359 lines
41 KiB
XML
359 lines
41 KiB
XML
<document id="4E88657C3BD4F5A6E94F8405989D4546" ID-DOI="10.1649/0010-065x(2001)055[0011:snsops]2.0.co;2" ID-ISSN="1938-4394" ID-Zenodo-Dep="4900311" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1622827801739" checkinUser="carolina" docAuthor="Morón, Miguel Angel & Solís, Angel" docDate="2001" docId="03A887FE5D46152A2D70D613FCBCFAA8" docLanguage="en" docName="TheColeopBull.55.1.11-29.pdf.imf" docOrigin="The Coleopterists Bulletin 55 (1)" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Phyllophaga (Phyllophaga) tilarana Moron and Solis, new species" docType="treatment" docVersion="1" lastPageNumber="14" masterDocId="FF91FF865D4715292C3FD44FFF9EFFD8" masterDocTitle="Seven New Species Of Phyllophaga (S. Str.) Harris From Costa Rica (Coleoptera: Melolonthidae: Melolonthinae)" masterLastPageNumber="29" masterPageNumber="11" pageNumber="12" updateTime="1730139982923" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:titleInfo id="15DF41BCC8696E8DF2E2392ECF90CC5A">
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<mods:title id="5766B8A1B8508BC64D10BD2E0D3B4E18">Seven New Species Of Phyllophaga (S. Str.) Harris From Costa Rica (Coleoptera: Melolonthidae: Melolonthinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="B5126C5A62E4A59C1965F98BC63290F2" type="personal">
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<mods:role id="89CE43A088B1A51B40C7EFD86AEB2255">
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<mods:roleTerm id="2FEA114845FC2A5B768CCD78DF8463A2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EF4D3E3DBF2C64A16DC8886A383FF04A">Morón, Miguel Angel</mods:namePart>
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<mods:affiliation id="63219CD585E1022F92CFDFDECA23D140">Departamento de Entomología Instituto de Ecología, A. C. (sector SEPCONACYT) Apartado Postal 63 Xalapa, Veracruz 91000, MÉXICO</mods:affiliation>
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</mods:name>
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<mods:name id="5F4568EE010BBC074BD23A31E5DDB51E" type="personal">
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<mods:role id="5D07A6A353AF7A4776CEAC3E97F020C3">
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<mods:roleTerm id="1DFCC79F57194DE1B629501736DE904D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="82EEBD3A3EB184C520AEF3965CD4CCAA">Solís, Angel</mods:namePart>
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<mods:affiliation id="8C3F63FE64B2EDD614D0115C27FD4E2F">Departamento de Entomología Instituto Nacional de Biodiversidad (INBio) Apartado Postal 22 3100 Santo Domingo de Heredia, COSTA RICA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="183D83439079FA2F4261F92A419660E5">text</mods:typeOfResource>
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<mods:relatedItem id="3B3271E5C028C994792606B03954E383" type="host">
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<mods:titleInfo id="FC41CF6620D258226DEBC83537C5E643">
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<mods:title id="D955A6E86977F385730D00BAE7B22300">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="B3ABBFCF3E78D6501497837BBBB54461">
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<mods:date id="ADC2D60D9C51484AA36E00F5628035E7">2001</mods:date>
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<mods:detail id="12C2F24F89B7EAB6FE1FFFC251904326" type="pubDate">
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<mods:number id="9696209F7A7DE7FA51476603CB40A099">2001-03-01</mods:number>
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</mods:detail>
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<mods:detail id="1D515229DDCA96221CC82A2AAA8014B4" type="volume">
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<mods:number id="BF9A4D314B89BC268619FC508B9CDC38">55</mods:number>
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</mods:detail>
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<mods:detail id="F13D2C7FA62011F4E379FB6F9EE290ED" type="issue">
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<mods:number id="861215B74ED3C39B76DAD42B8F07CE41">1</mods:number>
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<mods:start id="3ECE7DCF5CE11CF13E4FFE356CF9A731">11</mods:start>
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<mods:end id="A1EF9F9C2C753E9DB1B08538D875ED1E">29</mods:end>
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</mods:extent>
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<mods:classification id="D3648225D51008D92DE58F40D0B8F131">journal article</mods:classification>
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<mods:identifier id="7669E4DD7AFD9EB5E80B7EE849957A2B" type="DOI">10.1649/0010-065x(2001)055[0011:snsops]2.0.co;2</mods:identifier>
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<mods:identifier id="6BA3B20594995D2C39F48C87ABB9D7BA" type="ISSN">1938-4394</mods:identifier>
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<mods:identifier id="F3D0A4E64C1F42182DE7F288C6096DA1" type="Zenodo-Dep">4900311</mods:identifier>
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</mods:mods>
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<treatment id="03A887FE5D46152A2D70D613FCBCFAA8" LSID="urn:lsid:plazi:treatment:03A887FE5D46152A2D70D613FCBCFAA8" httpUri="http://treatment.plazi.org/id/03A887FE5D46152A2D70D613FCBCFAA8" lastPageId="3" lastPageNumber="14" pageId="1" pageNumber="12">
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<subSubSection id="C31B65635D4615282D70D613FBBEFDA8" box="[335,1056,603,625]" pageId="1" pageNumber="12" type="nomenclature">
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<paragraph id="8BBE36E85D4615282D70D613FBBEFDA8" blockId="1.[335,1056,603,651]" box="[335,1056,603,625]" pageId="1" pageNumber="12">
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<taxonomicName id="4C014D6B5D4615282D70D613FC0EFDA8" authority="Moron and Solis" authorityName="Moron and Solis" box="[335,912,603,625]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="12" phylum="Arthropoda" rank="species" species="tilarana" status="sp. nov." subGenus="Phyllophaga">
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<emphasis id="B975EAFA5D4615282D70D613FE4BFDA9" box="[335,469,604,625]" italics="true" pageId="1" pageNumber="12">Phyllophaga</emphasis>
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(
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<emphasis id="B975EAFA5D4615282DD8D613FDF2FDA9" box="[487,620,604,625]" italics="true" pageId="1" pageNumber="12">Phyllophaga</emphasis>
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)
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<emphasis id="B975EAFA5D4615282E42D613FD4FFDA9" box="[637,721,604,625]" italics="true" pageId="1" pageNumber="12">tilarana</emphasis>
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Morón and Solís
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</taxonomicName>
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,
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<emphasis id="B975EAFA5D4615282FA2D614FBBEFDA8" bold="true" box="[925,1056,603,624]" pageId="1" pageNumber="12">
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<taxonomicNameLabel id="A24657815D4615282FA2D614FBBEFDA8" box="[925,1056,603,624]" pageId="1" pageNumber="12" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31B65635D46152A2EB9D639FDFFFDB2" lastPageId="3" lastPageNumber="14" pageId="1" pageNumber="12" type="description">
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<paragraph id="8BBE36E85D4615282EB9D639FD77FD53" blockId="1.[335,1056,603,651]" box="[646,745,630,651]" pageId="1" pageNumber="12">
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<figureCitation id="133A2A6D5D4615282EB9D639FD77FD53" box="[646,745,630,651]" captionStart="Figs" captionStartId="2.[330,373,1228,1247]" captionTargetBox="[309,1134,273,1204]" captionTargetId="figure-12@2.[308,1137,272,1208]" captionTargetPageId="2" captionText="Figs. 1–5. Phyllophaga tilarana. 1) protarsal claw, male; 2) genital capsule, lateral view; 3) paramera, distal view; 4) protarsal claw, female; 5) genital plates, female. Scale lines 5 1 mm, except Figs. 1, 4 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900313" httpUri="https://zenodo.org/record/4900313/files/figure.png" pageId="1" pageNumber="12">Figs. 1–5</figureCitation>
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</paragraph>
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<paragraph id="8BBE36E85D46152B2D10D6E8FBEAFA40" blockId="1.[279,1113,679,1658]" lastBlockId="2.[307,1140,1389,1432]" lastPageId="2" lastPageNumber="13" pageId="1" pageNumber="12">
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<emphasis id="B975EAFA5D4615282D10D6E8FE11FD62" bold="true" box="[303,399,679,698]" pageId="1" pageNumber="12">
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<typeStatus id="54BA884A5D4615282D10D6E8FE15FD62" box="[303,395,679,698]" pageId="1" pageNumber="12" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Male. Clypeus, frons and pronotum shiny dark reddish brown; elytra silky, light brown, without macroscopic vestiture; mouthparts, sterna, pygidium and legs shiny dark reddish brown. Clypeus 3.2× wider than long, anterior border scarcely sinuate, with elevated margin, surface slightly irregular, with many uniformly distributed, deep, round shaped punctures and some scattered short setae. Frontoclypeal suture sinuate and deeply impressed. Frons 1.6× wider than long, convex, irregularly and shallowly punctate, with short slender setae at sides and on disk. Antenna 10segmented, with 3segmented club, lamellae of 8th to 10th segments 2.0× longer than length of preceeding 6 segments combined. Frons 3.2× wider than dorsal diameter of eye. Eye canthus long and narrow, with 16–17 setae. Labrum bilobed, deeply sinuate, with scattered long slender setae. Mentum slightly concave, impunctate, with scarce lateral setae, anterior border briefly sinuate. Pronotum 1.7× wider than long and 3.0× wider than frons. Pronotal disk shiny, with round, shallow punctures regularly separated from one another by 1–2 diameters; lateral borders widely angulated, lateral marginal bead nearly entire at anterior half and slightly crenulate at basal half, with scattered, long, slender setae; basal bead strongly indicated, except in front of scutellum where it is indicated only by punctures; anterior angles obtuse, not prominent; posterior angles obtuse, not prominent. Scutellum 1.6× wider than long, with some minute punctures. Elytron 2.9× longer than wide, silkshiny, densely rugopunctate; epipleural border very narrow, extended along complete margin, provided with scattered, short, slender setae; humeral callus rounded, prominent; apical callus rounded. Metathoracic wings completely developed. Propygidium shiny, with scattered, short setae, and dense punctuation. Pygidium very convex, shiny, with round, shallow punctures regularly distributed, with erect setae, shortened toward the basal border; apical margin with 22 long, slender setae; basal margin effaced medially. Pterosterna with long, dense, yellowish setae. Visible abdominal sternites II and IV slightly depressed, with numerous short setae near the middline; sternite V with a transverse, granulated, setose ridge, and scattered setae toward the sides; anal plate large, with wide transverse concavity, with numerous erect setae, posterior border thickened with a transverse row of long setae. Protibia shorter than protarsus (1:1.7), with external border tridentate, proximal tooth much shortened, preapical spur nearly straight, apex rounded, half as long as 2th protarsomere. Mesotibia with one oblique, sharp, setiferous carina on external side; upper apical spur straight, narrow, and 1.3× longer than lower spur. Metatibia one fifth shorter than metatarsus, with one oblique, sharp, setiferous carina on external side; upper apical spur articulated, recurved, apex rounded, slightly longer than basal metatarsomere, and 1.2× longer than lower spur; lower apical spur articulated with tibial border, with rounded apex. Tarsomeres semicylindrical, elongate, with enlarged apices, with some setae apically and two dense lines of long, thick setae on ventral side. Tarsal claws symmetrical, similar on all legs, with an acute short tooth located close to basal dilation (
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<figureCitation id="133A2A6D5D4615282D9ED278FE46F992" box="[417,472,1591,1610]" captionStart="Figs" captionStartId="2.[330,373,1228,1247]" captionTargetBox="[309,1134,273,1204]" captionTargetId="figure-12@2.[308,1137,272,1208]" captionTargetPageId="2" captionText="Figs. 1–5. Phyllophaga tilarana. 1) protarsal claw, male; 2) genital capsule, lateral view; 3) paramera, distal view; 4) protarsal claw, female; 5) genital plates, female. Scale lines 5 1 mm, except Figs. 1, 4 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900313" httpUri="https://zenodo.org/record/4900313/files/figure.png" pageId="1" pageNumber="12">Fig. 1</figureCitation>
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). Genital capsule with large parameres, dorsally and ventrally fused, symmetrical, pentagonal shaped, apex of each elongated, acute, with toothlike preapical projections. Aedeagus narrowed, with scarce sclerotized preapical structures (
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<figureCitation id="133A2A6D5D4615282FCCD228FBD3F9A2" box="[1011,1101,1639,1658]" captionStart="Figs" captionStartId="2.[330,373,1228,1247]" captionTargetBox="[309,1134,273,1204]" captionTargetId="figure-12@2.[308,1137,272,1208]" captionTargetPageId="2" captionText="Figs. 1–5. Phyllophaga tilarana. 1) protarsal claw, male; 2) genital capsule, lateral view; 3) paramera, distal view; 4) protarsal claw, female; 5) genital plates, female. Scale lines 5 1 mm, except Figs. 1, 4 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900313" httpUri="https://zenodo.org/record/4900313/files/figure.png" pageId="1" pageNumber="12">Figs. 2–3</figureCitation>
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||
). Tectum shortened, irregularly convex. Length of genital capsule from apex of parameres to border of basal piece:
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||
<quantity id="4CF99B0D5D45152B2E1FD1CAFDF2FA40" box="[544,620,1413,1432]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="2" pageNumber="13" unit="mm" value="5.1">5.1 mm</quantity>
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. Total body length: 23.0 mm. Humeral width:
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<quantity id="4CF99B0D5D45152B281AD1CAFBEEFA40" box="[1061,1136,1413,1432]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="2" pageNumber="13" unit="mm" value="9.3">9.3 mm</quantity>
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.
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</paragraph>
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<caption id="DF7E66605D45152B2D75D083FD48FAC8" ID-DOI="http://doi.org/10.5281/zenodo.4900313" ID-Zenodo-Dep="4900313" httpUri="https://zenodo.org/record/4900313/files/figure.png" pageId="2" pageNumber="13" startId="2.[330,373,1228,1247]" targetBox="[309,1134,273,1204]" targetPageId="2">
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<paragraph id="8BBE36E85D45152B2D75D083FD48FAC8" blockId="2.[307,1139,1228,1296]" pageId="2" pageNumber="13">
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||
<emphasis id="B975EAFA5D45152B2D75D083FE34FB07" bold="true" box="[330,426,1228,1247]" pageId="2" pageNumber="13">Figs. 1–5.</emphasis>
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<emphasis id="B975EAFA5D45152B2DFED082FD0CFB38" box="[449,658,1229,1248]" italics="true" pageId="2" pageNumber="13">
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<taxonomicName id="4C014D6B5D45152B2DFED082FD10FB38" box="[449,654,1229,1248]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="species" species="tilarana">Phyllophaga tilarana</taxonomicName>
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.
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</emphasis>
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<emphasis id="B975EAFA5D45152B2EA6D083FD3BFB07" bold="true" box="[665,677,1228,1247]" pageId="2" pageNumber="13">1</emphasis>
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) protarsal claw, male;
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<emphasis id="B975EAFA5D45152B2FBCD083FC11FB07" bold="true" box="[899,911,1228,1247]" pageId="2" pageNumber="13">2</emphasis>
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) genital capsule, lateral view;
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<emphasis id="B975EAFA5D45152B2D52D0ABFEE7FB2F" bold="true" box="[365,377,1252,1271]" pageId="2" pageNumber="13">3</emphasis>
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) paramera, distal view;
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<emphasis id="B975EAFA5D45152B2E61D0ABFDF4FB2F" bold="true" box="[606,618,1252,1271]" pageId="2" pageNumber="13">4</emphasis>
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) protarsal claw, female;
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<emphasis id="B975EAFA5D45152B2F6BD0ABFCFEFB2F" bold="true" box="[852,864,1252,1271]" pageId="2" pageNumber="13">5</emphasis>
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) genital plates, female. Scale lines 5 1 mm, except Figs. 1, 4 5 0.5 mm.
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</paragraph>
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</caption>
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<paragraph id="8BBE36E85D45152A2D72D1E5FC11FE12" blockId="2.[307,1139,1450,1658]" lastBlockId="3.[279,1112,277,1392]" lastPageId="3" lastPageNumber="14" pageId="2" pageNumber="13">
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||
<emphasis id="B975EAFA5D45152B2D72D1E5FE2CFA67" bold="true" box="[333,434,1450,1471]" pageId="2" pageNumber="13">
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<typeStatus id="54BA884A5D45152B2D72D1E5FE33FA67" box="[333,429,1450,1471]" pageId="2" pageNumber="13" type="allotype">Allotype</typeStatus>
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.
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</emphasis>
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Female. Similar to the male except as follows: antennae with lamellae of 8th to 10th segments slightly longer than the length of six preceeding segments combined (1:0.9). Visible abdominal sternites II to V convex, with scattered setiferous punctures; anal plate 1.2× longer than male anal plate, convex, with scattered setiferous punctures, with 22 slender setae at the posterior border. Pygidium slightly convex, nearly flattened toward the apex, apical border slightly sinuated. Protibiae with the teeth of external border wider and longer than in the male. Both apical spurs of metatibia articulated, wide, lanceolate and curved. Tarsomeres only with two single lines of long, slender setae on ventral side. Tarsal claws each with a large tooth at the middle of ventral border (
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<figureCitation id="133A2A6D5D44152A2DFCD504FD97FEB8" box="[451,521,331,352]" captionStart="Figs" captionStartId="2.[330,373,1228,1247]" captionTargetBox="[309,1134,273,1204]" captionTargetId="figure-12@2.[308,1137,272,1208]" captionTargetPageId="2" captionText="Figs. 1–5. Phyllophaga tilarana. 1) protarsal claw, male; 2) genital capsule, lateral view; 3) paramera, distal view; 4) protarsal claw, female; 5) genital plates, female. Scale lines 5 1 mm, except Figs. 1, 4 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900313" httpUri="https://zenodo.org/record/4900313/files/figure.png" pageId="3" pageNumber="14">Fig. 4</figureCitation>
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). Ventral genital plates moderately sclerotized, nearly symmetrical, elongate ovate shaped with shallow depression near the apex; dorsal plates wider than ventral plates with scattered setae on the distal border, each plate partially fused with ventral plate along the laterobasal border (
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<figureCitation id="133A2A6D5D44152A2812D5D4FEB8FE12" captionStart="Figs" captionStartId="2.[330,373,1228,1247]" captionTargetBox="[309,1134,273,1204]" captionTargetId="figure-12@2.[308,1137,272,1208]" captionTargetPageId="2" captionText="Figs. 1–5. Phyllophaga tilarana. 1) protarsal claw, male; 2) genital capsule, lateral view; 3) paramera, distal view; 4) protarsal claw, female; 5) genital plates, female. Scale lines 5 1 mm, except Figs. 1, 4 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900313" httpUri="https://zenodo.org/record/4900313/files/figure.png" pageId="3" pageNumber="14">Fig. 5</figureCitation>
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). Total body length:
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<quantity id="4CF99B0D5D44152A2E35D5FAFDF2FE12" box="[522,620,437,458]" metricMagnitude="-2" metricUnit="m" metricValue="2.46" pageId="3" pageNumber="14" unit="mm" value="24.6">24.6 mm</quantity>
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. Humeral width: 10.0 mm.
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</paragraph>
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<paragraph id="8BBE36E85D44152A2D0DD59FFDFFFDB2" blockId="3.[279,1112,277,1392]" pageId="3" pageNumber="14">
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<emphasis id="B975EAFA5D44152A2D0DD59FFD8FFE3D" bold="true" box="[306,529,464,485]" pageId="3" pageNumber="14">
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<typeStatus id="54BA884A5D44152A2D0DD59FFE06FE3D" box="[306,408,464,485]" pageId="3" pageNumber="14" type="paratype">Paratype</typeStatus>
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Variation.
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</emphasis>
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Males. Similar to
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<typeStatus id="54BA884A5D44152A2EEAD59FFCAEFE3D" box="[725,816,464,485]" pageId="3" pageNumber="14" type="holotype">holotype</typeStatus>
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except in total body length: 20.0–24.0 mm, humeral width:
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<quantity id="4CF99B0D5D44152A2E43D5A4FC9CFDD8" box="[636,770,491,512]" metricMagnitude="-3" metricUnit="m" metricValue="8.649999999999999" metricValueMax="9.1" metricValueMin="8.2" pageId="3" pageNumber="14" unit="mm" value="8.649999999999999" valueMax="9.1" valueMin="8.2">8.2–9.1 mm</quantity>
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, pronotum of some specimens darker than
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<typeStatus id="54BA884A5D44152A2DA1D64AFE62FDC2" box="[414,508,517,538]" pageId="3" pageNumber="14" type="holotype">holotype</typeStatus>
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, other specimens with antennal club 2.2× longer than length of preceeding six segments combined. Female similar to
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<typeStatus id="54BA884A5D44152A2F86D66FFB93FDED" box="[953,1037,544,565]" pageId="3" pageNumber="14" type="allotype">allotype</typeStatus>
|
||
except as follows: elytra and pygidium with more punctures; total body length:
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<quantity id="4CF99B0D5D44152A2815D674FEDDFDB2" metricMagnitude="-2" metricUnit="m" metricValue="2.43" pageId="3" pageNumber="14" unit="mm" value="24.3">24.3 mm</quantity>
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; humeral width: 10.0 mm.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31B65635D44152A2D0DD63FFE38FC72" pageId="3" pageNumber="14" type="materials_examined">
|
||
<paragraph id="8BBE36E85D44152A2D0DD63FFDD1FC82" blockId="3.[279,1112,277,1392]" pageId="3" pageNumber="14">
|
||
<emphasis id="B975EAFA5D44152A2D0DD63FFE20FD5D" bold="true" box="[306,446,624,645]" pageId="3" pageNumber="14">Type Series.</emphasis>
|
||
<collectingCounty id="62DF4E645D44152A2DF7D63FFDACFD5D" box="[456,562,624,645]" pageId="3" pageNumber="14">Described</collectingCounty>
|
||
from
|
||
<specimenCount id="9D07FD615D44152A2E45D63FFD50FD5D" box="[634,718,624,645]" pageId="3" pageNumber="14" type="male">9 males</specimenCount>
|
||
and
|
||
<specimenCount id="9D07FD615D44152A2F36D63FFCEBFD5D" box="[777,885,624,645]" pageId="3" pageNumber="14" type="female">2 females</specimenCount>
|
||
.
|
||
<materialsCitation id="3B693CB55D44152A2FBDD63FFD48FD62" collectingDate="1990-11" collectionCode="IN- BIO" collectorName="M. Ramirez." country="Costa Rica" elevation="1520" latitude="10.18" location="Reserva Biologica Monteverde" longitude="-84.47" municipality="Estacion La Casona" pageId="3" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="holotype">
|
||
<typeStatus id="54BA884A5D44152A2FBDD63FFC7DFD5D" box="[898,995,624,645]" pageId="3" pageNumber="14" type="holotype">Holotype</typeStatus>
|
||
<specimenCount id="9D07FD615D44152A2FD1D63FFBBFFD5D" box="[1006,1057,624,645]" pageId="3" pageNumber="14" type="male">male</specimenCount>
|
||
(
|
||
<collectionCode id="ED10AE2D5D44152A280DD63FFED7FD78" pageId="3" pageNumber="14">INBIO</collectionCode>
|
||
): ‘‘
|
||
<collectingCountry id="F31676785D44152A2D57D6C4FD8EFD78" box="[360,528,651,672]" name="Costa Rica" pageId="3" pageNumber="14">COSTA RICA</collectingCountry>
|
||
:
|
||
<collectingRegion id="49C5F80A5D44152A2E22D6C4FD0CFD78" box="[541,658,651,672]" country="Costa Rica" name="Puntarenas" pageId="3" pageNumber="14">Puntarenas</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BDAAC925D44152A2E9FD6C4FC6EFD78" box="[672,1008,651,672]" pageId="3" pageNumber="14">Reserva Biológica Monteverde</collectingMunicipality>
|
||
,
|
||
<location id="8EDE60335D44152A2FC1D6C4FE14FD62" LSID="urn:lsid:plazi:treatment:03A887FE5D46152A2D70D613FCBCFAA8:8EDE60335D44152A2FC1D6C4FE14FD62" country="Costa Rica" county="Described" municipality="Reserva Biologica Monteverde" name="Estacion La Casona" pageId="3" pageNumber="14" stateProvince="Puntarenas">Estación La Casona</location>
|
||
,
|
||
<quantity id="4CF99B0D5D44152A2DABD6EAFE73FD62" box="[404,493,677,698]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="3" pageNumber="14" unit="m" value="1520.0">
|
||
<elevation id="002CD1DB5D44152A2DABD6EAFE73FD62" box="[404,493,677,698]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="3" pageNumber="14" unit="m" value="1520.0">1,520 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFBF10285D44152A2DC8D6EAFDA6FD62" box="[503,568,677,698]" pageId="3" pageNumber="14" value="1990-11">
|
||
<collectingDate id="EFFBE9C05D44152A2DC8D6EAFDA6FD62" box="[503,568,677,698]" pageId="3" pageNumber="14" value="1990-11">XI90</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26F4533E5D44152A2E7CD6EAFD53FD62" box="[579,717,677,698]" pageId="3" pageNumber="14">M. Ramírez.</collectorName>
|
||
’’
|
||
</materialsCitation>
|
||
<materialsCitation id="3B693CB55D44152A2EE2D6EAFDC5FD28" collectingDate="1995-04-24" collectionCode="INBIO" collectorName="A. Azofeifa." country="Costa Rica" elevation="1520" latitude="10.18" location="Reserva Biologica Monteverde" longitude="-84.47" municipality="estacion La Casona" pageId="3" pageNumber="14" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="allotype">
|
||
<typeStatus id="54BA884A5D44152A2EE2D6EAFCA6FD62" box="[733,824,677,698]" pageId="3" pageNumber="14" type="allotype">Allotype</typeStatus>
|
||
<specimenCount id="9D07FD615D44152A2F00D6EAFC18FD62" box="[831,902,677,698]" pageId="3" pageNumber="14" type="female">female</specimenCount>
|
||
(
|
||
<collectionCode id="ED10AE2D5D44152A2FABD6EAFC7DFD62" box="[916,995,677,698]" pageId="3" pageNumber="14">INBIO</collectionCode>
|
||
): ‘‘
|
||
<collectingCountry id="F31676785D44152A2FC2D6EAFEC4FD0D" name="Costa Rica" pageId="3" pageNumber="14">COSTA RICA</collectingCountry>
|
||
:
|
||
<collectingRegion id="49C5F80A5D44152A2D5AD68FFE44FD0D" box="[357,474,704,725]" country="Costa Rica" name="Puntarenas" pageId="3" pageNumber="14">Puntarenas</collectingRegion>
|
||
,
|
||
<location id="8EDE60335D44152A2DD9D68FFCACFD0D" LSID="urn:lsid:plazi:treatment:03A887FE5D46152A2D70D613FCBCFAA8:8EDE60335D44152A2DD9D68FFCACFD0D" box="[486,818,704,725]" country="Costa Rica" county="Described" municipality="Reserva Biologica Monteverde" name="Reserva Biologica Monteverde" pageId="3" pageNumber="14" stateProvince="Puntarenas">Reserva Biológica Monteverde</location>
|
||
,
|
||
<location id="8EDE60335D44152A2F02D68FFB8CFD0D" LSID="urn:lsid:plazi:treatment:03A887FE5D46152A2D70D613FCBCFAA8:8EDE60335D44152A2F02D68FFB8CFD0D" box="[829,1042,704,725]" country="Costa Rica" county="Described" municipality="Reserva Biologica Monteverde" name="Estacion La Casona" pageId="3" pageNumber="14" stateProvince="Puntarenas">Estación La Casona</location>
|
||
,
|
||
<quantity id="4CF99B0D5D44152A2822D68FFEB0FD28" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="3" pageNumber="14" unit="m" value="1520.0">
|
||
<elevation id="002CD1DB5D44152A2822D68FFEB0FD28" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="3" pageNumber="14" unit="m" value="1520.0">1,520 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFBF10285D44152A2D01D694FE2AFD28" box="[318,436,731,752]" pageId="3" pageNumber="14" value="1995-04-24">
|
||
<collectingDate id="EFFBE9C05D44152A2D01D694FE2AFD28" box="[318,436,731,752]" pageId="3" pageNumber="14">3/24IV95</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26F4533E5D44152A2DFBD694FDCCFD28" box="[452,594,731,752]" pageId="3" pageNumber="14">A. Azofeifa.</collectorName>
|
||
’’
|
||
</materialsCitation>
|
||
<materialsCitation id="3B693CB55D44152A2E58D694FE51FCD2" collectionCode="CNC, INBIO, MXAL" collectorName="E. Bello" country="Costa Rica" elevation="1520" latitude="10.18" location="Reserva Biologica Monteverde" longitude="-84.47" municipality="Estacion La Casona" pageId="3" pageNumber="14" specimenCount="2" specimenCount-male="2" stateProvince="Puntarenas" typeStatus="paratype">
|
||
<typeStatus id="54BA884A5D44152A2E58D694FD4FFD28" box="[615,721,731,752]" pageId="3" pageNumber="14" type="paratype">Paratypes</typeStatus>
|
||
: same data as holotype except ‘‘
|
||
<collectorName id="26F4533E5D44152A2801D694FEC5FCD2" pageId="3" pageNumber="14">E. Bello’</collectorName>
|
||
’ (
|
||
<specimenCount id="9D07FD615D44152A2D4FD6BAFE56FCD2" box="[368,456,757,778]" count="2" pageId="3" pageNumber="14" type="male">2 males</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B693CB55D44152A2DE2D6BAFD33FCD2" box="[477,685,757,778]" collectingDate="1990-12" collectorName="de Tilaran" country="Costa Rica" latitude="10.18" location="Reserva Biologica Monteverde" longitude="-84.47" municipality="Estacion La Casona" pageId="3" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
|
||
‘‘
|
||
<date id="FFBF10285D44152A2DD8D6BAFDA7FCD2" box="[487,569,757,778]" pageId="3" pageNumber="14" value="1990-12">
|
||
<collectingDate id="EFFBE9C05D44152A2DD8D6BAFDA7FCD2" box="[487,569,757,778]" pageId="3" pageNumber="14" value="1990-12">XII90</collectingDate>
|
||
</date>
|
||
’’ (
|
||
<specimenCount id="9D07FD615D44152A2E69D6BAFD3FFCD2" box="[598,673,757,778]" count="1" pageId="3" pageNumber="14" type="male">1 male</specimenCount>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B693CB55D44152A2E8AD6BAFBC6FCD2" box="[693,1112,757,778]" collectingDate="1992-03" collectionCode="CNC, INBIO, MXAL" collectorName="K. Flores'" country="Costa Rica" elevation="1520" latitude="10.18" location="Reserva Biologica Monteverde" longitude="-84.47" municipality="Estacion La Casona" pageId="3" pageNumber="14" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
|
||
‘‘
|
||
<date id="FFBF10285D44152A2EFFD6BAFC9AFCD2" box="[704,772,757,778]" pageId="3" pageNumber="14" value="1992-03">
|
||
<collectingDate id="EFFBE9C05D44152A2EFFD6BAFC9AFCD2" box="[704,772,757,778]" pageId="3" pageNumber="14" value="1992-03">III92</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26F4533E5D44152A2F2ED6BAFC1EFCD2" box="[785,896,757,778]" pageId="3" pageNumber="14">K. Flores’</collectorName>
|
||
’ (
|
||
<specimenCount id="9D07FD615D44152A2FA9D6BAFC41FCD2" box="[918,991,757,778]" count="1" pageId="3" pageNumber="14" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="9D07FD615D44152A2FD3D6BAFBD3FCD2" box="[1004,1101,757,778]" count="1" pageId="3" pageNumber="14" type="female">1 female</specimenCount>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B693CB55D44152A2D28D75FFDC7FCFD" box="[279,601,784,805]" collectingDate="1993-01" collectionCode="CNC, INBIO, MXAL" collectorName="N. Obando'" country="Costa Rica" elevation="1520" latitude="10.18" location="Reserva Biologica Monteverde" longitude="-84.47" municipality="Estacion La Casona" pageId="3" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
|
||
‘‘
|
||
<date id="FFBF10285D44152A2D1CD75FFECBFCFD" box="[291,341,784,805]" pageId="3" pageNumber="14" value="1993-01">
|
||
<collectingDate id="EFFBE9C05D44152A2D1CD75FFECBFCFD" box="[291,341,784,805]" pageId="3" pageNumber="14" value="1993-01">I93</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26F4533E5D44152A2D5BD75FFE78FCFD" box="[356,486,784,805]" pageId="3" pageNumber="14">N. Obando’</collectorName>
|
||
’ (
|
||
<specimenCount id="9D07FD615D44152A2E3FD75FFDD0FCFD" box="[512,590,784,805]" count="1" pageId="3" pageNumber="14" type="male">1 male</specimenCount>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B693CB55D44152A2E5BD75FFC2EFCFD" box="[612,944,784,805]" collectingDate="1993-10" collectionCode="CNC, INBIO, MXAL" collectorName="N. Obando'" country="Costa Rica" elevation="1520" latitude="10.18" location="Reserva Biologica Monteverde" longitude="-84.47" municipality="Estacion La Casona" pageId="3" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
|
||
‘‘
|
||
<date id="FFBF10285D44152A2E50D75FFD32FCFD" box="[623,684,784,805]" pageId="3" pageNumber="14" value="1993-10">
|
||
<collectingDate id="EFFBE9C05D44152A2E50D75FFD32FCFD" box="[623,684,784,805]" pageId="3" pageNumber="14" value="1993-10">X93</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26F4533E5D44152A2E84D75FFCA3FCFD" box="[699,829,784,805]" pageId="3" pageNumber="14">N. Obando’</collectorName>
|
||
’ (
|
||
<specimenCount id="9D07FD615D44152A2F68D75FFC3AFCFD" box="[855,932,784,805]" count="1" pageId="3" pageNumber="14" type="male">1 male</specimenCount>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B693CB55D44152A2F84D75FFE6FFC98" collectingDate="1995-01-22" collectionCode="CNC, INBIO, MXAL" collectorName="K. Martinez'" country="Costa Rica" elevation="1520" latitude="10.18" location="Reserva Biologica Monteverde" longitude="-84.47" municipality="Estacion La Casona" pageId="3" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
|
||
‘‘
|
||
<date id="FFBF10285D44152A2FFAD75FFBB1FCFD" box="[965,1071,784,805]" pageId="3" pageNumber="14" value="1995-01-22">
|
||
<collectingDate id="EFFBE9C05D44152A2FFAD75FFBB1FCFD" box="[965,1071,784,805]" pageId="3" pageNumber="14">3/22I95</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26F4533E5D44152A2801D75FFE1FFC98" pageId="3" pageNumber="14">K. Martínez’</collectorName>
|
||
’ (
|
||
<specimenCount id="9D07FD615D44152A2DA6D764FE7BFC98" box="[409,485,811,832]" count="1" pageId="3" pageNumber="14" type="male">1 male</specimenCount>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B693CB55D44152A2DC4D764FCE2FC98" box="[507,892,811,832]" collectingDate="1995-04-17" collectionCode="CNC, INBIO, MXAL" collectorName="M. Madrigal'" country="Costa Rica" elevation="1520" latitude="10.18" location="Reserva Biologica Monteverde" longitude="-84.47" municipality="Estacion La Casona" pageId="3" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
|
||
‘‘
|
||
<date id="FFBF10285D44152A2E3AD764FDF5FC98" box="[517,619,811,832]" pageId="3" pageNumber="14" value="1995-04-17">
|
||
<collectingDate id="EFFBE9C05D44152A2E3AD764FDF5FC98" box="[517,619,811,832]" pageId="3" pageNumber="14" value="1995-04-17">17IV95</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26F4533E5D44152A2E46D764FC92FC98" box="[633,780,811,832]" pageId="3" pageNumber="14">M. Madrigal’</collectorName>
|
||
’ (
|
||
<specimenCount id="9D07FD615D44152A2F1BD764FCEEFC98" box="[804,880,811,832]" count="1" pageId="3" pageNumber="14" type="male">1 male</specimenCount>
|
||
).
|
||
</materialsCitation>
|
||
Paratypes deposited in
|
||
<collectionCode id="ED10AE2D5D44152A2D0AD70AFEF3FC82" box="[309,365,837,858]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" pageId="3" pageNumber="14">CNC</collectionCode>
|
||
,
|
||
<collectionCode id="ED10AE2D5D44152A2D45D70AFE5DFC82" box="[378,451,837,858]" pageId="3" pageNumber="14">INBIO</collectionCode>
|
||
and
|
||
<collectionCode id="ED10AE2D5D44152A2DC5D70AFDD4FC82" box="[506,586,837,858]" pageId="3" pageNumber="14">MXAL</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBE36E85D44152A2D0DD72FFE38FC72" blockId="3.[279,1112,277,1392]" pageId="3" pageNumber="14">
|
||
<emphasis id="B975EAFA5D44152A2D0DD72FFE48FCAD" bold="true" box="[306,470,864,885]" pageId="3" pageNumber="14">Type Locality.</emphasis>
|
||
<collectingMunicipality id="6BDAAC925D44152A2DE0D72FFD29FCAD" box="[479,695,864,885]" pageId="3" pageNumber="14">Estación La Casona</collectingMunicipality>
|
||
,
|
||
<location id="8EDE60335D44152A2EFBD72FFB8CFCAD" LSID="urn:lsid:plazi:treatment:03A887FE5D46152A2D70D613FCBCFAA8:8EDE60335D44152A2EFBD72FFB8CFCAD" box="[708,1042,864,885]" country="Costa Rica" municipality="Estacion La Casona" name="Reserva Biologica Monteverde" pageId="3" pageNumber="14" stateProvince="Puntarenas">Reserva Biológica Monteverde</location>
|
||
, province of
|
||
<collectingRegion id="49C5F80A5D44152A2DBBD734FE60FC48" box="[388,510,891,912]" country="Costa Rica" name="Puntarenas" pageId="3" pageNumber="14">Puntarenas</collectingRegion>
|
||
, cordillera de
|
||
<location id="8EDE60335D44152A2EFED734FC8CFC48" LSID="urn:lsid:plazi:treatment:03A887FE5D46152A2D70D613FCBCFAA8:8EDE60335D44152A2EFED734FC8CFC48" box="[705,786,891,912]" country="Costa Rica" municipality="Estacion La Casona" name="Tilaran" pageId="3" pageNumber="14" stateProvince="Puntarenas">Tilarán</location>
|
||
,
|
||
<collectingCountry id="F31676785D44152A2F14D734FC2FFC48" box="[811,945,891,912]" name="Costa Rica" pageId="3" pageNumber="14">Costa Rica</collectingCountry>
|
||
. (
|
||
<geoCoordinate id="EE35502F5D44152A2FEED734FBCAFC48" box="[977,1108,891,912]" direction="north" orientation="latitude" pageId="3" pageNumber="14" value="10.18">10°18'150N</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE35502F5D44152A2D28D7DAFE05FC72" box="[279,411,917,938]" direction="west" orientation="longitude" pageId="3" pageNumber="14" value="-84.47">84°47'460W</geoCoordinate>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B65635D44152A2D0DD7FFFDB2FB92" pageId="3" pageNumber="14" type="biology_ecology">
|
||
<paragraph id="8BBE36E85D44152A2D0DD7FFFDB2FB92" blockId="3.[279,1112,277,1392]" pageId="3" pageNumber="14">
|
||
<emphasis id="B975EAFA5D44152A2D0DD7FFFE79FC1D" bold="true" box="[306,487,944,965]" pageId="3" pageNumber="14">Biological Data.</emphasis>
|
||
Males and females of
|
||
<taxonomicName id="4C014D6B5D44152A2EDAD7FFFCC4FC1D" box="[741,858,944,965]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="14" phylum="Arthropoda" rank="species" species="tilarana" status="sp. nov.">
|
||
<emphasis id="B975EAFA5D44152A2EDAD7FFFCC4FC1D" box="[741,858,944,965]" italics="true" pageId="3" pageNumber="14">P. tilarana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24657815D44152A2F5BD7FFFC08FC1D" box="[868,918,944,965]" pageId="3" pageNumber="14" rank="species">n.sp.</taxonomicNameLabel>
|
||
were collected at lights in cloud forest located near
|
||
<quantity id="4CF99B0D5D44152A2EA2D784FD69FC38" box="[669,759,971,992]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="3" pageNumber="14" unit="m" value="1520.0">1,520 m</quantity>
|
||
of altitude,
|
||
<quantity id="4CF99B0D5D44152A2FB8D784FC5FFC38" box="[903,961,971,992]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="14" unit="km" value="1.0">1 km</quantity>
|
||
southwest of Atlantic/Pacific montane rainfall line. Phenology: January (2), March (2), April (2), October (1), November (3), December (1). Other species of
|
||
<taxonomicName id="4C014D6B5D44152A2FEDD04FFBC6FBCD" box="[978,1112,1024,1045]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B975EAFA5D44152A2FEDD04FFBC6FBCD" box="[978,1112,1024,1045]" italics="true" pageId="3" pageNumber="14">Phyllophaga</emphasis>
|
||
</taxonomicName>
|
||
flying at the same time and place were
|
||
<taxonomicName id="4C014D6B5D44152A2EF9D054FCD4FBE8" box="[710,842,1051,1072]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="14" phylum="Arthropoda" rank="species" species="tapantina" status="sp. nov.">
|
||
<emphasis id="B975EAFA5D44152A2EF9D054FCD4FBE8" box="[710,842,1051,1072]" italics="true" pageId="3" pageNumber="14">P. tapantina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24657815D44152A2F6CD054FC1BFBE8" box="[851,901,1051,1072]" pageId="3" pageNumber="14" rank="species">n.sp.</taxonomicNameLabel>
|
||
and two other new species under description.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B65635D44152A2D0DD01FFBCCFAE2" pageId="3" pageNumber="14" type="discussion">
|
||
<paragraph id="8BBE36E85D44152A2D0DD01FFBCCFAE2" blockId="3.[279,1112,277,1392]" pageId="3" pageNumber="14">
|
||
<emphasis id="B975EAFA5D44152A2D0DD01FFE00FBBD" bold="true" box="[306,414,1104,1125]" pageId="3" pageNumber="14">Remarks.</emphasis>
|
||
<taxonomicName id="4C014D6B5D44152A2D99D01FFCB6FBBD" box="[422,808,1104,1125]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="14" phylum="Arthropoda" rank="species" species="tilarana" subGenus="Phyllophaga">
|
||
<emphasis id="B975EAFA5D44152A2D99D01FFDB2FBBD" box="[422,556,1104,1125]" italics="true" pageId="3" pageNumber="14">Phyllophaga</emphasis>
|
||
(
|
||
<emphasis id="B975EAFA5D44152A2E02D01FFD5CFBBD" box="[573,706,1104,1125]" italics="true" pageId="3" pageNumber="14">Phyllophaga</emphasis>
|
||
)
|
||
<emphasis id="B975EAFA5D44152A2EEBD01FFCB6FBBD" box="[724,808,1104,1125]" italics="true" pageId="3" pageNumber="14">tilarana</emphasis>
|
||
</taxonomicName>
|
||
does not belong to any species group described by
|
||
<bibRefCitation id="EF904B195D44152A2E1BD024FD20FB58" author="Moron" box="[548,702,1131,1152]" firstAuthor="Moron" pageId="3" pageNumber="14" refId="ref11224" refString="Moron, M. A. 1986. El genero Phyllophaga en Mexico. Morfologia, distribucion y sistematica supraespecifica (Insecta: Coleoptera). Publ. 20, Instituto de Ecologia, Mexico, D. F. 341 pp." type="book" year="1986">Morón (1986)</bibRefCitation>
|
||
. Male tarsal claw structure resembles ‘‘
|
||
<emphasis id="B975EAFA5D44152A2D18D0C9FE09FB43" box="[295,407,1158,1179]" italics="true" pageId="3" pageNumber="14">rugipennis</emphasis>
|
||
’’ group, but female tarsal claws have typical dentate structure, found in many other groups.
|
||
<taxonomicName id="4C014D6B5D44152A2E71D0EFFCB1FB6D" box="[590,815,1184,1205]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="14" phylum="Arthropoda" rank="species" species="tilarana">
|
||
<emphasis id="B975EAFA5D44152A2E71D0EFFCB1FB6D" box="[590,815,1184,1205]" italics="true" pageId="3" pageNumber="14">Phyllophaga tilarana</emphasis>
|
||
</taxonomicName>
|
||
may be related with
|
||
<emphasis id="B975EAFA5D44152A282DD0EFFBB7FB6D" box="[1042,1065,1184,1205]" italics="true" pageId="3" pageNumber="14">P.</emphasis>
|
||
(
|
||
<emphasis id="B975EAFA5D44152A2807D0EFFBD1FB6D" box="[1080,1103,1184,1205]" italics="true" pageId="3" pageNumber="14">P.</emphasis>
|
||
)
|
||
<emphasis id="B975EAFA5D44152A2D28D0F4FE04FB08" box="[279,410,1211,1232]" italics="true" pageId="3" pageNumber="14">nevermanni,</emphasis>
|
||
described by
|
||
<bibRefCitation id="EF904B195D44152A2E09D0F4FD67FB08" author="Saylor, L. W." box="[566,761,1211,1232]" journalOrPublisher="Revista de Entomologia (Rio de Janeiro)" pageId="3" pageNumber="14" pagination="496 - 501" part="5" refId="ref11267" refString="Saylor, L. W. 1935. New neotropical Scarabaeidae of the genus Phyllophaga (Col.). Revista de Entomologia (Rio de Janeiro) 5: 496 - 501." title="New neotropical Scarabaeidae of the genus Phyllophaga (Col.)" type="journal article" year="1935">Saylor (1935:501)</bibRefCitation>
|
||
from Guápiles,
|
||
<collectingCountry id="F31676785D44152A2F92D0F4FBBCFB08" box="[941,1058,1211,1232]" name="Costa Rica" pageId="3" pageNumber="14">Costa Rica</collectingCountry>
|
||
(
|
||
<typeStatus id="54BA884A5D44152A280CD0F4FEC6FB32" pageId="3" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
USNM 54567) by the body shape, color, dorsal vestiture, antennal structure and the shape of tarsal claws; but the shape and details of the male genital capsule, the accessorial structures of male sternites V and VI, vestiture and shape of the pygidium in males and females aid to separate the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B65635D44152A2D0DD10FFCBCFAA8" pageId="3" pageNumber="14" type="etymology">
|
||
<paragraph id="8BBE36E85D44152A2D0DD10FFCBCFAA8" blockId="3.[279,1112,277,1392]" pageId="3" pageNumber="14">
|
||
<emphasis id="B975EAFA5D44152A2D0DD10FFE2FFA8D" bold="true" box="[306,433,1344,1365]" pageId="3" pageNumber="14">Etymology.</emphasis>
|
||
Derived from Tilarán Mountain System of northwestern
|
||
<collectingCountry id="F31676785D44152A2823D10FFED9FAA8" name="Costa Rica" pageId="3" pageNumber="14">Costa Rica</collectingCountry>
|
||
where this species was originally collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |