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<document id="8E84DA38C280657A17BA4054A156803C" ID-DOI="10.1649/0010-065x(2001)055[0011:snsops]2.0.co;2" ID-ISSN="1938-4394" ID-Zenodo-Dep="4900311" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1622827801739" checkinUser="carolina" docAuthor="Morón, Miguel Angel &amp; Solís, Angel" docDate="2001" docId="03A887FE5D4115212D5AD18CFE1BFB08" docLanguage="en" docName="TheColeopBull.55.1.11-29.pdf.imf" docOrigin="The Coleopterists Bulletin 55 (1)" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Phyllophaga (Phyllophaga) lorencita Moron and Solis, new species" docType="treatment" docVersion="1" lastPageNumber="19" masterDocId="FF91FF865D4715292C3FD44FFF9EFFD8" masterDocTitle="Seven New Species Of Phyllophaga (S. Str.) Harris From Costa Rica (Coleoptera: Melolonthidae: Melolonthinae)" masterLastPageNumber="29" masterPageNumber="11" pageNumber="17" updateTime="1730139982923" updateUser="juliana" zenodo-license-document="CLOSED">
<mods:mods id="0A88CB565590A489634EBE27ECEBA458" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="90476B19F9B074935AD2CE7AA69E5163">
<mods:title id="76B3913B7D2CB7078D56AF7DBFC90EAD">Seven New Species Of Phyllophaga (S. Str.) Harris From Costa Rica (Coleoptera: Melolonthidae: Melolonthinae)</mods:title>
</mods:titleInfo>
<mods:name id="EAC7566204CB2A3DD53B05849A4E9B08" type="personal">
<mods:role id="F1BF0A4D834E508907167E132BF540F0">
<mods:roleTerm id="0A3B714C50019515F18C68B0573F214D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="13BCB34F92F614564974FC09109431A5">Morón, Miguel Angel</mods:namePart>
<mods:affiliation id="0B28448CFBB0A4DDF3AECD8BF85D45B1">Departamento de Entomología Instituto de Ecología, A. C. (sector SEP­CONACYT) Apartado Postal 63 Xalapa, Veracruz 91000, MÉXICO</mods:affiliation>
</mods:name>
<mods:name id="7A784DF93870C5982156E3C5E7AFC885" type="personal">
<mods:role id="70D2E2268D64E6DBDE43B6442A905DBD">
<mods:roleTerm id="7792992FBC6C1BC504EF5FC1C8097F65">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="401404D22739A1C4064841C8457D5189">Solís, Angel</mods:namePart>
<mods:affiliation id="B549CBE0B6ECD45B96FB5B39C80850A4">Departamento de Entomología Instituto Nacional de Biodiversidad (INBio) Apartado Postal 22 ­ 3100 Santo Domingo de Heredia, COSTA RICA</mods:affiliation>
</mods:name>
<mods:typeOfResource id="B35F22929D9113751F369D4453D73455">text</mods:typeOfResource>
<mods:relatedItem id="E6FEC3887703DAD4BCFD9CC12EB55C1E" type="host">
<mods:titleInfo id="3CF98365A823C971C48C1809184B6013">
<mods:title id="DA6517591C6D9FE5ED5FEBB51FCAEBA2">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="CC6CAD1AAB544D1CC5D73AD75BF8DD05">
<mods:date id="7E6D81007FF2F8FC9A99C19EAFDCBFE8">2001</mods:date>
<mods:detail id="8C23B556D6ED9CBCBA25838613821E57" type="pubDate">
<mods:number id="B4E24C1747A92108E03848688D92CF12">2001-03-01</mods:number>
</mods:detail>
<mods:detail id="8C454AA897222A449FF4A61C27B25CBF" type="volume">
<mods:number id="03B6BD35C407B37ED6EDA689ACF2DDAD">55</mods:number>
</mods:detail>
<mods:detail id="0FBF85D794ED58DD8E8F7440EEB2181C" type="issue">
<mods:number id="7B0BA1A472C1BAA9347EDA0BA1C79B34">1</mods:number>
</mods:detail>
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<mods:start id="64E678BE44882A371CBBA37096142B1F">11</mods:start>
<mods:end id="CFEE8C7D326E4C8769C02F6863FC0C10">29</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="93A2E7E602945F1048584187099C4369">journal article</mods:classification>
<mods:identifier id="0889FDE1B1B91A8B55A4E4719082CD42" type="DOI">10.1649/0010-065x(2001)055[0011:snsops]2.0.co;2</mods:identifier>
<mods:identifier id="64096A2F71077D6FE6C8FF1EC78349A9" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="95C90C8169C61DF6F5D6C0BB82CB4102" type="Zenodo-Dep">4900311</mods:identifier>
</mods:mods>
<treatment id="03A887FE5D4115212D5AD18CFE1BFB08" LSID="urn:lsid:plazi:treatment:03A887FE5D4115212D5AD18CFE1BFB08" httpUri="http://treatment.plazi.org/id/03A887FE5D4115212D5AD18CFE1BFB08" lastPageId="8" lastPageNumber="19" pageId="6" pageNumber="17">
<subSubSection id="C31B65635D41152F2D5AD18CFBDEFA0F" box="[357,1088,1474,1496]" pageId="6" pageNumber="17" type="nomenclature">
<paragraph id="8BBE36E85D41152F2D5AD18CFBDEFA0F" blockId="6.[357,1088,1474,1522]" box="[357,1088,1474,1496]" pageId="6" pageNumber="17">
<taxonomicName id="4C014D6B5D41152F2D5AD18CFC2FFA00" authority="Moron and Solis" authorityName="Moron and Solis" box="[357,945,1475,1496]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="lorencita" status="sp. nov." subGenus="Phyllophaga">
<emphasis id="B975EAFA5D41152F2D5AD18CFE75FA00" box="[357,491,1475,1496]" italics="true" pageId="6" pageNumber="17">Phyllophaga</emphasis>
(
<emphasis id="B975EAFA5D41152F2DC2D18CFD1CFA00" box="[509,642,1475,1496]" italics="true" pageId="6" pageNumber="17">Phyllophaga</emphasis>
)
<emphasis id="B975EAFA5D41152F2EABD18CFD6CFA00" box="[660,754,1475,1496]" italics="true" pageId="6" pageNumber="17">lorencita</emphasis>
Morón and Solís
</taxonomicName>
,
<emphasis id="B975EAFA5D41152F2F82D18DFBDEFA0F" bold="true" box="[957,1088,1474,1495]" pageId="6" pageNumber="17">
<taxonomicNameLabel id="A24657815D41152F2F82D18DFBDEFA0F" box="[957,1088,1474,1495]" pageId="6" pageNumber="17" rank="species">new species</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C31B65635D4115212EACD192FE12FE68" lastPageId="8" lastPageNumber="19" pageId="6" pageNumber="17" type="description">
<paragraph id="8BBE36E85D41152F2EACD192FC8CFA2A" blockId="6.[357,1088,1474,1522]" box="[659,786,1501,1522]" pageId="6" pageNumber="17">
<figureCitation id="133A2A6D5D41152F2EACD192FC8CFA2A" box="[659,786,1501,1522]" captionStart="Figs" captionStartId="6.[330,373,1116,1135]" captionTargetBox="[336,1117,273,1093]" captionTargetId="figure-12@6.[328,1117,272,1096]" captionTargetPageId="6" captionText="Figs. 1013. Phyllophaga lorencita. 10) protarsal claw, male; 11) genital capsule, lateral view; 12) paramera, ventro­distal view; 13) genital plates, female. Scale lines 5 1 mm, except Fig. 10 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900317" httpUri="https://zenodo.org/record/4900317/files/figure.png" pageId="6" pageNumber="17">Figs. 1013</figureCitation>
</paragraph>
<paragraph id="8BBE36E85D41152E2D75D248FC2FFB35" blockId="6.[307,1140,1543,1658]" lastBlockId="7.[279,1113,277,1261]" lastPageId="7" lastPageNumber="18" pageId="6" pageNumber="17">
<emphasis id="B975EAFA5D41152F2D75D248FE34F9C2" bold="true" box="[330,426,1543,1562]" pageId="6" pageNumber="17">
<typeStatus id="54BA884A5D41152F2D75D248FE38F9C2" box="[330,422,1543,1562]" pageId="6" pageNumber="17" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male. Clypeus, frons and pronotum shiny reddish yellow; elytra shiny testaceus, with two erect long setae at the base of each elytron and row of setae along the elytral suture, setae located toward the apex of elytra longer than preceding elytral setae; mouthparts, sterna, pygidium and legs shiny testaceous. Clypeus 4.2× wider than long, anterior border scarcely sinuate, with elevated margin, surface scarcely concave, coarsely rugo­punctate and with some scattered short setae. Frontoclypeal suture straight and clearly impressed. Frons 1.3× wider than long, convex, coarsely punctate rugose, with short setae on disk. Antenna 9­segmented, with 3­segmented club, lamellae of 7th to 9th segments 1.2× longer than length of preceeding 5 segments combined. Frons 3.2× wider than dorsal diameter of eye. Eye canthus long and narrowed, with 810 setae. Labrum bilobed, wide and shallowly sinuated, with scattered slender setae. Mentum slightly convex, finely punctate, with long slender setae at sides, anterior border shallowly notched. Pronotum 1.8× wider than long and 2.7× wider than frons. Pronotal disk shiny, moderately punctate­rugose, punctures irregularly separated from one another by 15 diameters, midline with shallow longitudinal depression spaced anteriorly; lateral borders clearly angled, lateral marginal bead irregularly crenulate, with scattered, long, slender setae; complete basal bead indicated by punctures; anterior angles slightly acute, prominent; posterior angles widely obtuse, rounded. Scutellum 1.7× wider than long, with some small punctures. Elytron 2.5× longer than wide, shiny, densely punctate; epipleural border very narrow, extended along complete margin, provided with a row of very short, slender setae; humeral callus rounded, prominent; apical callus rounded. Metathoracic wings completely developed. Propygidium shiny, densely punctate­setiferous. Pygidium scarcely convex, shiny, rugo­punctate, with round punctures, with very short setae covering entire surface; apical margin with 16 long, slender setae; basal margin effaced medially. Pterosterna with long, dense, yellowish setae. Visible abdominal sternites II and IV slightly depressed, smooth and setiferous near the middline; sternite V granulate setiferous, with a shallow transverse depression on posterior half and some scattered short setae toward the sides; anal plate narrowed with a shallow transverse depression, with posterior border elevate and notched at midline, with 20 erect setae. Protibia slightly shorter than protarsus (1:1.3), with external border clearly tridentate, preapical spur small, slighly curved, acute, shorter than 2nd protarsomere. Mesotibia with one oblique, sharp, setiferous carina on external side; upper apical spur straight, narrow, and 0.1× shorter than lower spur. Metatibia shorter than metatarsus (1:1.2), with one oblique, sharp, setiferous carina on external side; upper apical spur articulated, curved, lanceolate, apex rounded, as long as basal metatarsomere, and 1.3× longer than lower spur; lower apical spur articulated with tibial border, with rounded apex. Tarsomeres semicylindrical, elongate, with enlarged apex, some setae apically and one longitudinal thin carina with a line of scattered, stout setae and other line of small setae at side. Protarsomere 14 with two carinae all along the ventral side. Tarsal claws symmetrical, similar on all legs, with curved tooth on the ventral border, slightly displaced toward the basal process (
<figureCitation id="133A2A6D5D40152E2838D006FBD3FB84" box="[1031,1101,1097,1116]" captionStart="Figs" captionStartId="6.[330,373,1116,1135]" captionTargetBox="[336,1117,273,1093]" captionTargetId="figure-12@6.[328,1117,272,1096]" captionTargetPageId="6" captionText="Figs. 1013. Phyllophaga lorencita. 10) protarsal claw, male; 11) genital capsule, lateral view; 12) paramera, ventro­distal view; 13) genital plates, female. Scale lines 5 1 mm, except Fig. 10 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900317" httpUri="https://zenodo.org/record/4900317/files/figure.png" pageId="7" pageNumber="18">Fig. 10</figureCitation>
). Genital capsule with short parameres, dorsally and ventrally fused, preapical inner borders of the parameres turned out as a blade­like structure, apex with very short, toothlike ventral projections. Aedeagus short, with the apex of sclerotized support bifurcate and dorso­basal crest­like brush of progressively enlarged setae (
<figureCitation id="133A2A6D5D40152E2FB4D0E5FB9EFB65" box="[907,1024,1194,1213]" captionStart="Figs" captionStartId="6.[330,373,1116,1135]" captionTargetBox="[336,1117,273,1093]" captionTargetId="figure-12@6.[328,1117,272,1096]" captionTargetPageId="6" captionText="Figs. 1013. Phyllophaga lorencita. 10) protarsal claw, male; 11) genital capsule, lateral view; 12) paramera, ventro­distal view; 13) genital plates, female. Scale lines 5 1 mm, except Fig. 10 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900317" httpUri="https://zenodo.org/record/4900317/files/figure.png" pageId="7" pageNumber="18">Figs. 1112</figureCitation>
). Tectum slighly convex. Length of genital capsule from apex of parameres to border of basal piece:
<quantity id="4CF99B0D5D40152E2D68D095FE3DFB35" box="[343,419,1242,1261]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="7" pageNumber="18" unit="mm" value="3.9">3.9 mm</quantity>
. Total body length:
<quantity id="4CF99B0D5D40152E2E5CD095FD25FB35" box="[611,699,1242,1261]" metricMagnitude="-2" metricUnit="m" metricValue="1.58" pageId="7" pageNumber="18" unit="mm" value="15.8">15.8 mm</quantity>
. Humeral width:
<quantity id="4CF99B0D5D40152E2F5ED095FC33FB35" box="[865,941,1242,1261]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="7" pageNumber="18" unit="mm" value="6.6">6.6 mm</quantity>
.
</paragraph>
<paragraph id="8BBE36E85D40152E2D0DD148FDBFF9B8" blockId="7.[279,1112,1287,1658]" pageId="7" pageNumber="18">
<emphasis id="B975EAFA5D40152E2D0DD148FE09FAC4" bold="true" box="[306,407,1287,1308]" pageId="7" pageNumber="18">
<typeStatus id="54BA884A5D40152E2D0DD148FE0CFAC4" box="[306,402,1287,1308]" pageId="7" pageNumber="18" type="allotype">Allotype</typeStatus>
.
</emphasis>
Female. Similar to the male except as follows: anterior border of frons elevated, disk of frons coarsely rugo­punctate; antennae with lamellae of 7th to 9th segments shorter than the length of five preceeding ones (1:1.2). The longitudinal depression at the anterior part of the midline deeper than male; and also with coarser punctures. Visible abdominal sternites V convex, densely setiferous punctuated mainly toward the sides; anal plate 1.3× longer than male anal plate, convex, punctate setiferous, with 16 long slender setae near the posterior border. Meso­ and metatibiae each with one oblique, sharp, setiferous carina on external side. Both apical spurs of metatibia articulated, wide, lanceolate and slightly curved. Ventral genital plates slightly sclerotized, symmetrical, widened, ovate, without setae; dorsal genital plates narrowed, with short setae on the rounded apex (
<figureCitation id="133A2A6D5D40152E2E84D27FFC92F99D" box="[699,780,1584,1605]" captionStart="Figs" captionStartId="6.[330,373,1116,1135]" captionTargetBox="[336,1117,273,1093]" captionTargetId="figure-12@6.[328,1117,272,1096]" captionTargetPageId="6" captionText="Figs. 1013. Phyllophaga lorencita. 10) protarsal claw, male; 11) genital capsule, lateral view; 12) paramera, ventro­distal view; 13) genital plates, female. Scale lines 5 1 mm, except Fig. 10 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900317" httpUri="https://zenodo.org/record/4900317/files/figure.png" pageId="7" pageNumber="18">Fig. 13</figureCitation>
). Total body length:
<quantity id="4CF99B0D5D40152E2FCED27FFBCAF99D" box="[1009,1108,1584,1605]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="7" pageNumber="18" unit="mm" value="16.2">16.2 mm</quantity>
. Humeral width: 7.0 mm.
</paragraph>
<paragraph id="8BBE36E85D4015212D0DD22AFE12FE68" blockId="7.[279,1112,1287,1658]" lastBlockId="8.[307,1140,277,1232]" lastPageId="8" lastPageNumber="19" pageId="7" pageNumber="18">
<emphasis id="B975EAFA5D40152E2D0DD22AFD8FF9A2" bold="true" box="[306,529,1637,1658]" pageId="7" pageNumber="18">
<typeStatus id="54BA884A5D40152E2D0DD22AFE06F9A2" box="[306,408,1637,1658]" pageId="7" pageNumber="18" type="paratype">Paratype</typeStatus>
Variation.
</emphasis>
Males. Similar to
<typeStatus id="54BA884A5D40152E2EEAD22AFCAEF9A2" box="[725,816,1637,1658]" pageId="7" pageNumber="18" type="holotype">holotype</typeStatus>
except in total body length:
<quantity id="4CF99B0D5D4F15212D0CD55AFE4AFEF2" box="[307,468,277,298]" metricMagnitude="-2" metricUnit="m" metricValue="1.3800000000000001" metricValueMax="1.6199999999999999" metricValueMin="1.1400000000000001" pageId="8" pageNumber="19" unit="mm" value="13.8" valueMax="16.2" valueMin="11.4">11.416.2 mm</quantity>
, humeral width: 4.96.0 mm, pronotum of some specimens darker or lighter than
<typeStatus id="54BA884A5D4F15212E1ED57FFDE1FE9D" box="[545,639,304,325]" pageId="8" pageNumber="19" type="holotype">holotype</typeStatus>
, other specimens with antennal club 1.01.2× longer than length of preceeding five segments combined. Females similar to
<typeStatus id="54BA884A5D4F15212D0CD52AFE19FEA2" box="[307,391,357,378]" pageId="8" pageNumber="19" type="allotype">allotype</typeStatus>
except as follows: anterior border of frons less prominent, body punctation coarser or finer, total body length:
<quantity id="4CF99B0D5D4F15212ED0D5CFFC10FE4D" box="[751,910,384,405]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.61" metricValueMin="1.29" pageId="8" pageNumber="19" unit="mm" value="14.5" valueMax="16.1" valueMin="12.9">12.916.1 mm</quantity>
; humeral width:
<quantity id="4CF99B0D5D4F1521287AD5CFFE16FE68" metricMagnitude="-3" metricUnit="m" metricValue="5.8" metricValueMax="6.5" metricValueMin="5.1" pageId="8" pageNumber="19" unit="mm" value="5.8" valueMax="6.5" valueMin="5.1">5.1 6.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C31B65635D4F15212D72D5FAFD3EFD0D" pageId="8" pageNumber="19" type="materials_examined">
<paragraph id="8BBE36E85D4F15212D72D5FAFD8FFD78" blockId="8.[307,1140,277,1232]" pageId="8" pageNumber="19">
<emphasis id="B975EAFA5D4F15212D72D5FAFE49FE12" bold="true" box="[333,471,437,458]" pageId="8" pageNumber="19">Type Series.</emphasis>
<location id="8EDE60335D4F15212DE0D5FAFDD7FE12" LSID="urn:lsid:plazi:treatment:03A887FE5D4115212D5AD18CFE1BFB08:8EDE60335D4F15212DE0D5FAFDD7FE12" box="[479,585,437,458]" name="Described" pageId="8" pageNumber="19">Described</location>
from
<specimenCount id="9D07FD615D4F15212EB3D5FAFD75FE12" box="[652,747,437,458]" pageId="8" pageNumber="19" type="male">67 males</specimenCount>
and
<specimenCount id="9D07FD615D4F15212F1ED5FAFC06FE12" box="[801,920,437,458]" pageId="8" pageNumber="19" type="female">25 females</specimenCount>
.
<materialsCitation id="3B693CB55D4F15212F9DD5FAFD78FDD8" collectingDate="1987-05-04" collectionCode="IN- BIO" collectorName="A. Solis" country="Costa Rica" county="San Ramon" elevation="800" latitude="10.35" location="Rio San Lorencito" longLatPrecision="1298" longitude="-84.416664" municipality="Rio San Lorencito" pageId="8" pageNumber="19" specimenCode="INBIO CR 1001 115762" specimenCount="1" specimenCount-male="1" stateProvince="Alajuela" typeStatus="holotype">
<typeStatus id="54BA884A5D4F15212F9DD5FAFB9DFE12" box="[930,1027,437,458]" pageId="8" pageNumber="19" type="holotype">Holotype</typeStatus>
<specimenCount id="9D07FD615D4F15212834D5FAFBA0FE12" box="[1035,1086,437,458]" pageId="8" pageNumber="19" type="male">male</specimenCount>
(
<collectionCode id="ED10AE2D5D4F15212872D5FAFEFBFE3D" pageId="8" pageNumber="19">INBIO</collectionCode>
):
<collectingCountry id="F31676785D4F15212DBCD59FFDB5FE3D" box="[387,555,464,485]" name="Costa Rica" pageId="8" pageNumber="19">COSTA RICA</collectingCountry>
:
<collectingRegion id="49C5F80A5D4F15212E07D59FFD0DFE3D" box="[568,659,464,485]" country="Costa Rica" name="Alajuela" pageId="8" pageNumber="19">Alajuela</collectingRegion>
,
<collectingCounty id="62DF4E645D4F15212E9DD59FFCBFFE3D" box="[674,801,464,485]" pageId="8" pageNumber="19">San Ramón</collectingCounty>
,
<location id="8EDE60335D4F15212F10D59FFC66FE3D" LSID="urn:lsid:plazi:treatment:03A887FE5D4115212D5AD18CFE1BFB08:8EDE60335D4F15212F10D59FFC66FE3D" box="[815,1016,464,485]" country="Costa Rica" county="San Ramon" municipality="Rio San Lorencito" name="Rio San Lorencito" pageId="8" pageNumber="19" stateProvince="Alajuela">Río San Lorencito</location>
,
<quantity id="4CF99B0D5D4F15212838D59FFBD1FE3D" box="[1031,1103,464,485]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="19" unit="m" value="800.0">
<elevation id="002CD1DB5D4F15212838D59FFBD1FE3D" box="[1031,1103,464,485]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="19" unit="m" value="800.0">800 m</elevation>
</quantity>
,
<date id="FFBF10285D4F15212862D59FFEF7FDD8" pageId="8" pageNumber="19" value="1987-05-04">
<collectingDate id="EFFBE9C05D4F15212862D59FFEF7FDD8" pageId="8" pageNumber="19" value="1987-05-04">4­ V­87</collectingDate>
</date>
,
<collectorName id="26F4533E5D4F15212D4BD5A4FE52FDD8" box="[372,460,491,512]" pageId="8" pageNumber="19">A. Solís</collectorName>
,
<specimenCode id="DBA79E935D4F15212DE8D5A4FD7CFDD8" box="[471,738,491,512]" pageId="8" pageNumber="19">INBIO CR 1001 115762</specimenCode>
.
</materialsCitation>
<materialsCitation id="3B693CB55D4F15212ED9D5A4FC63FDED" collectingDate="1993-06-30" collectionCode="INBIO" collectorName="P. Campos" country="Costa Rica" county="San Ramon" elevation="1120" latitude="10.35" location="Chirripo" longLatPrecision="1298" longitude="-84.416664" municipality="Turrialba" pageId="8" pageNumber="19" specimenCode="INBIO CR 1001 848913" specimenCount="1" specimenCount-female="1" stateProvince="Cartago" typeStatus="allotype">
<typeStatus id="54BA884A5D4F15212EC8D5A4FCCCFDD8" box="[759,850,491,512]" pageId="8" pageNumber="19" type="allotype">Allotype</typeStatus>
<specimenCount id="9D07FD615D4F15212F66D5A4FC3EFDD8" box="[857,928,491,512]" pageId="8" pageNumber="19" type="female">female</specimenCount>
(
<collectionCode id="ED10AE2D5D4F15212F90D5A4FC60FDD8" box="[943,1022,491,512]" pageId="8" pageNumber="19">INBIO</collectionCode>
):
<collectingCountry id="F31676785D4F15212827D5A4FEE8FDC2" name="Costa Rica" pageId="8" pageNumber="19">COSTA RICA</collectingCountry>
:
<collectingRegion id="49C5F80A5D4F15212DBFD64AFE4BFDC2" box="[384,469,517,538]" country="Costa Rica" name="Cartago" pageId="8" pageNumber="19">Cartago</collectingRegion>
,
<collectingMunicipality id="6BDAAC925D4F15212DDDD64AFDDAFDC2" box="[482,580,517,538]" pageId="8" pageNumber="19">Turrialba</collectingMunicipality>
,
<location id="8EDE60335D4F15212E6FD64AFD37FDC2" LSID="urn:lsid:plazi:treatment:03A887FE5D4115212D5AD18CFE1BFB08:8EDE60335D4F15212E6FD64AFD37FDC2" box="[592,681,517,538]" country="Costa Rica" county="San Ramon" municipality="Turrialba" name="Chirripo" pageId="8" pageNumber="19" stateProvince="Cartago">Chirripó</location>
,
<location id="8EDE60335D4F15212E88D64AFB98FDC2" LSID="urn:lsid:plazi:treatment:03A887FE5D4115212D5AD18CFE1BFB08:8EDE60335D4F15212E88D64AFB98FDC2" box="[695,1030,517,538]" country="Costa Rica" county="San Ramon" municipality="Turrialba" name="Area de Conservacion Amistad" pageId="8" pageNumber="19" stateProvince="Cartago">Area de Conservación Amistad</location>
,
<location id="8EDE60335D4F1521282ED64AFEC0FDED" LSID="urn:lsid:plazi:treatment:03A887FE5D4115212D5AD18CFE1BFB08:8EDE60335D4F1521282ED64AFEC0FDED" country="Costa Rica" county="San Ramon" municipality="Turrialba" name="Grano de Oro" pageId="8" pageNumber="19" stateProvince="Cartago">Grano de Oro</location>
,
<quantity id="4CF99B0D5D4F15212D54D66FFE58FDED" box="[363,454,544,565]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="8" pageNumber="19" unit="m" value="1120.0">
<elevation id="002CD1DB5D4F15212D54D66FFE58FDED" box="[363,454,544,565]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="8" pageNumber="19" unit="m" value="1120.0">1,120 m</elevation>
</quantity>
, 19/
<date id="FFBF10285D4F15212DCAD66FFDC7FDED" box="[501,601,544,565]" pageId="8" pageNumber="19" value="1993-06-30">
<collectingDate id="EFFBE9C05D4F15212DCAD66FFDC7FDED" box="[501,601,544,565]" pageId="8" pageNumber="19" value="1993-06-30">30­VI­93</collectingDate>
</date>
,
<collectorName id="26F4533E5D4F15212E5AD66FFD4BFDED" box="[613,725,544,565]" pageId="8" pageNumber="19">P. Campos</collectorName>
<specimenCode id="DBA79E935D4F15212EE1D66FFC6EFDED" box="[734,1008,544,565]" pageId="8" pageNumber="19">INBIO CR 1001 848913</specimenCode>
.
</materialsCitation>
<materialsCitation id="3B693CB55D4F15212839D66FFD13FD88" collectionCode="CASC, CNC, INBIO, MXAL, TAMU, ZMHU" collectorName="A. Solis" country="Costa Rica" county="San Ramon" elevation="800" latitude="10.35" location="Turrialba" longLatPrecision="1298" longitude="-84.416664" municipality="Rio San Lorencito" pageId="8" pageNumber="19" specimenCount="2" specimenCount-female="24" specimenCount-male="2" stateProvince="Alajuela" typeStatus="paratype">
<typeStatus id="54BA884A5D4F15212839D66FFBEEFDED" box="[1030,1136,544,565]" pageId="8" pageNumber="19" type="paratype">Paratypes</typeStatus>
: same data as holotype (
<specimenCount id="9D07FD615D4F15212E14D674FD1FFD88" box="[555,641,571,592]" count="2" pageId="8" pageNumber="19" type="male">2 males</specimenCount>
);
</materialsCitation>
<materialsCitation id="3B693CB55D4F15212EABD674FBEDFD88" box="[660,1139,571,592]" collectingDate="1993-06-30" collectionCode="CASC, CNC, INBIO, MXAL, TAMU, ZMHU" collectorName="P. Campos" country="Costa Rica" county="San Ramon" elevation="800" latitude="10.35" location="Turrialba" longLatPrecision="1298" longitude="-84.416664" municipality="Rio San Lorencito" pageId="8" pageNumber="19" specimenCode="INBIO CR 1001 848913" specimenCount="53" specimenCount-female="14" specimenCount-male="39" stateProvince="Alajuela" typeStatus="paratype">
same data as allotype (
<specimenCount id="9D07FD615D4F15212FBBD674FC78FD88" box="[900,998,571,592]" count="39" pageId="8" pageNumber="19" type="male">39 males</specimenCount>
,
<specimenCount id="9D07FD615D4F15212FCED674FBF6FD88" box="[1009,1128,571,592]" count="14" pageId="8" pageNumber="19" type="female">14 females</specimenCount>
);
</materialsCitation>
<materialsCitation id="3B693CB55D4F15212D0CD61AFD93FD5D" collectingDate="1989-07-10" collectionCode="CASC, CNC, INBIO, MXAL, TAMU, ZMHU" collectorName="A. Solis" country="Costa Rica" county="San Ramon" elevation="800" location="Rio San Lorencito" municipality="San Ramon" pageId="8" pageNumber="19" specimenCount="35" specimenCount-female="10" specimenCount-male="25" stateProvince="Alajuela" typeStatus="holotype">
<collectingRegion id="49C5F80A5D4F15212D0CD61AFE10FDB2" box="[307,398,597,618]" country="Costa Rica" name="Alajuela" pageId="8" pageNumber="19">Alajuela</collectingRegion>
,
<collectingMunicipality id="6BDAAC925D4F15212DA0D61AFDBFFDB2" box="[415,545,597,618]" pageId="8" pageNumber="19">San Ramón</collectingMunicipality>
,
<location id="8EDE60335D4F15212E0ED61AFC9EFDB2" LSID="urn:lsid:plazi:treatment:03A887FE5D4115212D5AD18CFE1BFB08:8EDE60335D4F15212E0ED61AFC9EFDB2" box="[561,768,597,618]" country="Costa Rica" county="San Ramon" municipality="San Ramon" name="Rio San Lorencito" pageId="8" pageNumber="19" stateProvince="Alajuela">Río San Lorencito</location>
,
<quantity id="4CF99B0D5D4F15212F2FD61AFCC5FDB2" box="[784,859,597,618]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="19" unit="m" value="800.0">
<elevation id="002CD1DB5D4F15212F2FD61AFCC5FDB2" box="[784,859,597,618]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="19" unit="m" value="800.0">800 m</elevation>
</quantity>
,
<date id="FFBF10285D4F15212F53D61AFC46FDB2" box="[876,984,597,618]" pageId="8" pageNumber="19" value="1989-07-10">
<collectingDate id="EFFBE9C05D4F15212F53D61AFC46FDB2" box="[876,984,597,618]" pageId="8" pageNumber="19" value="1989-07-10">10­VII­89</collectingDate>
</date>
,
<collectorName id="26F4533E5D4F15212FD6D61AFBDDFDB2" box="[1001,1091,597,618]" pageId="8" pageNumber="19">A. Solís</collectorName>
(
<specimenCount id="9D07FD615D4F15212867D61AFEEDFD5D" count="25" pageId="8" pageNumber="19" type="male">25 males</specimenCount>
,
<specimenCount id="9D07FD615D4F15212DBBD63FFD9CFD5D" box="[388,514,624,645]" count="10" pageId="8" pageNumber="19" type="female">10 females</specimenCount>
).
</materialsCitation>
Paratypes deposited in
<collectionCode id="ED10AE2D5D4F15212F1ED63FFCF6FD5D" box="[801,872,624,645]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="8" pageNumber="19">CASC</collectionCode>
,
<collectionCode id="ED10AE2D5D4F15212F45D63FFC2CFD5D" box="[890,946,624,645]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" pageId="8" pageNumber="19">CNC</collectionCode>
,
<collectionCode id="ED10AE2D5D4F15212FFCD63FFB90FD5D" box="[963,1038,624,645]" pageId="8" pageNumber="19">INBIO</collectionCode>
,
<collectionCode id="ED10AE2D5D4F15212820D63FFBF1FD5D" box="[1055,1135,624,645]" pageId="8" pageNumber="19">MXAL</collectionCode>
,
<collectionCode id="ED10AE2D5D4F15212D0CD6C4FE1CFD78" box="[307,386,651,672]" collectionName="USA, Texas, College Station, Texas A &amp; M University" pageId="8" pageNumber="19">TAMU</collectionCode>
, and
<collectionCode id="ED10AE2D5D4F15212D82D6C4FD93FD78" box="[445,525,651,672]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="8" pageNumber="19">ZMHU</collectionCode>
.
</paragraph>
<paragraph id="8BBE36E85D4F15212D72D6EAFD3EFD0D" blockId="8.[307,1140,277,1232]" pageId="8" pageNumber="19">
<emphasis id="B975EAFA5D4F15212D72D6EAFE6EFD62" bold="true" box="[333,496,677,698]" pageId="8" pageNumber="19">Type Locality.</emphasis>
<location id="8EDE60335D4F15212DC8D6EAFD22FD62" LSID="urn:lsid:plazi:treatment:03A887FE5D4115212D5AD18CFE1BFB08:8EDE60335D4F15212DC8D6EAFD22FD62" box="[503,700,677,698]" country="Costa Rica" latitude="10.35" longLatPrecision="1298" longitude="-84.416664" municipality="San Ramon" name="Rio San Lorencito" pageId="8" pageNumber="19" stateProvince="Alajuela">Río San Lorencito</location>
,
<collectingMunicipality id="6BDAAC925D4F15212EF7D6EAFCDBFD62" box="[712,837,677,698]" pageId="8" pageNumber="19">San Ramón</collectingMunicipality>
, province of
<collectingRegion id="49C5F80A5D4F15212FEED6EAFBB2FD62" box="[977,1068,677,698]" country="Costa Rica" name="Alajuela" pageId="8" pageNumber="19">Alajuela</collectingRegion>
,
<collectingCountry id="F31676785D4F15212808D6EAFEFDFD0D" name="Costa Rica" pageId="8" pageNumber="19">Costa Rica</collectingCountry>
(approx.
<geoCoordinate id="EE35502F5D4F15212DF3D68FFDB9FD0D" box="[460,551,704,725]" degrees="10" direction="north" minutes="21" orientation="latitude" pageId="8" pageNumber="19" precision="925" value="10.35">10°21'N</geoCoordinate>
;
<geoCoordinate id="EE35502F5D4F15212E0CD68FFD0BFD0D" box="[563,661,704,725]" degrees="84" direction="west" minutes="25" orientation="longitude" pageId="8" pageNumber="19" precision="925" value="-84.416664">84°25'W</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection id="C31B65635D4F15212D72D695FDE2FC1D" pageId="8" pageNumber="19" type="biology_ecology">
<paragraph id="8BBE36E85D4F15212D72D695FDE2FC1D" blockId="8.[307,1140,277,1232]" pageId="8" pageNumber="19">
<emphasis id="B975EAFA5D4F15212D72D695FD9EFD37" bold="true" box="[333,512,730,751]" pageId="8" pageNumber="19">Biological Data.</emphasis>
Males and females of
<taxonomicName id="4C014D6B5D4F15212EC8D694FCEAFD28" box="[759,884,731,752]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="lorencita" status="sp. nov.">
<emphasis id="B975EAFA5D4F15212EC8D694FCEAFD28" box="[759,884,731,752]" italics="true" pageId="8" pageNumber="19">P. lorencita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24657815D4F15212F43D694FC29FD28" box="[892,951,731,752]" pageId="8" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
were collected at lights at two sites separated by
<quantity id="4CF99B0D5D4F15212EB3D6BAFD7EFCD2" box="[652,736,757,778]" metricMagnitude="5" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="19" unit="km" value="130.0">130 km</quantity>
, near San Lorencito river, located on Tilaran ridge, between
<quantity id="4CF99B0D5D4F15212E13D75FFD38FCFD" box="[556,678,784,805]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="8" pageNumber="19" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">800900 m</quantity>
of altitude, and Grano de Oro, on the Talamanca ridge, at
<quantity id="4CF99B0D5D4F15212DD8D764FDA3FC98" box="[487,573,811,832]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="8" pageNumber="19" unit="m" value="1120.0">1,120 m</quantity>
of altitude. Phenology: May (3), June (54), July (35). Other species of
<taxonomicName id="4C014D6B5D4F15212DCBD709FDE4FC83" box="[500,634,838,859]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B975EAFA5D4F15212DCBD709FDE4FC83" box="[500,634,838,859]" italics="true" pageId="8" pageNumber="19">Phyllophaga</emphasis>
</taxonomicName>
flying at the same time and places were:
<emphasis id="B975EAFA5D4F15212863D709FEE3FCAD" italics="true" pageId="8" pageNumber="19">P. caraga</emphasis>
Saylor,
<emphasis id="B975EAFA5D4F15212DE5D72FFDF6FCAD" box="[474,616,864,885]" italics="true" pageId="8" pageNumber="19">P. chiriquina</emphasis>
Bates,
<emphasis id="B975EAFA5D4F15212E82D72FFCB1FCAD" box="[701,815,864,885]" italics="true" pageId="8" pageNumber="19">P. densata</emphasis>
Moser,
<emphasis id="B975EAFA5D4F15212FB4D72FFB99FCAD" box="[907,1031,864,885]" italics="true" pageId="8" pageNumber="19">P. gigantea</emphasis>
Bates,
<taxonomicName id="4C014D6B5D4F15212863D72FFE67FC48" authority="Saylor, P." authorityName="Saylor, P." class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="guapiles">
<emphasis id="B975EAFA5D4F15212863D72FFE12FC48" italics="true" pageId="8" pageNumber="19">P. guapiles</emphasis>
Saylor,
<emphasis id="B975EAFA5D4F15212DDDD734FE67FC48" box="[482,505,891,912]" italics="true" pageId="8" pageNumber="19">P.</emphasis>
</taxonomicName>
<emphasis id="B975EAFA5D4F15212E3FD734FDEFFC48" box="[512,625,891,912]" italics="true" pageId="8" pageNumber="19">guapilesea</emphasis>
Saylor,
<emphasis id="B975EAFA5D4F15212EF7D734FCFFFC48" box="[712,865,891,912]" italics="true" pageId="8" pageNumber="19">P. nevermanni</emphasis>
Saylor,
<emphasis id="B975EAFA5D4F15212F87D734FBBBFC48" box="[952,1061,891,912]" italics="true" pageId="8" pageNumber="19">P. orosina</emphasis>
Moser,
<emphasis id="B975EAFA5D4F15212D0CD7D9FE03FC73" box="[307,413,918,939]" italics="true" pageId="8" pageNumber="19">P. prolixa</emphasis>
Bates,
<taxonomicName id="4C014D6B5D4F15212DCFD7D9FC96FC73" authority="Blanchard, P." authorityName="Blanchard, P." box="[496,776,917,939]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="pruinosa">
<emphasis id="B975EAFA5D4F15212DCFD7D9FDF3FC73" box="[496,621,918,939]" italics="true" pageId="8" pageNumber="19">P. pruinosa</emphasis>
Blanchard,
<emphasis id="B975EAFA5D4F15212ECED7D9FC96FC73" box="[753,776,918,939]" italics="true" pageId="8" pageNumber="19">P.</emphasis>
</taxonomicName>
<emphasis id="B975EAFA5D4F15212F2FD7D9FCEAFC73" box="[784,884,918,939]" italics="true" pageId="8" pageNumber="19">setidorsis</emphasis>
Bates, and another two new species under description.
</paragraph>
</subSubSection>
<subSubSection id="C31B65635D4F15212D72D785FD6FFB42" pageId="8" pageNumber="19" type="discussion">
<paragraph id="8BBE36E85D4F15212D72D785FD6FFB42" blockId="8.[307,1140,277,1232]" pageId="8" pageNumber="19">
<emphasis id="B975EAFA5D4F15212D72D785FE27FC07" bold="true" box="[333,441,970,991]" pageId="8" pageNumber="19">Remarks.</emphasis>
Is not possible to include
<taxonomicName id="4C014D6B5D4F15212EDCD784FBEDFC38" box="[739,1139,971,992]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="lorencita" status="sp. nov." subGenus="Phyllophaga">
<emphasis id="B975EAFA5D4F15212EDCD784FCF7FC38" box="[739,873,971,992]" italics="true" pageId="8" pageNumber="19">Phyllophaga</emphasis>
(
<emphasis id="B975EAFA5D4F15212F43D784FB9FFC38" box="[892,1025,971,992]" italics="true" pageId="8" pageNumber="19">Phyllophaga</emphasis>
)
<emphasis id="B975EAFA5D4F1521282AD784FBEDFC38" box="[1045,1139,971,992]" italics="true" pageId="8" pageNumber="19">lorencita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24657815D4F15212D0CD7AAFEF2FC22" box="[307,364,997,1018]" pageId="8" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
in any known species group (
<emphasis id="B975EAFA5D4F15212E92D7A9FD79FC23" box="[685,743,998,1019]" italics="true" pageId="8" pageNumber="19">sensu</emphasis>
<bibRefCitation id="EF904B195D4F15212ED0D7AAFCE8FC22" author="Moron" box="[751,886,997,1018]" firstAuthor="Moron" pageId="8" pageNumber="19" refId="ref11224" refString="Moron, M. A. 1986. El genero Phyllophaga en Mexico. Morfologia, distribucion y sistematica supraespecifica (Insecta: Coleoptera). Publ. 20, Instituto de Ecologia, Mexico, D. F. 341 pp." type="book" year="1986">Morón 1986</bibRefCitation>
). This new species may be related to
<taxonomicName id="4C014D6B5D4F15212DFED04FFC85FBCD" authority="Saylor" authorityName="Saylor" authorityYear="1936" box="[449,795,1024,1045]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="reventazona">
<emphasis id="B975EAFA5D4F15212DFED04FFD51FBCD" box="[449,719,1024,1045]" italics="true" pageId="8" pageNumber="19">Phyllophaga reventazona</emphasis>
Saylor
</taxonomicName>
(Atlantic lowlands,
<collectingCountry id="F31676785D4F15212FC8D04FFBF2FBCD" box="[1015,1132,1024,1045]" name="Costa Rica" pageId="8" pageNumber="19">Costa Rica</collectingCountry>
) by the number and proportions of antenomeres, as well as the shape of the parameres and the dorsal vestiture of aedeagus, but the dense and erect pronotal vestiture, the absence of shallow longitudinal depression on the anterior half of the pronotum, the dark reddish brown body color of
<emphasis id="B975EAFA5D4F15212FF3D024FBEDFB58" box="[972,1139,1131,1152]" italics="true" pageId="8" pageNumber="19">
<taxonomicName id="4C014D6B5D4F15212FF3D024FBF0FB58" authorityName="Saylor" authorityYear="1936" box="[972,1134,1131,1152]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="reventazona">P. reventazona</taxonomicName>
,
</emphasis>
will aid in the separation of both species.
</paragraph>
</subSubSection>
<subSubSection id="C31B65635D4F15212D72D0EFFE1BFB08" pageId="8" pageNumber="19" type="etymology">
<paragraph id="8BBE36E85D4F15212D72D0EFFE1BFB08" blockId="8.[307,1140,277,1232]" pageId="8" pageNumber="19">
<emphasis id="B975EAFA5D4F15212D72D0EFFE52FB6D" bold="true" box="[333,460,1184,1205]" pageId="8" pageNumber="19">Etymology.</emphasis>
Derived from Río San Lorencito, at the eastern Cordillera de Tilarán.
</paragraph>
</subSubSection>
</treatment>
</document>