A new species of Alycaeidae, Pincerna yanseni n. sp. from Sumatra, with the resurrection of the genus Pincerna Preston, 1907 (Gastropoda: Cyclophoroidea)
Author
Páll-Gergely, Barna
text
Raffles Bulletin of Zoology
2017
2017-06-29
65
213
219
journal article
6569
10.5281/zenodo.4502847
3fbb2cbb-07dc-47ac-ae8a-257f70c83339
2345-7600
4502847
4CC08D61-7547-4F4B-BCE7-036ABF61D402
Genus
Pincerna
Preston, 1907
Pincerna
Preston, 1907: 206
. (introduced as a subgenus of
Alycaeus
, but apparently used on genus level).
Alycaeus
(
Cycloryx
) Godwin-Austen, 1914: 334
.
New synonym
Type
species.
Pincerna liratula
Preston, 1907
, by monotypy.
Alycaeus thieroti
Morgan, 1885
(
Type
locality: “G. Lano” near
Perak
) is very similar to
Pincerna liratula
(“Ke-lantan” in the Malay Peninsula); the latter might be a junior synonym of the former.
Remarks.
The genus-level revision of the family
Alycaeidae
is in progress and beyond the scope of the present paper. However, to place the new species in a genus correctly, some information on the relationship of the genus
Cycloryx
Godwin-Austen, 1914
, and
Pincerna
Preston, 1907
, must be added.
The discovery of
Pincerna yanseni
n. sp.
is surprising because it shows only superficial similarity with most other
Alycaeidae
of the Malay Archipelago, but it is similar to many species from northeastern
India
and
Myanmar
classified in the genus
Cycloryx
. The genus
Cycloryx
(
type
species:
Alycaeus constrictus
Benson, 1851
by original designation,
Fig. 1A
) was erected as a subgenus of
Alycaeus
Baird, 1850
, and was diagnosed on the basis of the ovately conoid shell shape, the regular ribbing on the upper whorls, and the extremely short, often clubbed or pear-shaped sutural tube (Godwin-Austen, 1914). Godwin-Austen (1914) only included species from northeastern
India
and
Burma
(Rakhin =
Arakan
, and the
Shan
States). However, the diagnosis of
Cycloryx
matches several species extralimital to the distributional range as defined by Godwin-Austen:
Alycaeus costulosus
Bavay & Dautzenberg, 1912
(northern
Vietnam
),
Alycaeus globosus
H.
Adams, 1870
and its subspecies (Borneo),
Dioryx maolanensis
Luo et al., 2009
(
Guizhou
,
China
),
Alycaeus thieroti
Morgan, 1885
(Perak, Malay Peninsula), and
Pincerna liratula
Preston, 1907
(Malay Peninsula and Sumatra). Some populations of the latter species have a slightly longer tube than the other species, but have the typical rather globular shell shape with strong radial sculpture.
Originally, the subgenus
Pincerna
was diagnosed on the basis of a “circular cup” on the outer surface of the operculum. The outer surface of operculum, however, has limited taxonomic value on the genus level in the
Alycaeidae
, especially that outer rings have been developed in multiple alycaeid genera (Páll-Gergely et al., in press). Consequently, no important shell characters distinguish
Cycloryx
and
Pincerna
, and they should be synonymised. Because
Pincerna
has been introduced earlier,
Cycloryx
is a junior synonym.