284 lines
30 KiB
XML
284 lines
30 KiB
XML
<document ID-DOI="10.11646/zootaxa.5231.5.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="7609734" ID-ZooBank="4EC66405-7706-4810-B516-BCAEB6A2DA6A" checkinTime="1675674648572" checkinUser="plazi" docAuthor="Nabozhenko, Maxim V., Perkovsky, Evgeny E. & Nazarenko, Vitaliy Yu." docDate="2023" docId="03DA87946652FFB3FF5FFBB2FABDF7CC" docLanguage="en" docName="zootaxa.5231.5.7.pdf" docOrigin="Zootaxa 5231 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5231.5.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Archaeoluprops Nabozhenko & Perkovsky & Nazarenko 2023, gen. n." docType="treatment" docVersion="3" lastPageNumber="587" masterDocId="FFE3FFEC6653FFB0FFC8FFA7FF97F661" masterDocTitle="A new tribe, genus and species of the tenebrionid subfamily Lagriinae (Coleoptera: Tenebrionidae) from the Eocene Baltic amber" masterLastPageNumber="590" masterPageNumber="584" pageNumber="585" updateTime="1675764477379" updateUser="tatiana" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new tribe, genus and species of the tenebrionid subfamily Lagriinae (Coleoptera: Tenebrionidae) from the Eocene Baltic amber</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Nabozhenko, Maxim V.</mods:namePart>
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<mods:affiliation>Dagestan State University, M. Gadzhiev str., 43 a, Makhachkala, Republic of Dagestan 367000, Russia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Perkovsky, Evgeny E.</mods:namePart>
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<mods:affiliation>I. I. Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, B. Khmelnitskogo 15, 01030 Kiev, Ukraine & A. A. Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow, 117647 Russia</mods:affiliation>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Nazarenko, Vitaliy Yu.</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0003-4245-5049</mods:nameIdentifier>
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<mods:affiliation>I. I. Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, B. Khmelnitskogo 15, 01030 Kiev, Ukraine & nazarenko @ izan. kiev. ua; https: // orcid. org / 0000 - 0003 - 4245 - 5049</mods:affiliation>
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<mods:nameIdentifier type="email">nazarenko@izan.kiev.ua</mods:nameIdentifier>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-02-02</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5231</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>5</mods:number>
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<mods:extent unit="page">
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<mods:start>584</mods:start>
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<mods:end>590</mods:end>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5231.5.7</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5231.5.7</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7609734</mods:identifier>
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<mods:identifier type="ZooBank">4EC66405-7706-4810-B516-BCAEB6A2DA6A</mods:identifier>
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<paragraph blockId="1.[151,522,1045,1108]" box="[151,522,1045,1072]" pageId="1" pageNumber="585">
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<heading bold="true" box="[151,522,1045,1072]" fontSize="11" level="1" pageId="1" pageNumber="585" reason="1">
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<emphasis bold="true" box="[151,522,1045,1072]" pageId="1" pageNumber="585">
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Genus
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<taxonomicName authority="Nabozhenko & Perkovsky & Nazarenko, 2023" authorityName="Nabozhenko & Perkovsky & Nazarenko" authorityYear="2023" box="[238,432,1045,1072]" class="Insecta" family="Tenebrionidae" genus="Archaeoluprops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis bold="true" box="[238,432,1045,1072]" italics="true" pageId="1" pageNumber="585">Archaeoluprops</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[440,522,1046,1072]" pageId="1" pageNumber="585" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,268,1082,1108]" pageId="1" pageNumber="585" type="description">
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<paragraph blockId="1.[151,522,1045,1108]" box="[151,268,1082,1108]" pageId="1" pageNumber="585">
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(
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<figureCitation box="[159,230,1082,1108]" captionStart="FIGURE 1" captionStartId="2.[152,255,1978,2002]" captionTargetBox="[151,1436,849,1955]" captionTargetId="figure-249@2.[151,1436,849,1955]" captionTargetPageId="2" captionText="FIGURE 1. Archaeoluprops groehni, gen. and sp. n., habitus: A—dorsal view; B—ventral view." figureDoi="http://doi.org/10.5281/zenodo.7609738" httpUri="https://zenodo.org/record/7609738/files/figure.png" pageId="1" pageNumber="585">Figs 1</figureCitation>
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,
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<figureCitation box="[244,259,1082,1108]" captionStart="FIGURE 2" captionStartId="3.[152,255,1171,1195]" captionTargetBox="[151,1436,453,1148]" captionTargetId="figure-126@3.[151,1436,453,1148]" captionTargetPageId="3" captionText="FIGURE 2. Archaeoluprops groehni, gen. and sp. n., details of structure:A—head and pronotum, dorsal view; B—pterothorax, ventral view (shows that mesocoxal cavities open, and trochantine is absent): Ms—mesoventrite, Ms-est—mesepisternum, Msepm—mesepimeron, Ms-cox—mesocoxae, Mt—metaventrite, Mt-est—metepisternum, Mt-epm—metepimeron." figureDoi="http://doi.org/10.5281/zenodo.7609740" httpUri="https://zenodo.org/record/7609740/files/figure.png" pageId="1" pageNumber="585">2</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,603,1150,1175]" pageId="1" pageNumber="585" type="type_taxon">
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<paragraph blockId="1.[151,603,1150,1175]" box="[151,603,1150,1175]" pageId="1" pageNumber="585">
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<typeStatus box="[151,201,1151,1175]" pageId="1" pageNumber="585">Type</typeStatus>
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species:
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<taxonomicName authority="Nabozhenko & Perkovsky & Nazarenko, 2023" authorityName="Nabozhenko & Perkovsky & Nazarenko" authorityYear="2023" box="[294,540,1150,1174]" class="Insecta" family="Tenebrionidae" genus="Archaeoluprops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="species" species="groehni" status="sp. n.">
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<emphasis box="[294,540,1150,1174]" italics="true" pageId="1" pageNumber="585">Archaeoluprops groehni</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[546,603,1150,1174]" pageId="1" pageNumber="585">
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<taxonomicNameLabel box="[546,603,1150,1174]" pageId="1" pageNumber="585" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="585" type="diagnosis">
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<paragraph blockId="1.[151,1437,1220,1751]" pageId="1" pageNumber="585">
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<emphasis bold="true" box="[151,625,1220,1247]" pageId="1" pageNumber="585">Diagnosis of the new tribe and genus.</emphasis>
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Tarsal formula 5-5-4. Trochanters heteromeroid. Body robust, shiny, pubescent with erected and suberected comparatively long setae at least dorsally (
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<figureCitation box="[1089,1180,1256,1283]" captionStart="FIGURE 1" captionStartId="2.[152,255,1978,2002]" captionTargetBox="[151,1436,849,1955]" captionTargetId="figure-249@2.[151,1436,849,1955]" captionTargetPageId="2" captionText="FIGURE 1. Archaeoluprops groehni, gen. and sp. n., habitus: A—dorsal view; B—ventral view." figureDoi="http://doi.org/10.5281/zenodo.7609738" httpUri="https://zenodo.org/record/7609738/files/figure.png" pageId="1" pageNumber="585">Fig. 1A</figureCitation>
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). Eyes large, reniform (
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<figureCitation box="[159,254,1292,1319]" captionStart="FIGURE 2" captionStartId="3.[152,255,1171,1195]" captionTargetBox="[151,1436,453,1148]" captionTargetId="figure-126@3.[151,1436,453,1148]" captionTargetPageId="3" captionText="FIGURE 2. Archaeoluprops groehni, gen. and sp. n., details of structure:A—head and pronotum, dorsal view; B—pterothorax, ventral view (shows that mesocoxal cavities open, and trochantine is absent): Ms—mesoventrite, Ms-est—mesepisternum, Msepm—mesepimeron, Ms-cox—mesocoxae, Mt—metaventrite, Mt-est—metepisternum, Mt-epm—metepimeron." figureDoi="http://doi.org/10.5281/zenodo.7609740" httpUri="https://zenodo.org/record/7609740/files/figure.png" pageId="1" pageNumber="585">Fig. 2A</figureCitation>
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). Maxillary apical palpomere triangular, elongate. Antennae moniliform. Lateral margins and base of pronotum coarsely beaded. Prosternum long before procoxae, prosternal process comparatively wide, not convex (
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<figureCitation box="[159,245,1364,1391]" captionStart="FIGURE 1" captionStartId="2.[152,255,1978,2002]" captionTargetBox="[151,1436,849,1955]" captionTargetId="figure-249@2.[151,1436,849,1955]" captionTargetPageId="2" captionText="FIGURE 1. Archaeoluprops groehni, gen. and sp. n., habitus: A—dorsal view; B—ventral view." figureDoi="http://doi.org/10.5281/zenodo.7609738" httpUri="https://zenodo.org/record/7609738/files/figure.png" pageId="1" pageNumber="585">Fig. 1B</figureCitation>
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). Mesocoxal cavities open (partly closed by an extension of the mesepimeron) (
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<figureCitation box="[1131,1218,1364,1391]" captionStart="FIGURE 2" captionStartId="3.[152,255,1171,1195]" captionTargetBox="[151,1436,453,1148]" captionTargetId="figure-126@3.[151,1436,453,1148]" captionTargetPageId="3" captionText="FIGURE 2. Archaeoluprops groehni, gen. and sp. n., details of structure:A—head and pronotum, dorsal view; B—pterothorax, ventral view (shows that mesocoxal cavities open, and trochantine is absent): Ms—mesoventrite, Ms-est—mesepisternum, Msepm—mesepimeron, Ms-cox—mesocoxae, Mt—metaventrite, Mt-est—metepisternum, Mt-epm—metepimeron." figureDoi="http://doi.org/10.5281/zenodo.7609740" httpUri="https://zenodo.org/record/7609740/files/figure.png" pageId="1" pageNumber="585">Fig. 2B</figureCitation>
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). Metaventrite with long median groove. Elytra non-striate, densely confusedly punctuate (
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<figureCitation box="[950,1040,1400,1427]" captionStart="FIGURE 1" captionStartId="2.[152,255,1978,2002]" captionTargetBox="[151,1436,849,1955]" captionTargetId="figure-249@2.[151,1436,849,1955]" captionTargetPageId="2" captionText="FIGURE 1. Archaeoluprops groehni, gen. and sp. n., habitus: A—dorsal view; B—ventral view." figureDoi="http://doi.org/10.5281/zenodo.7609738" httpUri="https://zenodo.org/record/7609738/files/figure.png" pageId="1" pageNumber="585">Fig. 1A</figureCitation>
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). Epipleura almost reaching elytral suture angle. Intersegmental membranes not visible between 3–5 ventrites, hinging tentyrioid (
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<figureCitation box="[1219,1306,1436,1463]" captionStart="FIGURE 1" captionStartId="2.[152,255,1978,2002]" captionTargetBox="[151,1436,849,1955]" captionTargetId="figure-249@2.[151,1436,849,1955]" captionTargetPageId="2" captionText="FIGURE 1. Archaeoluprops groehni, gen. and sp. n., habitus: A—dorsal view; B—ventral view." figureDoi="http://doi.org/10.5281/zenodo.7609738" httpUri="https://zenodo.org/record/7609738/files/figure.png" pageId="1" pageNumber="585">Fig. 1B</figureCitation>
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), defensive glands absent. Outer edge of the ventrite 5 is formed into a groove which receives the elytral edges. Preapical tarsomeres cupuliform.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="587" pageId="1" pageNumber="585" type="etymology">
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<paragraph blockId="1.[151,1437,1220,1751]" pageId="1" pageNumber="585">
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<emphasis bold="true" box="[199,334,1545,1571]" pageId="1" pageNumber="585">Etymology.</emphasis>
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The name is combined from archaeo- (ancient—Greek origin) and
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<taxonomicName authorityName="Hope" authorityYear="1833" box="[1153,1245,1545,1570]" class="Insecta" family="Tenebrionidae" genus="Luprops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="genus">
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<emphasis box="[1153,1245,1545,1570]" italics="true" pageId="1" pageNumber="585">Luprops</emphasis>
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</taxonomicName>
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, the name of a tenebrionid genus within the tribe
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<taxonomicName authority="Ardoin, 1958" authorityName="Ardoin" authorityYear="1958" box="[536,806,1580,1607]" pageId="1" pageNumber="585" rank="tribe" tribe="Lupropini">Lupropini Ardoin, 1958</taxonomicName>
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.
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</paragraph>
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<paragraph blockId="1.[151,1437,1220,1751]" pageId="1" pageNumber="585">
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<emphasis bold="true" box="[199,354,1616,1642]" pageId="1" pageNumber="585">Comparison.</emphasis>
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<taxonomicName authority="Nabozhenko & Perkovsky & Nazarenko, 2023" authorityName="Nabozhenko & Perkovsky & Nazarenko" authorityYear="2023" box="[359,629,1616,1642]" class="Insecta" family="Tenebrionidae" genus="Archaeoluprops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="species" species="groehni" status="sp. n.">
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<emphasis box="[359,629,1616,1642]" italics="true" pageId="1" pageNumber="585">Archaeoluprops groehni</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[635,697,1617,1643]" pageId="1" pageNumber="585">
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<taxonomicNameLabel box="[635,697,1617,1643]" pageId="1" pageNumber="585" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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is externally similar to the representatives of the tribes
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<taxonomicName authorityName="Lacordaire" authorityYear="1859" box="[1303,1437,1616,1642]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="tribe" tribe="Goniaderini">Goniaderini</taxonomicName>
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and
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<taxonomicName authorityName="Ardoin" authorityYear="1958" box="[197,309,1652,1678]" pageId="1" pageNumber="585" rank="tribe" tribe="Lupropini">Lupropini</taxonomicName>
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by the robust body and estriate elytra, but differs in the absence of intersegmental membranes between abdominal ventrites 3–5 (tentyrioid hinging) and accordingly the primary absence of defensive glands.
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</paragraph>
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<paragraph blockId="1.[151,1437,1220,1751]" box="[199,635,1724,1751]" pageId="1" pageNumber="585">
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<taxonomicName authority="Nabozhenko & Perkovsky & Nazarenko, 2023" authorityName="Nabozhenko & Perkovsky & Nazarenko" authorityYear="2023" box="[199,397,1724,1750]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="tribe" status="trib. n." tribe="Archaeolupropini">Archaeolupropini</taxonomicName>
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<emphasis bold="true" box="[404,486,1724,1751]" pageId="1" pageNumber="585">
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<taxonomicNameLabel box="[404,486,1724,1751]" pageId="1" pageNumber="585" rank="tribe">trib. n.</taxonomicNameLabel>
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</emphasis>
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differs from:
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</paragraph>
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<paragraph blockId="1.[151,1436,1796,2039]" pageId="1" pageNumber="585">
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<emphasis box="[151,165,1797,1822]" italics="true" pageId="1" pageNumber="585">–</emphasis>
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<taxonomicName authority="W. Kirby, 1828" authorityName="W. Kirby" authorityYear="1828" box="[198,466,1796,1823]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="tribe" tribe="Adeliini">
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Adeliini W. Kirby,
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</taxonomicName>
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in the tentyrioid hinging of the abdomen (tenebrioid in
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<taxonomicName authorityName="W. Kirby" authorityYear="1828" box="[1086,1182,1796,1822]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="tribe" tribe="Adeliini">Adeliini</taxonomicName>
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), estriate elytra (striate in
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<taxonomicName authorityName="W. Kirby" authorityYear="1828" box="[225,321,1832,1858]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="tribe" tribe="Adeliini">Adeliini</taxonomicName>
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||
), the weakly widened apical maxillary palpomere (wide, triangle to securiform in
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<taxonomicName authorityName="W. Kirby" authorityYear="1828" box="[1231,1327,1832,1858]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="tribe" tribe="Adeliini">Adeliini</taxonomicName>
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), the long metaventrite (short in almost all
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||
<taxonomicName authorityName="W. Kirby" authorityYear="1828" box="[560,652,1868,1894]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="tribe" tribe="Adeliini">Adeliini</taxonomicName>
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except several winged species (
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<bibRefCitation author="Matthews, E. G. & Bouchard, P." box="[1008,1330,1868,1895]" pageId="1" pageNumber="585" refId="ref3970" refString="Matthews, E. G. & Bouchard, P. (2008) Tenebrionid Beetles of Australia: Descriptions of Tribes, Keys to Genera, Catalogue of Species. Australian Biological Resources Study, Canberra, 410 pp." type="book" year="2008">Matthews & Bouchard 2008</bibRefCitation>
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)), the not expressed trochantine (well visible in
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<taxonomicName authorityName="W. Kirby" authorityYear="1828" box="[622,719,1904,1930]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="tribe" tribe="Adeliini">Adeliini</taxonomicName>
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);
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</paragraph>
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<paragraph blockId="1.[151,1436,1796,2039]" pageId="1" pageNumber="585">
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–
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<taxonomicName authority="Reitter, 1917" authorityName="Reitter" authorityYear="1917" box="[198,457,1940,1967]" class="Insecta" family="Tenebrionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="tribe" tribe="Belopini">
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Belopini Reitter,
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||
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||
</taxonomicName>
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in estriate elytra, the pubescent body (body is bare in
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<taxonomicName authorityName="Reitter" authorityYear="1917" box="[1122,1221,1940,1966]" class="Insecta" family="Tenebrionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="tribe" tribe="Belopini">Belopini</taxonomicName>
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, excluding extinct
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<taxonomicName authorityName="Nabozhenko, Kirejtshuk & Merkl" authorityYear="2016" box="[198,351,1977,2003]" class="Insecta" family="Lagriidae" genus="Yantaroxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="genus">Yantaroxenos</taxonomicName>
|
||
with very dense scaled setae), cupuliform preapical tarsomeres (simple in
|
||
<taxonomicName authorityName="Reitter" authorityYear="1917" box="[1187,1289,1976,2002]" class="Insecta" family="Tenebrionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="tribe" tribe="Belopini">Belopini</taxonomicName>
|
||
);
|
||
</paragraph>
|
||
<paragraph blockId="1.[151,1436,1796,2039]" lastBlockId="2.[151,1437,151,825]" lastPageId="2" lastPageNumber="586" pageId="1" pageNumber="585">
|
||
– Chaerodini Doyen, Matthews & Lawrence,
|
||
<quantity box="[715,805,2012,2039]" metricMagnitude="1" metricUnit="m" metricValue="5.0546" pageId="1" pageNumber="585" unit="in" value="1990.0">1990 in</quantity>
|
||
simple legs (short and fossorial Chaerodini), the not semispherical body, the prosternum long before procoxae (short in Chaerodini), hinging tentyrioid (tenebrionoid in Chaerodini);
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,825]" pageId="2" pageNumber="586">
|
||
–
|
||
<taxonomicName authority="Latreille, 1802" authorityName="Latreille" authorityYear="1802" box="[198,500,223,249]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="586" phylum="Arthropoda" rank="tribe" tribe="Cossyphini">
|
||
Cossyphini Latreille,
|
||
<quantity box="[444,500,223,249]" metricMagnitude="1" metricUnit="m" metricValue="4.57708" pageId="2" pageNumber="586" unit="in" value="1802.0">1802</quantity>
|
||
</taxonomicName>
|
||
in the not explanate and stongly flattened body and not concealed head (pronotal flange covering head in Cossiphyni), the extensions of the hypomeron are divided by the prosternal process behind procoxae (extensions of the hypomeron are connected behind procoxae in Cossiphini), open mesocoxal cavities (closed by meso- and metaventrite in
|
||
<taxonomicName authorityName="Latreille" authorityYear="1802" box="[712,843,331,357]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="586" phylum="Arthropoda" rank="tribe" tribe="Cossyphini">Cossyphini</taxonomicName>
|
||
);
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,825]" pageId="2" pageNumber="586">
|
||
– Eschatoporiini Blaisdell, 1906 (see characters in
|
||
<bibRefCitation author="Aalbu, R. L. & Kanda, K. & Smith, A. D." box="[774,1011,367,393]" pageId="2" pageNumber="586" refId="ref3141" refString="Aalbu, R. L., Kanda, K. & Smith, A. D. (2017) Reinstatement of Eschatoporiini Blaisdell, 1906, a unique tribe of blind cavernicolous Tenebrionidae from California, with a new species from Napa County (Coleoptera, Tenebrionidae, Lagriinae). ZooKeys, 688, 151 - 149. https: // doi. org / 10.3897 / zookeys. 688.13575" type="book" year="2017">
|
||
Aalbu
|
||
<emphasis box="[854,908,367,393]" italics="true" pageId="2" pageNumber="586">et al</emphasis>
|
||
. (2017))
|
||
</bibRefCitation>
|
||
in the estriate elytra, the concealed membrane between labrum and epistoma (the membrane exposed in Eschatoporiini), the scutellum concealed by the pronotum, with only scutellar shield visible (scutellum open in Eschatoporiini), large eyes (Eschatoporiini are without eyes);
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,825]" pageId="2" pageNumber="586">
|
||
–
|
||
<taxonomicName authority="Lacordaire, 1859" authorityName="Lacordaire" authorityYear="1859" box="[198,532,510,537]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="586" phylum="Arthropoda" rank="tribe" tribe="Goniaderini">
|
||
Goniaderini Lacordaire,
|
||
<quantity box="[476,532,510,537]" metricMagnitude="1" metricUnit="m" metricValue="4.7218599999999995" pageId="2" pageNumber="586" unit="in" value="1859.0">1859</quantity>
|
||
</taxonomicName>
|
||
in the tentyrioid hinging of the abdomen (tenebrioid in
|
||
<taxonomicName authorityName="W. Kirby" authorityYear="1828" box="[1169,1263,511,537]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="586" phylum="Arthropoda" rank="tribe" tribe="Adeliini">Adeliini</taxonomicName>
|
||
) and the apical maxillary palpomere weakly widened (distinctly triangle in
|
||
<taxonomicName authorityName="Lacordaire" authorityYear="1859" box="[869,1008,547,573]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="586" phylum="Arthropoda" rank="tribe" tribe="Goniaderini">Goniaderini</taxonomicName>
|
||
);
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,825]" pageId="2" pageNumber="586">
|
||
–
|
||
<taxonomicName box="[198,343,583,609]" pageId="2" pageNumber="584" rank="tribe" tribe="Gonialaenini">Gonialaenini</taxonomicName>
|
||
<bibRefCitation author="Nabozhenko, M. V. & Bukejs, A. & Telnov, D." box="[348,756,583,609]" pageId="2" pageNumber="586" pagination="253 - 260" refId="ref4238" refString="Nabozhenko, M. V., Bukejs, A. & Telnov, D. (2019) Gonialaenini, a new tribe of Lagriinae (Coleoptera: Tenebrionidae) from Eocene Baltic Amber. Zootaxa, 4565 (2), 253 - 260. https: // doi. org / 10.11646 / zootaxa. 4565.2.9" type="journal article" year="2019">Nabozhenko, Bukejs et Telnov, 2019</bibRefCitation>
|
||
(extinct) in the tentyrioid hinging of the abdomen (tenebrioid in
|
||
<taxonomicName authorityName="W. Kirby" authorityYear="1828" box="[229,325,619,645]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="586" phylum="Arthropoda" rank="tribe" tribe="Adeliini">Adeliini</taxonomicName>
|
||
), estriate elytra (striae are visible in
|
||
<taxonomicName box="[755,904,619,645]" pageId="2" pageNumber="584" rank="tribe" tribe="Gonialaenini">Gonialaenini</taxonomicName>
|
||
), weakly widened maxillary apical palpomere (securiform in
|
||
<taxonomicName box="[373,522,655,681]" pageId="2" pageNumber="584" rank="tribe" tribe="Gonialaenini">Gonialaenini</taxonomicName>
|
||
), coarsely bordered pronotum (unbordered in
|
||
<taxonomicName box="[1063,1212,655,681]" pageId="2" pageNumber="584" rank="tribe" tribe="Gonialaenini">Gonialaenini</taxonomicName>
|
||
), long metaventrite (moderate metaventrite, wings probably reduced in
|
||
<taxonomicName box="[777,927,691,717]" pageId="2" pageNumber="584" rank="tribe" tribe="Gonialaenini">Gonialaenini</taxonomicName>
|
||
);
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,825]" pageId="2" pageNumber="586">
|
||
– Laenini Seidlitz,
|
||
<quantity box="[391,477,726,753]" metricMagnitude="1" metricUnit="m" metricValue="4.8133" pageId="2" pageNumber="586" unit="in" value="1895.0">1895 in</quantity>
|
||
the tentyrioid hinging (tenebrionoid in
|
||
<taxonomicName authorityName="Latreille" authorityYear="1825" box="[923,1018,727,753]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="586" phylum="Arthropoda" rank="tribe" tribe="Lagriini">Lagriini</taxonomicName>
|
||
); the presence of humeral angles, the absence of setiferous pores on elytra, large and reniform eyes (more or less rounded in Laenini), in estriate elytra, cupuliform preapical tarsomeres;
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7609738" ID-Zenodo-Dep="7609738" box="[152,1142,1978,2003]" httpUri="https://zenodo.org/record/7609738/files/figure.png" pageId="2" pageNumber="586" startId="2.[152,255,1978,2002]" targetBox="[151,1436,849,1955]" targetPageId="2">
|
||
<paragraph blockId="2.[152,1142,1978,2003]" box="[152,1142,1978,2003]" pageId="2" pageNumber="586">
|
||
<emphasis bold="true" box="[152,281,1978,2002]" pageId="2" pageNumber="586">FIGURE 1.</emphasis>
|
||
<emphasis box="[287,539,1978,2002]" italics="true" pageId="2" pageNumber="586">
|
||
<taxonomicName authority="Nabozhenko & Perkovsky & Nazarenko, 2023" authorityName="Nabozhenko & Perkovsky & Nazarenko" authorityYear="2023" box="[287,534,1978,2002]" class="Insecta" family="Tenebrionidae" genus="Archaeoluprops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="586" phylum="Arthropoda" rank="species" species="groehni" status="gen. and sp. n.">Archaeoluprops groehni</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicNameLabel box="[545,696,1978,2003]" pageId="2" pageNumber="586" rank="species">
|
||
<emphasis bold="true" box="[545,590,1979,2003]" pageId="2" pageNumber="586">gen.</emphasis>
|
||
and
|
||
<emphasis bold="true" box="[639,696,1979,2003]" pageId="2" pageNumber="586">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, habitus: A—dorsal view; B—ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="3.[151,1437,150,429]" pageId="3" pageNumber="587">
|
||
–
|
||
<taxonomicName authority="Latreille, 1825" authorityName="Latreille" authorityYear="1825" box="[198,458,150,177]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="tribe" tribe="Lagriini">Lagriini Latreille, 1825</taxonomicName>
|
||
the tentyrioid hinging (tenebrionoid in
|
||
<taxonomicName authorityName="Latreille" authorityYear="1825" box="[895,990,151,177]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="tribe" tribe="Lagriini">Lagriini</taxonomicName>
|
||
), the pronotal and elytral bases subequal (pronotum much narrower than elytral base in
|
||
<taxonomicName authorityName="Latreille" authorityYear="1825" box="[763,858,187,213]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="tribe" tribe="Lagriini">Lagriini</taxonomicName>
|
||
), anterior margin of epistoma straight (arquately emarginated in
|
||
<taxonomicName authorityName="Latreille" authorityYear="1825" box="[372,467,223,249]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="tribe" tribe="Lagriini">Lagriini</taxonomicName>
|
||
);
|
||
</paragraph>
|
||
<paragraph blockId="3.[151,1437,150,429]" pageId="3" pageNumber="587">
|
||
–
|
||
<taxonomicName authorityName="Ardoin" authorityYear="1958" box="[198,310,259,285]" pageId="3" pageNumber="585" rank="tribe" tribe="Lupropini">Lupropini</taxonomicName>
|
||
Lesne,
|
||
<quantity box="[398,483,259,285]" metricMagnitude="1" metricUnit="m" metricValue="4.89204" pageId="3" pageNumber="587" unit="in" value="1926.0">1926 in</quantity>
|
||
the tentyrioid hinging (tenebrionoid in
|
||
<taxonomicName authorityName="Ardoin" authorityYear="1958" box="[926,1043,259,285]" pageId="3" pageNumber="585" rank="tribe" tribe="Lupropini">Lupropini</taxonomicName>
|
||
), eyes large and reniform (round in
|
||
<taxonomicName authorityName="Ardoin" authorityYear="1958" box="[198,315,295,321]" pageId="3" pageNumber="585" rank="tribe" tribe="Lupropini">Lupropini</taxonomicName>
|
||
), open mesocoxal cavities (closed by meso- and metaventrite in
|
||
<taxonomicName authorityName="Ardoin" authorityYear="1958" box="[1035,1152,295,321]" pageId="3" pageNumber="585" rank="tribe" tribe="Lupropini">Lupropini</taxonomicName>
|
||
);
|
||
</paragraph>
|
||
<paragraph blockId="3.[151,1437,150,429]" pageId="3" pageNumber="587">
|
||
– Pycnocerini Lacordaire,
|
||
<quantity box="[481,570,330,357]" metricMagnitude="1" metricUnit="m" metricValue="4.7218599999999995" pageId="3" pageNumber="587" unit="in" value="1859.0">1859 in</quantity>
|
||
the tentyrioid hinging (tenebrionoid in Pycnocerini), body densely setated at least dorsally (completely bare in Pycnocerini), the pronotal and elytral bases subequal (pronotum much narrower than elytral base in Pycnocerini), cupuliform preapical tarsomeres (simple in Pycnocerini).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |