681 lines
88 KiB
XML
681 lines
88 KiB
XML
<document id="6DE3BA53A33D8437ECAA81BCB47AE3D8" ID-CLB-Dataset="46773" ID-DOI="10.11646/zootaxa.4078.1.6" ID-GBIF-Dataset="598cb955-c4b7-4c2b-8c74-6ee0348c7f7e" ID-ISSN="1175-5326" ID-Zenodo-Dep="270853" ID-ZooBank="3D0D79C2-2583-47F4-BC01-5CF347614B44" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460362976381" checkinUser="plazi" docAuthor="Pinto, Ângelo Parise & Almeida, Marcus Vinícius Oliveira De" docDate="2016" docId="495E274CFFDAFF91FF43FA9CECE2F945" docLanguage="en" docName="zootaxa.4078.1.6.pdf" docOrigin="Zootaxa 4078 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tibiagomphus noval Rodrigues Capitulo 1985" docType="treatment" docVersion="8" masterDocId="B5675F34FFC9FF87FFD4FFFCE97EFD39" masterDocTitle="A taxonomic synopsis of South American Cyanogomphini Carle with description of Cyanogomphus angelomachadoi sp. nov. from the Cerrado of Brazil (Odonata: Gomphidae)" updateTime="1698294756334" updateUser="ExternalLinkService">
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<mods:title id="0E5FF0B17505271923F1D80C0BD56A5B">A taxonomic synopsis of South American Cyanogomphini Carle with description of Cyanogomphus angelomachadoi sp. nov. from the Cerrado of Brazil (Odonata: Gomphidae)</mods:title>
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<mods:namePart id="7BE941831FF3036D52DC8733C3FFF6F8">Pinto, Ângelo Parise</mods:namePart>
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<mods:namePart id="5570DC7708990364CCDB9D95EA836B2B">Almeida, Marcus Vinícius Oliveira De</mods:namePart>
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<mods:title id="1D72773C8BE51BB82312D3C7B7CA7086">Zootaxa</mods:title>
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<treatment id="495E274CFFDAFF91FF43FA9CECE2F945" ID-DOI="http://doi.org/10.5281/zenodo.6077755" ID-GBIF-Taxon="119410161" ID-Zenodo-Dep="6077755" LSID="urn:lsid:plazi:treatment:495E274CFFDAFF91FF43FA9CECE2F945" httpUri="http://treatment.plazi.org/id/495E274CFFDAFF91FF43FA9CECE2F945" lastPageId="22" pageId="19">
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<paragraph id="C148965AFFDAFF94FF43FA9CEB99F843" blockId="19.[151,983,1376,1467]" box="[151,743,1376,1402]" pageId="19">
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<heading id="9A002136FFDAFF94FF43FA9CEB99F843" bold="true" box="[151,743,1376,1402]" fontSize="11" level="1" pageId="19" reason="1">
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<taxonomicName id="06F7EDD9FFDAFF94FF43FA9CEB99F843" ID-CoL="7BYMM" authority="Rodrigues Capitulo, 1985" authorityName="Rodrigues Capitulo" authorityYear="1985" box="[151,743,1376,1402]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="noval">
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<emphasis id="F3834A48FFDAFF94FF43FA9CEB99F843" bold="true" box="[151,743,1376,1402]" pageId="19">
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<emphasis id="F3834A48FFDAFF94FF43FA9CE8F0F843" bold="true" box="[151,398,1376,1402]" italics="true" pageId="19">Tibiagomphus noval</emphasis>
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(Rodrigues Capítulo, 1985)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="C148965AFFDAFF94FF43FA7EEABCF882" blockId="19.[151,983,1376,1467]" pageId="19">
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LSID urn:lsid:zoobank.org:act:645A61F2-F59F-42A7-8B73-4270D67477AB (
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<figureCitation id="59CC8ADFFFDAFF94FF4AFA5EE874F882" box="[158,266,1442,1467]" captionStart="FIGURE 2" captionStartId="6.[151,250,1639,1661]" captionTargetBox="[151,1436,193,1617]" captionTargetId="figure@6.[151,1436,193,1618]" captionTargetPageId="6" captionText="FIGURE 2. Wings of Cyanogomphini in ventral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449), B. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448); C. Cyanogomphus comparabilis Belle, 1994, Holotype ♂ (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, ♀ (Brazil. RJ: Reserva Ecológica de Guapiaçu, DZRJ 2495); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), ♀ (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), ♂ (Brazil. SC: Nova Teutônia village, MZSP)." httpUri="https://zenodo.org/record/270855/files/figure.png" pageId="19">Figures 2</figureCitation>
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E, 3C, 4E, 5A–B, F–H, 6F–G, 7J–L, 8G–H, 9E, 10E, 11D, 13)
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</paragraph>
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</subSubSection>
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<subSubSection id="89EDC5D1FFDAFF93FF43FA16EB00FCC6" lastPageId="20" pageId="19" type="reference_group">
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<paragraph id="C148965AFFDAFF94FF43FA16EDE2FB81" blockId="19.[151,1436,1514,2027]" pageId="19">
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<emphasis id="F3834A48FFDAFF94FF43FA16E8C3FB39" box="[151,445,1514,1536]" italics="true" pageId="19">
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<taxonomicName id="06F7EDD9FFDAFF94FF43FA16E8F0FB39" ID-CoL="9WYQS" box="[151,398,1514,1536]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="uncatus">Cyanogomphus uncatus</taxonomicName>
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<emphasis id="F3834A48FFDAFF94FE4DFA17E8C3FB39" bold="true" box="[409,445,1515,1536]" italics="true" pageId="19">nec</emphasis>
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</emphasis>
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<bibRefCitation id="A566EBABFFDAFF94FE1CFA17EB33FB39" author="Fraser" box="[456,589,1514,1536]" pageId="19" refString="Fraser, F. C. (1947) The Odonata of the Argentine Republic I. Acta Zoologica Lilloana del Instituto Miguel Lillo, 4, 427 - 462." type="journal article" year="1947">Fraser, 1947</bibRefCitation>
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:—St.
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<bibRefCitation id="A566EBABFFDAFF94FD45FA16EAFDFB26" author="Quentin" pageId="19" refString="St. Quentin, D. (1973) Die Gomphidenfauna Sudamerikas (Ordn.: Odonata). Annalen des Naturhistorischen Museums in Wien, 77, 335 - 363." type="journal article" year="1973">
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Quentin (1973: 358, record from Rio Grande do Sul State,
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<collectingCountry id="B9E0D6CAFFDAFF94FAC7FA16EC2DFB39" box="[1299,1363,1514,1536]" name="Brazil" pageId="19">Brazil</collectingCountry>
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, likely pertains to this species; comparison with the
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<typeStatus id="1E4C28F8FFDAFF94FD43F9F5EB8EFB26" box="[663,752,1545,1567]" pageId="19" type="holotype">holotype</typeStatus>
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of
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<taxonomicName id="06F7EDD9FFDAFF94FCC0F9F6EA03FB26" ID-CoL="9WYQS" box="[788,893,1546,1567]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="uncatus">
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<emphasis id="F3834A48FFDAFF94FCC0F9F6EA03FB26" box="[788,893,1546,1567]" italics="true" pageId="19">T. uncatus</emphasis>
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</taxonomicName>
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)
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</bibRefCitation>
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;—
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<bibRefCitation id="A566EBABFFDAFF94FC77F9F5EB64FBA3" author="Belle" pageId="19" refString="Belle, J. (1980) Notes on three species of Cyanogomphus Selys, 1873 (Odonata: Gomphidae). Entomologische Berichten, 40 (10), 151 - 155." type="journal article" year="1980">
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Belle (1980: 153–155, figs 5–13, in part, records from Entre Rios Province,
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<collectingCountry id="B9E0D6CAFFDAFF94FE0DF9D4EB3FFB07" box="[473,577,1576,1598]" name="Argentina" pageId="19">Argentina</collectingCountry>
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, and Florida and Durazno Departments,
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<collectingCountry id="B9E0D6CAFFDAFF94FC34F9D5ED47FB07" box="[992,1081,1577,1598]" name="Uruguay" pageId="19">Uruguay</collectingCountry>
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, illustrations of metathoracic tibia, tarsi and pretarsal claws in lateral view,
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<date id="B549B09AFFDAFF94FDB3F9BAEBE1FB65" box="[615,671,1606,1628]" pageId="19">
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V
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<quantity id="060F3BBFFFDAFF94FDAFF9BAEBE1FB65" box="[635,671,1606,1628]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="10.16" metricValueMin="2.54" pageId="19" unit="in" value="2.5" valueMax="4.0" valueMin="1.0">1–4</quantity>
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</date>
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in lateral view,
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<date id="B549B09AFFDAFF94FC9CF9BAEA18FB65" box="[840,870,1606,1628]" pageId="19">
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V
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<quantity id="060F3BBFFFDAFF94FC8EF9BAEA18FB65" box="[858,870,1606,1628]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="19" unit="in" value="4.0">4</quantity>
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</date>
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in ventral view, ligula ‘penis guard’ in anteroposterior and lateral views, anterior hamules in ventral view and secondary genitalia in lateral view, comparison with male
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<typeStatus id="1E4C28F8FFDAFF94FA97F999ECE2FB42" box="[1347,1436,1637,1659]" pageId="19" type="holotype">holotype</typeStatus>
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and female and with
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<taxonomicName id="06F7EDD9FFDAFF94FE74F978EB6CFBA3" ID-CoL="32NMT" box="[416,530,1668,1690]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="waltheri">
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<emphasis id="F3834A48FFDAFF94FE74F978EB6CFBA3" box="[416,530,1668,1690]" italics="true" pageId="19">C. waltheri</emphasis>
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</taxonomicName>
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||
)
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</bibRefCitation>
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;—
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<bibRefCitation id="A566EBABFFDAFF94FDEEF978EAF8FBA3" author="Belle" box="[570,902,1668,1690]" pageId="19" refString="Belle, J. (1981) An annotated list of the Gomphidae of Uruguay (Odonata). Revista de la Facultad de Humanidades y Ciencias, Serie Ciencias Biologicas, 1 (16), 261 - 264." type="journal article" year="1981">Belle (1981: 261–262, mention)</bibRefCitation>
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||
;—
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<bibRefCitation id="A566EBABFFDAFF94FC70F978ED9AFBA3" author="De" box="[932,1252,1668,1690]" pageId="19" refString="De Abenante, Y. P. & Philippi, M. E. (1982) Lista preliminar de los odonatos del Uruguay. Notulae Odonatologicae, 1 (9), 151." type="journal article" year="1982">
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<collectingCountry id="B9E0D6CAFFDAFF94FC70F978EABCFBA3" box="[932,962,1668,1690]" name="South Georgia and the South Sandwich Islands" pageId="19">De</collectingCountry>
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||
<bibRefCitation id="A566EBABFFDAFF94FC1EF978ED9AFBA3" author="Abenante" box="[970,1252,1668,1690]" pageId="19" refString="De Abenante, Y. P. & Philippi, M. E. (1982) Lista preliminar de los odonatos del Uruguay. Notulae Odonatologicae, 1 (9), 151." type="journal article" year="1982">Abenante & Philippi (1982</bibRefCitation>
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</bibRefCitation>
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: 151, record from
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<collectingCountry id="B9E0D6CAFFDAFF94FF13F95EE85FFB81" box="[199,289,1698,1720]" name="Uruguay" pageId="19">Uruguay</collectingCountry>
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);—
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<bibRefCitation id="A566EBABFFDAFF94FE92F95EEB2DFB81" author="Rodrigues" box="[326,595,1698,1720]" pageId="19" refString="Rodrigues Capitulo, A. (1992) Los Odonata de la Republica Argentina (Insecta). Fauna de agua dulce de la Republica Argentina, 34 (1), 1 - 91." type="journal article" year="1992">
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Rodrigues
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<bibRefCitation id="A566EBABFFDAFF94FE62F95EEB2DFB81" author="Capitulo" box="[438,595,1698,1720]" pageId="19" refString="Rodrigues Capitulo, A. (1992) Los Odonata de la Republica Argentina (Insecta). Fauna de agua dulce de la Republica Argentina, 34 (1), 1 - 91." type="journal article" year="1992">Capítulo (1992</bibRefCitation>
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</bibRefCitation>
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: 58, in part, record from Entre Rios Province,
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<collectingCountry id="B9E0D6CAFFDAFF94FBF3F95EEDEFFB81" box="[1063,1169,1698,1720]" name="Argentina" pageId="19">Argentina</collectingCountry>
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||
).
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</paragraph>
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<paragraph id="C148965AFFDAFF94FF43F93DEAAAFA97" blockId="19.[151,1436,1514,2027]" pageId="19">
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||
<taxonomicName id="06F7EDD9FFDAFF94FF43F93DEBC0FBEE" ID-CoL="9WYQR" authority="Rodrigues Capitulo, 1985: 329" authorityName="Rodrigues Capitulo" authorityPageNumber="329" authorityYear="1985" box="[151,702,1729,1751]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="noval">
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||
<emphasis id="F3834A48FFDAFF94FF43F93DE80BFBEE" box="[151,373,1729,1751]" italics="true" pageId="19">Cyanogomphus noval</emphasis>
|
||
Rodrigues Capítulo, 1985: 329
|
||
</taxonomicName>
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–335, figs 2–14 (description of ♂
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||
<typeStatus id="1E4C28F8FFDAFF94FBF0F93DED01FBEE" box="[1060,1151,1729,1751]" pageId="19" type="holotype">holotype</typeStatus>
|
||
,
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||
<collectingCountry id="B9E0D6CAFFDAFF94FB5FF93DEC5CFBEE" box="[1163,1314,1729,1751]" name="Argentina" pageId="19">ARGENTINA</collectingCountry>
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||
. Entre Rios Province: Colón Department, Parque Nacional El Palmar,
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<date id="B549B09AFFDAFF94FCC6F91CEA16FBCF" box="[786,872,1760,1782]" pageId="19" value="1982-11">XI.1982</date>
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, Rodrigues Capítulo leg. in ILPLA [sic MLP], ♀ and F- 0 larva, illustrations of postfrons in dorsal view, secondary genitalia in lateral and ventral views, caudal appendages in dorsal, lateral and ventral views, S
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||
<quantity id="060F3BBFFFDAFF94FDF7F8E1EB29FA0A" box="[547,599,1821,1843]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="19" unit="in" value="10.0">10 in</quantity>
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||
dorsal view of ♂
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||
<typeStatus id="1E4C28F8FFDAFF94FCD9F8E1EA16FA0A" box="[781,872,1821,1843]" pageId="19" type="holotype">holotype</typeStatus>
|
||
, subgenital plate and caudal appendages of ♀ in ventral view, habitus of F-0 larvae in dorsal view, head and labium in dorsal view, caudal appendages in dorsolateral view, dorsal margin of abdomen in lateral view comparison with
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||
<taxonomicName id="06F7EDD9FFDAFF94FD27F8A7EA23FA49" ID-CoL="9WYQS" box="[755,861,1883,1904]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDAFF94FD27F8A7EA23FA49" box="[755,861,1883,1904]" italics="true" pageId="19">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="06F7EDD9FFDAFF94FC41F8A7EA8DFA49" ID-CoL="B7Q8" box="[917,1011,1883,1904]" class="Insecta" family="Gomphidae" genus="Agriogomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="ericae">
|
||
<emphasis id="F3834A48FFDAFF94FC41F8A7EA8DFA49" box="[917,1011,1883,1904]" italics="true" pageId="19">A. ericae</emphasis>
|
||
</taxonomicName>
|
||
);—
|
||
<bibRefCitation id="A566EBABFFDAFF94FBCEF8A6EC03FA49" author="Santos" box="[1050,1405,1882,1904]" pageId="19" refString="Santos, N. D. (1988) Catalogo Bibliografico de Ninfas de odonatos neotropicais. (Acompanhado de relacao alfabetica de autores e seus trabalhos). Acta Amazonica, 18 (1 - 2), 265 - 350." type="journal article" year="1988">Santos (1988: 267, 286, mention)</bibRefCitation>
|
||
;—
|
||
<bibRefCitation id="A566EBABFFDAFF94FF13F885E8A5FAB6" author="Rodrigues" box="[199,475,1913,1935]" pageId="19" refString="Rodrigues Capitulo, A. (1992) Los Odonata de la Republica Argentina (Insecta). Fauna de agua dulce de la Republica Argentina, 34 (1), 1 - 91." type="journal article" year="1992">Rodrigues Capítulo (1992</bibRefCitation>
|
||
: 58, 67, 81, 83, 89, figs 1–2, 4–5, 110, 132, 134, 180–181, distribution, reproduction of illustrations from Rodrigues Capítulo 1985);—
|
||
<bibRefCitation id="A566EBABFFDAFF94FD4BF864EAAEFA97" author="Garrison" box="[671,976,1944,1966]" pageId="19" refString="Garrison, R. W. (1991) A synonymic list of the New World Odonata. Argia, 3 (2), 1 - 30." type="journal article" year="1991">Garrison (1991: 17, mention)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="C148965AFFDAFF93FF43F84AEA4EFC33" blockId="19.[151,1436,1514,2027]" lastBlockId="20.[151,1436,152,511]" lastPageId="20" pageId="19">
|
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<taxonomicName id="06F7EDD9FFDAFF94FF43F84AEB01FAF5" ID-CoL="7BYMM" authority="Rodrigues Capitulo, 1985" authorityName="Rodrigues Capitulo" authorityYear="1985" box="[151,639,1974,1996]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="noval">
|
||
<emphasis id="F3834A48FFDAFF94FF43F84AE81AFAF5" box="[151,356,1974,1996]" italics="true" pageId="19">Tibiagomphus noval</emphasis>
|
||
(Rodrigues Capítulo, 1985)
|
||
</taxonomicName>
|
||
:—
|
||
<bibRefCitation id="A566EBABFFDAFF94FD4AF84AEA58FAF5" author="Belle" box="[670,806,1974,1996]" pageId="19" refString="Belle, J. (1992 a) Studies on ultimate instar larvae of Neotropical Gomphidae, with the description of Tibiagomphus gen. nov. (Anisoptera). Odonatologica, 21 (1), 1 - 24." type="journal article" year="1992" yearSuffix="a">Belle (1992a</bibRefCitation>
|
||
: 1, comb. nov., implicit by the context ICZN 1999, Art. 11.9.3 of the Code);—
|
||
<bibRefCitation id="A566EBABFFDAFF94FEB9F829EBF0FAD2" author="Belle" box="[365,654,2005,2027]" pageId="19" refString="Belle, J. (1993) Annotated checklist and bibliography of the immature stages of Neotropical Gomphidae, published up to September 1993 (Anisoptera). Odonatologica, 22 (4), 399 - 409." type="journal article" year="1993">Belle (1993: 406, mention)</bibRefCitation>
|
||
;—
|
||
<bibRefCitation id="A566EBABFFDAFF94FD78F829ED15FAD2" author="Bridges" box="[684,1131,2005,2027]" pageId="19" refString="Bridges, C. A. (1994) Catalogue of the family-group, genus-group and species-group names of the Odonata of the world (Third Edition). Privately published, Urbana, Illinois, 905 pp." type="book" year="1994">
|
||
Bridges (1994: VII.168,
|
||
<date id="B549B09AFFDAFF94FC67F829ED7EFAD2" box="[947,1024,2005,2027]" pageId="19" value="1970-08">VIII.70</date>
|
||
, mention)
|
||
</bibRefCitation>
|
||
;—Muzón & von
|
||
<bibRefCitation id="A566EBABFFDAFF93FAFFF829EB70FD97" author="Ellenrieder" lastPageId="20" pageId="19" refString="Muzon, J. & von Ellenrieder, N. (1999) Status and distribution of Odonata (Insecta) within natural protected areas in Argentina. Biogeographica, 75 (3), 119 - 128." type="journal article" year="1999">Ellenrieder (1999: 125, distributional data)</bibRefCitation>
|
||
;—
|
||
<bibRefCitation id="A566EBABFFDDFF93FDF9FF64EA6DFD97" author="Garrison" box="[557,787,152,174]" pageId="20" refString="Garrison, R. W., von Ellenrieder, N. & Louton, J. A. (2006) Dragonfly genera of the New World, an illustrated and annotated key to the Anisoptera. The Johns Hopkins University Press, Baltimore, xiv + 368 pp." type="book" year="2006">
|
||
Garrison
|
||
<emphasis id="F3834A48FFDDFF93FD46FF65EBB4FD97" box="[658,714,152,174]" italics="true" pageId="20">et al.</emphasis>
|
||
(2006
|
||
</bibRefCitation>
|
||
: 129–130, comparison, suggest synonym with
|
||
<taxonomicName id="06F7EDD9FFDDFF93FAD3FF65EC0DFD97" ID-CoL="9WYQS" box="[1287,1395,153,174]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDDFF93FAD3FF65EC0DFD97" box="[1287,1395,153,174]" italics="true" pageId="20">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
);—
|
||
<bibRefCitation id="A566EBABFFDDFF93FF13FF4AE813FDF5" author="Heckman" box="[199,365,182,204]" pageId="20" refString="Heckman, C. W. (2006) Encyclopedia of South American aquatic insects: Odonata Anisoptera. Illustrated keys to known families, genera, and species in South America. Springer, Dordrecht, viii + 725 pp." type="book" year="2006">Heckman (2006</bibRefCitation>
|
||
: 525, 534, 649, figs 3.2.589, 3.2.603, key, reproduction of illustrations from Rodrigues Capítulo 1985);— von
|
||
<bibRefCitation id="A566EBABFFDDFF93FF20FF29ED70FDD2" author="Ellenrieder" box="[244,1038,213,235]" pageId="20" refString="von Ellenrieder, N. & Muzon, J. (2008) An updated checklist of the Odonata from Argentina. Odonatologica, 37 (1), 55 - 68." type="journal article" year="2008">
|
||
Ellenrieder & Muzón (2008: 61, record from Entre Rios Province,
|
||
<collectingCountry id="B9E0D6CAFFDDFF93FC4BFF29ED79FDD2" box="[927,1031,213,235]" name="Argentina" pageId="20">Argentina</collectingCountry>
|
||
)
|
||
</bibRefCitation>
|
||
;—Muzón
|
||
<emphasis id="F3834A48FFDDFF93FBA8FF2AEDCEFDD2" box="[1148,1200,213,235]" italics="true" pageId="20">et al.</emphasis>
|
||
(2007: 302, 305, notes on the
|
||
<typeStatus id="1E4C28F8FFDDFF93FED8FF09E846FC33" box="[268,312,245,266]" pageId="20">type</typeStatus>
|
||
material);—
|
||
<bibRefCitation id="A566EBABFFDDFF93FE63FF08EA52FC33" author="Hamalainen" box="[439,812,244,266]" pageId="20" refString="Hamalainen, M. (2015) Catalogue of individuals commemorated in the scientific names of extant dragonflies, including lists of all available eponymous species-group and genus-group names. International Dragonfly Fund Report, 80, 1 - 168." type="journal article" year="2015">Hämäläinen (2015: 114, etymology)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="C148965AFFDDFF93FF43FEEEEB00FCC6" blockId="20.[151,1436,152,511]" pageId="20">
|
||
<emphasis id="F3834A48FFDDFF93FF43FEEEE8D7FC11" box="[151,425,274,296]" italics="true" pageId="20">
|
||
<taxonomicName id="06F7EDD9FFDDFF93FF43FEEEE803FC11" ID-CoL="7BYMN" box="[151,381,274,296]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="uncatus">Tibiagomphus uncatus</taxonomicName>
|
||
<emphasis id="F3834A48FFDDFF93FE51FEEFE8D7FC11" bold="true" box="[389,425,275,296]" italics="true" pageId="20">nec</emphasis>
|
||
</emphasis>
|
||
(
|
||
<bibRefCitation id="A566EBABFFDDFF93FE6DFEEEEB44FC11" author="Fraser" box="[441,570,274,296]" pageId="20" refString="Fraser, F. C. (1947) The Odonata of the Argentine Republic I. Acta Zoologica Lilloana del Instituto Miguel Lillo, 4, 427 - 462." type="journal article" year="1947">Fraser, 1947</bibRefCitation>
|
||
):—
|
||
<bibRefCitation id="A566EBABFFDDFF93FDB4FEEEEC64FC11" author="Belle" box="[608,1306,274,296]" pageId="20" refString="Belle, J. (1996) Higher classification of the South-American Gomphidae (Odonata). Zoologische Mededelingen, 70, 297 - 324." type="journal article" year="1996">
|
||
Belle (1996: 323, fig. 8, in part, pair of wings of ♂ from
|
||
<collectingCountry id="B9E0D6CAFFDDFF93FB6DFEEEEC6DFC11" box="[1209,1299,274,296]" name="Uruguay" pageId="20">Uruguay</collectingCountry>
|
||
)
|
||
</bibRefCitation>
|
||
;—
|
||
<bibRefCitation id="A566EBABFFDDFF93FAECFEEEE878FC7E" author="Heckman" pageId="20" refString="Heckman, C. W. (2006) Encyclopedia of South American aquatic insects: Odonata Anisoptera. Illustrated keys to known families, genera, and species in South America. Springer, Dordrecht, viii + 725 pp." type="book" year="2006">Heckman (2006</bibRefCitation>
|
||
: 649–650, fig. 3.2.778, in part, reproduction of illustrations from
|
||
<bibRefCitation id="A566EBABFFDDFF93FC1DFECDED43FC7E" author="Belle" box="[969,1085,305,327]" pageId="20" refString="Belle, J. (1980) Notes on three species of Cyanogomphus Selys, 1873 (Odonata: Gomphidae). Entomologische Berichten, 40 (10), 151 - 155." type="journal article" year="1980">Belle 1980</bibRefCitation>
|
||
of ♂ from Entre Rios Province,
|
||
<collectingCountry id="B9E0D6CAFFDDFF93FF13FEACE851FC5F" box="[199,303,336,358]" name="Argentina" pageId="20">Argentina</collectingCountry>
|
||
, and Florida Department,
|
||
<collectingCountry id="B9E0D6CAFFDDFF93FD93FEACEBDFFC5F" box="[583,673,336,358]" name="Uruguay" pageId="20">Uruguay</collectingCountry>
|
||
);—
|
||
<bibRefCitation id="A566EBABFFDDFF93FD12FEACEAD5FC5F" author="Garrison" box="[710,939,336,358]" pageId="20" refString="Garrison, R. W., von Ellenrieder, N. & Louton, J. A. (2006) Dragonfly genera of the New World, an illustrated and annotated key to the Anisoptera. The Johns Hopkins University Press, Baltimore, xiv + 368 pp." type="book" year="2006">
|
||
Garrison
|
||
<emphasis id="F3834A48FFDDFF93FCFFFEADEA1CFC5F" box="[811,866,336,358]" italics="true" pageId="20">et al.</emphasis>
|
||
(2006
|
||
</bibRefCitation>
|
||
: 71, 81–83, 129–130, 329, 331, 337, figs 727, 729a,b, 730, 732, in part, illustrations of mesepisternum in dorsal view and subgenital plate in ventral view of ♀ from Entre Rios Province,
|
||
<collectingCountry id="B9E0D6CAFFDDFF93FE74FE71EB76FC9A" box="[416,520,397,419]" name="Argentina" pageId="20">Argentina</collectingCountry>
|
||
, metathoracic tibia, and secondary genitalia in lateral view from ♂ of Florida Department,
|
||
<collectingCountry id="B9E0D6CAFFDDFF93FF13FE51E85CFCFB" box="[199,290,429,450]" name="Uruguay" pageId="20">Uruguay</collectingCountry>
|
||
);—von
|
||
<bibRefCitation id="A566EBABFFDDFF93FEA1FE50EDB8FCFB" author="Ellenrieder" box="[373,1222,428,450]" pageId="20" refString="von Ellenrieder, N. & Muzon, J. (2008) An updated checklist of the Odonata from Argentina. Odonatologica, 37 (1), 55 - 68." type="journal article" year="2008">
|
||
Ellenrieder & Muzón (2008: 61, in part, record to Entre Rios Province,
|
||
<collectingCountry id="B9E0D6CAFFDDFF93FB83FE50EDC1FCFB" box="[1111,1215,428,450]" name="Argentina" pageId="20">Argentina</collectingCountry>
|
||
)
|
||
</bibRefCitation>
|
||
;—von Ellenrieder
|
||
<emphasis id="F3834A48FFDDFF93FA5EFE51E99CFCD9" italics="true" pageId="20">et al.</emphasis>
|
||
(2009: 229, record from
|
||
<collectingCountry id="B9E0D6CAFFDDFF93FE35FE37EB42FCD9" box="[481,572,459,480]" name="Uruguay" pageId="20">Uruguay</collectingCountry>
|
||
);—von
|
||
<bibRefCitation id="A566EBABFFDDFF93FD5BFE36EAC3FCD9" author="Ellenrieder" box="[655,957,458,480]" pageId="20" refString="von Ellenrieder, N. & Garrison, R. W. (2009) Odonata. In: Dominguez, E. & Fernandez, H. R. (Eds.), Macroinvertebrados bentonicos sudamericanos. Sistematica y biologia. Fundacion Miguel Lillo, Tucuman, pp. 95 - 143." type="book chapter" year="2009">Ellenrieder & Garrison (2009</bibRefCitation>
|
||
: 123, fig. 275, reproduction of illustration of ♀ subgenital plate from
|
||
<bibRefCitation id="A566EBABFFDDFF93FE70FE15EB0DFCC6" author="Garrison" box="[420,627,489,511]" pageId="20" refString="Garrison, R. W., von Ellenrieder, N. & Louton, J. A. (2006) Dragonfly genera of the New World, an illustrated and annotated key to the Anisoptera. The Johns Hopkins University Press, Baltimore, xiv + 368 pp." type="book" year="2006">
|
||
Garrison
|
||
<emphasis id="F3834A48FFDDFF93FDD7FE16EB48FCC6" box="[515,566,489,511]" italics="true" pageId="20">et al.</emphasis>
|
||
2006
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="89EDC5D1FFDDFF93FF43FDD0ED35FE24" pageId="20" type="materials_examined">
|
||
<paragraph id="C148965AFFDDFF93FF43FDD0EA97FFED" blockId="20.[151,1437,556,2021]" pageId="20">
|
||
<emphasis id="F3834A48FFDDFF93FF43FDD0E88CFF7C" bold="true" box="[151,498,556,581]" pageId="20">
|
||
Material examined (
|
||
<specimenCount id="D7F15DD3FFDDFF93FE58FDD0E8CEFF7C" box="[396,432,556,581]" pageId="20" type="generic">8♂</specimenCount>
|
||
, 1♀).
|
||
</emphasis>
|
||
<collectingCountry id="B9E0D6CAFFDDFF93FE2EFDD0EB1DFF7D" box="[506,611,556,580]" name="Brazil" pageId="20">BRAZIL</collectingCountry>
|
||
. Rio Grande do Sul State:
|
||
<specimenCount id="D7F15DD3FFDDFF93FC40FDD0EAC6FF7D" box="[916,952,556,580]" pageId="20" type="generic">1♂</specimenCount>
|
||
, [Guaíba municipality], Arroio [stream] at BR 116 Km 310 [
|
||
<geoCoordinate id="A4C3F09DFFDDFF93FEB6FDB3EB73FF51" box="[354,525,591,616]" direction="south" orientation="latitude" pageId="20" precision="1" value="-30.195915">30°11’45.30”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A4C3F09DFFDDFF93FDCEFDB3EBB2FF51" box="[538,716,591,616]" direction="west" orientation="longitude" pageId="20" precision="1" value="-51.399048">51°23’56.56”W</geoCoordinate>
|
||
,
|
||
<quantity id="060F3BBFFFDDFF93FD0EFDACEA6AFF5E" box="[730,788,592,616]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="20" unit="m" value="20.0">20 m</quantity>
|
||
a.s.l.],
|
||
<date id="B549B09AFFDDFF93FCBDFDACEAE4FF51" box="[873,922,592,616]" pageId="20">17.I.</date>
|
||
[19]79, J.M. Costa leg. (
|
||
<collectionCode id="A7E60E9FFFDDFF93FB67FDACEC7AFF51" box="[1203,1284,592,616]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="20">MNRJ</collectionCode>
|
||
);
|
||
<specimenCount id="D7F15DD3FFDDFF93FACDFDACEC43FF51" box="[1305,1341,592,616]" pageId="20" type="generic">1♂</specimenCount>
|
||
, Pelotas municipality [
|
||
<geoCoordinate id="A4C3F09DFFDDFF93FEE2FD88E89CFFB4" box="[310,482,628,653]" direction="south" orientation="latitude" pageId="20" precision="1" value="-31.772">31°46’19.20”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A4C3F09DFFDDFF93FE24FD88EBDDFFB4" box="[496,675,628,653]" direction="west" orientation="longitude" pageId="20" precision="1" value="-52.3428">52°20’34.08”W</geoCoordinate>
|
||
,
|
||
<quantity id="060F3BBFFFDDFF93FD65FD88EB93FFB5" box="[689,749,628,652]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="20" unit="m" value="14.0">14 m</quantity>
|
||
a.s.l.],
|
||
<date id="B549B09AFFDDFF93FC92FD88EAF1FFB4" box="[838,911,628,653]" pageId="20">09.III.</date>
|
||
[19]51, C. Biezanko leg. (
|
||
<collectionCode id="A7E60E9FFFDDFF93FB6EFD88EC78FFB5" box="[1210,1286,628,652]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="20">MNRJ</collectionCode>
|
||
22082);
|
||
<specimenCount id="D7F15DD3FFDDFF93FAA6FD88ECE9FFB4" box="[1394,1431,628,653]" pageId="20" type="generic">6♂</specimenCount>
|
||
, Jaguarão municipality [
|
||
<geoCoordinate id="A4C3F09DFFDDFF93FE4AFD6BEB34FF89" box="[414,586,663,688]" direction="south" orientation="latitude" pageId="20" precision="1" value="-32.56585">32°33’57.06”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A4C3F09DFFDDFF93FD82FD6BEA77FF89" box="[598,777,663,688]" direction="west" orientation="longitude" pageId="20" precision="1" value="-53.376">53°22’33.60”W</geoCoordinate>
|
||
,
|
||
<quantity id="060F3BBFFFDDFF93FCC0FD64EA30FF96" box="[788,846,664,688]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="20" unit="m" value="19.0">19 m</quantity>
|
||
a.s.l.], Stream at
|
||
<quantity id="060F3BBFFFDDFF93FBC7FD6BED24FF89" box="[1043,1114,663,688]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="20" unit="km" value="15.0">15 km</quantity>
|
||
from highway,
|
||
<date id="B549B09AFFDDFF93FAD8FD64ECE6FF89" box="[1292,1432,664,688]" pageId="20" value="1978-12-12">12.XII.1978</date>
|
||
, J.M. Costa leg. (
|
||
<collectionCode id="A7E60E9FFFDDFF93FE9AFD40E8E1FFED" box="[334,415,700,724]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="20">MNRJ</collectionCode>
|
||
); 1♀, same data but 12.XIII [sic XII].1978 (
|
||
<collectionCode id="A7E60E9FFFDDFF93FC58FD40EAA3FFED" box="[908,989,700,724]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="20">MNRJ</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C148965AFFDDFF93FF13FD23ED35FE24" blockId="20.[151,1437,556,2021]" pageId="20">
|
||
<emphasis id="F3834A48FFDDFF93FF13FD23E8F1FFC1" bold="true" box="[199,399,735,760]" pageId="20">
|
||
<typeStatus id="1E4C28F8FFDDFF93FF13FD23E87CFFC1" box="[199,258,735,760]" pageId="20">Type</typeStatus>
|
||
repository.
|
||
</emphasis>
|
||
<typeStatus id="1E4C28F8FFDDFF93FE48FD23EB7AFFC1" box="[412,516,735,760]" pageId="20" type="holotype">Holotype</typeStatus>
|
||
♂ by original designation (see remarks) at Museo de La Plata, Universidad Nacional de La Plata, La Plata, Buenos Aires,
|
||
<collectingCountry id="B9E0D6CAFFDDFF93FD4BFCF8EA70FE24" box="[671,782,772,797]" name="Argentina" pageId="20">Argentina</collectingCountry>
|
||
(
|
||
<collectionCode id="A7E60E9FFFDDFF93FCC9FCF8EA26FE25" box="[797,856,772,796]" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="20">MLP</collectionCode>
|
||
, examined by photo).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="89EDC5D1FFDDFF93FF13FCDBED13F9BC" pageId="20" type="description">
|
||
<paragraph id="C148965AFFDDFF93FF13FCDBED6AFEE9" blockId="20.[151,1437,556,2021]" pageId="20">
|
||
<emphasis id="F3834A48FFDDFF93FF13FCDBE803FE79" bold="true" box="[199,381,807,832]" pageId="20">Measurements.</emphasis>
|
||
Males (n = 8). Total length (including caudal appendages) 45.4–47.0; abdomen length (excluding caudal appendages) 30.8–33.9; head maximum width 5.8–6.3; Fw length 25.8–30.5; Hw length 24.7–26.3; Fw maximum width 4.9–5.5, in Hw 6.1–7.0; pt length
|
||
<quantity id="060F3BBFFFDDFF93FC63FC93ED50FEB1" box="[951,1070,879,904]" metricMagnitude="-2" metricUnit="m" metricValue="7.493" metricValueMax="8.382" metricValueMin="6.604" pageId="20" unit="in" value="2.95" valueMax="3.3" valueMin="2.6">2.6–3.3 in</quantity>
|
||
Fw, 3.0–
|
||
<quantity id="060F3BBFFFDDFF93FB76FC93ED96FEB1" box="[1186,1256,879,904]" metricMagnitude="-2" metricUnit="m" metricValue="9.144" pageId="20" unit="in" value="3.6">3.6 in</quantity>
|
||
Hw; length of metathoracic femur 7.4–8.2; metathoracic tibia 5.7–6.2; length of S9+
|
||
<quantity id="060F3BBFFFDDFF93FC6AFC68EA84FE94" box="[958,1018,916,941]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="20" unit="in" value="10.0">10 in</quantity>
|
||
lateral view 3.2–3.8; total length of cercus in lateral view 1.6–1.9. Ratio cercus length / maximum width 0.30–0.36.
|
||
</paragraph>
|
||
<paragraph id="C148965AFFDDFF93FF13FC20EBB4F959" blockId="20.[151,1437,556,2021]" pageId="20">
|
||
Females (n = 1). Total length (including caudal appendages) 45.2; abdomen length (excluding caudal appendages) 31.8; head maximum width 6.3; Fw length 28.0; Hw length 27.1; Fw maximum width 5.6, in Hw 7.3; pt length
|
||
<quantity id="060F3BBFFFDDFF93FED5FBD8E83CF904" box="[257,322,1060,1085]" metricMagnitude="-2" metricUnit="m" metricValue="7.874" pageId="20" unit="in" value="3.1">3.1 in</quantity>
|
||
Fw,
|
||
<quantity id="060F3BBFFFDDFF93FEADFBD8E8C4F904" box="[377,442,1060,1085]" metricMagnitude="-2" metricUnit="m" metricValue="8.382" pageId="20" unit="in" value="3.3">3.3 in</quantity>
|
||
Hw; length of metathoracic femur 8.1; metathoracic tibia 6.2; length of S9+
|
||
<quantity id="060F3BBFFFDDFF93FAC1FBD8EC31F904" box="[1301,1359,1060,1085]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="20" unit="in" value="10.0">10 in</quantity>
|
||
lateral view 2.7; total length of cercus in lateral view 1.3.
|
||
</paragraph>
|
||
<paragraph id="C148965AFFDDFF93FF13FB90ED13F9BC" blockId="20.[151,1437,556,2021]" box="[199,1133,1132,1157]" pageId="20">
|
||
<emphasis id="F3834A48FFDDFF93FF13FB90E869F9BC" bold="true" box="[199,279,1132,1157]" pageId="20">Larva.</emphasis>
|
||
Rodrigues Capítulo (1985, p. 332) based on specimens reared in laboratory.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="89EDC5D1FFDDFF93FF13FB73EBD8FA09" pageId="20" type="diagnosis">
|
||
<paragraph id="C148965AFFDDFF93FF13FB73EA11F8D5" blockId="20.[151,1437,556,2021]" pageId="20">
|
||
<emphasis id="F3834A48FFDDFF93FF13FB73E844F991" bold="true" box="[199,314,1167,1192]" pageId="20">Diagnosis</emphasis>
|
||
. This pale species, as well as
|
||
<taxonomicName id="06F7EDD9FFDDFF93FD71FB6DEA64F991" box="[677,794,1169,1192]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDDFF93FD71FB6DEA64F991" box="[677,794,1169,1192]" italics="true" pageId="20">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
, can be separated from
|
||
<taxonomicName id="06F7EDD9FFDDFF93FBE8FB73ED94F991" box="[1084,1258,1167,1192]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3834A48FFDDFF93FBE8FB73ED94F991" box="[1084,1258,1167,1192]" italics="true" pageId="20">Cyanogomphus</emphasis>
|
||
</taxonomicName>
|
||
spp. except
|
||
<taxonomicName id="06F7EDD9FFDDFF93FA56FB6CE855F9F4" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="comparabilis">
|
||
<emphasis id="F3834A48FFDDFF93FA56FB6CE855F9F4" italics="true" pageId="20">C. comparabilis</emphasis>
|
||
</taxonomicName>
|
||
by the mesepisternum close to the dorsal carina largely pale (dark in
|
||
<taxonomicName id="06F7EDD9FFDDFF93FBE6FB49EC73F9F4" box="[1074,1293,1204,1229]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
|
||
<emphasis id="F3834A48FFDDFF93FBE6FB49EC73F9F4" box="[1074,1293,1204,1229]" italics="true" pageId="20">C. angelomachadoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="F3834A48FFDDFF93FAC0FB48EC13F9F4" bold="true" box="[1300,1389,1204,1229]" pageId="20">
|
||
<taxonomicNameLabel id="E8B0F733FFDDFF93FAC0FB48EC13F9F4" box="[1300,1389,1204,1229]" pageId="20" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="06F7EDD9FFDDFF93FF43FB24E866F9C9" box="[151,280,1239,1264]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="waltheri">
|
||
<emphasis id="F3834A48FFDDFF93FF43FB24E866F9C9" box="[151,280,1239,1264]" italics="true" pageId="20">C. waltheri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="59CC8ADFFFDDFF93FEFDFB2BE80CF9C9" box="[297,370,1239,1264]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="20">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="59CC8ADFFFDDFF93FE57FB24E8E0F9C9" box="[387,414,1240,1264]" captionStart="FIGURE 10" captionStartId="21.[151,250,1646,1668]" captionTargetBox="[151,1436,195,1620]" captionTargetId="figure@21.[151,1436,195,1622]" captionTargetPageId="21" captionText="FIGURE 10. Male habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Paratype (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270863/files/figure.png" pageId="20">10</figureCitation>
|
||
A–B, D, 11A, C). The narrow cornet-like distal portion of
|
||
<date id="B549B09AFFDDFF93FB84FB24ED0DF9C9" box="[1104,1139,1240,1264]" pageId="20">
|
||
V
|
||
<quantity id="060F3BBFFFDDFF93FBB1FB24ED0DF9C9" box="[1125,1139,1240,1264]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="20" unit="in" value="4.0">4</quantity>
|
||
</date>
|
||
in ventral view (
|
||
<figureCitation id="59CC8ADFFFDDFF93FAEAFB2BECFDF9C9" box="[1342,1411,1239,1264]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="20">Fig. 6</figureCitation>
|
||
G, maximum width <0.6 of
|
||
<date id="B549B09AFFDDFF93FE62FB00E8A6F82D" box="[438,472,1276,1300]" pageId="20">V3</date>
|
||
width vs.>
|
||
<quantity id="060F3BBFFFDDFF93FDB1FB00EBCDF82C" box="[613,691,1276,1301]" metricMagnitude="-2" metricUnit="m" metricValue="1.905" pageId="20" unit="in" value="0.75">0.75 in</quantity>
|
||
<taxonomicName id="06F7EDD9FFDDFF93FD6EFB00EA16F82C" box="[698,872,1276,1301]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3834A48FFDDFF93FD6EFB00EA16F82C" box="[698,872,1276,1301]" italics="true" pageId="20">Cyanogomphus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="59CC8ADFFFDDFF93FCA2FB00EAC2F82C" box="[886,956,1276,1301]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="20">Figs 6</figureCitation>
|
||
B, E), blade-like cercus distinctly flattened with apex directed posteriorly (
|
||
<figureCitation id="59CC8ADFFFDDFF93FE20FAE3EB44F801" box="[500,570,1311,1336]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="20">Figs 7</figureCitation>
|
||
J–L, 8G–H, cercus cylindrical, curved toward to the upturned tapered apex in
|
||
<taxonomicName id="06F7EDD9FFDDFF93FF43FAB8E83BF864" box="[151,325,1348,1373]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3834A48FFDDFF93FF43FAB8E83BF864" box="[151,325,1348,1373]" italics="true" pageId="20">Cyanogomphus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="59CC8ADFFFDDFF93FE80FAB8E8E4F865" box="[340,410,1348,1373]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="20">Figs 7</figureCitation>
|
||
A–I, 8A–F), the dorsally smooth epiproct, lacking a distal concavity on the epiproct (
|
||
<figureCitation id="59CC8ADFFFDDFF93FABEFAB8E9DAF8B9" captionStart="FIGURE 8" captionStartId="17.[151,250,1860,1882]" captionTargetBox="[151,1435,197,1827]" captionTargetId="figure@17.[151,1436,195,1835]" captionTargetPageId="17" captionText="FIGURE 8. Male caudal appendages of Cyanogomphini in lateral view (A, C, E, G, I), and detail of epiproct in laterodorsal view (B, D, F, H, J). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato, DZRJ 2493); E – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); I – J. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270861/files/figure.png" pageId="20">Figs 8</figureCitation>
|
||
H, epiproct with a distal concavity in
|
||
<taxonomicName id="06F7EDD9FFDDFF93FD98FA9BEB84F8B9" box="[588,762,1383,1408]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3834A48FFDDFF93FD98FA9BEB84F8B9" box="[588,762,1383,1408]" italics="true" pageId="20">Cyanogomphus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="59CC8ADFFFDDFF93FCDCFA9BEA30F8B9" box="[776,846,1383,1408]" captionStart="FIGURE 8" captionStartId="17.[151,250,1860,1882]" captionTargetBox="[151,1435,197,1827]" captionTargetId="figure@17.[151,1436,195,1835]" captionTargetPageId="17" captionText="FIGURE 8. Male caudal appendages of Cyanogomphini in lateral view (A, C, E, G, I), and detail of epiproct in laterodorsal view (B, D, F, H, J). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato, DZRJ 2493); E – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); I – J. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270861/files/figure.png" pageId="20">Figs 8</figureCitation>
|
||
B, D, F) of males, and subtriangular, long subgenital plate (length> 0.5 of S9) with a narrow and deep mesal incision (
|
||
<figureCitation id="59CC8ADFFFDDFF93FCA4FA70EACCF89D" box="[880,946,1420,1445]" captionStart="FIGURE 9" captionStartId="18.[151,250,1160,1182]" captionTargetBox="[151,1434,193,1138]" captionTargetId="figure@18.[151,1436,193,1139]" captionTargetPageId="18" captionText="FIGURE 9. Female S 8 – 10 of Cyanogomphini in ventral, inset an outline of subgenital plate under camera lucida. A – B. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ, Cachoeiras de Macacu), A. Reserva Ecológica de Guapiaçu (DZRJ 2495), B. Boca do Mato (DZRJ 2494); C. Cyanogomphus angelomachadoi sp. nov. additional specimen (Brazil. MG: Serra do Salitre, ABMM); D. Cyanogomphus sp. (Brazil. GO: Jataí, ABMM); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars for photographs = 1 mm." httpUri="https://zenodo.org/record/270862/files/figure.png" pageId="20">Fig. 9</figureCitation>
|
||
E, subgenital plate short, length <0.4 of S9, not incised, with a wide V- or U-shaped posteromesal concavity in
|
||
<taxonomicName id="06F7EDD9FFDDFF93FC16FA53ED0EF8F1" box="[962,1136,1455,1480]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3834A48FFDDFF93FC16FA53ED0EF8F1" box="[962,1136,1455,1480]" italics="true" pageId="20">Cyanogomphus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="59CC8ADFFFDDFF93FB57FA53EDB3F8F1" box="[1155,1229,1455,1480]" captionStart="FIGURE 9" captionStartId="18.[151,250,1160,1182]" captionTargetBox="[151,1434,193,1138]" captionTargetId="figure@18.[151,1436,193,1139]" captionTargetPageId="18" captionText="FIGURE 9. Female S 8 – 10 of Cyanogomphini in ventral, inset an outline of subgenital plate under camera lucida. A – B. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ, Cachoeiras de Macacu), A. Reserva Ecológica de Guapiaçu (DZRJ 2495), B. Boca do Mato (DZRJ 2494); C. Cyanogomphus angelomachadoi sp. nov. additional specimen (Brazil. MG: Serra do Salitre, ABMM); D. Cyanogomphus sp. (Brazil. GO: Jataí, ABMM); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars for photographs = 1 mm." httpUri="https://zenodo.org/record/270862/files/figure.png" pageId="20">Figs 9</figureCitation>
|
||
A–D) of females, separates both species of
|
||
<taxonomicName id="06F7EDD9FFDDFF93FE67FA28EB2CF8D4" box="[435,594,1492,1517]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3834A48FFDDFF93FE67FA28EB2CF8D4" box="[435,594,1492,1517]" italics="true" pageId="20">Tibiagomphus</emphasis>
|
||
</taxonomicName>
|
||
from all
|
||
<taxonomicName id="06F7EDD9FFDDFF93FD6DFA28EA19F8D4" box="[697,871,1492,1517]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3834A48FFDDFF93FD6DFA28EA19F8D4" box="[697,871,1492,1517]" italics="true" pageId="20">Cyanogomphus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="C148965AFFDDFF93FF13FA0BEBD8FA09" blockId="20.[151,1437,556,2021]" pageId="20">
|
||
Both sexes of
|
||
<taxonomicName id="06F7EDD9FFDDFF93FEB1FA05E8C1FB29" box="[357,447,1527,1552]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="noval">
|
||
<emphasis id="F3834A48FFDDFF93FEB1FA05E8C1FB29" box="[357,447,1527,1552]" italics="true" pageId="20">T. noval</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from it sibling congener
|
||
<taxonomicName id="06F7EDD9FFDDFF93FC10FA05ED4AFB29" box="[964,1076,1529,1552]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDDFF93FC10FA05ED4AFB29" box="[964,1076,1529,1552]" italics="true" pageId="20">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
by its paler generally yellowish color (
|
||
<figureCitation id="59CC8ADFFFDDFF93FF34F9E0E848FB0D" box="[224,310,1564,1589]" captionStart="FIGURE 10" captionStartId="21.[151,250,1646,1668]" captionTargetBox="[151,1436,195,1620]" captionTargetId="figure@21.[151,1436,195,1622]" captionTargetPageId="21" captionText="FIGURE 10. Male habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Paratype (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270863/files/figure.png" pageId="20">Figs 10</figureCitation>
|
||
E, 11D, distinctly darker in
|
||
<taxonomicName id="06F7EDD9FFDDFF93FDA5F9E1EB9AFB0D" box="[625,740,1565,1588]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDDFF93FDA5F9E1EB9AFB0D" box="[625,740,1565,1588]" italics="true" pageId="20">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="59CC8ADFFFDDFF93FD27F9E0EA3AFB0D" box="[755,836,1564,1589]" captionStart="FIGURE 10" captionStartId="21.[151,250,1646,1668]" captionTargetBox="[151,1436,195,1620]" captionTargetId="figure@21.[151,1436,195,1622]" captionTargetPageId="21" captionText="FIGURE 10. Male habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Paratype (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270863/files/figure.png" pageId="20">Fig. 10</figureCitation>
|
||
F), and by dorsally pale metathoracic tibia and tarsus (
|
||
<figureCitation id="59CC8ADFFFDDFF93FF4BF9C3E99FFB61" box="[159,225,1599,1624]" captionStart="FIGURE 3" captionStartId="7.[151,250,894,916]" captionTargetBox="[151,1436,193,871]" captionTargetId="figure@7.[151,1436,193,873]" captionTargetPageId="7" captionText="FIGURE 3. Details of metathoracic tibia and tarsus of males of Cyanogomphini in lateral view. A. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); C. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); D. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270856/files/figure.png" pageId="20">Fig. 3</figureCitation>
|
||
C, dark in
|
||
<taxonomicName id="06F7EDD9FFDDFF93FE82F9BDE8B6FB61" box="[342,456,1601,1624]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDDFF93FE82F9BDE8B6FB61" box="[342,456,1601,1624]" italics="true" pageId="20">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="59CC8ADFFFDDFF93FE02F9C3EB69FB61" box="[470,535,1599,1624]" captionStart="FIGURE 3" captionStartId="7.[151,250,894,916]" captionTargetBox="[151,1436,193,871]" captionTargetId="figure@7.[151,1436,193,873]" captionTargetPageId="7" captionText="FIGURE 3. Details of metathoracic tibia and tarsus of males of Cyanogomphini in lateral view. A. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); C. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); D. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270856/files/figure.png" pageId="20">Fig. 3</figureCitation>
|
||
D). The posterior hamule varies greatly, being longer and slender with the “heel” distinctly less pronounced in
|
||
<taxonomicName id="06F7EDD9FFDDFF93FE32F999EB3CFB44" box="[486,578,1636,1661]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="noval">
|
||
<emphasis id="F3834A48FFDDFF93FE32F999EB3CFB44" box="[486,578,1636,1661]" italics="true" pageId="20">T. noval</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="59CC8ADFFFDDFF93FD87F998EBE2FB45" box="[595,668,1636,1661]" captionStart="FIGURE 4" captionStartId="10.[151,250,1598,1620]" captionTargetBox="[154,1436,151,1575]" captionTargetId="figure@10.[154,1439,151,1576]" captionTargetPageId="10" captionText="FIGURE 4. Males of Cyanogomphini, head in dorsal view (A) and secondary genitalia in lateral view (B – F). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270857/files/figure.png" pageId="20">Figs 4</figureCitation>
|
||
E, 5F–H, shorter and stouter in
|
||
<taxonomicName id="06F7EDD9FFDDFF93FBDCF999ED04FB45" box="[1032,1146,1637,1660]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDDFF93FBDCF999ED04FB45" box="[1032,1146,1637,1660]" italics="true" pageId="20">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
with a more pronounced “heel”,
|
||
<figureCitation id="59CC8ADFFFDDFF93FF3BF97BE849FB99" box="[239,311,1671,1696]" captionStart="FIGURE 4" captionStartId="10.[151,250,1598,1620]" captionTargetBox="[154,1436,151,1575]" captionTargetId="figure@10.[154,1439,151,1576]" captionTargetPageId="10" captionText="FIGURE 4. Males of Cyanogomphini, head in dorsal view (A) and secondary genitalia in lateral view (B – F). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270857/files/figure.png" pageId="20">Figs 4</figureCitation>
|
||
F,
|
||
<date id="B549B09AFFDDFF93FE81F97BE812FB99" box="[341,364,1671,1696]" pageId="20">5I</date>
|
||
–L), male cercus robust (
|
||
<figureCitation id="59CC8ADFFFDDFF93FD5AF97BEBA9FB99" box="[654,727,1671,1696]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="20">Figs 7</figureCitation>
|
||
G, 8G–H, ratio cercus length / maximum width 0.30–0.36 vs. more slender,
|
||
<quantity id="060F3BBFFFDDFF93FEEAF950E8B1FBFC" box="[318,463,1708,1733]" metricMagnitude="-3" metricUnit="m" metricValue="6.985" metricValueMax="7.62" metricValueMin="6.35" pageId="20" unit="in" value="0.275" valueMax="0.3" valueMin="0.25">0.25–0.30 in</quantity>
|
||
<taxonomicName id="06F7EDD9FFDDFF93FE0EF951EB2FFBFD" box="[474,593,1709,1732]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDDFF93FE0EF951EB2FFBFD" box="[474,593,1709,1732]" italics="true" pageId="20">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="59CC8ADFFFDDFF93FDB0F950EBD3FBFD" box="[612,685,1708,1733]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="20">Figs 7</figureCitation>
|
||
M,
|
||
<date id="B549B09AFFDDFF93FD0DF950EB8EFBFD" box="[729,752,1708,1732]" pageId="20">8I</date>
|
||
–J). Further, the smaller size of
|
||
<taxonomicName id="06F7EDD9FFDDFF93FBB9F951EDB7FBFC" box="[1133,1225,1708,1733]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="noval">
|
||
<emphasis id="F3834A48FFDDFF93FBB9F951EDB7FBFC" box="[1133,1225,1708,1733]" italics="true" pageId="20">T. noval</emphasis>
|
||
</taxonomicName>
|
||
(male Hw length 24.7–26.3 vs. 26.8–29.7 of
|
||
<taxonomicName id="06F7EDD9FFDDFF93FE09F92DEB2CFBD1" box="[477,594,1745,1768]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDDFF93FE09F92DEB2CFBD1" box="[477,594,1745,1768]" italics="true" pageId="20">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
) and allopatric distribution, with
|
||
<taxonomicName id="06F7EDD9FFDDFF93FC30F92DED3CFBD1" box="[996,1090,1743,1768]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="noval">
|
||
<emphasis id="F3834A48FFDDFF93FC30F92DED3CFBD1" box="[996,1090,1743,1768]" italics="true" pageId="20">T. noval</emphasis>
|
||
</taxonomicName>
|
||
occurring in biogeographical Pampean province of Chacoan dominion and
|
||
<taxonomicName id="06F7EDD9FFDDFF93FD61F909EA55FA35" box="[693,811,1781,1804]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDDFF93FD61F909EA55FA35" box="[693,811,1781,1804]" italics="true" pageId="20">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
in Araucaria and Parana Forest provinces in Parana dominion (
|
||
<figureCitation id="59CC8ADFFFDDFF93FEC6F8EBE81AFA09" box="[274,356,1815,1840]" captionStart="FIGURE 13" captionStartId="25.[151,250,1528,1550]" captionTargetBox="[159,1434,488,1498]" captionTargetId="figure@25.[151,1436,483,1503]" captionTargetPageId="25" captionText="FIGURE 13. Map of southeastern South America with distribution records of Tibiagomphus Belle, 1992, species. Colored areas correspond to the Neotropical regionalization of biotic elements by Morrone (2014). Abbreviations: ER, Entre Rios and MI, Misiones for Argentine provinces; PR, Paraná, RS, Rio Grande do Sul, and SC, Santa Catarina for Brazilian States." httpUri="https://zenodo.org/record/270866/files/figure.png" pageId="20">Fig. 13</figureCitation>
|
||
), separate these two species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="89EDC5D1FFDDFF93FF13F8C0EC00FADC" pageId="20" type="distribution">
|
||
<paragraph id="C148965AFFDDFF93FF13F8C0ED0BFA41" blockId="20.[151,1437,556,2021]" pageId="20">
|
||
<emphasis id="F3834A48FFDDFF93FF13F8C0E827FA6C" bold="true" box="[199,345,1852,1877]" pageId="20">Distribution</emphasis>
|
||
. From Rio Grande do Sul State,
|
||
<collectingCountry id="B9E0D6CAFFDDFF93FD30F8C0EA55FA6C" box="[740,811,1852,1877]" name="Brazil" pageId="20">Brazil</collectingCountry>
|
||
, and Entre Rios Province,
|
||
<collectingCountry id="B9E0D6CAFFDDFF93FBBCF8C0EDA6FA6C" box="[1128,1240,1852,1877]" name="Argentina" pageId="20">Argentina</collectingCountry>
|
||
south to Florida Department,
|
||
<collectingCountry id="B9E0D6CAFFDDFF93FEFDF89DE8F0FA40" box="[297,398,1889,1913]" name="Uruguay" pageId="20">Uruguay</collectingCountry>
|
||
; an inhabitant of the Pampean biogeographical province (
|
||
<figureCitation id="59CC8ADFFFDDFF93FBC0F8A3ED19FA41" box="[1044,1127,1887,1912]" captionStart="FIGURE 13" captionStartId="25.[151,250,1528,1550]" captionTargetBox="[159,1434,488,1498]" captionTargetId="figure@25.[151,1436,483,1503]" captionTargetPageId="25" captionText="FIGURE 13. Map of southeastern South America with distribution records of Tibiagomphus Belle, 1992, species. Colored areas correspond to the Neotropical regionalization of biotic elements by Morrone (2014). Abbreviations: ER, Entre Rios and MI, Misiones for Argentine provinces; PR, Paraná, RS, Rio Grande do Sul, and SC, Santa Catarina for Brazilian States." httpUri="https://zenodo.org/record/270866/files/figure.png" pageId="20">Fig. 13</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C148965AFFDDFF93FF13F878EC00FADC" blockId="20.[151,1437,556,2021]" pageId="20">
|
||
<emphasis id="F3834A48FFDDFF93FF13F878EB4FFAA4" bold="true" box="[199,561,1924,1949]" pageId="20">Biological and ecological data.</emphasis>
|
||
Adults are found in lotic environments such as riffles and deep zones in slow streams surrounded by gallery forest, where adults perch in protected sites, from sea level to 107 meters of elevation. Larvae are shallow burrowers and were found with some adhered detritus (Rodrigues Capítulo 1985).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="9588C6D2FFDCFF92FF43F992EC40FBC6" httpUri="https://zenodo.org/record/270863/files/figure.png" pageId="21" targetBox="[151,1436,195,1620]" targetPageId="21">
|
||
<paragraph id="C148965AFFDCFF92FF43F992EC40FBC6" blockId="21.[151,1436,1646,1791]" pageId="21">
|
||
<emphasis id="F3834A48FFDCFF92FF43F992E865FBBA" bold="true" box="[151,283,1646,1668]" pageId="21">FIGURE 10</emphasis>
|
||
. Male habitus of Cyanogomphini in lateral view. A–B.
|
||
<taxonomicName id="06F7EDD9FFDCFF92FCB7F992EDCCFBBD" box="[867,1202,1646,1668]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="21" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
|
||
<emphasis id="F3834A48FFDCFF92FCB7F992EDCCFBBD" box="[867,1202,1646,1668]" italics="true" pageId="21">Cyanogomphus angelomachadoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="F3834A48FFDCFF92FB6EF993EC73FBBD" bold="true" box="[1210,1293,1647,1668]" pageId="21">
|
||
<taxonomicNameLabel id="E8B0F733FFDCFF92FB6EF993EC73FBBD" box="[1210,1293,1647,1668]" pageId="21" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, A. Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Paratype (Brazil. MG: Serra do Salitre, ABMM); C.
|
||
<taxonomicName id="06F7EDD9FFDCFF92FF43F950EB39FBFB" authority="Belle, 1994" authorityName="Belle" authorityYear="1994" box="[151,583,1708,1730]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="21" phylum="Arthropoda" rank="species" species="comparabilis">
|
||
<emphasis id="F3834A48FFDCFF92FF43F950E8BAFBFB" box="[151,452,1708,1730]" italics="true" pageId="21">Cyanogomphus comparabilis</emphasis>
|
||
Belle, 1994
|
||
</taxonomicName>
|
||
, Holotype (Brazil. MT: Diamantino, ABMM); D.
|
||
<taxonomicName id="06F7EDD9FFDCFF92FB8EF950E9B2FBD9" authority="Selys, 1873" authorityName="Selys" authorityYear="1873" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="21" phylum="Arthropoda" rank="species" species="waltheri">
|
||
<emphasis id="F3834A48FFDCFF92FB8EF950EC2BFBFB" box="[1114,1365,1708,1730]" italics="true" pageId="21">Cyanogomphus waltheri</emphasis>
|
||
Selys, 1873
|
||
</taxonomicName>
|
||
, (Brazil. RJ: Santana de Japuíba district, MNRJ); E.
|
||
<taxonomicName id="06F7EDD9FFDCFF92FCDAF936EC71FBD9" authority="Rodrigues Capitulo, 1985" authorityName="Rodrigues Capitulo" authorityYear="1985" box="[782,1295,1738,1760]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="21" phylum="Arthropoda" rank="species" species="noval">
|
||
<emphasis id="F3834A48FFDCFF92FCDAF936EA9FFBD9" box="[782,993,1738,1760]" italics="true" pageId="21">Tibiagomphus noval</emphasis>
|
||
(Rodrigues Capítulo, 1985)
|
||
</taxonomicName>
|
||
, (Brazil. RS: Jaguarão, MNRJ); F.
|
||
<taxonomicName id="06F7EDD9FFDCFF92FEBBF915EB95FBC6" authority="Fraser, 1947" authorityName="Fraser" authorityYear="1947" box="[367,747,1769,1791]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="21" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDCFF92FEBBF915EB2AFBC6" box="[367,596,1769,1791]" italics="true" pageId="21">Tibiagomphus uncatus</emphasis>
|
||
(Fraser, 1947)
|
||
</taxonomicName>
|
||
, (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="89EDC5D1FFDCFF91FF13F8DAECE2F945" lastPageId="22" pageId="21" type="discussion">
|
||
<paragraph id="C148965AFFDCFF91FF13F8DAE8DFFDC1" blockId="21.[151,1436,1830,2035]" lastBlockId="22.[151,1436,151,1149]" lastPageId="22" pageId="21">
|
||
<emphasis id="F3834A48FFDCFF92FF13F8DAE845FA06" bold="true" box="[199,315,1830,1855]" pageId="21">Remarks.</emphasis>
|
||
The typification by Rodrigues Capítulo (1985) is misleading and imprecise. In describing
|
||
<taxonomicName id="06F7EDD9FFDCFF92FAE8F8DBECE2FA06" box="[1340,1436,1830,1855]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="21" phylum="Arthropoda" rank="species" species="noval">
|
||
<emphasis id="F3834A48FFDCFF92FAE8F8DBECE2FA06" box="[1340,1436,1830,1855]" italics="true" pageId="21">C. noval</emphasis>
|
||
</taxonomicName>
|
||
he designated an adult male emerged in the laboratory as
|
||
<typeStatus id="1E4C28F8FFDCFF92FC9DF8B6EAD5FA5A" box="[841,939,1866,1891]" pageId="21" type="holotype">holotype</typeStatus>
|
||
without mention of its exuvia (Rodrigues Capítulo 1985, p. 329). Furthermore, the other cited specimens, an adult female and six larvae, were not designated as
|
||
<typeStatus id="1E4C28F8FFDCFF92FF61F86FE85DFA92" box="[181,291,1939,1963]" pageId="21" type="paratype">paratypes</typeStatus>
|
||
, therefore this meets the requirement that “expressly excludes them from the
|
||
<typeStatus id="1E4C28F8FFDCFF92FB56F86FEDCCFA92" box="[1154,1202,1939,1963]" pageId="21">type</typeStatus>
|
||
series” (ICZN 1999, Art. 72.4.6), even though an allotype was mentioned in the abstract. The subsequent citation of the female as allotype by Muzón
|
||
<emphasis id="F3834A48FFDCFF92FEA1F820E8D0FACA" box="[373,430,2010,2035]" italics="true" pageId="21">et al.</emphasis>
|
||
(2007) should be considered an invalid nomenclatural act. The date of publication of all papers in that volume is stated to be 1984, but the cover states 1985. For this reason, without further investigation, we follow
|
||
<bibRefCitation id="A566EBABFFDFFF91FEC4FF40EB7BFDEC" box="[272,517,188,213]" pageId="22" refString="Garrison, R. W., von Ellenrieder, N. & Louton, J. A. (2006) Dragonfly genera of the New World, an illustrated and annotated key to the Anisoptera. The Johns Hopkins University Press, Baltimore, xiv + 368 pp." type="book">
|
||
Garrison
|
||
<emphasis id="F3834A48FFDFFF91FEAEFF41E8CDFDEC" box="[378,435,188,213]" italics="true" pageId="22">et al.</emphasis>
|
||
(2006)
|
||
</bibRefCitation>
|
||
considering 1985 as the correct year of publication, while the respective day and month still are pending.
|
||
</paragraph>
|
||
<paragraph id="C148965AFFDFFF91FF13FEF9E9A1F90D" blockId="22.[151,1436,151,1149]" pageId="22">
|
||
Except by the paler color (“pardo amarillenta”), all characters mentioned by Rodrigues Capítulo (1985, p. 329–332) are not diagnostic of
|
||
<taxonomicName id="06F7EDD9FFDFFF91FDDEFED5EB1BFC79" box="[522,613,295,320]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="noval">
|
||
<emphasis id="F3834A48FFDFFF91FDDEFED5EB1BFC79" box="[522,613,295,320]" italics="true" pageId="22">T. noval</emphasis>
|
||
</taxonomicName>
|
||
in accordance with its new status recognized here, and his characters should be considered as examples of individual variation, as they are shared with some
|
||
<taxonomicName id="06F7EDD9FFDFFF91FB75FEB1EC6BFC5D" box="[1185,1301,333,356]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDFFF91FB75FEB1EC6BFC5D" box="[1185,1301,333,356]" italics="true" pageId="22">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
. Rodrigues Capítulo (1985) provided no diagnosis, instead comparing a few selected characters with
|
||
<taxonomicName id="06F7EDD9FFDFFF91FB55FE8DED8DFCB1" box="[1153,1267,369,392]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDFFF91FB55FE8DED8DFCB1" box="[1153,1267,369,392]" italics="true" pageId="22">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
as follows: (1) anterior margin of the postfrons in dorsal view sinuous with a small mesal convexity, while in
|
||
<taxonomicName id="06F7EDD9FFDFFF91FB3BFE69EC1DFC95" box="[1263,1379,405,428]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDFFF91FB3BFE69EC1DFC95" box="[1263,1379,405,428]" italics="true" pageId="22">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
it is entirely convex; (2) male S9 with a V-shaped mesodorsal invagination vs. lacking invagination in
|
||
<taxonomicName id="06F7EDD9FFDFFF91FB23FE45EC15FCE9" box="[1271,1387,441,464]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDFFF91FB23FE45EC15FCE9" box="[1271,1387,441,464]" italics="true" pageId="22">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
; (3) spines along posterior carina of S9 extend downward two-thirds of the segment height in the male vs. only over dorsal surface in
|
||
<taxonomicName id="06F7EDD9FFDFFF91FEB6FDFDE8A9FF21" box="[354,471,513,536]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDFFF91FEB6FDFDE8A9FF21" box="[354,471,513,536]" italics="true" pageId="22">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
; (4) posterior margin of male epiproct almost straight vs. ovoid in
|
||
<taxonomicName id="06F7EDD9FFDFFF91FB26FDFDEC19FF21" box="[1266,1383,513,536]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDFFF91FB26FDFDEC19FF21" box="[1266,1383,513,536]" italics="true" pageId="22">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
; (5) female distal margin of sternites of S9–10 distinct from
|
||
<taxonomicName id="06F7EDD9FFDFFF91FCECFDD9EAD0FF05" box="[824,942,549,572]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDFFF91FCECFDD9EAD0FF05" box="[824,942,549,572]" italics="true" pageId="22">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
. The shape of the anterior margin of the postfrons (character 1) is variable, specimens of
|
||
<taxonomicName id="06F7EDD9FFDFFF91FD1CFDB5EA5CFF59" box="[712,802,583,608]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="noval">
|
||
<emphasis id="F3834A48FFDFFF91FD1CFDB5EA5CFF59" box="[712,802,583,608]" italics="true" pageId="22">T. noval</emphasis>
|
||
</taxonomicName>
|
||
collected from the same site and day showing either a small mesal convexity (
|
||
<figureCitation id="59CC8ADFFFDFFF91FE71FD90E897FFBC" box="[421,489,620,645]" captionStart="FIGURE 5" captionStartId="11.[151,250,1535,1557]" captionTargetBox="[151,1436,195,1507]" captionTargetId="figure@11.[151,1436,195,1511]" captionTargetPageId="11" captionText="FIGURE 5. Males of Cyanogomphini, details of variations of postfrons in dorsal view (A – D) and posterior hamule in lateral view (E – L). A – B, F – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); C – D, I – L. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). E. Cyanogomphus comparabilis Belle, 1994, Paratype (Brazil. SP: Igarapava, ABMM). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270858/files/figure.png" pageId="22">Fig. 5</figureCitation>
|
||
B) or regularly convex postfrons (
|
||
<figureCitation id="59CC8ADFFFDFFF91FCB8FD90EAD1FFBC" box="[876,943,620,645]" captionStart="FIGURE 5" captionStartId="11.[151,250,1535,1557]" captionTargetBox="[151,1436,195,1507]" captionTargetId="figure@11.[151,1436,195,1511]" captionTargetPageId="11" captionText="FIGURE 5. Males of Cyanogomphini, details of variations of postfrons in dorsal view (A – D) and posterior hamule in lateral view (E – L). A – B, F – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); C – D, I – L. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). E. Cyanogomphus comparabilis Belle, 1994, Paratype (Brazil. SP: Igarapava, ABMM). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270858/files/figure.png" pageId="22">Fig. 5</figureCitation>
|
||
A) such as illustrated for the
|
||
<typeStatus id="1E4C28F8FFDFFF91FB2AFD90EC1EFFBC" box="[1278,1376,620,645]" pageId="22" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="06F7EDD9FFDFFF91FA53FD91E990FF91" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDFFF91FA53FD91E990FF91" italics="true" pageId="22">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
by Rodrigues Capítulo (1985, fig. 1). Additionally, variation in the postfrons in
|
||
<taxonomicName id="06F7EDD9FFDFFF91FB58FD6DEC7FFF91" box="[1164,1281,657,680]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDFFF91FB58FD6DEC7FFF91" box="[1164,1281,657,680]" italics="true" pageId="22">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
also shows a slightly mesal projection along the anterior margin of postfrons (
|
||
<figureCitation id="59CC8ADFFFDFFF91FCA0FD48EAC5FFF4" box="[884,955,692,717]" captionStart="FIGURE 5" captionStartId="11.[151,250,1535,1557]" captionTargetBox="[151,1436,195,1507]" captionTargetId="figure@11.[151,1436,195,1511]" captionTargetPageId="11" captionText="FIGURE 5. Males of Cyanogomphini, details of variations of postfrons in dorsal view (A – D) and posterior hamule in lateral view (E – L). A – B, F – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); C – D, I – L. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). E. Cyanogomphus comparabilis Belle, 1994, Paratype (Brazil. SP: Igarapava, ABMM). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270858/files/figure.png" pageId="22">Figs 5</figureCitation>
|
||
C–D). A small mesodorsal invagination in S9 (character 2) is common to all Cyanogomphini including
|
||
<taxonomicName id="06F7EDD9FFDFFF91FC91FD25EAC6FFC9" box="[837,952,729,752]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDFFF91FC91FD25EAC6FFC9" box="[837,952,729,752]" italics="true" pageId="22">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="59CC8ADFFFDFFF91FC1CFD2BED6EFFD6" box="[968,1040,727,752]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="22">Figs 7</figureCitation>
|
||
B, E, H, K, N). We did not find the shape of invagination to be a diagnostic character. The distribution of spines along S9 (character 3) also is variable among specimens, and in both
|
||
<taxonomicName id="06F7EDD9FFDFFF91FE27FCDDEB35FE01" box="[499,587,799,824]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="noval">
|
||
<emphasis id="F3834A48FFDFFF91FE27FCDDEB35FE01" box="[499,587,799,824]" italics="true" pageId="22">T. noval</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="06F7EDD9FFDFFF91FD50FCDDEB88FE01" box="[644,758,801,824]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDFFF91FD50FCDDEB88FE01" box="[644,758,801,824]" italics="true" pageId="22">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
they can extend down two-thirds of the segment height (see
|
||
<figureCitation id="59CC8ADFFFDFFF91FF43FCB8E9A3FE65" box="[151,221,836,861]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="22">Figs 7</figureCitation>
|
||
J, M). This variation might have lead J. Belle to misidentify some
|
||
<taxonomicName id="06F7EDD9FFDFFF91FC6AFCB9ED6BFE64" box="[958,1045,836,861]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="noval">
|
||
<emphasis id="F3834A48FFDFFF91FC6AFCB9ED6BFE64" box="[958,1045,836,861]" italics="true" pageId="22">T. noval</emphasis>
|
||
</taxonomicName>
|
||
as
|
||
<taxonomicName id="06F7EDD9FFDFFF91FBE8FCB9EDD3FE65" box="[1084,1197,837,860]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDFFF91FBE8FCB9EDD3FE65" box="[1084,1197,837,860]" italics="true" pageId="22">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
because they had the spines extending to the ventral half. Even though in
|
||
<taxonomicName id="06F7EDD9FFDFFF91FD35FC95EA47FEB9" box="[737,825,871,896]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="noval">
|
||
<emphasis id="F3834A48FFDFFF91FD35FC95EA47FEB9" box="[737,825,871,896]" italics="true" pageId="22">T. noval</emphasis>
|
||
</taxonomicName>
|
||
the distal margin of the epiproct (character 4) appears be straighter than that of
|
||
<taxonomicName id="06F7EDD9FFDFFF91FE68FC71EB4EFE9D" box="[444,560,909,932]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDFFF91FE68FC71EB4EFE9D" box="[444,560,909,932]" italics="true" pageId="22">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="59CC8ADFFFDFFF91FD96FC70EBF2FE9D" box="[578,652,908,933]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="22">Figs 7</figureCitation>
|
||
J, L, M,
|
||
<collectingCountry id="B9E0D6CAFFDFFF91FD25FC70EA79FE9D" box="[753,775,908,932]" name="Iceland" pageId="22">O</collectingCountry>
|
||
, 8G–J), it is not ovoid in
|
||
<taxonomicName id="06F7EDD9FFDFFF91FBE1FC71EDD4FE9D" box="[1077,1194,909,932]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDFFF91FBE1FC71EDD4FE9D" box="[1077,1194,909,932]" italics="true" pageId="22">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
, and this structure is similar for both species. Finally, character 5 on female sternites of S9–10 cannot be verified, as differences were not depicted. Although Rodrigues Capítulo (1985) borrowed the
|
||
<typeStatus id="1E4C28F8FFDFFF91FCBDFC28EAB5FED4" box="[873,971,980,1005]" pageId="22" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="06F7EDD9FFDFFF91FC24FC29ED1EFED5" box="[1008,1120,981,1004]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDFFF91FC24FC29ED1EFED5" box="[1008,1120,981,1004]" italics="true" pageId="22">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
, he adopted the illustrations by
|
||
<bibRefCitation id="A566EBABFFDFFF91FF69FC04E829F929" author="Fraser" box="[189,343,1016,1040]" pageId="22" refString="Fraser, F. C. (1947) The Odonata of the Argentine Republic I. Acta Zoologica Lilloana del Instituto Miguel Lillo, 4, 427 - 462." type="journal article" year="1947">Fraser (1947)</bibRefCitation>
|
||
for his comparisons, possibly due to the poor condition of the
|
||
<typeStatus id="1E4C28F8FFDFFF91FBE0FC0BEDE8F929" box="[1076,1174,1015,1040]" pageId="22" type="holotype">holotype</typeStatus>
|
||
(S4–10 lost, see
|
||
<bibRefCitation id="A566EBABFFDFFF91FAB5FC0BE9ADF90D" author="Belle" pageId="22" refString="Belle, J. (1980) Notes on three species of Cyanogomphus Selys, 1873 (Odonata: Gomphidae). Entomologische Berichten, 40 (10), 151 - 155." type="journal article" year="1980">Belle 1980</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C148965AFFDFFF91FF13FBC3ECE2F945" blockId="22.[151,1436,151,1149]" pageId="22">
|
||
The males from municipalities of Guaíba and Pelotas were determined as
|
||
<taxonomicName id="06F7EDD9FFDFFF91FBEEFBBDEDE9F961" box="[1082,1175,1087,1112]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="noval">
|
||
<emphasis id="F3834A48FFDFFF91FBEEFBBDEDE9F961" box="[1082,1175,1087,1112]" italics="true" pageId="22">T. noval</emphasis>
|
||
</taxonomicName>
|
||
, while a couple from Jaguarão municipality, all from Rio Grande do Sul State,
|
||
<collectingCountry id="B9E0D6CAFFDFFF91FCCCFB98EA21F944" box="[792,863,1124,1149]" name="Brazil" pageId="22">Brazil</collectingCountry>
|
||
, were determined as
|
||
<taxonomicName id="06F7EDD9FFDFFF91FB9EFB99EDC2F945" box="[1098,1212,1125,1148]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFDFFF91FB9EFB99EDC2F945" box="[1098,1212,1125,1148]" italics="true" pageId="22">T. uncatus</emphasis>
|
||
</taxonomicName>
|
||
by J. Belle in 1992.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |