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<document id="8E84DA38C280657A17BA4054A156803C" ID-DOI="10.1649/0010-065x(2001)055[0011:snsops]2.0.co;2" ID-ISSN="1938-4394" ID-Zenodo-Dep="4900311" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1622827801739" checkinUser="carolina" docAuthor="Morón, Miguel Angel & Solís, Angel" docDate="2001" docId="03A887FE5D4115212D5AD18CFE1BFB08" docLanguage="en" docName="TheColeopBull.55.1.11-29.pdf.imf" docOrigin="The Coleopterists Bulletin 55 (1)" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Phyllophaga (Phyllophaga) lorencita Moron and Solis, new species" docType="treatment" docVersion="1" lastPageNumber="19" masterDocId="FF91FF865D4715292C3FD44FFF9EFFD8" masterDocTitle="Seven New Species Of Phyllophaga (S. Str.) Harris From Costa Rica (Coleoptera: Melolonthidae: Melolonthinae)" masterLastPageNumber="29" masterPageNumber="11" pageNumber="17" updateTime="1730139982923" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:mods id="0A88CB565590A489634EBE27ECEBA458" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="90476B19F9B074935AD2CE7AA69E5163">
|
||||
<mods:title id="76B3913B7D2CB7078D56AF7DBFC90EAD">Seven New Species Of Phyllophaga (S. Str.) Harris From Costa Rica (Coleoptera: Melolonthidae: Melolonthinae)</mods:title>
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||||
</mods:titleInfo>
|
||||
<mods:name id="EAC7566204CB2A3DD53B05849A4E9B08" type="personal">
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<mods:role id="F1BF0A4D834E508907167E132BF540F0">
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<mods:roleTerm id="0A3B714C50019515F18C68B0573F214D">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="13BCB34F92F614564974FC09109431A5">Morón, Miguel Angel</mods:namePart>
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||||
<mods:affiliation id="0B28448CFBB0A4DDF3AECD8BF85D45B1">Departamento de Entomología Instituto de Ecología, A. C. (sector SEPCONACYT) Apartado Postal 63 Xalapa, Veracruz 91000, MÉXICO</mods:affiliation>
|
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</mods:name>
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<mods:name id="7A784DF93870C5982156E3C5E7AFC885" type="personal">
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<mods:roleTerm id="7792992FBC6C1BC504EF5FC1C8097F65">Author</mods:roleTerm>
|
||||
</mods:role>
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||||
<mods:namePart id="401404D22739A1C4064841C8457D5189">Solís, Angel</mods:namePart>
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||||
<mods:affiliation id="B549CBE0B6ECD45B96FB5B39C80850A4">Departamento de Entomología Instituto Nacional de Biodiversidad (INBio) Apartado Postal 22 3100 Santo Domingo de Heredia, COSTA RICA</mods:affiliation>
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||||
</mods:name>
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||||
<mods:typeOfResource id="B35F22929D9113751F369D4453D73455">text</mods:typeOfResource>
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<mods:relatedItem id="E6FEC3887703DAD4BCFD9CC12EB55C1E" type="host">
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<mods:titleInfo id="3CF98365A823C971C48C1809184B6013">
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||||
<mods:title id="DA6517591C6D9FE5ED5FEBB51FCAEBA2">The Coleopterists Bulletin</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part id="CC6CAD1AAB544D1CC5D73AD75BF8DD05">
|
||||
<mods:date id="7E6D81007FF2F8FC9A99C19EAFDCBFE8">2001</mods:date>
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<mods:detail id="8C23B556D6ED9CBCBA25838613821E57" type="pubDate">
|
||||
<mods:number id="B4E24C1747A92108E03848688D92CF12">2001-03-01</mods:number>
|
||||
</mods:detail>
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||||
<mods:detail id="8C454AA897222A449FF4A61C27B25CBF" type="volume">
|
||||
<mods:number id="03B6BD35C407B37ED6EDA689ACF2DDAD">55</mods:number>
|
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</mods:detail>
|
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<mods:detail id="0FBF85D794ED58DD8E8F7440EEB2181C" type="issue">
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<mods:number id="7B0BA1A472C1BAA9347EDA0BA1C79B34">1</mods:number>
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</mods:detail>
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<mods:extent id="7080D445C2BBB233BDE1CC12A2D68925" unit="page">
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<mods:start id="64E678BE44882A371CBBA37096142B1F">11</mods:start>
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<mods:end id="CFEE8C7D326E4C8769C02F6863FC0C10">29</mods:end>
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</mods:relatedItem>
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<mods:classification id="93A2E7E602945F1048584187099C4369">journal article</mods:classification>
|
||||
<mods:identifier id="0889FDE1B1B91A8B55A4E4719082CD42" type="DOI">10.1649/0010-065x(2001)055[0011:snsops]2.0.co;2</mods:identifier>
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<mods:identifier id="64096A2F71077D6FE6C8FF1EC78349A9" type="ISSN">1938-4394</mods:identifier>
|
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<mods:identifier id="95C90C8169C61DF6F5D6C0BB82CB4102" type="Zenodo-Dep">4900311</mods:identifier>
|
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</mods:mods>
|
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<treatment id="03A887FE5D4115212D5AD18CFE1BFB08" LSID="urn:lsid:plazi:treatment:03A887FE5D4115212D5AD18CFE1BFB08" httpUri="http://treatment.plazi.org/id/03A887FE5D4115212D5AD18CFE1BFB08" lastPageId="8" lastPageNumber="19" pageId="6" pageNumber="17">
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<subSubSection id="C31B65635D41152F2D5AD18CFBDEFA0F" box="[357,1088,1474,1496]" pageId="6" pageNumber="17" type="nomenclature">
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<paragraph id="8BBE36E85D41152F2D5AD18CFBDEFA0F" blockId="6.[357,1088,1474,1522]" box="[357,1088,1474,1496]" pageId="6" pageNumber="17">
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||||
<taxonomicName id="4C014D6B5D41152F2D5AD18CFC2FFA00" authority="Moron and Solis" authorityName="Moron and Solis" box="[357,945,1475,1496]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="lorencita" status="sp. nov." subGenus="Phyllophaga">
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||||
<emphasis id="B975EAFA5D41152F2D5AD18CFE75FA00" box="[357,491,1475,1496]" italics="true" pageId="6" pageNumber="17">Phyllophaga</emphasis>
|
||||
(
|
||||
<emphasis id="B975EAFA5D41152F2DC2D18CFD1CFA00" box="[509,642,1475,1496]" italics="true" pageId="6" pageNumber="17">Phyllophaga</emphasis>
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||||
)
|
||||
<emphasis id="B975EAFA5D41152F2EABD18CFD6CFA00" box="[660,754,1475,1496]" italics="true" pageId="6" pageNumber="17">lorencita</emphasis>
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||||
Morón and Solís
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</taxonomicName>
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||||
,
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||||
<emphasis id="B975EAFA5D41152F2F82D18DFBDEFA0F" bold="true" box="[957,1088,1474,1495]" pageId="6" pageNumber="17">
|
||||
<taxonomicNameLabel id="A24657815D41152F2F82D18DFBDEFA0F" box="[957,1088,1474,1495]" pageId="6" pageNumber="17" rank="species">new species</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D4115212EACD192FE12FE68" lastPageId="8" lastPageNumber="19" pageId="6" pageNumber="17" type="description">
|
||||
<paragraph id="8BBE36E85D41152F2EACD192FC8CFA2A" blockId="6.[357,1088,1474,1522]" box="[659,786,1501,1522]" pageId="6" pageNumber="17">
|
||||
<figureCitation id="133A2A6D5D41152F2EACD192FC8CFA2A" box="[659,786,1501,1522]" captionStart="Figs" captionStartId="6.[330,373,1116,1135]" captionTargetBox="[336,1117,273,1093]" captionTargetId="figure-12@6.[328,1117,272,1096]" captionTargetPageId="6" captionText="Figs. 10–13. Phyllophaga lorencita. 10) protarsal claw, male; 11) genital capsule, lateral view; 12) paramera, ventrodistal view; 13) genital plates, female. Scale lines 5 1 mm, except Fig. 10 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900317" httpUri="https://zenodo.org/record/4900317/files/figure.png" pageId="6" pageNumber="17">Figs. 10–13</figureCitation>
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||||
</paragraph>
|
||||
<paragraph id="8BBE36E85D41152E2D75D248FC2FFB35" blockId="6.[307,1140,1543,1658]" lastBlockId="7.[279,1113,277,1261]" lastPageId="7" lastPageNumber="18" pageId="6" pageNumber="17">
|
||||
<emphasis id="B975EAFA5D41152F2D75D248FE34F9C2" bold="true" box="[330,426,1543,1562]" pageId="6" pageNumber="17">
|
||||
<typeStatus id="54BA884A5D41152F2D75D248FE38F9C2" box="[330,422,1543,1562]" pageId="6" pageNumber="17" type="holotype">Holotype</typeStatus>
|
||||
.
|
||||
</emphasis>
|
||||
Male. Clypeus, frons and pronotum shiny reddish yellow; elytra shiny testaceus, with two erect long setae at the base of each elytron and row of setae along the elytral suture, setae located toward the apex of elytra longer than preceding elytral setae; mouthparts, sterna, pygidium and legs shiny testaceous. Clypeus 4.2× wider than long, anterior border scarcely sinuate, with elevated margin, surface scarcely concave, coarsely rugopunctate and with some scattered short setae. Frontoclypeal suture straight and clearly impressed. Frons 1.3× wider than long, convex, coarsely punctate rugose, with short setae on disk. Antenna 9segmented, with 3segmented club, lamellae of 7th to 9th segments 1.2× longer than length of preceeding 5 segments combined. Frons 3.2× wider than dorsal diameter of eye. Eye canthus long and narrowed, with 8–10 setae. Labrum bilobed, wide and shallowly sinuated, with scattered slender setae. Mentum slightly convex, finely punctate, with long slender setae at sides, anterior border shallowly notched. Pronotum 1.8× wider than long and 2.7× wider than frons. Pronotal disk shiny, moderately punctaterugose, punctures irregularly separated from one another by 1–5 diameters, midline with shallow longitudinal depression spaced anteriorly; lateral borders clearly angled, lateral marginal bead irregularly crenulate, with scattered, long, slender setae; complete basal bead indicated by punctures; anterior angles slightly acute, prominent; posterior angles widely obtuse, rounded. Scutellum 1.7× wider than long, with some small punctures. Elytron 2.5× longer than wide, shiny, densely punctate; epipleural border very narrow, extended along complete margin, provided with a row of very short, slender setae; humeral callus rounded, prominent; apical callus rounded. Metathoracic wings completely developed. Propygidium shiny, densely punctatesetiferous. Pygidium scarcely convex, shiny, rugopunctate, with round punctures, with very short setae covering entire surface; apical margin with 16 long, slender setae; basal margin effaced medially. Pterosterna with long, dense, yellowish setae. Visible abdominal sternites II and IV slightly depressed, smooth and setiferous near the middline; sternite V granulate setiferous, with a shallow transverse depression on posterior half and some scattered short setae toward the sides; anal plate narrowed with a shallow transverse depression, with posterior border elevate and notched at midline, with 20 erect setae. Protibia slightly shorter than protarsus (1:1.3), with external border clearly tridentate, preapical spur small, slighly curved, acute, shorter than 2nd protarsomere. Mesotibia with one oblique, sharp, setiferous carina on external side; upper apical spur straight, narrow, and 0.1× shorter than lower spur. Metatibia shorter than metatarsus (1:1.2), with one oblique, sharp, setiferous carina on external side; upper apical spur articulated, curved, lanceolate, apex rounded, as long as basal metatarsomere, and 1.3× longer than lower spur; lower apical spur articulated with tibial border, with rounded apex. Tarsomeres semicylindrical, elongate, with enlarged apex, some setae apically and one longitudinal thin carina with a line of scattered, stout setae and other line of small setae at side. Protarsomere 1–4 with two carinae all along the ventral side. Tarsal claws symmetrical, similar on all legs, with curved tooth on the ventral border, slightly displaced toward the basal process (
|
||||
<figureCitation id="133A2A6D5D40152E2838D006FBD3FB84" box="[1031,1101,1097,1116]" captionStart="Figs" captionStartId="6.[330,373,1116,1135]" captionTargetBox="[336,1117,273,1093]" captionTargetId="figure-12@6.[328,1117,272,1096]" captionTargetPageId="6" captionText="Figs. 10–13. Phyllophaga lorencita. 10) protarsal claw, male; 11) genital capsule, lateral view; 12) paramera, ventrodistal view; 13) genital plates, female. Scale lines 5 1 mm, except Fig. 10 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900317" httpUri="https://zenodo.org/record/4900317/files/figure.png" pageId="7" pageNumber="18">Fig. 10</figureCitation>
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). Genital capsule with short parameres, dorsally and ventrally fused, preapical inner borders of the parameres turned out as a bladelike structure, apex with very short, toothlike ventral projections. Aedeagus short, with the apex of sclerotized support bifurcate and dorsobasal crestlike brush of progressively enlarged setae (
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<figureCitation id="133A2A6D5D40152E2FB4D0E5FB9EFB65" box="[907,1024,1194,1213]" captionStart="Figs" captionStartId="6.[330,373,1116,1135]" captionTargetBox="[336,1117,273,1093]" captionTargetId="figure-12@6.[328,1117,272,1096]" captionTargetPageId="6" captionText="Figs. 10–13. Phyllophaga lorencita. 10) protarsal claw, male; 11) genital capsule, lateral view; 12) paramera, ventrodistal view; 13) genital plates, female. Scale lines 5 1 mm, except Fig. 10 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900317" httpUri="https://zenodo.org/record/4900317/files/figure.png" pageId="7" pageNumber="18">Figs. 11–12</figureCitation>
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||||
). Tectum slighly convex. Length of genital capsule from apex of parameres to border of basal piece:
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||||
<quantity id="4CF99B0D5D40152E2D68D095FE3DFB35" box="[343,419,1242,1261]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="7" pageNumber="18" unit="mm" value="3.9">3.9 mm</quantity>
|
||||
. Total body length:
|
||||
<quantity id="4CF99B0D5D40152E2E5CD095FD25FB35" box="[611,699,1242,1261]" metricMagnitude="-2" metricUnit="m" metricValue="1.58" pageId="7" pageNumber="18" unit="mm" value="15.8">15.8 mm</quantity>
|
||||
. Humeral width:
|
||||
<quantity id="4CF99B0D5D40152E2F5ED095FC33FB35" box="[865,941,1242,1261]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="7" pageNumber="18" unit="mm" value="6.6">6.6 mm</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BBE36E85D40152E2D0DD148FDBFF9B8" blockId="7.[279,1112,1287,1658]" pageId="7" pageNumber="18">
|
||||
<emphasis id="B975EAFA5D40152E2D0DD148FE09FAC4" bold="true" box="[306,407,1287,1308]" pageId="7" pageNumber="18">
|
||||
<typeStatus id="54BA884A5D40152E2D0DD148FE0CFAC4" box="[306,402,1287,1308]" pageId="7" pageNumber="18" type="allotype">Allotype</typeStatus>
|
||||
.
|
||||
</emphasis>
|
||||
Female. Similar to the male except as follows: anterior border of frons elevated, disk of frons coarsely rugopunctate; antennae with lamellae of 7th to 9th segments shorter than the length of five preceeding ones (1:1.2). The longitudinal depression at the anterior part of the midline deeper than male; and also with coarser punctures. Visible abdominal sternites V convex, densely setiferous punctuated mainly toward the sides; anal plate 1.3× longer than male anal plate, convex, punctate setiferous, with 16 long slender setae near the posterior border. Meso and metatibiae each with one oblique, sharp, setiferous carina on external side. Both apical spurs of metatibia articulated, wide, lanceolate and slightly curved. Ventral genital plates slightly sclerotized, symmetrical, widened, ovate, without setae; dorsal genital plates narrowed, with short setae on the rounded apex (
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<figureCitation id="133A2A6D5D40152E2E84D27FFC92F99D" box="[699,780,1584,1605]" captionStart="Figs" captionStartId="6.[330,373,1116,1135]" captionTargetBox="[336,1117,273,1093]" captionTargetId="figure-12@6.[328,1117,272,1096]" captionTargetPageId="6" captionText="Figs. 10–13. Phyllophaga lorencita. 10) protarsal claw, male; 11) genital capsule, lateral view; 12) paramera, ventrodistal view; 13) genital plates, female. Scale lines 5 1 mm, except Fig. 10 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900317" httpUri="https://zenodo.org/record/4900317/files/figure.png" pageId="7" pageNumber="18">Fig. 13</figureCitation>
|
||||
). Total body length:
|
||||
<quantity id="4CF99B0D5D40152E2FCED27FFBCAF99D" box="[1009,1108,1584,1605]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="7" pageNumber="18" unit="mm" value="16.2">16.2 mm</quantity>
|
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. Humeral width: 7.0 mm.
|
||||
</paragraph>
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||||
<paragraph id="8BBE36E85D4015212D0DD22AFE12FE68" blockId="7.[279,1112,1287,1658]" lastBlockId="8.[307,1140,277,1232]" lastPageId="8" lastPageNumber="19" pageId="7" pageNumber="18">
|
||||
<emphasis id="B975EAFA5D40152E2D0DD22AFD8FF9A2" bold="true" box="[306,529,1637,1658]" pageId="7" pageNumber="18">
|
||||
<typeStatus id="54BA884A5D40152E2D0DD22AFE06F9A2" box="[306,408,1637,1658]" pageId="7" pageNumber="18" type="paratype">Paratype</typeStatus>
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Variation.
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||||
</emphasis>
|
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Males. Similar to
|
||||
<typeStatus id="54BA884A5D40152E2EEAD22AFCAEF9A2" box="[725,816,1637,1658]" pageId="7" pageNumber="18" type="holotype">holotype</typeStatus>
|
||||
except in total body length:
|
||||
<quantity id="4CF99B0D5D4F15212D0CD55AFE4AFEF2" box="[307,468,277,298]" metricMagnitude="-2" metricUnit="m" metricValue="1.3800000000000001" metricValueMax="1.6199999999999999" metricValueMin="1.1400000000000001" pageId="8" pageNumber="19" unit="mm" value="13.8" valueMax="16.2" valueMin="11.4">11.4–16.2 mm</quantity>
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, humeral width: 4.9–6.0 mm, pronotum of some specimens darker or lighter than
|
||||
<typeStatus id="54BA884A5D4F15212E1ED57FFDE1FE9D" box="[545,639,304,325]" pageId="8" pageNumber="19" type="holotype">holotype</typeStatus>
|
||||
, other specimens with antennal club 1.0–1.2× longer than length of preceeding five segments combined. Females similar to
|
||||
<typeStatus id="54BA884A5D4F15212D0CD52AFE19FEA2" box="[307,391,357,378]" pageId="8" pageNumber="19" type="allotype">allotype</typeStatus>
|
||||
except as follows: anterior border of frons less prominent, body punctation coarser or finer, total body length:
|
||||
<quantity id="4CF99B0D5D4F15212ED0D5CFFC10FE4D" box="[751,910,384,405]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.61" metricValueMin="1.29" pageId="8" pageNumber="19" unit="mm" value="14.5" valueMax="16.1" valueMin="12.9">12.9–16.1 mm</quantity>
|
||||
; humeral width:
|
||||
<quantity id="4CF99B0D5D4F1521287AD5CFFE16FE68" metricMagnitude="-3" metricUnit="m" metricValue="5.8" metricValueMax="6.5" metricValueMin="5.1" pageId="8" pageNumber="19" unit="mm" value="5.8" valueMax="6.5" valueMin="5.1">5.1– 6.5 mm</quantity>
|
||||
.
|
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</paragraph>
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</subSubSection>
|
||||
<subSubSection id="C31B65635D4F15212D72D5FAFD3EFD0D" pageId="8" pageNumber="19" type="materials_examined">
|
||||
<paragraph id="8BBE36E85D4F15212D72D5FAFD8FFD78" blockId="8.[307,1140,277,1232]" pageId="8" pageNumber="19">
|
||||
<emphasis id="B975EAFA5D4F15212D72D5FAFE49FE12" bold="true" box="[333,471,437,458]" pageId="8" pageNumber="19">Type Series.</emphasis>
|
||||
<location id="8EDE60335D4F15212DE0D5FAFDD7FE12" LSID="urn:lsid:plazi:treatment:03A887FE5D4115212D5AD18CFE1BFB08:8EDE60335D4F15212DE0D5FAFDD7FE12" box="[479,585,437,458]" name="Described" pageId="8" pageNumber="19">Described</location>
|
||||
from
|
||||
<specimenCount id="9D07FD615D4F15212EB3D5FAFD75FE12" box="[652,747,437,458]" pageId="8" pageNumber="19" type="male">67 males</specimenCount>
|
||||
and
|
||||
<specimenCount id="9D07FD615D4F15212F1ED5FAFC06FE12" box="[801,920,437,458]" pageId="8" pageNumber="19" type="female">25 females</specimenCount>
|
||||
.
|
||||
<materialsCitation id="3B693CB55D4F15212F9DD5FAFD78FDD8" collectingDate="1987-05-04" collectionCode="IN- BIO" collectorName="A. Solis" country="Costa Rica" county="San Ramon" elevation="800" latitude="10.35" location="Rio San Lorencito" longLatPrecision="1298" longitude="-84.416664" municipality="Rio San Lorencito" pageId="8" pageNumber="19" specimenCode="INBIO CR 1001 115762" specimenCount="1" specimenCount-male="1" stateProvince="Alajuela" typeStatus="holotype">
|
||||
<typeStatus id="54BA884A5D4F15212F9DD5FAFB9DFE12" box="[930,1027,437,458]" pageId="8" pageNumber="19" type="holotype">Holotype</typeStatus>
|
||||
<specimenCount id="9D07FD615D4F15212834D5FAFBA0FE12" box="[1035,1086,437,458]" pageId="8" pageNumber="19" type="male">male</specimenCount>
|
||||
(
|
||||
<collectionCode id="ED10AE2D5D4F15212872D5FAFEFBFE3D" pageId="8" pageNumber="19">INBIO</collectionCode>
|
||||
): ‘‘
|
||||
<collectingCountry id="F31676785D4F15212DBCD59FFDB5FE3D" box="[387,555,464,485]" name="Costa Rica" pageId="8" pageNumber="19">COSTA RICA</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49C5F80A5D4F15212E07D59FFD0DFE3D" box="[568,659,464,485]" country="Costa Rica" name="Alajuela" pageId="8" pageNumber="19">Alajuela</collectingRegion>
|
||||
,
|
||||
<collectingCounty id="62DF4E645D4F15212E9DD59FFCBFFE3D" box="[674,801,464,485]" pageId="8" pageNumber="19">San Ramón</collectingCounty>
|
||||
,
|
||||
<location id="8EDE60335D4F15212F10D59FFC66FE3D" LSID="urn:lsid:plazi:treatment:03A887FE5D4115212D5AD18CFE1BFB08:8EDE60335D4F15212F10D59FFC66FE3D" box="[815,1016,464,485]" country="Costa Rica" county="San Ramon" municipality="Rio San Lorencito" name="Rio San Lorencito" pageId="8" pageNumber="19" stateProvince="Alajuela">Río San Lorencito</location>
|
||||
,
|
||||
<quantity id="4CF99B0D5D4F15212838D59FFBD1FE3D" box="[1031,1103,464,485]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="19" unit="m" value="800.0">
|
||||
<elevation id="002CD1DB5D4F15212838D59FFBD1FE3D" box="[1031,1103,464,485]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="19" unit="m" value="800.0">800 m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<date id="FFBF10285D4F15212862D59FFEF7FDD8" pageId="8" pageNumber="19" value="1987-05-04">
|
||||
<collectingDate id="EFFBE9C05D4F15212862D59FFEF7FDD8" pageId="8" pageNumber="19" value="1987-05-04">4 V87</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F4533E5D4F15212D4BD5A4FE52FDD8" box="[372,460,491,512]" pageId="8" pageNumber="19">A. Solís</collectorName>
|
||||
,
|
||||
<specimenCode id="DBA79E935D4F15212DE8D5A4FD7CFDD8" box="[471,738,491,512]" pageId="8" pageNumber="19">INBIO CR 1001 115762</specimenCode>
|
||||
.
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D4F15212ED9D5A4FC63FDED" collectingDate="1993-06-30" collectionCode="INBIO" collectorName="P. Campos" country="Costa Rica" county="San Ramon" elevation="1120" latitude="10.35" location="Chirripo" longLatPrecision="1298" longitude="-84.416664" municipality="Turrialba" pageId="8" pageNumber="19" specimenCode="INBIO CR 1001 848913" specimenCount="1" specimenCount-female="1" stateProvince="Cartago" typeStatus="allotype">
|
||||
’’
|
||||
<typeStatus id="54BA884A5D4F15212EC8D5A4FCCCFDD8" box="[759,850,491,512]" pageId="8" pageNumber="19" type="allotype">Allotype</typeStatus>
|
||||
<specimenCount id="9D07FD615D4F15212F66D5A4FC3EFDD8" box="[857,928,491,512]" pageId="8" pageNumber="19" type="female">female</specimenCount>
|
||||
(
|
||||
<collectionCode id="ED10AE2D5D4F15212F90D5A4FC60FDD8" box="[943,1022,491,512]" pageId="8" pageNumber="19">INBIO</collectionCode>
|
||||
): ‘‘
|
||||
<collectingCountry id="F31676785D4F15212827D5A4FEE8FDC2" name="Costa Rica" pageId="8" pageNumber="19">COSTA RICA</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49C5F80A5D4F15212DBFD64AFE4BFDC2" box="[384,469,517,538]" country="Costa Rica" name="Cartago" pageId="8" pageNumber="19">Cartago</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="6BDAAC925D4F15212DDDD64AFDDAFDC2" box="[482,580,517,538]" pageId="8" pageNumber="19">Turrialba</collectingMunicipality>
|
||||
,
|
||||
<location id="8EDE60335D4F15212E6FD64AFD37FDC2" LSID="urn:lsid:plazi:treatment:03A887FE5D4115212D5AD18CFE1BFB08:8EDE60335D4F15212E6FD64AFD37FDC2" box="[592,681,517,538]" country="Costa Rica" county="San Ramon" municipality="Turrialba" name="Chirripo" pageId="8" pageNumber="19" stateProvince="Cartago">Chirripó</location>
|
||||
,
|
||||
<location id="8EDE60335D4F15212E88D64AFB98FDC2" LSID="urn:lsid:plazi:treatment:03A887FE5D4115212D5AD18CFE1BFB08:8EDE60335D4F15212E88D64AFB98FDC2" box="[695,1030,517,538]" country="Costa Rica" county="San Ramon" municipality="Turrialba" name="Area de Conservacion Amistad" pageId="8" pageNumber="19" stateProvince="Cartago">Area de Conservación Amistad</location>
|
||||
,
|
||||
<location id="8EDE60335D4F1521282ED64AFEC0FDED" LSID="urn:lsid:plazi:treatment:03A887FE5D4115212D5AD18CFE1BFB08:8EDE60335D4F1521282ED64AFEC0FDED" country="Costa Rica" county="San Ramon" municipality="Turrialba" name="Grano de Oro" pageId="8" pageNumber="19" stateProvince="Cartago">Grano de Oro</location>
|
||||
,
|
||||
<quantity id="4CF99B0D5D4F15212D54D66FFE58FDED" box="[363,454,544,565]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="8" pageNumber="19" unit="m" value="1120.0">
|
||||
<elevation id="002CD1DB5D4F15212D54D66FFE58FDED" box="[363,454,544,565]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="8" pageNumber="19" unit="m" value="1120.0">1,120 m</elevation>
|
||||
</quantity>
|
||||
, 19/
|
||||
<date id="FFBF10285D4F15212DCAD66FFDC7FDED" box="[501,601,544,565]" pageId="8" pageNumber="19" value="1993-06-30">
|
||||
<collectingDate id="EFFBE9C05D4F15212DCAD66FFDC7FDED" box="[501,601,544,565]" pageId="8" pageNumber="19" value="1993-06-30">30VI93</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F4533E5D4F15212E5AD66FFD4BFDED" box="[613,725,544,565]" pageId="8" pageNumber="19">P. Campos</collectorName>
|
||||
<specimenCode id="DBA79E935D4F15212EE1D66FFC6EFDED" box="[734,1008,544,565]" pageId="8" pageNumber="19">INBIO CR 1001 848913</specimenCode>
|
||||
.’’
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D4F15212839D66FFD13FD88" collectionCode="CASC, CNC, INBIO, MXAL, TAMU, ZMHU" collectorName="A. Solis" country="Costa Rica" county="San Ramon" elevation="800" latitude="10.35" location="Turrialba" longLatPrecision="1298" longitude="-84.416664" municipality="Rio San Lorencito" pageId="8" pageNumber="19" specimenCount="2" specimenCount-female="24" specimenCount-male="2" stateProvince="Alajuela" typeStatus="paratype">
|
||||
<typeStatus id="54BA884A5D4F15212839D66FFBEEFDED" box="[1030,1136,544,565]" pageId="8" pageNumber="19" type="paratype">Paratypes</typeStatus>
|
||||
: same data as holotype (
|
||||
<specimenCount id="9D07FD615D4F15212E14D674FD1FFD88" box="[555,641,571,592]" count="2" pageId="8" pageNumber="19" type="male">2 males</specimenCount>
|
||||
);
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D4F15212EABD674FBEDFD88" box="[660,1139,571,592]" collectingDate="1993-06-30" collectionCode="CASC, CNC, INBIO, MXAL, TAMU, ZMHU" collectorName="P. Campos" country="Costa Rica" county="San Ramon" elevation="800" latitude="10.35" location="Turrialba" longLatPrecision="1298" longitude="-84.416664" municipality="Rio San Lorencito" pageId="8" pageNumber="19" specimenCode="INBIO CR 1001 848913" specimenCount="53" specimenCount-female="14" specimenCount-male="39" stateProvince="Alajuela" typeStatus="paratype">
|
||||
same data as allotype (
|
||||
<specimenCount id="9D07FD615D4F15212FBBD674FC78FD88" box="[900,998,571,592]" count="39" pageId="8" pageNumber="19" type="male">39 males</specimenCount>
|
||||
,
|
||||
<specimenCount id="9D07FD615D4F15212FCED674FBF6FD88" box="[1009,1128,571,592]" count="14" pageId="8" pageNumber="19" type="female">14 females</specimenCount>
|
||||
);
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D4F15212D0CD61AFD93FD5D" collectingDate="1989-07-10" collectionCode="CASC, CNC, INBIO, MXAL, TAMU, ZMHU" collectorName="A. Solis" country="Costa Rica" county="San Ramon" elevation="800" location="Rio San Lorencito" municipality="San Ramon" pageId="8" pageNumber="19" specimenCount="35" specimenCount-female="10" specimenCount-male="25" stateProvince="Alajuela" typeStatus="holotype">
|
||||
<collectingRegion id="49C5F80A5D4F15212D0CD61AFE10FDB2" box="[307,398,597,618]" country="Costa Rica" name="Alajuela" pageId="8" pageNumber="19">Alajuela</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="6BDAAC925D4F15212DA0D61AFDBFFDB2" box="[415,545,597,618]" pageId="8" pageNumber="19">San Ramón</collectingMunicipality>
|
||||
,
|
||||
<location id="8EDE60335D4F15212E0ED61AFC9EFDB2" LSID="urn:lsid:plazi:treatment:03A887FE5D4115212D5AD18CFE1BFB08:8EDE60335D4F15212E0ED61AFC9EFDB2" box="[561,768,597,618]" country="Costa Rica" county="San Ramon" municipality="San Ramon" name="Rio San Lorencito" pageId="8" pageNumber="19" stateProvince="Alajuela">Río San Lorencito</location>
|
||||
,
|
||||
<quantity id="4CF99B0D5D4F15212F2FD61AFCC5FDB2" box="[784,859,597,618]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="19" unit="m" value="800.0">
|
||||
<elevation id="002CD1DB5D4F15212F2FD61AFCC5FDB2" box="[784,859,597,618]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="19" unit="m" value="800.0">800 m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<date id="FFBF10285D4F15212F53D61AFC46FDB2" box="[876,984,597,618]" pageId="8" pageNumber="19" value="1989-07-10">
|
||||
<collectingDate id="EFFBE9C05D4F15212F53D61AFC46FDB2" box="[876,984,597,618]" pageId="8" pageNumber="19" value="1989-07-10">10VII89</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F4533E5D4F15212FD6D61AFBDDFDB2" box="[1001,1091,597,618]" pageId="8" pageNumber="19">A. Solís</collectorName>
|
||||
(
|
||||
<specimenCount id="9D07FD615D4F15212867D61AFEEDFD5D" count="25" pageId="8" pageNumber="19" type="male">25 males</specimenCount>
|
||||
,
|
||||
<specimenCount id="9D07FD615D4F15212DBBD63FFD9CFD5D" box="[388,514,624,645]" count="10" pageId="8" pageNumber="19" type="female">10 females</specimenCount>
|
||||
).
|
||||
</materialsCitation>
|
||||
Paratypes deposited in
|
||||
<collectionCode id="ED10AE2D5D4F15212F1ED63FFCF6FD5D" box="[801,872,624,645]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="8" pageNumber="19">CASC</collectionCode>
|
||||
,
|
||||
<collectionCode id="ED10AE2D5D4F15212F45D63FFC2CFD5D" box="[890,946,624,645]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" pageId="8" pageNumber="19">CNC</collectionCode>
|
||||
,
|
||||
<collectionCode id="ED10AE2D5D4F15212FFCD63FFB90FD5D" box="[963,1038,624,645]" pageId="8" pageNumber="19">INBIO</collectionCode>
|
||||
,
|
||||
<collectionCode id="ED10AE2D5D4F15212820D63FFBF1FD5D" box="[1055,1135,624,645]" pageId="8" pageNumber="19">MXAL</collectionCode>
|
||||
,
|
||||
<collectionCode id="ED10AE2D5D4F15212D0CD6C4FE1CFD78" box="[307,386,651,672]" collectionName="USA, Texas, College Station, Texas A & M University" pageId="8" pageNumber="19">TAMU</collectionCode>
|
||||
, and
|
||||
<collectionCode id="ED10AE2D5D4F15212D82D6C4FD93FD78" box="[445,525,651,672]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="8" pageNumber="19">ZMHU</collectionCode>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BBE36E85D4F15212D72D6EAFD3EFD0D" blockId="8.[307,1140,277,1232]" pageId="8" pageNumber="19">
|
||||
<emphasis id="B975EAFA5D4F15212D72D6EAFE6EFD62" bold="true" box="[333,496,677,698]" pageId="8" pageNumber="19">Type Locality.</emphasis>
|
||||
<location id="8EDE60335D4F15212DC8D6EAFD22FD62" LSID="urn:lsid:plazi:treatment:03A887FE5D4115212D5AD18CFE1BFB08:8EDE60335D4F15212DC8D6EAFD22FD62" box="[503,700,677,698]" country="Costa Rica" latitude="10.35" longLatPrecision="1298" longitude="-84.416664" municipality="San Ramon" name="Rio San Lorencito" pageId="8" pageNumber="19" stateProvince="Alajuela">Río San Lorencito</location>
|
||||
,
|
||||
<collectingMunicipality id="6BDAAC925D4F15212EF7D6EAFCDBFD62" box="[712,837,677,698]" pageId="8" pageNumber="19">San Ramón</collectingMunicipality>
|
||||
, province of
|
||||
<collectingRegion id="49C5F80A5D4F15212FEED6EAFBB2FD62" box="[977,1068,677,698]" country="Costa Rica" name="Alajuela" pageId="8" pageNumber="19">Alajuela</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="F31676785D4F15212808D6EAFEFDFD0D" name="Costa Rica" pageId="8" pageNumber="19">Costa Rica</collectingCountry>
|
||||
(approx.
|
||||
<geoCoordinate id="EE35502F5D4F15212DF3D68FFDB9FD0D" box="[460,551,704,725]" degrees="10" direction="north" minutes="21" orientation="latitude" pageId="8" pageNumber="19" precision="925" value="10.35">10°21'N</geoCoordinate>
|
||||
;
|
||||
<geoCoordinate id="EE35502F5D4F15212E0CD68FFD0BFD0D" box="[563,661,704,725]" degrees="84" direction="west" minutes="25" orientation="longitude" pageId="8" pageNumber="19" precision="925" value="-84.416664">84°25'W</geoCoordinate>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D4F15212D72D695FDE2FC1D" pageId="8" pageNumber="19" type="biology_ecology">
|
||||
<paragraph id="8BBE36E85D4F15212D72D695FDE2FC1D" blockId="8.[307,1140,277,1232]" pageId="8" pageNumber="19">
|
||||
<emphasis id="B975EAFA5D4F15212D72D695FD9EFD37" bold="true" box="[333,512,730,751]" pageId="8" pageNumber="19">Biological Data.</emphasis>
|
||||
Males and females of
|
||||
<taxonomicName id="4C014D6B5D4F15212EC8D694FCEAFD28" box="[759,884,731,752]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="lorencita" status="sp. nov.">
|
||||
<emphasis id="B975EAFA5D4F15212EC8D694FCEAFD28" box="[759,884,731,752]" italics="true" pageId="8" pageNumber="19">P. lorencita</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A24657815D4F15212F43D694FC29FD28" box="[892,951,731,752]" pageId="8" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
|
||||
were collected at lights at two sites separated by
|
||||
<quantity id="4CF99B0D5D4F15212EB3D6BAFD7EFCD2" box="[652,736,757,778]" metricMagnitude="5" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="19" unit="km" value="130.0">130 km</quantity>
|
||||
, near San Lorencito river, located on Tilaran ridge, between
|
||||
<quantity id="4CF99B0D5D4F15212E13D75FFD38FCFD" box="[556,678,784,805]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="8" pageNumber="19" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">800–900 m</quantity>
|
||||
of altitude, and Grano de Oro, on the Talamanca ridge, at
|
||||
<quantity id="4CF99B0D5D4F15212DD8D764FDA3FC98" box="[487,573,811,832]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="8" pageNumber="19" unit="m" value="1120.0">1,120 m</quantity>
|
||||
of altitude. Phenology: May (3), June (54), July (35). Other species of
|
||||
<taxonomicName id="4C014D6B5D4F15212DCBD709FDE4FC83" box="[500,634,838,859]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B975EAFA5D4F15212DCBD709FDE4FC83" box="[500,634,838,859]" italics="true" pageId="8" pageNumber="19">Phyllophaga</emphasis>
|
||||
</taxonomicName>
|
||||
flying at the same time and places were:
|
||||
<emphasis id="B975EAFA5D4F15212863D709FEE3FCAD" italics="true" pageId="8" pageNumber="19">P. caraga</emphasis>
|
||||
Saylor,
|
||||
<emphasis id="B975EAFA5D4F15212DE5D72FFDF6FCAD" box="[474,616,864,885]" italics="true" pageId="8" pageNumber="19">P. chiriquina</emphasis>
|
||||
Bates,
|
||||
<emphasis id="B975EAFA5D4F15212E82D72FFCB1FCAD" box="[701,815,864,885]" italics="true" pageId="8" pageNumber="19">P. densata</emphasis>
|
||||
Moser,
|
||||
<emphasis id="B975EAFA5D4F15212FB4D72FFB99FCAD" box="[907,1031,864,885]" italics="true" pageId="8" pageNumber="19">P. gigantea</emphasis>
|
||||
Bates,
|
||||
<taxonomicName id="4C014D6B5D4F15212863D72FFE67FC48" authority="Saylor, P." authorityName="Saylor, P." class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="guapiles">
|
||||
<emphasis id="B975EAFA5D4F15212863D72FFE12FC48" italics="true" pageId="8" pageNumber="19">P. guapiles</emphasis>
|
||||
Saylor,
|
||||
<emphasis id="B975EAFA5D4F15212DDDD734FE67FC48" box="[482,505,891,912]" italics="true" pageId="8" pageNumber="19">P.</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B975EAFA5D4F15212E3FD734FDEFFC48" box="[512,625,891,912]" italics="true" pageId="8" pageNumber="19">guapilesea</emphasis>
|
||||
Saylor,
|
||||
<emphasis id="B975EAFA5D4F15212EF7D734FCFFFC48" box="[712,865,891,912]" italics="true" pageId="8" pageNumber="19">P. nevermanni</emphasis>
|
||||
Saylor,
|
||||
<emphasis id="B975EAFA5D4F15212F87D734FBBBFC48" box="[952,1061,891,912]" italics="true" pageId="8" pageNumber="19">P. orosina</emphasis>
|
||||
Moser,
|
||||
<emphasis id="B975EAFA5D4F15212D0CD7D9FE03FC73" box="[307,413,918,939]" italics="true" pageId="8" pageNumber="19">P. prolixa</emphasis>
|
||||
Bates,
|
||||
<taxonomicName id="4C014D6B5D4F15212DCFD7D9FC96FC73" authority="Blanchard, P." authorityName="Blanchard, P." box="[496,776,917,939]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="pruinosa">
|
||||
<emphasis id="B975EAFA5D4F15212DCFD7D9FDF3FC73" box="[496,621,918,939]" italics="true" pageId="8" pageNumber="19">P. pruinosa</emphasis>
|
||||
Blanchard,
|
||||
<emphasis id="B975EAFA5D4F15212ECED7D9FC96FC73" box="[753,776,918,939]" italics="true" pageId="8" pageNumber="19">P.</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B975EAFA5D4F15212F2FD7D9FCEAFC73" box="[784,884,918,939]" italics="true" pageId="8" pageNumber="19">setidorsis</emphasis>
|
||||
Bates, and another two new species under description.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D4F15212D72D785FD6FFB42" pageId="8" pageNumber="19" type="discussion">
|
||||
<paragraph id="8BBE36E85D4F15212D72D785FD6FFB42" blockId="8.[307,1140,277,1232]" pageId="8" pageNumber="19">
|
||||
<emphasis id="B975EAFA5D4F15212D72D785FE27FC07" bold="true" box="[333,441,970,991]" pageId="8" pageNumber="19">Remarks.</emphasis>
|
||||
Is not possible to include
|
||||
<taxonomicName id="4C014D6B5D4F15212EDCD784FBEDFC38" box="[739,1139,971,992]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="lorencita" status="sp. nov." subGenus="Phyllophaga">
|
||||
<emphasis id="B975EAFA5D4F15212EDCD784FCF7FC38" box="[739,873,971,992]" italics="true" pageId="8" pageNumber="19">Phyllophaga</emphasis>
|
||||
(
|
||||
<emphasis id="B975EAFA5D4F15212F43D784FB9FFC38" box="[892,1025,971,992]" italics="true" pageId="8" pageNumber="19">Phyllophaga</emphasis>
|
||||
)
|
||||
<emphasis id="B975EAFA5D4F1521282AD784FBEDFC38" box="[1045,1139,971,992]" italics="true" pageId="8" pageNumber="19">lorencita</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A24657815D4F15212D0CD7AAFEF2FC22" box="[307,364,997,1018]" pageId="8" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
|
||||
in any known species group (
|
||||
<emphasis id="B975EAFA5D4F15212E92D7A9FD79FC23" box="[685,743,998,1019]" italics="true" pageId="8" pageNumber="19">sensu</emphasis>
|
||||
<bibRefCitation id="EF904B195D4F15212ED0D7AAFCE8FC22" author="Moron" box="[751,886,997,1018]" firstAuthor="Moron" pageId="8" pageNumber="19" refId="ref11224" refString="Moron, M. A. 1986. El genero Phyllophaga en Mexico. Morfologia, distribucion y sistematica supraespecifica (Insecta: Coleoptera). Publ. 20, Instituto de Ecologia, Mexico, D. F. 341 pp." type="book" year="1986">Morón 1986</bibRefCitation>
|
||||
). This new species may be related to
|
||||
<taxonomicName id="4C014D6B5D4F15212DFED04FFC85FBCD" authority="Saylor" authorityName="Saylor" authorityYear="1936" box="[449,795,1024,1045]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="reventazona">
|
||||
<emphasis id="B975EAFA5D4F15212DFED04FFD51FBCD" box="[449,719,1024,1045]" italics="true" pageId="8" pageNumber="19">Phyllophaga reventazona</emphasis>
|
||||
Saylor
|
||||
</taxonomicName>
|
||||
(Atlantic lowlands,
|
||||
<collectingCountry id="F31676785D4F15212FC8D04FFBF2FBCD" box="[1015,1132,1024,1045]" name="Costa Rica" pageId="8" pageNumber="19">Costa Rica</collectingCountry>
|
||||
) by the number and proportions of antenomeres, as well as the shape of the parameres and the dorsal vestiture of aedeagus, but the dense and erect pronotal vestiture, the absence of shallow longitudinal depression on the anterior half of the pronotum, the dark reddish brown body color of
|
||||
<emphasis id="B975EAFA5D4F15212FF3D024FBEDFB58" box="[972,1139,1131,1152]" italics="true" pageId="8" pageNumber="19">
|
||||
<taxonomicName id="4C014D6B5D4F15212FF3D024FBF0FB58" authorityName="Saylor" authorityYear="1936" box="[972,1134,1131,1152]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="reventazona">P. reventazona</taxonomicName>
|
||||
,
|
||||
</emphasis>
|
||||
will aid in the separation of both species.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D4F15212D72D0EFFE1BFB08" pageId="8" pageNumber="19" type="etymology">
|
||||
<paragraph id="8BBE36E85D4F15212D72D0EFFE1BFB08" blockId="8.[307,1140,277,1232]" pageId="8" pageNumber="19">
|
||||
<emphasis id="B975EAFA5D4F15212D72D0EFFE52FB6D" bold="true" box="[333,460,1184,1205]" pageId="8" pageNumber="19">Etymology.</emphasis>
|
||||
Derived from Río San Lorencito, at the eastern Cordillera de Tilarán.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
373
data/03/A8/87/03A887FE5D44152F2D78D1DEFE52FA7A.xml
Normal file
373
data/03/A8/87/03A887FE5D44152F2D78D1DEFE52FA7A.xml
Normal file
|
|
@ -0,0 +1,373 @@
|
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<document id="88DE0F976562B46A53A5F4AE6258DCAB" ID-DOI="10.1649/0010-065x(2001)055[0011:snsops]2.0.co;2" ID-ISSN="1938-4394" ID-Zenodo-Dep="4900311" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1622827801739" checkinUser="carolina" docAuthor="Morón, Miguel Angel & Solís, Angel" docDate="2001" docId="03A887FE5D44152F2D78D1DEFE52FA7A" docLanguage="en" docName="TheColeopBull.55.1.11-29.pdf.imf" docOrigin="The Coleopterists Bulletin 55 (1)" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Phyllophaga (Phyllophaga) tapantina Moron and Solis, new species" docType="treatment" docVersion="1" lastPageNumber="17" masterDocId="FF91FF865D4715292C3FD44FFF9EFFD8" masterDocTitle="Seven New Species Of Phyllophaga (S. Str.) Harris From Costa Rica (Coleoptera: Melolonthidae: Melolonthinae)" masterLastPageNumber="29" masterPageNumber="11" pageNumber="14" updateTime="1730139982923" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:mods id="1C547EF2D22252D52B27AF5901DFE73C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2719A1E82640A7C2D1B562A924B586CB">
|
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<mods:title id="ECA2C230C05A3247119B27CA55A0F21C">Seven New Species Of Phyllophaga (S. Str.) Harris From Costa Rica (Coleoptera: Melolonthidae: Melolonthinae)</mods:title>
|
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</mods:titleInfo>
|
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<mods:name id="4FE52B4C965EBDFA4DA9F0B13066C328" type="personal">
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<mods:role id="4EA5FAF4BC867A54E80BBBE7DF8D291E">
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<mods:roleTerm id="D2403A95DC9A329A6798130B7C4029BB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1ABD07C2B869F10BE306B833FEBBD7FE">Morón, Miguel Angel</mods:namePart>
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<mods:affiliation id="182F90029378207CFCAD08990301725A">Departamento de Entomología Instituto de Ecología, A. C. (sector SEPCONACYT) Apartado Postal 63 Xalapa, Veracruz 91000, MÉXICO</mods:affiliation>
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</mods:name>
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</mods:role>
|
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<mods:namePart id="4B52E10918D2887D103BF73BBCDEC611">Solís, Angel</mods:namePart>
|
||||
<mods:affiliation id="215B736F9FE647251FE455E724CD7B4A">Departamento de Entomología Instituto Nacional de Biodiversidad (INBio) Apartado Postal 22 3100 Santo Domingo de Heredia, COSTA RICA</mods:affiliation>
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</mods:name>
|
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<mods:typeOfResource id="9630F9EF7F325B16E515EB336F8EF6ED">text</mods:typeOfResource>
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<mods:title id="6CCF8F37D3408669D017A32723B54FA8">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="A4C7D6387AC8FAFEFC317F95C2847EF7">
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<mods:date id="95D38989A31D9CCB7F097AD69E60812D">2001</mods:date>
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|
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|
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<mods:number id="F36B4F967694081C5EE9FDA73B361B36">55</mods:number>
|
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|
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<mods:identifier id="13AE99091F8E6EEDD34FA01746E88630" type="ISSN">1938-4394</mods:identifier>
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</mods:mods>
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<treatment id="03A887FE5D44152F2D78D1DEFE52FA7A" LSID="urn:lsid:plazi:treatment:03A887FE5D44152F2D78D1DEFE52FA7A" httpUri="http://treatment.plazi.org/id/03A887FE5D44152F2D78D1DEFE52FA7A" lastPageId="6" lastPageNumber="17" pageId="3" pageNumber="14">
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<subSubSection id="C31B65635D44152A2D78D1DEFBB6FA7E" box="[327,1064,1425,1446]" pageId="3" pageNumber="14" type="nomenclature">
|
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<paragraph id="8BBE36E85D44152A2D78D1DEFBB6FA7E" blockId="3.[327,1064,1425,1473]" box="[327,1064,1425,1446]" pageId="3" pageNumber="14">
|
||||
<taxonomicName id="4C014D6B5D44152A2D78D1DEFC06FA7E" authority="Moron and Solis" authorityName="Moron and Solis" box="[327,920,1425,1446]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="14" phylum="Arthropoda" rank="species" species="tapantina" status="sp. nov." subGenus="Phyllophaga">
|
||||
<emphasis id="B975EAFA5D44152A2D78D1DEFE53FA7E" box="[327,461,1425,1446]" italics="true" pageId="3" pageNumber="14">Phyllophaga</emphasis>
|
||||
(
|
||||
<emphasis id="B975EAFA5D44152A2DE0D1DEFDFAFA7E" box="[479,612,1425,1446]" italics="true" pageId="3" pageNumber="14">Phyllophaga</emphasis>
|
||||
)
|
||||
<emphasis id="B975EAFA5D44152A2E4AD1DEFD47FA7E" box="[629,729,1425,1446]" italics="true" pageId="3" pageNumber="14">tapantina</emphasis>
|
||||
Morón and Solís
|
||||
</taxonomicName>
|
||||
,
|
||||
<emphasis id="B975EAFA5D44152A2F9AD1DEFBB6FA7E" bold="true" box="[933,1064,1425,1446]" pageId="3" pageNumber="14">
|
||||
<taxonomicNameLabel id="A24657815D44152A2F9AD1DEFBB6FA7E" box="[933,1064,1425,1446]" pageId="3" pageNumber="14" rank="species">new species</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D44152C2EB9D1E3FE34FBD7" lastPageId="5" lastPageNumber="16" pageId="3" pageNumber="14" type="description">
|
||||
<paragraph id="8BBE36E85D44152A2EB9D1E3FD77FA19" blockId="3.[327,1064,1425,1473]" box="[646,745,1452,1473]" pageId="3" pageNumber="14">
|
||||
<figureCitation id="133A2A6D5D44152A2EB9D1E3FD77FA19" box="[646,745,1452,1473]" captionStart="Figs" captionStartId="4.[330,373,836,855]" captionTargetBox="[340,1107,276,815]" captionTargetId="figure-12@4.[337,1108,272,816]" captionTargetPageId="4" captionText="Figs. 6–9. Phyllophaga tapantina. 6) protarsal claw, male; 7) genital capsule, lateral view; 8) paramera, distal view; 9) genital plates, female. Scale lines 5 1 mm, except Fig. 6 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900315" httpUri="https://zenodo.org/record/4900315/files/figure.png" pageId="3" pageNumber="14">Figs. 6–9</figureCitation>
|
||||
</paragraph>
|
||||
<paragraph id="8BBE36E85D44152C2D10D198FC23FDC0" blockId="3.[279,1113,1495,1658]" lastBlockId="5.[279,1112,277,536]" lastPageId="5" lastPageNumber="16" pageId="3" pageNumber="14">
|
||||
<emphasis id="B975EAFA5D44152A2D10D198FE11FA32" bold="true" box="[303,399,1495,1514]" pageId="3" pageNumber="14">
|
||||
<typeStatus id="54BA884A5D44152A2D10D198FE15FA32" box="[303,395,1495,1514]" pageId="3" pageNumber="14" type="holotype">Holotype</typeStatus>
|
||||
.
|
||||
</emphasis>
|
||||
Male. Clypeus, frons, pronotum and elytra shiny dark reddish brown; pronotum with scarce, erect, long, setae; elytra with greenish vitreus luster, with long setae around the scutellum and scattered minute setae only toward the apex; mouthparts, sterna, pygidium and legs shiny reddish brown. Clypeus 3.0× wider than long, anterior border scarcely sinuate, with poorly elevated margin, surface nearly flat, with scarce irregularlydistributed, deep, round punctures and some scattered microscopic setae. Frontoclypeal suture sinuate and deeply impressed. Frons 1.8× wider than long, convex, with reduced number of coarse wide punctures, long slender setae at sides and short setae scattered on disk. Antenna 10segmented, with 3segmented club, lamellae of 8th to 10th segments 1.3× longer than length of preceeding 6 segments combined. Frons 2.8× wider than dorsal diameter of eye. Eye canthus long and narrowed, with 77 setae. Labrum bilobed, widely sinuated, with scattered slender setae. Mentum slightly convex, impunctate, with scarce slender setae at sides, anterior border widely sinuate. Pronotum 1.6× wider than long and 2.8× wider than frons. Pronotal disk shiny, with some deep, round punctures irregularly separated from one another by 1–5 diameters; lateral borders widely angulated, lateral marginal bead irregularly crenulate, with scattered, long, slender setae; basal bead vaguely indicated toward the posterior angles; anterior angles acute, clearly prominent; posterior angles obtuse, slighly prominent. Scutellum 1.3× wider than long, smooth, without punctures. Elytron 2.8× longer than wide, smooth shiny, with scattered punctures on disk and dense uniformly punctate at external sides; epipleural border very narrow, extended along the complete margin, provided with fringe of mediumsize setae; humeral callus rounded, prominent; apical callus rounded. Metathoracic wings completely developed. Propygidium shiny, densely punctate setose. Pygidium moderately convex, shiny, shallowly punctate rugose with scattered erect setae; apical margin with 14 long, slender setae; basal margin effaced medially. Pterosterna with long, dense, yellowish setae. Visible abdominal sternites II and IV convex, smooth and glabrous near the middline; sternite V convex, shiny, with setiferous punctures at the posterior half and numerous setae toward the sides; anal plate narrowed, with a transverse shallow concavity and longitudinal furrow at midline, anterior and posterior borders slightly elevated, with 24 erect setae near the posterior border. Protibia as long as protarsus (1:1), with external border tridentate, apical and subapical teeth long, wide, apices rounded, preapical spur acute, nearly straight, as long as 1st protarsomere. Mesotibia with one oblique, sharp, setiferous carina on external side; upper apical spur straight, narrow, as long as lower spur. Metatibia 0.8× shorter than metatarsus, with one oblique, sharp, setiferous carina on external side; upper apical spur articulated, curved, with apex rounded, 1.3× longer than basal metatarsomere, and 1.3× longer than lower spur; lower apical spur articulated with tibial border, with rounded apex. Tarsomeres semicylindrical, not much elongated, with enlarged apex, some setae apically and one line of short setae close to fine longitudinal carina on ventral side. Protarsomeres I–IV with a subapical, small spine. Tarsal claws symmetrical, similar on all legs, each with a large acute tooth at the middle of ventral border (
|
||||
<figureCitation id="133A2A6D5D42152C2D8ED5C2FE6EFE78" box="[433,496,397,416]" captionStart="Figs" captionStartId="4.[330,373,836,855]" captionTargetBox="[340,1107,276,815]" captionTargetId="figure-12@4.[337,1108,272,816]" captionTargetPageId="4" captionText="Figs. 6–9. Phyllophaga tapantina. 6) protarsal claw, male; 7) genital capsule, lateral view; 8) paramera, distal view; 9) genital plates, female. Scale lines 5 1 mm, except Fig. 6 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900315" httpUri="https://zenodo.org/record/4900315/files/figure.png" pageId="5" pageNumber="16">Fig. 6</figureCitation>
|
||||
). Genital capsule with short parameres, dorsally and ventrally fused, ringshaped, apex below with short, triangular bladelike projection. Aedeagus short, wide, with preapical ventral dense patches of spinules, dorsal sclerotized plates, narrowed sclerotized support, and basidorsal brush of long setae (
|
||||
<figureCitation id="133A2A6D5D42152C2F9ED59AFC61FE30" box="[929,1023,469,488]" captionStart="Figs" captionStartId="4.[330,373,836,855]" captionTargetBox="[340,1107,276,815]" captionTargetId="figure-12@4.[337,1108,272,816]" captionTargetPageId="4" captionText="Figs. 6–9. Phyllophaga tapantina. 6) protarsal claw, male; 7) genital capsule, lateral view; 8) paramera, distal view; 9) genital plates, female. Scale lines 5 1 mm, except Fig. 6 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900315" httpUri="https://zenodo.org/record/4900315/files/figure.png" pageId="5" pageNumber="16">Figs. 7–8</figureCitation>
|
||||
). Tectum uniformly convex. Length of genital capsule from apex of parameres to border of basal piece: 6.0 mm. Total body length: 23.0 mm. Humeral width:
|
||||
<quantity id="4CF99B0D5D42152C2F5ED64AFC27FDC0" box="[865,953,517,536]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="5" pageNumber="16" unit="mm" value="13.2">13.2 mm</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
<caption id="DF7E66605D43152D2D75D70BFE42FC50" ID-DOI="http://doi.org/10.5281/zenodo.4900315" ID-Zenodo-Dep="4900315" httpUri="https://zenodo.org/record/4900315/files/figure.png" pageId="4" pageNumber="15" startId="4.[330,373,836,855]" targetBox="[340,1107,276,815]" targetPageId="4">
|
||||
<paragraph id="8BBE36E85D43152D2D75D70BFE42FC50" blockId="4.[307,1139,836,904]" pageId="4" pageNumber="15">
|
||||
<emphasis id="B975EAFA5D43152D2D75D70BFE36FC8F" bold="true" box="[330,424,836,855]" pageId="4" pageNumber="15">Figs. 6–9.</emphasis>
|
||||
<emphasis id="B975EAFA5D43152D2DFFD70AFD03FC80" box="[448,669,837,856]" italics="true" pageId="4" pageNumber="15">
|
||||
<taxonomicName id="4C014D6B5D43152D2DFFD70AFD07FC80" box="[448,665,837,856]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="15" phylum="Arthropoda" rank="species" species="tapantina">Phyllophaga tapantina</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
<emphasis id="B975EAFA5D43152D2E9CD70BFD31FC8F" bold="true" box="[675,687,836,855]" pageId="4" pageNumber="15">6</emphasis>
|
||||
) protarsal claw, male;
|
||||
<emphasis id="B975EAFA5D43152D2FB8D70BFC0DFC8F" bold="true" box="[903,915,836,855]" pageId="4" pageNumber="15">7</emphasis>
|
||||
) genital capsule, lateral view;
|
||||
<emphasis id="B975EAFA5D43152D2D4FD713FEE2FCB7" bold="true" box="[368,380,860,879]" pageId="4" pageNumber="15">8</emphasis>
|
||||
) paramera, distal view;
|
||||
<emphasis id="B975EAFA5D43152D2E55D713FDE8FCB7" bold="true" box="[618,630,860,879]" pageId="4" pageNumber="15">9</emphasis>
|
||||
) genital plates, female. Scale lines 5 1 mm, except Fig. 6 5 0.5 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BBE36E85D42152C2D0DD67EFEEFFCE0" blockId="5.[279,1113,561,1658]" pageId="5" pageNumber="16">
|
||||
<emphasis id="B975EAFA5D42152C2D0DD67EFE09FD9E" bold="true" box="[306,407,561,582]" pageId="5" pageNumber="16">
|
||||
<typeStatus id="54BA884A5D42152C2D0DD67EFE0CFD9E" box="[306,402,561,582]" pageId="5" pageNumber="16" type="allotype">Allotype</typeStatus>
|
||||
.
|
||||
</emphasis>
|
||||
Female. Similar to the male except as follows: antennae with lamellae of 8th to 10th segments subequal to the length of six preceeding segments combined (1:1). The setae located toward the apex of elytra longer than preceeding elytral setae. Anal plate convex, without concavity or furrow, punctate, with 60 scattered slender setae. Both apical spurs of metatibia shorter than in the male. Pygidium less convex. Ventral genital plates slightly sclerotized, nearly symmetrical, with a narrow lateral bridge toward each dorsal plate; dorsal plates with scattered setae at the apex and apparently fused laterally with ventral ones (
|
||||
<figureCitation id="133A2A6D5D42152C2E1CD747FDFBFCC5" box="[547,613,776,797]" captionStart="Figs" captionStartId="4.[330,373,836,855]" captionTargetBox="[340,1107,276,815]" captionTargetId="figure-12@4.[337,1108,272,816]" captionTargetPageId="4" captionText="Figs. 6–9. Phyllophaga tapantina. 6) protarsal claw, male; 7) genital capsule, lateral view; 8) paramera, distal view; 9) genital plates, female. Scale lines 5 1 mm, except Fig. 6 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900315" httpUri="https://zenodo.org/record/4900315/files/figure.png" pageId="5" pageNumber="16">Fig. 9</figureCitation>
|
||||
). Total body length:
|
||||
<quantity id="4CF99B0D5D42152C2F7AD747FC38FCC5" box="[837,934,776,797]" metricMagnitude="-2" metricUnit="m" metricValue="2.18" pageId="5" pageNumber="16" unit="mm" value="21.8">21.8 mm</quantity>
|
||||
. Humeral width: 9.0 mm.
|
||||
</paragraph>
|
||||
<paragraph id="8BBE36E85D42152C2D0DD772FE34FBD7" blockId="5.[279,1113,561,1658]" pageId="5" pageNumber="16">
|
||||
<emphasis id="B975EAFA5D42152C2D0DD772FD8FFC8A" bold="true" box="[306,529,829,850]" pageId="5" pageNumber="16">
|
||||
<typeStatus id="54BA884A5D42152C2D0DD772FE06FC8A" box="[306,408,829,850]" pageId="5" pageNumber="16" type="paratype">Paratype</typeStatus>
|
||||
Variation.
|
||||
</emphasis>
|
||||
Males. Similar to
|
||||
<typeStatus id="54BA884A5D42152C2EEAD772FCAEFC8A" box="[725,816,829,850]" pageId="5" pageNumber="16" type="holotype">holotype</typeStatus>
|
||||
except in total body length: 19.0–
|
||||
<quantity id="4CF99B0D5D42152C2D6CD717FE2AFCB5" box="[339,436,856,877]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" pageId="5" pageNumber="16" unit="mm" value="22.5">22.5 mm</quantity>
|
||||
, humeral width:
|
||||
<quantity id="4CF99B0D5D42152C2E57D717FD77FCB5" box="[616,745,856,877]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" metricValueMax="9.4" metricValueMin="8.2" pageId="5" pageNumber="16" unit="mm" value="8.8" valueMax="9.4" valueMin="8.2">8.2–9.4 mm</quantity>
|
||||
, body and legs in some specimens are shiny reddish brown, some specimens present scattered setae along the elytral suture, other specimens with antennal club 1.2–1.4× longer than length of preceeding six segments combined. Females similar to
|
||||
<typeStatus id="54BA884A5D42152C2FAAD7E6FC77FC66" box="[917,1001,937,958]" pageId="5" pageNumber="16" type="allotype">allotype</typeStatus>
|
||||
except as follows: total body length: 21.5–
|
||||
<quantity id="4CF99B0D5D42152C2E46D78BFD45FC01" box="[633,731,964,985]" metricMagnitude="-2" metricUnit="m" metricValue="1.89" pageId="5" pageNumber="16" unit="mm" value="18.9">18.9 mm</quantity>
|
||||
; humeral width: 8.0–
|
||||
<quantity id="4CF99B0D5D42152C2FFDD78BFB88FC01" box="[962,1046,964,985]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="5" pageNumber="16" unit="mm" value="9.1">9.1 mm</quantity>
|
||||
, body and legs shiny reddish brown, some specimens with more setae along the elytral suture.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D42152C2D0DD05AFC4EFA8A" pageId="5" pageNumber="16" type="materials_examined">
|
||||
<paragraph id="8BBE36E85D42152C2D0DD05AFCABFAC4" blockId="5.[279,1113,561,1658]" pageId="5" pageNumber="16">
|
||||
<emphasis id="B975EAFA5D42152C2D0DD05AFE22FBF2" bold="true" box="[306,444,1045,1066]" pageId="5" pageNumber="16">Type Series.</emphasis>
|
||||
<location id="8EDE60335D42152C2DFBD05AFDB0FBF2" LSID="urn:lsid:plazi:treatment:03A887FE5D44152F2D78D1DEFE52FA7A:8EDE60335D42152C2DFBD05AFDB0FBF2" box="[452,558,1045,1066]" name="Described" pageId="5" pageNumber="16">Described</location>
|
||||
from
|
||||
<specimenCount id="9D07FD615D42152C2E4FD05AFD4EFBF2" box="[624,720,1045,1066]" pageId="5" pageNumber="16" type="male">38 males</specimenCount>
|
||||
and
|
||||
<specimenCount id="9D07FD615D42152C2F3AD05AFCE2FBF2" box="[773,892,1045,1066]" pageId="5" pageNumber="16" type="female">23 females</specimenCount>
|
||||
.
|
||||
<materialsCitation id="3B693CB55D42152C2FB8D05AFDE7FBB8" collectingDate="1987-04-10" collectionCode="IN- BIO" collectorName="J. Sanchez." country="Costa Rica" elevation="1200" latitude="9.75" location="Parque Nacional Tapanti´" longLatPrecision="1299" longitude="-83.816666" municipality="Quebrada Segunda" pageId="5" pageNumber="16" specimenCount="1" specimenCount-male="1" stateProvince="Cartago" typeStatus="holotype">
|
||||
<typeStatus id="54BA884A5D42152C2FB8D05AFC76FBF2" box="[903,1000,1045,1066]" pageId="5" pageNumber="16" type="holotype">Holotype</typeStatus>
|
||||
<specimenCount id="9D07FD615D42152C2FCFD05AFBBDFBF2" box="[1008,1059,1045,1066]" pageId="5" pageNumber="16" type="male">male</specimenCount>
|
||||
(
|
||||
<collectionCode id="ED10AE2D5D42152C280DD05AFED7FB9D" pageId="5" pageNumber="16">INBIO</collectionCode>
|
||||
): ‘‘
|
||||
<collectingCountry id="F31676785D42152C2D59D07FFD92FB9D" box="[358,524,1072,1093]" name="Costa Rica" pageId="5" pageNumber="16">COSTA RICA</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49C5F80A5D42152C2E29D07FFDF5FB9D" box="[534,619,1072,1093]" country="Costa Rica" name="Cartago" pageId="5" pageNumber="16">Cartago</collectingRegion>
|
||||
,
|
||||
<location id="8EDE60335D42152C2E47D07FFCE1FB9D" LSID="urn:lsid:plazi:treatment:03A887FE5D44152F2D78D1DEFE52FA7A:8EDE60335D42152C2E47D07FFCE1FB9D" box="[632,895,1072,1093]" country="Costa Rica" county="Parque Nacional Tapanti'" municipality="Quebrada Segunda" name="Parque Nacional Tapanti´" pageId="5" pageNumber="16" stateProvince="Cartago">Parque Nacional Tapantı´</location>
|
||||
,
|
||||
<collectingMunicipality id="6BDAAC925D42152C2FB5D07FFBCAFB9D" box="[906,1108,1072,1093]" pageId="5" pageNumber="16">Quebrada Segunda</collectingMunicipality>
|
||||
,
|
||||
<quantity id="4CF99B0D5D42152C2D28D004FEEDFBB8" box="[279,371,1099,1120]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="16" unit="m" value="1200.0">
|
||||
<elevation id="002CD1DB5D42152C2D28D004FEEDFBB8" box="[279,371,1099,1120]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="16" unit="m" value="1200.0">1,200 m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<date id="FFBF10285D42152C2DBED004FE7CFBB8" box="[385,482,1099,1120]" pageId="5" pageNumber="16" value="1987-04-10">
|
||||
<collectingDate id="EFFBE9C05D42152C2DBED004FE7CFBB8" box="[385,482,1099,1120]" pageId="5" pageNumber="16" value="1987-04-10">10IV87</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F4533E5D42152C2DCED004FDF1FBB8" box="[497,623,1099,1120]" pageId="5" pageNumber="16">J. Sánchez.</collectorName>
|
||||
’’
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D42152C2EBCD004FD73FB4E" collectingDate="1995-12" collectionCode="INBIO, INBio" collectorName="G. Mora'" country="Costa Rica" county="Parque Nacional Tapanti'" elevation="1375" location="Parque Nacional Tapanti´" municipality="Sendero La Pava" pageId="5" pageNumber="16" specimenCount="1" specimenCount-female="1" stateProvince="Cartago" typeStatus="allotype">
|
||||
<typeStatus id="54BA884A5D42152C2EBCD004FD40FBB8" box="[643,734,1099,1120]" pageId="5" pageNumber="16" type="allotype">Allotype</typeStatus>
|
||||
<specimenCount id="9D07FD615D42152C2ED7D004FCB1FBB8" box="[744,815,1099,1120]" pageId="5" pageNumber="16" type="female">female</specimenCount>
|
||||
(
|
||||
<collectionCode id="ED10AE2D5D42152C2F7ED004FC0EFBB8" box="[833,912,1099,1120]" pageId="5" pageNumber="16">INBIO</collectionCode>
|
||||
): ‘‘
|
||||
<collectingCountry id="F31676785D42152C2F91D004FBCBFBB8" box="[942,1109,1099,1120]" name="Costa Rica" pageId="5" pageNumber="16">COSTA RICA</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49C5F80A5D42152C2D28D029FEF2FBA3" box="[279,364,1126,1147]" country="Costa Rica" name="Cartago" pageId="5" pageNumber="16">Cartago</collectingRegion>
|
||||
,
|
||||
<location id="8EDE60335D42152C2D42D029FD12FBA3" LSID="urn:lsid:plazi:treatment:03A887FE5D44152F2D78D1DEFE52FA7A:8EDE60335D42152C2D42D029FD12FBA3" box="[381,652,1126,1147]" country="Costa Rica" county="Parque Nacional Tapanti'" municipality="Sendero La Pava" name="Parque Nacional Tapanti´" pageId="5" pageNumber="16" stateProvince="Cartago">Parque Nacional Tapantı´</location>
|
||||
,
|
||||
<location id="8EDE60335D42152C2EA4D029FC18FBA3" LSID="urn:lsid:plazi:treatment:03A887FE5D44152F2D78D1DEFE52FA7A:8EDE60335D42152C2EA4D029FC18FBA3" box="[667,902,1126,1147]" country="Costa Rica" county="Parque Nacional Tapanti'" municipality="Sendero La Pava" name="Rio Grande de Orosi" pageId="5" pageNumber="16" stateProvince="Cartago">Río Grande de Orosi</location>
|
||||
,
|
||||
<collectingMunicipality id="6BDAAC925D42152C2FA9D029FBCDFBA3" box="[918,1107,1126,1147]" pageId="5" pageNumber="16">Sendero La Pava</collectingMunicipality>
|
||||
,
|
||||
<quantity id="4CF99B0D5D42152C2D28D0CEFE22FB4E" box="[279,444,1153,1174]" metricMagnitude="3" metricUnit="m" metricValue="1.375" metricValueMax="1.6" metricValueMin="1.15" pageId="5" pageNumber="16" unit="m" value="1375.0" valueMax="1600.0" valueMin="1150.0">
|
||||
<elevation id="002CD1DB5D42152C2D28D0CEFE22FB4E" box="[279,444,1153,1174]" metricMagnitude="3" metricUnit="m" metricValue="1.375" metricValueMax="1.6" metricValueMin="1.15" pageId="5" pageNumber="16" unit="m" value="1375.0" valueMax="1600.0" valueMin="1150.0">1,150–1,600 m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<date id="FFBF10285D42152C2DF6D0CEFD8CFB4E" box="[457,530,1153,1174]" pageId="5" pageNumber="16" value="1995-12">
|
||||
<collectingDate id="EFFBE9C05D42152C2DF6D0CEFD8CFB4E" box="[457,530,1153,1174]" pageId="5" pageNumber="16" value="1995-12">XII95</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F4533E5D42152C2E20D0CEFD18FB4E" box="[543,646,1153,1174]" pageId="5" pageNumber="16">G. Mora’</collectorName>
|
||||
’ (
|
||||
<collectionCode id="ED10AE2D5D42152C2EA4D0CEFD7CFB4E" box="[667,738,1153,1174]" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)" pageId="5" pageNumber="16">INBio</collectionCode>
|
||||
).
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D42152C2ECAD0CEFE0DFB69" collectingDate="1987-04-10" collectionCode="CASC, CNC, INBio, MXAL, TAMU, ZMHU" collectorName="J. Sanchez" country="Costa Rica" elevation="1200" latitude="9.75" location="Parque Nacional Tapanti'" longLatPrecision="1299" longitude="-83.816666" municipality="Quebrada Segunda" pageId="5" pageNumber="16" specimenCount="19" specimenCount-male="19" stateProvince="Cartago" typeStatus="paratype">
|
||||
<typeStatus id="54BA884A5D42152C2ECAD0CEFCC1FB4E" box="[757,863,1153,1174]" pageId="5" pageNumber="16" type="paratype">Paratypes</typeStatus>
|
||||
: same data as holotype (
|
||||
<specimenCount id="9D07FD615D42152C2D1FD0D3FE19FB69" box="[288,391,1180,1201]" count="19" pageId="5" pageNumber="16" type="male">19 males</specimenCount>
|
||||
);
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D42152C2DA0D0D3FE52FB14" collectingDate="1992-04" collectionCode="INBIO" collectorName="R. Vargas" country="Costa Rica" elevation="1250" latitude="9.75" location="Parque Nacional Tapanti'" longLatPrecision="1299" longitude="-83.816666" municipality="Quebrada Segunda" pageId="5" pageNumber="16" specimenCode="INBIO CR 1000 459238" specimenCount="1" specimenCount-male="1" stateProvince="Cartago" typeStatus="paratype">
|
||||
same data except:
|
||||
<date id="FFBF10285D42152C2E51D0D3FD30FB69" box="[622,686,1180,1201]" pageId="5" pageNumber="16" value="1992-04">
|
||||
<collectingDate id="EFFBE9C05D42152C2E51D0D3FD30FB69" box="[622,686,1180,1201]" pageId="5" pageNumber="16" value="1992-04">IV92</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<quantity id="4CF99B0D5D42152C2E82D0D3FC84FB69" box="[701,794,1180,1201]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="5" pageNumber="16" unit="m" value="1250.0">
|
||||
<elevation id="002CD1DB5D42152C2E82D0D3FC84FB69" box="[701,794,1180,1201]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="5" pageNumber="16" unit="m" value="1250.0">1,250 m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<collectorName id="26F4533E5D42152C2F15D0D3FC0BFB69" box="[810,917,1180,1201]" pageId="5" pageNumber="16">R. Vargas</collectorName>
|
||||
<specimenCode id="DBA79E935D42152C2F9ED0D3FEFBFB14" pageId="5" pageNumber="16">INBIO CR 1000 459238</specimenCode>
|
||||
(
|
||||
<specimenCount id="9D07FD615D42152C2D48D0F8FE5FFB14" box="[375,449,1207,1228]" count="1" pageId="5" pageNumber="16" type="male">1 male</specimenCount>
|
||||
);
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D42152C2DECD0F8FC09FB14" box="[467,919,1207,1228]" collectingDate="1995-12" collectionCode="INBIO, INBio" collectorName="G. Mora'" country="Costa Rica" elevation="1200" location="Parque Nacional Tapanti'" municipality="Quebrada Segunda" pageId="5" pageNumber="16" specimenCount="16" specimenCount-female="9" specimenCount-male="7" stateProvince="Cartago" typeStatus="paratype">
|
||||
same data as allotype (
|
||||
<specimenCount id="9D07FD615D42152C2EFCD0F8FC89FB14" box="[707,791,1207,1228]" count="7" pageId="5" pageNumber="16" type="male">7 males</specimenCount>
|
||||
,
|
||||
<specimenCount id="9D07FD615D42152C2F1DD0F8FC12FB14" box="[802,908,1207,1228]" count="9" pageId="5" pageNumber="16" type="female">9 females</specimenCount>
|
||||
);
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D42152C2FA1D0F8FD71FB3E" collectingDate="1991-10" collectingDateMax="1996-06" collectingDateMin="1991-10" collectionCode="INBIO" collectorName="A. Solis'" country="Costa Rica" county="Parque Nacional Tapanti'" elevation="1200" location="Rio San Lorencito" municipality="San Ramon" pageId="5" pageNumber="16" specimenCode="INBIO CR 1002 443314" specimenCount="1" specimenCount-female="1" stateProvince="Alajuela" typeStatus="paratype">
|
||||
same data except:
|
||||
<date id="FFBF10285D42152C2D28D09EFEC6FB3E" box="[279,344,1233,1254]" pageId="5" pageNumber="16" value="1996-06">
|
||||
<collectingDate id="EFFBE9C05D42152C2D28D09EFEC6FB3E" box="[279,344,1233,1254]" pageId="5" pageNumber="16" value="1996-06">VI96</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<specimenCode id="DBA79E935D42152C2D58D09EFDEAFB3E" box="[359,628,1233,1254]" pageId="5" pageNumber="16">INBIO CR 1002 443314</specimenCode>
|
||||
(
|
||||
<specimenCount id="9D07FD615D42152C2EB6D09EFD76FB3E" box="[649,744,1233,1254]" count="1" pageId="5" pageNumber="16" type="female">1 female</specimenCount>
|
||||
)
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D42152C2EC6D09EFCC8FAD9" collectingDate="1991-10" collectionCode="INBIO" collectorName="A. Solis'" country="Costa Rica" elevation="1200" location="Rio San Lorencito" municipality="San Ramon" pageId="5" pageNumber="16" specimenCode="INBIO CR 1002 443314" specimenCount="22" specimenCount-female="12" specimenCount-male="10" stateProvince="Alajuela" typeStatus="paratype">
|
||||
‘‘
|
||||
<collectingRegion id="49C5F80A5D42152C2F3CD09EFCFAFB3E" box="[771,868,1233,1254]" country="Costa Rica" name="Alajuela" pageId="5" pageNumber="16">Alajuela</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="6BDAAC925D42152C2F4CD09EFC6CFB3E" box="[883,1010,1233,1254]" pageId="5" pageNumber="16">San Ramón</collectingMunicipality>
|
||||
,
|
||||
<location id="8EDE60335D42152C283FD09EFEE1FAD9" LSID="urn:lsid:plazi:treatment:03A887FE5D44152F2D78D1DEFE52FA7A:8EDE60335D42152C283FD09EFEE1FAD9" country="Costa Rica" county="Parque Nacional Tapanti'" municipality="San Ramon" name="Rio San Lorencito" pageId="5" pageNumber="16" stateProvince="Alajuela">Río San Lorencito</location>
|
||||
,
|
||||
<date id="FFBF10285D42152C2DAFD0A3FE56FAD9" box="[400,456,1260,1281]" pageId="5" pageNumber="16" value="1991-10">
|
||||
<collectingDate id="EFFBE9C05D42152C2DAFD0A3FE56FAD9" box="[400,456,1260,1281]" pageId="5" pageNumber="16" value="1991-10">X91</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F4533E5D42152C2DE6D0A3FDA0FAD9" box="[473,574,1260,1281]" pageId="5" pageNumber="16">A. Solís’</collectorName>
|
||||
’ (
|
||||
<specimenCount id="9D07FD615D42152C2E68D0A3FD20FAD9" box="[599,702,1260,1281]" count="10" pageId="5" pageNumber="16" type="male">10 males</specimenCount>
|
||||
,
|
||||
<specimenCount id="9D07FD615D42152C2EF1D0A3FCD4FAD9" box="[718,842,1260,1281]" count="12" pageId="5" pageNumber="16" type="female">12 females</specimenCount>
|
||||
).
|
||||
</materialsCitation>
|
||||
Paratypes deposited in
|
||||
<collectionCode id="ED10AE2D5D42152C2D28D148FEC0FAC4" box="[279,350,1287,1308]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="5" pageNumber="16">CASC</collectionCode>
|
||||
,
|
||||
<collectionCode id="ED10AE2D5D42152C2D54D148FE3DFAC4" box="[363,419,1287,1308]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" pageId="5" pageNumber="16">CNC</collectionCode>
|
||||
,
|
||||
<collectionCode id="ED10AE2D5D42152C2D8FD148FE6AFAC4" box="[432,500,1287,1308]" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)" pageId="5" pageNumber="16">INBio</collectionCode>
|
||||
,
|
||||
<collectionCode id="ED10AE2D5D42152C2E3ED148FDCFFAC4" box="[513,593,1287,1308]" pageId="5" pageNumber="16">MXAL</collectionCode>
|
||||
,
|
||||
<collectionCode id="ED10AE2D5D42152C2E62D148FD37FAC4" box="[605,681,1287,1308]" collectionName="USA, Texas, College Station, Texas A & M University" pageId="5" pageNumber="16">TAMU</collectionCode>
|
||||
and
|
||||
<collectionCode id="ED10AE2D5D42152C2EDED148FCAFFAC4" box="[737,817,1287,1308]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="5" pageNumber="16">ZMHU</collectionCode>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BBE36E85D42152C2D0DD16DFC4EFA8A" blockId="5.[279,1113,561,1658]" pageId="5" pageNumber="16">
|
||||
<emphasis id="B975EAFA5D42152C2D0DD16DFE4BFAEF" bold="true" box="[306,469,1314,1335]" pageId="5" pageNumber="16">Type Locality.</emphasis>
|
||||
<location id="8EDE60335D42152C2DE1D16DFD78FAEF" LSID="urn:lsid:plazi:treatment:03A887FE5D44152F2D78D1DEFE52FA7A:8EDE60335D42152C2DE1D16DFD78FAEF" box="[478,742,1314,1335]" country="Costa Rica" latitude="9.75" longLatPrecision="1299" longitude="-83.816666" municipality="Cordillera de Talamanca" name="Parque Nacional Tapanti" pageId="5" pageNumber="16">Parque Nacional Tapanti</location>
|
||||
,
|
||||
<location id="8EDE60335D42152C2ECCD16DFBCAFAEF" LSID="urn:lsid:plazi:treatment:03A887FE5D44152F2D78D1DEFE52FA7A:8EDE60335D42152C2ECCD16DFBCAFAEF" box="[755,1108,1314,1335]" country="Costa Rica" latitude="9.75" longLatPrecision="1299" longitude="-83.816666" municipality="Cordillera de Talamanca" name="Cuenca del Rio Grande de Orosi" pageId="5" pageNumber="16">Cuenca del Río Grande de Orosi</location>
|
||||
,
|
||||
<collectingMunicipality id="6BDAAC925D42152C2D28D172FD81FA8A" box="[279,543,1341,1362]" pageId="5" pageNumber="16">Cordillera de Talamanca</collectingMunicipality>
|
||||
,
|
||||
<collectingCountry id="F31676785D42152C2E13D172FD3EFA8A" box="[556,672,1341,1362]" name="Costa Rica" pageId="5" pageNumber="16">Costa Rica</collectingCountry>
|
||||
(approx.
|
||||
<geoCoordinate id="EE35502F5D42152C2F35D172FCC9FA8A" box="[778,855,1341,1362]" degrees="9" direction="north" minutes="45" orientation="latitude" pageId="5" pageNumber="16" precision="925" value="9.75">9°45'N</geoCoordinate>
|
||||
;
|
||||
<geoCoordinate id="EE35502F5D42152C2F5BD172FC5BFA8A" box="[868,965,1341,1362]" degrees="83" direction="west" minutes="49" orientation="longitude" pageId="5" pageNumber="16" precision="925" value="-83.816666">83°49'W</geoCoordinate>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D42152C2D0DD117FD72F9A2" pageId="5" pageNumber="16" type="biology_ecology">
|
||||
<paragraph id="8BBE36E85D42152C2D0DD117FD72F9A2" blockId="5.[279,1113,561,1658]" pageId="5" pageNumber="16">
|
||||
<emphasis id="B975EAFA5D42152C2D0DD117FE79FAB5" bold="true" box="[306,487,1368,1389]" pageId="5" pageNumber="16">Biological Data.</emphasis>
|
||||
Males and females of
|
||||
<taxonomicName id="4C014D6B5D42152C2ED8D117FCF2FAB5" box="[743,876,1368,1389]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="16" phylum="Arthropoda" rank="species" species="tapantina" status="sp. nov.">
|
||||
<emphasis id="B975EAFA5D42152C2ED8D117FCF2FAB5" box="[743,876,1368,1389]" italics="true" pageId="5" pageNumber="16">P. tapantina</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A24657815D42152C2F49D117FC2CFAB5" box="[886,946,1368,1389]" pageId="5" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
|
||||
were collected at lights between
|
||||
<quantity id="4CF99B0D5D42152C2DE5D13CFD81FA50" box="[474,543,1395,1416]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="16" unit="m" value="800.0">800 m</quantity>
|
||||
altitude on Río San Lorencito and
|
||||
<quantity id="4CF99B0D5D42152C2F99D13CFC61FA50" box="[934,1023,1395,1416]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="16" unit="m" value="1500.0">1,500 m</quantity>
|
||||
altitude at shore of Río Grande de Orosi. These localities are placed along the Atlantic slopes from eastern Cordillera de Tilarán to western Cordillera de Talamanca, and present cloud forests and subtropical evergreen forests with species of
|
||||
<emphasis id="B975EAFA5D42152C2D28D190FE2DF9D7" italics="true" pageId="5" pageNumber="16">
|
||||
<taxonomicName id="4C014D6B5D42152C2D28D190FB82FA2C" authority=", Alnus, Nectandra, Ocotea, Podocarpus, Didymopanax, Ulmus" authorityName="Alnus, Nectandra, Ocotea, Podocarpus, Didymopanax, Ulmus" box="[279,1052,1503,1524]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="16" phylum="Tracheophyta" rank="genus">Quercus, Alnus, Nectandra, Ocotea, Podocarpus, Didymopanax, Ulmus</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C014D6B5D42152C2816D190FE2DF9D7" authority=", Vismia" authorityName="Vismia" class="Magnoliopsida" family="Fabaceae" genus="Erythrina" kingdom="Animalia" order="Fabales" pageId="5" pageNumber="16" phylum="Tracheophyta" rank="genus">Erythrina, Vismia</taxonomicName>
|
||||
</emphasis>
|
||||
and
|
||||
<taxonomicName id="4C014D6B5D42152C2DD0D1B5FD7BF9D7" authority="(Boza 1988)" baseAuthorityName="Boza" baseAuthorityYear="1988" box="[495,741,1530,1551]" class="Magnoliopsida" family="Siparunaceae" genus="Siparuna" kingdom="Plantae" order="Laurales" pageId="5" pageNumber="16" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="B975EAFA5D42152C2DD0D1B5FDD1F9D7" box="[495,591,1530,1551]" italics="true" pageId="5" pageNumber="16">Siparuna</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF904B195D42152C2E5DD1B5FD43F9D7" author="Boza" box="[610,733,1530,1551]" firstAuthor="Boza" pageId="5" pageNumber="16" refId="ref11168" refString="Boza, M. 1988. Costa Rica National Parks. Heliconia. San Jose', Costa Rica. 323 pp." type="book" year="1988">Boza 1988</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
. Phenology: April (25), June (1), October (22), December (16). Other species of
|
||||
<taxonomicName id="4C014D6B5D42152C2F31D25AFC0AF9F2" box="[782,916,1557,1578]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B975EAFA5D42152C2F31D25AFC0AF9F2" box="[782,916,1557,1578]" italics="true" pageId="5" pageNumber="16">Phyllophaga</emphasis>
|
||||
</taxonomicName>
|
||||
flying at the same time and places were:
|
||||
<emphasis id="B975EAFA5D42152C2E33D27FFD27F99D" box="[524,697,1584,1605]" italics="true" pageId="5" pageNumber="16">P. nevermannea</emphasis>
|
||||
Saylor,
|
||||
<emphasis id="B975EAFA5D42152C2F2BD27FFC31F99D" box="[788,943,1584,1605]" italics="true" pageId="5" pageNumber="16">P. nevermanni</emphasis>
|
||||
Saylor,
|
||||
<emphasis id="B975EAFA5D42152C2834D27FFEDDF9B8" italics="true" pageId="5" pageNumber="16">P. orosina</emphasis>
|
||||
Moser,
|
||||
<emphasis id="B975EAFA5D42152C2DA3D204FD83F9B8" box="[412,541,1611,1632]" italics="true" pageId="5" pageNumber="16">P. lissopyge</emphasis>
|
||||
Bates,
|
||||
<taxonomicName id="4C014D6B5D42152C2E4FD204FC89F9B8" box="[624,791,1611,1632]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="16" phylum="Arthropoda" rank="species" species="talamancana" status="sp. nov.">
|
||||
<emphasis id="B975EAFA5D42152C2E4FD204FC89F9B8" box="[624,791,1611,1632]" italics="true" pageId="5" pageNumber="16">P. talamancana</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A24657815D42152C2F1ED204FCC3F9B8" box="[801,861,1611,1632]" pageId="5" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
|
||||
,
|
||||
<taxonomicName id="4C014D6B5D42152C2F54D204FC41F9B8" box="[875,991,1611,1632]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="16" phylum="Arthropoda" rank="species" species="tilarana" status="sp. nov.">
|
||||
<emphasis id="B975EAFA5D42152C2F54D204FC41F9B8" box="[875,991,1611,1632]" italics="true" pageId="5" pageNumber="16">P. tilarana</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A24657815D42152C2FD7D204FBBAF9B8" box="[1000,1060,1611,1632]" pageId="5" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
|
||||
, and another four new species under description.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF7E66605D41152F2D75D013FDECFB78" ID-DOI="http://doi.org/10.5281/zenodo.4900317" ID-Zenodo-Dep="4900317" httpUri="https://zenodo.org/record/4900317/files/figure.png" pageId="6" pageNumber="17" startId="6.[330,373,1116,1135]" targetBox="[336,1117,273,1093]" targetPageId="6">
|
||||
<paragraph id="8BBE36E85D41152F2D75D013FDECFB78" blockId="6.[307,1139,1116,1184]" pageId="6" pageNumber="17">
|
||||
<emphasis id="B975EAFA5D41152F2D75D013FE5CFBB7" bold="true" box="[330,450,1116,1135]" pageId="6" pageNumber="17">Figs. 10–13.</emphasis>
|
||||
<emphasis id="B975EAFA5D41152F2DE6D012FD2AFBA8" box="[473,692,1117,1136]" italics="true" pageId="6" pageNumber="17">
|
||||
<taxonomicName id="4C014D6B5D41152F2DE6D012FD2FFBA8" box="[473,689,1117,1136]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="lorencita">Phyllophaga lorencita</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
<emphasis id="B975EAFA5D41152F2E82D013FD4AFBB7" bold="true" box="[701,724,1116,1135]" pageId="6" pageNumber="17">10</emphasis>
|
||||
) protarsal claw, male;
|
||||
<emphasis id="B975EAFA5D41152F2F88D013FC50FBB7" bold="true" box="[951,974,1116,1135]" pageId="6" pageNumber="17">11</emphasis>
|
||||
) genital capsule, lateral view;
|
||||
<emphasis id="B975EAFA5D41152F2D8ED03BFE56FB5F" bold="true" box="[433,456,1140,1159]" pageId="6" pageNumber="17">12</emphasis>
|
||||
) paramera, ventrodistal view;
|
||||
<emphasis id="B975EAFA5D41152F2ECAD03BFC92FB5F" bold="true" box="[757,780,1140,1159]" pageId="6" pageNumber="17">13</emphasis>
|
||||
) genital plates, female. Scale lines 5 1 mm, except Fig. 10 5 0.5 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="C31B65635D41152F2D72D0A2FD50FAB5" pageId="6" pageNumber="17" type="discussion">
|
||||
<paragraph id="8BBE36E85D41152F2D72D0A2FD50FAB5" blockId="6.[307,1139,1261,1442]" pageId="6" pageNumber="17">
|
||||
<emphasis id="B975EAFA5D41152F2D72D0A2FE27FADA" bold="true" box="[333,441,1261,1282]" pageId="6" pageNumber="17">Remarks.</emphasis>
|
||||
Is not possible to include
|
||||
<taxonomicName id="4C014D6B5D41152F2EE0D0A1FBEDFADB" box="[735,1139,1261,1283]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="tapantina" status="sp. nov." subGenus="Phyllophaga">
|
||||
<emphasis id="B975EAFA5D41152F2EE0D0A1FCFBFADB" box="[735,869,1262,1283]" italics="true" pageId="6" pageNumber="17">Phyllophaga</emphasis>
|
||||
(
|
||||
<emphasis id="B975EAFA5D41152F2F48D0A1FC62FADB" box="[887,1020,1262,1283]" italics="true" pageId="6" pageNumber="17">Phyllophaga</emphasis>
|
||||
)
|
||||
<emphasis id="B975EAFA5D41152F2830D0A1FBEDFADB" box="[1039,1139,1262,1283]" italics="true" pageId="6" pageNumber="17">tapantina</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A24657815D41152F2D0CD147FEF3FAC5" box="[307,365,1288,1309]" pageId="6" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
|
||||
in any known species group (
|
||||
<emphasis id="B975EAFA5D41152F2E8FD147FD74FAC5" box="[688,746,1288,1309]" italics="true" pageId="6" pageNumber="17">sensu</emphasis>
|
||||
<bibRefCitation id="EF904B195D41152F2ECCD147FCE4FAC5" author="Moron" box="[755,890,1288,1309]" firstAuthor="Moron" pageId="6" pageNumber="17" refId="ref11224" refString="Moron, M. A. 1986. El genero Phyllophaga en Mexico. Morfologia, distribucion y sistematica supraespecifica (Insecta: Coleoptera). Publ. 20, Instituto de Ecologia, Mexico, D. F. 341 pp." type="book" year="1986">Morón 1986</bibRefCitation>
|
||||
). The greenish vitreous luster on the elytra, the shortened tarsomeres in both sexes, the spines on protarsomeres and the shape and details of the male genital capsule will aid in the recognition of this new species.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D41152F2D72D13DFE52FA7A" pageId="6" pageNumber="17" type="etymology">
|
||||
<paragraph id="8BBE36E85D41152F2D72D13DFE52FA7A" blockId="6.[307,1139,1261,1442]" pageId="6" pageNumber="17">
|
||||
<emphasis id="B975EAFA5D41152F2D72D13DFE52FA5F" bold="true" box="[333,460,1394,1415]" pageId="6" pageNumber="17">Etymology.</emphasis>
|
||||
Derived from Tapantí National Park in the western Cordillera de Talamanca.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
359
data/03/A8/87/03A887FE5D46152A2D70D613FCBCFAA8.xml
Normal file
359
data/03/A8/87/03A887FE5D46152A2D70D613FCBCFAA8.xml
Normal file
|
|
@ -0,0 +1,359 @@
|
|||
<document id="4E88657C3BD4F5A6E94F8405989D4546" ID-DOI="10.1649/0010-065x(2001)055[0011:snsops]2.0.co;2" ID-ISSN="1938-4394" ID-Zenodo-Dep="4900311" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1622827801739" checkinUser="carolina" docAuthor="Morón, Miguel Angel & Solís, Angel" docDate="2001" docId="03A887FE5D46152A2D70D613FCBCFAA8" docLanguage="en" docName="TheColeopBull.55.1.11-29.pdf.imf" docOrigin="The Coleopterists Bulletin 55 (1)" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Phyllophaga (Phyllophaga) tilarana Moron and Solis, new species" docType="treatment" docVersion="1" lastPageNumber="14" masterDocId="FF91FF865D4715292C3FD44FFF9EFFD8" masterDocTitle="Seven New Species Of Phyllophaga (S. Str.) Harris From Costa Rica (Coleoptera: Melolonthidae: Melolonthinae)" masterLastPageNumber="29" masterPageNumber="11" pageNumber="12" updateTime="1730139982923" updateUser="juliana" zenodo-license-document="CLOSED">
|
||||
<mods:mods id="6D302F1C3989876DC3ABAF66EF67F63A" xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo id="15DF41BCC8696E8DF2E2392ECF90CC5A">
|
||||
<mods:title id="5766B8A1B8508BC64D10BD2E0D3B4E18">Seven New Species Of Phyllophaga (S. Str.) Harris From Costa Rica (Coleoptera: Melolonthidae: Melolonthinae)</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name id="B5126C5A62E4A59C1965F98BC63290F2" type="personal">
|
||||
<mods:role id="89CE43A088B1A51B40C7EFD86AEB2255">
|
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<mods:roleTerm id="2FEA114845FC2A5B768CCD78DF8463A2">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="EF4D3E3DBF2C64A16DC8886A383FF04A">Morón, Miguel Angel</mods:namePart>
|
||||
<mods:affiliation id="63219CD585E1022F92CFDFDECA23D140">Departamento de Entomología Instituto de Ecología, A. C. (sector SEPCONACYT) Apartado Postal 63 Xalapa, Veracruz 91000, MÉXICO</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name id="5F4568EE010BBC074BD23A31E5DDB51E" type="personal">
|
||||
<mods:role id="5D07A6A353AF7A4776CEAC3E97F020C3">
|
||||
<mods:roleTerm id="1DFCC79F57194DE1B629501736DE904D">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="82EEBD3A3EB184C520AEF3965CD4CCAA">Solís, Angel</mods:namePart>
|
||||
<mods:affiliation id="8C3F63FE64B2EDD614D0115C27FD4E2F">Departamento de Entomología Instituto Nacional de Biodiversidad (INBio) Apartado Postal 22 3100 Santo Domingo de Heredia, COSTA RICA</mods:affiliation>
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||||
</mods:name>
|
||||
<mods:typeOfResource id="183D83439079FA2F4261F92A419660E5">text</mods:typeOfResource>
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<mods:titleInfo id="FC41CF6620D258226DEBC83537C5E643">
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<mods:title id="D955A6E86977F385730D00BAE7B22300">The Coleopterists Bulletin</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part id="B3ABBFCF3E78D6501497837BBBB54461">
|
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<mods:date id="ADC2D60D9C51484AA36E00F5628035E7">2001</mods:date>
|
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<mods:detail id="12C2F24F89B7EAB6FE1FFFC251904326" type="pubDate">
|
||||
<mods:number id="9696209F7A7DE7FA51476603CB40A099">2001-03-01</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="1D515229DDCA96221CC82A2AAA8014B4" type="volume">
|
||||
<mods:number id="BF9A4D314B89BC268619FC508B9CDC38">55</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="F13D2C7FA62011F4E379FB6F9EE290ED" type="issue">
|
||||
<mods:number id="861215B74ED3C39B76DAD42B8F07CE41">1</mods:number>
|
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|
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|
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|
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<mods:classification id="D3648225D51008D92DE58F40D0B8F131">journal article</mods:classification>
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|
||||
<mods:identifier id="6BA3B20594995D2C39F48C87ABB9D7BA" type="ISSN">1938-4394</mods:identifier>
|
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<mods:identifier id="F3D0A4E64C1F42182DE7F288C6096DA1" type="Zenodo-Dep">4900311</mods:identifier>
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</mods:mods>
|
||||
<treatment id="03A887FE5D46152A2D70D613FCBCFAA8" LSID="urn:lsid:plazi:treatment:03A887FE5D46152A2D70D613FCBCFAA8" httpUri="http://treatment.plazi.org/id/03A887FE5D46152A2D70D613FCBCFAA8" lastPageId="3" lastPageNumber="14" pageId="1" pageNumber="12">
|
||||
<subSubSection id="C31B65635D4615282D70D613FBBEFDA8" box="[335,1056,603,625]" pageId="1" pageNumber="12" type="nomenclature">
|
||||
<paragraph id="8BBE36E85D4615282D70D613FBBEFDA8" blockId="1.[335,1056,603,651]" box="[335,1056,603,625]" pageId="1" pageNumber="12">
|
||||
<taxonomicName id="4C014D6B5D4615282D70D613FC0EFDA8" authority="Moron and Solis" authorityName="Moron and Solis" box="[335,912,603,625]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="12" phylum="Arthropoda" rank="species" species="tilarana" status="sp. nov." subGenus="Phyllophaga">
|
||||
<emphasis id="B975EAFA5D4615282D70D613FE4BFDA9" box="[335,469,604,625]" italics="true" pageId="1" pageNumber="12">Phyllophaga</emphasis>
|
||||
(
|
||||
<emphasis id="B975EAFA5D4615282DD8D613FDF2FDA9" box="[487,620,604,625]" italics="true" pageId="1" pageNumber="12">Phyllophaga</emphasis>
|
||||
)
|
||||
<emphasis id="B975EAFA5D4615282E42D613FD4FFDA9" box="[637,721,604,625]" italics="true" pageId="1" pageNumber="12">tilarana</emphasis>
|
||||
Morón and Solís
|
||||
</taxonomicName>
|
||||
,
|
||||
<emphasis id="B975EAFA5D4615282FA2D614FBBEFDA8" bold="true" box="[925,1056,603,624]" pageId="1" pageNumber="12">
|
||||
<taxonomicNameLabel id="A24657815D4615282FA2D614FBBEFDA8" box="[925,1056,603,624]" pageId="1" pageNumber="12" rank="species">new species</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D46152A2EB9D639FDFFFDB2" lastPageId="3" lastPageNumber="14" pageId="1" pageNumber="12" type="description">
|
||||
<paragraph id="8BBE36E85D4615282EB9D639FD77FD53" blockId="1.[335,1056,603,651]" box="[646,745,630,651]" pageId="1" pageNumber="12">
|
||||
<figureCitation id="133A2A6D5D4615282EB9D639FD77FD53" box="[646,745,630,651]" captionStart="Figs" captionStartId="2.[330,373,1228,1247]" captionTargetBox="[309,1134,273,1204]" captionTargetId="figure-12@2.[308,1137,272,1208]" captionTargetPageId="2" captionText="Figs. 1–5. Phyllophaga tilarana. 1) protarsal claw, male; 2) genital capsule, lateral view; 3) paramera, distal view; 4) protarsal claw, female; 5) genital plates, female. Scale lines 5 1 mm, except Figs. 1, 4 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900313" httpUri="https://zenodo.org/record/4900313/files/figure.png" pageId="1" pageNumber="12">Figs. 1–5</figureCitation>
|
||||
</paragraph>
|
||||
<paragraph id="8BBE36E85D46152B2D10D6E8FBEAFA40" blockId="1.[279,1113,679,1658]" lastBlockId="2.[307,1140,1389,1432]" lastPageId="2" lastPageNumber="13" pageId="1" pageNumber="12">
|
||||
<emphasis id="B975EAFA5D4615282D10D6E8FE11FD62" bold="true" box="[303,399,679,698]" pageId="1" pageNumber="12">
|
||||
<typeStatus id="54BA884A5D4615282D10D6E8FE15FD62" box="[303,395,679,698]" pageId="1" pageNumber="12" type="holotype">Holotype</typeStatus>
|
||||
.
|
||||
</emphasis>
|
||||
Male. Clypeus, frons and pronotum shiny dark reddish brown; elytra silky, light brown, without macroscopic vestiture; mouthparts, sterna, pygidium and legs shiny dark reddish brown. Clypeus 3.2× wider than long, anterior border scarcely sinuate, with elevated margin, surface slightly irregular, with many uniformly distributed, deep, round shaped punctures and some scattered short setae. Frontoclypeal suture sinuate and deeply impressed. Frons 1.6× wider than long, convex, irregularly and shallowly punctate, with short slender setae at sides and on disk. Antenna 10segmented, with 3segmented club, lamellae of 8th to 10th segments 2.0× longer than length of preceeding 6 segments combined. Frons 3.2× wider than dorsal diameter of eye. Eye canthus long and narrow, with 16–17 setae. Labrum bilobed, deeply sinuate, with scattered long slender setae. Mentum slightly concave, impunctate, with scarce lateral setae, anterior border briefly sinuate. Pronotum 1.7× wider than long and 3.0× wider than frons. Pronotal disk shiny, with round, shallow punctures regularly separated from one another by 1–2 diameters; lateral borders widely angulated, lateral marginal bead nearly entire at anterior half and slightly crenulate at basal half, with scattered, long, slender setae; basal bead strongly indicated, except in front of scutellum where it is indicated only by punctures; anterior angles obtuse, not prominent; posterior angles obtuse, not prominent. Scutellum 1.6× wider than long, with some minute punctures. Elytron 2.9× longer than wide, silkshiny, densely rugopunctate; epipleural border very narrow, extended along complete margin, provided with scattered, short, slender setae; humeral callus rounded, prominent; apical callus rounded. Metathoracic wings completely developed. Propygidium shiny, with scattered, short setae, and dense punctuation. Pygidium very convex, shiny, with round, shallow punctures regularly distributed, with erect setae, shortened toward the basal border; apical margin with 22 long, slender setae; basal margin effaced medially. Pterosterna with long, dense, yellowish setae. Visible abdominal sternites II and IV slightly depressed, with numerous short setae near the middline; sternite V with a transverse, granulated, setose ridge, and scattered setae toward the sides; anal plate large, with wide transverse concavity, with numerous erect setae, posterior border thickened with a transverse row of long setae. Protibia shorter than protarsus (1:1.7), with external border tridentate, proximal tooth much shortened, preapical spur nearly straight, apex rounded, half as long as 2th protarsomere. Mesotibia with one oblique, sharp, setiferous carina on external side; upper apical spur straight, narrow, and 1.3× longer than lower spur. Metatibia one fifth shorter than metatarsus, with one oblique, sharp, setiferous carina on external side; upper apical spur articulated, recurved, apex rounded, slightly longer than basal metatarsomere, and 1.2× longer than lower spur; lower apical spur articulated with tibial border, with rounded apex. Tarsomeres semicylindrical, elongate, with enlarged apices, with some setae apically and two dense lines of long, thick setae on ventral side. Tarsal claws symmetrical, similar on all legs, with an acute short tooth located close to basal dilation (
|
||||
<figureCitation id="133A2A6D5D4615282D9ED278FE46F992" box="[417,472,1591,1610]" captionStart="Figs" captionStartId="2.[330,373,1228,1247]" captionTargetBox="[309,1134,273,1204]" captionTargetId="figure-12@2.[308,1137,272,1208]" captionTargetPageId="2" captionText="Figs. 1–5. Phyllophaga tilarana. 1) protarsal claw, male; 2) genital capsule, lateral view; 3) paramera, distal view; 4) protarsal claw, female; 5) genital plates, female. Scale lines 5 1 mm, except Figs. 1, 4 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900313" httpUri="https://zenodo.org/record/4900313/files/figure.png" pageId="1" pageNumber="12">Fig. 1</figureCitation>
|
||||
). Genital capsule with large parameres, dorsally and ventrally fused, symmetrical, pentagonal shaped, apex of each elongated, acute, with toothlike preapical projections. Aedeagus narrowed, with scarce sclerotized preapical structures (
|
||||
<figureCitation id="133A2A6D5D4615282FCCD228FBD3F9A2" box="[1011,1101,1639,1658]" captionStart="Figs" captionStartId="2.[330,373,1228,1247]" captionTargetBox="[309,1134,273,1204]" captionTargetId="figure-12@2.[308,1137,272,1208]" captionTargetPageId="2" captionText="Figs. 1–5. Phyllophaga tilarana. 1) protarsal claw, male; 2) genital capsule, lateral view; 3) paramera, distal view; 4) protarsal claw, female; 5) genital plates, female. Scale lines 5 1 mm, except Figs. 1, 4 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900313" httpUri="https://zenodo.org/record/4900313/files/figure.png" pageId="1" pageNumber="12">Figs. 2–3</figureCitation>
|
||||
). Tectum shortened, irregularly convex. Length of genital capsule from apex of parameres to border of basal piece:
|
||||
<quantity id="4CF99B0D5D45152B2E1FD1CAFDF2FA40" box="[544,620,1413,1432]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="2" pageNumber="13" unit="mm" value="5.1">5.1 mm</quantity>
|
||||
. Total body length: 23.0 mm. Humeral width:
|
||||
<quantity id="4CF99B0D5D45152B281AD1CAFBEEFA40" box="[1061,1136,1413,1432]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="2" pageNumber="13" unit="mm" value="9.3">9.3 mm</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
<caption id="DF7E66605D45152B2D75D083FD48FAC8" ID-DOI="http://doi.org/10.5281/zenodo.4900313" ID-Zenodo-Dep="4900313" httpUri="https://zenodo.org/record/4900313/files/figure.png" pageId="2" pageNumber="13" startId="2.[330,373,1228,1247]" targetBox="[309,1134,273,1204]" targetPageId="2">
|
||||
<paragraph id="8BBE36E85D45152B2D75D083FD48FAC8" blockId="2.[307,1139,1228,1296]" pageId="2" pageNumber="13">
|
||||
<emphasis id="B975EAFA5D45152B2D75D083FE34FB07" bold="true" box="[330,426,1228,1247]" pageId="2" pageNumber="13">Figs. 1–5.</emphasis>
|
||||
<emphasis id="B975EAFA5D45152B2DFED082FD0CFB38" box="[449,658,1229,1248]" italics="true" pageId="2" pageNumber="13">
|
||||
<taxonomicName id="4C014D6B5D45152B2DFED082FD10FB38" box="[449,654,1229,1248]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="species" species="tilarana">Phyllophaga tilarana</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
<emphasis id="B975EAFA5D45152B2EA6D083FD3BFB07" bold="true" box="[665,677,1228,1247]" pageId="2" pageNumber="13">1</emphasis>
|
||||
) protarsal claw, male;
|
||||
<emphasis id="B975EAFA5D45152B2FBCD083FC11FB07" bold="true" box="[899,911,1228,1247]" pageId="2" pageNumber="13">2</emphasis>
|
||||
) genital capsule, lateral view;
|
||||
<emphasis id="B975EAFA5D45152B2D52D0ABFEE7FB2F" bold="true" box="[365,377,1252,1271]" pageId="2" pageNumber="13">3</emphasis>
|
||||
) paramera, distal view;
|
||||
<emphasis id="B975EAFA5D45152B2E61D0ABFDF4FB2F" bold="true" box="[606,618,1252,1271]" pageId="2" pageNumber="13">4</emphasis>
|
||||
) protarsal claw, female;
|
||||
<emphasis id="B975EAFA5D45152B2F6BD0ABFCFEFB2F" bold="true" box="[852,864,1252,1271]" pageId="2" pageNumber="13">5</emphasis>
|
||||
) genital plates, female. Scale lines 5 1 mm, except Figs. 1, 4 5 0.5 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BBE36E85D45152A2D72D1E5FC11FE12" blockId="2.[307,1139,1450,1658]" lastBlockId="3.[279,1112,277,1392]" lastPageId="3" lastPageNumber="14" pageId="2" pageNumber="13">
|
||||
<emphasis id="B975EAFA5D45152B2D72D1E5FE2CFA67" bold="true" box="[333,434,1450,1471]" pageId="2" pageNumber="13">
|
||||
<typeStatus id="54BA884A5D45152B2D72D1E5FE33FA67" box="[333,429,1450,1471]" pageId="2" pageNumber="13" type="allotype">Allotype</typeStatus>
|
||||
.
|
||||
</emphasis>
|
||||
Female. Similar to the male except as follows: antennae with lamellae of 8th to 10th segments slightly longer than the length of six preceeding segments combined (1:0.9). Visible abdominal sternites II to V convex, with scattered setiferous punctures; anal plate 1.2× longer than male anal plate, convex, with scattered setiferous punctures, with 22 slender setae at the posterior border. Pygidium slightly convex, nearly flattened toward the apex, apical border slightly sinuated. Protibiae with the teeth of external border wider and longer than in the male. Both apical spurs of metatibia articulated, wide, lanceolate and curved. Tarsomeres only with two single lines of long, slender setae on ventral side. Tarsal claws each with a large tooth at the middle of ventral border (
|
||||
<figureCitation id="133A2A6D5D44152A2DFCD504FD97FEB8" box="[451,521,331,352]" captionStart="Figs" captionStartId="2.[330,373,1228,1247]" captionTargetBox="[309,1134,273,1204]" captionTargetId="figure-12@2.[308,1137,272,1208]" captionTargetPageId="2" captionText="Figs. 1–5. Phyllophaga tilarana. 1) protarsal claw, male; 2) genital capsule, lateral view; 3) paramera, distal view; 4) protarsal claw, female; 5) genital plates, female. Scale lines 5 1 mm, except Figs. 1, 4 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900313" httpUri="https://zenodo.org/record/4900313/files/figure.png" pageId="3" pageNumber="14">Fig. 4</figureCitation>
|
||||
). Ventral genital plates moderately sclerotized, nearly symmetrical, elongate ovate shaped with shallow depression near the apex; dorsal plates wider than ventral plates with scattered setae on the distal border, each plate partially fused with ventral plate along the laterobasal border (
|
||||
<figureCitation id="133A2A6D5D44152A2812D5D4FEB8FE12" captionStart="Figs" captionStartId="2.[330,373,1228,1247]" captionTargetBox="[309,1134,273,1204]" captionTargetId="figure-12@2.[308,1137,272,1208]" captionTargetPageId="2" captionText="Figs. 1–5. Phyllophaga tilarana. 1) protarsal claw, male; 2) genital capsule, lateral view; 3) paramera, distal view; 4) protarsal claw, female; 5) genital plates, female. Scale lines 5 1 mm, except Figs. 1, 4 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900313" httpUri="https://zenodo.org/record/4900313/files/figure.png" pageId="3" pageNumber="14">Fig. 5</figureCitation>
|
||||
). Total body length:
|
||||
<quantity id="4CF99B0D5D44152A2E35D5FAFDF2FE12" box="[522,620,437,458]" metricMagnitude="-2" metricUnit="m" metricValue="2.46" pageId="3" pageNumber="14" unit="mm" value="24.6">24.6 mm</quantity>
|
||||
. Humeral width: 10.0 mm.
|
||||
</paragraph>
|
||||
<paragraph id="8BBE36E85D44152A2D0DD59FFDFFFDB2" blockId="3.[279,1112,277,1392]" pageId="3" pageNumber="14">
|
||||
<emphasis id="B975EAFA5D44152A2D0DD59FFD8FFE3D" bold="true" box="[306,529,464,485]" pageId="3" pageNumber="14">
|
||||
<typeStatus id="54BA884A5D44152A2D0DD59FFE06FE3D" box="[306,408,464,485]" pageId="3" pageNumber="14" type="paratype">Paratype</typeStatus>
|
||||
Variation.
|
||||
</emphasis>
|
||||
Males. Similar to
|
||||
<typeStatus id="54BA884A5D44152A2EEAD59FFCAEFE3D" box="[725,816,464,485]" pageId="3" pageNumber="14" type="holotype">holotype</typeStatus>
|
||||
except in total body length: 20.0–24.0 mm, humeral width:
|
||||
<quantity id="4CF99B0D5D44152A2E43D5A4FC9CFDD8" box="[636,770,491,512]" metricMagnitude="-3" metricUnit="m" metricValue="8.649999999999999" metricValueMax="9.1" metricValueMin="8.2" pageId="3" pageNumber="14" unit="mm" value="8.649999999999999" valueMax="9.1" valueMin="8.2">8.2–9.1 mm</quantity>
|
||||
, pronotum of some specimens darker than
|
||||
<typeStatus id="54BA884A5D44152A2DA1D64AFE62FDC2" box="[414,508,517,538]" pageId="3" pageNumber="14" type="holotype">holotype</typeStatus>
|
||||
, other specimens with antennal club 2.2× longer than length of preceeding six segments combined. Female similar to
|
||||
<typeStatus id="54BA884A5D44152A2F86D66FFB93FDED" box="[953,1037,544,565]" pageId="3" pageNumber="14" type="allotype">allotype</typeStatus>
|
||||
except as follows: elytra and pygidium with more punctures; total body length:
|
||||
<quantity id="4CF99B0D5D44152A2815D674FEDDFDB2" metricMagnitude="-2" metricUnit="m" metricValue="2.43" pageId="3" pageNumber="14" unit="mm" value="24.3">24.3 mm</quantity>
|
||||
; humeral width: 10.0 mm.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D44152A2D0DD63FFE38FC72" pageId="3" pageNumber="14" type="materials_examined">
|
||||
<paragraph id="8BBE36E85D44152A2D0DD63FFDD1FC82" blockId="3.[279,1112,277,1392]" pageId="3" pageNumber="14">
|
||||
<emphasis id="B975EAFA5D44152A2D0DD63FFE20FD5D" bold="true" box="[306,446,624,645]" pageId="3" pageNumber="14">Type Series.</emphasis>
|
||||
<collectingCounty id="62DF4E645D44152A2DF7D63FFDACFD5D" box="[456,562,624,645]" pageId="3" pageNumber="14">Described</collectingCounty>
|
||||
from
|
||||
<specimenCount id="9D07FD615D44152A2E45D63FFD50FD5D" box="[634,718,624,645]" pageId="3" pageNumber="14" type="male">9 males</specimenCount>
|
||||
and
|
||||
<specimenCount id="9D07FD615D44152A2F36D63FFCEBFD5D" box="[777,885,624,645]" pageId="3" pageNumber="14" type="female">2 females</specimenCount>
|
||||
.
|
||||
<materialsCitation id="3B693CB55D44152A2FBDD63FFD48FD62" collectingDate="1990-11" collectionCode="IN- BIO" collectorName="M. Ramirez." country="Costa Rica" elevation="1520" latitude="10.18" location="Reserva Biologica Monteverde" longitude="-84.47" municipality="Estacion La Casona" pageId="3" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="holotype">
|
||||
<typeStatus id="54BA884A5D44152A2FBDD63FFC7DFD5D" box="[898,995,624,645]" pageId="3" pageNumber="14" type="holotype">Holotype</typeStatus>
|
||||
<specimenCount id="9D07FD615D44152A2FD1D63FFBBFFD5D" box="[1006,1057,624,645]" pageId="3" pageNumber="14" type="male">male</specimenCount>
|
||||
(
|
||||
<collectionCode id="ED10AE2D5D44152A280DD63FFED7FD78" pageId="3" pageNumber="14">INBIO</collectionCode>
|
||||
): ‘‘
|
||||
<collectingCountry id="F31676785D44152A2D57D6C4FD8EFD78" box="[360,528,651,672]" name="Costa Rica" pageId="3" pageNumber="14">COSTA RICA</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49C5F80A5D44152A2E22D6C4FD0CFD78" box="[541,658,651,672]" country="Costa Rica" name="Puntarenas" pageId="3" pageNumber="14">Puntarenas</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="6BDAAC925D44152A2E9FD6C4FC6EFD78" box="[672,1008,651,672]" pageId="3" pageNumber="14">Reserva Biológica Monteverde</collectingMunicipality>
|
||||
,
|
||||
<location id="8EDE60335D44152A2FC1D6C4FE14FD62" LSID="urn:lsid:plazi:treatment:03A887FE5D46152A2D70D613FCBCFAA8:8EDE60335D44152A2FC1D6C4FE14FD62" country="Costa Rica" county="Described" municipality="Reserva Biologica Monteverde" name="Estacion La Casona" pageId="3" pageNumber="14" stateProvince="Puntarenas">Estación La Casona</location>
|
||||
,
|
||||
<quantity id="4CF99B0D5D44152A2DABD6EAFE73FD62" box="[404,493,677,698]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="3" pageNumber="14" unit="m" value="1520.0">
|
||||
<elevation id="002CD1DB5D44152A2DABD6EAFE73FD62" box="[404,493,677,698]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="3" pageNumber="14" unit="m" value="1520.0">1,520 m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<date id="FFBF10285D44152A2DC8D6EAFDA6FD62" box="[503,568,677,698]" pageId="3" pageNumber="14" value="1990-11">
|
||||
<collectingDate id="EFFBE9C05D44152A2DC8D6EAFDA6FD62" box="[503,568,677,698]" pageId="3" pageNumber="14" value="1990-11">XI90</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F4533E5D44152A2E7CD6EAFD53FD62" box="[579,717,677,698]" pageId="3" pageNumber="14">M. Ramírez.</collectorName>
|
||||
’’
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D44152A2EE2D6EAFDC5FD28" collectingDate="1995-04-24" collectionCode="INBIO" collectorName="A. Azofeifa." country="Costa Rica" elevation="1520" latitude="10.18" location="Reserva Biologica Monteverde" longitude="-84.47" municipality="estacion La Casona" pageId="3" pageNumber="14" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="allotype">
|
||||
<typeStatus id="54BA884A5D44152A2EE2D6EAFCA6FD62" box="[733,824,677,698]" pageId="3" pageNumber="14" type="allotype">Allotype</typeStatus>
|
||||
<specimenCount id="9D07FD615D44152A2F00D6EAFC18FD62" box="[831,902,677,698]" pageId="3" pageNumber="14" type="female">female</specimenCount>
|
||||
(
|
||||
<collectionCode id="ED10AE2D5D44152A2FABD6EAFC7DFD62" box="[916,995,677,698]" pageId="3" pageNumber="14">INBIO</collectionCode>
|
||||
): ‘‘
|
||||
<collectingCountry id="F31676785D44152A2FC2D6EAFEC4FD0D" name="Costa Rica" pageId="3" pageNumber="14">COSTA RICA</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49C5F80A5D44152A2D5AD68FFE44FD0D" box="[357,474,704,725]" country="Costa Rica" name="Puntarenas" pageId="3" pageNumber="14">Puntarenas</collectingRegion>
|
||||
,
|
||||
<location id="8EDE60335D44152A2DD9D68FFCACFD0D" LSID="urn:lsid:plazi:treatment:03A887FE5D46152A2D70D613FCBCFAA8:8EDE60335D44152A2DD9D68FFCACFD0D" box="[486,818,704,725]" country="Costa Rica" county="Described" municipality="Reserva Biologica Monteverde" name="Reserva Biologica Monteverde" pageId="3" pageNumber="14" stateProvince="Puntarenas">Reserva Biológica Monteverde</location>
|
||||
,
|
||||
<location id="8EDE60335D44152A2F02D68FFB8CFD0D" LSID="urn:lsid:plazi:treatment:03A887FE5D46152A2D70D613FCBCFAA8:8EDE60335D44152A2F02D68FFB8CFD0D" box="[829,1042,704,725]" country="Costa Rica" county="Described" municipality="Reserva Biologica Monteverde" name="Estacion La Casona" pageId="3" pageNumber="14" stateProvince="Puntarenas">Estación La Casona</location>
|
||||
,
|
||||
<quantity id="4CF99B0D5D44152A2822D68FFEB0FD28" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="3" pageNumber="14" unit="m" value="1520.0">
|
||||
<elevation id="002CD1DB5D44152A2822D68FFEB0FD28" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="3" pageNumber="14" unit="m" value="1520.0">1,520 m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<date id="FFBF10285D44152A2D01D694FE2AFD28" box="[318,436,731,752]" pageId="3" pageNumber="14" value="1995-04-24">
|
||||
<collectingDate id="EFFBE9C05D44152A2D01D694FE2AFD28" box="[318,436,731,752]" pageId="3" pageNumber="14">3/24IV95</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F4533E5D44152A2DFBD694FDCCFD28" box="[452,594,731,752]" pageId="3" pageNumber="14">A. Azofeifa.</collectorName>
|
||||
’’
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D44152A2E58D694FE51FCD2" collectionCode="CNC, INBIO, MXAL" collectorName="E. Bello" country="Costa Rica" elevation="1520" latitude="10.18" location="Reserva Biologica Monteverde" longitude="-84.47" municipality="Estacion La Casona" pageId="3" pageNumber="14" specimenCount="2" specimenCount-male="2" stateProvince="Puntarenas" typeStatus="paratype">
|
||||
<typeStatus id="54BA884A5D44152A2E58D694FD4FFD28" box="[615,721,731,752]" pageId="3" pageNumber="14" type="paratype">Paratypes</typeStatus>
|
||||
: same data as holotype except ‘‘
|
||||
<collectorName id="26F4533E5D44152A2801D694FEC5FCD2" pageId="3" pageNumber="14">E. Bello’</collectorName>
|
||||
’ (
|
||||
<specimenCount id="9D07FD615D44152A2D4FD6BAFE56FCD2" box="[368,456,757,778]" count="2" pageId="3" pageNumber="14" type="male">2 males</specimenCount>
|
||||
)
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B693CB55D44152A2DE2D6BAFD33FCD2" box="[477,685,757,778]" collectingDate="1990-12" collectorName="de Tilaran" country="Costa Rica" latitude="10.18" location="Reserva Biologica Monteverde" longitude="-84.47" municipality="Estacion La Casona" pageId="3" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
|
||||
‘‘
|
||||
<date id="FFBF10285D44152A2DD8D6BAFDA7FCD2" box="[487,569,757,778]" pageId="3" pageNumber="14" value="1990-12">
|
||||
<collectingDate id="EFFBE9C05D44152A2DD8D6BAFDA7FCD2" box="[487,569,757,778]" pageId="3" pageNumber="14" value="1990-12">XII90</collectingDate>
|
||||
</date>
|
||||
’’ (
|
||||
<specimenCount id="9D07FD615D44152A2E69D6BAFD3FFCD2" box="[598,673,757,778]" count="1" pageId="3" pageNumber="14" type="male">1 male</specimenCount>
|
||||
);
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D44152A2E8AD6BAFBC6FCD2" box="[693,1112,757,778]" collectingDate="1992-03" collectionCode="CNC, INBIO, MXAL" collectorName="K. Flores'" country="Costa Rica" elevation="1520" latitude="10.18" location="Reserva Biologica Monteverde" longitude="-84.47" municipality="Estacion La Casona" pageId="3" pageNumber="14" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
|
||||
‘‘
|
||||
<date id="FFBF10285D44152A2EFFD6BAFC9AFCD2" box="[704,772,757,778]" pageId="3" pageNumber="14" value="1992-03">
|
||||
<collectingDate id="EFFBE9C05D44152A2EFFD6BAFC9AFCD2" box="[704,772,757,778]" pageId="3" pageNumber="14" value="1992-03">III92</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F4533E5D44152A2F2ED6BAFC1EFCD2" box="[785,896,757,778]" pageId="3" pageNumber="14">K. Flores’</collectorName>
|
||||
’ (
|
||||
<specimenCount id="9D07FD615D44152A2FA9D6BAFC41FCD2" box="[918,991,757,778]" count="1" pageId="3" pageNumber="14" type="male">1 male</specimenCount>
|
||||
,
|
||||
<specimenCount id="9D07FD615D44152A2FD3D6BAFBD3FCD2" box="[1004,1101,757,778]" count="1" pageId="3" pageNumber="14" type="female">1 female</specimenCount>
|
||||
);
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D44152A2D28D75FFDC7FCFD" box="[279,601,784,805]" collectingDate="1993-01" collectionCode="CNC, INBIO, MXAL" collectorName="N. Obando'" country="Costa Rica" elevation="1520" latitude="10.18" location="Reserva Biologica Monteverde" longitude="-84.47" municipality="Estacion La Casona" pageId="3" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
|
||||
‘‘
|
||||
<date id="FFBF10285D44152A2D1CD75FFECBFCFD" box="[291,341,784,805]" pageId="3" pageNumber="14" value="1993-01">
|
||||
<collectingDate id="EFFBE9C05D44152A2D1CD75FFECBFCFD" box="[291,341,784,805]" pageId="3" pageNumber="14" value="1993-01">I93</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F4533E5D44152A2D5BD75FFE78FCFD" box="[356,486,784,805]" pageId="3" pageNumber="14">N. Obando’</collectorName>
|
||||
’ (
|
||||
<specimenCount id="9D07FD615D44152A2E3FD75FFDD0FCFD" box="[512,590,784,805]" count="1" pageId="3" pageNumber="14" type="male">1 male</specimenCount>
|
||||
);
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D44152A2E5BD75FFC2EFCFD" box="[612,944,784,805]" collectingDate="1993-10" collectionCode="CNC, INBIO, MXAL" collectorName="N. Obando'" country="Costa Rica" elevation="1520" latitude="10.18" location="Reserva Biologica Monteverde" longitude="-84.47" municipality="Estacion La Casona" pageId="3" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
|
||||
‘‘
|
||||
<date id="FFBF10285D44152A2E50D75FFD32FCFD" box="[623,684,784,805]" pageId="3" pageNumber="14" value="1993-10">
|
||||
<collectingDate id="EFFBE9C05D44152A2E50D75FFD32FCFD" box="[623,684,784,805]" pageId="3" pageNumber="14" value="1993-10">X93</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F4533E5D44152A2E84D75FFCA3FCFD" box="[699,829,784,805]" pageId="3" pageNumber="14">N. Obando’</collectorName>
|
||||
’ (
|
||||
<specimenCount id="9D07FD615D44152A2F68D75FFC3AFCFD" box="[855,932,784,805]" count="1" pageId="3" pageNumber="14" type="male">1 male</specimenCount>
|
||||
);
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D44152A2F84D75FFE6FFC98" collectingDate="1995-01-22" collectionCode="CNC, INBIO, MXAL" collectorName="K. Martinez'" country="Costa Rica" elevation="1520" latitude="10.18" location="Reserva Biologica Monteverde" longitude="-84.47" municipality="Estacion La Casona" pageId="3" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
|
||||
‘‘
|
||||
<date id="FFBF10285D44152A2FFAD75FFBB1FCFD" box="[965,1071,784,805]" pageId="3" pageNumber="14" value="1995-01-22">
|
||||
<collectingDate id="EFFBE9C05D44152A2FFAD75FFBB1FCFD" box="[965,1071,784,805]" pageId="3" pageNumber="14">3/22I95</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F4533E5D44152A2801D75FFE1FFC98" pageId="3" pageNumber="14">K. Martínez’</collectorName>
|
||||
’ (
|
||||
<specimenCount id="9D07FD615D44152A2DA6D764FE7BFC98" box="[409,485,811,832]" count="1" pageId="3" pageNumber="14" type="male">1 male</specimenCount>
|
||||
);
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D44152A2DC4D764FCE2FC98" box="[507,892,811,832]" collectingDate="1995-04-17" collectionCode="CNC, INBIO, MXAL" collectorName="M. Madrigal'" country="Costa Rica" elevation="1520" latitude="10.18" location="Reserva Biologica Monteverde" longitude="-84.47" municipality="Estacion La Casona" pageId="3" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
|
||||
‘‘
|
||||
<date id="FFBF10285D44152A2E3AD764FDF5FC98" box="[517,619,811,832]" pageId="3" pageNumber="14" value="1995-04-17">
|
||||
<collectingDate id="EFFBE9C05D44152A2E3AD764FDF5FC98" box="[517,619,811,832]" pageId="3" pageNumber="14" value="1995-04-17">17IV95</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F4533E5D44152A2E46D764FC92FC98" box="[633,780,811,832]" pageId="3" pageNumber="14">M. Madrigal’</collectorName>
|
||||
’ (
|
||||
<specimenCount id="9D07FD615D44152A2F1BD764FCEEFC98" box="[804,880,811,832]" count="1" pageId="3" pageNumber="14" type="male">1 male</specimenCount>
|
||||
).
|
||||
</materialsCitation>
|
||||
Paratypes deposited in
|
||||
<collectionCode id="ED10AE2D5D44152A2D0AD70AFEF3FC82" box="[309,365,837,858]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" pageId="3" pageNumber="14">CNC</collectionCode>
|
||||
,
|
||||
<collectionCode id="ED10AE2D5D44152A2D45D70AFE5DFC82" box="[378,451,837,858]" pageId="3" pageNumber="14">INBIO</collectionCode>
|
||||
and
|
||||
<collectionCode id="ED10AE2D5D44152A2DC5D70AFDD4FC82" box="[506,586,837,858]" pageId="3" pageNumber="14">MXAL</collectionCode>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BBE36E85D44152A2D0DD72FFE38FC72" blockId="3.[279,1112,277,1392]" pageId="3" pageNumber="14">
|
||||
<emphasis id="B975EAFA5D44152A2D0DD72FFE48FCAD" bold="true" box="[306,470,864,885]" pageId="3" pageNumber="14">Type Locality.</emphasis>
|
||||
<collectingMunicipality id="6BDAAC925D44152A2DE0D72FFD29FCAD" box="[479,695,864,885]" pageId="3" pageNumber="14">Estación La Casona</collectingMunicipality>
|
||||
,
|
||||
<location id="8EDE60335D44152A2EFBD72FFB8CFCAD" LSID="urn:lsid:plazi:treatment:03A887FE5D46152A2D70D613FCBCFAA8:8EDE60335D44152A2EFBD72FFB8CFCAD" box="[708,1042,864,885]" country="Costa Rica" municipality="Estacion La Casona" name="Reserva Biologica Monteverde" pageId="3" pageNumber="14" stateProvince="Puntarenas">Reserva Biológica Monteverde</location>
|
||||
, province of
|
||||
<collectingRegion id="49C5F80A5D44152A2DBBD734FE60FC48" box="[388,510,891,912]" country="Costa Rica" name="Puntarenas" pageId="3" pageNumber="14">Puntarenas</collectingRegion>
|
||||
, cordillera de
|
||||
<location id="8EDE60335D44152A2EFED734FC8CFC48" LSID="urn:lsid:plazi:treatment:03A887FE5D46152A2D70D613FCBCFAA8:8EDE60335D44152A2EFED734FC8CFC48" box="[705,786,891,912]" country="Costa Rica" municipality="Estacion La Casona" name="Tilaran" pageId="3" pageNumber="14" stateProvince="Puntarenas">Tilarán</location>
|
||||
,
|
||||
<collectingCountry id="F31676785D44152A2F14D734FC2FFC48" box="[811,945,891,912]" name="Costa Rica" pageId="3" pageNumber="14">Costa Rica</collectingCountry>
|
||||
. (
|
||||
<geoCoordinate id="EE35502F5D44152A2FEED734FBCAFC48" box="[977,1108,891,912]" direction="north" orientation="latitude" pageId="3" pageNumber="14" value="10.18">10°18'150N</geoCoordinate>
|
||||
;
|
||||
<geoCoordinate id="EE35502F5D44152A2D28D7DAFE05FC72" box="[279,411,917,938]" direction="west" orientation="longitude" pageId="3" pageNumber="14" value="-84.47">84°47'460W</geoCoordinate>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D44152A2D0DD7FFFDB2FB92" pageId="3" pageNumber="14" type="biology_ecology">
|
||||
<paragraph id="8BBE36E85D44152A2D0DD7FFFDB2FB92" blockId="3.[279,1112,277,1392]" pageId="3" pageNumber="14">
|
||||
<emphasis id="B975EAFA5D44152A2D0DD7FFFE79FC1D" bold="true" box="[306,487,944,965]" pageId="3" pageNumber="14">Biological Data.</emphasis>
|
||||
Males and females of
|
||||
<taxonomicName id="4C014D6B5D44152A2EDAD7FFFCC4FC1D" box="[741,858,944,965]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="14" phylum="Arthropoda" rank="species" species="tilarana" status="sp. nov.">
|
||||
<emphasis id="B975EAFA5D44152A2EDAD7FFFCC4FC1D" box="[741,858,944,965]" italics="true" pageId="3" pageNumber="14">P. tilarana</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A24657815D44152A2F5BD7FFFC08FC1D" box="[868,918,944,965]" pageId="3" pageNumber="14" rank="species">n.sp.</taxonomicNameLabel>
|
||||
were collected at lights in cloud forest located near
|
||||
<quantity id="4CF99B0D5D44152A2EA2D784FD69FC38" box="[669,759,971,992]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="3" pageNumber="14" unit="m" value="1520.0">1,520 m</quantity>
|
||||
of altitude,
|
||||
<quantity id="4CF99B0D5D44152A2FB8D784FC5FFC38" box="[903,961,971,992]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="14" unit="km" value="1.0">1 km</quantity>
|
||||
southwest of Atlantic/Pacific montane rainfall line. Phenology: January (2), March (2), April (2), October (1), November (3), December (1). Other species of
|
||||
<taxonomicName id="4C014D6B5D44152A2FEDD04FFBC6FBCD" box="[978,1112,1024,1045]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B975EAFA5D44152A2FEDD04FFBC6FBCD" box="[978,1112,1024,1045]" italics="true" pageId="3" pageNumber="14">Phyllophaga</emphasis>
|
||||
</taxonomicName>
|
||||
flying at the same time and place were
|
||||
<taxonomicName id="4C014D6B5D44152A2EF9D054FCD4FBE8" box="[710,842,1051,1072]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="14" phylum="Arthropoda" rank="species" species="tapantina" status="sp. nov.">
|
||||
<emphasis id="B975EAFA5D44152A2EF9D054FCD4FBE8" box="[710,842,1051,1072]" italics="true" pageId="3" pageNumber="14">P. tapantina</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A24657815D44152A2F6CD054FC1BFBE8" box="[851,901,1051,1072]" pageId="3" pageNumber="14" rank="species">n.sp.</taxonomicNameLabel>
|
||||
and two other new species under description.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D44152A2D0DD01FFBCCFAE2" pageId="3" pageNumber="14" type="discussion">
|
||||
<paragraph id="8BBE36E85D44152A2D0DD01FFBCCFAE2" blockId="3.[279,1112,277,1392]" pageId="3" pageNumber="14">
|
||||
<emphasis id="B975EAFA5D44152A2D0DD01FFE00FBBD" bold="true" box="[306,414,1104,1125]" pageId="3" pageNumber="14">Remarks.</emphasis>
|
||||
<taxonomicName id="4C014D6B5D44152A2D99D01FFCB6FBBD" box="[422,808,1104,1125]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="14" phylum="Arthropoda" rank="species" species="tilarana" subGenus="Phyllophaga">
|
||||
<emphasis id="B975EAFA5D44152A2D99D01FFDB2FBBD" box="[422,556,1104,1125]" italics="true" pageId="3" pageNumber="14">Phyllophaga</emphasis>
|
||||
(
|
||||
<emphasis id="B975EAFA5D44152A2E02D01FFD5CFBBD" box="[573,706,1104,1125]" italics="true" pageId="3" pageNumber="14">Phyllophaga</emphasis>
|
||||
)
|
||||
<emphasis id="B975EAFA5D44152A2EEBD01FFCB6FBBD" box="[724,808,1104,1125]" italics="true" pageId="3" pageNumber="14">tilarana</emphasis>
|
||||
</taxonomicName>
|
||||
does not belong to any species group described by
|
||||
<bibRefCitation id="EF904B195D44152A2E1BD024FD20FB58" author="Moron" box="[548,702,1131,1152]" firstAuthor="Moron" pageId="3" pageNumber="14" refId="ref11224" refString="Moron, M. A. 1986. El genero Phyllophaga en Mexico. Morfologia, distribucion y sistematica supraespecifica (Insecta: Coleoptera). Publ. 20, Instituto de Ecologia, Mexico, D. F. 341 pp." type="book" year="1986">Morón (1986)</bibRefCitation>
|
||||
. Male tarsal claw structure resembles ‘‘
|
||||
<emphasis id="B975EAFA5D44152A2D18D0C9FE09FB43" box="[295,407,1158,1179]" italics="true" pageId="3" pageNumber="14">rugipennis</emphasis>
|
||||
’’ group, but female tarsal claws have typical dentate structure, found in many other groups.
|
||||
<taxonomicName id="4C014D6B5D44152A2E71D0EFFCB1FB6D" box="[590,815,1184,1205]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="14" phylum="Arthropoda" rank="species" species="tilarana">
|
||||
<emphasis id="B975EAFA5D44152A2E71D0EFFCB1FB6D" box="[590,815,1184,1205]" italics="true" pageId="3" pageNumber="14">Phyllophaga tilarana</emphasis>
|
||||
</taxonomicName>
|
||||
may be related with
|
||||
<emphasis id="B975EAFA5D44152A282DD0EFFBB7FB6D" box="[1042,1065,1184,1205]" italics="true" pageId="3" pageNumber="14">P.</emphasis>
|
||||
(
|
||||
<emphasis id="B975EAFA5D44152A2807D0EFFBD1FB6D" box="[1080,1103,1184,1205]" italics="true" pageId="3" pageNumber="14">P.</emphasis>
|
||||
)
|
||||
<emphasis id="B975EAFA5D44152A2D28D0F4FE04FB08" box="[279,410,1211,1232]" italics="true" pageId="3" pageNumber="14">nevermanni,</emphasis>
|
||||
described by
|
||||
<bibRefCitation id="EF904B195D44152A2E09D0F4FD67FB08" author="Saylor, L. W." box="[566,761,1211,1232]" journalOrPublisher="Revista de Entomologia (Rio de Janeiro)" pageId="3" pageNumber="14" pagination="496 - 501" part="5" refId="ref11267" refString="Saylor, L. W. 1935. New neotropical Scarabaeidae of the genus Phyllophaga (Col.). Revista de Entomologia (Rio de Janeiro) 5: 496 - 501." title="New neotropical Scarabaeidae of the genus Phyllophaga (Col.)" type="journal article" year="1935">Saylor (1935:501)</bibRefCitation>
|
||||
from Guápiles,
|
||||
<collectingCountry id="F31676785D44152A2F92D0F4FBBCFB08" box="[941,1058,1211,1232]" name="Costa Rica" pageId="3" pageNumber="14">Costa Rica</collectingCountry>
|
||||
(
|
||||
<typeStatus id="54BA884A5D44152A280CD0F4FEC6FB32" pageId="3" pageNumber="14" type="holotype">holotype</typeStatus>
|
||||
USNM 54567) by the body shape, color, dorsal vestiture, antennal structure and the shape of tarsal claws; but the shape and details of the male genital capsule, the accessorial structures of male sternites V and VI, vestiture and shape of the pygidium in males and females aid to separate the new species.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D44152A2D0DD10FFCBCFAA8" pageId="3" pageNumber="14" type="etymology">
|
||||
<paragraph id="8BBE36E85D44152A2D0DD10FFCBCFAA8" blockId="3.[279,1112,277,1392]" pageId="3" pageNumber="14">
|
||||
<emphasis id="B975EAFA5D44152A2D0DD10FFE2FFA8D" bold="true" box="[306,433,1344,1365]" pageId="3" pageNumber="14">Etymology.</emphasis>
|
||||
Derived from Tilarán Mountain System of northwestern
|
||||
<collectingCountry id="F31676785D44152A2823D10FFED9FAA8" name="Costa Rica" pageId="3" pageNumber="14">Costa Rica</collectingCountry>
|
||||
where this species was originally collected.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
286
data/03/A8/87/03A887FE5D4815382D03D0A1FC9DF9D6.xml
Normal file
286
data/03/A8/87/03A887FE5D4815382D03D0A1FC9DF9D6.xml
Normal file
|
|
@ -0,0 +1,286 @@
|
|||
<document id="A30EC588A67FA02E627BFEBD6956A053" ID-DOI="10.1649/0010-065x(2001)055[0011:snsops]2.0.co;2" ID-ISSN="1938-4394" ID-Zenodo-Dep="4900311" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1622827801739" checkinUser="carolina" docAuthor="Morón, Miguel Angel & Solís, Angel" docDate="2001" docId="03A887FE5D4815382D03D0A1FC9DF9D6" docLanguage="en" docName="TheColeopBull.55.1.11-29.pdf.imf" docOrigin="The Coleopterists Bulletin 55 (1)" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Phyllophaga (Phyllophaga) guapiloides Moron and Solis, new species" docType="treatment" docVersion="1" lastPageNumber="28" masterDocId="FF91FF865D4715292C3FD44FFF9EFFD8" masterDocTitle="Seven New Species Of Phyllophaga (S. Str.) Harris From Costa Rica (Coleoptera: Melolonthidae: Melolonthinae)" masterLastPageNumber="29" masterPageNumber="11" pageNumber="26" updateTime="1730139982923" updateUser="juliana" zenodo-license-document="CLOSED">
|
||||
<mods:mods id="95FB120637ED8D17FA283F3562FB8A60" xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo id="B933A77CE125AE3536C5119A2AE69192">
|
||||
<mods:title id="305F567F01DCB2B56B67827DCF99C845">Seven New Species Of Phyllophaga (S. Str.) Harris From Costa Rica (Coleoptera: Melolonthidae: Melolonthinae)</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name id="2D59EB587554A12C8FA6E81AD1F41822" type="personal">
|
||||
<mods:role id="D7C05A901127A432771F55C9F8B414F4">
|
||||
<mods:roleTerm id="4E834789EB5582318E56AE169658977E">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="D3067EEE2B33650A5B26876864B23BFE">Morón, Miguel Angel</mods:namePart>
|
||||
<mods:affiliation id="D494B4FA4217D655DA1DF1588691E455">Departamento de Entomología Instituto de Ecología, A. C. (sector SEPCONACYT) Apartado Postal 63 Xalapa, Veracruz 91000, MÉXICO</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name id="E23315D869D7DD59D57F0B1A1550A9FD" type="personal">
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<mods:role id="802AFC9C307AFC087DC94596DD2E404D">
|
||||
<mods:roleTerm id="7DB57396A9B1608EDEE71E1B90BDA3B3">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="AEB7F70E97F3014F488F18CBC8D116D2">Solís, Angel</mods:namePart>
|
||||
<mods:affiliation id="DC7B51928057815D6353C4159BBDDAC4">Departamento de Entomología Instituto Nacional de Biodiversidad (INBio) Apartado Postal 22 3100 Santo Domingo de Heredia, COSTA RICA</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:typeOfResource id="57C24F732584ABB8776E98FC73BFC3E3">text</mods:typeOfResource>
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<mods:titleInfo id="2861DA605E06AB14ABF8CC5B580CEDE6">
|
||||
<mods:title id="F32ECAD589053F8AF9E15B7F89127A6C">The Coleopterists Bulletin</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part id="B1ECD1FA92C72FF975983CD22D523B81">
|
||||
<mods:date id="49AD67AE9C329FFFA7F54939AC7EB1E2">2001</mods:date>
|
||||
<mods:detail id="E0D9882B7B7D5F00F24BD7EDFD066E74" type="pubDate">
|
||||
<mods:number id="EB9720A1FB9BF8FF3B9CBA977D8A8C65">2001-03-01</mods:number>
|
||||
</mods:detail>
|
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<mods:detail id="30D9600B729059CF4BC7E42E2A562C79" type="volume">
|
||||
<mods:number id="0B9EA9FCC6426B3B86DF5289E6261117">55</mods:number>
|
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</mods:detail>
|
||||
<mods:detail id="531E1F5C90AFD6E4161D2CBF7AA8FF28" type="issue">
|
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<mods:number id="24006F5D958D9CFB0F096AB84A7A53DC">1</mods:number>
|
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</mods:detail>
|
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<mods:start id="B77B9546564566EBC2DC1AB3069A14A1">11</mods:start>
|
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<mods:end id="CBED72D0453498182B4D01CD63BE84AC">29</mods:end>
|
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</mods:extent>
|
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|
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</mods:relatedItem>
|
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<mods:classification id="80FFD55E915F9632D6AB9FFA0551042E">journal article</mods:classification>
|
||||
<mods:identifier id="8BDA437877788A397E0292C78A9C333E" type="DOI">10.1649/0010-065x(2001)055[0011:snsops]2.0.co;2</mods:identifier>
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<mods:identifier id="270A2655C19B9CCA2B7527F9DD001772" type="ISSN">1938-4394</mods:identifier>
|
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<mods:identifier id="2CCF109CAB04819108B1CD3255AB7ED7" type="Zenodo-Dep">4900311</mods:identifier>
|
||||
</mods:mods>
|
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<treatment id="03A887FE5D4815382D03D0A1FC9DF9D6" LSID="urn:lsid:plazi:treatment:03A887FE5D4815382D03D0A1FC9DF9D6" httpUri="http://treatment.plazi.org/id/03A887FE5D4815382D03D0A1FC9DF9D6" lastPageId="17" lastPageNumber="28" pageId="15" pageNumber="26">
|
||||
<subSubSection id="C31B65635D4815262D03D0A1FBADFADA" box="[316,1075,1261,1283]" pageId="15" pageNumber="26" type="nomenclature">
|
||||
<paragraph id="8BBE36E85D4815262D03D0A1FBADFADA" blockId="15.[316,1075,1261,1309]" box="[316,1075,1261,1283]" pageId="15" pageNumber="26">
|
||||
<taxonomicName id="4C014D6B5D4815262D03D0A1FC3DFADB" authority="Moron and Solis" authorityName="Moron and Solis" box="[316,931,1262,1283]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="26" phylum="Arthropoda" rank="species" species="guapiloides" status="sp. nov." subGenus="Phyllophaga">
|
||||
<emphasis id="B975EAFA5D4815262D03D0A1FE5CFADB" box="[316,450,1262,1283]" italics="true" pageId="15" pageNumber="26">Phyllophaga</emphasis>
|
||||
(
|
||||
<emphasis id="B975EAFA5D4815262DEBD0A1FDC7FADB" box="[468,601,1262,1283]" italics="true" pageId="15" pageNumber="26">Phyllophaga</emphasis>
|
||||
)
|
||||
<emphasis id="B975EAFA5D4815262E54D0A1FD7BFADB" box="[619,741,1262,1283]" italics="true" pageId="15" pageNumber="26">guapiloides</emphasis>
|
||||
Morón and Solís
|
||||
</taxonomicName>
|
||||
,
|
||||
<emphasis id="B975EAFA5D4815262F8FD0A2FBADFADA" bold="true" box="[944,1075,1261,1282]" pageId="15" pageNumber="26">
|
||||
<taxonomicNameLabel id="A24657815D4815262F8FD0A2FBADFADA" box="[944,1075,1261,1282]" pageId="15" pageNumber="26" rank="species">new species</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D4815382E46D147FCDAFCA0" lastPageId="17" lastPageNumber="28" pageId="15" pageNumber="26" type="description">
|
||||
<paragraph id="8BBE36E85D4815262E46D147FD69FAC5" blockId="15.[316,1075,1261,1309]" box="[633,759,1288,1309]" pageId="15" pageNumber="26">
|
||||
<figureCitation id="133A2A6D5D4815262E46D147FD69FAC5" box="[633,759,1288,1309]" captionStart="Figs" captionStartId="16.[330,373,903,922]" captionTargetBox="[331,1116,273,880]" captionTargetId="figure-12@16.[325,1120,272,883]" captionTargetPageId="16" captionText="Figs. 28–31. Phyllophaga guapiloides. 28) Protarsal claw, male; 29) genital capsule, lateral view; 30) paramera, distal view; 31) genital plates, female. Scale lines 5 1 mm, except Fig. 28 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900325" httpUri="https://zenodo.org/record/4900325/files/figure.png" pageId="15" pageNumber="26">Figs. 28–31</figureCitation>
|
||||
</paragraph>
|
||||
<paragraph id="8BBE36E85D4815382D10D160FC2FFDE8" blockId="15.[279,1113,1327,1658]" lastBlockId="17.[279,1113,277,1550]" lastPageId="17" lastPageNumber="28" pageId="15" pageNumber="26">
|
||||
<emphasis id="B975EAFA5D4815262D10D160FE11FA9A" bold="true" box="[303,399,1327,1346]" pageId="15" pageNumber="26">
|
||||
<typeStatus id="54BA884A5D4815262D10D160FE15FA9A" box="[303,395,1327,1346]" pageId="15" pageNumber="26" type="holotype">Holotype</typeStatus>
|
||||
.
|
||||
</emphasis>
|
||||
Male. Clypeus, frons and pronotum shiny dark brown; elytra shiny dark brown, head and pronotum with erect, mixed long and short setae on entire surface; elytra with abundant decumbent short setae on entire surface and some scattered erect setae along the suture and toward the sides of scutellum; mouth parts, sterna, pygidium and legs shiny reddish brown. Clypeus 3.8× wider than long, anterior border slightly sinuate, with clearly elevated margin, surface slightly concave, densely rugopunctate, with many short erect setae. Frontoclypeal suture sinuate and partially hidden by punctures. Frons 1.7× wider than long, convex,densely rugopunctate, with mixed short and long setae on entire surface. Antenna 10segmented, with 3segmented club, lamellae of 8th to 10th segments 1.2× longer than length of preceeding 6 segments combined. Frons 5.0× wider than dorsal diameter of eye. Eye canthus long and wide, with 12–15 setae. Labrum bilobed, deeply notched, with abundant slender setae. Mentum slightly convex, with slender setae on each side, moderately sinuate anteriorly. Pronotum 1.8× wider than long and 2.2× wider than frons. Pronotal disk shiny, with deep, round punctures irregularly separated from one another by less than 1 diameter; lateral borders strongly angled, lateral marginal bead crenulate and progressively dentate toward posterior corner, with regularly located, long, slender setae; basal bead indicated by punctures, mainly toward the sides; anterior angles right, clearly prominent; posterior angles obtuse, clearly prominent. Scutellum 1.7× wider than long, with numerous shallow setiferous punctures. Elytron 2.4× longer than wider, shiny, densely setiferous punctate; epipleural border very narrow, extended along the complete margin, provided with dense fringe of short setae for entire length; humeral callus rounded, prominent; apical callus rounded. Metathoracic wings completely developed. Propygidium shiny, with fine punctures densely placed and setiferous punctures. Pygidium moderately convex, shiny, with fine punctures densely placed and setiferous punctures, with scattered long erect setae on distal half of disk; apical margin with 16 long, slender setae; basal margin effaced medially. Pterosterna with long, dense, yellowish setae. Visible abdominal sternites II–IV convex, with scattered fine setiferous punctures; sternite V convex, finely rugose with setiferous punctures on the middle, with a shallow transverse sulcus before the posterior border; anal plate moderately narrowed, slightly concave, with scattered setiferous punctures, anterior and posterior border clearly elevated nearly all along the complete width, with 14 long setae near the posterior border. Protibia slightly shorter than protarsus (0.8:1), with external border tridentate, basal tooth short, preapical spur acute, straight, as long as 2nd protarsomere. Mesotibia with one oblique, sharp, setiferous carina on external side; upper apical spur straight, narrow, and as long as lower spur. Metatibia 0.8× shorter than metatarsus, with one oblique, feeble, setiferous carina on external side; upper apical spur articulated, slightly curved, with apex acute, 0.9× shorter than basal metatarsomere, and 1.6× longer than lower spur; lower apical spur articulated with tibial border, slightly curved, with rounded apex. Tarsomeres semicylindrical, not much elongate, with enlarged apex and some setae apically; meso and metatarsomeres 2–4 ventrally with a serrulate longitudinal carina and one line of short setae at each side; protarsomeres 1–3 each with subapical small toothlike blade. Tarsal claws symmetrical, similar on all legs, each with a large acute tooth near the middle of ventral border with some minute incisions before the base (
|
||||
<figureCitation id="133A2A6D5D5615382DC5D53AFDDAFE50" box="[506,580,373,392]" captionStart="Figs" captionStartId="16.[330,373,903,922]" captionTargetBox="[331,1116,273,880]" captionTargetId="figure-12@16.[325,1120,272,883]" captionTargetPageId="16" captionText="Figs. 28–31. Phyllophaga guapiloides. 28) Protarsal claw, male; 29) genital capsule, lateral view; 30) paramera, distal view; 31) genital plates, female. Scale lines 5 1 mm, except Fig. 28 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900325" httpUri="https://zenodo.org/record/4900325/files/figure.png" pageId="17" pageNumber="28">Fig. 28</figureCitation>
|
||||
). Genital capsule with short and narrowed parameres, dorsally and ventrally fused, ringshaped, apex straight, progresively narrowed, with toothlike apex finely notched. Aedeagus long with preapical patches of strongly sclerotized short spines and one tuft of 7–8 very long setae on the dorsoapical area of sclerotized support, this support is a canaliculated wide structure with the base articulated with membranous parts of trema; ventral border of the apex slightly bisinuate (
|
||||
<figureCitation id="133A2A6D5D5615382FC0D5A2FEAFFDC0" captionStart="Figs" captionStartId="16.[330,373,903,922]" captionTargetBox="[331,1116,273,880]" captionTargetId="figure-12@16.[325,1120,272,883]" captionTargetPageId="16" captionText="Figs. 28–31. Phyllophaga guapiloides. 28) Protarsal claw, male; 29) genital capsule, lateral view; 30) paramera, distal view; 31) genital plates, female. Scale lines 5 1 mm, except Fig. 28 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900325" httpUri="https://zenodo.org/record/4900325/files/figure.png" pageId="17" pageNumber="28">Figs. 29– 30</figureCitation>
|
||||
). Tectum convex. Length of genital capsule from apex of parameres to border of basal piece:
|
||||
<quantity id="4CF99B0D5D5615382D68D652FE3DFDE8" box="[343,419,541,560]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="17" pageNumber="28" unit="mm" value="4.7">4.7 mm</quantity>
|
||||
. Total body length:
|
||||
<quantity id="4CF99B0D5D5615382E5CD652FD25FDE8" box="[611,699,541,560]" metricMagnitude="-2" metricUnit="m" metricValue="1.92" pageId="17" pageNumber="28" unit="mm" value="19.2">19.2 mm</quantity>
|
||||
. Humeral width:
|
||||
<quantity id="4CF99B0D5D5615382F5ED652FC33FDE8" box="[865,941,541,560]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="17" pageNumber="28" unit="mm" value="8.9">8.9 mm</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
<caption id="DF7E66605D5715392D75D7C8FDB3FC12" ID-DOI="http://doi.org/10.5281/zenodo.4900325" ID-Zenodo-Dep="4900325" httpUri="https://zenodo.org/record/4900325/files/figure.png" pageId="16" pageNumber="27" startId="16.[330,373,903,922]" targetBox="[331,1116,273,880]" targetPageId="16">
|
||||
<paragraph id="8BBE36E85D5715392D75D7C8FDB3FC12" blockId="16.[307,1140,903,970]" pageId="16" pageNumber="27">
|
||||
<emphasis id="B975EAFA5D5715392D75D7C8FE21FC42" bold="true" box="[330,447,903,922]" pageId="16" pageNumber="27">Figs. 28–31.</emphasis>
|
||||
<emphasis id="B975EAFA5D5715392DE8D7C8FD59FC42" box="[471,711,903,922]" italics="true" pageId="16" pageNumber="27">
|
||||
<taxonomicName id="4C014D6B5D5715392DE8D7C8FD5DFC42" box="[471,707,903,922]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="27" phylum="Arthropoda" rank="species" species="guapiloides">Phyllophaga guapiloides</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
<emphasis id="B975EAFA5D5715392EF2D7C8FD7AFC42" bold="true" box="[717,740,903,922]" pageId="16" pageNumber="27">28</emphasis>
|
||||
) Protarsal claw, male;
|
||||
<emphasis id="B975EAFA5D5715392F82D7C8FC4AFC42" bold="true" box="[957,980,903,922]" pageId="16" pageNumber="27">29</emphasis>
|
||||
) genital capsule, lateral view;
|
||||
<emphasis id="B975EAFA5D5715392D8FD7D0FE59FC6A" bold="true" box="[432,455,927,946]" pageId="16" pageNumber="27">30</emphasis>
|
||||
) paramera, distal view;
|
||||
<emphasis id="B975EAFA5D5715392E8ED7D0FD56FC6A" bold="true" box="[689,712,927,946]" pageId="16" pageNumber="27">31</emphasis>
|
||||
) genital plates, female. Scale lines 5 1 mm, except Fig. 28 5 0.5 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BBE36E85D5615382D0DD672FBACFCD5" blockId="17.[279,1113,277,1550]" pageId="17" pageNumber="28">
|
||||
<emphasis id="B975EAFA5D5615382D0DD672FE09FD8A" bold="true" box="[306,407,573,594]" pageId="17" pageNumber="28">
|
||||
<typeStatus id="54BA884A5D5615382D0DD672FE0CFD8A" box="[306,402,573,594]" pageId="17" pageNumber="28" type="allotype">Allotype</typeStatus>
|
||||
.
|
||||
</emphasis>
|
||||
Female. Similar to the male except as follows: antennal club 0.8× shorter than the preceeding 6 segments; visible abdominal sternites II to V convex with more setae; anal plate convex. Metafemur 1.4× wider than femur of male. Metatibia at apex 1.4× wider than male metatibia. Both apical spurs of metatibia wider than in male. Pygidium less convex. Ventral genital plates strongly sclerotized, nearly symmetrical, round with wide, slightly elongated, setiferous apex; dorsal genital plates sclerotized, shortened, with wide setiferous apex (
|
||||
<figureCitation id="133A2A6D5D5615382DB8D6B7FE49FCD5" box="[391,471,760,781]" captionStart="Figs" captionStartId="16.[330,373,903,922]" captionTargetBox="[331,1116,273,880]" captionTargetId="figure-12@16.[325,1120,272,883]" captionTargetPageId="16" captionText="Figs. 28–31. Phyllophaga guapiloides. 28) Protarsal claw, male; 29) genital capsule, lateral view; 30) paramera, distal view; 31) genital plates, female. Scale lines 5 1 mm, except Fig. 28 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900325" httpUri="https://zenodo.org/record/4900325/files/figure.png" pageId="17" pageNumber="28">Fig. 31</figureCitation>
|
||||
). Total body length:
|
||||
<quantity id="4CF99B0D5D5615382E86D6B7FC82FCD5" box="[697,796,760,781]" metricMagnitude="-2" metricUnit="m" metricValue="2.19" pageId="17" pageNumber="28" unit="mm" value="21.9">21.9 mm</quantity>
|
||||
. Humeral width:
|
||||
<quantity id="4CF99B0D5D5615382FE7D6B7FBB0FCD5" box="[984,1070,760,781]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="17" pageNumber="28" unit="mm" value="9.2">9.2 mm</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BBE36E85D5615382D0DD75DFCDAFCA0" blockId="17.[279,1113,277,1550]" pageId="17" pageNumber="28">
|
||||
<emphasis id="B975EAFA5D5615382D0DD75DFD8FFCFF" bold="true" box="[306,529,786,807]" pageId="17" pageNumber="28">
|
||||
<typeStatus id="54BA884A5D5615382D0DD75DFE06FCFF" box="[306,408,786,807]" pageId="17" pageNumber="28" type="paratype">Paratype</typeStatus>
|
||||
Variation.
|
||||
</emphasis>
|
||||
Males. Similar to
|
||||
<typeStatus id="54BA884A5D5615382EEAD75CFCAEFCF0" box="[725,816,787,808]" pageId="17" pageNumber="28" type="holotype">holotype</typeStatus>
|
||||
except in total body length: 18.0–
|
||||
<quantity id="4CF99B0D5D5615382D6DD762FE28FC9A" box="[338,438,813,834]" metricMagnitude="-2" metricUnit="m" metricValue="2.12" pageId="17" pageNumber="28" unit="mm" value="21.2">21.2 mm</quantity>
|
||||
, humeral width: 8.0–
|
||||
<quantity id="4CF99B0D5D5615382E9FD762FD69FC9A" box="[672,759,813,834]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="17" pageNumber="28" unit="mm" value="9.5">9.5 mm</quantity>
|
||||
; some specimens slightly darker or lighter than
|
||||
<typeStatus id="54BA884A5D5615382DFBD707FDBDFC85" box="[452,547,840,861]" pageId="17" pageNumber="28" type="holotype">holotype</typeStatus>
|
||||
. Females. Similar to
|
||||
<typeStatus id="54BA884A5D5615382F28D707FCF5FC85" box="[791,875,840,861]" pageId="17" pageNumber="28" type="allotype">allotype</typeStatus>
|
||||
except in total body length: 18.2–21.0 mm, humeral width:
|
||||
<quantity id="4CF99B0D5D5615382E82D72CFCDEFCA0" box="[701,832,867,888]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" metricValueMax="9.1" metricValueMin="8.3" pageId="17" pageNumber="28" unit="mm" value="8.7" valueMax="9.1" valueMin="8.3">8.3–9.1 mm</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D5615382D0DD732FC97FB95" pageId="17" pageNumber="28" type="materials_examined">
|
||||
<paragraph id="8BBE36E85D5615382D0DD732FC35FBC0" blockId="17.[279,1113,277,1550]" pageId="17" pageNumber="28">
|
||||
<emphasis id="B975EAFA5D5615382D0DD732FE22FC4A" bold="true" box="[306,444,893,914]" pageId="17" pageNumber="28">Type Series.</emphasis>
|
||||
Described from
|
||||
<specimenCount id="9D07FD615D5615382E4FD732FD4EFC4A" box="[624,720,893,914]" pageId="17" pageNumber="28" type="male">19 males</specimenCount>
|
||||
and
|
||||
<specimenCount id="9D07FD615D5615382F3AD732FCE2FC4A" box="[773,892,893,914]" pageId="17" pageNumber="28" type="female">13 females</specimenCount>
|
||||
.
|
||||
<materialsCitation id="3B693CB55D5615382FB8D732FEE2FC3A" collectingDate="1992-03-23" collectingDateMax="1992-05-02" collectingDateMin="1992-03-23" collectionCode="IN- BIO" collectorName="F. Araya" country="Costa Rica" elevation="1500" latitude="8.966666" location="Estacion Biologica Las Alturas" longLatPrecision="1300" longitude="-83.1" municipality="Coto Brus" pageId="17" pageNumber="28" specimenCode="INBIO CR 1000 909226" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="holotype">
|
||||
<typeStatus id="54BA884A5D5615382FB8D732FC76FC4A" box="[903,1000,893,914]" pageId="17" pageNumber="28" type="holotype">Holotype</typeStatus>
|
||||
<specimenCount id="9D07FD615D5615382FCFD732FBBDFC4A" box="[1008,1059,893,914]" pageId="17" pageNumber="28" type="male">male</specimenCount>
|
||||
(
|
||||
<collectionCode id="ED10AE2D5D561538280DD732FED7FC75" pageId="17" pageNumber="28">INBIO</collectionCode>
|
||||
): ‘‘
|
||||
<collectingCountry id="F31676785D5615382D55D7D7FE77FC75" box="[362,489,920,941]" name="Costa Rica" pageId="17" pageNumber="28">Costa Rica</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="49C5F80A5D5615382DC7D7D7FDF3FC75" box="[504,621,920,941]" country="Costa Rica" name="Puntarenas" pageId="17" pageNumber="28">Puntarenas</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="6BDAAC925D5615382E42D7D7FD71FC75" box="[637,751,920,941]" pageId="17" pageNumber="28">Coto Brus</collectingMunicipality>
|
||||
,
|
||||
<location id="8EDE60335D5615382EC1D7D7FBCAFC75" LSID="urn:lsid:plazi:treatment:03A887FE5D4815382D03D0A1FC9DF9D6:8EDE60335D5615382EC1D7D7FBCAFC75" box="[766,1108,920,941]" country="Costa Rica" municipality="Coto Brus" name="Estacion Biologica Las Alturas" pageId="17" pageNumber="28" stateProvince="Puntarenas">Estación Biológica Las Alturas</location>
|
||||
,
|
||||
<date id="FFBF10285D5615382D28D7FCFE50FC10" box="[279,462,947,968]" pageId="17" pageNumber="28" value="1992-03-23" valueMax="1992-05-02" valueMin="1992-03-23">
|
||||
<collectingDate id="EFFBE9C05D5615382D28D7FCFE50FC10" box="[279,462,947,968]" pageId="17" pageNumber="28" value="1992-03-23" valueMax="1992-05-02" valueMin="1992-03-23">23III to 2V92</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<quantity id="4CF99B0D5D5615382DDFD7FCFDA1FC10" box="[480,575,947,968]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="28" unit="m" value="1500.0">
|
||||
<elevation id="002CD1DB5D5615382DDFD7FCFDA1FC10" box="[480,575,947,968]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="28" unit="m" value="1500.0">1,500 m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<collectorName id="26F4533E5D5615382E6FD7FCFD30FC10" box="[592,686,947,968]" pageId="17" pageNumber="28">F. Araya</collectorName>
|
||||
LS 322500, 591300/
|
||||
<specimenCode id="DBA79E935D5615382FA0D7FCFEF0FC3A" pageId="17" pageNumber="28">INBIO CR 1000 909226</specimenCode>
|
||||
.’’
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D5615382DBAD782FE3CFC25" accessionNumber="LS 322500" collectingDate="1992-03-23" collectingDateMax="1992-05-02" collectingDateMin="1992-03-23" collectionCode="INBIO" collectorName="F. Araya" country="Costa Rica" elevation="1500" latitude="8.966666" location="El Rosario de Naranjo" longLatPrecision="1300" longitude="-83.1" municipality="Coto Brus" pageId="17" pageNumber="28" specimenCode="INBIO CR I000 909233" specimenCount="1" specimenCount-female="1" stateProvince="Alajuela" typeStatus="allotype">
|
||||
<typeStatus id="54BA884A5D5615382DBAD782FE7EFC3A" box="[389,480,973,994]" pageId="17" pageNumber="28" type="allotype">Allotype</typeStatus>
|
||||
<specimenCount id="9D07FD615D5615382DD6D782FDAEFC3A" box="[489,560,973,994]" pageId="17" pageNumber="28" type="female">female</specimenCount>
|
||||
(
|
||||
<collectionCode id="ED10AE2D5D5615382E7FD782FD11FC3A" box="[576,655,973,994]" pageId="17" pageNumber="28">INBIO</collectionCode>
|
||||
): same data as holotype except
|
||||
<specimenCode id="DBA79E935D5615382FDBD782FE3CFC25" pageId="17" pageNumber="28">INBIO CR I000 909233</specimenCode>
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3B693CB55D5615382D90D7A7FB87FC25" accessionNumber="LS 322500" box="[431,1049,1000,1021]" collectingDate="1992-03-23" collectingDateMax="1992-05-02" collectingDateMin="1992-03-23" collectionCode="CASC, CNC, INBIO, MXAL, ZMHU" collectorName="F. Araya" country="Costa Rica" elevation="1500" latitude="8.966666" location="El Rosario de Naranjo" longLatPrecision="1300" longitude="-83.1" municipality="Coto Brus" pageId="17" pageNumber="28" specimenCode="INBIO CR 1000 909226" specimenCount="30" specimenCount-female="12" specimenCount-male="18" stateProvince="Alajuela" typeStatus="paratype">
|
||||
<typeStatus id="54BA884A5D5615382D90D7A7FD87FC25" box="[431,537,1000,1021]" pageId="17" pageNumber="28" type="paratype">Paratypes</typeStatus>
|
||||
: same data as holotype (
|
||||
<specimenCount id="9D07FD615D5615382F1CD7A7FC19FC25" box="[803,903,1000,1021]" count="18" pageId="17" pageNumber="28" type="male">18 males</specimenCount>
|
||||
,
|
||||
<specimenCount id="9D07FD615D5615382FABD7A7FB93FC25" box="[916,1037,1000,1021]" count="12" pageId="17" pageNumber="28" type="female">12 females</specimenCount>
|
||||
).
|
||||
</materialsCitation>
|
||||
<typeStatus id="54BA884A5D561538281ED7A7FED0FBC0" pageId="17" pageNumber="28" type="paratype">Paratypes</typeStatus>
|
||||
deposited in
|
||||
<collectionCode id="ED10AE2D5D5615382DDDD04CFDB7FBC0" box="[482,553,1027,1048]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="17" pageNumber="28">CASC</collectionCode>
|
||||
,
|
||||
<collectionCode id="ED10AE2D5D5615382E09D04CFDF0FBC0" box="[566,622,1027,1048]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" pageId="17" pageNumber="28">CNC</collectionCode>
|
||||
,
|
||||
<collectionCode id="ED10AE2D5D5615382E44D04CFD58FBC0" box="[635,710,1027,1048]" pageId="17" pageNumber="28">INBIO</collectionCode>
|
||||
,
|
||||
<collectionCode id="ED10AE2D5D5615382EECD04CFCBEFBC0" box="[723,800,1027,1048]" pageId="17" pageNumber="28">MXAL</collectionCode>
|
||||
and
|
||||
<collectionCode id="ED10AE2D5D5615382F68D04CFC39FBC0" box="[855,935,1027,1048]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="17" pageNumber="28">ZMHU</collectionCode>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BBE36E85D5615382D0DD052FC97FB95" blockId="17.[279,1113,277,1550]" pageId="17" pageNumber="28">
|
||||
<emphasis id="B975EAFA5D5615382D0DD052FE4AFBEA" bold="true" box="[306,468,1053,1074]" pageId="17" pageNumber="28">Type Locality.</emphasis>
|
||||
<location id="8EDE60335D5615382DE5D052FCBBFBEA" LSID="urn:lsid:plazi:treatment:03A887FE5D4815382D03D0A1FC9DF9D6:8EDE60335D5615382DE5D052FCBBFBEA" box="[474,805,1053,1074]" country="Costa Rica" latitude="8.966666" longLatPrecision="1300" longitude="-83.1" municipality="Coto Brus" name="Estacion Biologica Las Alturas" pageId="17" pageNumber="28" stateProvince="Puntarenas">Estación Biológica Las Alturas</location>
|
||||
,
|
||||
<collectingMunicipality id="6BDAAC925D5615382F10D052FC03FBEA" box="[815,925,1053,1074]" pageId="17" pageNumber="28">Coto Brus</collectingMunicipality>
|
||||
, province of
|
||||
<collectingRegion id="49C5F80A5D5615382819D052FEFAFB95" country="Costa Rica" name="Puntarenas" pageId="17" pageNumber="28">Puntarenas</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="F31676785D5615382D4ED077FE7BFB95" box="[369,485,1080,1101]" name="Costa Rica" pageId="17" pageNumber="28">Costa Rica</collectingCountry>
|
||||
(approx.
|
||||
<geoCoordinate id="EE35502F5D5615382E70D077FD02FB95" box="[591,668,1080,1101]" degrees="8" direction="north" minutes="58" orientation="latitude" pageId="17" pageNumber="28" precision="925" value="8.966666">8°58'N</geoCoordinate>
|
||||
;
|
||||
<geoCoordinate id="EE35502F5D5615382E96D077FD63FB95" box="[681,765,1080,1101]" degrees="83" direction="west" minutes="6" orientation="longitude" pageId="17" pageNumber="28" precision="925" value="-83.1">83°6'W</geoCoordinate>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D5615382D0DD01DFD0FFB44" pageId="17" pageNumber="28" type="biology_ecology">
|
||||
<paragraph id="8BBE36E85D5615382D0DD01DFD0FFB44" blockId="17.[279,1113,277,1550]" pageId="17" pageNumber="28">
|
||||
<emphasis id="B975EAFA5D5615382D0DD01DFE78FBBF" bold="true" box="[306,486,1106,1127]" pageId="17" pageNumber="28">Biological Data.</emphasis>
|
||||
All males and females of
|
||||
<taxonomicName id="4C014D6B5D5615382F35D01CFC3AFBB0" box="[778,932,1107,1128]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="28" phylum="Arthropoda" rank="species" species="guapiloides" status="sp. nov.">
|
||||
<emphasis id="B975EAFA5D5615382F35D01CFC3AFBB0" box="[778,932,1107,1128]" italics="true" pageId="17" pageNumber="28">P. guapiloides</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A24657815D5615382F91D01CFC77FBB0" box="[942,1001,1107,1128]" pageId="17" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
|
||||
were collected at lights on the border of coffee plantations located at
|
||||
<quantity id="4CF99B0D5D5615382F92D022FB9BFB5A" box="[941,1029,1133,1154]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="28" unit="m" value="1500.0">1,500 m</quantity>
|
||||
of altitude. Phenology: March–May (22).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D5615382D0DD0EEFD1DFA67" pageId="17" pageNumber="28" type="discussion">
|
||||
<paragraph id="8BBE36E85D5615382D0DD0EEFD1DFA67" blockId="17.[279,1113,277,1550]" pageId="17" pageNumber="28">
|
||||
<emphasis id="B975EAFA5D5615382D0DD0EEFE00FB6E" bold="true" box="[306,414,1185,1206]" pageId="17" pageNumber="28">Remarks.</emphasis>
|
||||
<taxonomicName id="4C014D6B5D5615382D96D0EDFCC8FB6F" box="[425,854,1186,1207]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="28" phylum="Arthropoda" rank="species" species="guapiloides" status="sp. nov." subGenus="Phyllophaga">
|
||||
<emphasis id="B975EAFA5D5615382D96D0EDFDB1FB6F" box="[425,559,1186,1207]" italics="true" pageId="17" pageNumber="28">Phyllophaga</emphasis>
|
||||
(
|
||||
<emphasis id="B975EAFA5D5615382E7DD0EDFD59FB6F" box="[578,711,1186,1207]" italics="true" pageId="17" pageNumber="28">Phyllophaga</emphasis>
|
||||
)
|
||||
<emphasis id="B975EAFA5D5615382EE3D0EDFCC8FB6F" box="[732,854,1186,1207]" italics="true" pageId="17" pageNumber="28">guapiloides</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A24657815D5615382F5ED0EDFC00FB6F" box="[865,926,1186,1207]" pageId="17" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
|
||||
probably belong to ‘‘
|
||||
<emphasis id="B975EAFA5D5615382D78D0F3FE35FB09" box="[327,427,1212,1233]" italics="true" pageId="17" pageNumber="28">setidorsis</emphasis>
|
||||
’’ species group (
|
||||
<emphasis id="B975EAFA5D5615382E50D0F3FD37FB09" box="[623,681,1212,1233]" italics="true" pageId="17" pageNumber="28">sensu</emphasis>
|
||||
<bibRefCitation id="EF904B195D5615382E8AD0F3FCA1FB09" author="Moron" box="[693,831,1212,1233]" firstAuthor="Moron" pageId="17" pageNumber="28" refId="ref11224" refString="Moron, M. A. 1986. El genero Phyllophaga en Mexico. Morfologia, distribucion y sistematica supraespecifica (Insecta: Coleoptera). Publ. 20, Instituto de Ecologia, Mexico, D. F. 341 pp." type="book" year="1986">Morón 1986</bibRefCitation>
|
||||
), but the structure of the aedeagus is much different from that species. It is closely related to
|
||||
<taxonomicName id="4C014D6B5D5615382834D098FD2FFADE" authority="Saylor" authorityName="Saylor" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="28" phylum="Arthropoda" rank="species" species="guapiles" subGenus="Phyllophaga">
|
||||
<emphasis id="B975EAFA5D5615382834D098FEC6FADE" italics="true" pageId="17" pageNumber="28">Phyllophaga</emphasis>
|
||||
(
|
||||
<emphasis id="B975EAFA5D5615382D51D0BEFE6AFADE" box="[366,500,1265,1286]" italics="true" pageId="17" pageNumber="28">Phyllophaga</emphasis>
|
||||
)
|
||||
<emphasis id="B975EAFA5D5615382E37D0BEFDFFFADE" box="[520,609,1265,1286]" italics="true" pageId="17" pageNumber="28">guapiles</emphasis>
|
||||
Saylor
|
||||
</taxonomicName>
|
||||
from
|
||||
<collectingRegion id="49C5F80A5D5615382EC3D0BEFCEAFADE" box="[764,884,1265,1286]" country="Costa Rica" name="Guanacaste" pageId="17" pageNumber="28">Guanacaste</collectingRegion>
|
||||
area and is without external differences, but the structure of parameres and the sclerotized support of aedeagus are clearly different (
|
||||
<figureCitation id="133A2A6D5D5615382EBCD169FC98FAE3" box="[643,774,1318,1339]" captionStart="Figs" captionStartId="18.[330,373,687,706]" captionTargetBox="[341,1102,275,666]" captionTargetId="figure-12@18.[340,1105,272,667]" captionTargetPageId="18" captionText="Figs. 32–34. Phyllophaga guapiles. 32) apex of genital capsule, lateral view; 33) apex of paramera and sclerotized support of aedeagus, ventrodistal view; 34) genital plates, female. Scale lines 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4900327" httpUri="https://zenodo.org/record/4900327/files/figure.png" pageId="17" pageNumber="28">Figs. 32–33</figureCitation>
|
||||
). Female genital plates are also slightly different from
|
||||
<taxonomicName id="4C014D6B5D5615382E2BD10FFD10FA8D" box="[532,654,1344,1365]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="28" phylum="Arthropoda" rank="species" species="guapiles">
|
||||
<emphasis id="B975EAFA5D5615382E2BD10FFD10FA8D" box="[532,654,1344,1365]" italics="true" pageId="17" pageNumber="28">P. guapiles</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="133A2A6D5D5615382E9FD10FFD6AFA8D" box="[672,756,1344,1365]" captionStart="Figs" captionStartId="18.[330,373,687,706]" captionTargetBox="[341,1102,275,666]" captionTargetId="figure-12@18.[340,1105,272,667]" captionTargetPageId="18" captionText="Figs. 32–34. Phyllophaga guapiles. 32) apex of genital capsule, lateral view; 33) apex of paramera and sclerotized support of aedeagus, ventrodistal view; 34) genital plates, female. Scale lines 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4900327" httpUri="https://zenodo.org/record/4900327/files/figure.png" pageId="17" pageNumber="28">Fig. 34</figureCitation>
|
||||
). Density, size and shape of the punctures on the head, pronotum, elytra and pygidium; body vestiture; shape of hind tibial spurs, and tarsal claws; structure of tarsomeres and general pattern of aedeagus, will aid to separate this pair of sister species from other species in the ‘‘
|
||||
<emphasis id="B975EAFA5D5615382DFDD1E5FDB8FA67" box="[450,550,1450,1471]" italics="true" pageId="17" pageNumber="28">setidorsis</emphasis>
|
||||
’’ group.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D5615382D0DD18BFC9DF9D6" pageId="17" pageNumber="28" type="etymology">
|
||||
<paragraph id="8BBE36E85D5615382D0DD18BFC9DF9D6" blockId="17.[279,1113,277,1550]" pageId="17" pageNumber="28">
|
||||
<emphasis id="B975EAFA5D5615382D0DD18BFE2FFA01" bold="true" box="[306,433,1476,1497]" pageId="17" pageNumber="28">Etymology.</emphasis>
|
||||
Derived from latin sufix
|
||||
<emphasis id="B975EAFA5D5615382EE2D18BFC85FA01" box="[733,795,1476,1497]" italics="true" pageId="17" pageNumber="28">oides,</emphasis>
|
||||
similar, false (Jaeger 1955), meaning ‘‘similar to
|
||||
<taxonomicName id="4C014D6B5D5615382DCCD190FDD2FA2C" box="[499,588,1503,1524]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="28" phylum="Arthropoda" rank="species" species="guapiles">
|
||||
<emphasis id="B975EAFA5D5615382DCCD190FDD2FA2C" box="[499,588,1503,1524]" italics="true" pageId="17" pageNumber="28">guapiles</emphasis>
|
||||
</taxonomicName>
|
||||
.’’ In the popular Spanish language of
|
||||
<collectingCountry id="F31676785D5615382FE0D190FBCAFA2C" box="[991,1108,1503,1524]" name="Costa Rica" pageId="17" pageNumber="28">Costa Rica</collectingCountry>
|
||||
, the term ‘‘
|
||||
<emphasis id="B975EAFA5D5615382DB6D1B6FE52F9D6" box="[393,460,1529,1550]" italics="true" pageId="17" pageNumber="28">guapil</emphasis>
|
||||
’’ is applied to paired things.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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data/03/A8/87/03A887FE5D4A15262D79D703FEC5FB08.xml
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|
|
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<document id="7AF7B015E32D20C77F9FA2B83CE05BDF" ID-DOI="10.1649/0010-065x(2001)055[0011:snsops]2.0.co;2" ID-ISSN="1938-4394" ID-Zenodo-Dep="4900311" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1622827801739" checkinUser="carolina" docAuthor="Morón, Miguel Angel & Solís, Angel" docDate="2001" docId="03A887FE5D4A15262D79D703FEC5FB08" docLanguage="en" docName="TheColeopBull.55.1.11-29.pdf.imf" docOrigin="The Coleopterists Bulletin 55 (1)" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Phyllophaga (Phyllophaga) naranjina Moron and Solis, new species" docType="treatment" docVersion="1" lastPageNumber="26" masterDocId="FF91FF865D4715292C3FD44FFF9EFFD8" masterDocTitle="Seven New Species Of Phyllophaga (S. Str.) Harris From Costa Rica (Coleoptera: Melolonthidae: Melolonthinae)" masterLastPageNumber="29" masterPageNumber="11" pageNumber="24" updateTime="1730139982923" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:titleInfo id="F4B43F42FEC0E36AE4E6C2836FEC0DF0">
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<mods:title id="FCDEC0335B3C4AC75F25BAE63F06E49F">Seven New Species Of Phyllophaga (S. Str.) Harris From Costa Rica (Coleoptera: Melolonthidae: Melolonthinae)</mods:title>
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<mods:name id="022409E7DE7BE3126AD19552DEECE77C" type="personal">
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<mods:roleTerm id="BAD647F05E8DF97A7596447FB636AC58">Author</mods:roleTerm>
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<mods:namePart id="3F084BE7C55CCB9324FB8FE0E5E773B8">Morón, Miguel Angel</mods:namePart>
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||||
<mods:affiliation id="FECC272E78C4F760E31806B845E97A97">Departamento de Entomología Instituto de Ecología, A. C. (sector SEPCONACYT) Apartado Postal 63 Xalapa, Veracruz 91000, MÉXICO</mods:affiliation>
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</mods:role>
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<mods:namePart id="093367A218D76CB02AB3A9F4F6C2D14F">Solís, Angel</mods:namePart>
|
||||
<mods:affiliation id="08276D8BB905287122D9D8DAB9380264">Departamento de Entomología Instituto Nacional de Biodiversidad (INBio) Apartado Postal 22 3100 Santo Domingo de Heredia, COSTA RICA</mods:affiliation>
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</mods:name>
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<mods:title id="64073F10DF6F5AA60028F2C39C63D634">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="EA1E0194581EEB00F3D7B5397C159072">
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<mods:date id="82ADE12A5FB7CBC67F3F9ADE4B8069D9">2001</mods:date>
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<mods:number id="29FC8E8D8E47091E64341F7E293E2F9D">2001-03-01</mods:number>
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<mods:number id="15ED8276F00A7F0FF22BF9077B526F07">55</mods:number>
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<mods:classification id="03BB215FC16572F4337A42C4F0636041">journal article</mods:classification>
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<mods:identifier id="92F8D4013EBA2D667FD19714942890C7" type="DOI">10.1649/0010-065x(2001)055[0011:snsops]2.0.co;2</mods:identifier>
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<mods:identifier id="59D72BD72ECB18E71FA735D4437AA59B" type="ISSN">1938-4394</mods:identifier>
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<mods:identifier id="325544E7730877742235A625F550A9AB" type="Zenodo-Dep">4900311</mods:identifier>
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</mods:mods>
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<treatment id="03A887FE5D4A15262D79D703FEC5FB08" LSID="urn:lsid:plazi:treatment:03A887FE5D4A15262D79D703FEC5FB08" httpUri="http://treatment.plazi.org/id/03A887FE5D4A15262D79D703FEC5FB08" lastPageId="15" lastPageNumber="26" pageId="13" pageNumber="24">
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<subSubSection id="C31B65635D4A15242D79D703FBB7FCB8" box="[326,1065,843,865]" pageId="13" pageNumber="24" type="nomenclature">
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<paragraph id="8BBE36E85D4A15242D79D703FBB7FCB8" blockId="13.[326,1065,843,891]" box="[326,1065,843,865]" pageId="13" pageNumber="24">
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<taxonomicName id="4C014D6B5D4A15242D79D703FC04FCB8" authority="Moron and Solis" authorityName="Moron and Solis" box="[326,922,843,865]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="24" phylum="Arthropoda" rank="species" species="naranjina" status="sp. nov." subGenus="Phyllophaga">
|
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<emphasis id="B975EAFA5D4A15242D79D703FE52FCB9" box="[326,460,844,865]" italics="true" pageId="13" pageNumber="24">Phyllophaga</emphasis>
|
||||
(
|
||||
<emphasis id="B975EAFA5D4A15242DE2D703FDFCFCB9" box="[477,610,844,865]" italics="true" pageId="13" pageNumber="24">Phyllophaga</emphasis>
|
||||
)
|
||||
<emphasis id="B975EAFA5D4A15242E4BD703FD45FCB9" box="[628,731,844,865]" italics="true" pageId="13" pageNumber="24">naranjina</emphasis>
|
||||
Morón and Solís
|
||||
</taxonomicName>
|
||||
,
|
||||
<emphasis id="B975EAFA5D4A15242F98D704FBB7FCB8" bold="true" box="[935,1065,843,864]" pageId="13" pageNumber="24">
|
||||
<taxonomicNameLabel id="A24657815D4A15242F98D704FBB7FCB8" box="[935,1065,843,864]" pageId="13" pageNumber="24" rank="species">new species</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D4A15262E46D729FDFDFD0D" lastPageId="15" lastPageNumber="26" pageId="13" pageNumber="24" type="description">
|
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<paragraph id="8BBE36E85D4A15242E46D729FD68FCA3" blockId="13.[326,1065,843,891]" box="[633,758,870,891]" pageId="13" pageNumber="24">
|
||||
<figureCitation id="133A2A6D5D4A15242E46D729FD68FCA3" box="[633,758,870,891]" captionStart="Figs" captionStartId="14.[330,373,1095,1114]" captionTargetBox="[336,1117,273,1072]" captionTargetId="figure-12@14.[328,1117,272,1075]" captionTargetPageId="14" captionText="Figs. 22–27. Phyllophaga naranjina. 22) protarsal claw, male; 23) genital capsule, lateral view; 24) paramera, distal view; 25) paramera, ventrodistal view; 26) protarsal claw, female; 27) genital plates, female. Scale lines 5 0.5 mm, except Fig. 23 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4900323" httpUri="https://zenodo.org/record/4900323/files/figure.png" pageId="13" pageNumber="24">Figs. 22–27</figureCitation>
|
||||
</paragraph>
|
||||
<paragraph id="8BBE36E85D4A15272D10D7D8FCF6F9A2" blockId="13.[279,1113,919,1658]" lastBlockId="14.[307,1141,1255,1658]" lastPageId="14" lastPageNumber="25" pageId="13" pageNumber="24">
|
||||
<emphasis id="B975EAFA5D4A15242D10D7D8FE11FC72" bold="true" box="[303,399,919,938]" pageId="13" pageNumber="24">
|
||||
<typeStatus id="54BA884A5D4A15242D10D7D8FE15FC72" box="[303,395,919,938]" pageId="13" pageNumber="24" type="holotype">Holotype</typeStatus>
|
||||
.
|
||||
</emphasis>
|
||||
Male. Clypeus, frons and pronotum shiny reddish testaceus; elytra light yellowish testaceus, head with erect, short setae; pronotum shiny, with short setae around the margins and very scarce short setae on the disk; elytra shiny, nearly glabrous with scattered setae along the suture and toward the posterior half; mouthparts, sterna, pygidium and legs shiny reddish testaceus. Clypeus 5.0× wider than long, anterior border widely rounded, with clearly elevated margin, surface concave, with irregularly distributed, shallow, round punctures, glabrous. Frontoclypeal suture sinuate and effaced at the median third. Frons 1.3× wider than long, convex, coarsely rugopunctate, with short slender setae on entire surface. Antenna 9segmented, with 3segmented club, lamellae of 7th to 9th segments 1.5× longer than length of preceeding 5 segments combined. Frons 2.4× wider than dorsal diameter of eye. Eye canthus long and wide, with 12 setae. Labrum bilobed, shallow and widely sinuate, with scattered slender setae. Mentum slightly convex, with slender setae on each side, anterior border nearly straight. Pronotum 1.6× wider than long and 2.9× wider than frons. Pronotal disk shiny, with deep, round punctures irregularly separated from one another by 1–6 diameters; lateral borders strongly angled, lateral marginal bead crenulate with regularly located, long, slender setae; basal bead indicated by punctures, mainly toward the sides; anterior angles right, clearly prominent; posterior angles obtuse, slighly prominent. Scutellum 1.6× wider than long, with some deep punctures, glabrous. Elytron 2.9× longer than wider, shiny, densely punctate; epipleural border very narrowed, extended along the complete margin, provided with dense fringe of short setae all along; humeral callus rounded, prominent; apical callus rounded. Metathoracic wings completely developed. Propygidium shiny, densely punctate rugose with scattered short setae. Pygidium moderately convex, shiny, deeply punctate rugose toward the midline with scattered erect short setae; apical margin with 12 long, slender setae; basal margin effaced medially. Pterosterna with long, dense, yellowish setae. Visible abdominal sternites II–IV concave, with scattered setiferous granules toward the midline; sternite V nearly flat, shiny, moderately granulose setiferous at the middle, with fine ridge on the posterior border; anal plate moderately narrowed, slightly punctate, convex, with coarse setiferous punctures, a vage longitudinal furrow at midline, anterior border slightly elevated at sides, posterior border not elevated, without setae. Protibia slightly shorter than protarsus (0.7:1), with external border tridentate, basal tooth short, preapical spur acute, slightly curved, shorter than 2nd protarsomere. Mesotibia with one oblique, sharp, setiferous carina on external side; upper apical spur straight, narrow, and as long as lower spur. Metatibia 0.6× shorter than metatarsus, with one oblique, sharp, setiferous carina on external side; upper apical spur articulated, curved, with apex acute, 0.6× shorter than basal metatarsomere, and 1.6× longer than lower spur; lower apical spur articulated with tibial border, with acute apex. Tarsomeres semicylindrical, not much elongate, with enlarged apices and some setae apically; mesoand metatarsomeres 2–4 each with two longitudinal carinae with lines of short setae on ventral side; protarsomeres 1–3 each with subapical small hooklike tubercle. Tarsal claws symmetrical, similar on all legs, with a small acute tooth near the middle of ventral border (
|
||||
<figureCitation id="133A2A6D5D4915272D41D198FE58FA32" box="[382,454,1495,1514]" captionStart="Figs" captionStartId="14.[330,373,1095,1114]" captionTargetBox="[336,1117,273,1072]" captionTargetId="figure-12@14.[328,1117,272,1075]" captionTargetPageId="14" captionText="Figs. 22–27. Phyllophaga naranjina. 22) protarsal claw, male; 23) genital capsule, lateral view; 24) paramera, distal view; 25) paramera, ventrodistal view; 26) protarsal claw, female; 27) genital plates, female. Scale lines 5 0.5 mm, except Fig. 23 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4900323" httpUri="https://zenodo.org/record/4900323/files/figure.png" pageId="14" pageNumber="25">Fig. 22</figureCitation>
|
||||
). Genital capsule with short and wide parameres, dorsally and ventrally fused, ring shaped, apex curved, progressively narrowed to acute apex, ventrally with a transverse strongly sclerotized wide blade with sinuate border directed backward. Aedeagus short, with a long curved bristle and tuft of yellowish setae on the dorsoapical area of the sclerotized support (
|
||||
<figureCitation id="133A2A6D5D4915272E5ED278FD4AF992" box="[609,724,1591,1610]" captionStart="Figs" captionStartId="14.[330,373,1095,1114]" captionTargetBox="[336,1117,273,1072]" captionTargetId="figure-12@14.[328,1117,272,1075]" captionTargetPageId="14" captionText="Figs. 22–27. Phyllophaga naranjina. 22) protarsal claw, male; 23) genital capsule, lateral view; 24) paramera, distal view; 25) paramera, ventrodistal view; 26) protarsal claw, female; 27) genital plates, female. Scale lines 5 0.5 mm, except Fig. 23 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4900323" httpUri="https://zenodo.org/record/4900323/files/figure.png" pageId="14" pageNumber="25">Figs. 23–25</figureCitation>
|
||||
). Tectum with two vague tumosities toward the base. Length of genital capsule from apex of parameres to border of basal piece:
|
||||
<quantity id="4CF99B0D5D4915272869D200FEC4F9A2" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="14" pageNumber="25" unit="mm" value="2.7">2.7 mm</quantity>
|
||||
. Total body length:
|
||||
<quantity id="4CF99B0D5D4915272E25D228FDECF9A2" box="[538,626,1639,1658]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="14" pageNumber="25" unit="mm" value="11.8">11.8 mm</quantity>
|
||||
. Humeral width:
|
||||
<quantity id="4CF99B0D5D4915272F27D228FCFAF9A2" box="[792,868,1639,1658]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="14" pageNumber="25" unit="mm" value="4.6">4.6 mm</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
<caption id="DF7E66605D4915272D75D008FBEAFB52" ID-DOI="http://doi.org/10.5281/zenodo.4900323" ID-Zenodo-Dep="4900323" httpUri="https://zenodo.org/record/4900323/files/figure.png" pageId="14" pageNumber="25" startId="14.[330,373,1095,1114]" targetBox="[336,1117,273,1072]" targetPageId="14">
|
||||
<paragraph id="8BBE36E85D4915272D75D008FBEAFB52" blockId="14.[307,1140,1095,1162]" pageId="14" pageNumber="25">
|
||||
<emphasis id="B975EAFA5D4915272D75D008FE5FFB82" bold="true" box="[330,449,1095,1114]" pageId="14" pageNumber="25">Figs. 22–27.</emphasis>
|
||||
<emphasis id="B975EAFA5D4915272DE6D008FD24FB82" box="[473,698,1095,1114]" italics="true" pageId="14" pageNumber="25">
|
||||
<taxonomicName id="4C014D6B5D4915272DE6D008FD29FB82" box="[473,695,1095,1114]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="25" phylum="Arthropoda" rank="species" species="naranjina">Phyllophaga naranjina</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
<emphasis id="B975EAFA5D4915272EFDD008FD47FB82" bold="true" box="[706,729,1095,1114]" pageId="14" pageNumber="25">22</emphasis>
|
||||
) protarsal claw, male;
|
||||
<emphasis id="B975EAFA5D4915272F86D008FC4EFB82" bold="true" box="[953,976,1095,1114]" pageId="14" pageNumber="25">23</emphasis>
|
||||
) genital capsule, lateral view;
|
||||
<emphasis id="B975EAFA5D4915272D8DD010FE57FBAA" bold="true" box="[434,457,1119,1138]" pageId="14" pageNumber="25">24</emphasis>
|
||||
) paramera, distal view;
|
||||
<emphasis id="B975EAFA5D4915272E8AD010FD52FBAA" bold="true" box="[693,716,1119,1138]" pageId="14" pageNumber="25">25</emphasis>
|
||||
) paramera, ventrodistal view;
|
||||
<emphasis id="B975EAFA5D4915272FC4D010FB8CFBAA" bold="true" box="[1019,1042,1119,1138]" pageId="14" pageNumber="25">26</emphasis>
|
||||
) protarsal claw, female;
|
||||
<emphasis id="B975EAFA5D4915272D87D038FE51FB52" bold="true" box="[440,463,1143,1162]" pageId="14" pageNumber="25">27</emphasis>
|
||||
) genital plates, female. Scale lines 5 0.5 mm, except Fig. 23 5 1 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BBE36E85D4815262D0DD55AFC11FDED" blockId="15.[279,1113,277,1232]" pageId="15" pageNumber="26">
|
||||
<emphasis id="B975EAFA5D4815262D0DD55AFE09FEF2" bold="true" box="[306,407,277,298]" pageId="15" pageNumber="26">
|
||||
<typeStatus id="54BA884A5D4815262D0DD55AFE0CFEF2" box="[306,402,277,298]" pageId="15" pageNumber="26" type="allotype">Allotype</typeStatus>
|
||||
.
|
||||
</emphasis>
|
||||
Female. Similar to the male except as follows: antennal club as long as the preceeding 5 segments; visible abdominal sternites II to V convex; anal plate 1.2× longer than male anal plate, convex, punctate, with 25 scattered slender setae. Protarsomeres with subapical hooklike tubercles reduced. Metafemur 1.1× wider than metafemur of male. Metatibia at apex 1.2× wider than male metatibia. Both apical spurs of metatibia articulated, wider than in male. Tarsal claws each with large tooth near middle of ventral border (
|
||||
<figureCitation id="133A2A6D5D4815262812D5FAFEABFE3D" captionStart="Figs" captionStartId="14.[330,373,1095,1114]" captionTargetBox="[336,1117,273,1072]" captionTargetId="figure-12@14.[328,1117,272,1075]" captionTargetPageId="14" captionText="Figs. 22–27. Phyllophaga naranjina. 22) protarsal claw, male; 23) genital capsule, lateral view; 24) paramera, distal view; 25) paramera, ventrodistal view; 26) protarsal claw, female; 27) genital plates, female. Scale lines 5 0.5 mm, except Fig. 23 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4900323" httpUri="https://zenodo.org/record/4900323/files/figure.png" pageId="15" pageNumber="26">Fig. 26</figureCitation>
|
||||
). Pygidium less convex. Ventral genital plates slightly sclerotized, nearly symmetrical, round with elongate apex; dorsal genital plates articulated with lateral borders of ventral plates, apices rounded with 3 setae on each side (
|
||||
<figureCitation id="133A2A6D5D4815262811D64AFEABFDED" captionStart="Figs" captionStartId="14.[330,373,1095,1114]" captionTargetBox="[336,1117,273,1072]" captionTargetId="figure-12@14.[328,1117,272,1075]" captionTargetPageId="14" captionText="Figs. 22–27. Phyllophaga naranjina. 22) protarsal claw, male; 23) genital capsule, lateral view; 24) paramera, distal view; 25) paramera, ventrodistal view; 26) protarsal claw, female; 27) genital plates, female. Scale lines 5 0.5 mm, except Fig. 23 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4900323" httpUri="https://zenodo.org/record/4900323/files/figure.png" pageId="15" pageNumber="26">Fig. 27</figureCitation>
|
||||
). Total body length:
|
||||
<quantity id="4CF99B0D5D4815262E28D66FFDE7FDED" box="[535,633,544,565]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="15" pageNumber="26" unit="mm" value="12.9">12.9 mm</quantity>
|
||||
. Humeral width:
|
||||
<quantity id="4CF99B0D5D4815262F0AD66FFC15FDED" box="[821,907,544,565]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="15" pageNumber="26" unit="mm" value="4.5">4.5 mm</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BBE36E85D4815262D0DD675FDFDFD0D" blockId="15.[279,1113,277,1232]" pageId="15" pageNumber="26">
|
||||
<emphasis id="B975EAFA5D4815262D0DD675FD8FFD97" bold="true" box="[306,529,570,591]" pageId="15" pageNumber="26">
|
||||
<typeStatus id="54BA884A5D4815262D0DD675FE06FD97" box="[306,408,570,591]" pageId="15" pageNumber="26" type="paratype">Paratype</typeStatus>
|
||||
Variation.
|
||||
</emphasis>
|
||||
Males. Similar to
|
||||
<typeStatus id="54BA884A5D4815262EEAD674FCAEFD88" box="[725,816,571,592]" pageId="15" pageNumber="26" type="holotype">holotype</typeStatus>
|
||||
except in total body length: 10.0–
|
||||
<quantity id="4CF99B0D5D4815262D6AD61AFE2BFDB2" box="[341,437,597,618]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="15" pageNumber="26" unit="mm" value="13.1">13.1 mm</quantity>
|
||||
, humeral width:
|
||||
<quantity id="4CF99B0D5D4815262E53D61AFD71FDB2" box="[620,751,597,618]" metricMagnitude="-3" metricUnit="m" metricValue="5.35" metricValueMax="5.9" metricValueMin="4.8" pageId="15" pageNumber="26" unit="mm" value="5.35" valueMax="5.9" valueMin="4.8">4.8–5.9 mm</quantity>
|
||||
; antennal club 1.3–1.5× preceeding 5 segments;
|
||||
<specimenCount id="9D07FD615D4815262DEDD63FFDDDFD5D" box="[466,579,624,645]" pageId="15" pageNumber="26" type="male">two males</specimenCount>
|
||||
with scattered short, erect setae at the sides of scutellum. Females. Similar to
|
||||
<typeStatus id="54BA884A5D4815262E5BD6C4FD26FD78" box="[612,696,651,672]" pageId="15" pageNumber="26" type="allotype">allotype</typeStatus>
|
||||
except in total body length: 11.9–13.0 mm, humeral width:
|
||||
<quantity id="4CF99B0D5D4815262E3BD6EAFD12FD62" box="[516,652,677,698]" metricMagnitude="-3" metricUnit="m" metricValue="5.05" metricValueMax="5.2" metricValueMin="4.9" pageId="15" pageNumber="26" unit="mm" value="5.05" valueMax="5.2" valueMin="4.9">4.9–5.2 mm</quantity>
|
||||
;
|
||||
<specimenCount id="9D07FD615D4815262EA3D6EAFCB0FD62" box="[668,814,677,698]" pageId="15" pageNumber="26" type="female">three females</specimenCount>
|
||||
with scattered short, erect setae at the sides of scutellum.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D4815262D0DD695FC92FC48" pageId="15" pageNumber="26" type="materials_examined">
|
||||
<paragraph id="8BBE36E85D4815262D0DD695FDE8FC82" blockId="15.[279,1113,277,1232]" pageId="15" pageNumber="26">
|
||||
<emphasis id="B975EAFA5D4815262D0DD695FE20FD37" bold="true" box="[306,446,730,751]" pageId="15" pageNumber="26">Type Series.</emphasis>
|
||||
Described from
|
||||
<specimenCount id="9D07FD615D4815262E45D694FD50FD28" box="[634,718,731,752]" pageId="15" pageNumber="26" type="male">9 males</specimenCount>
|
||||
and
|
||||
<specimenCount id="9D07FD615D4815262F36D694FCEBFD28" box="[777,885,731,752]" pageId="15" pageNumber="26" type="female">7 females</specimenCount>
|
||||
.
|
||||
<materialsCitation id="3B693CB55D4815262FBDD694FD55FCFD" accessionNumber="LN 226000" collectingDate="1998-04-06" collectionCode="IN- BIO" collectorName="A. Solis." country="Costa Rica" county="Prov." latitude="10.166667" location="Estac. de Peaje." longLatPrecision="1298" longitude="-84.36667" municipality="El Rosario de Naranjo" pageId="15" pageNumber="26" specimenCount="1" specimenCount-male="1" stateProvince="Prov. Alajuela" typeStatus="holotype">
|
||||
<typeStatus id="54BA884A5D4815262FBDD694FC7DFD28" box="[898,995,731,752]" pageId="15" pageNumber="26" type="holotype">Holotype</typeStatus>
|
||||
<specimenCount id="9D07FD615D4815262FD1D694FBBFFD28" box="[1006,1057,731,752]" pageId="15" pageNumber="26" type="male">male</specimenCount>
|
||||
(
|
||||
<collectionCode id="ED10AE2D5D481526280DD694FED7FCD2" pageId="15" pageNumber="26">INBIO</collectionCode>
|
||||
): ‘‘
|
||||
<collectingCountry id="F31676785D4815262D56D6BAFE77FCD2" box="[361,489,757,778]" name="Costa Rica" pageId="15" pageNumber="26">Costa Rica</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="49C5F80A5D4815262DC8D6BAFD0AFCD2" box="[503,660,757,778]" pageId="15" pageNumber="26">Prov. Alajuela</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="6BDAAC925D4815262E9CD6BAFC02FCD2" box="[675,924,757,778]" pageId="15" pageNumber="26">El Rosario de Naranjo</collectingMunicipality>
|
||||
,
|
||||
<location id="8EDE60335D4815262F94D6BAFBC7FCD2" LSID="urn:lsid:plazi:treatment:03A887FE5D4A15262D79D703FEC5FB08:8EDE60335D4815262F94D6BAFBC7FCD2" box="[939,1113,757,778]" country="Costa Rica" county="Prov." municipality="El Rosario de Naranjo" name="Estac. de Peaje." pageId="15" pageNumber="26" stateProvince="Prov. Alajuela">Estac. de Peaje.</location>
|
||||
LN 226000, 496700,
|
||||
<date id="FFBF10285D4815262DC2D75FFDCCFCFD" box="[509,594,784,805]" pageId="15" pageNumber="26" value="1998-04-06">
|
||||
<collectingDate id="EFFBE9C05D4815262DC2D75FFDCCFCFD" box="[509,594,784,805]" pageId="15" pageNumber="26" value="1998-04-06">6IV98</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F4533E5D4815262E61D75FFD5EFCFD" box="[606,704,784,805]" pageId="15" pageNumber="26">A. Solís.</collectorName>
|
||||
’’
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D4815262EECD75FFE0BFC98" accessionNumber="LN 226000" collectingDate="1998-04-06" collectionCode="INBIO" collectorName="A. Solis." country="Costa Rica" county="Prov." latitude="10.166667" location="Estac. de Peaje." longLatPrecision="1298" longitude="-84.36667" municipality="El Rosario de Naranjo" pageId="15" pageNumber="26" specimenCount="1" specimenCount-female="1" stateProvince="Alajuela" typeStatus="allotype">
|
||||
<typeStatus id="54BA884A5D4815262EECD75FFCB0FCFD" box="[723,814,784,805]" pageId="15" pageNumber="26" type="allotype">Allotype</typeStatus>
|
||||
<specimenCount id="9D07FD615D4815262F09D75FFC1FFCFD" box="[822,897,784,805]" pageId="15" pageNumber="26" type="female">female</specimenCount>
|
||||
: (
|
||||
<collectionCode id="ED10AE2D5D4815262FABD75FFC7EFCFD" box="[916,992,784,805]" pageId="15" pageNumber="26">INBIO</collectionCode>
|
||||
) same data as holotype
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3B693CB55D4815262D9FD764FC75FC98" accessionNumber="LN 226000" box="[416,1003,811,832]" collectingDate="1998-04-06" collectionCode="INBIO, MXAL" collectorName="A. Solis." country="Costa Rica" county="Prov." latitude="10.166667" location="Estac. de Peaje." longLatPrecision="1298" longitude="-84.36667" municipality="El Rosario de Naranjo" pageId="15" pageNumber="26" specimenCount="14" specimenCount-female="6" specimenCount-male="8" stateProvince="Alajuela" typeStatus="paratype">
|
||||
<typeStatus id="54BA884A5D4815262D9FD764FD94FC98" box="[416,522,811,832]" pageId="15" pageNumber="26" type="paratype">Paratypes</typeStatus>
|
||||
: same data as holotype (
|
||||
<specimenCount id="9D07FD615D4815262F2CD764FCF7FC98" box="[787,873,811,832]" count="8" pageId="15" pageNumber="26" type="male">8 males</specimenCount>
|
||||
,
|
||||
<specimenCount id="9D07FD615D4815262F4BD764FC7EFC98" box="[884,992,811,832]" count="6" pageId="15" pageNumber="26" type="female">6 females</specimenCount>
|
||||
).
|
||||
</materialsCitation>
|
||||
<typeStatus id="54BA884A5D4815262FCDD764FBC6FC98" box="[1010,1112,811,832]" pageId="15" pageNumber="26" type="paratype">Paratypes</typeStatus>
|
||||
deposited in
|
||||
<collectionCode id="ED10AE2D5D4815262D9DD70AFE75FC82" box="[418,491,837,858]" pageId="15" pageNumber="26">INBIO</collectionCode>
|
||||
and
|
||||
<collectionCode id="ED10AE2D5D4815262E1DD70AFDECFC82" box="[546,626,837,858]" pageId="15" pageNumber="26">MXAL</collectionCode>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BBE36E85D4815262D0DD72FFC92FC48" blockId="15.[279,1113,277,1232]" pageId="15" pageNumber="26">
|
||||
<emphasis id="B975EAFA5D4815262D0DD72FFE48FCAD" bold="true" box="[306,470,864,885]" pageId="15" pageNumber="26">Type Locality.</emphasis>
|
||||
<location id="8EDE60335D4815262DDFD72FFD4BFCAD" LSID="urn:lsid:plazi:treatment:03A887FE5D4A15262D79D703FEC5FB08:8EDE60335D4815262DDFD72FFD4BFCAD" box="[480,725,864,885]" country="Costa Rica" latitude="10.166667" longLatPrecision="1298" longitude="-84.36667" name="El Rosario de Naranjo" pageId="15" pageNumber="26" stateProvince="Alajuela">El Rosario de Naranjo</location>
|
||||
,
|
||||
<locationDeviation id="9A0C2FCA5D4815262EDDD72FFCA9FCAD" box="[738,823,864,885]" pageId="15" pageNumber="26">
|
||||
<quantity id="4CF99B0D5D4815262EDDD72FFC84FCAD" box="[738,794,864,885]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="26" unit="km" value="3.0">3 km</quantity>
|
||||
N
|
||||
</locationDeviation>
|
||||
<location id="8EDE60335D4815262F7FD72FFC09FCAD" LSID="urn:lsid:plazi:treatment:03A887FE5D4A15262D79D703FEC5FB08:8EDE60335D4815262F7FD72FFC09FCAD" box="[832,919,864,885]" country="Costa Rica" latitude="10.166667" longLatPrecision="1298" longitude="-84.36667" name="Naranjo" pageId="15" pageNumber="26" stateProvince="Alajuela">Naranjo</location>
|
||||
, province of
|
||||
<collectingRegion id="49C5F80A5D4815262816D72FFED3FC48" country="Costa Rica" name="Alajuela" pageId="15" pageNumber="26">Alajuela</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="F31676785D4815262D65D734FE50FC48" box="[346,462,891,912]" name="Costa Rica" pageId="15" pageNumber="26">Costa Rica</collectingCountry>
|
||||
(approx.
|
||||
<geoCoordinate id="EE35502F5D4815262E07D734FD0CFC48" box="[568,658,891,912]" degrees="10" direction="north" minutes="10" orientation="latitude" pageId="15" pageNumber="26" precision="925" value="10.166667">10°10'N</geoCoordinate>
|
||||
;
|
||||
<geoCoordinate id="EE35502F5D4815262EA0D734FC9EFC48" box="[671,768,891,912]" degrees="84" direction="west" minutes="22" orientation="longitude" pageId="15" pageNumber="26" precision="925" value="-84.36667">84°22'W</geoCoordinate>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D4815262D0DD7DAFD94FC38" pageId="15" pageNumber="26" type="biology_ecology">
|
||||
<paragraph id="8BBE36E85D4815262D0DD7DAFD94FC38" blockId="15.[279,1113,277,1232]" pageId="15" pageNumber="26">
|
||||
<emphasis id="B975EAFA5D4815262D0DD7DAFE7DFC72" bold="true" box="[306,483,917,938]" pageId="15" pageNumber="26">Biological Data.</emphasis>
|
||||
All males and females of
|
||||
<taxonomicName id="4C014D6B5D4815262EC8D7D9FCE5FC73" box="[759,891,918,939]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="26" phylum="Arthropoda" rank="species" species="naranjina" status="sp. nov.">
|
||||
<emphasis id="B975EAFA5D4815262EC8D7D9FCE5FC73" box="[759,891,918,939]" italics="true" pageId="15" pageNumber="26">P. naranjina</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A24657815D4815262FBDD7DAFC24FC72" box="[898,954,917,938]" pageId="15" pageNumber="26" rank="species">n. sp.</taxonomicNameLabel>
|
||||
were collected at lights at the border of coffee plantations located at
|
||||
<quantity id="4CF99B0D5D4815262F45D7FFFC4BFC1D" box="[890,981,944,965]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="15" pageNumber="26" unit="m" value="1050.0">1,050 m</quantity>
|
||||
of altitude. Phenology: April (16).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D4815262D0DD7AAFBD4FB42" pageId="15" pageNumber="26" type="discussion">
|
||||
<paragraph id="8BBE36E85D4815262D0DD7AAFBD4FB42" blockId="15.[279,1113,277,1232]" pageId="15" pageNumber="26">
|
||||
<emphasis id="B975EAFA5D4815262D0DD7AAFE00FC22" bold="true" box="[306,414,997,1018]" pageId="15" pageNumber="26">Remarks.</emphasis>
|
||||
<taxonomicName id="4C014D6B5D4815262D97D7A9FCA1FC23" box="[424,831,997,1019]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="26" phylum="Arthropoda" rank="species" species="naranjina" status="sp. nov." subGenus="Phyllophaga">
|
||||
<emphasis id="B975EAFA5D4815262D97D7A9FDB0FC23" box="[424,558,998,1019]" italics="true" pageId="15" pageNumber="26">Phyllophaga</emphasis>
|
||||
(
|
||||
<emphasis id="B975EAFA5D4815262E7FD7A9FD5BFC23" box="[576,709,998,1019]" italics="true" pageId="15" pageNumber="26">Phyllophaga</emphasis>
|
||||
)
|
||||
<emphasis id="B975EAFA5D4815262EE7D7A9FCA1FC23" box="[728,831,998,1019]" italics="true" pageId="15" pageNumber="26">naranjina</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A24657815D4815262F77D7AAFC1AFC22" box="[840,900,997,1018]" pageId="15" pageNumber="26" rank="species">n. sp.</taxonomicNameLabel>
|
||||
does not belong to any known species group (
|
||||
<emphasis id="B975EAFA5D4815262E0DD04FFDF2FBCD" box="[562,620,1024,1045]" italics="true" pageId="15" pageNumber="26">sensu</emphasis>
|
||||
<bibRefCitation id="EF904B195D4815262E4CD04FFD64FBCD" author="Moron" box="[627,762,1024,1045]" firstAuthor="Moron" pageId="15" pageNumber="26" refId="ref11224" refString="Moron, M. A. 1986. El genero Phyllophaga en Mexico. Morfologia, distribucion y sistematica supraespecifica (Insecta: Coleoptera). Publ. 20, Instituto de Ecologia, Mexico, D. F. 341 pp." type="book" year="1986">Morón 1986</bibRefCitation>
|
||||
), but the nine segmented antenna may related it with the ‘‘
|
||||
<emphasis id="B975EAFA5D4815262E1FD054FD1FFBE8" box="[544,641,1051,1072]" italics="true" pageId="15" pageNumber="26">porodera</emphasis>
|
||||
’’ group, and the shape of tarsal claws, male anal plate and protarsal spurs are similar to some species in the ‘‘
|
||||
<emphasis id="B975EAFA5D4815262FEDD079FBD6FB93" box="[978,1096,1078,1099]" italics="true" pageId="15" pageNumber="26">anodentata</emphasis>
|
||||
’’ group. The very short clypeus, number of antennal segments, scattered granules near the middle line of the sternites, shape and details of the male genital capsule, and small body size will aid in the recognition of this new species.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D4815262D0DD0EFFEC5FB08" pageId="15" pageNumber="26" type="etymology">
|
||||
<paragraph id="8BBE36E85D4815262D0DD0EFFEC5FB08" blockId="15.[279,1113,277,1232]" pageId="15" pageNumber="26">
|
||||
<emphasis id="B975EAFA5D4815262D0DD0EFFE2FFB6D" bold="true" box="[306,433,1184,1205]" pageId="15" pageNumber="26">Etymology.</emphasis>
|
||||
Derived from the name of
|
||||
<typeStatus id="54BA884A5D4815262ED8D0EFFC8AFB6D" box="[743,788,1184,1205]" pageId="15" pageNumber="26">type</typeStatus>
|
||||
locality ‘‘El Rosario de Naranjo.’’
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
352
data/03/A8/87/03A887FE5D4C15242D0AD03CFC1BFCFD.xml
Normal file
352
data/03/A8/87/03A887FE5D4C15242D0AD03CFC1BFCFD.xml
Normal file
|
|
@ -0,0 +1,352 @@
|
|||
<document id="9FD23803774666A14A86DFAE54E656D2" ID-DOI="10.1649/0010-065x(2001)055[0011:snsops]2.0.co;2" ID-ISSN="1938-4394" ID-Zenodo-Dep="4900311" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1622827801739" checkinUser="carolina" docAuthor="Morón, Miguel Angel & Solís, Angel" docDate="2001" docId="03A887FE5D4C15242D0AD03CFC1BFCFD" docLanguage="en" docName="TheColeopBull.55.1.11-29.pdf.imf" docOrigin="The Coleopterists Bulletin 55 (1)" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Phyllophaga (Phyllophaga) talamancana Moron and Solis, new species" docType="treatment" docVersion="1" lastPageNumber="24" masterDocId="FF91FF865D4715292C3FD44FFF9EFFD8" masterDocTitle="Seven New Species Of Phyllophaga (S. Str.) Harris From Costa Rica (Coleoptera: Melolonthidae: Melolonthinae)" masterLastPageNumber="29" masterPageNumber="11" pageNumber="22" updateTime="1730139982923" updateUser="juliana" zenodo-license-document="CLOSED">
|
||||
<mods:mods id="559ACA7FA70FAFD42EAA8C8F87DF3A45" xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo id="8ACF0D118B15F00F9510DB63A9A12E35">
|
||||
<mods:title id="2213881540FEC2B7EA8AD12F74BA8B19">Seven New Species Of Phyllophaga (S. Str.) Harris From Costa Rica (Coleoptera: Melolonthidae: Melolonthinae)</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name id="14F5994351B5E8B8B0AC4182A2F08F23" type="personal">
|
||||
<mods:role id="9EC9F5E298BAC700E5174A0829C9197B">
|
||||
<mods:roleTerm id="A284940A7BB9E5371F2849CE0277D281">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="7B2234BA5603B150C224280DC0707CB1">Morón, Miguel Angel</mods:namePart>
|
||||
<mods:affiliation id="0D617EC914BD0CAD7D6998ADA7222E3A">Departamento de Entomología Instituto de Ecología, A. C. (sector SEPCONACYT) Apartado Postal 63 Xalapa, Veracruz 91000, MÉXICO</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name id="40E4D8EC01EE30005730EED4C48B7F59" type="personal">
|
||||
<mods:role id="FDF4F75AA5B2432DA9DCF825D3944E06">
|
||||
<mods:roleTerm id="6C875FC6763D9EC9CEF94DA34E8C9393">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="7837FCBEC8810CF5AE5A81362723ED35">Solís, Angel</mods:namePart>
|
||||
<mods:affiliation id="8C4737AA148D87C20AFA91610EA2E4AE">Departamento de Entomología Instituto Nacional de Biodiversidad (INBio) Apartado Postal 22 3100 Santo Domingo de Heredia, COSTA RICA</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:typeOfResource id="11FFF6E92A06E35C328A169D166FBA7F">text</mods:typeOfResource>
|
||||
<mods:relatedItem id="7ADE4BD762558A93A7E1B067B5F46F5C" type="host">
|
||||
<mods:titleInfo id="877792B93757F4967BA851F9F0890ED6">
|
||||
<mods:title id="0AFC44CDE6BDC2F349F831386AF7C963">The Coleopterists Bulletin</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part id="A2CBC30DF363872B3ED04453BDAE4041">
|
||||
<mods:date id="B871EE556745076916866D1A35D181DD">2001</mods:date>
|
||||
<mods:detail id="AC0D7800B1E67A3CCAB1AB0C849999B6" type="pubDate">
|
||||
<mods:number id="34037430E04299507F1AEF9A91D442EF">2001-03-01</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="D64DDD4BB4AE9B8C4DBA7A2EC36340F8" type="volume">
|
||||
<mods:number id="DF05666CD6311F1EC3D5F3E8910C1E0E">55</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="5EDE30C9E83BCC463DB7DD2028FE937A" type="issue">
|
||||
<mods:number id="8B9AB655DCD701F5D1A241243EE0ABC5">1</mods:number>
|
||||
</mods:detail>
|
||||
<mods:extent id="53D8821C3CA62D9125E90EC1DC41C34D" unit="page">
|
||||
<mods:start id="16A81D157141876F6161E6872EC8E7FC">11</mods:start>
|
||||
<mods:end id="36CAEC7D57E9CDDC6087A9E3DD84BD01">29</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:classification id="B08C9CEC838940B67D0D98726419F3C9">journal article</mods:classification>
|
||||
<mods:identifier id="F0A11BAB9E768FFC80BE532E4FD89FA0" type="DOI">10.1649/0010-065x(2001)055[0011:snsops]2.0.co;2</mods:identifier>
|
||||
<mods:identifier id="E9740E164D7EA7D9CFCFAF5FFFDE25BE" type="ISSN">1938-4394</mods:identifier>
|
||||
<mods:identifier id="CBC02CFCCF2044FF7221DD63B6CE4F99" type="Zenodo-Dep">4900311</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="03A887FE5D4C15242D0AD03CFC1BFCFD" LSID="urn:lsid:plazi:treatment:03A887FE5D4C15242D0AD03CFC1BFCFD" httpUri="http://treatment.plazi.org/id/03A887FE5D4C15242D0AD03CFC1BFCFD" lastPageId="13" lastPageNumber="24" pageId="11" pageNumber="22">
|
||||
<subSubSection id="C31B65635D4C15222D0AD03CFBA4FB5F" box="[309,1082,1138,1160]" pageId="11" pageNumber="22" type="nomenclature">
|
||||
<paragraph id="8BBE36E85D4C15222D0AD03CFBA4FB5F" blockId="11.[309,1082,1138,1186]" box="[309,1082,1138,1160]" pageId="11" pageNumber="22">
|
||||
<taxonomicName id="4C014D6B5D4C15222D0AD03CFC35FB50" authority="Moron and Solis" authorityName="Moron and Solis" box="[309,939,1139,1160]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="species" species="talamancana" status="sp. nov." subGenus="Phyllophaga">
|
||||
<emphasis id="B975EAFA5D4C15222D0AD03CFE25FB50" box="[309,443,1139,1160]" italics="true" pageId="11" pageNumber="22">Phyllophaga</emphasis>
|
||||
(
|
||||
<emphasis id="B975EAFA5D4C15222DF3D03CFDCFFB50" box="[460,593,1139,1160]" italics="true" pageId="11" pageNumber="22">Phyllophaga</emphasis>
|
||||
)
|
||||
<emphasis id="B975EAFA5D4C15222E5CD03CFD75FB50" box="[611,747,1139,1160]" italics="true" pageId="11" pageNumber="22">talamancana</emphasis>
|
||||
Morón and Solís
|
||||
</taxonomicName>
|
||||
,
|
||||
<emphasis id="B975EAFA5D4C15222F88D03DFBA4FB5F" bold="true" box="[951,1082,1138,1159]" pageId="11" pageNumber="22">
|
||||
<taxonomicNameLabel id="A24657815D4C15222F88D03DFBA4FB5F" box="[951,1082,1138,1159]" pageId="11" pageNumber="22" rank="species">new species</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D4C15252E46D0C2FD8EFAB0" lastPageId="12" lastPageNumber="23" pageId="11" pageNumber="22" type="description">
|
||||
<paragraph id="8BBE36E85D4C15222E46D0C2FD69FB7A" blockId="11.[309,1082,1138,1186]" box="[633,759,1165,1186]" pageId="11" pageNumber="22">
|
||||
<figureCitation id="133A2A6D5D4C15222E46D0C2FD69FB7A" box="[633,759,1165,1186]" captionStart="Figs" captionStartId="11.[303,346,988,1007]" captionTargetBox="[305,1093,273,963]" captionTargetId="figure-12@11.[297,1094,272,968]" captionTargetPageId="11" captionText="Figs. 18–21. Phyllophaga talamancana. 18) protarsal claw, male; 19) genital capsule, lateral view; 20) paramera, distal view; 21) genital plates, female. Scale lines 5 1 mm, except Fig. 18 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900321" httpUri="https://zenodo.org/record/4900321/files/figure.png" pageId="11" pageNumber="22">Figs. 18–21</figureCitation>
|
||||
</paragraph>
|
||||
<paragraph id="8BBE36E85D4C15252D10D0F8FCF6FC37" blockId="11.[279,1113,1207,1658]" lastBlockId="12.[307,1140,277,1007]" lastPageId="12" lastPageNumber="23" pageId="11" pageNumber="22">
|
||||
<emphasis id="B975EAFA5D4C15222D10D0F8FE11FB12" bold="true" box="[303,399,1207,1226]" pageId="11" pageNumber="22">
|
||||
<typeStatus id="54BA884A5D4C15222D10D0F8FE15FB12" box="[303,395,1207,1226]" pageId="11" pageNumber="22" type="holotype">Holotype</typeStatus>
|
||||
.
|
||||
</emphasis>
|
||||
Male. Clypeus, frons and pronotum shiny dark reddish brown; elytra light brown with iridescent luster, head and pronotum with abundant, erect, long setae; elytra with abundant, erect, long setae around scutellum and toward the apex, near suture, and vestiture of uniformly distributed short setae on rest of surface; mouthparts, sterna, pygidium and legs shiny reddish brown. Clypeus 4.2× wider than long, anterior border very scarcely sinuate, with poorly elevated margin, surface shallowly concave, with scarce irregularly distributed, deep, round shaped punctures, each with short setae. Frontoclypeal suture sinuate and deeply impressed. Frons 2.3× wider than long, convex, with reduced number of coarse wide punctures, with long slender setae at sides and on disk. Antenna 10segmented, with 3segmented club, lamellae of 8th to 10th segments 1.1× longer than length of preceeding 6 segments combined. Frons 4.0× wider than dorsal diameter of eye. Eye canthus long and wide, with 6–7 setae. Labrum bilobed, widely sinuate, with scattered slender setae. Mentum slightly convex, with 4 punctures in a transverse row, and long slender setae near the borders; anterior border briefly sinuate. Pronotum 1.4× wider than long and 2.7× wider than frons. Pronotal disk shiny, with deep, round punctures irregularly separated from one another by 1–7 diameters; lateral borders widely angled, lateral marginal bead crenulate, with regularly located, long, slender setae; basal bead indicated by punctures mainly toward the sides; anterior angles acute, clearly prominent; posterior angles obtuse, slighly prominent. Scutellum 1.5× wider than long, with some deep punctures each with short setae. Elytron 3.0× longer than wider, shiny, densely punctate; epipleural border very narrow, extended along the complete margin, provided with setae along first half, setae diminishing in size anteroposteriorly from long to minute; humeral callus rounded, prominent; apical callus rounded. Metathoracic wings completely developed. Propygidium shiny, densely punctate, setose. Pygidium moderately convex, shiny, shallowly punctate rugose with scattered erect setae; apical margin with 19 long, slender setae; basal margin effaced medially. Pterosternon with long, dense, yellowish setae. Visible abdominal sternites II–IV convex, smooth and glabrous near midline; sternite V convex, shiny, densely punctate with short setae at midline and sparse punctures with mixed long and short setae toward sides; anal plate narrowed, with a transverse shallow concavity and longitudinal furrow at midline, anterior and posterior borders slightly elevated, near the posterior border with 23 erect setae. Protibia slightly shorter than protarsus (0.9:1), with external border tridentate, basal tooth short, preapical spur acute, nearly straight, as long as 2nd protarsomere. Mesotibia with one oblique, sharp, setiferous carina on external side; upper apical spur straight, narrow, and 1.1× longer than lower spur. Metatibia 0.9× shorter than metatarsus, with one oblique, sharp, setiferous carina on external side; upper apical spur articulated, curved, with apex rounded, 1.1× longer than basal metatarsomere, and 1.4× longer than lower spur; lower apical spur articulated with tibial border, with rounded apex. Tarsomeres semicylindrical, not much elongate, with enlarged apex, some setae apically and ventrally one longitudinal carina with a line of short setae at each side. Protaromeres 1– 4 with poorly developed subapical tubercles. Tarsal claws symmetrical, similar on all legs, each with large acute tooth at middle of ventral border and with finely serrate border toward the base (
|
||||
<figureCitation id="133A2A6D5D4B15252E1AD706FDF0FC84" box="[549,622,841,860]" captionStart="Figs" captionStartId="11.[303,346,988,1007]" captionTargetBox="[305,1093,273,963]" captionTargetId="figure-12@11.[297,1094,272,968]" captionTargetPageId="11" captionText="Figs. 18–21. Phyllophaga talamancana. 18) protarsal claw, male; 19) genital capsule, lateral view; 20) paramera, distal view; 21) genital plates, female. Scale lines 5 1 mm, except Fig. 18 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900321" httpUri="https://zenodo.org/record/4900321/files/figure.png" pageId="12" pageNumber="23">Fig. 18</figureCitation>
|
||||
). Genital capsule with short parameres, dorsally and ventrally fused, ringshaped, apex widely notched, ventrally with preapical minute, toothlike projections. Aedeagus long, with large preapical patches of spinules and strong sclerotized support with dorsal brush of long setae directed to the apex; basally with a pair of dorsal rounded, membranous saclike structures (
|
||||
<figureCitation id="133A2A6D5D4B15252F79D7E4FC24FC66" box="[838,954,939,958]" captionStart="Figs" captionStartId="11.[303,346,988,1007]" captionTargetBox="[305,1093,273,963]" captionTargetId="figure-12@11.[297,1094,272,968]" captionTargetPageId="11" captionText="Figs. 18–21. Phyllophaga talamancana. 18) protarsal claw, male; 19) genital capsule, lateral view; 20) paramera, distal view; 21) genital plates, female. Scale lines 5 1 mm, except Fig. 18 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900321" httpUri="https://zenodo.org/record/4900321/files/figure.png" pageId="12" pageNumber="23">Figs. 19–20</figureCitation>
|
||||
). Tectum uniformly convex. Length of genital capsule from apex of parameres to border of basal piece:
|
||||
<quantity id="4CF99B0D5D4B15252858D78CFEC4FC37" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="23" unit="mm" value="5.0">5 mm</quantity>
|
||||
. Total body length:
|
||||
<quantity id="4CF99B0D5D4B15252E25D793FDECFC37" box="[538,626,988,1007]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="12" pageNumber="23" unit="mm" value="19.1">19.1 mm</quantity>
|
||||
. Humeral width: 8.0 mm.
|
||||
</paragraph>
|
||||
<paragraph id="8BBE36E85D4B15252D72D043FCA4FB23" blockId="12.[307,1140,1036,1658]" pageId="12" pageNumber="23">
|
||||
<emphasis id="B975EAFA5D4B15252D72D043FE2CFBF9" bold="true" box="[333,434,1036,1057]" pageId="12" pageNumber="23">
|
||||
<typeStatus id="54BA884A5D4B15252D72D043FE33FBF9" box="[333,429,1036,1057]" pageId="12" pageNumber="23" type="allotype">Allotype</typeStatus>
|
||||
.
|
||||
</emphasis>
|
||||
Female. Similar to the male except as follows: visible abdominal sternites II to V less convex; anal plate 1.5× longer than male anal plate, convex, punctate, with 60 scattered slender setae. Metafemur 1.3× wider than femur of male. Metatibia at the apex 1.4× wider than male metatibia. Both apical spurs of metatibia articulated, wider than in the male. Pygidium less convex. Ventral genital plates moderately sclerotized, nearly symmetrical, ovate; dorsal plates narrowed, curved, with scattered setae at apex and distal border, apparently articulated basolaterally with ventral plates (
|
||||
<figureCitation id="133A2A6D5D4B15252FE4D084FBB4FB38" box="[987,1066,1227,1248]" captionStart="Figs" captionStartId="11.[303,346,988,1007]" captionTargetBox="[305,1093,273,963]" captionTargetId="figure-12@11.[297,1094,272,968]" captionTargetPageId="11" captionText="Figs. 18–21. Phyllophaga talamancana. 18) protarsal claw, male; 19) genital capsule, lateral view; 20) paramera, distal view; 21) genital plates, female. Scale lines 5 1 mm, except Fig. 18 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900321" httpUri="https://zenodo.org/record/4900321/files/figure.png" pageId="12" pageNumber="23">Fig. 21</figureCitation>
|
||||
). Total body length:
|
||||
<quantity id="4CF99B0D5D4B15252DFDD0A9FDBAFB23" box="[450,548,1254,1275]" metricMagnitude="-2" metricUnit="m" metricValue="1.93" pageId="12" pageNumber="23" unit="mm" value="19.3">19.3 mm</quantity>
|
||||
. Humeral width:
|
||||
<quantity id="4CF99B0D5D4B15252EDFD0A9FCA8FB23" box="[736,822,1254,1275]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="12" pageNumber="23" unit="mm" value="8.3">8.3 mm</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BBE36E85D4B15252D72D14EFD8EFAB0" blockId="12.[307,1140,1036,1658]" pageId="12" pageNumber="23">
|
||||
<emphasis id="B975EAFA5D4B15252D72D14EFDB3FACE" bold="true" box="[333,557,1281,1302]" pageId="12" pageNumber="23">
|
||||
<typeStatus id="54BA884A5D4B15252D72D14EFE2DFACE" box="[333,435,1281,1302]" pageId="12" pageNumber="23" type="paratype">Paratype</typeStatus>
|
||||
Variation.
|
||||
</emphasis>
|
||||
Male. Similar to
|
||||
<typeStatus id="54BA884A5D4B15252ED4D14EFCD8FACE" box="[747,838,1281,1302]" pageId="12" pageNumber="23" type="holotype">holotype</typeStatus>
|
||||
except in total body length:
|
||||
<quantity id="4CF99B0D5D4B15252D0CD153FE0AFAE9" box="[307,404,1308,1329]" metricMagnitude="-2" metricUnit="m" metricValue="1.8199999999999998" pageId="12" pageNumber="23" unit="mm" value="18.2">18.2 mm</quantity>
|
||||
, humeral width: 8.0 mm. Female. Similar to
|
||||
<typeStatus id="54BA884A5D4B15252F41D153FC4CFAE9" box="[894,978,1308,1329]" pageId="12" pageNumber="23" type="allotype">allotype</typeStatus>
|
||||
except in total body length:
|
||||
<quantity id="4CF99B0D5D4B15252DF6D178FDB0FA94" box="[457,558,1335,1356]" metricMagnitude="-2" metricUnit="m" metricValue="1.8699999999999999" pageId="12" pageNumber="23" unit="mm" value="18.7">18.7 mm</quantity>
|
||||
, humeral width: 8.0 mm, elytra of some specimens darker than
|
||||
<typeStatus id="54BA884A5D4B15252D8BD11CFD92FAB0" box="[436,524,1363,1384]" pageId="12" pageNumber="23" type="allotype">allotype</typeStatus>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D4B15242D72D121FDFAFEA2" lastPageId="13" lastPageNumber="24" pageId="12" pageNumber="23" type="materials_examined">
|
||||
<paragraph id="8BBE36E85D4B15242D72D121FCAEFE9D" blockId="12.[307,1140,1036,1658]" lastBlockId="13.[279,1113,277,805]" lastPageId="13" lastPageNumber="24" pageId="12" pageNumber="23">
|
||||
<emphasis id="B975EAFA5D4B15252D72D121FE44FA5B" bold="true" box="[333,474,1390,1411]" pageId="12" pageNumber="23">Type Series.</emphasis>
|
||||
<location id="8EDE60335D4B15252DDCD121FDD3FA5B" LSID="urn:lsid:plazi:treatment:03A887FE5D4C15242D0AD03CFC1BFCFD:8EDE60335D4B15252DDCD121FDD3FA5B" box="[483,589,1390,1411]" name="Described" pageId="12" pageNumber="23">Described</location>
|
||||
from
|
||||
<specimenCount id="9D07FD615D4B15252EAAD121FD77FA5B" box="[661,745,1390,1411]" pageId="12" pageNumber="23" type="male">4 males</specimenCount>
|
||||
and
|
||||
<specimenCount id="9D07FD615D4B15252F1BD121FC0EFA5B" box="[804,912,1390,1411]" pageId="12" pageNumber="23" type="female">4 females</specimenCount>
|
||||
.
|
||||
<materialsCitation id="3B693CB55D4B15252FA2D121FC15FA62" collectingDate="1996-07-15" collectionCode="IN- BIO" collectorName="M. Moraga." country="Costa Rica" elevation="600" location="Cerro Pittier" municipality="Sendero" pageId="12" pageNumber="23" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="holotype">
|
||||
<typeStatus id="54BA884A5D4B15252FA2D121FC60FA5B" box="[925,1022,1390,1411]" pageId="12" pageNumber="23" type="holotype">Holotype</typeStatus>
|
||||
<specimenCount id="9D07FD615D4B15252836D121FBA2FA5B" box="[1033,1084,1390,1411]" pageId="12" pageNumber="23" type="male">male</specimenCount>
|
||||
(
|
||||
<collectionCode id="ED10AE2D5D4B15252872D121FEFBFA47" pageId="12" pageNumber="23">INBIO</collectionCode>
|
||||
): ‘‘
|
||||
<collectingCountry id="F31676785D4B15252DBED1C5FDB8FA47" box="[385,550,1418,1439]" name="Costa Rica" pageId="12" pageNumber="23">COSTA RICA</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49C5F80A5D4B15252E0FD1C5FD3BFA47" box="[560,677,1418,1439]" country="Costa Rica" name="Puntarenas" pageId="12" pageNumber="23">Puntarenas</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="6BDAAC925D4B15252E8ED1C5FC99FA47" box="[689,775,1418,1439]" pageId="12" pageNumber="23">Sendero</collectingMunicipality>
|
||||
a
|
||||
<location id="8EDE60335D4B15252F1DD1C5FC38FA47" LSID="urn:lsid:plazi:treatment:03A887FE5D4C15242D0AD03CFC1BFCFD:8EDE60335D4B15252F1DD1C5FC38FA47" box="[802,934,1418,1439]" country="Costa Rica" municipality="Sendero" name="Cerro Pittier" pageId="12" pageNumber="23" stateProvince="Puntarenas">Cerro Pittier</location>
|
||||
,
|
||||
<quantity id="4CF99B0D5D4B15252F8FD1C5FC6DFA47" box="[944,1011,1418,1439]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="23" unit="m" value="600.0">
|
||||
<elevation id="002CD1DB5D4B15252F8FD1C5FC6DFA47" box="[944,1011,1418,1439]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="23" unit="m" value="600.0">600 m</elevation>
|
||||
</quantity>
|
||||
al Noroeste de la
|
||||
<location id="8EDE60335D4B15252D50D1EAFD8DFA62" LSID="urn:lsid:plazi:treatment:03A887FE5D4C15242D0AD03CFC1BFCFD:8EDE60335D4B15252D50D1EAFD8DFA62" box="[367,531,1445,1466]" country="Costa Rica" municipality="Sendero" name="Estacion Pittier" pageId="12" pageNumber="23" stateProvince="Puntarenas">Estación Pittier</location>
|
||||
, 1,750 m,
|
||||
<date id="FFBF10285D4B15252EBAD1EAFD6FFA62" box="[645,753,1445,1466]" pageId="12" pageNumber="23" value="1996-07-15">
|
||||
<collectingDate id="EFFBE9C05D4B15252EBAD1EAFD6FFA62" box="[645,753,1445,1466]" pageId="12" pageNumber="23" value="1996-07-15">15VII96</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F4533E5D4B15252EC1D1EAFC1CFA62" box="[766,898,1445,1466]" pageId="12" pageNumber="23">M. Moraga.</collectorName>
|
||||
’’
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D4B15252FACD1EAFC28FA29" collectingDate="1994-10" collectionCode="IN- BIO" collectorName="Z. Fuentes" country="Costa Rica" elevation="1250" latitude="1.15" location="Sector Altamira" longLatPrecision="78" longitude="-1.35" municipality="Buenos Aires" pageId="12" pageNumber="23" specimenCount="1" stateProvince="Puntarenas" typeStatus="allotype">
|
||||
<typeStatus id="54BA884A5D4B15252FACD1EAFC70FA62" box="[915,1006,1445,1466]" pageId="12" pageNumber="23" type="allotype">Allotype</typeStatus>
|
||||
<specimenCount id="9D07FD615D4B15252FC8D1EAFBA0FA62" box="[1015,1086,1445,1466]" pageId="12" pageNumber="23" type="female">female</specimenCount>
|
||||
(
|
||||
<collectionCode id="ED10AE2D5D4B15252872D1EAFEFBFA0E" pageId="12" pageNumber="23">INBIO</collectionCode>
|
||||
): ‘‘
|
||||
<collectingCountry id="F31676785D4B15252DBDD18EFDB6FA0E" box="[386,552,1473,1494]" name="Costa Rica" pageId="12" pageNumber="23">COSTA RICA</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49C5F80A5D4B15252E0CD18EFD36FA0E" box="[563,680,1473,1494]" country="Costa Rica" name="Puntarenas" pageId="12" pageNumber="23">Puntarenas</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="6BDAAC925D4B15252E8BD18EFCD8FA0E" box="[692,838,1473,1494]" pageId="12" pageNumber="23">Buenos Aires</collectingMunicipality>
|
||||
,
|
||||
<location id="8EDE60335D4B15252F6DD18EFC63FA0E" LSID="urn:lsid:plazi:treatment:03A887FE5D4C15242D0AD03CFC1BFCFD:8EDE60335D4B15252F6DD18EFC63FA0E" box="[850,1021,1473,1494]" country="Costa Rica" latitude="1.15" longLatPrecision="78" longitude="-1.35" municipality="Buenos Aires" name="Sector Altamira" pageId="12" pageNumber="23" stateProvince="Puntarenas">Sector Altamira</location>
|
||||
,
|
||||
<locationDeviation id="9A0C2FCA5D4B15252836D18EFEF7FA29" pageId="12" pageNumber="23">1 km Suroeste</locationDeviation>
|
||||
del
|
||||
<location id="8EDE60335D4B15252DA8D193FDB6FA29" LSID="urn:lsid:plazi:treatment:03A887FE5D4C15242D0AD03CFC1BFCFD:8EDE60335D4B15252DA8D193FDB6FA29" box="[407,552,1500,1521]" country="Costa Rica" latitude="1.15" longLatPrecision="78" longitude="-1.35" municipality="Buenos Aires" name="Cerro Biolley" pageId="12" pageNumber="23" stateProvince="Puntarenas">Cerro Biolley</location>
|
||||
,
|
||||
<quantity id="4CF99B0D5D4B15252E0CD193FD48FA29" box="[563,726,1500,1521]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.35" metricValueMin="1.15" pageId="12" pageNumber="23" unit="m" value="1250.0" valueMax="1350.0" valueMin="1150.0">
|
||||
<elevation id="002CD1DB5D4B15252E0CD193FD48FA29" box="[563,726,1500,1521]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.35" metricValueMin="1.15" pageId="12" pageNumber="23" unit="m" value="1250.0" valueMax="1350.0" valueMin="1150.0">1,150–1,350 m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<date id="FFBF10285D4B15252EDED193FCADFA29" box="[737,819,1500,1521]" pageId="12" pageNumber="23" value="1994-10">
|
||||
<collectingDate id="EFFBE9C05D4B15252EDED193FCADFA29" box="[737,819,1500,1521]" pageId="12" pageNumber="23" value="1994-10">X1994</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F4533E5D4B15252F01D193FC28FA29" box="[830,950,1500,1521]" pageId="12" pageNumber="23">Z. Fuentes</collectorName>
|
||||
</materialsCitation>
|
||||
.’’
|
||||
<materialsCitation id="3B693CB55D4B15252FF5D193FEEBF9F0" collectingDate="1994-11" collectionCode="INBIO, MXAL" collectorName="M. Segura'" country="Costa Rica" county="Sendero Altamira" elevation="1375" latitude="1.15" location="Refugio Nacional de Fauna" longLatPrecision="78" longitude="-1.35" municipality="Estacion Pittier" pageId="12" pageNumber="23" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
|
||||
<typeStatus id="54BA884A5D4B15252FF5D193FBAAFA29" box="[970,1076,1500,1521]" pageId="12" pageNumber="23" type="paratype">Paratypes</typeStatus>
|
||||
: same data as allotype except ‘‘
|
||||
<quantity id="4CF99B0D5D4B15252E7DD1B7FD70F9D5" box="[578,750,1528,1549]" metricMagnitude="3" metricUnit="m" metricValue="1.375" metricValueMax="1.45" metricValueMin="1.3" pageId="12" pageNumber="23" unit="m" value="1375.0" valueMax="1450.0" valueMin="1300.0">
|
||||
<elevation id="002CD1DB5D4B15252E7DD1B7FD70F9D5" box="[578,750,1528,1549]" metricMagnitude="3" metricUnit="m" metricValue="1.375" metricValueMax="1.45" metricValueMin="1.3" pageId="12" pageNumber="23" unit="m" value="1375.0" valueMax="1450.0" valueMin="1300.0">1,300–1,450 m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<date id="FFBF10285D4B15252EC2D1B7FCA0F9D5" box="[765,830,1528,1549]" pageId="12" pageNumber="23" value="1994-11">
|
||||
<collectingDate id="EFFBE9C05D4B15252EC2D1B7FCA0F9D5" box="[765,830,1528,1549]" pageId="12" pageNumber="23" value="1994-11">XI94</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F4533E5D4B15252F72D1B7FC57F9D5" box="[845,969,1528,1549]" pageId="12" pageNumber="23">M. Segura’</collectorName>
|
||||
’ (
|
||||
<specimenCount id="9D07FD615D4B15252FDCD1B7FBB0F9D5" box="[995,1070,1528,1549]" count="1" pageId="12" pageNumber="23" type="male">1 male</specimenCount>
|
||||
,
|
||||
<specimenCount id="9D07FD615D4B15252801D1B7FEF4F9F0" count="1" pageId="12" pageNumber="23" type="female">1 female</specimenCount>
|
||||
);
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D4B15252DBCD25CFC3CF99B" collectingDate="1995-10-04" collectionCode="INBIO, MXAL" collectorName="M. Moraga" country="Costa Rica" county="Sendero Altamira" elevation="1720" latitude="1.15" location="Estacion Pittier" longLatPrecision="78" longitude="-1.35" municipality="Estacion Pittier" pageId="12" pageNumber="23" specimenCode="INBIO CR 1002 468647" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
|
||||
<collectingRegion id="49C5F80A5D4B15252DBCD25CFE66F9F0" box="[387,504,1555,1576]" country="Costa Rica" name="Puntarenas" pageId="12" pageNumber="23">Puntarenas</collectingRegion>
|
||||
,
|
||||
<locationDeviation id="9A0C2FCA5D4B15252E35D25CFDAEF9F0" box="[522,560,1555,1576]" pageId="12" pageNumber="23">NO</locationDeviation>
|
||||
<location id="8EDE60335D4B15252E02D25CFD78F9F0" LSID="urn:lsid:plazi:treatment:03A887FE5D4C15242D0AD03CFC1BFCFD:8EDE60335D4B15252E02D25CFD78F9F0" box="[573,742,1555,1576]" country="Costa Rica" county="Sendero Altamira" latitude="1.15" longLatPrecision="78" longitude="-1.35" municipality="Estacion Pittier" name="Estacion Pittier" pageId="12" pageNumber="23" stateProvince="Puntarenas">Estación Pittier</location>
|
||||
,
|
||||
<collectingCounty id="62DF4E645D4B15252EC8D25CFC27F9F0" box="[759,953,1555,1576]" pageId="12" pageNumber="23">Sendero Altamira</collectingCounty>
|
||||
,
|
||||
<date id="FFBF10285D4B15252FF5D25CFB86F9F0" box="[970,1048,1555,1576]" pageId="12" pageNumber="23" value="1995-10-04">
|
||||
<collectingDate id="EFFBE9C05D4B15252FF5D25CFB86F9F0" box="[970,1048,1555,1576]" pageId="12" pageNumber="23" value="1995-10-04">4X95</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<quantity id="4CF99B0D5D4B15252816D25CFE11F99B" metricMagnitude="3" metricUnit="m" metricValue="1.72" metricValueMax="1.7399999999999998" metricValueMin="1.7" pageId="12" pageNumber="23" unit="m" value="1720.0" valueMax="1740.0" valueMin="1700.0">
|
||||
<elevation id="002CD1DB5D4B15252816D25CFE11F99B" metricMagnitude="3" metricUnit="m" metricValue="1.72" metricValueMax="1.7399999999999998" metricValueMin="1.7" pageId="12" pageNumber="23" unit="m" value="1720.0" valueMax="1740.0" valueMin="1700.0">1,700– 1,740 m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<collectorName id="26F4533E5D4B15252DA2D261FD84F99B" box="[413,538,1582,1603]" pageId="12" pageNumber="23">M. Moraga</collectorName>
|
||||
,
|
||||
<specimenCode id="DBA79E935D4B15252E17D261FCABF99B" box="[552,821,1582,1603]" pageId="12" pageNumber="23">INBIO CR 1002 468647</specimenCode>
|
||||
(
|
||||
<specimenCount id="9D07FD615D4B15252F75D261FC09F99B" box="[842,919,1582,1603]" count="1" pageId="12" pageNumber="23" type="male">1 male</specimenCount>
|
||||
);
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D4B15252F92D261FE61F9A2" collectingDate="1995-01-18" collectionCode="INBIO, MXAL" collectorName="R. Villalobos" country="Costa Rica" county="Sendero Altamira" elevation="1670" latitude="1.15" location="PILA- ACLA" longLatPrecision="78" longitude="-1.35" municipality="Estacion Pittier" pageId="12" pageNumber="23" specimenCode="INBIO CR 1002 183335" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="paratype">
|
||||
<collectingRegion id="49C5F80A5D4B15252F92D261FBBCF99B" box="[941,1058,1582,1603]" country="Costa Rica" name="Puntarenas" pageId="12" pageNumber="23">Puntarenas</collectingRegion>
|
||||
,
|
||||
<location id="8EDE60335D4B1525280FD261FEE3F987" LSID="urn:lsid:plazi:treatment:03A887FE5D4C15242D0AD03CFC1BFCFD:8EDE60335D4B1525280FD261FEE3F987" country="Costa Rica" county="Sendero Altamira" latitude="1.15" longLatPrecision="78" longitude="-1.35" municipality="Estacion Pittier" name="PILA- ACLA" pageId="12" pageNumber="23" stateProvince="Puntarenas">PILAACLA</location>
|
||||
,
|
||||
<collectingMunicipality id="6BDAAC925D4B15252DB4D205FDAFF987" box="[395,561,1610,1631]" pageId="12" pageNumber="23">Estación Pittier</collectingMunicipality>
|
||||
, 5/
|
||||
<date id="FFBF10285D4B15252E6AD205FD3AF987" box="[597,676,1610,1631]" pageId="12" pageNumber="23" value="1995-01-18">
|
||||
<collectingDate id="EFFBE9C05D4B15252E6AD205FD3AF987" box="[597,676,1610,1631]" pageId="12" pageNumber="23" value="1995-01-18">18I95</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<quantity id="4CF99B0D5D4B15252E8CD205FC91F987" box="[691,783,1610,1631]" metricMagnitude="3" metricUnit="m" metricValue="1.67" pageId="12" pageNumber="23" unit="m" value="1670.0">
|
||||
<elevation id="002CD1DB5D4B15252E8CD205FC91F987" box="[691,783,1610,1631]" metricMagnitude="3" metricUnit="m" metricValue="1.67" pageId="12" pageNumber="23" unit="m" value="1670.0">1,670 m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<collectorName id="26F4533E5D4B15252F21D205FC2EF987" box="[798,944,1610,1631]" pageId="12" pageNumber="23">R. Villalobos</collectorName>
|
||||
,
|
||||
<specimenCode id="DBA79E935D4B15252F81D205FE1FF9A2" pageId="12" pageNumber="23">INBIO CR 1002 183335</specimenCode>
|
||||
(
|
||||
<specimenCount id="9D07FD615D4B15252DABD22AFE6DF9A2" box="[404,499,1637,1658]" count="1" pageId="12" pageNumber="23" type="female">1 female</specimenCount>
|
||||
);
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D4B15242E38D22AFEC4FE9D" collectingDate="1992-04" collectionCode="INBIO, MXAL" collectorName="G. Mora'" country="Costa Rica" county="Sendero Altamira" elevation="1250" lastPageId="13" lastPageNumber="24" latitude="1.15" location="Refugio Nacional de Fauna Silvestre Tapanti´" longLatPrecision="78" longitude="-1.35" municipality="Quebrada Segunda" pageId="12" pageNumber="23" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Cartago" typeStatus="paratype">
|
||||
‘‘
|
||||
<collectingCountry id="F31676785D4B15252E2FD22AFD28F9A2" box="[528,694,1637,1658]" name="Costa Rica" pageId="12" pageNumber="23">COSTA RICA</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49C5F80A5D4B15252EFED22AFC88F9A2" box="[705,790,1637,1658]" country="Costa Rica" name="Cartago" pageId="12" pageNumber="23">Cartago</collectingRegion>
|
||||
,
|
||||
<location id="8EDE60335D4B15242F1DD22AFE2FFEF2" LSID="urn:lsid:plazi:treatment:03A887FE5D4C15242D0AD03CFC1BFCFD:8EDE60335D4B15242F1DD22AFE2FFEF2" country="Costa Rica" county="Sendero Altamira" lastPageId="13" lastPageNumber="24" latitude="1.15" longLatPrecision="78" longitude="-1.35" municipality="Quebrada Segunda" name="Refugio Nacional de Fauna Sil- vestre Tapanti´" pageId="12" pageNumber="23" stateProvince="Cartago">Refugio Nacional de Fauna Silvestre Tapantı´</location>
|
||||
,
|
||||
<collectingMunicipality id="6BDAAC925D4A15242D81D55AFD14FEF2" box="[446,650,277,298]" pageId="13" pageNumber="24">Quebrada Segunda</collectingMunicipality>
|
||||
,
|
||||
<quantity id="4CF99B0D5D4A15242EA8D55AFD6DFEF2" box="[663,755,277,298]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="13" pageNumber="24" unit="m" value="1250.0">
|
||||
<elevation id="002CD1DB5D4A15242EA8D55AFD6DFEF2" box="[663,755,277,298]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="13" pageNumber="24" unit="m" value="1250.0">1,250 m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<date id="FFBF10285D4A15242F3FD55AFCA1FEF2" box="[768,831,277,298]" pageId="13" pageNumber="24" value="1992-04">
|
||||
<collectingDate id="EFFBE9C05D4A15242F3FD55AFCA1FEF2" box="[768,831,277,298]" pageId="13" pageNumber="24" value="1992-04">IV92</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F4533E5D4A15242F73D55AFC2DFEF2" box="[844,947,277,298]" pageId="13" pageNumber="24">G. Mora’</collectorName>
|
||||
’ (
|
||||
<specimenCount id="9D07FD615D4A15242FF3D55AFB88FEF2" box="[972,1046,277,298]" count="1" pageId="13" pageNumber="24" type="male">1 male</specimenCount>
|
||||
,
|
||||
<specimenCount id="9D07FD615D4A1524281BD55AFED0FE9D" count="1" pageId="13" pageNumber="24" type="female">1 female</specimenCount>
|
||||
).
|
||||
</materialsCitation>
|
||||
Paratypes deposited in
|
||||
<collectionCode id="ED10AE2D5D4A15242E64D57FFD3AFE9D" box="[603,676,304,325]" pageId="13" pageNumber="24">INBIO</collectionCode>
|
||||
and
|
||||
<collectionCode id="ED10AE2D5D4A15242EE4D57FFCB5FE9D" box="[731,811,304,325]" pageId="13" pageNumber="24">MXAL</collectionCode>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BBE36E85D4A15242D0DD505FDFAFEA2" blockId="13.[279,1113,277,805]" pageId="13" pageNumber="24">
|
||||
<emphasis id="B975EAFA5D4A15242D0DD505FE48FE87" bold="true" box="[306,470,330,351]" pageId="13" pageNumber="24">Type Locality.</emphasis>
|
||||
<location id="8EDE60335D4A15242DE1D504FD63FEB8" LSID="urn:lsid:plazi:treatment:03A887FE5D4C15242D0AD03CFC1BFCFD:8EDE60335D4A15242DE1D504FD63FEB8" box="[478,765,331,352]" country="Costa Rica" municipality="Cordillera de Talamanca" name="Falda Sur del Cerro Pittier" pageId="13" pageNumber="24">Falda Sur del Cerro Pittier</location>
|
||||
,
|
||||
<collectingMunicipality id="6BDAAC925D4A15242F36D504FB8EFEB8" box="[777,1040,331,352]" pageId="13" pageNumber="24">Cordillera de Talamanca</collectingMunicipality>
|
||||
,
|
||||
<collectingCountry id="F31676785D4A15242823D504FED9FEA2" name="Costa Rica" pageId="13" pageNumber="24">Costa Rica</collectingCountry>
|
||||
(
|
||||
<geoCoordinate id="EE35502F5D4A15242D67D52AFE56FEA2" box="[344,456,357,378]" pageId="13" pageNumber="24">9°01'450N</geoCoordinate>
|
||||
;
|
||||
<geoCoordinate id="EE35502F5D4A15242DEAD52AFDC6FEA2" box="[469,600,357,378]" pageId="13" pageNumber="24">82°57'550W</geoCoordinate>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D4A15242D0DD5CFFCCAFDC2" pageId="13" pageNumber="24" type="biology_ecology">
|
||||
<paragraph id="8BBE36E85D4A15242D0DD5CFFCCAFDC2" blockId="13.[279,1113,277,805]" pageId="13" pageNumber="24">
|
||||
<emphasis id="B975EAFA5D4A15242D0DD5CFFE7DFE4D" bold="true" box="[306,483,384,405]" pageId="13" pageNumber="24">Biological Data.</emphasis>
|
||||
Four males and
|
||||
<specimenCount id="9D07FD615D4A15242EADD5CFFC80FE4D" box="[658,798,384,405]" pageId="13" pageNumber="24" type="female">three females</specimenCount>
|
||||
of
|
||||
<taxonomicName id="4C014D6B5D4A15242F7ED5CFFC78FE4D" box="[833,998,384,405]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="24" phylum="Arthropoda" rank="species" species="talamancana" status="sp. nov.">
|
||||
<emphasis id="B975EAFA5D4A15242F7ED5CFFC78FE4D" box="[833,998,384,405]" italics="true" pageId="13" pageNumber="24">P. talamancana</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A24657815D4A15242FD2D5CFFB81FE4D" box="[1005,1055,384,405]" pageId="13" pageNumber="24" rank="species">n.sp.</taxonomicNameLabel>
|
||||
were collected at lights in grasslands, near the forest, and
|
||||
<specimenCount id="9D07FD615D4A15242F74D5D4FC5EFE68" box="[843,960,411,432]" pageId="13" pageNumber="24" type="female">one female</specimenCount>
|
||||
was collected at lights in premontane rain forest, located between
|
||||
<quantity id="4CF99B0D5D4A15242F01D5FAFC40FE12" box="[830,990,437,458]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="1.75" metricValueMin="1.25" pageId="13" pageNumber="24" unit="m" value="1500.0" valueMax="1750.0" valueMin="1250.0">
|
||||
<geoCoordinate id="EE35502F5D4A15242F01D5FAFCE7FE12" box="[830,889,437,458]" degrees="1,250" direction="north" orientation="latitude" pageId="13" pageNumber="24" precision="55" value="1.25">1,250</geoCoordinate>
|
||||
<geoCoordinate id="EE35502F5D4A15242F46D5FAFC5CFE12" box="[889,962,437,458]" degrees="1,750" direction="west" orientation="longitude" pageId="13" pageNumber="24" precision="55" value="-1.75">–1,750</geoCoordinate>
|
||||
m
|
||||
</quantity>
|
||||
of altitude. Phenology: January (1), April (2), July (1), October (2), November (2). Other species of
|
||||
<taxonomicName id="4C014D6B5D4A15242DAFD5A4FD88FDD8" box="[400,534,491,512]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B975EAFA5D4A15242DAFD5A4FD88FDD8" box="[400,534,491,512]" italics="true" pageId="13" pageNumber="24">Phyllophaga</emphasis>
|
||||
</taxonomicName>
|
||||
flying at the same time and places were:
|
||||
<emphasis id="B975EAFA5D4A15242FCCD5A4FEF4FDC3" italics="true" pageId="13" pageNumber="24">P. nevermannea</emphasis>
|
||||
Saylor,
|
||||
<emphasis id="B975EAFA5D4A15242DFAD649FDAAFDC3" box="[453,564,518,539]" italics="true" pageId="13" pageNumber="24">P. orosina</emphasis>
|
||||
Moser,
|
||||
<taxonomicName id="4C014D6B5D4A15242EB2D649FC8EFDC3" box="[653,784,518,539]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="24" phylum="Arthropoda" rank="species" species="tapantina" status="sp. nov.">
|
||||
<emphasis id="B975EAFA5D4A15242EB2D649FC8EFDC3" box="[653,784,518,539]" italics="true" pageId="13" pageNumber="24">P. tapantina</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A24657815D4A15242F25D64AFCCAFDC2" box="[794,852,517,538]" pageId="13" pageNumber="24" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D4A15242D0DD66FFC21FD28" pageId="13" pageNumber="24" type="discussion">
|
||||
<paragraph id="8BBE36E85D4A15242D0DD66FFC21FD28" blockId="13.[279,1113,277,805]" pageId="13" pageNumber="24">
|
||||
<emphasis id="B975EAFA5D4A15242D0DD66FFE00FDED" bold="true" box="[306,414,544,565]" pageId="13" pageNumber="24">Remarks.</emphasis>
|
||||
<taxonomicName id="4C014D6B5D4A15242D94D66FFCF7FDED" box="[427,873,544,565]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="24" phylum="Arthropoda" rank="species" species="talamancana" status="sp. nov." subGenus="Phyllophaga">
|
||||
<emphasis id="B975EAFA5D4A15242D94D66FFDAFFDED" box="[427,561,544,565]" italics="true" pageId="13" pageNumber="24">Phyllophaga</emphasis>
|
||||
(
|
||||
<emphasis id="B975EAFA5D4A15242E79D66FFD55FDED" box="[582,715,544,565]" italics="true" pageId="13" pageNumber="24">Phyllophaga</emphasis>
|
||||
)
|
||||
<emphasis id="B975EAFA5D4A15242EDED66FFCF7FDED" box="[737,873,544,565]" italics="true" pageId="13" pageNumber="24">talamancana</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A24657815D4A15242F49D66FFC2BFDED" box="[886,949,544,565]" pageId="13" pageNumber="24" rank="species">n. sp.</taxonomicNameLabel>
|
||||
is not closely related with any known species group (
|
||||
<emphasis id="B975EAFA5D4A15242E84D674FD6BFD88" box="[699,757,571,592]" italics="true" pageId="13" pageNumber="24">sensu</emphasis>
|
||||
<bibRefCitation id="EF904B195D4A15242EC2D674FC1AFD88" author="Moron" box="[765,900,571,592]" firstAuthor="Moron" pageId="13" pageNumber="24" refId="ref11224" refString="Moron, M. A. 1986. El genero Phyllophaga en Mexico. Morfologia, distribucion y sistematica supraespecifica (Insecta: Coleoptera). Publ. 20, Instituto de Ecologia, Mexico, D. F. 341 pp." type="book" year="1986">Morón 1986</bibRefCitation>
|
||||
). The general structure of the last sternites, shape of tarsal claws and the length of antennal club in the males suggest a vague relation with some species in the ‘‘
|
||||
<emphasis id="B975EAFA5D4A15242FF1D63FFBD6FD5D" box="[974,1096,624,645]" italics="true" pageId="13" pageNumber="24">schizorhina</emphasis>
|
||||
’’ group, and the sculpture of the head and pronotum resemble some species in the ‘‘
|
||||
<emphasis id="B975EAFA5D4A15242D6DD6E9FE5BFD63" box="[338,453,678,699]" italics="true" pageId="13" pageNumber="24">blanchardi</emphasis>
|
||||
’’ group. The shape and details of the male genital capsule, sculpture of the pronotum, pygidium and sternites, body luster and color will aid to separate this species from other Costarican
|
||||
<emphasis id="B975EAFA5D4A15242F0DD694FC21FD28" box="[818,959,731,752]" italics="true" pageId="13" pageNumber="24">
|
||||
<taxonomicName id="4C014D6B5D4A15242F0DD694FC25FD28" box="[818,955,731,752]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="24" phylum="Arthropoda" rank="genus">Phyllophaga</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D4A15242D0DD6BAFC1BFCFD" pageId="13" pageNumber="24" type="etymology">
|
||||
<paragraph id="8BBE36E85D4A15242D0DD6BAFC1BFCFD" blockId="13.[279,1113,277,805]" pageId="13" pageNumber="24">
|
||||
<emphasis id="B975EAFA5D4A15242D0DD6BAFE2FFCD2" bold="true" box="[306,433,757,778]" pageId="13" pageNumber="24">Etymology.</emphasis>
|
||||
Derived from ‘‘Cordillera de Talamanca’’ that comprises the majority of the geographic range where this species lives.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
304
data/03/A8/87/03A887FE5D4F15232D6CD0A1FBEAF9A2.xml
Normal file
304
data/03/A8/87/03A887FE5D4F15232D6CD0A1FBEAF9A2.xml
Normal file
|
|
@ -0,0 +1,304 @@
|
|||
<document id="C8C1C0A7BB13D2ED784AC60601B02A6D" ID-DOI="10.1649/0010-065x(2001)055[0011:snsops]2.0.co;2" ID-ISSN="1938-4394" ID-Zenodo-Dep="4900311" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1622827801739" checkinUser="carolina" docAuthor="Morón, Miguel Angel & Solís, Angel" docDate="2001" docId="03A887FE5D4F15232D6CD0A1FBEAF9A2" docLanguage="en" docName="TheColeopBull.55.1.11-29.pdf.imf" docOrigin="The Coleopterists Bulletin 55 (1)" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Phyllophaga (Phyllophaga) puntarenosa Moron and Solis, new species" docType="treatment" docVersion="1" lastPageNumber="21" masterDocId="FF91FF865D4715292C3FD44FFF9EFFD8" masterDocTitle="Seven New Species Of Phyllophaga (S. Str.) Harris From Costa Rica (Coleoptera: Melolonthidae: Melolonthinae)" masterLastPageNumber="29" masterPageNumber="11" pageNumber="19" updateTime="1730139982923" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:titleInfo id="A4C6EFFA9DB634FE6A9A232FCFF451B7">
|
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<mods:title id="678C85510E34E05DC85D090C411ADA9F">Seven New Species Of Phyllophaga (S. Str.) Harris From Costa Rica (Coleoptera: Melolonthidae: Melolonthinae)</mods:title>
|
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</mods:titleInfo>
|
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<mods:name id="2F46E5ABBDE869C72FE14FFBAE750B09" type="personal">
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<mods:role id="A7C7B4556C9AA3D75A31345725722161">
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<mods:roleTerm id="FC8508B93DD5ED3C68AC9D22BF7589FC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B8531F45C615EDFC9C594636039654E0">Morón, Miguel Angel</mods:namePart>
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<mods:affiliation id="C596289D21A6E969CA6F9342FD92C5DB">Departamento de Entomología Instituto de Ecología, A. C. (sector SEPCONACYT) Apartado Postal 63 Xalapa, Veracruz 91000, MÉXICO</mods:affiliation>
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</mods:name>
|
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<mods:roleTerm id="74E0A210BF609008D580D8474B9D903E">Author</mods:roleTerm>
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||||
</mods:role>
|
||||
<mods:namePart id="8E525748580FCF1D3988BD5DD08E70E6">Solís, Angel</mods:namePart>
|
||||
<mods:affiliation id="447BA7602D9CC37FBC8EC89924AA25F0">Departamento de Entomología Instituto Nacional de Biodiversidad (INBio) Apartado Postal 22 3100 Santo Domingo de Heredia, COSTA RICA</mods:affiliation>
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</mods:name>
|
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<mods:typeOfResource id="73E1FAF13F0B429628A040A3BAF60DB4">text</mods:typeOfResource>
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<mods:title id="CB509EA1E3D3450432653E627782CB4A">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="D5F42803C9709E104DED1168E1C0CC3E">
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<mods:date id="0601917F44D720AB4B9A1BEAA1730D9F">2001</mods:date>
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<mods:number id="D5446AAD54BC4CBF85C675FC20701843">2001-03-01</mods:number>
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<mods:number id="0647CED8A9C8E25A5F55EB1176C4E212">55</mods:number>
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|
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<mods:classification id="2974464F295D437D9A9520A9B36C33C9">journal article</mods:classification>
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<mods:identifier id="D44F23728DDF30B8364A926B77644C38" type="DOI">10.1649/0010-065x(2001)055[0011:snsops]2.0.co;2</mods:identifier>
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<mods:identifier id="C4E66567D47CB9D001FD17A989E7FCF5" type="ISSN">1938-4394</mods:identifier>
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<mods:identifier id="F1EA6543EEBF685DCCF388AB1192D82E" type="Zenodo-Dep">4900311</mods:identifier>
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</mods:mods>
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<treatment id="03A887FE5D4F15232D6CD0A1FBEAF9A2" LSID="urn:lsid:plazi:treatment:03A887FE5D4F15232D6CD0A1FBEAF9A2" httpUri="http://treatment.plazi.org/id/03A887FE5D4F15232D6CD0A1FBEAF9A2" lastPageId="10" lastPageNumber="21" pageId="8" pageNumber="19">
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<subSubSection id="C31B65635D4F15212D6CD0A1FBCCFADA" box="[339,1106,1261,1283]" pageId="8" pageNumber="19" type="nomenclature">
|
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<paragraph id="8BBE36E85D4F15212D6CD0A1FBCCFADA" blockId="8.[339,1106,1261,1309]" box="[339,1106,1261,1283]" pageId="8" pageNumber="19">
|
||||
<taxonomicName id="4C014D6B5D4F15212D6CD0A1FC5DFADB" authority="Moron and Solis" authorityName="Moron and Solis" box="[339,963,1262,1283]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="puntarenosa" status="sp. nov." subGenus="Phyllophaga">
|
||||
<emphasis id="B975EAFA5D4F15212D6CD0A1FE47FADB" box="[339,473,1262,1283]" italics="true" pageId="8" pageNumber="19">Phyllophaga</emphasis>
|
||||
(
|
||||
<emphasis id="B975EAFA5D4F15212DD5D0A1FDF1FADB" box="[490,623,1262,1283]" italics="true" pageId="8" pageNumber="19">Phyllophaga</emphasis>
|
||||
)
|
||||
<emphasis id="B975EAFA5D4F15212EBED0A1FC9AFADB" box="[641,772,1262,1283]" italics="true" pageId="8" pageNumber="19">puntarenosa</emphasis>
|
||||
Morón and Solís
|
||||
</taxonomicName>
|
||||
,
|
||||
<emphasis id="B975EAFA5D4F15212FF0D0A2FBCCFADA" bold="true" box="[975,1106,1261,1282]" pageId="8" pageNumber="19">
|
||||
<taxonomicNameLabel id="A24657815D4F15212FF0D0A2FBCCFADA" box="[975,1106,1261,1282]" pageId="8" pageNumber="19" rank="species">new species</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D4F15232EACD147FC55FC66" lastPageId="10" lastPageNumber="21" pageId="8" pageNumber="19" type="description">
|
||||
<paragraph id="8BBE36E85D4F15212EACD147FC8CFAC5" blockId="8.[339,1106,1261,1309]" box="[659,786,1288,1309]" pageId="8" pageNumber="19">
|
||||
<figureCitation id="133A2A6D5D4F15212EACD147FC8CFAC5" box="[659,786,1288,1309]" captionStart="Figs" captionStartId="9.[303,346,836,855]" captionTargetBox="[318,1075,273,816]" captionTargetId="figure-12@9.[313,1078,272,816]" captionTargetPageId="9" captionText="Figs. 14–17. Phyllophaga puntarenosa. 14) protarsal claw, male; 15) genital capsule, lateral view; 16) paramera, distal view; 17) genital plates, female. Scale lines 5 1 mm, except Fig. 14 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900319" httpUri="https://zenodo.org/record/4900319/files/figure.png" pageId="8" pageNumber="19">Figs. 14–17</figureCitation>
|
||||
</paragraph>
|
||||
<paragraph id="8BBE36E85D4F15232D75D160FE58FE99" blockId="8.[307,1140,1327,1658]" lastBlockId="10.[307,1139,277,321]" lastPageId="10" lastPageNumber="21" pageId="8" pageNumber="19">
|
||||
<emphasis id="B975EAFA5D4F15212D75D160FE34FA9A" bold="true" box="[330,426,1327,1346]" pageId="8" pageNumber="19">
|
||||
<typeStatus id="54BA884A5D4F15212D75D160FE38FA9A" box="[330,422,1327,1346]" pageId="8" pageNumber="19" type="holotype">Holotype</typeStatus>
|
||||
.
|
||||
</emphasis>
|
||||
Male. Clypeus, frons, pronotum and basal sixth of elytra, shiny dark reddish brown; remaining surface of elytra shiny reddish brown, without macroscopic vestiture; mouthparts, sterna, pygidium and legs shiny dark reddish brown. Clypeus 3.5× wider than long, anterior border widely sinuate, with elevated margin, surface slightly concave toward sides, with many uniformly distributed, shallow, round punctures and without setae. Frontoclypeal suture sinuate and shallowly impressed. Frons 2.1× wider than long, convex, densely punctate, slightly rugose at center of disk, without setae. Antenna 10 segmented, with 3segmented club, lamellae of 8th to 10th segments 1.8× longer than length of preceeding 6 segments combined. Frons 2.4× wider than dorsal diameter of eye. Eye canthus long and narrow, with 10–11 setae. Labrum bilobed, widely sinuate, with scattered slender setae. Mentum slightly concave, impunctate, with scarce slender setae at sides, anterior border narrowly notched. Pronotum 1.8× wider than long and 3.0× wider than frons. Pronotal disk shiny, with shallow, round punctures irregularly separated from one another by 0.3–2 diameters; lateral borders widely angled, lateral marginal bead irregularly crenulate, with scattered, short, slender setae; basal bead clearly indicated, reinforced with punctures throughout; anterior angles obtuse, not prominent; posterior angles widely obtuse, not prominent. Scutellum 1.5× wider than long, with some small punctures. Elytron 2.3× longer than wide, shiny, densely punctate; epipleural border very narrow, extended along complete margin, provided with very short, scattered, slender setae; humeral callus rounded, prominent; apical callus rounded. Metathoracic wings completely developed. Propygidium shiny, densely punctate with sparse short setae. Pygidium scarcely convex, shiny, punctaterugose with round punctures, glabrous; apical margin with 12 long, slender setae; basal margin effaced medially. Pterosterna with long, moderately dense, yellowish setae. Visible abdominal sternites II and IV slightly depressed, polished and with scarce short setae near the midline; sternite V convex, shiny, with sparse punctures and some scattered setae toward the sides; anal plate narrow, sligthly prominent, irregularly punctate, with transverse carina and 24 slender setae near the posterior border. Protibia slightly shorter than protarsus (0.8:1), with external border tridentate, basal tooth small, preapical spur slightly curved with rounded apex, as long as 2th protarsomere. Mesotibia with one oblique, sharp, setiferous carina on external side; upper apical spur straight, narrow, and 1.4× longer than lower spur. Metatibia as long as metatarsus, with one oblique, sharp, setiferous carina on external side; upper apical spur articulated, curved, apex rounded, as long as basal metatarsomere, and 2.6× longer than lower spur; lower apical spur fused with tibial border, acute apex. Tarsomeres semicylindrical, elongate, with enlarged apices, some setae apically and two longitudinal fine carinae usually with lines of scattered, stout setae. Protarsomeres 1–4 with subapical toothlike or rounded process. Tarsal claws symmetrical, similar on all legs, with long tooth at middle of ventral border directed slightly towards apex (
|
||||
<figureCitation id="133A2A6D5D4E1520280DD251FEAFF991" captionStart="Figs" captionStartId="9.[303,346,836,855]" captionTargetBox="[318,1075,273,816]" captionTargetId="figure-12@9.[313,1078,272,816]" captionTargetPageId="9" captionText="Figs. 14–17. Phyllophaga puntarenosa. 14) protarsal claw, male; 15) genital capsule, lateral view; 16) paramera, distal view; 17) genital plates, female. Scale lines 5 1 mm, except Fig. 14 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900319" httpUri="https://zenodo.org/record/4900319/files/figure.png" pageId="9" pageNumber="20">Fig. 14</figureCitation>
|
||||
). Genital capsule with short parameres, dorsally fused, apices ventrally curved, sharply pointed. Aedeagus long, with preapical dorsal patches of spinules and wide sclerotized support (
|
||||
<figureCitation id="133A2A6D5D4E15202D52D228FE7FF9A2" box="[365,481,1639,1658]" captionStart="Figs" captionStartId="9.[303,346,836,855]" captionTargetBox="[318,1075,273,816]" captionTargetId="figure-12@9.[313,1078,272,816]" captionTargetPageId="9" captionText="Figs. 14–17. Phyllophaga puntarenosa. 14) protarsal claw, male; 15) genital capsule, lateral view; 16) paramera, distal view; 17) genital plates, female. Scale lines 5 1 mm, except Fig. 14 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900319" httpUri="https://zenodo.org/record/4900319/files/figure.png" pageId="9" pageNumber="20">Figs. 15–16</figureCitation>
|
||||
). Tectum uniformly convex. Length of genital capsule from apex of parameres to border of basal piece: 4.0 mm. Total body length:
|
||||
<quantity id="4CF99B0D5D4D15232F81D55AFB88FEF0" box="[958,1046,277,296]" metricMagnitude="-2" metricUnit="m" metricValue="1.72" pageId="10" pageNumber="21" unit="mm" value="17.2">17.2 mm</quantity>
|
||||
. Humeral width:
|
||||
<quantity id="4CF99B0D5D4D15232D49D561FE5CFE99" box="[374,450,302,321]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="10" pageNumber="21" unit="mm" value="7.8">7.8 mm</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
<caption id="DF7E66605D4E15202D10D70BFD8FFC50" ID-DOI="http://doi.org/10.5281/zenodo.4900319" ID-Zenodo-Dep="4900319" httpUri="https://zenodo.org/record/4900319/files/figure.png" pageId="9" pageNumber="20" startId="9.[303,346,836,855]" targetBox="[318,1075,273,816]" targetPageId="9">
|
||||
<paragraph id="8BBE36E85D4E15202D10D70BFD8FFC50" blockId="9.[279,1112,836,904]" pageId="9" pageNumber="20">
|
||||
<emphasis id="B975EAFA5D4E15202D10D70BFE3DFC8F" bold="true" box="[303,419,836,855]" pageId="9" pageNumber="20">Figs. 14–17.</emphasis>
|
||||
<emphasis id="B975EAFA5D4E15202D84D70AFD2DFC80" box="[443,691,837,856]" italics="true" pageId="9" pageNumber="20">
|
||||
<taxonomicName id="4C014D6B5D4E15202D84D70AFD30FC80" box="[443,686,837,856]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="20" phylum="Arthropoda" rank="species" species="puntarenosa">Phyllophaga puntarenosa</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
<emphasis id="B975EAFA5D4E15202E88D70BFD50FC8F" bold="true" box="[695,718,836,855]" pageId="9" pageNumber="20">14</emphasis>
|
||||
) protarsal claw, male;
|
||||
<emphasis id="B975EAFA5D4E15202F9CD70BFC24FC8F" bold="true" box="[931,954,836,855]" pageId="9" pageNumber="20">15</emphasis>
|
||||
) genital capsule, lateral view;
|
||||
<emphasis id="B975EAFA5D4E15202DAAD713FE32FCB7" bold="true" box="[405,428,860,879]" pageId="9" pageNumber="20">16</emphasis>
|
||||
) paramera, distal view;
|
||||
<emphasis id="B975EAFA5D4E15202EAAD713FD32FCB7" bold="true" box="[661,684,860,879]" pageId="9" pageNumber="20">17</emphasis>
|
||||
) genital plates, female. Scale lines 5 1 mm, except Fig. 14 5 0.5 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BBE36E85D4D15232D72D52DFCA4FD38" blockId="10.[307,1140,354,1658]" pageId="10" pageNumber="21">
|
||||
<emphasis id="B975EAFA5D4D15232D72D52DFE2CFEAF" bold="true" box="[333,434,354,375]" pageId="10" pageNumber="21">
|
||||
<typeStatus id="54BA884A5D4D15232D72D52DFE33FEAF" box="[333,429,354,375]" pageId="10" pageNumber="21" type="allotype">Allotype</typeStatus>
|
||||
.
|
||||
</emphasis>
|
||||
Female. Similar to the male except as follows: anterior border of clypeus more sinuate; antennae with lamellae of 8th to 10th segments shorter than the length of six preceeding one (0.9:1). Pygidium moderadely convex, 1.2× longer than in male, distal half nearly flattened, the center of disk shiny and sides slightly pruinose. Visible abdominal sternites II to V convex, with scattered, setiferous punctures; anal plate 1.6× longer than male anal plate, convex, moderately punctate setiferous, with 26 slender setae near the posterior border. Meso and metatibiae each with one oblique, strong, setiferous carina on external side. Both apical spurs of metatibia articulated, wide, lanceolate and curved. Ventral genital plates slightly sclerotized, sligthly asymmetrical, elongated and vaguely angulate, with scarce short setae at mesodistal angle; dorsal genital plates slightly sclerotized, completely fused to each other, distal border widely sinuate with five stout, long setae at each side (
|
||||
<figureCitation id="133A2A6D5D4D15232FE8D6E0FBB6FD1C" box="[983,1064,687,708]" captionStart="Figs" captionStartId="9.[303,346,836,855]" captionTargetBox="[318,1075,273,816]" captionTargetId="figure-12@9.[313,1078,272,816]" captionTargetPageId="9" captionText="Figs. 14–17. Phyllophaga puntarenosa. 14) protarsal claw, male; 15) genital capsule, lateral view; 16) paramera, distal view; 17) genital plates, female. Scale lines 5 1 mm, except Fig. 14 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900319" httpUri="https://zenodo.org/record/4900319/files/figure.png" pageId="10" pageNumber="21">Fig. 17</figureCitation>
|
||||
). Total body length:
|
||||
<quantity id="4CF99B0D5D4D15232DFDD684FDBAFD38" box="[450,548,715,736]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="10" pageNumber="21" unit="mm" value="19.5">19.5 mm</quantity>
|
||||
. Humeral width:
|
||||
<quantity id="4CF99B0D5D4D15232EDFD684FCA8FD38" box="[736,822,715,736]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="10" pageNumber="21" unit="mm" value="9.2">9.2 mm</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BBE36E85D4D15232D72D6A9FC55FC66" blockId="10.[307,1140,354,1658]" pageId="10" pageNumber="21">
|
||||
<emphasis id="B975EAFA5D4D15232D72D6A9FDB2FD23" bold="true" box="[333,556,742,763]" pageId="10" pageNumber="21">
|
||||
<typeStatus id="54BA884A5D4D15232D72D6A9FE2DFD23" box="[333,435,742,763]" pageId="10" pageNumber="21" type="paratype">Paratype</typeStatus>
|
||||
Variation.
|
||||
</emphasis>
|
||||
Males. Similar to
|
||||
<typeStatus id="54BA884A5D4D15232ECFD6A8FCD5FD24" box="[752,843,743,764]" pageId="10" pageNumber="21" type="holotype">holotype</typeStatus>
|
||||
except in total body length:
|
||||
<quantity id="4CF99B0D5D4D15232D0CD74DFE4CFCCF" box="[307,466,770,791]" metricMagnitude="-2" metricUnit="m" metricValue="1.815" metricValueMax="2.1399999999999997" metricValueMin="1.49" pageId="10" pageNumber="21" unit="mm" value="18.15" valueMax="21.4" valueMin="14.9">14.9–21.4 mm</quantity>
|
||||
, humeral width: 6.3–9.0 mm, anterior dorsal surface of some specimens darker or lighter than
|
||||
<typeStatus id="54BA884A5D4D15232EADD751FD6EFCEB" box="[658,752,798,819]" pageId="10" pageNumber="21" type="holotype">holotype</typeStatus>
|
||||
, other specimens with antennal club 1.5–1.6× longer than length of preceeding six segments combined. Female similar to
|
||||
<typeStatus id="54BA884A5D4D15232D99D71AFE64FCB2" box="[422,506,853,874]" pageId="10" pageNumber="21" type="allotype">allotype</typeStatus>
|
||||
except as follows: total body length: 17.8–24.0 mm; humeral width:
|
||||
<quantity id="4CF99B0D5D4D15232DFED73EFDCEFC5E" box="[449,592,881,902]" metricMagnitude="-3" metricUnit="m" metricValue="9.25" metricValueMax="10.7" metricValueMin="7.8" pageId="10" pageNumber="21" unit="mm" value="9.25" valueMax="10.7" valueMin="7.8">7.8–10.7 mm</quantity>
|
||||
, some specimens do not have the pruinosity on the pygidium, but others have the pygidial disk completely pruinose. Anterior dorsal surface of some specimens darker or lighter than
|
||||
<typeStatus id="54BA884A5D4D15232F4FD7E6FC56FC66" box="[880,968,937,958]" pageId="10" pageNumber="21" type="allotype">allotype</typeStatus>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D4D15232D72D78AFC96FB62" pageId="10" pageNumber="21" type="materials_examined">
|
||||
<paragraph id="8BBE36E85D4D15232D72D78AFC9AFB5A" blockId="10.[307,1140,354,1658]" pageId="10" pageNumber="21">
|
||||
<emphasis id="B975EAFA5D4D15232D72D78AFE49FC02" bold="true" box="[333,471,965,986]" pageId="10" pageNumber="21">Type Series.</emphasis>
|
||||
Described from
|
||||
<specimenCount id="9D07FD615D4D15232EB3D78AFD75FC02" box="[652,747,965,986]" pageId="10" pageNumber="21" type="male">30 males</specimenCount>
|
||||
and
|
||||
<specimenCount id="9D07FD615D4D15232F1ED78AFC06FC02" box="[801,920,965,986]" pageId="10" pageNumber="21" type="female">24 females</specimenCount>
|
||||
.
|
||||
<materialsCitation id="3B693CB55D4D15232F9DD78AFD77FBCA" accessionNumber="CR 292500" collectingDate="1991-02" collectionCode="IN- BIO" collectorName="F. Quezada" country="Costa Rica" latitude="8.566667" location="Rancho Quemado" longLatPrecision="1301" longitude="-83.583336" municipality="Peninsula Osa" pageId="10" pageNumber="21" specimenCode="INBIO CR 292500 511000" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="holotype">
|
||||
<typeStatus id="54BA884A5D4D15232F9DD78AFB9DFC02" box="[930,1027,965,986]" pageId="10" pageNumber="21" type="holotype">Holotype</typeStatus>
|
||||
<specimenCount id="9D07FD615D4D15232834D78AFBA0FC02" box="[1035,1086,965,986]" pageId="10" pageNumber="21" type="male">male</specimenCount>
|
||||
(
|
||||
<collectionCode id="ED10AE2D5D4D15232872D78AFEFBFC2E" pageId="10" pageNumber="21">INBIO</collectionCode>
|
||||
): ‘‘
|
||||
<collectingCountry id="F31676785D4D15232DBBD7AEFDB2FC2E" box="[388,556,993,1014]" name="Costa Rica" pageId="10" pageNumber="21">COSTA RICA</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49C5F80A5D4D15232E06D7AEFD30FC2E" box="[569,686,993,1014]" country="Costa Rica" name="Puntarenas" pageId="10" pageNumber="21">Puntarenas</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="6BDAAC925D4D15232E82D7AEFCC6FC2E" box="[701,856,993,1014]" pageId="10" pageNumber="21">Península Osa</collectingMunicipality>
|
||||
,
|
||||
<location id="8EDE60335D4D15232F58D7AEFBB7FC2E" LSID="urn:lsid:plazi:treatment:03A887FE5D4F15232D6CD0A1FBEAF9A2:8EDE60335D4D15232F58D7AEFBB7FC2E" box="[871,1065,993,1014]" country="Costa Rica" municipality="Peninsula Osa" name="Rancho Quemado" pageId="10" pageNumber="21" stateProvince="Puntarenas">Rancho Quemado</location>
|
||||
,
|
||||
<date id="FFBF10285D4D15232807D7AEFBF0FC2E" box="[1080,1134,993,1014]" pageId="10" pageNumber="21" value="1991-02">
|
||||
<collectingDate id="EFFBE9C05D4D15232807D7AEFBF0FC2E" box="[1080,1134,993,1014]" pageId="10" pageNumber="21" value="1991-02">II91</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F4533E5D4D15232D0CD7B2FE37FBCA" box="[307,425,1021,1042]" pageId="10" pageNumber="21">F. Quezada</collectorName>
|
||||
,
|
||||
<specimenCode id="DBA79E935D4D15232D8AD7B2FD42FBCA" box="[437,732,1021,1042]" pageId="10" pageNumber="21">INBIO CR 292500 511000</specimenCode>
|
||||
.’’
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D4D15232ECED7B2FE2CFBF6" accessionNumber="CR 292500" collectingDate="1991-02" collectionCode="INBIO" collectorName="F. Quezada" country="Costa Rica" latitude="8.566667" location="Peninsula Osa" longLatPrecision="1301" longitude="-83.583336" municipality="Peninsula Osa" pageId="10" pageNumber="21" specimenCode="INBIO CR 292500 511000" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="allotype">
|
||||
<typeStatus id="54BA884A5D4D15232ECED7B2FCD2FBCA" box="[753,844,1021,1042]" pageId="10" pageNumber="21" type="allotype">Allotype</typeStatus>
|
||||
<specimenCount id="9D07FD615D4D15232F6BD7B2FC05FBCA" box="[852,923,1021,1042]" pageId="10" pageNumber="21" type="female">female</specimenCount>
|
||||
(
|
||||
<collectionCode id="ED10AE2D5D4D15232F95D7B2FC67FBCA" box="[938,1017,1021,1042]" pageId="10" pageNumber="21">INBIO</collectionCode>
|
||||
): same data as holotype
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3B693CB55D4D15232D80D056FBB6FBF6" accessionNumber="CR 292500" box="[447,1064,1049,1070]" collectingDate="1991-02" collectionCode="CASC, CNC, INBIO, MXAL, TAMU, ZMHU" collectorName="F. Quezada" country="Costa Rica" latitude="8.566667" location="Peninsula Osa" longLatPrecision="1301" longitude="-83.583336" municipality="Peninsula Osa" pageId="10" pageNumber="21" specimenCode="INBIO CR 292500 511000" specimenCount="18" specimenCount-female="10" specimenCount-male="8" stateProvince="Puntarenas" typeStatus="paratype">
|
||||
<typeStatus id="54BA884A5D4D15232D80D056FDB7FBF6" box="[447,553,1049,1070]" pageId="10" pageNumber="21" type="paratype">Paratypes</typeStatus>
|
||||
: same data as holotype (
|
||||
<specimenCount id="9D07FD615D4D15232F03D056FC0AFBF6" box="[828,916,1049,1070]" count="8" pageId="10" pageNumber="21" type="male">8 males</specimenCount>
|
||||
,
|
||||
<specimenCount id="9D07FD615D4D15232F9DD056FB83FBF6" box="[930,1053,1049,1070]" count="10" pageId="10" pageNumber="21" type="female">10 females</specimenCount>
|
||||
);
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B693CB55D4D1523280DD056FC8DFBBE" accessionNumber="CR 292500" collectingDate="1991-04" collectionCode="INBIO" collectorName="G. Mora" country="Costa Rica" location="Est. Las Mellizas" municipality="P. N. Amistad" pageId="10" pageNumber="21" specimenCode="INBIO CR 1000 659021" specimenCount="34" specimenCount-female="13" specimenCount-male="21" stateProvince="Puntarenas" typeStatus="paratype">
|
||||
‘‘
|
||||
<collectingRegion id="49C5F80A5D4D15232803D056FE1EFB92" country="Costa Rica" name="Puntarenas" pageId="10" pageNumber="21">Puntarenas</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="6BDAAC925D4D15232DB5D07AFD89FB92" box="[394,535,1077,1098]" pageId="10" pageNumber="21">P.N. Amistad</collectingMunicipality>
|
||||
,
|
||||
<location id="8EDE60335D4D15232E1DD07AFD44FB92" LSID="urn:lsid:plazi:treatment:03A887FE5D4F15232D6CD0A1FBEAF9A2:8EDE60335D4D15232E1DD07AFD44FB92" box="[546,730,1077,1098]" country="Costa Rica" municipality="P. N. Amistad" name="Est. Las Mellizas" pageId="10" pageNumber="21" stateProvince="Puntarenas">Est. Las Mellizas</location>
|
||||
,
|
||||
<location id="8EDE60335D4D15232EDAD07AFCEEFB92" LSID="urn:lsid:plazi:treatment:03A887FE5D4F15232D6CD0A1FBEAF9A2:8EDE60335D4D15232EDAD07AFCEEFB92" box="[741,880,1077,1098]" country="Costa Rica" municipality="P. N. Amistad" name="Finca Cafosa" pageId="10" pageNumber="21" stateProvince="Puntarenas">Finca Cafosa</location>
|
||||
,
|
||||
<date id="FFBF10285D4D15232F44D07AFC24FB92" box="[891,954,1077,1098]" pageId="10" pageNumber="21" value="1991-04">
|
||||
<collectingDate id="EFFBE9C05D4D15232F44D07AFC24FB92" box="[891,954,1077,1098]" pageId="10" pageNumber="21" value="1991-04">IV91</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F4533E5D4D15232FFAD07AFBBEFB92" box="[965,1056,1077,1098]" pageId="10" pageNumber="21">G. Mora</collectorName>
|
||||
,
|
||||
<specimenCode id="DBA79E935D4D15232815D07AFE66FBBE" pageId="10" pageNumber="21">INBIO CR 1000 659021</specimenCode>
|
||||
’’ (
|
||||
<specimenCount id="9D07FD615D4D15232E28D01EFDE1FBBE" box="[535,639,1105,1126]" count="21" pageId="10" pageNumber="21" type="male">21 males</specimenCount>
|
||||
,
|
||||
<specimenCount id="9D07FD615D4D15232EB0D01EFC92FBBE" box="[655,780,1105,1126]" count="13" pageId="10" pageNumber="21" type="female">13 females</specimenCount>
|
||||
)
|
||||
</materialsCitation>
|
||||
. Paratypes deposited in
|
||||
<collectionCode id="ED10AE2D5D4D15232817D01EFBF1FBBE" box="[1064,1135,1105,1126]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="10" pageNumber="21">CASC</collectionCode>
|
||||
,
|
||||
<collectionCode id="ED10AE2D5D4D15232D0CD022FEF5FB5A" box="[307,363,1133,1154]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" pageId="10" pageNumber="21">CNC</collectionCode>
|
||||
,
|
||||
<collectionCode id="ED10AE2D5D4D15232D47D022FE5DFB5A" box="[376,451,1133,1154]" pageId="10" pageNumber="21">INBIO</collectionCode>
|
||||
,
|
||||
<collectionCode id="ED10AE2D5D4D15232DEFD022FDBEFB5A" box="[464,544,1133,1154]" pageId="10" pageNumber="21">MXAL</collectionCode>
|
||||
,
|
||||
<collectionCode id="ED10AE2D5D4D15232E13D022FDE6FB5A" box="[556,632,1133,1154]" collectionName="USA, Texas, College Station, Texas A & M University" pageId="10" pageNumber="21">TAMU</collectionCode>
|
||||
and
|
||||
<collectionCode id="ED10AE2D5D4D15232E8FD022FC9EFB5A" box="[688,768,1133,1154]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="10" pageNumber="21">ZMHU</collectionCode>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BBE36E85D4D15232D72D0C6FC96FB62" blockId="10.[307,1140,354,1658]" pageId="10" pageNumber="21">
|
||||
<emphasis id="B975EAFA5D4D15232D72D0C6FE6DFB46" bold="true" box="[333,499,1161,1182]" pageId="10" pageNumber="21">Type Locality.</emphasis>
|
||||
<collectingMunicipality id="6BDAAC925D4D15232DC1D0C6FD5FFB46" box="[510,705,1161,1182]" pageId="10" pageNumber="21">Rancho Quemado</collectingMunicipality>
|
||||
,
|
||||
<location id="8EDE60335D4D15232EEFD0C6FC0EFB46" LSID="urn:lsid:plazi:treatment:03A887FE5D4F15232D6CD0A1FBEAF9A2:8EDE60335D4D15232EEFD0C6FC0EFB46" box="[720,912,1161,1182]" country="Costa Rica" latitude="8.566667" longLatPrecision="1301" longitude="-83.583336" municipality="Rancho Quemado" name="Peninsula de Osa" pageId="10" pageNumber="21" stateProvince="Puntarenas">Península de Osa</location>
|
||||
, province of
|
||||
<collectingRegion id="49C5F80A5D4D15232818D0C6FEFAFB62" country="Costa Rica" name="Puntarenas" pageId="10" pageNumber="21">Puntarenas</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="F31676785D4D15232D4ED0EAFE7BFB62" box="[369,485,1189,1210]" name="Costa Rica" pageId="10" pageNumber="21">Costa Rica</collectingCountry>
|
||||
(aprox.
|
||||
<geoCoordinate id="EE35502F5D4D15232E7DD0EAFD11FB62" box="[578,655,1189,1210]" degrees="8" direction="north" minutes="34" orientation="latitude" pageId="10" pageNumber="21" precision="925" value="8.566667">8°34'N</geoCoordinate>
|
||||
;
|
||||
<geoCoordinate id="EE35502F5D4D15232EA3D0EAFD63FB62" box="[668,765,1189,1210]" degrees="83" direction="west" minutes="35" orientation="longitude" pageId="10" pageNumber="21" precision="925" value="-83.583336">83°35'W</geoCoordinate>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D4D15232D72D08EFC27FAA6" pageId="10" pageNumber="21" type="biology_ecology">
|
||||
<paragraph id="8BBE36E85D4D15232D72D08EFC27FAA6" blockId="10.[307,1140,354,1658]" pageId="10" pageNumber="21">
|
||||
<emphasis id="B975EAFA5D4D15232D72D08EFE61FB0E" bold="true" box="[333,511,1217,1238]" pageId="10" pageNumber="21">Biological Data.</emphasis>
|
||||
Males and females of
|
||||
<taxonomicName id="4C014D6B5D4D15232ED3D08DFC65FB0F" box="[748,1019,1218,1239]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="puntarenosa" status="sp. nov.">
|
||||
<emphasis id="B975EAFA5D4D15232ED3D08DFC65FB0F" box="[748,1019,1218,1239]" italics="true" pageId="10" pageNumber="21">Phyllophaga puntarenosa</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A24657815D4D1523283DD08EFBA4FB0E" box="[1026,1082,1217,1238]" pageId="10" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
|
||||
were collected at lights at localities between
|
||||
<quantity id="4CF99B0D5D4D15232EE4D092FCC2FB2A" box="[731,860,1245,1266]" metricMagnitude="2" metricUnit="m" metricValue="6.25" metricValueMax="12.0" metricValueMin="0.5" pageId="10" pageNumber="21" unit="m" value="625.0" valueMax="1200.0" valueMin="50.0">50–1,200 m</quantity>
|
||||
of altitude, including disturbed habitat as well as tropical deciduous forests. Phenology: February (18), April (34). Other species of
|
||||
<taxonomicName id="4C014D6B5D4D15232E54D159FD6FFAF3" box="[619,753,1302,1323]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B975EAFA5D4D15232E54D159FD6FFAF3" box="[619,753,1302,1323]" italics="true" pageId="10" pageNumber="21">Phyllophaga</emphasis>
|
||||
</taxonomicName>
|
||||
flying at the same time and places were:
|
||||
<emphasis id="B975EAFA5D4D15232D49D17DFE44FA9F" box="[374,474,1330,1351]" italics="true" pageId="10" pageNumber="21">P. favosa</emphasis>
|
||||
Saylor,
|
||||
<emphasis id="B975EAFA5D4D15232E09D17DFD2FFA9F" box="[566,689,1330,1351]" italics="true" pageId="10" pageNumber="21">P. gigantea</emphasis>
|
||||
Bates,
|
||||
<emphasis id="B975EAFA5D4D15232F3AD17DFC16FA9F" box="[773,904,1330,1351]" italics="true" pageId="10" pageNumber="21">P. hemilissa</emphasis>
|
||||
Bates,
|
||||
<emphasis id="B975EAFA5D4D15232FE4D17DFBEDFA9F" box="[987,1139,1330,1351]" italics="true" pageId="10" pageNumber="21">P. morganella</emphasis>
|
||||
Saylor,
|
||||
<emphasis id="B975EAFA5D4D15232DB9D101FD8BFABB" box="[390,533,1358,1379]" italics="true" pageId="10" pageNumber="21">P. menetriesi</emphasis>
|
||||
Blanchard,
|
||||
<emphasis id="B975EAFA5D4D15232EA2D101FCA6FABB" box="[669,824,1358,1379]" italics="true" pageId="10" pageNumber="21">P. schizorhina</emphasis>
|
||||
Bates,
|
||||
<emphasis id="B975EAFA5D4D15232FB0D101FC6EFABB" box="[911,1008,1358,1379]" italics="true" pageId="10" pageNumber="21">P. vicina</emphasis>
|
||||
Moser, one new species under description and four unidentified species.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D4D15232D72D1CAFD69F99A" pageId="10" pageNumber="21" type="discussion">
|
||||
<paragraph id="8BBE36E85D4D15232D72D1CAFD69F99A" blockId="10.[307,1140,354,1658]" pageId="10" pageNumber="21">
|
||||
<emphasis id="B975EAFA5D4D15232D72D1CAFE27FA42" bold="true" box="[333,441,1413,1434]" pageId="10" pageNumber="21">Remarks.</emphasis>
|
||||
<taxonomicName id="4C014D6B5D4D15232DFFD1C9FCF0FA43" box="[448,878,1413,1435]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="puntarenosa" status="sp. nov." subGenus="Phyllophaga">
|
||||
<emphasis id="B975EAFA5D4D15232DFFD1C9FDD8FA43" box="[448,582,1414,1435]" italics="true" pageId="10" pageNumber="21">Phyllophaga</emphasis>
|
||||
(
|
||||
<emphasis id="B975EAFA5D4D15232E69D1C9FD45FA43" box="[598,731,1414,1435]" italics="true" pageId="10" pageNumber="21">Phyllophaga</emphasis>
|
||||
)
|
||||
<emphasis id="B975EAFA5D4D15232ED4D1C9FCF0FA43" box="[747,878,1414,1435]" italics="true" pageId="10" pageNumber="21">puntarenosa</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A24657815D4D15232F4AD1CAFC38FA42" box="[885,934,1413,1434]" pageId="10" pageNumber="21" rank="species">n.sp.</taxonomicNameLabel>
|
||||
belongs to the species group ‘‘
|
||||
<emphasis id="B975EAFA5D4D15232D85D1EDFD83FA6F" box="[442,541,1442,1463]" italics="true" pageId="10" pageNumber="21">rorulenta</emphasis>
|
||||
’’ (
|
||||
<emphasis id="B975EAFA5D4D15232E00D1EDFDE7FA6F" box="[575,633,1442,1463]" italics="true" pageId="10" pageNumber="21">sensu</emphasis>
|
||||
<bibRefCitation id="EF904B195D4D15232EBDD1EEFC94FA6E" author="Moron" box="[642,778,1441,1462]" firstAuthor="Moron" pageId="10" pageNumber="21" refId="ref11224" refString="Moron, M. A. 1986. El genero Phyllophaga en Mexico. Morfologia, distribucion y sistematica supraespecifica (Insecta: Coleoptera). Publ. 20, Instituto de Ecologia, Mexico, D. F. 341 pp." type="book" year="1986">Morón 1986</bibRefCitation>
|
||||
). It may be related to
|
||||
<emphasis id="B975EAFA5D4D15232FC1D1EDFB8BFA6F" box="[1022,1045,1442,1463]" italics="true" pageId="10" pageNumber="21">P.</emphasis>
|
||||
(
|
||||
<taxonomicName id="4C014D6B5D4D15232819D1EDFEEAFA0B" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B975EAFA5D4D15232819D1EDFEEAFA0B" italics="true" pageId="10" pageNumber="21">Phyllophaga</emphasis>
|
||||
</taxonomicName>
|
||||
)
|
||||
<emphasis id="B975EAFA5D4D15232DB9D1F1FE6DFA0B" box="[390,499,1470,1491]" italics="true" pageId="10" pageNumber="21">chiriquina</emphasis>
|
||||
(Bates) (
|
||||
<collectingCountry id="F31676785D4D15232E69D1F2FD5DFA0A" box="[598,707,1469,1490]" name="Nicaragua" pageId="10" pageNumber="21">Nicaragua</collectingCountry>
|
||||
to
|
||||
<collectingCountry id="F31676785D4D15232ED5D1F2FCCBFA0A" box="[746,853,1469,1490]" name="Colombia" pageId="10" pageNumber="21">Colombia</collectingCountry>
|
||||
;
|
||||
<bibRefCitation id="EF904B195D4D15232F5ED1F2FC7AFA0A" author="Saylor, L. W." box="[865,996,1469,1490]" journalOrPublisher="Proceedings Biological Society Washington" pageId="10" pageNumber="21" pagination="129 - 142" part="56" refId="ref11301" refString="Saylor, L. W. 1943. Revision of the rorulenta group of the scarab beetle genus Phyllophaga. Proceedings Biological Society Washington, 56: 129 - 142." title="Revision of the rorulenta group of the scarab beetle genus Phyllophaga" type="journal article" year="1943">Saylor 1943</bibRefCitation>
|
||||
), by the general shape of the apical half of parameres, but the bicolorate dorsum, lenght of male antennal club, the flattened female pygidial disk, the lenght of the lower male metatibial spur, and the apex of parameres sharply pointed, will aid in the recognition of this new species.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31B65635D4D15232D72D206FBEAF9A2" pageId="10" pageNumber="21" type="etymology">
|
||||
<paragraph id="8BBE36E85D4D15232D72D206FBEAF9A2" blockId="10.[307,1140,354,1658]" pageId="10" pageNumber="21">
|
||||
<emphasis id="B975EAFA5D4D15232D72D206FE52F986" bold="true" box="[333,460,1609,1630]" pageId="10" pageNumber="21">Etymology.</emphasis>
|
||||
Derived from combination of words ‘‘Puntarenas’’ province and ‘‘Osa’’ peninsula, that comprise the geographic range where this species lives.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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