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<document ID-DOI="10.5252/a2015n2a2" ID-ISSN="1639-4798" ID-Zenodo-Dep="5208903" approvalRequired="50" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="44" approvalRequired_for_treatments="5" checkinTime="1629150308770" checkinUser="carolina" docAuthor="Daly, Douglas C., Raharimampionona, Jeannie &amp; Federman, Sarah" docDate="2015" docId="03F5FA39BC4FFFE7B267FD2C30BAF84D" docLanguage="en" docName="Adansonia.37.2.277-345.pdf" docOrigin="Adansonia 37 (2)" docSource="https://www.mendeley.com/catalogue/2522e05a-0ab0-3e53-96aa-d45147a2ca6c/" docStyle="DocumentStyle:E7A860AFEA63D44BCC37955D07FCAFC7.2:Adansonia.2015-2017.journal_article" docStyleId="E7A860AFEA63D44BCC37955D07FCAFC7" docStyleName="Adansonia.2015-2017.journal_article" docStyleVersion="2" docTitle="Canarium egregium Daly, Raharim. &amp; Federman 2015, sp. nov." docType="treatment" docVersion="3" lastPageId="25" lastPageNumber="302" masterDocId="FFCC8241BC58FFFEB157FF99310AFFA1" masterDocTitle="A revision of Canarium L. (Burseraceae) in Madagascar" masterLastPageNumber="345" masterPageNumber="277" pageId="23" pageNumber="300" updateTime="1629151310460" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title>A revision of Canarium L. (Burseraceae) in Madagascar</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Daly, Douglas C.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Raharimampionona, Jeannie</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Federman, Sarah</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Adansonia</mods:title>
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<mods:date>2015</mods:date>
<mods:detail type="pubDate">
<mods:number>2015-12-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>37</mods:number>
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<mods:detail type="issue">
<mods:number>2</mods:number>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5252/a2015n2a2</mods:identifier>
<mods:identifier type="ISSN">1639-4798</mods:identifier>
<mods:identifier type="Zenodo-Dep">5208903</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03F5FA39BC4FFFE7B267FD2C30BAF84D" httpUri="http://treatment.plazi.org/id/03F5FA39BC4FFFE7B267FD2C30BAF84D" lastPageId="25" lastPageNumber="302" pageId="23" pageNumber="300">
<subSubSection box="[816,1453,693,720]" pageId="23" pageNumber="300" type="nomenclature">
<paragraph blockId="23.[816,1453,693,752]" box="[816,1453,693,720]" pageId="23" pageNumber="300">
<heading box="[816,1453,693,720]" centered="true" fontSize="11" level="2" pageId="23" pageNumber="300" reason="3">
<taxonomicName authority="Daly, Raharim. &amp; Federman" authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" box="[816,1357,693,720]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="23" pageNumber="300" phylum="Tracheophyta" rank="species" species="egregium" status="sp. nov.">
<emphasis bold="true" box="[816,1034,693,719]" italics="true" pageId="23" pageNumber="300">Canarium egregium</emphasis>
Daly, Raharim. &amp; Federman
</taxonomicName>
,
<taxonomicNameLabel box="[1367,1453,694,720]" pageId="23" pageNumber="300" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="300" type="description">
<paragraph blockId="23.[816,1453,693,752]" box="[1068,1199,725,752]" pageId="23" pageNumber="300">
(
<figureCitation box="[1075,1144,725,751]" captionStart="FIG" captionStartId="15.[132,143,1228,1246]" captionTargetBox="[131,775,215,1191]" captionTargetId="figure-739@15.[132,775,216,1191]" captionTargetPageId="15" captionText="FIG. 6. — Distributions of Canarium betamponae Daly, Raharim. &amp; Federman, sp. nov., C. bullatum (Leenh.) Daly, Raharim. &amp; Federman, comb. et stat. nov., C. egregium Daly, Raharim. &amp; Federman, sp. nov., and C. indistinctum Daly, Raharim. &amp; Federman, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5208913" httpUri="https://zenodo.org/record/5208913/files/figure.png" pageId="23" pageNumber="300">Figs 6</figureCitation>
;
<figureCitation box="[1156,1186,726,752]" captionStart="FIG" captionStartId="24.[132,143,1939,1957]" captionTargetBox="[199,1381,214,1868]" captionTargetId="figure-19@24.[193,1394,209,1872]" captionTargetPageId="24" captionText="FIG. 11. — Canarium egregium Daly, Raharim. &amp; Federman, sp. nov.: A, branchlet with pistillate flowers and immature fruits; B, pistillate flower with bracteoles and petal (left); C, young developing fruit with persistent bracteole, calyx and staminodes; D, adaxial and abaxial views of staminode; E, basal portion of leaf with detail of stipule; F, portion of infructescence; A-D, Daly et al. 13103 (NY sheet and field photos by D. Daly); E, F, Antilahimena 834 (MO). Scale bars: A, E, F, 2 cm; B, C, 5 mm; D, 1 mm; E, not to scale." figureDoi="http://doi.org/10.5281/zenodo.5209003" httpUri="https://zenodo.org/record/5209003/files/figure.png" pageId="23" pageNumber="300">11</figureCitation>
).
</paragraph>
<paragraph blockId="23.[811,1458,782,1526]" pageId="23" pageNumber="300">
<emphasis italics="true" pageId="23" pageNumber="300">Small to medium-sized trees, leaves 6-8-jugate, leaflet apex gradually acuminate, and lateral leaflet base truncate to slightly cordate; distinguished from</emphasis>
<taxonomicName authority="Engl." authorityName="Engl." box="[863,1110,835,859]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="23" pageNumber="300" phylum="Tracheophyta" rank="species" species="madagascariense">
C. madagascariense
<emphasis box="[1060,1110,835,859]" italics="true" pageId="23" pageNumber="300">Engl.</emphasis>
</taxonomicName>
<emphasis italics="true" pageId="23" pageNumber="300">
by the stipules inserted
<quantity box="[1328,1405,835,859]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="23" pageNumber="300" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">2-5 mm</quantity>
from petiole insertion (vs at petiole base in
</emphasis>
<taxonomicName box="[1174,1366,862,886]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="23" pageNumber="300" phylum="Tracheophyta" rank="species" species="madagascariense">C. madagascariense</taxonomicName>
<emphasis italics="true" pageId="23" pageNumber="300">), stipules symmetrical (vs oblique), inflorescence with darker ferrugineous pubescence, infructescences shorter (</emphasis>
c.
<emphasis italics="true" pageId="23" pageNumber="300">
11 vs
<quantity box="[1108,1168,915,939]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="23" pageNumber="300" unit="cm" value="18.0">18 cm</quantity>
) and fruits slightly obovoid (vs narrowly ovoid).
</emphasis>
</paragraph>
<paragraph blockId="23.[811,1458,782,1526]" pageId="23" pageNumber="300">
<materialsCitation collectingDate="2006-02-10" collectionCode="NY, MO, TEF" collectorName="D. C. Daly &amp; J. Raharimampionona &amp; R. Ranaivojaona" country="Madagascar" county="Diana" elevation="350" latitude="-13.588611" location="Region Anaborano" longLatPrecision="21" longitude="48.709167" municipality="Ambilobe" pageId="23" pageNumber="300" specimenCount="1" stateProvince="Antsiranana">
TYPUS.
<emphasis box="[888,913,990,1013]" italics="true" pageId="23" pageNumber="300"></emphasis>
<emphasis bold="true" box="[920,1038,989,1013]" pageId="23" pageNumber="300">
<collectingCountry box="[920,1038,989,1013]" name="Madagascar" pageId="23" pageNumber="300">Madagascar</collectingCountry>
</emphasis>
. Prov.
<collectingRegion box="[1107,1227,989,1013]" country="Madagascar" name="Antsiranana" pageId="23" pageNumber="300">Antsiranana</collectingRegion>
,
<collectingCounty box="[1238,1300,989,1013]" pageId="23" pageNumber="300">Diana</collectingCounty>
,
<collectingMunicipality box="[1311,1410,989,1013]" pageId="23" pageNumber="300">Ambilobe</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03F5FA39BC4FFFE7B267FD2C30BAF84D:8E831DF4BC4FFFE9B4DAFC4432C1FBAE" country="Madagascar" county="Diana" latitude="-13.588611" longLatPrecision="21" longitude="48.709167" municipality="Ambilobe" name="Region Anaborano" pageId="23" pageNumber="300" stateProvince="Antsiranana">Région Anaborano</location>
,
<location LSID="urn:lsid:plazi:treatment:03F5FA39BC4FFFE7B267FD2C30BAF84D:8E831DF4BC4FFFE9B284FC6E3586FBAE" box="[979,1164,1015,1039]" country="Madagascar" county="Diana" latitude="-13.588611" longLatPrecision="21" longitude="48.709167" municipality="Ambilobe" name="Galoko Mountains" pageId="23" pageNumber="300" stateProvince="Antsiranana">Galoko Mountains</location>
,
<quantity box="[1173,1281,1015,1039]" metricMagnitude="2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="23" pageNumber="300" unit="m" value="350.0" valueMax="400.0" valueMin="300.0">
<elevation box="[1173,1281,1015,1039]" metricMagnitude="2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="23" pageNumber="300" unit="m" value="350.0" valueMax="400.0" valueMin="300.0">300-400 m</elevation>
</quantity>
elev.,
<geoCoordinate box="[1338,1452,1015,1039]" degrees="13" direction="south" minutes="35" orientation="latitude" pageId="23" pageNumber="300" precision="15" seconds="19" value="-13.588611">13°3519”S</geoCoordinate>
,
<geoCoordinate box="[811,926,1042,1066]" degrees="48" direction="east" minutes="42" orientation="longitude" pageId="23" pageNumber="300" precision="15" seconds="33" value="48.709167">48°4233”E</geoCoordinate>
,
<date box="[935,1042,1042,1066]" pageId="23" pageNumber="300" value="2006-02-10">
<collectingDate box="[935,1042,1042,1066]" pageId="23" pageNumber="300" value="2006-02-10">10.II.2006</collectingDate>
</date>
,
<emphasis italics="true" pageId="23" pageNumber="300">
<collectorName box="[1050,1153,1042,1066]" pageId="23" pageNumber="300">D. C. Daly</collectorName>
,
<collectorName box="[1160,1366,1042,1066]" pageId="23" pageNumber="300">J. Raharimampionona</collectorName>
&amp;
<collectorName pageId="23" pageNumber="300">R. Ranaivojaona</collectorName>
13103
</emphasis>
(holo-,
<collectionCode box="[1065,1104,1068,1092]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="23" pageNumber="300" type="Herbarium">NY</collectionCode>
!; iso-, G!,
<collectionCode box="[1203,1251,1068,1092]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="23" pageNumber="300" type="Herbarium">MO</collectionCode>
!,
<collectionCode box="[1263,1314,1068,1092]" country="Madagascar" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" pageId="23" pageNumber="300" type="Herbarium">TEF</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<paragraph blockId="23.[811,1458,782,1526]" pageId="23" pageNumber="300">
<materialsCitation collectingDate="2001-11-17" collectingDateMax="2006-02-10" collectingDateMin="2001-11-17" collectionCode="MO, NY, TEF" collectorName="P. Antilahimena &amp; D. C. Daly &amp; J. Raharimampionona &amp; R. Ranaivojaona" country="Madagascar" county="Diana" elevation="561" latitude="-13.588611" location="Reserve Naturelle Integrale de Tsaratanana" longLatPrecision="21" longitude="48.709167" municipality="Ambanja" pageId="23" pageNumber="300" specimenCount="1" stateProvince="Antsiranana" typeStatus="paratype">
<typeStatus box="[813,902,1116,1139]" pageId="23" pageNumber="300">PARATYPI</typeStatus>
.
<emphasis box="[915,940,1117,1140]" italics="true" pageId="23" pageNumber="300"></emphasis>
<emphasis bold="true" box="[947,1070,1116,1140]" pageId="23" pageNumber="300">
<collectingCountry box="[947,1066,1116,1140]" name="Madagascar" pageId="23" pageNumber="300">Madagascar</collectingCountry>
.
</emphasis>
<collectingRegion box="[1076,1196,1116,1140]" country="Madagascar" name="Antsiranana" pageId="23" pageNumber="300">Antsiranana</collectingRegion>
,
<collectingCounty box="[1207,1269,1116,1140]" pageId="23" pageNumber="300">Diana</collectingCounty>
,
<collectingMunicipality box="[1279,1370,1116,1140]" pageId="23" pageNumber="300">Ambanja</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03F5FA39BC4FFFE7B267FD2C30BAF84D:8E831DF4BC4FFFE9B432FBC53562FB2F" country="Madagascar" county="Diana" latitude="-13.588611" longLatPrecision="21" longitude="48.709167" municipality="Ambanja" name="Reserve Naturelle Integrale de Tsaratanana" pageId="23" pageNumber="300" stateProvince="Antsiranana">Réserve Naturelle Intégrale de Tsaratanana</location>
,
<quantity box="[1136,1198,1142,1166]" metricMagnitude="2" metricUnit="m" metricValue="5.61" pageId="23" pageNumber="300" unit="m" value="561.0">
<elevation box="[1136,1198,1142,1166]" metricMagnitude="2" metricUnit="m" metricValue="5.61" pageId="23" pageNumber="300" unit="m" value="561.0">561 m</elevation>
</quantity>
,
<geoCoordinate box="[1206,1316,1142,1166]" degrees="13" direction="south" minutes="53" orientation="latitude" pageId="23" pageNumber="300" precision="15" seconds="47" value="-13.896389">13°5347”S</geoCoordinate>
,
<geoCoordinate box="[1323,1452,1142,1166]" degrees="048" direction="east" minutes="51" orientation="longitude" pageId="23" pageNumber="300" precision="15" seconds="57" value="48.865833">048 °5157”E</geoCoordinate>
,
<date box="[811,927,1169,1193]" pageId="23" pageNumber="300" value="2001-11-17">
<collectingDate box="[811,927,1169,1193]" pageId="23" pageNumber="300" value="2001-11-17">17.XI.2001</collectingDate>
</date>
,
<emphasis box="[941,1140,1169,1193]" italics="true" pageId="23" pageNumber="300">
<collectorName box="[941,1095,1169,1193]" pageId="23" pageNumber="300">P. Antilahimena</collectorName>
834
</emphasis>
(
<collectionCode box="[1156,1202,1169,1193]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="23" pageNumber="300" type="Herbarium">MO</collectionCode>
); Diana, Ambilobe, Anaborano,
<location LSID="urn:lsid:plazi:treatment:03F5FA39BC4FFFE7B267FD2C30BAF84D:8E831DF4BC4FFFE9B22AFB323533FB62" box="[893,1081,1195,1219]" country="Madagascar" county="Diana" latitude="-13.588611" longLatPrecision="21" longitude="48.709167" municipality="Ambanja" name="Galoko Mountains" pageId="23" pageNumber="300" stateProvince="Antsiranana">Galoko Mountains</location>
,
<quantity box="[1089,1201,1195,1219]" metricMagnitude="2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="23" pageNumber="300" unit="m" value="350.0" valueMax="400.0" valueMin="300.0">
<elevation box="[1089,1201,1195,1219]" metricMagnitude="2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="23" pageNumber="300" unit="m" value="350.0" valueMax="400.0" valueMin="300.0">300-400 m</elevation>
</quantity>
,
<geoCoordinate box="[1210,1325,1195,1219]" degrees="13" direction="south" minutes="35" orientation="latitude" pageId="23" pageNumber="300" precision="15" seconds="19" value="-13.588611">13°3519”S</geoCoordinate>
,
<geoCoordinate box="[1334,1452,1195,1219]" degrees="48" direction="east" minutes="42" orientation="longitude" pageId="23" pageNumber="300" precision="15" seconds="33" value="48.709167">48°4233”E</geoCoordinate>
,
<date box="[812,919,1221,1245]" pageId="23" pageNumber="300" value="2006-02-10">
<collectingDate box="[812,919,1221,1245]" pageId="23" pageNumber="300" value="2006-02-10">10.II.2006</collectingDate>
</date>
,
<emphasis italics="true" pageId="23" pageNumber="300">
<collectorName box="[932,1041,1222,1246]" pageId="23" pageNumber="300">D. C. Daly</collectorName>
,
<collectorName box="[1052,1263,1222,1246]" pageId="23" pageNumber="300">J. Raharimampionona</collectorName>
&amp;
<collectorName box="[1297,1455,1222,1245]" pageId="23" pageNumber="300">R. Ranaivojaona</collectorName>
13096, 130988
</emphasis>
(
<collectionCode box="[978,1014,1248,1272]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="23" pageNumber="300" type="Herbarium">NY</collectionCode>
,
<collectionCode box="[1023,1072,1248,1272]" country="Madagascar" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" pageId="23" pageNumber="300" type="Herbarium">TEF</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="300" type="distribution">
<paragraph blockId="23.[811,1458,782,1526]" pageId="23" pageNumber="300">
DISTRIBUTION AND ECOLOGY. —
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[1153,1342,1296,1319]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="23" pageNumber="300" phylum="Tracheophyta" rank="species" species="egregium" status="sp. nov.">
<emphasis box="[1153,1342,1296,1319]" italics="true" pageId="23" pageNumber="300">Canarium egregium</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[1356,1433,1296,1319]" pageId="23" pageNumber="300" rank="species">sp. nov.</taxonomicNameLabel>
is known only from the Sambirano region, in the upper Ramena and the Réserve Naturelle Intégrale de Tsaratanana, where it is a small to medium-sized tree in moist forests with a relatively closed canopy
<emphasis box="[1439,1454,1376,1399]" italics="true" pageId="23" pageNumber="300">c.</emphasis>
<quantity box="[813,863,1401,1425]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="23" pageNumber="300" unit="m" value="25.0">25 m</quantity>
high, on steep slopes with rock outcrops, between
<quantity box="[1346,1455,1401,1425]" metricMagnitude="2" metricUnit="m" metricValue="3.55" metricValueMax="5.6" metricValueMin="1.5" pageId="23" pageNumber="300" unit="m" value="355.0" valueMax="560.0" valueMin="150.0">150-560 m</quantity>
elevation. Flowering Feb., fruiting Nov.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5208923" ID-Zenodo-Dep="5208923" httpUri="https://zenodo.org/record/5208923/files/figure.png" pageId="23" pageNumber="300" startId="23.[132,142,1444,1462]" targetBox="[132,775,214,1402]" targetPageId="23">
<paragraph blockId="23.[132,777,1444,1485]" pageId="23" pageNumber="300">
FIG. 10.— Distributions of
<taxonomicName authority="Daly,Raharim.&amp; Federman" authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" box="[340,707,1444,1462]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="23" pageNumber="300" phylum="Tracheophyta" rank="species" species="elegans" status="sp.nov.">
<emphasis box="[340,484,1444,1462]" italics="true" pageId="23" pageNumber="300">Canarium elegans</emphasis>
Daly,Raharim.&amp; Federman
</taxonomicName>
,
<taxonomicNameLabel box="[713,777,1444,1462]" pageId="23" pageNumber="300" rank="species">sp.nov.</taxonomicNameLabel>
and
<taxonomicName authority="Daly, Raharim. &amp; Federman" authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" box="[169,537,1467,1485]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="23" pageNumber="300" phylum="Tracheophyta" rank="species" species="scholasticum" status="sp. nov.">
<emphasis box="[169,302,1467,1485]" italics="true" pageId="23" pageNumber="300">C. scholasticum</emphasis>
Daly, Raharim. &amp; Federman
</taxonomicName>
,
<taxonomicNameLabel box="[544,611,1467,1485]" pageId="23" pageNumber="300" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection pageId="23" pageNumber="300" type="etymology">
<paragraph blockId="23.[811,1458,782,1526]" pageId="23" pageNumber="300">
ETYMOLOGY. — The specific epithet
<taxonomicName authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[1153,1234,1476,1499]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="23" pageNumber="300" phylum="Tracheophyta" rank="species" species="egregium">
<emphasis box="[1153,1234,1476,1499]" italics="true" pageId="23" pageNumber="300">egregium</emphasis>
</taxonomicName>
means “to stand apart,” which is certainly true of this distinctive new species.
</paragraph>
</subSubSection>
<subSubSection lastPageId="25" lastPageNumber="302" pageId="23" pageNumber="300" type="description">
<paragraph blockId="23.[811,1456,1587,2028]" box="[813,957,1587,1613]" pageId="23" pageNumber="300">DESCRIPTION</paragraph>
<paragraph blockId="23.[811,1456,1587,2028]" pageId="23" pageNumber="300">
Trees, reproductive size
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<emphasis box="[1152,1168,1619,1644]" italics="true" pageId="23" pageNumber="300">×</emphasis>
<quantity box="[1170,1273,1618,1645]" metricMagnitude="-1" metricUnit="m" metricValue="3.1" metricValueMax="4.7" metricValueMin="1.5" pageId="23" pageNumber="300" unit="cm" value="31.0" valueMax="47.0" valueMin="15.0">15-47 cm</quantity>
diam, with plank buttresses to
<quantity box="[953,998,1650,1677]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="300" unit="m" value="1.0">1 m</quantity>
high. Outer bark gray, thin, rough, some raised lenticels, shed in thin, irregular, variable-sized plates; inner bark reddish(-tan). Leaves
<quantity box="[1163,1268,1714,1741]" metricMagnitude="-1" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="23" pageNumber="300" unit="cm" value="32.5" valueMax="35.0" valueMin="30.0">30-35 cm</quantity>
long, 6-8-jugate, petiole
<quantity box="[893,1013,1746,1773]" metricMagnitude="-2" metricUnit="m" metricValue="5.65" metricValueMax="6.8" metricValueMin="4.5" pageId="23" pageNumber="300" unit="cm" value="5.65" valueMax="6.8" valueMin="4.5">4.5-6.8 cm</quantity>
long, usually dilated at base, petiole and rachis on young leaves with dense fine straight (but variously oriented) hairs to
<quantity box="[999,1083,1810,1836]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="23" pageNumber="300" unit="mm" value="0.7">0.7 mm</quantity>
long (these sometimes deciduous), also with dense capitate glands and dense flexuous hairs to
<quantity box="[814,902,1874,1900]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="300" unit="mm" value="0.2">0.2 mm</quantity>
long; stipules inserted
<quantity box="[1156,1245,1873,1900]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="23" pageNumber="300" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">2-5 mm</quantity>
from petiole insertion,
<quantity box="[872,962,1905,1932]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="23" pageNumber="300" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5-8 mm</quantity>
long, broadly ovate or possibly suborbicular, with dense appressed to ascending ferrugineous hairs, stipule scar
<emphasis box="[904,921,1970,1995]" italics="true" pageId="23" pageNumber="300">c.</emphasis>
<quantity box="[927,994,1969,1996]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="23" pageNumber="300" unit="mm" value="4.0">4 mm</quantity>
long; basal petiolules
<quantity box="[1237,1327,1969,1996]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="23" pageNumber="300" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4-5 mm</quantity>
long, other laterals
<quantity box="[897,1000,2001,2028]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="11.0" metricValueMin="5.0" pageId="23" pageNumber="300" unit="mm" value="8.0" valueMax="11.0" valueMin="5.0">5-11 mm</quantity>
long, terminal
<quantity box="[1170,1286,2001,2028]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="23" pageNumber="300" unit="mm" value="22.5" valueMax="25.0" valueMin="20.0">20-25 mm</quantity>
long, pulvinuli
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5209003" ID-Zenodo-Dep="5209003" httpUri="https://zenodo.org/record/5209003/files/figure.png" pageId="24" pageNumber="301" startId="24.[132,143,1939,1957]" targetBox="[199,1381,214,1868]" targetPageId="24">
<paragraph blockId="24.[132,1457,1939,2025]" pageId="24" pageNumber="301">
FIG. 11. —
<taxonomicName authority="Daly, Raharim. &amp; Federman" authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" box="[224,625,1939,1957]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="24" pageNumber="301" phylum="Tracheophyta" rank="species" species="egregium" status="sp. nov.">
<emphasis box="[224,388,1939,1957]" italics="true" pageId="24" pageNumber="301">Canarium egregium</emphasis>
Daly, Raharim. &amp; Federman
</taxonomicName>
,
<taxonomicNameLabel box="[633,703,1939,1957]" pageId="24" pageNumber="301" rank="species">sp. nov.</taxonomicNameLabel>
:
<emphasis bold="true" box="[710,723,1939,1957]" pageId="24" pageNumber="301">A</emphasis>
, branchlet with pistillate flowers and immature fruits;
<emphasis bold="true" box="[1172,1185,1939,1957]" pageId="24" pageNumber="301">B</emphasis>
, pistillate flower with bracteoles and petal (left);
<emphasis bold="true" box="[259,273,1962,1980]" pageId="24" pageNumber="301">C</emphasis>
, young developing fruit with persistent bracteole, calyx and staminodes;
<emphasis bold="true" box="[875,894,1962,1980]" pageId="24" pageNumber="301">D,</emphasis>
adaxial and abaxial views of staminode;
<emphasis bold="true" box="[1231,1243,1962,1980]" pageId="24" pageNumber="301">E</emphasis>
, basal portion of leaf with detail of stipule;
<emphasis bold="true" box="[268,279,1985,2003]" pageId="24" pageNumber="301">F</emphasis>
, portion of infructescence;
<emphasis bold="true" box="[505,539,1985,2003]" pageId="24" pageNumber="301">A -D</emphasis>
,
<emphasis box="[548,685,1985,2003]" italics="true" pageId="24" pageNumber="301">Daly et al. 13103</emphasis>
(NY sheet and field photos by D. Daly);
<emphasis bold="true" box="[1012,1024,1985,2003]" pageId="24" pageNumber="301">E</emphasis>
,
<emphasis bold="true" box="[1033,1044,1985,2003]" pageId="24" pageNumber="301">F</emphasis>
,
<emphasis box="[1054,1197,1985,2003]" italics="true" pageId="24" pageNumber="301">Antilahimena 834</emphasis>
(MO). Scale bars: A, E, F, 2 cm; B, C, 5 mm; D, 1 mm; E, not to scale.
</paragraph>
</caption>
<paragraph blockId="25.[130,778,215,1549]" pageId="25" pageNumber="302">
inconspicuous; basal leaflets 3.5-4.5
<emphasis box="[546,562,216,241]" italics="true" pageId="25" pageNumber="302">×</emphasis>
<quantity box="[574,679,215,242]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" metricValueMax="3.0" metricValueMin="2.6" pageId="25" pageNumber="302" unit="cm" value="2.8" valueMax="3.0" valueMin="2.6">2.6-3 cm</quantity>
, broadly ovate, other laterals 6-19
<emphasis box="[408,424,248,273]" italics="true" pageId="25" pageNumber="302">×</emphasis>
<quantity box="[432,555,247,274]" metricMagnitude="-2" metricUnit="m" metricValue="4.699999999999999" metricValueMax="6.6" metricValueMin="2.8" pageId="25" pageNumber="302" unit="cm" value="4.699999999999999" valueMax="6.6" valueMin="2.8">2.8-6.6 cm</quantity>
, oblong(-lanceolate) to (broadly) ovate, terminal one 8-14
<emphasis box="[536,552,280,305]" italics="true" pageId="25" pageNumber="302">×</emphasis>
<quantity box="[558,679,279,306]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" metricValueMax="5.5" metricValueMin="4.1" pageId="25" pageNumber="302" unit="cm" value="4.8" valueMax="5.5" valueMin="4.1">4.1-5.5 cm</quantity>
, ovate to broadly elliptic; leaflet apex gradually acuminate (rarely acute), the acumen to 10 (20) mm long, base of laterals cordate to truncate, subsymmetric, less often slightly oblique; margin often slightly revolute, leaflets coriaceous, drying grayishbrown, abaxial side somewhat glossy and adaxial side less so; secondary venation weakly brochidodromous with some festooning, secondary veins in 13-19 pairs, slightly spreading (almost straight), insertion excurrent, spacing decreasing toward apex and especially toward base, angle slightly acute but increasing toward the base, some perpendicular intersecondary veins and (often multiple) perpendicular epimedial tertiaries, the intercostal tertiaries alternate-percurrent, quaternaries regular-polygonal; on leaflet abaxial side the midvein and secondaries prominent, the rest narrowly prominulous, the surface papillate and glabrous or the midvein with hairs as on the rachis; on adaxial side all venation narrowly prominulous except the midvein sunk in a groove, glabrous. Pistillate inflorescences (staminate unknown)
<quantity box="[504,630,853,880]" metricMagnitude="-1" metricUnit="m" metricValue="1.925" metricValueMax="2.3" metricValueMin="1.55" pageId="25" pageNumber="302" unit="cm" value="19.25" valueMax="23.0" valueMin="15.5">15.5-23 cm</quantity>
long, robust, the secondary axes 2-6 (13.5) cm long, axes with dense fine darkly ferrugineous hairs to
<quantity box="[448,539,917,943]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="25" pageNumber="302" unit="mm" value="0.6">0.6 mm</quantity>
; bracteoles
<quantity box="[670,774,917,943]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="11.0" metricValueMin="6.0" pageId="25" pageNumber="302" unit="mm" value="8.5" valueMax="11.0" valueMin="6.0">6-11 mm</quantity>
long, broadly ovate, slightly acuminate and with slightly constricted base, the scars broad and semi-clasping; pedicel
<quantity box="[133,221,1013,1039]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="25" pageNumber="302" unit="mm" value="1.0" valueMax="2.0" valueMin="0.0">0-2 mm</quantity>
long, slightly clavate. Pistillate flowers
<emphasis box="[637,654,1013,1038]" italics="true" pageId="25" pageNumber="302">c.</emphasis>
<quantity box="[660,713,1013,1039]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="25" pageNumber="302" unit="cm" value="1.0">1 cm</quantity>
long; calyx deeply cupular and fleshy,
<emphasis box="[476,493,1045,1070]" italics="true" pageId="25" pageNumber="302">c.</emphasis>
<quantity box="[499,564,1045,1071]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="25" pageNumber="302" unit="mm" value="7.0">7 mm</quantity>
long, red, the lobes rounded perdepressed-deltate,
<quantity box="[474,566,1076,1103]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="25" pageNumber="302" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1-3 mm</quantity>
long, surface with dense thick retrorse-appressed to descending ferrugineous hairs to
<quantity box="[217,304,1140,1167]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="25" pageNumber="302" unit="mm" value="0.2">0.2 mm</quantity>
; corolla (passed) red, exposed part negligible. Infructescences
<quantity box="[306,378,1172,1198]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="25" pageNumber="302" unit="cm" value="11.0">11 cm</quantity>
long, secondary axes to
<quantity box="[654,710,1172,1198]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="25" pageNumber="302" unit="cm" value="3.0">3 cm</quantity>
long; persistent bracts on primary axes
<emphasis box="[485,502,1205,1230]" italics="true" pageId="25" pageNumber="302">c.</emphasis>
<quantity box="[509,573,1204,1230]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="25" pageNumber="302" unit="mm" value="8.0">8 mm</quantity>
long, fleshy, ovate, semi-clasping, the margin long-ciliate. Fruiting pedicel
<emphasis box="[760,777,1236,1261]" italics="true" pageId="25" pageNumber="302">c.</emphasis>
<quantity box="[132,196,1268,1294]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="25" pageNumber="302" unit="mm" value="4.0">4 mm</quantity>
long, subcylindric, slender, sparsely raised-lenticellate; fruiting calyx large,
<emphasis box="[356,373,1300,1325]" italics="true" pageId="25" pageNumber="302">c.</emphasis>
<quantity box="[379,460,1300,1326]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="25" pageNumber="302" unit="mm" value="12.0">12 mm</quantity>
long, the lobes only slightly distinct, nearly patent, chartaceous, sparsely raised-lenticellate, with long, appressed, ferrugineous hairs. Fruits (immature) 4.3-5
<emphasis box="[200,216,1396,1421]" italics="true" pageId="25" pageNumber="302">×</emphasis>
<quantity box="[227,332,1395,1422]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="25" pageNumber="302" unit="cm" value="1.95" valueMax="2.0" valueMin="1.9">1.9-2 cm</quantity>
, narrowly ovoid (slightly obovoid when very young), apex slightly acuminate but apex tip truncate, base acute, surface smooth or sparsely and finely lenticellate, glabrous or with a few scattered ascending fine white hairs to
<quantity box="[163,228,1523,1549]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="25" pageNumber="302" unit="mm" value="1.0">1 mm</quantity>
long near apex.
</paragraph>
<paragraph blockId="25.[130,777,1587,2028]" box="[132,206,1587,1613]" pageId="25" pageNumber="302">NOTES</paragraph>
<paragraph blockId="25.[130,777,1587,2028]" pageId="25" pageNumber="302">
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[132,339,1619,1645]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="25" pageNumber="302" phylum="Tracheophyta" rank="species" species="egregium" status="sp. nov.">
<emphasis box="[132,339,1619,1645]" italics="true" pageId="25" pageNumber="302">Canarium egregium</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[351,433,1619,1645]" pageId="25" pageNumber="302" rank="species">sp. nov.</taxonomicNameLabel>
belongs to a group of 10 species that all have relatively large, broad leaflets with truncate to slightly cordate base. Like
<taxonomicName box="[416,616,1682,1709]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="25" pageNumber="302" phylum="Tracheophyta" rank="species" species="madagascariense">
<emphasis box="[416,616,1682,1709]" italics="true" pageId="25" pageNumber="302">C. madagascariense</emphasis>
</taxonomicName>
, it has numerous leaflets (6-8 vs 7-10 pairs), stipules conspicuous, leaflet gradually acuminate, primary bracts on inflorescence large and often persistent, calyx in fruit large, and glabrous fruit.
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[133,340,1810,1836]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="25" pageNumber="302" phylum="Tracheophyta" rank="species" species="egregium" status="sp. nov.">
<emphasis box="[133,340,1810,1836]" italics="true" pageId="25" pageNumber="302">Canarium egregium</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[356,439,1810,1836]" pageId="25" pageNumber="302" rank="species">sp. nov.</taxonomicNameLabel>
differs by the stipules
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from petiole insertion (vs at petiole base), the stipules symmetrical (vs oblique), leaflet secondary vein angle more acute (except at base), inflorescence with darker ferrugineous pubescence, infructescences shorter (
<emphasis box="[506,523,1938,1963]" italics="true" pageId="25" pageNumber="302">c.</emphasis>
11 vs
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), fruiting pedicel shorter (
<emphasis box="[299,315,1970,1995]" italics="true" pageId="25" pageNumber="302">c.</emphasis>
4 vs
<quantity box="[364,430,1969,1996]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="25" pageNumber="302" unit="mm" value="8.0">8 mm</quantity>
), and distinct fruit shape (slightly obovoid vs narrowly ovoid).
</paragraph>
</subSubSection>
</treatment>
</document>