 312, L). All scale bars 1 mm, oi = outer indusium, ii = inner indusium." figureDoi="10.3897/phytokeys.53.4955.figure3" httpUri="https://binary.pensoft.net/fig/47605" pageId="17" pageNumber="56">, 3t</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Sori of Orthiopteris seen from abaxial side. a, b Orthiopteris campylura var. campylura (Cuming 119, MICH) c Orthiopteris campylura var. kingii (Brass 12941, MICH) d Orthiopteris trichophylla (Brass 12239, L) e Orthiopteris tenuis (Smith 9154, L) f Orthiopteris tenuis (Smith 7274, L) g Orthiopteris campylura var. caudata (Carr 12518, L) h Orthiopteris cicutarioides (Carr 13257, K) i, j Orthiopteris campylura var. insularis (Braithwaite 4110, L) k Orthiopteris tenuis (small segment) (Degener 14648, L) l Orthiopteris ferulacea (Smith 8932, K) m, n Orthiopteris firma (Schlechter 14890, L) o, p Orthiopteris campylura var. laxa (Mueller s. n., L) q, r Orthiopteris henriettae (Lam & Meeuse 5892, L) s Orthiopteris henriettae. t Orthiopteris samoensis (Reinecke 97 a, L) u, v Orthiopteris samoensis (Vaupel 312, L). All scale bars 1 mm, oi = outer indusium, ii = inner indusium." figureDoi="10.3897/phytokeys.53.4955.figure3" httpUri="https://binary.pensoft.net/fig/47605" pageId="17" pageNumber="56">, 3u, v</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. a-d Rhizome scales and e-h rachis scales of Orthiopteris. a Orthiopteris cicutarioides (Carr 13257, L) b Orthiopteris ferulacea (Smith 8932) c Orthiopteris firma (Green 1793, K) d Orthiopteris campylura var. kingi (Hoogland & Craven 10162, L) e, f Orthiopteris campylura var. kingi (Larhing 6910, L) g, h Orthiopteris samoensis (Reinecke 97 a, L). Scale bars: 1 mm (a-d), 0.1 mm (e-h)." figureDoi="10.3897/phytokeys.53.4955.figure4" httpUri="https://binary.pensoft.net/fig/47606" pageId="17" pageNumber="56">, 4g, h</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Orthiopteris spores. a, b Orthiopteris firma (Guillaumin & Baumann-Bodenheim 10318, L) c Orthiopteris tenuis (Smith 9154, L) d Orthiopteris campylura var. campylura (Elmer 11717, L) e Orthiopteris campylura var. caudata (Ridsdale 30654, L) f Orthiopteris henriettae (Lam & Meeuse 5892, L) g, h Orthiopteris samoensis (Sledge 1563, L). All scale bars 10 µm." figureDoi="10.3897/phytokeys.53.4955.figure6" httpUri="https://binary.pensoft.net/fig/47608" pageId="17" pageNumber="56">, 6g, h</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="57" type="diagnosis">
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<paragraph pageId="18" pageNumber="57">
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<pageBreakToken pageId="18" pageNumber="57" start="start">Diagnosis</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="18" pageNumber="57">
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Differs from all other
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<taxonomicName class="Polypodiopsida" family="Saccolomataceae" genus="Orthiopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Orthiopteris" order="Polypodiales" pageId="18" pageNumber="57" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="18" pageNumber="57">Orthiopteris</emphasis>
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</taxonomicName>
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species in relatively dense scales on lower surface of the lamina.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="57" type="type">
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<paragraph pageId="18" pageNumber="57">Type.</paragraph>
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<paragraph pageId="18" pageNumber="57">
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SAMOA.
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<normalizedToken originalValue="Savai’i">Savai'i</normalizedToken>
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, 31 December 1905,
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<emphasis italics="true" pageId="18" pageNumber="57">Vaupel 312</emphasis>
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(holo: L, 3 sheets: L0319870, L0319927, L0319847, iso: MO, U).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="57" type="description">
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<paragraph pageId="18" pageNumber="57">Description.</paragraph>
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<paragraph pageId="18" pageNumber="57">
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Rhizome erect, rising at least 50 cm above ground, diameter unknown. Rhizome scales unknown. Fronds 150-200 cm (complete fronds not seen); stipes stout, ca. 70 cm long, 0.5-0.8 cm across (at base), light brown; lamina probably deltoid, quadripinnate, 100-150 cm long, at least 60 cm wide, firm- herbaceous, brownish green when dry, scaly with scattered scales on abaxial surface, denser at base of segments, and on veins and base of rachis; pinnae at 45-50° to rachis, separated or slightly overlapping, stalk 1.6 cm, including stalk up to 30-45
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<normalizedToken originalValue="×">x</normalizedToken>
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15-20 cm, lanceolate; ultimate segments 1.0-2.0
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<normalizedToken originalValue="×">x</normalizedToken>
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0.4-0.8 cm, sessile or very short stalked, narrowly trapezoid, apex obtuse to acute, margin with weak cartilaginous border; shallowly incised to 0.3-1.0 mm from veins; lobes obtuse to acute, veins in lobes form 3-6 forks, yellow green, weakly contrasting with lamina, percurrent. Scales on rachis irregularly triagular, basifixed, 0.4-1.4
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<normalizedToken originalValue="×">x</normalizedToken>
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0.4-0.9 mm. Sori lateral on lobes, symmetric, not reflexed, in one plane with lamina wings, 0.5-0.8
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<normalizedToken originalValue="×">x</normalizedToken>
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0.6-0.75 mm, suborbicular to funnelform, widest at mouth; inner indusium brown to dark brown, firm, slightly shorter or equal to outer indusium, rarely longer, apex with a rounded to truncate, 0.1-0.2 mm long lobe, eroded, ca.
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<normalizedToken originalValue="¼">1/4</normalizedToken>
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length of sorus; outer indusium truncate, shallowly lobed or eroded; sporangia 5-10 per sorus, capsule globose and rounded at apex, gradually narrowed toward base, indurated annulus cells 17-19, clearly unequal; spores in polar view 30-33
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, in lateral view 23-27
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<normalizedToken originalValue="µm">µm</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="57" type="distribution">
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<paragraph pageId="18" pageNumber="57">Distribution.</paragraph>
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<paragraph pageId="18" pageNumber="57">endemic to Samoa (Upolu, Savaii).</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="57" type="ecology">
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<paragraph pageId="18" pageNumber="57">Ecology.</paragraph>
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<paragraph pageId="18" pageNumber="57">Terrestrial, medium wet forest, at 600-900m altitude</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="57" type="etymology">
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<paragraph pageId="18" pageNumber="57">Etymology.</paragraph>
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<paragraph pageId="18" pageNumber="57">The specific epithet refers to the type locality.</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="57" type="discussion">
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<paragraph pageId="18" pageNumber="57">Discussion.</paragraph>
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<paragraph pageId="18" pageNumber="57">
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The key character to separate this newly recognised species from the others are the many scales covering the abaxial surface of lamina.
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<taxonomicName class="Polypodiopsida" family="Saccolomataceae" genus="Orthiopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Orthiopteris campylura" order="Polypodiales" pageId="18" pageNumber="57" phylum="Tracheophyta" rank="species" species="campylura">
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<emphasis italics="true" pageId="18" pageNumber="57">Orthiopteris campylura</emphasis>
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</taxonomicName>
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may have some scales on the lower lamina surface, but these are sparser, smaller and narrower. Apart from the laminal scales,
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<taxonomicName authorityName="P. H. Hovenkamp & T. T. Luong" authorityYear="2015" class="Polypodiopsida" family="Saccolomataceae" genus="Orthiopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Orthiopteris samoensis" order="Polypodiales" pageId="18" pageNumber="57" phylum="Tracheophyta" rank="species" species="samoensis">
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<emphasis italics="true" pageId="18" pageNumber="57">Orthiopteris samoensis</emphasis>
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</taxonomicName>
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most closely resembles
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<taxonomicName authorityName="P. H. Hovenkamp & T. T. Luong" authorityYear="2015" baseAuthorityName="P. H. Hovenkamp & T. T. Luong" baseAuthorityYear="2015" class="Polypodiopsida" family="Saccolomataceae" genus="Orthiopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Orthiopteris campylura subsp. var. var. caudata" order="Polypodiales" pageId="18" pageNumber="57" phylum="Tracheophyta" rank="variety" species="campylura" subSpecies="var." variety="caudata">Orthiopteris campylura var. caudata</taxonomicName>
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in terms of frond and sori shape, but the inner indusia of
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<taxonomicName authorityName="P. H. Hovenkamp & T. T. Luong" authorityYear="2015" class="Polypodiopsida" family="Saccolomataceae" genus="Orthiopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Orthiopteris samoensis" order="Polypodiales" pageId="18" pageNumber="57" phylum="Tracheophyta" rank="species" species="samoensis">
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<emphasis italics="true" pageId="18" pageNumber="57">Orthiopteris samoensis</emphasis>
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</taxonomicName>
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are not lobed.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |