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<document id="278D474838DC065CDF4CDCE7F6EE1B10" ID-CLB-Dataset="46530" ID-DOI="10.11646/zootaxa.3760.3.4" ID-GBIF-Dataset="d6365622-f1f8-4956-8699-f87ea35d8fe6" ID-ISSN="1175-5326" ID-Zenodo-Dep="228316" ID-ZooBank="1787FEE3-CDF8-4FA5-86E0-39FE5080CDF7" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460378704359" checkinUser="plazi" docAuthor="Jiang, Guo-Chen, Liu, Hong-Chang, Chan, Tin-Yam &amp; Chan, Benny K. K." docDate="2014" docId="03A887A31A59EA7559F28EF1FC47FB13" docLanguage="en" docName="zt03760p382.pdf" docOrigin="Zootaxa 3760 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ocypode cordimanus Latreille 1818" docType="treatment" docVersion="10" lastPageNumber="372" masterDocId="FF91FFDB1A5AEA7659658E66FFA0FFF4" masterDocTitle="First stage zoeal morphology of four ghost crabs Ocypode ceratophthalmus (Pallas, 1772), O. cordimanus Latreille, 1818, O. sinensis Dai, Song &amp; Yang, 1985 and O. stimpsoni Ortmann, 1897 (Crustacea, Decapoda, Ocypodidae)" masterLastPageNumber="382" masterPageNumber="369" pageNumber="372" updateTime="1698296540278" updateUser="ExternalLinkService">
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<mods:title id="365D50D408A2726771813D7929B7215E">First stage zoeal morphology of four ghost crabs Ocypode ceratophthalmus (Pallas, 1772), O. cordimanus Latreille, 1818, O. sinensis Dai, Song &amp; Yang, 1985 and O. stimpsoni Ortmann, 1897 (Crustacea, Decapoda, Ocypodidae)</mods:title>
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<mods:namePart id="8CCDECBB553A2C270B1A3CC4EB292F13">Jiang, Guo-Chen</mods:namePart>
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<mods:namePart id="A472704B4F41DBE786A81DFB2777C51E">Chan, Tin-Yam</mods:namePart>
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<treatment id="03A887A31A59EA7559F28EF1FC47FB13" ID-DOI="http://doi.org/10.5281/zenodo.5680588" ID-GBIF-Taxon="119413727" ID-Zenodo-Dep="5680588" LSID="urn:lsid:plazi:treatment:03A887A31A59EA7559F28EF1FC47FB13" httpUri="http://treatment.plazi.org/id/03A887A31A59EA7559F28EF1FC47FB13" lastPageNumber="372" pageId="3" pageNumber="372">
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<paragraph id="8BBE36B51A59EA7559F28EF1FDF1FF46" blockId="3.[151,593,151,211]" box="[151,593,151,178]" pageId="3" pageNumber="372">
<heading id="D0F681D91A59EA7559F28EF1FDF1FF46" bold="true" box="[151,593,151,178]" fontSize="11" level="1" pageId="3" pageNumber="372" reason="1">
<taxonomicName id="4C014D361A59EA7559F28EF1FDF1FF46" ID-CoL="8N3FC" authority="Latreille, 1818" authorityName="Latreille" authorityYear="1818" box="[151,593,151,178]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="cordimanus">
<emphasis id="B975EAA71A59EA7559F28EF1FDF1FF46" bold="true" box="[151,593,151,178]" pageId="3" pageNumber="372">
<emphasis id="B975EAA71A59EA7559F28EF1FE35FF45" bold="true" box="[151,405,151,177]" italics="true" pageId="3" pageNumber="372">Ocypode cordimanus</emphasis>
Latreille, 1818
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BBE36B51A59EA7559F28EDCFEB2FF27" blockId="3.[151,593,151,211]" box="[151,274,186,211]" pageId="3" pageNumber="372">
(
<figureCitation id="133A2A301A59EA7559FA8EDCFF4EFF27" box="[159,238,186,211]" captionStart="FIGURE 2" captionStartId="4.[151,250,1965,1987]" captionTargetBox="[187,1403,207,1942]" captionTargetId="figure@4.[175,1411,193,1946]" captionTargetPageId="4" captionText="FIGURE 2. Ocypode cordimanus first zoeal stage; A, carapace; B, antennule; C, antenna; D, mandible; E, maxillule; F, maxilla; G, first maxilliped; H, second maxilliped; I, dorsal view of abdomen; J, lateral view of abdomen." httpUri="https://zenodo.org/record/228318/files/figure.png" pageId="3" pageNumber="372">Figs. 2</figureCitation>
,
<figureCitation id="133A2A301A59EA75599F8EDCFEA9FF27" box="[250,265,186,211]" captionStart="FIGURE 6" captionStartId="9.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@9.[193,1394,193,1932]" captionTargetPageId="9" captionText="FIGURE 6. Scanning electron microscope on the first zoeal stage of Ocypode cordimanus; A, lateral view; B, frontal foramen; C, dorsal spine; D, E, ventral carapace; F, dorsal view of telson; G, part of telson showing minute lateral spine; H, part of telson furca." httpUri="https://zenodo.org/record/228322/files/figure.png" pageId="3" pageNumber="372">6</figureCitation>
)
</paragraph>
<paragraph id="8BBE36B51A59EA7559F28F65FF48FE5F" blockId="3.[151,1436,258,1255]" pageId="3" pageNumber="372">
<emphasis id="B975EAA71A59EA7559F28F65FEA5FEEE" box="[151,261,259,282]" italics="true" pageId="3" pageNumber="372">Carapace</emphasis>
(
<figureCitation id="133A2A301A59EA7558728F64FEC7FEEF" box="[279,359,258,283]" captionStart="FIGURE 2" captionStartId="4.[151,250,1965,1987]" captionTargetBox="[187,1403,207,1942]" captionTargetId="figure@4.[175,1411,193,1946]" captionTargetPageId="4" captionText="FIGURE 2. Ocypode cordimanus first zoeal stage; A, carapace; B, antennule; C, antenna; D, mandible; E, maxillule; F, maxilla; G, first maxilliped; H, second maxilliped; I, dorsal view of abdomen; J, lateral view of abdomen." httpUri="https://zenodo.org/record/228318/files/figure.png" pageId="3" pageNumber="372">Figs. 2</figureCitation>
A, 6AE): Pockmarked patterns on surface, with weak rectangular patterns on ventral margin (
<figureCitation id="133A2A301A59EA7559FA8F40FF46FECB" box="[159,230,294,319]" captionStart="FIGURE 6" captionStartId="9.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@9.[193,1394,193,1932]" captionTargetPageId="9" captionText="FIGURE 6. Scanning electron microscope on the first zoeal stage of Ocypode cordimanus; A, lateral view; B, frontal foramen; C, dorsal spine; D, E, ventral carapace; F, dorsal view of telson; G, part of telson showing minute lateral spine; H, part of telson furca." httpUri="https://zenodo.org/record/228322/files/figure.png" pageId="3" pageNumber="372">Fig. 6</figureCitation>
D, E); frontal foramen present (
<figureCitation id="133A2A301A59EA755B058F40FD08FECB" box="[608,680,294,319]" captionStart="FIGURE 6" captionStartId="9.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@9.[193,1394,193,1932]" captionTargetPageId="9" captionText="FIGURE 6. Scanning electron microscope on the first zoeal stage of Ocypode cordimanus; A, lateral view; B, frontal foramen; C, dorsal spine; D, E, ventral carapace; F, dorsal view of telson; G, part of telson showing minute lateral spine; H, part of telson furca." httpUri="https://zenodo.org/record/228322/files/figure.png" pageId="3" pageNumber="372">Fig. 6</figureCitation>
B); dorsal spine with some tubercles (
<figureCitation id="133A2A301A59EA755D0E8F40FB13FECB" box="[1131,1203,294,319]" captionStart="FIGURE 6" captionStartId="9.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@9.[193,1394,193,1932]" captionTargetPageId="9" captionText="FIGURE 6. Scanning electron microscope on the first zoeal stage of Ocypode cordimanus; A, lateral view; B, frontal foramen; C, dorsal spine; D, E, ventral carapace; F, dorsal view of telson; G, part of telson showing minute lateral spine; H, part of telson furca." httpUri="https://zenodo.org/record/228322/files/figure.png" pageId="3" pageNumber="372">Fig. 6</figureCitation>
C), slightly straight, relatively short, approximately same length as rostral spine; rostral spine slightly longer than antennal protopod, without distal spinulation; lateral spines shorter, slightly curved downward; 1 pair of posterodorsal setae; eyes sessile.
</paragraph>
<paragraph id="8BBE36B51A59EA7559A28FD0FE6EFE06" blockId="3.[151,1436,258,1255]" pageId="3" pageNumber="372">
<emphasis id="B975EAA71A59EA7559A28FD0FE99FE3B" box="[199,313,438,463]" italics="true" pageId="3" pageNumber="372">Antennule</emphasis>
(
<figureCitation id="133A2A301A59EA75582F8FD0FE30FE3B" box="[330,400,438,463]" captionStart="FIGURE 2" captionStartId="4.[151,250,1965,1987]" captionTargetBox="[187,1403,207,1942]" captionTargetId="figure@4.[175,1411,193,1946]" captionTargetPageId="4" captionText="FIGURE 2. Ocypode cordimanus first zoeal stage; A, carapace; B, antennule; C, antenna; D, mandible; E, maxillule; F, maxilla; G, first maxilliped; H, second maxilliped; I, dorsal view of abdomen; J, lateral view of abdomen." httpUri="https://zenodo.org/record/228318/files/figure.png" pageId="3" pageNumber="372">Fig. 2</figureCitation>
B): Uniramous, endopod absent; exopod unsegmented with 3 (2 broad, 1 slender) terminal asethetascs, 1 terminal seta.
</paragraph>
<paragraph id="8BBE36B51A59EA7559A28F99FB2FFDCF" blockId="3.[151,1436,258,1255]" pageId="3" pageNumber="372">
<emphasis id="B975EAA71A59EA7559A28F99FE85FDE2" box="[199,293,511,534]" italics="true" pageId="3" pageNumber="372">Antenna</emphasis>
(
<figureCitation id="133A2A301A59EA7558518F98FED9FDE3" box="[308,377,510,535]" captionStart="FIGURE 2" captionStartId="4.[151,250,1965,1987]" captionTargetBox="[187,1403,207,1942]" captionTargetId="figure@4.[175,1411,193,1946]" captionTargetPageId="4" captionText="FIGURE 2. Ocypode cordimanus first zoeal stage; A, carapace; B, antennule; C, antenna; D, mandible; E, maxillule; F, maxilla; G, first maxilliped; H, second maxilliped; I, dorsal view of abdomen; J, lateral view of abdomen." httpUri="https://zenodo.org/record/228318/files/figure.png" pageId="3" pageNumber="372">Fig. 2</figureCitation>
C): Protopod longer than antennule, bearing two rows of small lateral spines; endopod absent; exopod short, with 2 subterminal setae (1 long, almost reaching tip of protopod, 1 shorter).
</paragraph>
<paragraph id="8BBE36B51A59EA7559A28C20FD02FDAB" blockId="3.[151,1436,258,1255]" box="[199,674,582,607]" pageId="3" pageNumber="372">
<emphasis id="B975EAA71A59EA7559A28C20FE93FDAB" box="[199,307,582,607]" italics="true" pageId="3" pageNumber="372">Mandible</emphasis>
(
<figureCitation id="133A2A301A59EA7558248C20FE24FDAB" box="[321,388,582,607]" captionStart="FIGURE 2" captionStartId="4.[151,250,1965,1987]" captionTargetBox="[187,1403,207,1942]" captionTargetId="figure@4.[175,1411,193,1946]" captionTargetPageId="4" captionText="FIGURE 2. Ocypode cordimanus first zoeal stage; A, carapace; B, antennule; C, antenna; D, mandible; E, maxillule; F, maxilla; G, first maxilliped; H, second maxilliped; I, dorsal view of abdomen; J, lateral view of abdomen." httpUri="https://zenodo.org/record/228318/files/figure.png" pageId="3" pageNumber="372">Fig. 2</figureCitation>
D): Endopod palp absent.
</paragraph>
<paragraph id="8BBE36B51A59EA7559A28C0CFA2AFD53" blockId="3.[151,1436,258,1255]" pageId="3" pageNumber="372">
<emphasis id="B975EAA71A59EA7559A28C0CFE93FD77" box="[199,307,618,643]" italics="true" pageId="3" pageNumber="372">Maxillule</emphasis>
(
<figureCitation id="133A2A301A59EA7558248C0CFE23FD77" box="[321,387,618,643]" captionStart="FIGURE 2" captionStartId="4.[151,250,1965,1987]" captionTargetBox="[187,1403,207,1942]" captionTargetId="figure@4.[175,1411,193,1946]" captionTargetPageId="4" captionText="FIGURE 2. Ocypode cordimanus first zoeal stage; A, carapace; B, antennule; C, antenna; D, mandible; E, maxillule; F, maxilla; G, first maxilliped; H, second maxilliped; I, dorsal view of abdomen; J, lateral view of abdomen." httpUri="https://zenodo.org/record/228318/files/figure.png" pageId="3" pageNumber="372">Fig. 2</figureCitation>
E): Epipod seta absent; coxal endite with 5 plumose setae; basial endite with 5 setal processes; endopod 2-segmented, proximal segment without setae, distal segment with 4 terminal setae; exopod seta absent.
</paragraph>
<paragraph id="8BBE36B51A59EA7559A28CD4FDB2FCE6" blockId="3.[151,1436,258,1255]" pageId="3" pageNumber="372">
<emphasis id="B975EAA71A59EA7559A28CD4FEBEFD3F" box="[199,286,690,715]" italics="true" pageId="3" pageNumber="372">Maxilla</emphasis>
(
<figureCitation id="133A2A301A59EA75584A8CD4FED3FD3F" box="[303,371,690,715]" captionStart="FIGURE 2" captionStartId="4.[151,250,1965,1987]" captionTargetBox="[187,1403,207,1942]" captionTargetId="figure@4.[175,1411,193,1946]" captionTargetPageId="4" captionText="FIGURE 2. Ocypode cordimanus first zoeal stage; A, carapace; B, antennule; C, antenna; D, mandible; E, maxillule; F, maxilla; G, first maxilliped; H, second maxilliped; I, dorsal view of abdomen; J, lateral view of abdomen." httpUri="https://zenodo.org/record/228318/files/figure.png" pageId="3" pageNumber="372">Fig. 2</figureCitation>
F): Coxal endite bilobed, proximal lobe with 4 setae, distal lobe with 2 setae + 1 rudimentary seta; basial endite bilobed with 5+4 setae; endopod with 1+2 terminal setae; scaphognathite margin with 4 setae, 1 long stout plumose distal process.
</paragraph>
<paragraph id="8BBE36B51A59EA7559A28D79FE3FFC8A" blockId="3.[151,1436,258,1255]" pageId="3" pageNumber="372">
<emphasis id="B975EAA71A59EA7559A28D79FEDCFCC3" box="[199,380,798,823]" italics="true" pageId="3" pageNumber="372">First maxilliped</emphasis>
(
<figureCitation id="133A2A301A59EA7558E88D78FE72FCC3" box="[397,466,798,823]" captionStart="FIGURE 2" captionStartId="4.[151,250,1965,1987]" captionTargetBox="[187,1403,207,1942]" captionTargetId="figure@4.[175,1411,193,1946]" captionTargetPageId="4" captionText="FIGURE 2. Ocypode cordimanus first zoeal stage; A, carapace; B, antennule; C, antenna; D, mandible; E, maxillule; F, maxilla; G, first maxilliped; H, second maxilliped; I, dorsal view of abdomen; J, lateral view of abdomen." httpUri="https://zenodo.org/record/228318/files/figure.png" pageId="3" pageNumber="372">Fig. 2</figureCitation>
G): Coxa with 1 seta; basis with 9 setae arranged as 2, 2, 3, 2; endopod 5-segmented with 2, 2, 1, 2, 5 (1 subterminal + 4 terminal) setae; exopod 2-segmented, distal segment with 4 long terminal plumose natatory setae.
</paragraph>
<paragraph id="8BBE36B51A59EA7559A28DECFDA3FC1E" blockId="3.[151,1436,258,1255]" pageId="3" pageNumber="372">
<emphasis id="B975EAA71A59EA7559A28DECFE3AFC57" box="[199,410,906,931]" italics="true" pageId="3" pageNumber="372">Second maxilliped</emphasis>
(
<figureCitation id="133A2A301A59EA7558CA8DECFE58FC57" box="[431,504,906,931]" captionStart="FIGURE 2" captionStartId="4.[151,250,1965,1987]" captionTargetBox="[187,1403,207,1942]" captionTargetId="figure@4.[175,1411,193,1946]" captionTargetPageId="4" captionText="FIGURE 2. Ocypode cordimanus first zoeal stage; A, carapace; B, antennule; C, antenna; D, mandible; E, maxillule; F, maxilla; G, first maxilliped; H, second maxilliped; I, dorsal view of abdomen; J, lateral view of abdomen." httpUri="https://zenodo.org/record/228318/files/figure.png" pageId="3" pageNumber="372">Fig. 2</figureCitation>
H): Coxa without setae; basis with 4 setae arranged as 1, 1, 1, 1; endopod 3- segmented with 0, 0, 5 (3 subterminal + 2 terminal) setae; exopod unsegmented, distal segment with 4 long terminal plumose natatory setae.
</paragraph>
<paragraph id="8BBE36B51A59EA7559A28D90FE47FBFB" blockId="3.[151,1436,258,1255]" box="[199,487,1014,1039]" pageId="3" pageNumber="372">
<emphasis id="B975EAA71A59EA7559A28D90FE21FBFB" box="[199,385,1014,1039]" italics="true" pageId="3" pageNumber="372">Third maxilliped</emphasis>
: Absent.
</paragraph>
<paragraph id="8BBE36B51A59EA7559A28A7CFE0BFBC7" blockId="3.[151,1436,258,1255]" box="[199,427,1050,1075]" pageId="3" pageNumber="372">
<emphasis id="B975EAA71A59EA7559A28A7CFEE5FBC7" box="[199,325,1050,1075]" italics="true" pageId="3" pageNumber="372">Pereiopods</emphasis>
: Absent.
</paragraph>
<paragraph id="8BBE36B51A59EA7559A28A58FD28FB6B" blockId="3.[151,1436,258,1255]" pageId="3" pageNumber="372">
<emphasis id="B975EAA71A59EA7559A28A58FE91FBA3" box="[199,305,1086,1111]" italics="true" pageId="3" pageNumber="372">Abdomen</emphasis>
(
<figureCitation id="133A2A301A59EA7558248A58FE25FBA3" box="[321,389,1086,1111]" captionStart="FIGURE 2" captionStartId="4.[151,250,1965,1987]" captionTargetBox="[187,1403,207,1942]" captionTargetId="figure@4.[175,1411,193,1946]" captionTargetPageId="4" captionText="FIGURE 2. Ocypode cordimanus first zoeal stage; A, carapace; B, antennule; C, antenna; D, mandible; E, maxillule; F, maxilla; G, first maxilliped; H, second maxilliped; I, dorsal view of abdomen; J, lateral view of abdomen." httpUri="https://zenodo.org/record/228318/files/figure.png" pageId="3" pageNumber="372">
Fig.
<date id="FFBF10751A59EA7558138A59FE25FBA3" box="[374,389,1087,1111]" pageId="3" pageNumber="372">2</date>
</figureCitation>
I, J): With 5 somites; somites 2, 3 with pair of dorsolateral processes; somites 23 with short posterolateral spinous processes, somites 45 with expanded posterolateral spinous processes; somites 25 with 1 pair of posterodorsal setae; pleopods absent.
</paragraph>
<paragraph id="8BBE36B51A59EA7559A28ACCFC47FB13" blockId="3.[151,1436,258,1255]" pageId="3" pageNumber="372">
<emphasis id="B975EAA71A59EA7559A28ACCFEAFFB37" box="[199,271,1194,1219]" italics="true" pageId="3" pageNumber="372">Telson</emphasis>
(
<figureCitation id="133A2A301A59EA75587A8ACCFECDFB37" box="[287,365,1194,1219]" captionStart="FIGURE 2" captionStartId="4.[151,250,1965,1987]" captionTargetBox="[187,1403,207,1942]" captionTargetId="figure@4.[175,1411,193,1946]" captionTargetPageId="4" captionText="FIGURE 2. Ocypode cordimanus first zoeal stage; A, carapace; B, antennule; C, antenna; D, mandible; E, maxillule; F, maxilla; G, first maxilliped; H, second maxilliped; I, dorsal view of abdomen; J, lateral view of abdomen." httpUri="https://zenodo.org/record/228318/files/figure.png" pageId="3" pageNumber="372">
Figs.
<date id="FFBF10751A59EA75583B8ACDFECDFB37" box="[350,365,1195,1219]" pageId="3" pageNumber="372">2</date>
</figureCitation>
I, J, 6FH): Telson bifid, curved upward distally; with 1 pair of minute lateral spines; generally pointed on each furca; posterior margin with 3 pairs of stout pointed spines.
</paragraph>
</subSubSection>
</treatment>
</document>