234 lines
23 KiB
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234 lines
23 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.901.39432" ID-GBIF-Dataset="654566e8-947b-4dfe-8fbc-baea3d254fa8" ID-PMC="PMC6965333" ID-Pensoft-Pub="1313-2970-901-1" ID-Pensoft-UUID="D8C38C1C3844549F8BEC7C1D884CDF8E" ID-PubMed="31975782" ModsDocID="1313-2970-901-1" checkinTime="1578615027095" checkinUser="pensoft" docAuthor="Balkenohl, Michael" docDate="2020" docId="704FD76839B55F92ACDA9F7A31B9BF51" docLanguage="en" docName="ZooKeys 901: 1-81" docOrigin="ZooKeys 901" docSource="http://dx.doi.org/10.3897/zookeys.901.39432" docTitle="Salcedia tuberculata Balkenohl 2020, sp. nov." docType="treatment" docVersion="5" id="D8C38C1C3844549F8BEC7C1D884CDF8E" lastPageNumber="1" masterDocId="D8C38C1C3844549F8BEC7C1D884CDF8E" masterDocTitle="A genus in disguise. Revision of the genus Salcedia Fairmaire, 1899 with descriptions of nine new species (Coleoptera, Carabidae, Scaritinae, Salcediini)" masterLastPageNumber="81" masterPageNumber="1" pageNumber="1" updateTime="1668168267548" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A genus in disguise. Revision of the genus Salcedia Fairmaire, 1899 with descriptions of nine new species (Coleoptera, Carabidae, Scaritinae, Salcediini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Balkenohl, Michael</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>901</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>81</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.901.39432</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.901.39432</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-901-1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">D8C38C1C3844549F8BEC7C1D884CDF8E</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="161671755" LSID="urn:lsid:zoobank.org:act:7367647F-C04E-4326-A9AB-8502FC1A4AE2" httpUri="http://treatment.plazi.org/id/704FD76839B55F92ACDA9F7A31B9BF51" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="704FD768-39B5-5F92-ACDA-9F7A31B9BF51" authority="Balkenohl, 2020" authorityName="Balkenohl" authorityYear="2020" class="Insecta" family="Carabidae" genus="Salcedia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salcedia tuberculata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov.">Salcedia tuberculata</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figures 20–25" captionStartId="F9" captionText="Figures 20 - 25. Salcedia species, habitus, dorsal view, holotype 20 S. utetea sp. nov. 21 S. lukulua sp. nov. 22 S. robusta sp. nov. 23 S. tuberculata sp. nov. 24 S. miranda (Andrewes) 25 S. parallela Baehr." figureDoi="10.3897/zookeys.901.39432.figures20-25" httpUri="https://binary.pensoft.net/fig/370610" pageId="0" pageNumber="1">Figs 23</figureCitation>
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<figureCitation captionStart="Figures 36–43" captionStartId="F11" captionText="Figures 36 - 43. Salcedia species, pronotum, dorsal view 36 S. procera sp. nov. 37 S. baroensis sp. nov. 38 S. utetea sp. nov. 39 S. lukulua sp. nov. 40 S. robusta sp. nov. 41 S. tuberculata sp. nov. 42 S. miranda (Andrewes) 43 S. parallela Baehr." figureDoi="10.3897/zookeys.901.39432.figures36-43" httpUri="https://binary.pensoft.net/fig/370612" pageId="0" pageNumber="1">, 41</figureCitation>
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<figureCitation captionStart="Figures 64–67" captionStartId="F15" captionText="Figures 64 - 67. Salcedia species, male genitalia, dorsal view of aedeagus (a) and parameres (b, c), lateral view (d). Small sketches in black symbolising the apex in cross section 64 S. utetea sp. nov. 65 S. robusta sp. nov. 66 S. tuberculata sp. nov. 67 S. miranda (Andrewes)." figureDoi="10.3897/zookeys.901.39432.figures64-67" httpUri="https://binary.pensoft.net/fig/370616" pageId="0" pageNumber="1">, 66</figureCitation>
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<figureCitation captionStart="Figure 84" captionStartId="F18" captionText="Figure 84. Map of tropical Africa providing an overview of the occurrence of all African Salcedia species described to date (recorded localities plotted). 1 (black S. perrieri Fairmaire; 2 (white) S. unifoveata sp. nov.; 3 (black circle) S. faillei sp. nov.; 4 (red) S. coquilhati Alluaud; 5 (yellow) S. elongata Alluaud; 6 (blue) S. africana (Britton); 7 (orange) S. utetea sp. nov.; 8 (grey) S. procera sp. nov.; 9 (light green) S. robusta sp. nov.; 10 (turquoise) S. schoutedeni Alluaud; 11 (purple) S. lukulua sp. nov.; 12 (dark green) S. tuberculata sp. nov.; 13 (brown) S. baroensis sp. nov.; 14 (light blue) S. nigeriensis Alluaud; 16 (dark blue) S. putzeysi (Oberthuer); 17 (black) S. matsumotoi sp. nov.;? in a circle (twice): collection locality of a single female Salcedia specimen of uncertain species. Basic schematic map of Africa taken from SimpleMappr. net." figureDoi="10.3897/zookeys.901.39432.figure84" httpUri="https://binary.pensoft.net/fig/370619" pageId="0" pageNumber="1">, 84</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Holotype</emphasis>
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: ♂, with labels and data: white, printed "S. Afr; KrugerNat. Pk PundaMariaNgotsodam
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<geoCoordinate degrees="21.26" direction="south" orientation="latitude" precision="555" value="-21.26">21.26S</geoCoordinate>
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,
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<geoCoordinate degrees="31.14" direction="east" orientation="longitude" precision="555" value="31.14">31.14E</geoCoordinate>
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/ 7.2.1994; E-Y: 2984 shorewashing leg.
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<normalizedToken originalValue="Endrödy-Younga”">Endroedy-Younga"</normalizedToken>
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/ "Salcedia africana (BRITT.) det.M.
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<normalizedToken originalValue="Baehr’03”">Baehr'03"</normalizedToken>
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/ "SALCEDIA africana (
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<bibRefCitation author="Britton, EB" journalOrPublisher="Entomologist's monthly magazine, May" pageId="0" pageNumber="1" pagination="125 - 127" refId="B18" refString="Britton, EB, 1947. A new species of Zelma (Col., Carabidae) from Africa. Entomologist's monthly magazine, May 1947: 125 - 127" title="A new species of Zelma (Col., Carabidae) from Africa." volume="1947" year="1947">Britton 1947</bibRefCitation>
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) P. Bulirsch det. 2012" (SMNS).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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A medium sized species, with oblong-elongate outline of the elytra with maximum width at middle and the pronotum with three additional complete carinae. The pseudohumerus is nearly rectangular and not dentate. The antennomeres are sub-elongate. Distinguished most clearly from the similar species
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<taxonomicName lsidName="S. africana" pageId="0" pageNumber="1" rank="species" species="africana">
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<emphasis italics="true" pageId="0" pageNumber="1">S. africana</emphasis>
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</taxonomicName>
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by the pronotum with its base laterally with straight but obtuse lobe produced posteriorly and with a distinct peg-like tubercle. Distinguished at the first glance from all other species by the interneur four of the elytron which shows a distinct knob-like tubercle at base (in addition to the tubercle at base of the interneur two present in all species).
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</paragraph>
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<caption doi="10.3897/zookeys.901.39432.figures64-67" httpUri="https://binary.pensoft.net/fig/370616" pageId="0" pageNumber="1" start="Figures 64–67" startId="F15">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figures 64-67.</emphasis>
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<taxonomicName authorityName="Fairmaire" authorityYear="1899" class="Insecta" family="Carabidae" genus="Salcedia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salcedia" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Salcedia</emphasis>
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</taxonomicName>
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species, male genitalia, dorsal view of aedeagus (
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<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
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) and parameres (
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<emphasis bold="true" pageId="0" pageNumber="1">b, c</emphasis>
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), lateral view (
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<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
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). Small sketches in black symbolising the apex in cross section
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<emphasis bold="true" pageId="0" pageNumber="1">64</emphasis>
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<taxonomicName lsidName="S. utetea" pageId="0" pageNumber="1" rank="species" species="utetea">
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<emphasis italics="true" pageId="0" pageNumber="1">S. utetea</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="1">65</emphasis>
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<taxonomicName lsidName="S. robusta" pageId="0" pageNumber="1" rank="species" species="robusta">
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<emphasis italics="true" pageId="0" pageNumber="1">S. robusta</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="1">66</emphasis>
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<taxonomicName lsidName="S. tuberculata" pageId="0" pageNumber="1" rank="species" species="tuberculata">
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<emphasis italics="true" pageId="0" pageNumber="1">S. tuberculata</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="1">67</emphasis>
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<taxonomicName lsidName="S. miranda" pageId="0" pageNumber="1" rank="species" species="miranda">
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<emphasis italics="true" pageId="0" pageNumber="1">S. miranda</emphasis>
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</taxonomicName>
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(Andrewes).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Measurements in Table
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<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements and ratios of Salcedia species. N = number of specimens measured; x ̄ = arithmetic mean; " term used " refers to the shape of the elytra." httpUri="http://table.plazi.org/id/FDDA58A1B347C9F0593A738CF707345F" pageId="0" pageNumber="1" tableUuid="FDDA58A1B347C9F0593A738CF707345F">1</tableCitation>
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.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour and surface:</emphasis>
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Fuscous to piceous, areas between carinae and pits shiny; lateral fifth of pronotum fuscous; mandibles piceous; legs, antennae and palpi fuscous; pronotum with posterior and lateral margin densely covered with beige-grey pili, more intensive around posterior angles.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head:</emphasis>
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Four-fifths of the pronotum width. Outline shaped like a frustum of a pyramid. Clypeus wide, indistinctly convex anteriorly, fused with clypeal wings, separated from supra-antennal plates by distinct obtuse notches, with square elevated field at middle, separated step-like from frons. Frons with two raised paramedian carinae, joining anteriorly, prolonged anteriorly into a central keel, with two small acute glossy teeth bilaterally anterior to central keel, with two parallel carinae paralaterally near base, converging anteriorly, broadened basally; frons and clypeus separated from supra-antennal and supraorbital plates by deep broad furrows, each furrow with deep circular pit between supra-antennal plate and clypeus; supra-antennal and supraorbital plates margined, margin raised, carina-like, at supraorbital plates excised concavely, supra-antennal plates slightly vaulted, surface of supraorbital and supra-antennal plates with numerous small flat irregular impressions. Basal border indistinctly emarginated at middle, distinctly and obtuse angled laterally (angle 118-120°). Eyes convex, oval shaped in lateral view, with small part just visible in dorsal view; genae slightly convex, with small notch anterior angle. Antenna with segments five to ten sub-elongate (L/W 1.17), densely pubescent, segments two to four scarcely pubescent, scapus and pedicellus with indistinct reticulation. Labrum clearly visible from above, nearly straight anteriorly. Mandible moderately short, wide, regularly arcuate at apex. Mentum pentagonal; epilobe of mentum moderately wide, projecting and distinctly angled anteriorly, completely margined, somewhat hollowed out, surface covered regularly with pits.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Pronotum</emphasis>
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(Fig.
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<figureCitation captionStart="Figures 36–43" captionStartId="F11" captionText="Figures 36 - 43. Salcedia species, pronotum, dorsal view 36 S. procera sp. nov. 37 S. baroensis sp. nov. 38 S. utetea sp. nov. 39 S. lukulua sp. nov. 40 S. robusta sp. nov. 41 S. tuberculata sp. nov. 42 S. miranda (Andrewes) 43 S. parallela Baehr." figureDoi="10.3897/zookeys.901.39432.figures36-43" httpUri="https://binary.pensoft.net/fig/370612" pageId="0" pageNumber="1">41</figureCitation>
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): Outline rectangular, transverse, one fifth wider than long. Lateral margin convex, converging to anterior angles, maximum width at end of second third. Lateral margin distinctly crenulated, with thirteen tubercles, with distinct emargination at posterior angles, posterior angles acute. Base laterally with straight but obtuse lobe produced posteriorly, lobe with tubercle towards middle, with broad and deep emargination, with broad flat keel at middle. Disc, with two raised paramedian carinae parallel to median line, diverging posteriorly, finely tuberculate on top, with median line small, ending in pits anteriorly and posteriorly, with four additional shorter carinae bilaterally, the paralateral one joining with the paramedian carinae at base and forming tooth-like tubercle pointing posteriorly, extended anteriorly as less raised paralateral carina. With shorter not raised inner lateral carina and quite long more raised outer lateral carina. Carinae uneven on top. Lateral margin and space between lateral margin and paralateral carina wing-like bent up, with six large transverse pits.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Elytron:</emphasis>
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Indistinctly depressed in anterior half (lateral view), moderately convex in frontal view. Oblong-elongate, margin at middle straight but indistinctly diverging, maximum width at middle. Pseudohumerus nearly rectangular, not dentate. Apex long-oval, retracted. Disc with interneur six indistinctly crenulated, interneur two running up to apex as indistinctly convex line, moderately raised; interneur four running slightly convex at middle, distinctly convex towards apex, not reaching apex, reaching base, with distinct knob-like tubercle at base. Interneur five and six with two rows of serial pits.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Hind wings:</emphasis>
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Reduced by approximately 50%.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Lower surface:</emphasis>
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Antennal channel of pronotum with isodiametric reticulation. Pseudoepipleura with two rows of pits, lateral margin of elytron smooth. Metepisternum moderately long, with flat longitudinal groove. Last visible sternum with obtuse longitudinal carina at middle, with rough pits laterally.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Legs:</emphasis>
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Profemora with dorsal surface roughly reticulated. Protibia longer, with terminal spine curved regularly, laterally with four teeth of decreasing size, dorsally and ventrally with two carinae. Movable spur short, length a third of first tarsomere, slightly curved ventrally. Meso- and metafemora of moderate length, slender. First tarsomeres distinctly elongated, almost as long as tarsomeres two to five together.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">External sexual dimorphism:</emphasis>
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Not observed.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Male genitalia</emphasis>
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(Fig.
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<figureCitation captionStart="Figures 64–67" captionStartId="F15" captionText="Figures 64 - 67. Salcedia species, male genitalia, dorsal view of aedeagus (a) and parameres (b, c), lateral view (d). Small sketches in black symbolising the apex in cross section 64 S. utetea sp. nov. 65 S. robusta sp. nov. 66 S. tuberculata sp. nov. 67 S. miranda (Andrewes)." figureDoi="10.3897/zookeys.901.39432.figures64-67" httpUri="https://binary.pensoft.net/fig/370616" pageId="0" pageNumber="1">66</figureCitation>
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): Median lobe moderately short, in dorsal view cracked at end of basal third and beginning of apical third, in lateral view bisinuate, with fine scattered pili from basal third nearly up to apex, apex sub-spatulate, in cross section explanate dorsally, convex ventrally. Oroficium half as long as median lobe. Endophallus with elongated group of microtrichia conspicuously regularly arranged, with additional two small groups of finer and short trichia basally and apically. Dorsal paramere elongated, bisinuate, with elongated apophyses, broader at base; ventral one like spatulate, hyaline at apex; both parameres slightly distorted.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Female genitalia:</emphasis>
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Unknown.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Variation:</emphasis>
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Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">The name refers to the distinct tubercle at base of the interneur four of the elytra as well as to the distinct tubercles at the base of the pronotum.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution</paragraph>
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<paragraph pageId="0" pageNumber="1">
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(Fig.
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<figureCitation captionStart="Figure 84" captionStartId="F18" captionText="Figure 84. Map of tropical Africa providing an overview of the occurrence of all African Salcedia species described to date (recorded localities plotted). 1 (black S. perrieri Fairmaire; 2 (white) S. unifoveata sp. nov.; 3 (black circle) S. faillei sp. nov.; 4 (red) S. coquilhati Alluaud; 5 (yellow) S. elongata Alluaud; 6 (blue) S. africana (Britton); 7 (orange) S. utetea sp. nov.; 8 (grey) S. procera sp. nov.; 9 (light green) S. robusta sp. nov.; 10 (turquoise) S. schoutedeni Alluaud; 11 (purple) S. lukulua sp. nov.; 12 (dark green) S. tuberculata sp. nov.; 13 (brown) S. baroensis sp. nov.; 14 (light blue) S. nigeriensis Alluaud; 16 (dark blue) S. putzeysi (Oberthuer); 17 (black) S. matsumotoi sp. nov.;? in a circle (twice): collection locality of a single female Salcedia specimen of uncertain species. Basic schematic map of Africa taken from SimpleMappr. net." figureDoi="10.3897/zookeys.901.39432.figure84" httpUri="https://binary.pensoft.net/fig/370619" pageId="0" pageNumber="1">84</figureCitation>
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). Known from the type locality, in the Kruger National Park, Republic of South Africa.
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</paragraph>
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<caption doi="10.3897/zookeys.901.39432.figures68-76" httpUri="https://binary.pensoft.net/fig/370617" pageId="0" pageNumber="1" start="Figures 68–76" startId="F16">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figures 68-76.</emphasis>
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<taxonomicName authorityName="Fairmaire" authorityYear="1899" class="Insecta" family="Carabidae" genus="Salcedia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salcedia" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Salcedia</emphasis>
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</taxonomicName>
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species, female coxostyli (gonopod IX) and laterotergite IX (epipleutite IX)
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<emphasis bold="true" pageId="0" pageNumber="1">68</emphasis>
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<taxonomicName lsidName="S. perrieri" pageId="0" pageNumber="1" rank="species" species="perrieri">
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<emphasis italics="true" pageId="0" pageNumber="1">S. perrieri</emphasis>
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</taxonomicName>
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Fairmaire
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<emphasis bold="true" pageId="0" pageNumber="1">69</emphasis>
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<taxonomicName lsidName="S. unifoveata" pageId="0" pageNumber="1" rank="species" species="unifoveata">
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<emphasis italics="true" pageId="0" pageNumber="1">S. unifoveata</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="1">70</emphasis>
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<taxonomicName lsidName="S. coquilhati" pageId="0" pageNumber="1" rank="species" species="coquilhati">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. coquilhati</emphasis>
|
||
</taxonomicName>
|
||
Alluaud
|
||
<emphasis bold="true" pageId="0" pageNumber="1">71</emphasis>
|
||
<taxonomicName lsidName="S. elongata" pageId="0" pageNumber="1" rank="species" species="elongata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. elongata</emphasis>
|
||
</taxonomicName>
|
||
Alluaud
|
||
<emphasis bold="true" pageId="0" pageNumber="1">72</emphasis>
|
||
<taxonomicName lsidName="S. africana" pageId="0" pageNumber="1" rank="species" species="africana">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. africana</emphasis>
|
||
</taxonomicName>
|
||
(Britton)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">73</emphasis>
|
||
<taxonomicName lsidName="S. schoutedeni" pageId="0" pageNumber="1" rank="species" species="schoutedeni">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. schoutedeni</emphasis>
|
||
</taxonomicName>
|
||
Alluaud
|
||
<emphasis bold="true" pageId="0" pageNumber="1">74</emphasis>
|
||
<taxonomicName lsidName="S. nigeriensis" pageId="0" pageNumber="1" rank="species" species="nigeriensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. nigeriensis</emphasis>
|
||
</taxonomicName>
|
||
Alluaud
|
||
<emphasis bold="true" pageId="0" pageNumber="1">75</emphasis>
|
||
<taxonomicName lsidName="S. putzeysi" pageId="0" pageNumber="1" rank="species" species="putzeysi">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. putzeysi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<normalizedToken originalValue="Oberthür">Oberthuer</normalizedToken>
|
||
)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">76</emphasis>
|
||
<taxonomicName lsidName="S. matsumotoi" pageId="0" pageNumber="1" rank="species" species="matsumotoi">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. matsumotoi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |