First stage zoeal morphology of four ghost crabs Ocypode ceratophthalmus (Pallas, 1772), O. cordimanus Latreille, 1818, O. sinensis Dai, Song & Yang, 1985 and O. stimpsoni Ortmann, 1897 (Crustacea, Decapoda, Ocypodidae) Author Jiang, Guo-Chen Author Liu, Hong-Chang Author Chan, Tin-Yam Author Chan, Benny K. K. text Zootaxa 2014 3760 3 369 382 journal article 46530 10.11646/zootaxa.3760.3.4 d6365622-f1f8-4956-8699-f87ea35d8fe6 1175-5326 228316 1787FEE3-CDF8-4FA5-86E0-39FE5080CDF7 Ocypode cordimanus Latreille, 1818 ( Figs. 2 , 6 ) Carapace ( Figs. 2 A, 6A–E): Pockmarked patterns on surface, with weak rectangular patterns on ventral margin ( Fig. 6 D, E); frontal foramen present ( Fig. 6 B); dorsal spine with some tubercles ( Fig. 6 C), slightly straight, relatively short, approximately same length as rostral spine; rostral spine slightly longer than antennal protopod, without distal spinulation; lateral spines shorter, slightly curved downward; 1 pair of posterodorsal setae; eyes sessile. Antennule ( Fig. 2 B): Uniramous, endopod absent; exopod unsegmented with 3 (2 broad, 1 slender) terminal asethetascs, 1 terminal seta. Antenna ( Fig. 2 C): Protopod longer than antennule, bearing two rows of small lateral spines; endopod absent; exopod short, with 2 subterminal setae (1 long, almost reaching tip of protopod, 1 shorter). Mandible ( Fig. 2 D): Endopod palp absent. Maxillule ( Fig. 2 E): Epipod seta absent; coxal endite with 5 plumose setae; basial endite with 5 setal processes; endopod 2-segmented, proximal segment without setae, distal segment with 4 terminal setae; exopod seta absent. Maxilla ( Fig. 2 F): Coxal endite bilobed, proximal lobe with 4 setae, distal lobe with 2 setae + 1 rudimentary seta; basial endite bilobed with 5+4 setae; endopod with 1+2 terminal setae; scaphognathite margin with 4 setae, 1 long stout plumose distal process. First maxilliped ( Fig. 2 G): Coxa with 1 seta; basis with 9 setae arranged as 2, 2, 3, 2; endopod 5-segmented with 2, 2, 1, 2, 5 (1 subterminal + 4 terminal) setae; exopod 2-segmented, distal segment with 4 long terminal plumose natatory setae. Second maxilliped ( Fig. 2 H): Coxa without setae; basis with 4 setae arranged as 1, 1, 1, 1; endopod 3- segmented with 0, 0, 5 (3 subterminal + 2 terminal) setae; exopod unsegmented, distal segment with 4 long terminal plumose natatory setae. Third maxilliped : Absent. Pereiopods : Absent. Abdomen ( Fig. 2 I, J): With 5 somites; somites 2, 3 with pair of dorsolateral processes; somites 2–3 with short posterolateral spinous processes, somites 4–5 with expanded posterolateral spinous processes; somites 2–5 with 1 pair of posterodorsal setae; pleopods absent. Telson ( Figs. 2 I, J, 6F–H): Telson bifid, curved upward distally; with 1 pair of minute lateral spines; generally pointed on each furca; posterior margin with 3 pairs of stout pointed spines.