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<mods:title id="C57849FF144DBFEF9748AB6A4E17C8D4">A new species of Alycaeidae, Pincerna yanseni n. sp. from Sumatra, with the resurrection of the genus Pincerna Preston, 1907 (Gastropoda: Cyclophoroidea)</mods:title>
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<treatment id="03FD87E8FFDDA07F347CF985FAECFB1D" ID-DOI="http://doi.org/10.5281/zenodo.4776518" ID-GBIF-Taxon="182248001" ID-Zenodo-Dep="4776518" LSID="urn:lsid:plazi:treatment:03FD87E8FFDDA07F347CF985FAECFB1D" httpUri="http://treatment.plazi.org/id/03FD87E8FFDDA07F347CF985FAECFB1D" lastPageNumber="214" pageId="1" pageNumber="214">
<subSubSection id="C34E6575FFDDA07F347CF985FDF5F9F9" box="[259,605,1562,1586]" pageId="1" pageNumber="214" type="nomenclature">
<paragraph id="8BEB36FEFFDDA07F347CF985FDF5F9F9" blockId="1.[259,605,1562,1586]" box="[259,605,1562,1586]" pageId="1" pageNumber="214">
<heading id="D0A38192FFDDA07F347CF985FDF5F9F9" bold="true" box="[259,605,1562,1586]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="214" reason="2">
<emphasis id="B920EAECFFDDA07F347CF985FDF5F9F9" bold="true" box="[259,605,1562,1586]" pageId="1" pageNumber="214">
Genus
<taxonomicName id="4C544D7DFFDDA07F342BF985FDF5F9F9" authority="Preston, 1907" authorityName="Preston" authorityYear="1907" box="[340,605,1562,1586]" class="Gastropoda" family="Cyclophoridae" genus="Pincerna" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="genus">
<emphasis id="B920EAECFFDDA07F342BF985FE11F9F9" bold="true" box="[340,441,1562,1586]" italics="true" pageId="1" pageNumber="214">Pincerna</emphasis>
<bibRefCitation id="EFC54B0FFFDDA07F34BDF985FDF5F9F9" author="Preston HB" box="[450,605,1562,1586]" pageId="1" pageNumber="214" pagination="206" refId="ref4023" refString="Preston HB (1907) Description of a new subgenus and species of Alycaeus from Ke-Lan-Tan. Proceedings of the Malacological Society of London, 7: 206." type="journal article" year="1907">Preston, 1907</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E6575FFDDA07F350EF9C6FD40F96C" pageId="1" pageNumber="214" type="reference_group">
<paragraph id="8BEB36FEFFDDA07F350EF9C6FE53F947" blockId="1.[113,752,1625,1704]" pageId="1" pageNumber="214">
<treatmentCitationGroup id="AB4411D0FFDDA07F350EF9C6FE53F947" pageId="1" pageNumber="214">
<treatmentCitation id="0AF510EFFFDDA07F350EF9C6FED7F9A4" author="Preston HB" box="[113,383,1625,1647]" page="206" pageId="1" pageNumber="214" year="1907">
<taxonomicName id="4C544D7DFFDDA07F350EF9C6FED7F9A4" authority="Preston, 1907: 206" authorityName="Preston" authorityPageNumber="206" authorityYear="1907" box="[113,383,1625,1647]" class="Gastropoda" family="Cyclophoridae" genus="Pincerna" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="genus">
<emphasis id="B920EAECFFDDA07F350EF9C6FF60F9A5" box="[113,200,1625,1646]" italics="true" pageId="1" pageNumber="214">Pincerna</emphasis>
<bibRefCitation id="EFC54B0FFFDDA07F35B3F9C6FED7F9A4" author="Preston HB" box="[204,383,1625,1647]" pageId="1" pageNumber="214" pagination="206" refId="ref4023" refString="Preston HB (1907) Description of a new subgenus and species of Alycaeus from Ke-Lan-Tan. Proceedings of the Malacological Society of London, 7: 206." type="journal article" year="1907">Preston, 1907: 206</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. (introduced as a subgenus of
<taxonomicName id="4C544D7DFFDDA07F37ECF9C6FD41F9A5" authorityName="Baird" authorityYear="1850" box="[659,745,1625,1646]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="genus">
<emphasis id="B920EAECFFDDA07F37ECF9C6FD41F9A5" box="[659,745,1625,1646]" italics="true" pageId="1" pageNumber="214">Alycaeus</emphasis>
</taxonomicName>
, but apparently used on genus level).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEB36FEFFDDA07F350EF90DFD40F96C" blockId="1.[113,752,1625,1704]" box="[113,744,1682,1704]" pageId="1" pageNumber="214">
<taxonomicName id="4C544D7DFFDDA07F350EF90DFDE2F96C" authority="Godwin-Austen, 1914: 334" authorityName="Godwin-Austen" authorityPageNumber="334" authorityYear="1914" box="[113,586,1682,1704]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="subGenus" subGenus="Cycloryx">
<emphasis id="B920EAECFFDDA07F350EF90DFF60F96C" box="[113,200,1682,1703]" italics="true" pageId="1" pageNumber="214">Alycaeus</emphasis>
(
<emphasis id="B920EAECFFDDA07F35A8F90DFE86F96C" box="[215,302,1682,1703]" italics="true" pageId="1" pageNumber="214">Cycloryx</emphasis>
) Godwin-Austen, 1914: 334
</taxonomicName>
.
<emphasis id="B920EAECFFDDA07F3728F90DFD40F96C" bold="true" box="[599,744,1682,1703]" pageId="1" pageNumber="214">New synonym</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C34E6575FFDDA07F350EF94EFEDCF8A3" pageId="1" pageNumber="214" type="type_taxon">
<paragraph id="8BEB36FEFFDDA07F350EF94EFEDCF8A3" blockId="1.[112,753,1744,1897]" pageId="1" pageNumber="214">
<emphasis id="B920EAECFFDDA07F350EF94EFEACF923" bold="true" box="[113,260,1744,1769]" pageId="1" pageNumber="214">
<typeStatus id="54EF885CFFDDA07F350EF94EFF02F922" box="[113,170,1745,1769]" pageId="1" pageNumber="214">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C544D7DFFDDA07F3476F94EFDFDF922" authority="Preston, 1907" authorityName="Preston" authorityYear="1907" box="[265,597,1744,1769]" class="Gastropoda" family="Cyclophoridae" genus="Pincerna" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="species" species="liratula">
<emphasis id="B920EAECFFDDA07F3476F94EFE16F923" box="[265,446,1744,1768]" italics="true" pageId="1" pageNumber="214">Pincerna liratula</emphasis>
<bibRefCitation id="EFC54B0FFFDDA07F34BCF94EFDFDF922" author="Preston HB" box="[451,597,1745,1769]" pageId="1" pageNumber="214" pagination="206" refId="ref4023" refString="Preston HB (1907) Description of a new subgenus and species of Alycaeus from Ke-Lan-Tan. Proceedings of the Malacological Society of London, 7: 206." type="journal article" year="1907">Preston, 1907</bibRefCitation>
</taxonomicName>
, by monotypy.
<taxonomicName id="4C544D7DFFDDA07F350FF96FFE63F8C3" authority="Morgan, 1885" authorityName="Morgan" authorityYear="1885" box="[112,459,1776,1801]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="species" species="thieroti">
<emphasis id="B920EAECFFDDA07F350FF96FFE81F8C3" box="[112,297,1776,1800]" italics="true" pageId="1" pageNumber="214">Alycaeus thieroti</emphasis>
<bibRefCitation id="EFC54B0FFFDDA07F344DF96EFE63F8C3" author="Morgan J de" box="[306,459,1776,1801]" pageId="1" pageNumber="214" pagination="353 - 429" refId="ref3907" refString="Morgan J de (1885) Mollusques terrestres et fluviatiles du Royaume de Perak et des pais voisins (presque'Ile Malaise). Bulletin de la Societe Zoologique de France, 10: 353 - 429." type="journal article" year="1885">Morgan, 1885</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54EF885CFFDDA07F34A3F96EFDBCF8C2" box="[476,532,1777,1801]" pageId="1" pageNumber="214">Type</typeStatus>
locality: “G. Lano” near
<collectingRegion id="4990F81CFFDDA07F35D6F88FFF40F8E3" box="[169,232,1808,1832]" country="Malaysia" name="Perak" pageId="1" pageNumber="214">Perak</collectingRegion>
) is very similar to
<taxonomicName id="4C544D7DFFDDA07F34BDF88EFDD7F8E3" authorityName="Preston" authorityYear="1907" box="[450,639,1808,1832]" class="Gastropoda" family="Cyclophoridae" genus="Pincerna" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="species" species="liratula">
<emphasis id="B920EAECFFDDA07F34BDF88EFDD7F8E3" box="[450,639,1808,1832]" italics="true" pageId="1" pageNumber="214">Pincerna liratula</emphasis>
</taxonomicName>
(“Ke-lantan” in the Malay Peninsula); the latter might be a junior synonym of the former.
</paragraph>
</subSubSection>
<subSubSection id="C34E6575FFDDA07F350EF80EFAECFB1D" pageId="1" pageNumber="214" type="discussion">
<paragraph id="8BEB36FEFFDDA07F350EF80EFF72F783" blockId="1.[112,752,1936,2120]" pageId="1" pageNumber="214">
<emphasis id="B920EAECFFDDA07F350EF80EFF77F862" bold="true" box="[113,223,1937,1961]" pageId="1" pageNumber="214">Remarks.</emphasis>
The genus-level revision of the family
<taxonomicName id="4C544D7DFFDDA07F3706F80FFD58F863" authorityName="W. T. Blanford" authorityYear="1864" box="[633,752,1936,1960]" class="Gastropoda" family="Alycaeidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="family">Alycaeidae</taxonomicName>
is in progress and beyond the scope of the present paper. However, to place the new species in a genus correctly, some information on the relationship of the genus
<taxonomicName id="4C544D7DFFDDA07F37EFF86FFECEF7E2" authority="Godwin-Austen, 1914" authorityName="Godwin-Austen" authorityYear="1914" class="Gastropoda" family="Cyclophoridae" genus="Cycloryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="genus">
<emphasis id="B920EAECFFDDA07F37EFF86FFD58F7C3" box="[656,752,2032,2056]" italics="true" pageId="1" pageNumber="214">Cycloryx</emphasis>
Godwin-Austen, 1914
</taxonomicName>
, and
<taxonomicName id="4C544D7DFFDDA07F34D9F78EFD05F7E2" authority="Preston, 1907" authorityName="Preston" authorityYear="1907" box="[422,685,2065,2089]" class="Gastropoda" family="Cyclophoridae" genus="Pincerna" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="genus">
<emphasis id="B920EAECFFDDA07F34D9F78EFDA1F7E3" box="[422,521,2065,2088]" italics="true" pageId="1" pageNumber="214">Pincerna</emphasis>
<bibRefCitation id="EFC54B0FFFDDA07F376CF78EFD05F7E2" author="Preston HB" box="[531,685,2065,2089]" pageId="1" pageNumber="214" pagination="206" refId="ref4023" refString="Preston HB (1907) Description of a new subgenus and species of Alycaeus from Ke-Lan-Tan. Proceedings of the Malacological Society of London, 7: 206." type="journal article" year="1907">Preston, 1907</bibRefCitation>
</taxonomicName>
, must be added.
</paragraph>
<paragraph id="8BEB36FEFFDDA07F3653FF22FB8EFC5E" blockId="1.[811,1452,189,918]" pageId="1" pageNumber="214">
The discovery of
<taxonomicName id="4C544D7DFFDDA07F36A1FF21FB3BFF1E" authority="Páll-Gergely, 2017" authorityName="Páll-Gergely" authorityYear="2017" box="[990,1171,190,213]" class="Gastropoda" family="Cyclophoridae" genus="Pincerna" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="species" species="yanseni" status="sp. nov.">
<emphasis id="B920EAECFFDDA07F36A1FF21FB3BFF1E" box="[990,1171,190,213]" italics="true" pageId="1" pageNumber="214">Pincerna yanseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2135797FFDDA07F31E6FF21FB78FF1E" box="[1177,1232,190,213]" pageId="1" pageNumber="214" rank="species">n. sp.</taxonomicNameLabel>
is surprising because it shows only superficial similarity with most other
<taxonomicName id="4C544D7DFFDDA07F304AFF42FA02FF3E" authorityName="W. T. Blanford" authorityYear="1864" box="[1333,1450,221,245]" class="Gastropoda" family="Alycaeidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="family">Alycaeidae</taxonomicName>
of the Malay Archipelago, but it is similar to many species from northeastern
<collectingCountry id="F343766EFFDDA07F317AFE82FB96FEFE" box="[1029,1086,285,309]" name="India" pageId="1" pageNumber="214">India</collectingCountry>
and
<collectingCountry id="F343766EFFDDA07F3102FE81FB41FEFD" box="[1149,1257,286,310]" name="Myanmar" pageId="1" pageNumber="214">Myanmar</collectingCountry>
classified in the genus
<taxonomicName id="4C544D7DFFDDA07F360FFEA2FC78FE9E" authorityName="Godwin-Austen" authorityYear="1914" box="[880,976,317,341]" class="Gastropoda" family="Cyclophoridae" genus="Cycloryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="genus">
<emphasis id="B920EAECFFDDA07F360FFEA2FC78FE9E" box="[880,976,317,341]" italics="true" pageId="1" pageNumber="214">Cycloryx</emphasis>
</taxonomicName>
. The genus
<taxonomicName id="4C544D7DFFDDA07F3131FEA2FB06FE9E" authorityName="Godwin-Austen" authorityYear="1914" box="[1102,1198,317,341]" class="Gastropoda" family="Cyclophoridae" genus="Cycloryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="genus">
<emphasis id="B920EAECFFDDA07F3131FEA2FB06FE9E" box="[1102,1198,317,341]" italics="true" pageId="1" pageNumber="214">Cycloryx</emphasis>
</taxonomicName>
(
<typeStatus id="54EF885CFFDDA07F31C3FEA2FB43FE9E" box="[1212,1259,317,341]" pageId="1" pageNumber="214">type</typeStatus>
species:
<taxonomicName id="4C544D7DFFDDA07F3035FEA2FB96FEBE" authority="Benson, 1851" authorityName="Benson" authorityYear="1851" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="species" species="constrictus">
<emphasis id="B920EAECFFDDA07F3035FEA2FC09FEBE" italics="true" pageId="1" pageNumber="214">Alycaeus constrictus</emphasis>
<bibRefCitation id="EFC54B0FFFDDA07F36D5FEC1FB96FEBE" author="Benson WH" box="[938,1086,349,373]" pageId="1" pageNumber="214" pagination="184 - 195" refId="ref3386" refString="Benson WH (1851) Geographical notices, and characters of fourteen new species of Cyclostoma, from the East Indies. The Annals and Magazine of Natural History, Ser. 2 (8): 184 - 195." type="journal article" year="1851">Benson, 1851</bibRefCitation>
</taxonomicName>
by original designation,
<figureCitation id="136F2A7BFFDDA07F3030FEC2FA0AFEBE" box="[1359,1442,349,373]" captionStart="Fig" captionStartId="2.[136,171,1887,1909]" captionTargetBox="[136,1474,337,1871]" captionTargetId="figure-7@2.[136,1474,337,1871]" captionTargetPageId="2" captionText="Fig. 1. Shells of Pincerna species. A, Pincerna constricta (Benson 1851) (type species of Cycloryx), NHMUK 1906.4.4.41 (syntype); B, Pincernayanseni n. sp. MZB.Gst. 18.970 (holotype); C,Pincerna liratula Preston, 1907 (type species ofPincerna), NHMUK 1907.5.20.191, (syntype). Scale = 1 mm, upper scale refers to AB. Photos: H. Taylor (A, C) and B. Páll-Gergely (B)." figureDoi="http://doi.org/10.5281/zenodo.4502851" httpUri="https://zenodo.org/record/4502851/files/figure.png" pageId="1" pageNumber="214">Fig. 1A</figureCitation>
) was erected as a subgenus of
<taxonomicName id="4C544D7DFFDDA07F311CFEE2FAE3FE5E" authority="Baird, 1850" authorityName="Baird" authorityYear="1850" box="[1123,1355,381,405]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="genus">
<emphasis id="B920EAECFFDDA07F311CFEE2FB6BFE5E" box="[1123,1219,381,405]" italics="true" pageId="1" pageNumber="214">Alycaeus</emphasis>
<bibRefCitation id="EFC54B0FFFDDA07F31B5FEE2FAE3FE5E" author="Baird W" box="[1226,1355,381,405]" pageId="1" pageNumber="214" refId="ref3270" refString="Baird W (1850) Nomenclature of Molluscous Animals and Shells in the Collection of the British Museum. Part I. Cyclophoridae. Printed by order of the trustees [by Spottiswoodes and Shaw], London, 69 pp." type="book" year="1850">Baird, 1850</bibRefCitation>
</taxonomicName>
, and was diagnosed on the basis of the ovately conoid shell shape, the regular ribbing on the upper whorls, and the extremely short, often clubbed or pear-shaped sutural tube (Godwin-Austen, 1914). Godwin-Austen (1914) only included species from northeastern
<collectingCountry id="F343766EFFDDA07F36CDFD82FC40FDFE" box="[946,1000,541,565]" name="India" pageId="1" pageNumber="214">India</collectingCountry>
and
<collectingCountry id="F343766EFFDDA07F3168FD81FBF6FDFE" box="[1047,1118,542,565]" name="Myanmar" pageId="1" pageNumber="214">Burma</collectingCountry>
(Rakhin =
<collectingRegion id="4990F81CFFDDA07F31AFFD82FA88FDFE" box="[1232,1312,541,565]" country="Myanmar" name="Rakhine" pageId="1" pageNumber="214">Arakan</collectingRegion>
, and the
<collectingRegion id="4990F81CFFDDA07F3008FD82FA04FDFE" box="[1399,1452,541,565]" country="Myanmar" name="Shan" pageId="1" pageNumber="214">Shan</collectingRegion>
States). However, the diagnosis of
<taxonomicName id="4C544D7DFFDDA07F31E3FDA2FB54FD9E" authorityName="Godwin-Austen" authorityYear="1914" box="[1180,1276,573,597]" class="Gastropoda" family="Cyclophoridae" genus="Cycloryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="genus">
<emphasis id="B920EAECFFDDA07F31E3FDA2FB54FD9E" box="[1180,1276,573,597]" italics="true" pageId="1" pageNumber="214">Cycloryx</emphasis>
</taxonomicName>
matches several species extralimital to the distributional range as defined by Godwin-Austen:
<taxonomicName id="4C544D7DFFDDA07F369DFDE2FCC8FD7E" authority="Bavay &amp; Dautzenberg, 1912" authorityName="Bavay &amp; Dautzenberg" authorityYear="1912" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="species" species="costulosus">
<emphasis id="B920EAECFFDDA07F369DFDE2FB1DFD5E" box="[994,1205,637,661]" italics="true" pageId="1" pageNumber="214">Alycaeus costulosus</emphasis>
<bibRefCitation id="EFC54B0FFFDDA07F31C5FDE1FCC8FD7E" author="Bavay A &amp; Dautzenberg P" pageId="1" pageNumber="214" pagination="1 - 54" refId="ref3361" refString="Bavay A &amp; Dautzenberg P (1912) Description de Coquilles nouvelles de I'Indo-Chine. Journal de Conchyliologie, 60: 1 - 54." type="journal article" year="1912">Bavay &amp; Dautzenberg, 1912</bibRefCitation>
</taxonomicName>
(northern
<collectingCountry id="F343766EFFDDA07F36B4FD02FB82FD7E" box="[971,1066,669,693]" name="Vietnam" pageId="1" pageNumber="214">Vietnam</collectingCountry>
),
<taxonomicName id="4C544D7DFFDDA07F3145FD02FA02FD7E" authority="H. Adams, 1870" authorityName="H. Adams" authorityYear="1870" box="[1082,1450,669,693]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="species" species="globosus">
<emphasis id="B920EAECFFDDA07F3145FD02FB54FD7E" box="[1082,1276,669,693]" italics="true" pageId="1" pageNumber="214">Alycaeus globosus</emphasis>
H.
<bibRefCitation id="EFC54B0FFFDDA07F3060FD02FA02FD7E" author="Adams H" box="[1311,1450,669,693]" pageId="1" pageNumber="214" pagination="793 - 795" refId="ref3239" refString="Adams H (1870) Descriptions of two new genera and five new species of shells. Proceedings of the Zoological Society of London, 1870: 793 - 795." type="journal article" year="1870">Adams, 1870</bibRefCitation>
</taxonomicName>
and its subspecies (Borneo),
<taxonomicName id="4C544D7DFFDDA07F311EFD21FCCBFD3E" authority="Luo et al., 2009" authorityName="Luo" authorityYear="2009" class="Gastropoda" family="Cyclophoridae" genus="Dioryx" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="species" species="maolanensis">
<emphasis id="B920EAECFFDDA07F311EFD21FA9FFD1E" box="[1121,1335,701,725]" italics="true" pageId="1" pageNumber="214">Dioryx maolanensis</emphasis>
<bibRefCitation id="EFC54B0FFFDDA07F3040FD21FCCBFD3E" author="Luo T-C &amp; Zhang W-H &amp; Zhou W-C" pageId="1" pageNumber="214" pagination="862 - 864" refId="ref3735" refString="Luo T-C, Zhang W-H &amp; Zhou W-C (2009) A new species of the genus Dioryx Benson from China (Prosobranchia, Mesogastropoda, Cyclophoridae). Acta Zootaxonomica Sinica, 34 (4): 862 - 864. [In Mandarin Chinese]" type="journal article" year="2009">Luo et al., 2009</bibRefCitation>
</taxonomicName>
(
<collectingRegion id="4990F81CFFDDA07F360AFD42FC70FD3E" box="[885,984,733,757]" country="China" name="Guizhou" pageId="1" pageNumber="214">Guizhou</collectingRegion>
,
<collectingCountry id="F343766EFFDDA07F3698FD42FB85FD3E" box="[999,1069,733,757]" name="China" pageId="1" pageNumber="214">China</collectingCountry>
),
<taxonomicName id="4C544D7DFFDDA07F313CFD42FA03FD3E" authority="Morgan, 1885" authorityName="Morgan" authorityYear="1885" box="[1091,1451,733,758]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="species" species="thieroti">
<emphasis id="B920EAECFFDDA07F313CFD42FAAAFD3E" box="[1091,1282,733,757]" italics="true" pageId="1" pageNumber="214">Alycaeus thieroti</emphasis>
<bibRefCitation id="EFC54B0FFFDDA07F3073FD41FA03FD3E" author="Morgan J de" box="[1292,1451,733,758]" pageId="1" pageNumber="214" pagination="353 - 429" refId="ref3907" refString="Morgan J de (1885) Mollusques terrestres et fluviatiles du Royaume de Perak et des pais voisins (presque'Ile Malaise). Bulletin de la Societe Zoologique de France, 10: 353 - 429." type="journal article" year="1885">Morgan, 1885</bibRefCitation>
</taxonomicName>
(Perak, Malay Peninsula), and
<taxonomicName id="4C544D7DFFDDA07F31F6FD61FCC9FCFE" authority="Preston, 1907" authorityName="Preston" authorityYear="1907" class="Gastropoda" family="Cyclophoridae" genus="Pincerna" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="species" species="liratula">
<emphasis id="B920EAECFFDDA07F31F6FD61FAE1FCDE" box="[1161,1353,765,789]" italics="true" pageId="1" pageNumber="214">Pincerna liratula</emphasis>
<bibRefCitation id="EFC54B0FFFDDA07F302CFD61FCC9FCFE" author="Preston HB" pageId="1" pageNumber="214" pagination="206" refId="ref4023" refString="Preston HB (1907) Description of a new subgenus and species of Alycaeus from Ke-Lan-Tan. Proceedings of the Malacological Society of London, 7: 206." type="journal article" year="1907">Preston, 1907</bibRefCitation>
</taxonomicName>
(Malay Peninsula and Sumatra). Some populations of the latter species have a slightly longer tube than the other species, but have the typical rather globular shell shape with strong radial sculpture.
</paragraph>
<paragraph id="8BEB36FEFFDDA07F3653FC22FAECFB1D" blockId="1.[811,1451,957,1238]" pageId="1" pageNumber="214">
Originally, the subgenus
<taxonomicName id="4C544D7DFFDDA07F3151FC21FB26FC1E" authorityName="Preston" authorityYear="1907" box="[1070,1166,958,981]" class="Gastropoda" family="Cyclophoridae" genus="Pincerna" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="genus">
<emphasis id="B920EAECFFDDA07F3151FC21FB26FC1E" box="[1070,1166,958,981]" italics="true" pageId="1" pageNumber="214">Pincerna</emphasis>
</taxonomicName>
was diagnosed on the basis of a “circular cup” on the outer surface of the operculum. The outer surface of operculum, however, has limited taxonomic value on the genus level in the
<taxonomicName id="4C544D7DFFDDA07F31F8FB82FAAAFBFE" authorityName="W. T. Blanford" authorityYear="1864" box="[1159,1282,1053,1077]" class="Gastropoda" family="Alycaeidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="family">Alycaeidae</taxonomicName>
, especially that outer rings have been developed in multiple alycaeid genera (Páll-Gergely et al., in press). Consequently, no important shell characters distinguish
<taxonomicName id="4C544D7DFFDDA07F3113FBE2FB78FB5E" authorityName="Godwin-Austen" authorityYear="1914" box="[1132,1232,1149,1173]" class="Gastropoda" family="Cyclophoridae" genus="Cycloryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="genus">
<emphasis id="B920EAECFFDDA07F3113FBE2FB78FB5E" box="[1132,1232,1149,1173]" italics="true" pageId="1" pageNumber="214">Cycloryx</emphasis>
</taxonomicName>
and
<taxonomicName id="4C544D7DFFDDA07F3073FBE1FAD9FB5E" authorityName="Preston" authorityYear="1907" box="[1292,1393,1150,1173]" class="Gastropoda" family="Cyclophoridae" genus="Pincerna" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="genus">
<emphasis id="B920EAECFFDDA07F3073FBE1FAD9FB5E" box="[1292,1393,1150,1173]" italics="true" pageId="1" pageNumber="214">Pincerna</emphasis>
</taxonomicName>
, and they should be synonymised. Because
<taxonomicName id="4C544D7DFFDDA07F31A2FB01FAE8FB7E" authorityName="Preston" authorityYear="1907" box="[1245,1344,1182,1205]" class="Gastropoda" family="Cyclophoridae" genus="Pincerna" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="genus">
<emphasis id="B920EAECFFDDA07F31A2FB01FAE8FB7E" box="[1245,1344,1182,1205]" italics="true" pageId="1" pageNumber="214">Pincerna</emphasis>
</taxonomicName>
has been introduced earlier,
<taxonomicName id="4C544D7DFFDDA07F3683FB22FBF4FB1E" authorityName="Godwin-Austen" authorityYear="1914" box="[1020,1116,1213,1237]" class="Gastropoda" family="Cyclophoridae" genus="Cycloryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="214" phylum="Mollusca" rank="genus">
<emphasis id="B920EAECFFDDA07F3683FB22FBF4FB1E" box="[1020,1116,1213,1237]" italics="true" pageId="1" pageNumber="214">Cycloryx</emphasis>
</taxonomicName>
is a junior synonym.
</paragraph>
</subSubSection>
</treatment>
</document>