531 lines
71 KiB
XML
531 lines
71 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zse.99.99227" ID-GBIF-Dataset="7bf29efe-d807-401c-b61e-f35ba146bfc7" ID-Pensoft-Pub="1860-0743-1-273" ID-Pensoft-UUID="F3161813062A5D37A0C4DBE590620DF8" ID-ZooBank="86213715F1FC4838BC478464DC0B9F4B" ModsDocID="1860-0743-99-1-273" checkinTime="1680107811176" checkinUser="pensoft" docAuthor="Villarreal, Osvaldo & Chame-Vazquez, David" docDate="2023" docId="5A9658223F055671A90BD59088A40312" docLanguage="en" docName="ZoosystEvol 99(1): 273-280" docOrigin="Zoosystematics and Evolution 99 (1)" docPubDate="2023-03-29" docSource="http://dx.doi.org/10.3897/zse.99.99227" docTitle="Drymusa huberi Villarreal & Chamé-Vázquez 2023, sp. nov." docType="treatment" docUuid="32F46572-64B6-40C6-94E8-5126B59DA6BC" docUuidSource="ZooBank" docVersion="4" id="F3161813062A5D37A0C4DBE590620DF8" lastPageNumber="273" masterDocId="F3161813062A5D37A0C4DBE590620DF8" masterDocTitle="First record of the false violin spider of the family Drymusidae (Araneae, Synspermiata, Scytodoidea) from Venezuela, with the description of a new species" masterLastPageNumber="280" masterPageNumber="273" pageNumber="273" updateTime="1732480792725" updateUser="ExternalLinkService">
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<mods:titleInfo id="623809EB62B4B1EDF7EED3C0F42369AD">
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<mods:title id="DFFD757E913025A77251531438F69331">First record of the false violin spider of the family Drymusidae (Araneae, Synspermiata, Scytodoidea) from Venezuela, with the description of a new species</mods:title>
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<mods:namePart id="F8E27A709F76EE22C8E11BD7232CA946">Villarreal, Osvaldo</mods:namePart>
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<mods:nameIdentifier id="0B7E8DF764B5704A97B862286A3AEDBC" type="ORCID">https://orcid.org/0000-0001-5355-3723</mods:nameIdentifier>
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<mods:affiliation id="5D6A881D95ADB5298D9B62A81778B8BF">Departamento de Invertebrados - Laboratorio de Aracnologia, Campus de Ensino e Pesquisa do Museu Nacional - Predio Administrativo, Avenida Bartolomeu de Gusmao, 875, RJ, CEP 20941 - 160 Sao Cristovao, Rio de Janeiro, Brazil & Museo del Instituto de Zoologia Agricola, Facultad de Agronomia, Universidad Central de Venezuela, Av 19 de Abril c / c Av Casanova Godoy, Maracay, Aragua, Venezuela</mods:affiliation>
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<mods:name id="1EFA121C783035B81F69FE831514D8C2" type="personal">
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<mods:roleTerm id="170091EB3E78119646BEA53063F3C72C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C51C3B37B2BB29873B98D0A256B9D3A4">Chame-Vazquez, David</mods:namePart>
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<mods:nameIdentifier id="389C5222A5CF92661832CF979CC1BFE2" type="ORCID">https://orcid.org/0000-0003-3433-153X</mods:nameIdentifier>
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<mods:affiliation id="DBA583E5C54E899ED27F8B1B5E6AE2F3">Laboratorio de Aracnologia y Entomologia, Centro de Investigaciones Biologicas del Noroeste S. C., Instituto Politecnico Nacional 195, Col. Playa Palo de Santa Rita Sur, CP. 23096, La Paz, Baja California Sur, Mexico</mods:affiliation>
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<mods:nameIdentifier id="0F476D265C529740DF97A1E255598F45" type="email">chamevazquez@gmail.com</mods:nameIdentifier>
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<mods:title id="0B26E65EE0DD66177382463BDED9FB83">Zoosystematics and Evolution</mods:title>
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<mods:part id="6D761C908CC14FE5B5279D3FD23C500F">
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<mods:date id="9A8868F091CD73167ED1E6FAB540116D">2023</mods:date>
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<mods:number id="76E1D4783D612C4E6E59D04732F07EDB">2023-03-29</mods:number>
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<mods:number id="81E952833BB97E9B720539E345F3578B">99</mods:number>
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<mods:number id="EA098AC34F8AE340FDB1D9E493AC9A01">1</mods:number>
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<mods:start id="B3003761340A23E66BA3706A1ACD5DF8">273</mods:start>
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<mods:end id="AA9F3C1BDE78A0C923F7C5B96E03BE03">280</mods:end>
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<mods:url id="B008C85F79B2C02550306A68CD61198E">http://dx.doi.org/10.3897/zse.99.99227</mods:url>
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<mods:classification id="0D5F53893FE6D30B88AD67AA2FB8ABD8">journal article</mods:classification>
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<mods:identifier id="C4AE40530A21AD5787CC1E22BBB60199" type="DOI">http://dx.doi.org/10.3897/zse.99.99227</mods:identifier>
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<treatment id="5A9658223F055671A90BD59088A40312" ID-GBIF-Taxon="209474030" LSID="urn:lsid:zoobank.org:act:32F46572-64B6-40C6-94E8-5126B59DA6BC" httpUri="http://treatment.plazi.org/id/5A9658223F055671A90BD59088A40312" lastPageNumber="273" pageId="0" pageNumber="273" scope_family="Drymusidae" scope_order="Araneae">
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<subSubSection id="4D96D2C587D952235EAF112477895D16" pageId="0" pageNumber="273" type="nomenclature">
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<paragraph id="955E40FCED7930AEDD1B31BF0A35068D" pageId="0" pageNumber="273">
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<taxonomicName id="F035B972F56A0E0A22532691AD7F4FF3" ID-CoL="C75QQ" LSID="https://zoobank.org/32F46572-64B6-40C6-94E8-5126B59DA6BC" authority="Villarreal & Chamé-Vázquez, 2023" authorityName="Villarreal & Chamé-Vázquez" authorityYear="2023" class="Arachnida" family="Drymusidae" genus="Drymusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymusa huberi" order="Araneae" pageId="0" pageNumber="273" phylum="Arthropoda" rank="species" species="huberi" status="sp. nov.">Drymusa huberi</taxonomicName>
|
||
<taxonomicNameLabel id="4BDCDA9DBE7547135DDD2F554D07CACD" pageId="0" pageNumber="273">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F21EA174F7FFFC3B34E6A78372C48C43" pageId="0" pageNumber="273" type="description">
|
||
<paragraph id="446655A32A45CB7F88545F3A4E8597F4" pageId="0" pageNumber="273">
|
||
<figureCitation id="C8006B5A4AA07CD4B5D306B962805136" captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">Figs 1-12</figureCitation>
|
||
<figureCitation id="3BEAE53288519448D1E085BA24494DA3" captionStart="Figures 13–20" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">, 13-20</figureCitation>
|
||
<figureCitation id="6066BE8AB856E8DB5FAC1D7AE84E5AFA" captionStart="Figures 21–32" captionStartId="F3" captionText="Figures 21 - 32. Drymusa spp., females. 21. D. simoni, posterior part of opisthosoma, lateral view; 22. D. dinora, opisthosoma, ventral view; 23, 25 - 27. Female genitalia, ventral view (23. D. colligata; 25. D. spectata; 26. D. armasi; 27. D. serrana); 24, 28 - 32. Same, dorsal view (24. D. huberi sp. nov.; 28. D. serrana; 29. D. canhemabae; 30. D. colligata; 31. D. rengan; 32. D. spelunca)." figureDoi="10.3897/zse.99.99227.figures21-32" httpUri="https://binary.pensoft.net/fig/831283" pageId="0" pageNumber="273">, 24</figureCitation>
|
||
<figureCitation id="951E095CF44FCEC9B226DD2EB4AFBA70" captionStart="Figures 49–53" captionStartId="F5" captionText="Figures 49 - 53. 49, 50. Drymusa huberi sp. nov. live specimens from La Guaquira, Yaracuy State, Venezuela. 49. Male, paratype; 50. Female, holotype; 51. Habitat, tropical dry forest in La Guaquira, Yaracuy State; 52. Leaf litter on the forest floor, microhabitat of the species; 53. Known distribution of all species of the genus Drymusa. Photo credit: O. Villarreal: 49, 50, 51; D. Romero: 52." figureDoi="10.3897/zse.99.99227.figures49-53" httpUri="https://binary.pensoft.net/fig/831285" pageId="0" pageNumber="273">, 49</figureCitation>
|
||
<figureCitation id="F92E7F682BCF34CCDF0A959DF47380A5" captionStart="Figures 49–53" captionStartId="F5" captionText="Figures 49 - 53. 49, 50. Drymusa huberi sp. nov. live specimens from La Guaquira, Yaracuy State, Venezuela. 49. Male, paratype; 50. Female, holotype; 51. Habitat, tropical dry forest in La Guaquira, Yaracuy State; 52. Leaf litter on the forest floor, microhabitat of the species; 53. Known distribution of all species of the genus Drymusa. Photo credit: O. Villarreal: 49, 50, 51; D. Romero: 52." figureDoi="10.3897/zse.99.99227.figures49-53" httpUri="https://binary.pensoft.net/fig/831285" pageId="0" pageNumber="273">, 50</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F65BBA52255CB4432BA84F7E8AB2DE84" pageId="0" pageNumber="273" type="type data">
|
||
<paragraph id="D50926EB4DEA1C697AE431FAAB285C6C" pageId="0" pageNumber="273">Type data.</paragraph>
|
||
<paragraph id="B0F05F4B2A50856843A85581DFC8E5FF" pageId="0" pageNumber="273">
|
||
Venezuela • ♀
|
||
<emphasis id="56FEA2B5883B15358B287E9450F0363C" bold="true" italics="true" pageId="0" pageNumber="273">holotype</emphasis>
|
||
: MIZA0105853; Yaracuy, La
|
||
<normalizedToken id="89EF841B4B4B709583FB5949242F1AEF" originalValue="Guáquira">Guaquira</normalizedToken>
|
||
;
|
||
<geoCoordinate id="61ACA09F4263EE576177357FC692B6C0" degrees="10.2951" direction="north" orientation="latitude" precision="5" value="10.2951">10.2951°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A881927E99FAA1683D14690E1B9D1634" degrees="68.6535" direction="west" orientation="longitude" precision="5" value="-68.6535">68.6535°W</geoCoordinate>
|
||
; 120 m a.s.l.; 16 Feb 2020; O. Villarreal, B.A. Huber, Q. Arias C. leg.; forest along stream • Paratype: 1 ♂ same data as the holotype.
|
||
</paragraph>
|
||
<caption id="F610FDB245960028A9437DFF571D567D" doi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273" start="Figures 1–12" startId="F1">
|
||
<paragraph id="BA223666E041F35A9C6D63740D76D97E" pageId="0" pageNumber="273">
|
||
<emphasis id="EE53111855DBFF97DB053E8303C3A3F7" bold="true" pageId="0" pageNumber="273">Figures 1-12.</emphasis>
|
||
<taxonomicName id="4767B6BD4550CA65C15863972EC79F19" authorityName="Villarreal & Chamé-Vázquez" authorityYear="2023" class="Arachnida" family="Drymusidae" genus="Drymusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymusa huberi" order="Araneae" pageId="0" pageNumber="273" phylum="Arthropoda" rank="species" species="huberi">
|
||
<emphasis id="4FA1301B9B7399020E92C13723921D99" italics="true" pageId="0" pageNumber="273">Drymusa huberi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., female holotype (
|
||
<emphasis id="D2B672B67F4303A984E82064B565C030" bold="true" pageId="0" pageNumber="273">1-4, 9, 11</emphasis>
|
||
) and male paratype (
|
||
<emphasis id="A7E74A3C4D3BAAD2729C0D99C02C5637" bold="true" pageId="0" pageNumber="273">5-8, 10, 12</emphasis>
|
||
).
|
||
<emphasis id="569FDBD309392889BA2EFD3C3CC92C37" bold="true" pageId="0" pageNumber="273">1, 5.</emphasis>
|
||
Prosoma, dorsal view;
|
||
<emphasis id="6C6E3C249BF6663A2C9ED46D5EDD0543" bold="true" pageId="0" pageNumber="273">3, 7.</emphasis>
|
||
Same, ventral view;
|
||
<emphasis id="6D4A5DAC523B8DBFFBCC80659B589778" bold="true" pageId="0" pageNumber="273">2, 6.</emphasis>
|
||
Opisthosoma, dorsal view;
|
||
<emphasis id="0A92068C04F0B864363352CF8640DE13" bold="true" pageId="0" pageNumber="273">4, 8, 11</emphasis>
|
||
Same, ventral view (
|
||
<emphasis id="EA43CCAFCCE4F9543FA818B3614A6F68" bold="true" pageId="0" pageNumber="273">11</emphasis>
|
||
depilated);
|
||
<emphasis id="43067683161506B502C2C2B04B43D397" bold="true" pageId="0" pageNumber="273">9, 10.</emphasis>
|
||
Prosoma, frontal view;
|
||
<emphasis id="D1DAF99EC0B431D99AEEC4993E9BDA11" bold="true" pageId="0" pageNumber="273">12.</emphasis>
|
||
Same, lateral view. Scale bars: 0.5 mm (
|
||
<emphasis id="B6715406E0323452834B49C3D388D555" bold="true" pageId="0" pageNumber="273">1-12</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="E921558F600A0C31AA8E56912A18B30D" doi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273" start="Figures 13–20" startId="F2">
|
||
<paragraph id="9395A800930894D6EAD7FA7E087F756B" pageId="0" pageNumber="273">
|
||
<emphasis id="FD63480E807B5FC57E7497C75184DE86" bold="true" pageId="0" pageNumber="273">Figures 13-20.</emphasis>
|
||
<taxonomicName id="A08C390AA7A278AD0A7DEB9D85519D90" authorityName="Villarreal & Chamé-Vázquez" authorityYear="2023" class="Arachnida" family="Drymusidae" genus="Drymusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymusa huberi" order="Araneae" pageId="0" pageNumber="273" phylum="Arthropoda" rank="species" species="huberi">
|
||
<emphasis id="D3FC519680399C448EBCA6BDD7326EAE" italics="true" pageId="0" pageNumber="273">Drymusa huberi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis id="476675A87355CE4A3877414974DD9A72" bold="true" pageId="0" pageNumber="273">13.</emphasis>
|
||
Female genitalia, ventral view;
|
||
<emphasis id="2B15E75349974FA91FF916C5A16C477A" bold="true" pageId="0" pageNumber="273">14.</emphasis>
|
||
Same (cleared), dorsal view;
|
||
<emphasis id="8F15AF7E02BD82A19BE5EFC3FE3B77AA" bold="true" pageId="0" pageNumber="273">15.</emphasis>
|
||
Male palp, prolateral view;
|
||
<emphasis id="FF40FDE593C1EFBF21A90DC75F58ADDA" bold="true" pageId="0" pageNumber="273">16, 20.</emphasis>
|
||
Same, ventral view (
|
||
<emphasis id="1BF4EC136D8FB1D699F926C62557052F" bold="true" pageId="0" pageNumber="273">20</emphasis>
|
||
cleared);
|
||
<emphasis id="DCAA27D09596000209FCA22414474AF5" bold="true" pageId="0" pageNumber="273">17.</emphasis>
|
||
Detail of vulva (cleared), dorsal view;
|
||
<emphasis id="528A9EC78DF7D3F832492DB21D94D0D6" bold="true" pageId="0" pageNumber="273">18.</emphasis>
|
||
Male palpal bulb, apical view;
|
||
<emphasis id="B3B058BD9F96710B67A704A96F2F4988" bold="true" pageId="0" pageNumber="273">19.</emphasis>
|
||
Same, ventral view. Scale bars: 0.2 mm (
|
||
<emphasis id="004B31017A4DE5B6B5C0B9EE605F6B4C" bold="true" pageId="0" pageNumber="273">13-16, 18-20</emphasis>
|
||
). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="E2E7CB0989FA5E947039762DA32264D2" pageId="0" pageNumber="273" type="etymology">
|
||
<paragraph id="8561F15FD142470BB2E7619576FA6A61" pageId="0" pageNumber="273">Etymology.</paragraph>
|
||
<paragraph id="F4718949952CC11CF60760377889C263" pageId="0" pageNumber="273">The specific epithet is a name in apposition to honour the arachnologist Bernhard A. Huber in recognition of his prolific and excellent contributions to the taxonomy and systematics of the pholcid spiders of Venezuela and the world. Bernhard was also one of the collectors of the type specimens.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="53C668D8A6F1F311C7B65F5F0F6AC19C" pageId="0" pageNumber="273" type="diagnosis">
|
||
<paragraph id="9514A847A3AD0717F01931646A0A18BF" pageId="0" pageNumber="273">Diagnosis.</paragraph>
|
||
<paragraph id="9C9C7AD213CD7DD9C1DF5CBE6D91AE24" pageId="0" pageNumber="273">
|
||
The vulva of
|
||
<taxonomicName id="44C6F529A67D855FB6AB37278960B300" lsidName="D. huberi" pageId="0" pageNumber="273" rank="species" species="huberi">
|
||
<emphasis id="BE0F3673EE57A2BD1D1F5277FE0AFFCD" italics="true" pageId="0" pageNumber="273">D. huberi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. resembles those of
|
||
<taxonomicName id="8002C694EE36276026C1634C9763F2FB" lsidName="D. canhemabae" pageId="0" pageNumber="273" rank="species" species="canhemabae">
|
||
<emphasis id="72FCB503A5AE615007E35D8EEB85762A" italics="true" pageId="0" pageNumber="273">D. canhemabae</emphasis>
|
||
</taxonomicName>
|
||
Brescovit, Bonaldo & Rheims, 2004 and
|
||
<taxonomicName id="F0DEEBB6CB43051CACE15B57028EE54E" lsidName="D. colligata" pageId="0" pageNumber="273" rank="species" species="colligata">
|
||
<emphasis id="6BAB8B4B01EB7586FB1C280A757F1D42" italics="true" pageId="0" pageNumber="273">D. colligata</emphasis>
|
||
</taxonomicName>
|
||
Bonaldo, Rheims & Brescovit, 2006 by the unbranched, long spermathecae (Figs
|
||
<figureCitation id="F79F11D1BA4C0F1715923E2CE1B09B92" captionStart="Figures 13–20" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">14</figureCitation>
|
||
,
|
||
<figureCitation id="12B6DD12F15132D8A32ECCE02AB81A94" captionStart="Figures 13–20" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">17</figureCitation>
|
||
). They differ by the spermathecae with a long stalk that is not coiled and the distal part globular (Figs
|
||
<figureCitation id="E60A14B314AA57F5EEB78495319232EE" captionStart="Figures 13–20" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">14</figureCitation>
|
||
,
|
||
<figureCitation id="DC766314655EEC957BE67B5A47499B4B" captionStart="Figures 13–20" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">17</figureCitation>
|
||
), whereas
|
||
<taxonomicName id="533705F90236F9111E718912263721D7" lsidName="D. canhemabae" pageId="0" pageNumber="273" rank="species" species="canhemabae">
|
||
<emphasis id="4F79B0F7F3F7037E1147402FB88426AE" italics="true" pageId="0" pageNumber="273">D. canhemabae</emphasis>
|
||
</taxonomicName>
|
||
has coiled spermathecae (see
|
||
<bibRefCitation id="F8113BA5985BA1B085A3FD97DFA29895" DOI="https://doi.org/10.11646/zootaxa.697.1.1" author="Brescovit, AD" journalOrPublisher="Zootaxa" pageId="0" pageNumber="273" pagination="1 - 5" refId="B3" refString="Brescovit, AD, Bonaldo, AB, Rheims, CA, 2004. A new species of Drymusa Simon, 1891 (Araneae, Drymusidae) from Brazil. Zootaxa 697 (1): 1 - 5, DOI: https://doi.org/10.11646/zootaxa.697.1.1" title="A new species of Drymusa Simon, 1891 (Araneae, Drymusidae) from Brazil." url="https://doi.org/10.11646/zootaxa.697.1.1" volume="697" year="2004">Brescovit et al. (2004)</bibRefCitation>
|
||
: fig. 5; Fig.
|
||
<figureCitation id="05AF0A778E90A57A8494FF3CEA9E2F2F" captionStart="Figures 21–32" captionStartId="F3" captionText="Figures 21 - 32. Drymusa spp., females. 21. D. simoni, posterior part of opisthosoma, lateral view; 22. D. dinora, opisthosoma, ventral view; 23, 25 - 27. Female genitalia, ventral view (23. D. colligata; 25. D. spectata; 26. D. armasi; 27. D. serrana); 24, 28 - 32. Same, dorsal view (24. D. huberi sp. nov.; 28. D. serrana; 29. D. canhemabae; 30. D. colligata; 31. D. rengan; 32. D. spelunca)." figureDoi="10.3897/zse.99.99227.figures21-32" httpUri="https://binary.pensoft.net/fig/831283" pageId="0" pageNumber="273">29</figureCitation>
|
||
) and
|
||
<taxonomicName id="189FE43C86CA0D72E4A6B1DA3B131B29" lsidName="D. colligata" pageId="0" pageNumber="273" rank="species" species="colligata">
|
||
<emphasis id="E3AC584476D69B6FB49E049663F21C32" italics="true" pageId="0" pageNumber="273">D. colligata</emphasis>
|
||
</taxonomicName>
|
||
has shorter spermatheca, with its distal part swollen (not globular) (see
|
||
<bibRefCitation id="C337CD3C9542B7F7B6A6572E215419A0" DOI="https://doi.org/10.1590/S0101-81752006000200020" author="Bonaldo, AB" journalOrPublisher="Revista Brasileira de Zoologia" pageId="0" pageNumber="273" pagination="455 - 459" refId="B2" refString="Bonaldo, AB, Rheims, CA, Brescovit, AD, 2006. Four new species of Drymusa Simon (Araneae, Drymusidae) from Brazilian Oriental Amazonia. Revista Brasileira de Zoologia 23 (2): 455 - 459, DOI: https://doi.org/10.1590/S0101-81752006000200020" title="Four new species of Drymusa Simon (Araneae, Drymusidae) from Brazilian Oriental Amazonia." url="https://doi.org/10.1590/S0101-81752006000200020" volume="23" year="2006">Bonaldo et al. (2006)</bibRefCitation>
|
||
: fig. 6; Fig.
|
||
<figureCitation id="51352C598E51595C3B0A20B4A6B78ED0" captionStart="Figures 21–32" captionStartId="F3" captionText="Figures 21 - 32. Drymusa spp., females. 21. D. simoni, posterior part of opisthosoma, lateral view; 22. D. dinora, opisthosoma, ventral view; 23, 25 - 27. Female genitalia, ventral view (23. D. colligata; 25. D. spectata; 26. D. armasi; 27. D. serrana); 24, 28 - 32. Same, dorsal view (24. D. huberi sp. nov.; 28. D. serrana; 29. D. canhemabae; 30. D. colligata; 31. D. rengan; 32. D. spelunca)." figureDoi="10.3897/zse.99.99227.figures21-32" httpUri="https://binary.pensoft.net/fig/831283" pageId="0" pageNumber="273">30</figureCitation>
|
||
). Females of
|
||
<taxonomicName id="34BC208458DA8A772C4FA79234492058" lsidName="D. huberi" pageId="0" pageNumber="273" rank="species" species="huberi">
|
||
<emphasis id="35425EE211C3B01F9D22F3BF856BC209" italics="true" pageId="0" pageNumber="273">D. huberi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. also resemble those of
|
||
<taxonomicName id="4FED610605A91736E746D3DAEDC726FE" lsidName="D. dinora" pageId="0" pageNumber="273" rank="species" species="dinora">
|
||
<emphasis id="FA2B77E27685154223DEA37E4E0709E4" italics="true" pageId="0" pageNumber="273">D. dinora</emphasis>
|
||
</taxonomicName>
|
||
Valerio, 1971 by the long and broad epigastric plate, but
|
||
<taxonomicName id="F7ED91D75BF0776C13BF7E6369299999" lsidName="D. huberi" pageId="0" pageNumber="273" rank="species" species="huberi">
|
||
<emphasis id="4D98FDA08140F86FF5D6AE378ED1F6DB" italics="true" pageId="0" pageNumber="273">D. huberi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. has a median hood on the post-epigastric plate, absent in
|
||
<taxonomicName id="10F762592ABC806F80DCA23996A2C784" lsidName="D. dinora" pageId="0" pageNumber="273" rank="species" species="dinora">
|
||
<emphasis id="726F1EA02B8DBEF28168D02368211859" italics="true" pageId="0" pageNumber="273">D. dinora</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="8F35F4EA869E06EE3485097D13917B52" DOI="https://doi.org/10.2307/3493570" author="Valerio, CE" journalOrPublisher="The Florida Entomologist" pageId="0" pageNumber="273" pagination="193 - 200" refId="B10" refString="Valerio, CE, 1971. The spider genus Drymusa in the New World (Araneae: Scytodidae). The Florida Entomologist 54 (2): 193 - 200, DOI: https://doi.org/10.2307/3493570" title="The spider genus Drymusa in the New World (Araneae: Scytodidae)." url="https://doi.org/10.2307/3493570" volume="54" year="1971">Valerio (1971)</bibRefCitation>
|
||
: figs 8, 9; Fig.
|
||
<figureCitation id="FDD1DF2620AEDBA30A8948BFF72F05C0" captionStart="Figures 13–20" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">20</figureCitation>
|
||
),
|
||
<taxonomicName id="BA25FCCDD8BA32DB19B0BF9B07AE32F8" lsidName="D. canhemabae" pageId="0" pageNumber="273" rank="species" species="canhemabae">
|
||
<emphasis id="9D22450E06114F65A013A98F4D2EE6C5" italics="true" pageId="0" pageNumber="273">D. canhemabae</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="F273CECD6B0EE5D959393E4B476356FE" lsidName="D. colligata" pageId="0" pageNumber="273" rank="species" species="colligata">
|
||
<emphasis id="B08D75F3FD1C1F9902F5FC5140D37854" italics="true" pageId="0" pageNumber="273">D. colligata</emphasis>
|
||
</taxonomicName>
|
||
. Males of
|
||
<taxonomicName id="27358FFC5B2CFE29B7FED9D9D2B3184C" lsidName="D. huberi" pageId="0" pageNumber="273" rank="species" species="huberi">
|
||
<emphasis id="0ABC739970E74983F3B033A66F6E31B6" italics="true" pageId="0" pageNumber="273">D. huberi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be easily distinguished from all congeneric species by having the embolus shorter than the bulb length (Figs
|
||
<figureCitation id="38C1BE3D3AA0B30210B367A82BF6E3D5" captionStart="Figures 13–20" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">15</figureCitation>
|
||
,
|
||
<figureCitation id="F5F28BAF2E6C94858E17D3E02C8F221C" captionStart="Figures 13–20" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">16</figureCitation>
|
||
,
|
||
<figureCitation id="E2D6078B550E5A92334FF668FB5E628C" captionStart="Figures 13–20" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">18</figureCitation>
|
||
,
|
||
<figureCitation id="19EFA6A05D95A2E54991D2A4065EF1B8" captionStart="Figures 13–20" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">20</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="527DF77BC822347541438B22784E6C1A" pageId="0" pageNumber="273" type="description">
|
||
<paragraph id="770D9EF1F0F8FEF810C85BA3AA7ABDE6" pageId="0" pageNumber="273">Description.</paragraph>
|
||
<paragraph id="50D9C610CFEA82654A19E4FCC74BD99E" pageId="0" pageNumber="273">
|
||
<emphasis id="0B0A34182290524D8966C60B18241AC4" bold="true" pageId="0" pageNumber="273">
|
||
Female (Holotype).
|
||
<emphasis id="E1363A7B61DA65F2332BDC578748DB44" italics="true" pageId="0" pageNumber="273">Habitus</emphasis>
|
||
.
|
||
</emphasis>
|
||
Carapace pear-shaped, pars cephalica elongated, with many long, procumbent setae (Figs
|
||
<figureCitation id="A499ABE895E84B5D14718E1B74D03B7B" captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">1</figureCitation>
|
||
,
|
||
<figureCitation id="E3F352F8D55520F05C212ECA1BF27DDB" captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">9</figureCitation>
|
||
). Opisthosoma elongated, with posterior two thirds attenuated and covered sparsely with long, brown setae (Fig.
|
||
<figureCitation id="6A08C002FF386E2590ADE6586C1EA0A8" captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">2</figureCitation>
|
||
).
|
||
<emphasis id="7724D11709A4917E996E82D067A51149" bold="true" pageId="0" pageNumber="273">
|
||
<emphasis id="92290A4010697ED6AF7B42FDD0056E1A" italics="true" pageId="0" pageNumber="273">Colouration</emphasis>
|
||
.
|
||
</emphasis>
|
||
Carapace yellowish, margins on pars thoracica tinged with purple-brown and a median broad V-shaped pattern; pars cephalica reticulated with purple-brown maculations (Fig.
|
||
<figureCitation id="0DFFAF7A2A49F69A0DD9F83425FF4CD1" captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">1</figureCitation>
|
||
). Chelicerae orange, with brownish maculations. Endites and labium purple-brown, the latter darker than the former. Sternum purple-brown, except on lateral margins, which are yellowish, with one darker spot on the posterior margin (arrow in Fig.
|
||
<figureCitation id="687C9506ECFE33928543D492CA15E4D8" captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">3</figureCitation>
|
||
). Palp with femur and patella mostly yellowish, tinged with purple-brown on lateral sides; tibia and tarsus mostly purple-brown.
|
||
<emphasis id="7809C60972B44A4DABFCDA21581B4E29" bold="true" italics="true" pageId="0" pageNumber="273">Legs</emphasis>
|
||
: femora mostly purple-brown, with one basal and one distal yellowish ring; patellae mostly purple-brown; tibiae mostly yellowish, with one basal, thin purple-brown ring and one distal purple-brown ring; metatarsi and tarsi yellowish. Dorsum of prosoma grey-white with three W-shaped dark brown bands, posterior third mostly dark brown (Fig.
|
||
<figureCitation id="5107219C9267BE6F68D847F47370D069" captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">2</figureCitation>
|
||
). Venter purple-brown, with pale-yellowish spots on the epigastric area (Fig.
|
||
<figureCitation id="E5148E507DF9DF96215E586FC4B0673E" captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">4</figureCitation>
|
||
).
|
||
<emphasis id="92BBD413927228F56860E9B7445137CA" bold="true" pageId="0" pageNumber="273">
|
||
<emphasis id="8113F2E4F8F563D14858A1C88DE7524B" italics="true" pageId="0" pageNumber="273">Female genitalia</emphasis>
|
||
.
|
||
</emphasis>
|
||
Epigastric plate rectangular, large and about half the opisthosoma length (Figs
|
||
<figureCitation id="415641E55D596CDFAD0C055CB5184F3E" captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">4</figureCitation>
|
||
,
|
||
<figureCitation id="DDEC707FF4894B0DB8FD1DDC6930E7D8" captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">11</figureCitation>
|
||
); posterior corners are more sclerotised than the plate (arrow in Fig.
|
||
<figureCitation id="6BBE32284A5E2CAE1373899FEAE7D4D4" captionStart="Figures 13–20" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">13</figureCitation>
|
||
). Post-epigastric plate crescent-shaped, with a small median hood. Vulva with the distal part of spermatheca globular, on long, thin stalks, arising on lateral corners of the epigastric plate (Figs
|
||
<figureCitation id="8D7A41E64DE6F548C5C50C2D974BC77D" captionStart="Figures 13–20" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">14</figureCitation>
|
||
,
|
||
<figureCitation id="F88B26024D2E5377A2E7E3A7692231FA" captionStart="Figures 13–20" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">17</figureCitation>
|
||
). Spermatheca with gland ductules well-spaced on the distal half (Fig.
|
||
<figureCitation id="73E2FF8AD3F60BA4E05EFF86F83A5D4D" captionStart="Figures 13–20" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">17</figureCitation>
|
||
).
|
||
<emphasis id="F8D2BC99AAF241887C12B4522C75FC8A" bold="true" pageId="0" pageNumber="273">
|
||
<emphasis id="96A43788877A0FB346B2AF071081062E" italics="true" pageId="0" pageNumber="273">Measurements</emphasis>
|
||
.
|
||
</emphasis>
|
||
Carapace 1.32 long, 1.02 wide; eye diameters: PME 0.10, ALE 0.09, PLE 0.06; labium 0.30 long, 0.30 wide; sternum 0.69 long, 0.65 wide. Legs: I 10.8 (2.82, 0.36, 2.97, 3.06, 0.87), II 8.76 (2.46, 0.34, 2.49, 2.65, 0.82), III 6.87 (1.95, 0.39, 1.74, 2.04, 0.75), IV 8.59 (2.35, 0.33, 2.43, 2.61, 0.87). Leg formula: 1243.
|
||
</paragraph>
|
||
<caption id="FEE351AA6A1599D8336A5E652792408D" doi="10.3897/zse.99.99227.figures21-32" httpUri="https://binary.pensoft.net/fig/831283" pageId="0" pageNumber="273" start="Figures 21–32" startId="F3">
|
||
<paragraph id="79AB287055C4A99AEE99BA3FEE921DAF" pageId="0" pageNumber="273">
|
||
<emphasis id="0D84FB3B4F866BA57EC98580038BBBA1" bold="true" pageId="0" pageNumber="273">Figures 21-32.</emphasis>
|
||
<taxonomicName id="8630CD28B523FA39E43CCA0A81CA432E" class="Arachnida" family="Drymusidae" genus="Drymusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymusa" order="Araneae" pageId="0" pageNumber="273" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="07F8A44E5D3733C07AF912591057A65B" italics="true" pageId="0" pageNumber="273">Drymusa</emphasis>
|
||
</taxonomicName>
|
||
spp., females.
|
||
<emphasis id="98C28496532E17774A8B65D6A5B82E1F" bold="true" pageId="0" pageNumber="273">21.</emphasis>
|
||
<taxonomicName id="64224349E4E5F6935D066FB2A2056367" lsidName="D. simoni" pageId="0" pageNumber="273" rank="species" species="simoni">
|
||
<emphasis id="DA4931BF4A837F429B32181C7E406B0C" italics="true" pageId="0" pageNumber="273">D. simoni</emphasis>
|
||
</taxonomicName>
|
||
, posterior part of opisthosoma, lateral view;
|
||
<emphasis id="82A9BDCEB6C44FAB329C64525A995270" bold="true" pageId="0" pageNumber="273">22.</emphasis>
|
||
<taxonomicName id="8E490405E365BE6BB548ACAECF095825" lsidName="D. dinora" pageId="0" pageNumber="273" rank="species" species="dinora">
|
||
<emphasis id="152003BFE4942DE80D7FC2EE5FE008BD" italics="true" pageId="0" pageNumber="273">D. dinora</emphasis>
|
||
</taxonomicName>
|
||
, opisthosoma, ventral view;
|
||
<emphasis id="7E790CF35D0B2A39DCB1A2D014D97563" bold="true" pageId="0" pageNumber="273">23, 25-27.</emphasis>
|
||
Female genitalia, ventral view (
|
||
<emphasis id="296A5C127AA8390CE45B17CFB57AC770" bold="true" pageId="0" pageNumber="273">23.</emphasis>
|
||
<taxonomicName id="6707DF35C58FE4E6F40DB5610D0D1D75" lsidName="D. colligata" pageId="0" pageNumber="273" rank="species" species="colligata">
|
||
<emphasis id="5ABBECFCF61D1EDACC117F230C4F62B4" italics="true" pageId="0" pageNumber="273">D. colligata</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="5BFD8EFA72A743DF9DAF8B7B09664DF4" bold="true" pageId="0" pageNumber="273">25.</emphasis>
|
||
<taxonomicName id="09463F856DB6EEE08218AE3A6292CD99" lsidName="D. spectata" pageId="0" pageNumber="273" rank="species" species="spectata">
|
||
<emphasis id="CDC7434C5A1A1EF93970AAA7E8B3DC23" italics="true" pageId="0" pageNumber="273">D. spectata</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="74BE655930E3F06867A0B24D4D5EA6D4" bold="true" pageId="0" pageNumber="273">26.</emphasis>
|
||
<taxonomicName id="A8457C164F0A23E8C10B8A61DE204162" lsidName="D. armasi" pageId="0" pageNumber="273" rank="species" species="armasi">
|
||
<emphasis id="FE5CB009C2B4397741ABE1384C461BC7" italics="true" pageId="0" pageNumber="273">D. armasi</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="3A37B9E83FD74A00A814C35A1D06C638" bold="true" pageId="0" pageNumber="273">27.</emphasis>
|
||
<taxonomicName id="33BE84A7ABA673116D8F25DCCED6226F" lsidName="D. serrana" pageId="0" pageNumber="273" rank="species" species="serrana">
|
||
<emphasis id="D358308A40E4CC94290978E0239911F8" italics="true" pageId="0" pageNumber="273">D. serrana</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<emphasis id="30DFC9DEECDE02536D0CF770CD4C0C54" bold="true" pageId="0" pageNumber="273">24, 28-32.</emphasis>
|
||
Same, dorsal view (
|
||
<emphasis id="D85CFD675B5233666076FA165CA218BB" bold="true" pageId="0" pageNumber="273">24.</emphasis>
|
||
<taxonomicName id="AE52A519D2136379461823A12CA394E7" lsidName="D. huberi" pageId="0" pageNumber="273" rank="species" species="huberi">
|
||
<emphasis id="37A8660D3ED7A877706DD363E34F3A9F" italics="true" pageId="0" pageNumber="273">D. huberi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.;
|
||
<emphasis id="B7563EF3E1477F3780BDA9D6E9266A3A" bold="true" pageId="0" pageNumber="273">28.</emphasis>
|
||
<taxonomicName id="DCE46C0E6E0F085C1D8FB2277EF5914B" lsidName="D. serrana" pageId="0" pageNumber="273" rank="species" species="serrana">
|
||
<emphasis id="B87191EA43E2F7AC18EF758F5DD12E52" italics="true" pageId="0" pageNumber="273">D. serrana</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="1C36D96995028CCD867799C7F0AE8609" bold="true" pageId="0" pageNumber="273">29.</emphasis>
|
||
<taxonomicName id="271EADCD5D7166C19BC349F22DF3EA72" lsidName="D. canhemabae" pageId="0" pageNumber="273" rank="species" species="canhemabae">
|
||
<emphasis id="65FD8B14B69B38A46CAC009C4F9F1AAF" italics="true" pageId="0" pageNumber="273">D. canhemabae</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="E865D6238435FEEEA4EB77CE7A742FA2" bold="true" pageId="0" pageNumber="273">30.</emphasis>
|
||
<taxonomicName id="CD0CD7373A77011472CBC240350D44A1" lsidName="D. colligata" pageId="0" pageNumber="273" rank="species" species="colligata">
|
||
<emphasis id="5650063B01C9949AA2518C891517FA79" italics="true" pageId="0" pageNumber="273">D. colligata</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="A3F7F27DC78A7C75526A4DF49DD29990" bold="true" pageId="0" pageNumber="273">31.</emphasis>
|
||
<taxonomicName id="CF145BA58195CA6568950173AF611528" lsidName="D. rengan" pageId="0" pageNumber="273" rank="species" species="rengan">
|
||
<emphasis id="4911B9BE39989DA37124BE9A9A5F854A" italics="true" pageId="0" pageNumber="273">D. rengan</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="0AB8A8506C8A6DC5E4F8A28EB61AC232" bold="true" pageId="0" pageNumber="273">32.</emphasis>
|
||
<taxonomicName id="10AA0F6335447248BCBD5CF84D21A634" lsidName="D. spelunca" pageId="0" pageNumber="273" rank="species" species="spelunca">
|
||
<emphasis id="2D4D0AC3128E10D2DB553872E6CC9649" italics="true" pageId="0" pageNumber="273">D. spelunca</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="432880FEFFBCD0317D62EA510B5808A8" doi="10.3897/zse.99.99227.figures33-48" httpUri="https://binary.pensoft.net/fig/831284" pageId="0" pageNumber="273" start="Figures 33–48" startId="F4">
|
||
<paragraph id="6B8897107F455A763244EFE42CE70017" pageId="0" pageNumber="273">
|
||
<emphasis id="B2D2B4076CCA275C4B74812D38D6B1C6" bold="true" pageId="0" pageNumber="273">Figures 33-48.</emphasis>
|
||
<taxonomicName id="81F5344EE653A0040B05680BCBD65509" class="Arachnida" family="Drymusidae" genus="Drymusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymusa" order="Araneae" pageId="0" pageNumber="273" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64157AB0415F75FB206D85205DC4495F" italics="true" pageId="0" pageNumber="273">Drymusa</emphasis>
|
||
</taxonomicName>
|
||
spp., male palp (
|
||
<emphasis id="BB656DCB452DF6F439199D39D6C4617D" bold="true" pageId="0" pageNumber="273">33, 34.</emphasis>
|
||
<taxonomicName id="9D7CB2ADDA66DBD1C9AC78D54F3C1EAC" lsidName="D. rengan" pageId="0" pageNumber="273" rank="species" species="rengan">
|
||
<emphasis id="1B9255908024CF0CCB4C4C38C85D8BBF" italics="true" pageId="0" pageNumber="273">D. rengan</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="26E27E7155BE47A24707C8F1A0378FC9" bold="true" pageId="0" pageNumber="273">35, 36.</emphasis>
|
||
<taxonomicName id="10B5F0E2FDF7676090FC92E26D97AB39" lsidName="D. serrana" pageId="0" pageNumber="273" rank="species" species="serrana">
|
||
<emphasis id="57D38A78159D7694FFEE6E2BCBBCABFD" italics="true" pageId="0" pageNumber="273">D. serrana</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="609204BB85EB99195554D1360F8084C3" bold="true" pageId="0" pageNumber="273">37, 38.</emphasis>
|
||
<taxonomicName id="A75B8D903D34A4E1E4D6A1CD6651B7CE" lsidName="D. spelunca" pageId="0" pageNumber="273" rank="species" species="spelunca">
|
||
<emphasis id="A53FE4070FB78E70A0014FE26A29CADE" italics="true" pageId="0" pageNumber="273">D. spelunca</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="DC759C47F73C54AE5F5E1494084767AC" bold="true" pageId="0" pageNumber="273">39.</emphasis>
|
||
<taxonomicName id="5B834AF257DB60A588EDBE43DD552468" lsidName="D. dinora" pageId="0" pageNumber="273" rank="species" species="dinora">
|
||
<emphasis id="FA1EDF9E3C7856A9F8D5B96643CEABC7" italics="true" pageId="0" pageNumber="273">D. dinora</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="6E9D7626A2558CD2559BCEA5C3DCB1D3" bold="true" pageId="0" pageNumber="273">40.</emphasis>
|
||
<taxonomicName id="AFA8DA3D2A77C8EC0B43EE37E943B7BE" lsidName="D. armasi" pageId="0" pageNumber="273" rank="species" species="armasi">
|
||
<emphasis id="514F74D0B143BFB40E91EFD4FD402273" italics="true" pageId="0" pageNumber="273">D. armasi</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="AD55FFBD8FDEB0EB8189AEC5B3629080" bold="true" pageId="0" pageNumber="273">41.</emphasis>
|
||
<taxonomicName id="1AC43A86760C2CD25A9BB25E88FDDDD0" lsidName="D. spectata" pageId="0" pageNumber="273" rank="species" species="spectata">
|
||
<emphasis id="58CDEF63F0BDE8D613F98A0E6FA6D921" italics="true" pageId="0" pageNumber="273">D. spectata</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="09D7CCF2A2C36860CDC02B70FA8C83F5" bold="true" pageId="0" pageNumber="273">42.</emphasis>
|
||
<taxonomicName id="3282263E763A42B1BCCA1ADA1D223516" lsidName="D. tobyi" pageId="0" pageNumber="273" rank="species" species="tobyi">
|
||
<emphasis id="E1651FBA52785234990C0AB076FFDEFC" italics="true" pageId="0" pageNumber="273">D. tobyi</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="9B42A929D07CABFBBE5C09D3312F53C0" bold="true" pageId="0" pageNumber="273">43.</emphasis>
|
||
<taxonomicName id="CDAB0F17C5173DB997A3BEFEDE49BB17" lsidName="D. canhemabae" pageId="0" pageNumber="273" rank="species" species="canhemabae">
|
||
<emphasis id="168A0F1DA664CE16BC0B651CBE140F3D" italics="true" pageId="0" pageNumber="273">D. canhemabae</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="5FB3758B4662CA1EF2614851A05619FB" bold="true" pageId="0" pageNumber="273">44, 45.</emphasis>
|
||
<taxonomicName id="B13D3B80BE2E5FAF1BA9FB80DF19E284" lsidName="D. colligata" pageId="0" pageNumber="273" rank="species" species="colligata">
|
||
<emphasis id="157150F1B82F39FBFCB1F281F5FBF2E7" italics="true" pageId="0" pageNumber="273">D. colligata</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="67AF069DC5459F757AD499FC5DA1584B" bold="true" pageId="0" pageNumber="273">46, 47.</emphasis>
|
||
<taxonomicName id="9ADB74156C6E20F9E3AE2992679CDE21" lsidName="D. philomatica" pageId="0" pageNumber="273" rank="species" species="philomatica">
|
||
<emphasis id="F2E0BCD629240D7A0DA103459D38D82C" italics="true" pageId="0" pageNumber="273">D. philomatica</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="F84786306EBFF31F3092E8467E18C2AF" bold="true" pageId="0" pageNumber="273">48.</emphasis>
|
||
<taxonomicName id="A37352A110EE7E71090118A75D5FF51A" lsidName="D. simoni" pageId="0" pageNumber="273" rank="species" species="simoni">
|
||
<emphasis id="9F57D92F1130DA1E24622832D8E17BCC" italics="true" pageId="0" pageNumber="273">D. simoni</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<emphasis id="309BFA1A59F69278763ADF6B17BDAD6D" bold="true" pageId="0" pageNumber="273">33, 35, 37, 40, 41, 44, 46, 48.</emphasis>
|
||
Palp, prolateral view;
|
||
<emphasis id="617D80A2FA17ECB3D45E863AE85DC9FA" bold="true" pageId="0" pageNumber="273">34, 36, 38, 39, 42, 43, 45, 47.</emphasis>
|
||
Palp, retrolateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="92670CBC8DD807FD999F8517FEFBEBD1" doi="10.3897/zse.99.99227.figures49-53" httpUri="https://binary.pensoft.net/fig/831285" pageId="0" pageNumber="273" start="Figures 49–53" startId="F5">
|
||
<paragraph id="0552AFA07AD179BBFFEAF3E8FA3DC67E" pageId="0" pageNumber="273">
|
||
<emphasis id="68984795D5958CC5B467B24A26750877" bold="true" pageId="0" pageNumber="273">Figures 49-53.</emphasis>
|
||
<emphasis id="27B1735487D97DFF7FAB6159EF88D4B9" bold="true" pageId="0" pageNumber="273">49, 50.</emphasis>
|
||
<taxonomicName id="4903DE53BAC05DFACC7FE39349F3FB17" authorityName="Villarreal & Chamé-Vázquez" authorityYear="2023" class="Arachnida" family="Drymusidae" genus="Drymusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymusa huberi" order="Araneae" pageId="0" pageNumber="273" phylum="Arthropoda" rank="species" species="huberi">
|
||
<emphasis id="CEFF587A12005DB675C306D66A91EDA6" italics="true" pageId="0" pageNumber="273">Drymusa huberi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. live specimens from La
|
||
<normalizedToken id="79D184D6B5E18B9B675324875FB57486" originalValue="Guáquira">Guaquira</normalizedToken>
|
||
, Yaracuy State, Venezuela.
|
||
<emphasis id="C7C0145E390135FB4CE2DA0E7F9B85DC" bold="true" pageId="0" pageNumber="273">49.</emphasis>
|
||
Male, paratype;
|
||
<emphasis id="444A810BE4E903D0753FFE36019D1E36" bold="true" pageId="0" pageNumber="273">50.</emphasis>
|
||
Female, holotype;
|
||
<emphasis id="BCAF62D836AEE827C41F29CDC4D005C7" bold="true" pageId="0" pageNumber="273">51.</emphasis>
|
||
Habitat, tropical dry forest in La
|
||
<normalizedToken id="4E8CAF7B96017ACB2DE3A8A8C137FDE5" originalValue="Guáquira">Guaquira</normalizedToken>
|
||
, Yaracuy State;
|
||
<emphasis id="09A3E52DAD51A2BF1F233629D48D7C9C" bold="true" pageId="0" pageNumber="273">52.</emphasis>
|
||
Leaf litter on the forest floor, microhabitat of the species;
|
||
<emphasis id="23F8238D7ED135AEB4EE2EFC211B5B54" bold="true" pageId="0" pageNumber="273">53.</emphasis>
|
||
Known distribution of all species of the genus
|
||
<taxonomicName id="DBC5847C86AAE8A0ADF3EFCBE0835CFC" class="Arachnida" family="Drymusidae" genus="Drymusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymusa" order="Araneae" pageId="0" pageNumber="273" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E45C911139B4FAE90242D7BE77C30AF6" italics="true" pageId="0" pageNumber="273">Drymusa</emphasis>
|
||
</taxonomicName>
|
||
. Photo credit: O. Villarreal:
|
||
<emphasis id="8F2FA5C82DF3C2C5755D5165137CE7DC" bold="true" pageId="0" pageNumber="273">49, 50, 51</emphasis>
|
||
; D. Romero:
|
||
<emphasis id="38566DF62876C86802DC2CCE87D20D84" bold="true" pageId="0" pageNumber="273">52</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2931053C72CCA2CA2C94F837F8B79BA8" pageId="0" pageNumber="273">
|
||
<emphasis id="CD77EB92D98915321B0E70EF74469D00" bold="true" pageId="0" pageNumber="273">Male (paratype).</emphasis>
|
||
Habitus as the female (Figs
|
||
<figureCitation id="2DE00CBD3F8F877FFA67066EBB16C1F8" captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">5-8</figureCitation>
|
||
,
|
||
<figureCitation id="A86AD056D7DF7BE020897A33C05E80D4" captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">10</figureCitation>
|
||
,
|
||
<figureCitation id="12E968B3938B1E29E4C5AD9DD4C4DDE4" captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">12</figureCitation>
|
||
), except pars cephalic with few short setae, mostly on clypeal area, opisthosoma with posterior third attenuated.
|
||
<emphasis id="872F02AFF07179D5CB4134459094FE3E" bold="true" pageId="0" pageNumber="273">
|
||
<emphasis id="F04F0B6B8797C8E5C2F5C8A9A870A0FA" italics="true" pageId="0" pageNumber="273">Colouration</emphasis>
|
||
.
|
||
</emphasis>
|
||
As the female, but the purple-brown median pattern and lateral margins are narrower; the sternum maculation is light purple; palp yellowish with scattered purple-brown maculations, tarsus darker than other segments. Dorsum of opisthosoma with transversal bands narrower, elongated, fused with the next one; venter yellowish, except posterior third, which is light purple.
|
||
<emphasis id="10E97C87F4559B15991DF03175A27BF8" bold="true" pageId="0" pageNumber="273">
|
||
<emphasis id="22730264EF9A4A257155F2D721AF4284" italics="true" pageId="0" pageNumber="273">Palp</emphasis>
|
||
.
|
||
</emphasis>
|
||
Sub-cylindrical femur longer than tibia; tibia slightly more thickened than femur and slightly swollen at mid-length (Figs
|
||
<figureCitation id="5DE5DB6FC5134D2D05DAC9F812D4CD8C" captionStart="Figures 13–20" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">15</figureCitation>
|
||
,
|
||
<figureCitation id="B658829499E7823CAF98D11E6D807620" captionStart="Figures 13–20" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">16</figureCitation>
|
||
,
|
||
<figureCitation id="CE5E923F575972ACB9F2D2CE3D013476" captionStart="Figures 13–20" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">20</figureCitation>
|
||
). Bulb subspherical, slightly longer than wide (Figs
|
||
<figureCitation id="B935D0A46D88EC015290829D1BF27EB3" captionStart="Figures 13–20" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">15</figureCitation>
|
||
,
|
||
<figureCitation id="691501280D13042B797E951903203420" captionStart="Figures 13–20" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">16</figureCitation>
|
||
,
|
||
<figureCitation id="C441EA1EE868C585B2756D543BF11AA8" captionStart="Figures 13–20" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">18</figureCitation>
|
||
). Embolus spiniform, curved, directed ventral-retrolaterally and relatively short, about half the length of bulb (Figs
|
||
<figureCitation id="3A99F93F07EB4BEFDC23C718518B95AA" captionStart="Figures 13–20" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">18</figureCitation>
|
||
,
|
||
<figureCitation id="5DCA148ABE6011D97EB4DC094239468A" captionStart="Figures 13–20" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">19</figureCitation>
|
||
).
|
||
<emphasis id="6E6970725525891D1705414820DD388F" bold="true" pageId="0" pageNumber="273">
|
||
<emphasis id="088EB7869A30E0F4603E225ECC6412ED" italics="true" pageId="0" pageNumber="273">Measurements</emphasis>
|
||
.
|
||
</emphasis>
|
||
Carapace 1.26 long, 1.05 wide; eye diameters: PME 0.12, ALE 0.09, PLE 0.07; labium 0.24 long, 0.27 wide; sternum 0.72 long, 0.72 wide. Legs: I missing, II 10.62 (2.97, 0.39, 3.09, 3.24, 0.93), III 7.77 (2.13, 0.36, 2.04, 2.40, 0.84), IV 10.20 (2.82, 0.36, 2.82, 3.09, 0.93). Leg formula: (1)243.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1AC46AFD13E9078061B68D74617D8D18" pageId="0" pageNumber="273" type="distribution">
|
||
<paragraph id="1A5526D0414E8A9A13603C2691270AAE" pageId="0" pageNumber="273">Distribution.</paragraph>
|
||
<paragraph id="F8505F4CF6EBF8E7BF2FA37F14C0D5A5" pageId="0" pageNumber="273">
|
||
Only known from the type locality. See Fig.
|
||
<figureCitation id="B2D341439BA7C96BE21D1CE10EACE683" captionStart="Figures 49–53" captionStartId="F5" captionText="Figures 49 - 53. 49, 50. Drymusa huberi sp. nov. live specimens from La Guaquira, Yaracuy State, Venezuela. 49. Male, paratype; 50. Female, holotype; 51. Habitat, tropical dry forest in La Guaquira, Yaracuy State; 52. Leaf litter on the forest floor, microhabitat of the species; 53. Known distribution of all species of the genus Drymusa. Photo credit: O. Villarreal: 49, 50, 51; D. Romero: 52." figureDoi="10.3897/zse.99.99227.figures49-53" httpUri="https://binary.pensoft.net/fig/831285" pageId="0" pageNumber="273">53</figureCitation>
|
||
for the known distribution of all
|
||
<taxonomicName id="6BCE09C6603EE2F9A8092478C8158AE7" class="Arachnida" family="Drymusidae" genus="Drymusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymusa" order="Araneae" pageId="0" pageNumber="273" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7E75994FC2749D1433CFA8E344C28BCB" italics="true" pageId="0" pageNumber="273">Drymusa</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9B849F3467AA49F5C8AB6E21231B8D4F" pageId="0" pageNumber="273" type="natural history">
|
||
<paragraph id="8FBEF2159F4C003241E873511C79BCAE" pageId="0" pageNumber="273">Natural history.</paragraph>
|
||
<paragraph id="42593834E5816772C167EE8436D9BDBD" pageId="0" pageNumber="273">
|
||
Both specimens were found in the leaf litter in a tropical dry forest along a streambed (Figs
|
||
<figureCitation id="79C26EC42118FBEB3B1BAC697B5DC633" captionStart="Figures 49–53" captionStartId="F5" captionText="Figures 49 - 53. 49, 50. Drymusa huberi sp. nov. live specimens from La Guaquira, Yaracuy State, Venezuela. 49. Male, paratype; 50. Female, holotype; 51. Habitat, tropical dry forest in La Guaquira, Yaracuy State; 52. Leaf litter on the forest floor, microhabitat of the species; 53. Known distribution of all species of the genus Drymusa. Photo credit: O. Villarreal: 49, 50, 51; D. Romero: 52." figureDoi="10.3897/zse.99.99227.figures49-53" httpUri="https://binary.pensoft.net/fig/831285" pageId="0" pageNumber="273">51</figureCitation>
|
||
,
|
||
<figureCitation id="FCC7080809A5783C3FE196091CB9719F" captionStart="Figures 49–53" captionStartId="F5" captionText="Figures 49 - 53. 49, 50. Drymusa huberi sp. nov. live specimens from La Guaquira, Yaracuy State, Venezuela. 49. Male, paratype; 50. Female, holotype; 51. Habitat, tropical dry forest in La Guaquira, Yaracuy State; 52. Leaf litter on the forest floor, microhabitat of the species; 53. Known distribution of all species of the genus Drymusa. Photo credit: O. Villarreal: 49, 50, 51; D. Romero: 52." figureDoi="10.3897/zse.99.99227.figures49-53" httpUri="https://binary.pensoft.net/fig/831285" pageId="0" pageNumber="273">52</figureCitation>
|
||
); this species shares the microhabitat with
|
||
<taxonomicName id="9294314BCA6EAB40E5EEEC5F94838D0C" class="Arachnida" family="Ochyroceratidae" genus="Ochyrocera" kingdom="Animalia" lsidName="Ochyrocera" order="Araneae" pageId="0" pageNumber="273" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D4427FDFDF3AE9E681FC1EDA5B736FE5" italics="true" pageId="0" pageNumber="273">Ochyrocera</emphasis>
|
||
</taxonomicName>
|
||
sp. (
|
||
<taxonomicName id="FC482BA9D8C0EE092407DA360C8E8749" authorityName="Fage" authorityYear="1912" class="Arachnida" family="Ochyroceratidae" kingdom="Animalia" lsidName="" order="Araneae" pageId="0" pageNumber="273" phylum="Arthropoda" rank="family">Ochyroceratidae</taxonomicName>
|
||
). For live specimens of
|
||
<taxonomicName id="08B8751F01FE99CF78B5839BBA049FF8" lsidName="D. huberi" pageId="0" pageNumber="273" rank="species" species="huberi">
|
||
<emphasis id="F7ED4189FF031456D33B96325F1FE7A7" italics="true" pageId="0" pageNumber="273">D. huberi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., see Figs
|
||
<figureCitation id="E7F208A6AA86EC59548544C8447EA7EA" captionStart="Figures 49–53" captionStartId="F5" captionText="Figures 49 - 53. 49, 50. Drymusa huberi sp. nov. live specimens from La Guaquira, Yaracuy State, Venezuela. 49. Male, paratype; 50. Female, holotype; 51. Habitat, tropical dry forest in La Guaquira, Yaracuy State; 52. Leaf litter on the forest floor, microhabitat of the species; 53. Known distribution of all species of the genus Drymusa. Photo credit: O. Villarreal: 49, 50, 51; D. Romero: 52." figureDoi="10.3897/zse.99.99227.figures49-53" httpUri="https://binary.pensoft.net/fig/831285" pageId="0" pageNumber="273">49</figureCitation>
|
||
,
|
||
<figureCitation id="766966D9336E2FCFCA8733147F7823AF" captionStart="Figures 49–53" captionStartId="F5" captionText="Figures 49 - 53. 49, 50. Drymusa huberi sp. nov. live specimens from La Guaquira, Yaracuy State, Venezuela. 49. Male, paratype; 50. Female, holotype; 51. Habitat, tropical dry forest in La Guaquira, Yaracuy State; 52. Leaf litter on the forest floor, microhabitat of the species; 53. Known distribution of all species of the genus Drymusa. Photo credit: O. Villarreal: 49, 50, 51; D. Romero: 52." figureDoi="10.3897/zse.99.99227.figures49-53" httpUri="https://binary.pensoft.net/fig/831285" pageId="0" pageNumber="273">50</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |