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<document id="0264F64A8EB83E5949AE325D856A2D48" ID-CLB-Dataset="8975" ID-DOI="10.11646/zootaxa.4903.4.3" ID-GBIF-Dataset="0a98a7e5-af6f-40b6-821e-c56567584422" ID-ISSN="1175-5326" ID-Zenodo-Dep="4431309" ID-ZooBank="CF96683E-75B8-4AC4-90A2-20AD8CC2F14B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610349665964" checkinUser="plazi" docAuthor="Carrejo, Nancy, Obando, Ranulfo González, Aldrete, Alfonso Neri García &amp; Mendivil, Julián" docDate="2021" docId="03C49710FFFCFFA2598DFC47B3E6F830" docLanguage="en" docName="zootaxa.4903.4.3.pdf" docOrigin="Zootaxa 4903 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Goja munchiquensis Carrejo &amp; Obando &amp; Aldrete &amp; Mendivil 2021, n. sp." docType="treatment" docVersion="7" lastPageNumber="524" masterDocId="FFFDEF68FFEBFFB5591AFF94B216FFED" masterDocTitle="New species of Goja Navás (Psocodea: Psocoptera: Epipsocidae) from Colombia Mexico and Peru" masterLastPageNumber="541" masterPageNumber="501" pageNumber="524" updateTime="1698924142089" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3EC1EA1A83C127FD7F148AC0ECFE7667">New species of Goja Navás (Psocodea: Psocoptera: Epipsocidae) from Colombia Mexico and Peru</mods:title>
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<mods:namePart id="58A76C86FAD225BFCCFE40495D66F309">Carrejo, Nancy</mods:namePart>
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<mods:affiliation id="89A42B3A239684E483A4F8B85C698E84">Departamento de Biología, Facultad de Ciencias Naturales y Exactas, Universidad del Valle, Santiago de Cali, COLOMBIA. Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México, Apdo. Postal 70 - 153, CdMx, MÉXICO. anga @ ib. unam. mx; https: // orcid. org / 0000 - 0001 - 7214 - 7966 &amp; nancy. carrejo @ correounivalle. edu. co; https: // orcid. org / 0000 - 0002 - 2395 - 6887</mods:affiliation>
<mods:nameIdentifier id="0165758048122B9EB5705FB08184525B" type="email">nancy.carrejo@correounivalle.edu.co</mods:nameIdentifier>
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<mods:namePart id="59E54BF4B95896241518A30199CF145C">Obando, Ranulfo González</mods:namePart>
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<mods:affiliation id="FA82F3527F437496A3F268C7F0D40EC8">Departamento de Biología, Facultad de Ciencias Naturales y Exactas, Universidad del Valle, Santiago de Cali, COLOMBIA. Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México, Apdo. Postal 70 - 153, CdMx, MÉXICO. anga @ ib. unam. mx; https: // orcid. org / 0000 - 0001 - 7214 - 7966 &amp; ranulfo. gonzalez @ correounivalle. edu. co; https: // orcid. org / 0000 - 0003 - 4047 - 8677</mods:affiliation>
<mods:nameIdentifier id="3D44031454E04D2FF792CA8E75C5AE7E" type="email">ranulfo.gonzalez@correounivalle.edu.co</mods:nameIdentifier>
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<mods:namePart id="A80E9499AB5B4B072807C8CF41143F2C">Aldrete, Alfonso Neri García</mods:namePart>
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<mods:namePart id="269150947A9AD5703B631997381CD32D">Mendivil, Julián</mods:namePart>
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<mods:affiliation id="A844B4C5B039F416AD3059D6D4044833">Departamento de Biología, Facultad de Ciencias Naturales y Exactas, Universidad del Valle, Santiago de Cali, COLOMBIA. Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México, Apdo. Postal 70 - 153, CdMx, MÉXICO. anga @ ib. unam. mx; https: // orcid. org / 0000 - 0001 - 7214 - 7966 &amp; chalcidoidea 88 @ gmail. com; https: // orcid. org / 0000 - 0002 - 6339 - 2000</mods:affiliation>
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<mods:title id="D852B2C779CBA5D2B2759ACEEAB7E643">Zootaxa</mods:title>
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<mods:date id="2F25161A0E78DE885C7C987E8F9C13FD">2021</mods:date>
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<mods:number id="B9FA533FD29AA934F58FE0C37FB35CA2">4903</mods:number>
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<treatment id="03C49710FFFCFFA2598DFC47B3E6F830" ID-DOI="http://doi.org/10.5281/zenodo.4431353" ID-GBIF-Taxon="176570437" ID-Zenodo-Dep="4431353" LSID="urn:lsid:plazi:treatment:03C49710FFFCFFA2598DFC47B3E6F830" httpUri="http://treatment.plazi.org/id/03C49710FFFCFFA2598DFC47B3E6F830" lastPageNumber="524" pageId="23" pageNumber="524">
<subSubSection id="C377758DFFFCFFA2598DFC47B034FC00" box="[151,546,978,1005]" pageId="23" pageNumber="524" type="nomenclature">
<paragraph id="8BD22606FFFCFFA2598DFC47B034FC00" blockId="23.[151,546,978,1042]" box="[151,546,978,1005]" pageId="23" pageNumber="524">
<heading id="D09A916AFFFCFFA2598DFC47B034FC00" bold="true" box="[151,546,978,1005]" fontSize="11" level="1" pageId="23" pageNumber="524" reason="1">
<emphasis id="B919FA14FFFCFFA2598DFC47B034FC00" bold="true" box="[151,546,978,1005]" pageId="23" pageNumber="524">
<taxonomicName id="4C6D5D85FFFCFFA2598DFC47B387FC00" ID-CoL="9HZXV" authority="Carrejo &amp; Obando &amp; Aldrete &amp; Mendivil, 2021" authorityName="Carrejo &amp; Obando &amp; Aldrete &amp; Mendivil" authorityYear="2021" box="[151,401,978,1005]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="23" pageNumber="524" phylum="Arthropoda" rank="species" species="munchiquensis" status="sp. nov.">
<emphasis id="B919FA14FFFCFFA2598DFC47B387FC00" bold="true" box="[151,401,978,1005]" italics="true" pageId="23" pageNumber="524">Goja munchiquensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A476FFFFCFFA25883FC47B3CDFC00" box="[409,475,979,1005]" pageId="23" pageNumber="524" rank="species">n. sp.</taxonomicNameLabel>
Male
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C377758DFFFCFFA2598DFC63B331FBFF" box="[151,295,1015,1042]" pageId="23" pageNumber="524" type="description">
<paragraph id="8BD22606FFFCFFA2598DFC63B331FBFF" blockId="23.[151,546,978,1042]" box="[151,295,1015,1042]" pageId="23" pageNumber="524">
(
<figureCitation id="13563A83FFFCFFA25985FC63B336FBFF" box="[159,288,1015,1042]" captionStart="FIGURES 6772" captionStartId="24.[151,264,1789,1813]" captionTargetBox="[167,1423,191,1755]" captionTargetId="figure-22@24.[151,1435,181,1765]" captionTargetPageId="24" captionText="FIGURES 6772. G. munchiquensis n. sp. Male. 67. Forewing. 68. Hindwing. 69. Front view of head. 70. Left paraproct and epiproct. 71. Hypandrium. 72. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4431359" httpUri="https://zenodo.org/record/4431359/files/figure.png" pageId="23" pageNumber="524">Figs 6772</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C377758DFFFCFFA2598DFBAAB699F8C4" pageId="23" pageNumber="524" type="diagnosis">
<paragraph id="8BD22606FFFCFFA2598DFBAAB78AFB04" blockId="23.[151,1437,1086,2013]" pageId="23" pageNumber="524">
<emphasis id="B919FA14FFFCFFA2598DFBAAB304FBB5" bold="true" box="[151,274,1086,1112]" pageId="23" pageNumber="524">Diagnosis.</emphasis>
Forewing R2+3 unbranched, R4+5 dichotomously branched. M three- branched, M2 forked. Hindwing M two-branched. Phallosome concave anteriorly, aedeagal arch projected in the middle, truncate. Anterior endophallic sclerites broadly triangular, with a slender arm projected posteriorly, distally acuminate. Posterior endophallic sclerites broadly U-shaped, with a truncate mesal projection directed outward, and a posterior, curved acuminate projection, directed outward. Similar to
<taxonomicName id="4C6D5D85FFFCFFA25B42FB5BB0CFFB05" authority="Carrejo &amp; Obando &amp; Aldrete &amp; Mendivil, 2021" authorityName="Carrejo &amp; Obando &amp; Aldrete &amp; Mendivil" authorityYear="2021" box="[600,729,1231,1257]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="23" pageNumber="524" phylum="Arthropoda" rank="species" species="caucana" status="sp. nov.">
<emphasis id="B919FA14FFFCFFA25B42FB5BB0CFFB05" box="[600,729,1231,1257]" italics="true" pageId="23" pageNumber="524">G. caucana</emphasis>
</taxonomicName>
<emphasis id="B919FA14FFFCFFA25BFAFB5BB130FB04" bold="true" box="[736,806,1231,1257]" pageId="23" pageNumber="524">
<taxonomicNameLabel id="A22A476FFFFCFFA25BFAFB5BB109FB04" box="[736,799,1231,1257]" pageId="23" pageNumber="524" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
for differences with it, see diagnosis of the latter above.
</paragraph>
<paragraph id="8BD22606FFFCFFA259DDFB66B039F9E4" blockId="23.[151,1437,1086,2013]" pageId="23" pageNumber="524">
<emphasis id="B919FA14FFFCFFA259DDFB66B31DFAE1" bold="true" box="[199,267,1266,1292]" pageId="23" pageNumber="524">Color</emphasis>
(in 80% ethanol). Body pale brown. Head dark brown (
<figureCitation id="13563A83FFFCFFA25A80FB67B1E7FAE3" box="[922,1009,1267,1294]" captionStart="FIGURES 6772" captionStartId="24.[151,264,1789,1813]" captionTargetBox="[167,1423,191,1755]" captionTargetId="figure-22@24.[151,1435,181,1765]" captionTargetPageId="24" captionText="FIGURES 6772. G. munchiquensis n. sp. Male. 67. Forewing. 68. Hindwing. 69. Front view of head. 70. Left paraproct and epiproct. 71. Hypandrium. 72. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4431359" httpUri="https://zenodo.org/record/4431359/files/figure.png" pageId="23" pageNumber="524">Fig. 69</figureCitation>
). Compound eyes dark brown, ocelli hyaline, with ochre centripetal crescents; postclypeus without pigmented striae. Antennae pale brown. Maxillary palps pale brown, Mx4 dark brown. Thorax brown, tergal lobes of meso- and metathorax brown, with light brown spots, meso- and metapleura brown, with ochre spots. Legs: fore- coxae cream with pale brown spot basally, midcoxae brown; trochanters and femora cream; tibiae and tarsi brown. Forewings (
<figureCitation id="13563A83FFFCFFA25D05FA17B665FA73" box="[1055,1139,1411,1438]" captionStart="FIGURES 6772" captionStartId="24.[151,264,1789,1813]" captionTargetBox="[167,1423,191,1755]" captionTargetId="figure-22@24.[151,1435,181,1765]" captionTargetPageId="24" captionText="FIGURES 6772. G. munchiquensis n. sp. Male. 67. Forewing. 68. Hindwing. 69. Front view of head. 70. Left paraproct and epiproct. 71. Hypandrium. 72. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4431359" httpUri="https://zenodo.org/record/4431359/files/figure.png" pageId="23" pageNumber="524">Fig. 67</figureCitation>
), brown, with cream areas as illustrated. Hindwings (
<figureCitation id="13563A83FFFCFFA258DAFA33B002FA2F" box="[448,532,1447,1474]" captionStart="FIGURES 6772" captionStartId="24.[151,264,1789,1813]" captionTargetBox="[167,1423,191,1755]" captionTargetId="figure-22@24.[151,1435,181,1765]" captionTargetPageId="24" captionText="FIGURES 6772. G. munchiquensis n. sp. Male. 67. Forewing. 68. Hindwing. 69. Front view of head. 70. Left paraproct and epiproct. 71. Hypandrium. 72. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4431359" httpUri="https://zenodo.org/record/4431359/files/figure.png" pageId="23" pageNumber="524">Fig. 68</figureCitation>
), almost hyaline, light brown, veins brown. Abdomen pale brown. Clunium and hypandrium brown. Phallosome with aedeagal arch and endophallic sclerites dark brown, external parameres cream. Epiproct and paraprocts pale brown.
</paragraph>
<paragraph id="8BD22606FFFCFFA259DDF987B699F8C4" blockId="23.[151,1437,1086,2013]" pageId="23" pageNumber="524">
<emphasis id="B919FA14FFFCFFA259DDF987B377F9C0" bold="true" box="[199,353,1555,1581]" pageId="23" pageNumber="524">Morphology.</emphasis>
As in diagnosis, plus the following: Head: H/MxW: 1.47; compound eyes large, H/d: 4.54; IO/ MxW: 0.81. Outer cusp of lacinial tips broad, with nine denticles. Mx4/Mx2: 1.16. Forewings (
<figureCitation id="13563A83FFFCFFA25DC7F9A3B725F9BF" box="[1245,1331,1591,1618]" captionStart="FIGURES 6772" captionStartId="24.[151,264,1789,1813]" captionTargetBox="[167,1423,191,1755]" captionTargetId="figure-22@24.[151,1435,181,1765]" captionTargetPageId="24" captionText="FIGURES 6772. G. munchiquensis n. sp. Male. 67. Forewing. 68. Hindwing. 69. Front view of head. 70. Left paraproct and epiproct. 71. Hypandrium. 72. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4431359" httpUri="https://zenodo.org/record/4431359/files/figure.png" pageId="23" pageNumber="524">Fig. 67</figureCitation>
): FW/W: 2.74; pterostigma long, wider in the middle: lp/wp: 4.0, areola postica tall, rounded: al/ah: 2.62. Hindwings (
<figureCitation id="13563A83FFFCFFA25C75F9CFB2A1F977" captionStart="FIGURES 6772" captionStartId="24.[151,264,1789,1813]" captionTargetBox="[167,1423,191,1755]" captionTargetId="figure-22@24.[151,1435,181,1765]" captionTargetPageId="24" captionText="FIGURES 6772. G. munchiquensis n. sp. Male. 67. Forewing. 68. Hindwing. 69. Front view of head. 70. Left paraproct and epiproct. 71. Hypandrium. 72. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4431359" httpUri="https://zenodo.org/record/4431359/files/figure.png" pageId="23" pageNumber="524">Fig. 68</figureCitation>
): HW/w: 3.29. Hypandrium (
<figureCitation id="13563A83FFFCFFA25B16F9EBB076F974" box="[524,608,1663,1689]" captionStart="FIGURES 6772" captionStartId="24.[151,264,1789,1813]" captionTargetBox="[167,1423,191,1755]" captionTargetId="figure-22@24.[151,1435,181,1765]" captionTargetPageId="24" captionText="FIGURES 6772. G. munchiquensis n. sp. Male. 67. Forewing. 68. Hindwing. 69. Front view of head. 70. Left paraproct and epiproct. 71. Hypandrium. 72. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4431359" httpUri="https://zenodo.org/record/4431359/files/figure.png" pageId="23" pageNumber="524">Fig. 71</figureCitation>
) almost rectangular, membranous, rounded posteriorly, with setae as illustrated. Phallosome (
<figureCitation id="13563A83FFFCFFA25881F937B3E7F950" box="[411,497,1699,1725]" captionStart="FIGURES 6772" captionStartId="24.[151,264,1789,1813]" captionTargetBox="[167,1423,191,1755]" captionTargetId="figure-22@24.[151,1435,181,1765]" captionTargetPageId="24" captionText="FIGURES 6772. G. munchiquensis n. sp. Male. 67. Forewing. 68. Hindwing. 69. Front view of head. 70. Left paraproct and epiproct. 71. Hypandrium. 72. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4431359" httpUri="https://zenodo.org/record/4431359/files/figure.png" pageId="23" pageNumber="524">Fig. 72</figureCitation>
); side struts slender; aedeagal arch as illustrated; external parameres membranous, joined to the side struts, wide basally, rounded posteriorly, without pores. Paraprocts (
<figureCitation id="13563A83FFFCFFA25D60F953B6C6F90C" box="[1146,1232,1735,1761]" captionStart="FIGURES 6772" captionStartId="24.[151,264,1789,1813]" captionTargetBox="[167,1423,191,1755]" captionTargetId="figure-22@24.[151,1435,181,1765]" captionTargetPageId="24" captionText="FIGURES 6772. G. munchiquensis n. sp. Male. 67. Forewing. 68. Hindwing. 69. Front view of head. 70. Left paraproct and epiproct. 71. Hypandrium. 72. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4431359" httpUri="https://zenodo.org/record/4431359/files/figure.png" pageId="23" pageNumber="524">Fig. 70</figureCitation>
), oval, with setae and macrosetae on distal field; sensory fields with 28 trichobothria on basal rosettes. Epiproct (
<figureCitation id="13563A83FFFCFFA25DD4F97FB736F8E8" box="[1230,1312,1771,1797]" captionStart="FIGURES 6772" captionStartId="24.[151,264,1789,1813]" captionTargetBox="[167,1423,191,1755]" captionTargetId="figure-22@24.[151,1435,181,1765]" captionTargetPageId="24" captionText="FIGURES 6772. G. munchiquensis n. sp. Male. 67. Forewing. 68. Hindwing. 69. Front view of head. 70. Left paraproct and epiproct. 71. Hypandrium. 72. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4431359" httpUri="https://zenodo.org/record/4431359/files/figure.png" pageId="23" pageNumber="524">Fig. 70</figureCitation>
) semioval, rounded posteriorly, with two lateral macrosetae on posterior field, one macroseta mesally.
</paragraph>
</subSubSection>
<subSubSection id="C377758DFFFCFFA259DDF8A7B64EF878" pageId="23" pageNumber="524" type="description">
<paragraph id="8BD22606FFFCFFA259DDF8A7B716F8A0" blockId="23.[151,1437,1086,2013]" box="[199,1280,1842,1869]" pageId="23" pageNumber="524">
<emphasis id="B919FA14FFFCFFA259DDF8A7B368F8A0" bold="true" box="[199,382,1843,1869]" pageId="23" pageNumber="524">Measurements.</emphasis>
FW: 3700, HW: 2800, Mx4: 250, IO: 460, D: 262, d: 185, IO/d: 2.49, PO: 0.71.
</paragraph>
<paragraph id="8BD22606FFFCFFA259DDF8C2B64EF878" blockId="23.[151,1437,1086,2013]" pageId="23" pageNumber="524">
<emphasis id="B919FA14FFFCFFA259DDF8C2B388F89D" bold="true" box="[199,414,1878,1904]" pageId="23" pageNumber="524">Specimen studied.</emphasis>
<materialsCitation id="3B052C5BFFFCFFA258BEF8C3B64EF878" ID-GBIF-Occurrence="3014907324" collectingDate="2017-07-22" collectorName="National Natural Park Cerro Munchique. El Planchon Trail &amp; J. Panche. On" country="Colombia" elevation="2387" latitude="2.6896944" location="Cauca" longLatPrecision="1" longitude="-76.56841" pageId="23" pageNumber="524" specimenCount="1" stateProvince="Cauca" typeStatus="holotype">
<typeStatus id="54D698A4FFFCFFA258BEF8C3B01AF89C" box="[420,524,1879,1905]" pageId="23" pageNumber="524" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="F37A6696FFFCFFA25B4FF8C3B0F8F89C" box="[597,750,1879,1905]" name="Colombia" pageId="23" pageNumber="524">COLOMBIA</collectingCountry>
.
<collectingRegion id="49A9E8E4FFFCFFA25BEDF8C3B156F89C" box="[759,832,1879,1905]" country="Colombia" name="Cauca" pageId="23" pageNumber="524">Cauca</collectingRegion>
,
<collectorName id="269843D0FFFCFFA25A50F8C3B2D9F878" pageId="23" pageNumber="524">National Natural Park Cerro Munchique. El Planchón Trail</collectorName>
,
<geoCoordinate id="EE5940C1FFFCFFA259C1F8EFB39BF878" box="[219,397,1915,1941]" degrees="02" direction="north" minutes="41" orientation="latitude" pageId="23" pageNumber="524" precision="1" seconds="22.9" value="2.6896944">02° 41 22.9N</geoCoordinate>
:
<geoCoordinate id="EE5940C1FFFCFFA25882F8EFB028F878" box="[408,574,1915,1941]" degrees="76" direction="west" minutes="34" orientation="longitude" pageId="23" pageNumber="524" precision="1" seconds="6.3" value="-76.56841">76° 34 6.3W</geoCoordinate>
,
<quantity id="4C958BE3FFFCFFA25B50F8EFB0B4F878" box="[586,674,1915,1941]" metricMagnitude="3" metricUnit="m" metricValue="2.3869999999999996" pageId="23" pageNumber="524" unit="m" value="2387.0">
<elevation id="0040C135FFFCFFA25B50F8EFB0B4F878" box="[586,674,1915,1941]" metricMagnitude="3" metricUnit="m" metricValue="2.3869999999999996" pageId="23" pageNumber="524" unit="m" value="2387.0">2387 m</elevation>
</quantity>
.
<date id="FFD300C6FFFCFFA25BB6F8EFB12FF878" box="[684,825,1915,1941]" pageId="23" pageNumber="524" value="2017-07-22">
<collectingDate id="EF97F92EFFFCFFA25BB6F8EFB12FF878" box="[684,825,1915,1941]" pageId="23" pageNumber="524" value="2017-07-22">22.VII.2017</collectingDate>
</date>
.
<collectorName id="269843D0FFFCFFA25A59F8EFB1CBF878" box="[835,989,1915,1941]" pageId="23" pageNumber="524">J. Panche. On</collectorName>
tree trunk.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C377758DFFFCFFA259DDF80BB3E6F830" pageId="23" pageNumber="524" type="etymology">
<paragraph id="8BD22606FFFCFFA259DDF80BB3E6F830" blockId="23.[151,1437,1086,2013]" pageId="23" pageNumber="524">
<emphasis id="B919FA14FFFCFFA259DDF80BB35EF854" bold="true" box="[199,328,1951,1977]" pageId="23" pageNumber="524">Etymology</emphasis>
. The specific epithet is a name in apposition that refers to the NNP Munchique,
<collectingRegion id="49A9E8E4FFFCFFA25DD0F80BB78EF854" box="[1226,1432,1951,1977]" country="Colombia" name="Cauca" pageId="23" pageNumber="524">Cauca department</collectingRegion>
, where the
<typeStatus id="54D698A4FFFCFFA25816F857B378F830" box="[268,366,1987,2013]" pageId="23" pageNumber="524" type="holotype">holotype</typeStatus>
was found.
</paragraph>
</subSubSection>
</treatment>
</document>