250 lines
37 KiB
XML
250 lines
37 KiB
XML
<document id="762B7C9129B2A2FB8885AEBEE09B3B24" ID-DOI="10.1186/s13358-023-00296-0" ID-ISSN="1664-2384" ID-Zenodo-Dep="12003174" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1718654193459" checkinUser="felipe" docAuthor="El-Desouky, Heba, Herbig, Hans-Georg & Kora, Mahmoud" docDate="2023" docId="493869167F12FFF5FC91F953FABFFDEE" docLanguage="en" docName="SwissJPalaeontol.142.32.pdf" docOrigin="Swiss Journal of Palaeontology (32) 142 (1)" docSource="http://dx.doi.org/10.1186/s13358-023-00296-0" docStyle="DocumentStyle:DBDC680DC95E08776BDF6354C30A315B.1:SwissJPalaeontol.2022-.journal_article" docStyleId="DBDC680DC95E08776BDF6354C30A315B" docStyleName="SwissJPalaeontol.2022-.journal_article" docStyleVersion="1" docTitle="Lytvolasma aheimerensis El-Desouky & Herbig & Kora 2023, n. sp." docType="treatment" docVersion="3" lastPageNumber="14" masterDocId="B501116E7F19FFF8FFBCFFFFFFE2FF8B" masterDocTitle="Kasimovian (late Pennsylvanian) cornute rugose corals from Egypt: taxonomy, facies and palaeogeography of a cool-water fauna from northern Gondwana" masterLastPageNumber="39" masterPageNumber="1" pageNumber="12" updateTime="1721391550094" updateUser="guilherme" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="D4F271758A9FFAC7EE45D358CC1538CD">
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<mods:title id="719D1D9084549033FE89C855C0BF9FD4">Kasimovian (late Pennsylvanian) cornute rugose corals from Egypt: taxonomy, facies and palaeogeography of a cool-water fauna from northern Gondwana</mods:title>
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<mods:name id="1B019C99FA52F82BE49200879999BC26" type="personal">
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<mods:roleTerm id="B287D7D0D859D68678E2BBD802C63AA4">Author</mods:roleTerm>
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<mods:namePart id="0F394ECC97BFDA2D8391E7EE68191D9F">El-Desouky, Heba</mods:namePart>
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<mods:roleTerm id="FC888FAA46EC4E8FB93B910FDA609B87">Author</mods:roleTerm>
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<mods:namePart id="5E01D203C8D2678ACD4E5AAC7B536F80">Herbig, Hans-Georg</mods:namePart>
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</mods:name>
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<mods:name id="0564A3E323F60974501AE7CEE1EFD086" type="personal">
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<mods:roleTerm id="42035BEF6335C2E5AAB5C04A05A8C6EF">Author</mods:roleTerm>
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<mods:namePart id="B06E4CD970719E28258F7AA1439EC033">Kora, Mahmoud</mods:namePart>
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<mods:typeOfResource id="01333D3F944F1C337892798CC1B7891D">text</mods:typeOfResource>
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<mods:title id="60257E3DFD2FA042234BF5EBC3A39C94">Swiss Journal of Palaeontology</mods:title>
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<mods:part id="C227BC3301CC0825F4879444C0F35D43">
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<mods:date id="B567F4B12A222BF0A3E702B86AAEB0DE">2023</mods:date>
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<mods:title id="01D6619F0F020337FAA3AB8E9C580D04">32</mods:title>
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<mods:number id="768180E3805C5A40020E767E99EEDB2C">2023-11-27</mods:number>
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<mods:number id="321F772E53F5B356F5E5FD49C2CDABDA">142</mods:number>
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<mods:url id="58B160FBFBECCFC857E8E98B18D108E6">http://dx.doi.org/10.1186/s13358-023-00296-0</mods:url>
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<mods:classification id="D4E4859DE8F9F0B9E545A7E56BBE2948">journal article</mods:classification>
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<mods:identifier id="1788327AF2D2287F1785D7A2A77D563F" type="DOI">10.1186/s13358-023-00296-0</mods:identifier>
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<mods:identifier id="EC2D48A935DC10F26C921CA04DDD6DB6" type="ISSN">1664-2384</mods:identifier>
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<mods:identifier id="16F466375562FC74F220666C08022C46" type="Zenodo-Dep">12003174</mods:identifier>
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<treatment id="493869167F12FFF5FC91F953FABFFDEE" LSID="urn:lsid:plazi:treatment:493869167F12FFF5FC91F953FABFFDEE" httpUri="http://treatment.plazi.org/id/493869167F12FFF5FC91F953FABFFDEE" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12">
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<subSubSection id="898B8B8B7F12FFF3FC91F953FB73F94E" box="[813,1169,1708,1733]" pageId="11" pageNumber="12" type="nomenclature">
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<paragraph id="C12ED8007F12FFF3FC91F953FB73F94E" blockId="11.[813,1169,1708,1733]" box="[813,1169,1708,1733]" pageId="11" pageNumber="12">
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<heading id="9A666F6C7F12FFF3FC91F953FB73F94E" bold="true" box="[813,1169,1708,1733]" fontSize="10" level="1" pageId="11" pageNumber="12" reason="1">
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<emphasis id="F3E504127F12FFF3FC91F953FB73F94E" bold="true" box="[813,1169,1708,1733]" pageId="11" pageNumber="12">
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<taxonomicName id="0691A3837F12FFF3FC91F953FBACF94F" authority="El-Desouky & Herbig & Kora, 2023" authorityName="El-Desouky & Herbig & Kora" authorityYear="2023" box="[813,1102,1708,1732]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stauriida" pageId="11" pageNumber="12" phylum="Cnidaria" rank="species" species="aheimerensis" status="sp. nov.">
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<emphasis id="F3E504127F12FFF3FC91F953FBACF94F" bold="true" box="[813,1102,1708,1732]" italics="true" pageId="11" pageNumber="12">Lytvolasma aheimerensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E8D6B9697F12FFF3FBE8F952FB73F94E" box="[1108,1169,1709,1733]" pageId="11" pageNumber="12" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="898B8B8B7F12FFF3FC91F933FBF8F8AF" pageId="11" pageNumber="12" type="description">
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<paragraph id="C12ED8007F12FFF3FC91F933FC48F96E" blockId="11.[813,1437,1740,1892]" box="[813,938,1740,1765]" pageId="11" pageNumber="12">
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(
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<figureCitation id="59AAC4857F12FFF3FC88F933FC7CF96E" box="[820,926,1740,1765]" captionStart="Fig" captionStartId="12.[167,198,1342,1362]" captionTargetBox="[285,1299,238,1327]" captionTargetId="figure-189@12.[283,1303,236,1329]" captionTargetPageId="12" captionText="Fig. 7 Lytvolasma aheimerensis n. sp. A External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.B–D Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in D. E–G Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end. H–I Two transverse thin-sections below the calice, deformed due to rejuvenation in H and lateral compaction in I. Scale bars: 2.5 mm, except A: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003254" httpUri="https://zenodo.org/record/12003254/files/figure.png" pageId="11" pageNumber="12">Fig. 7A–I</figureCitation>
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).
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</paragraph>
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<paragraph id="C12ED8007F12FFF3FC91F913FBF8F8AF" blockId="11.[813,1437,1740,1892]" pageId="11" pageNumber="12">
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<emphasis id="F3E504127F12FFF3FC91F913FBA9F88E" bold="true" box="[813,1099,1772,1797]" pageId="11" pageNumber="12">Derivation of the name</emphasis>
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: From the bearing rock unit “Aheimer Formation".
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</paragraph>
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</subSubSection>
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<subSubSection id="898B8B8B7F12FFF4FC91F8D2FA7AF9D3" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="materials_examined">
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<paragraph id="C12ED8007F12FFF3FC91F8D2FAE1F8EF" blockId="11.[813,1437,1740,1892]" pageId="11" pageNumber="12">
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<materialsCitation id="71F9D25D7F12FFF3FC91F8D2FAE1F8EF" pageId="11" pageNumber="12" specimenCode="RAh 2" specimenCount="1" typeStatus="holotype">
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<emphasis id="F3E504127F12FFF3FC91F8D2FC7DF8CE" bold="true" box="[813,927,1837,1861]" pageId="11" pageNumber="12">
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<typeStatus id="1E2A66A27F12FFF3FC91F8D2FC79F8CE" box="[813,923,1837,1861]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<specimenCount id="D79713897F12FFF3FC11F8D3FBFAF8CF" box="[941,1048,1836,1860]" pageId="11" pageNumber="12" type="generic">Specimen</specimenCount>
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<specimenCode id="9137707B7F12FFF3FB9BF8D3FB94F8CE" box="[1063,1142,1836,1861]" pageId="11" pageNumber="12">RAh 2</specimenCode>
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, illustrated in
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<figureCitation id="59AAC4857F12FFF3FA94F8D3FA7BF8CE" box="[1320,1433,1836,1861]" captionStart="Fig" captionStartId="12.[167,198,1342,1362]" captionTargetBox="[285,1299,238,1327]" captionTargetId="figure-189@12.[283,1303,236,1329]" captionTargetPageId="12" captionText="Fig. 7 Lytvolasma aheimerensis n. sp. A External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.B–D Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in D. E–G Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end. H–I Two transverse thin-sections below the calice, deformed due to rejuvenation in H and lateral compaction in I. Scale bars: 2.5 mm, except A: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003254" httpUri="https://zenodo.org/record/12003254/files/figure.png" pageId="11" pageNumber="12">Fig. 7A–I</figureCitation>
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. Eight transverse thin-sections are available.
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</materialsCitation>
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</paragraph>
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<caption id="95EE88887F15FFF4FF1BFAC1FA8FFA5C" ID-DOI="http://doi.org/10.5281/zenodo.12003254" ID-Zenodo-Dep="12003254" httpUri="https://zenodo.org/record/12003254/files/figure.png" pageId="12" pageNumber="13" startId="12.[167,198,1342,1362]" targetBox="[285,1299,238,1327]" targetPageId="12" targetType="figure">
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<paragraph id="C12ED8007F15FFF4FF1BFAC1FA8FFA5C" blockId="12.[167,1399,1342,1495]" pageId="12" pageNumber="13">
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<emphasis id="F3E504127F15FFF4FF1BFAC1FF38FAD9" bold="true" box="[167,218,1342,1362]" pageId="12" pageNumber="13">Fig. 7</emphasis>
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<taxonomicName id="0691A3837F15FFF4FF58FAC1FE52FAD9" authority="El-Desouky & Herbig & Kora, 2023" authorityName="El-Desouky & Herbig & Kora" authorityYear="2023" box="[228,432,1342,1362]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="13" phylum="Cnidaria" rank="species" species="aheimerensis" status="sp. nov.">
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<emphasis id="F3E504127F15FFF4FF58FAC1FE52FAD9" box="[228,432,1342,1362]" italics="true" pageId="12" pageNumber="13">Lytvolasma aheimerensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E8D6B9697F15FFF4FE09FAC1FE03FAD9" box="[437,481,1342,1362]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
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<emphasis id="F3E504127F15FFF4FE58FAC1FE10FAD9" bold="true" box="[484,498,1342,1362]" pageId="12" pageNumber="13">A</emphasis>
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External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.
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<emphasis id="F3E504127F15FFF4FD5EFAA7FCEAFAE7" bold="true" box="[738,776,1368,1388]" pageId="12" pageNumber="13">B–D</emphasis>
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Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in
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<emphasis id="F3E504127F15FFF4FC56FA8CFC1BFA0C" bold="true" box="[1002,1017,1395,1415]" pageId="12" pageNumber="13">D</emphasis>
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.
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<emphasis id="F3E504127F15FFF4FBBDFA8CFBC4FA0C" bold="true" box="[1025,1062,1395,1415]" pageId="12" pageNumber="13">E–G</emphasis>
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Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end.
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<emphasis id="F3E504127F15FFF4FE1BFA57FE24FA37" bold="true" box="[423,454,1448,1468]" pageId="12" pageNumber="13">H–I</emphasis>
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Two transverse thin-sections below the calice, deformed due to rejuvenation in
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<emphasis id="F3E504127F15FFF4FBD1FA57FB99FA37" bold="true" box="[1133,1147,1448,1468]" pageId="12" pageNumber="13">H</emphasis>
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and lateral compaction in
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<emphasis id="F3E504127F15FFF4FAE2FA57FA86FA37" bold="true" box="[1374,1380,1448,1468]" pageId="12" pageNumber="13">I</emphasis>
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. Scale bars: 2.5 mm, except
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<emphasis id="F3E504127F15FFF4FE37FA3CFE7BFA5C" bold="true" box="[395,409,1475,1495]" pageId="12" pageNumber="13">A</emphasis>
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: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa
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</paragraph>
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</caption>
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<paragraph id="C12ED8007F15FFF4FF2BF9DFFE94F953" blockId="12.[151,775,1568,1881]" pageId="12" pageNumber="13">
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<emphasis id="F3E504127F15FFF4FF2BF9DFFEE0F9B3" bold="true" box="[151,258,1568,1592]" pageId="12" pageNumber="13">Material:</emphasis>
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In addition to the
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<typeStatus id="1E2A66A27F15FFF4FE5AF9DFFDAAF9B3" box="[486,584,1568,1592]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
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, two moderately preserved
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<typeStatus id="1E2A66A27F15FFF4FEACF9BEFE9BF9D2" box="[272,377,1601,1625]" pageId="12" pageNumber="13" type="paratype">paratypes</typeStatus>
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with compressed to crushed calices and eroded apical ends (RAh 10 and RAh 113). Five transverse thin-sections are available. They are collected from the basal shales of the lower member of the Aheimer Formation.
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</paragraph>
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<paragraph id="C12ED8007F15FFF4FF2BF91FFF3FF8D2" blockId="12.[151,775,1568,1881]" pageId="12" pageNumber="13">
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<emphasis id="F3E504127F15FFF4FF2BF91FFED5F973" bold="true" box="[151,311,1760,1784]" pageId="12" pageNumber="13">Type locality:</emphasis>
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<materialsCitation id="71F9D25D7F15FFF4FE83F91EFF3BF8D2" country="Egypt" county="Gulf" latitude="29.4725" location="Northern Galala" longLatPrecision="20" longitude="32.460556" municipality="northern Eastern Desert" pageId="12" pageNumber="13" specimenCount="1" stateProvince="As Suways">
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Eastern cliffs of the Northern Galala plateau (Lat. 29° 28
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<emphasis id="F3E504127F15FFF4FEF4F900FEB1F893" box="[328,339,1791,1816]" italics="true" pageId="12" pageNumber="13">′</emphasis>
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21
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<emphasis id="F3E504127F15FFF4FECFF900FE61F893" box="[371,387,1791,1816]" italics="true" pageId="12" pageNumber="13">″</emphasis>
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N and Long. 32° 27
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<emphasis id="F3E504127F15FFF4FDE6F900FD87F893" box="[602,613,1791,1816]" italics="true" pageId="12" pageNumber="13">′</emphasis>
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38
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<emphasis id="F3E504127F15FFF4FD3AF900FD74F893" box="[646,662,1791,1816]" italics="true" pageId="12" pageNumber="13">″</emphasis>
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E), northern Eastern Desert, western side of the Gulf of Suez, Egypt
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="C12ED8007F15FFF4FC91F9DFFA7AF9D3" blockId="12.[813,1437,1568,1912]" pageId="12" pageNumber="13">
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<emphasis id="F3E504127F15FFF4FC91F9DFFC32F9B3" bold="true" box="[813,976,1568,1592]" pageId="12" pageNumber="13">Type horizon:</emphasis>
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Basal shales of the lower member of the Aheimer Formation; Upper Pennsylvanian (Kasimovian).
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</paragraph>
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</subSubSection>
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<subSubSection id="898B8B8B7F15FFF4FC91F99FFCBEF973" pageId="12" pageNumber="13" type="diagnosis">
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<paragraph id="C12ED8007F15FFF4FC91F99FFCBEF973" blockId="12.[813,1437,1568,1912]" pageId="12" pageNumber="13">
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<emphasis id="F3E504127F15FFF4FC91F99FFC44F9F3" bold="true" box="[813,934,1632,1656]" pageId="12" pageNumber="13">Diagnosis:</emphasis>
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<taxonomicName id="0691A3837F15FFF4FC10F99FFBCAF9F3" box="[940,1064,1632,1656]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="13" phylum="Cnidaria" rank="genus">
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<emphasis id="F3E504127F15FFF4FC10F99FFBCAF9F3" box="[940,1064,1632,1656]" italics="true" pageId="12" pageNumber="13">Lytvolasma</emphasis>
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</taxonomicName>
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with thick wall, up to
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<quantity id="066975E57F15FFF4FAA5F99FFABBF9F3" box="[1305,1369,1632,1656]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="13" unit="mm" value="2.0">2 mm</quantity>
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thick.
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<taxonomicName id="0691A3837F15FFF4FC91F97FFC69F913" box="[813,907,1664,1688]" genus="Cardinal" pageId="12" pageNumber="8" rank="genus">Cardinal</taxonomicName>
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septum shortened from early maturity. Counter septum elongated up to late maturity without forming a columella. It enters the free axial area with its rhopaloid end.
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</paragraph>
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</subSubSection>
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<subSubSection id="898B8B8B7F15FFF5FC91F8FFFE55FAA8" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="description">
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<paragraph id="C12ED8007F15FFF5FC91F8FFFD95FEE8" blockId="12.[813,1437,1568,1912]" lastBlockId="13.[151,775,235,1571]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
|
||
<emphasis id="F3E504127F15FFF4FC91F8FFFC5FF893" bold="true" box="[813,957,1792,1816]" pageId="12" pageNumber="13">Description:</emphasis>
|
||
External characters: Solitary, small, conicocylindrical corallites. The preserved parts of the three corallites ranges in length between 2 and
|
||
<quantity id="066975E57F15FFF4FAAEF8BFFAADF8D3" box="[1298,1359,1856,1880]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="13" unit="cm" value="3.0">3 cm</quantity>
|
||
. Moderately preserved; their apexes and much of their early neanic stages are missing. Calices are deep. Growth lines, rugae, septal ridges and furrows are well-developed on the outer wall of the corallites. Rejuvenation is not common, only constrictions are visible (
|
||
<figureCitation id="59AAC4857F14FFF5FDA5FEB4FD89FEE8" box="[537,619,331,355]" captionStart="Fig" captionStartId="12.[167,198,1342,1362]" captionTargetBox="[285,1299,238,1327]" captionTargetId="figure-189@12.[283,1303,236,1329]" captionTargetPageId="12" captionText="Fig. 7 Lytvolasma aheimerensis n. sp. A External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.B–D Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in D. E–G Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end. H–I Two transverse thin-sections below the calice, deformed due to rejuvenation in H and lateral compaction in I. Scale bars: 2.5 mm, except A: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003254" httpUri="https://zenodo.org/record/12003254/files/figure.png" pageId="13" pageNumber="14">Fig. 7A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C12ED8007F14FFF5FF11FE94FE55FAA8" blockId="13.[151,775,235,1571]" pageId="13" pageNumber="14">
|
||
Internal characters: The ontogenetically earliest growth stage available has 23 septa in 5.9 mm (
|
||
<figureCitation id="59AAC4857F14FFF5FD0DFE74FF4EFE48" captionStart="Fig" captionStartId="12.[167,198,1342,1362]" captionTargetBox="[285,1299,238,1327]" captionTargetId="figure-189@12.[283,1303,236,1329]" captionTargetPageId="12" captionText="Fig. 7 Lytvolasma aheimerensis n. sp. A External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.B–D Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in D. E–G Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end. H–I Two transverse thin-sections below the calice, deformed due to rejuvenation in H and lateral compaction in I. Scale bars: 2.5 mm, except A: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003254" httpUri="https://zenodo.org/record/12003254/files/figure.png" pageId="13" pageNumber="14">Fig. 7B, C</figureCitation>
|
||
). Major septa are radially arranged. They are very thick and joined together in the centre forming a strong axial structure. The cardinal septum is somewhat thinner and the fossular break is noticeable. Further in the ontogeny, at a diameter of 8.8–13.3 mm with 24–26 septa (
|
||
<figureCitation id="59AAC4857F14FFF5FF5EFDB4FEBCFDE8" box="[226,350,587,611]" captionStart="Fig" captionStartId="12.[167,198,1342,1362]" captionTargetBox="[285,1299,238,1327]" captionTargetId="figure-189@12.[283,1303,236,1329]" captionTargetPageId="12" captionText="Fig. 7 Lytvolasma aheimerensis n. sp. A External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.B–D Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in D. E–G Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end. H–I Two transverse thin-sections below the calice, deformed due to rejuvenation in H and lateral compaction in I. Scale bars: 2.5 mm, except A: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003254" httpUri="https://zenodo.org/record/12003254/files/figure.png" pageId="13" pageNumber="14">Fig. 7D–H</figureCitation>
|
||
), the septa become thinner gradually with maturity, and are more radially arranged. They are united near the axis with their rhopaloid ends, forming a stereozone with a free axial area. A strange attitude of one of the lateral cardinal septa is shown in
|
||
<figureCitation id="59AAC4857F14FFF5FD29FD34FCE0FD68" box="[661,770,715,739]" captionStart="Fig" captionStartId="12.[167,198,1342,1362]" captionTargetBox="[285,1299,238,1327]" captionTargetId="figure-189@12.[283,1303,236,1329]" captionTargetPageId="12" captionText="Fig. 7 Lytvolasma aheimerensis n. sp. A External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.B–D Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in D. E–G Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end. H–I Two transverse thin-sections below the calice, deformed due to rejuvenation in H and lateral compaction in I. Scale bars: 2.5 mm, except A: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003254" httpUri="https://zenodo.org/record/12003254/files/figure.png" pageId="13" pageNumber="14">Fig. 7G–I</figureCitation>
|
||
. It started to diminish gradually with maturity. This led to the reduction of the number of septa below the calice (
|
||
<figureCitation id="59AAC4857F14FFF5FF23FCD4FF0BFCC8" box="[159,233,811,835]" captionStart="Fig" captionStartId="12.[167,198,1342,1362]" captionTargetBox="[285,1299,238,1327]" captionTargetId="figure-189@12.[283,1303,236,1329]" captionTargetPageId="12" captionText="Fig. 7 Lytvolasma aheimerensis n. sp. A External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.B–D Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in D. E–G Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end. H–I Two transverse thin-sections below the calice, deformed due to rejuvenation in H and lateral compaction in I. Scale bars: 2.5 mm, except A: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003254" httpUri="https://zenodo.org/record/12003254/files/figure.png" pageId="13" pageNumber="14">Fig. 7I</figureCitation>
|
||
) to 25 septa. This attitude might be attributed to the rejuvenation took place in this part of the corallite. The cardinal fossula is slightly triangular and open adaxially. Counter septum is the longest with swollen end, invading the free axial area (
|
||
<figureCitation id="59AAC4857F14FFF5FDA1FC54FD8DFC48" box="[541,623,939,963]" captionStart="Fig" captionStartId="12.[167,198,1342,1362]" captionTargetBox="[285,1299,238,1327]" captionTargetId="figure-189@12.[283,1303,236,1329]" captionTargetPageId="12" captionText="Fig. 7 Lytvolasma aheimerensis n. sp. A External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.B–D Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in D. E–G Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end. H–I Two transverse thin-sections below the calice, deformed due to rejuvenation in H and lateral compaction in I. Scale bars: 2.5 mm, except A: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003254" httpUri="https://zenodo.org/record/12003254/files/figure.png" pageId="13" pageNumber="14">Fig. 7F</figureCitation>
|
||
), and irregular in shape. Alar fossulae are not developed. The most adult stage, below the calice and the calice itself are compacted and the septa are somewhat deformed, the stereozone gradually diminishes and the septa retreat a little from the centre (
|
||
<figureCitation id="59AAC4857F14FFF5FE04FBB4FDE7FBE8" box="[440,517,1099,1123]" captionStart="Fig" captionStartId="12.[167,198,1342,1362]" captionTargetBox="[285,1299,238,1327]" captionTargetId="figure-189@12.[283,1303,236,1329]" captionTargetPageId="12" captionText="Fig. 7 Lytvolasma aheimerensis n. sp. A External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.B–D Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in D. E–G Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end. H–I Two transverse thin-sections below the calice, deformed due to rejuvenation in H and lateral compaction in I. Scale bars: 2.5 mm, except A: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003254" httpUri="https://zenodo.org/record/12003254/files/figure.png" pageId="13" pageNumber="14">Fig. 7I</figureCitation>
|
||
), the cardinal septum becomes distinctively shorter than the other major septa, whereas the counter is still long with thick rhopaloid end. The wall is thick (0.9–
|
||
<quantity id="066975E57F14FFF5FDA7FB54FDBDFB48" box="[539,607,1195,1219]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="14" unit="mm" value="2.0">2 mm</quantity>
|
||
) in all ontogenetic stages. Minor septa are absent. A white line in the midlines of the septa may indicate their original compound trabecular growth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="898B8B8B7F14FFF5FF2BFAD4FABFFDEE" pageId="13" pageNumber="14" type="discussion">
|
||
<paragraph id="C12ED8007F14FFF5FF2BFAD4FED3F9A8" blockId="13.[151,775,235,1571]" pageId="13" pageNumber="14">
|
||
<emphasis id="F3E504127F14FFF5FF2BFAD4FEC2FAC8" bold="true" box="[151,288,1323,1347]" pageId="13" pageNumber="14">Discussion:</emphasis>
|
||
The elongation of the counter septum makes this species similar to
|
||
<taxonomicName id="0691A3837F14FFF5FE53FAB4FF2FFA08" authority="Fomichev, 1953" authorityName="Fomichev, Shortened" authorityYear="1953" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="F3E504127F14FFF5FE53FAB4FD69FAE8" box="[495,651,1355,1379]" italics="true" pageId="13" pageNumber="14">Monophyllum</emphasis>
|
||
<bibRefCitation id="A500A5F17F14FFF5FD29FAB4FF2FFA08" author="Fomichev, V." pageId="13" pageNumber="14" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
|
||
</taxonomicName>
|
||
. It can be differentiated from
|
||
<taxonomicName id="0691A3837F14FFF5FD9EFA94FD5CFA08" authorityName="Fomichev, Shortened" authorityYear="1953" box="[546,702,1387,1411]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="F3E504127F14FFF5FD9EFA94FD5CFA08" box="[546,702,1387,1411]" italics="true" pageId="13" pageNumber="14">Monophyllum</emphasis>
|
||
</taxonomicName>
|
||
in the shape of the cardinal fossula and the lack of the incipient columella formation in the calice. The current species differs from
|
||
<taxonomicName id="0691A3837F14FFF5FEDDFA33FD74FA68" authority="Fedorowski, 1987" authorityName="Fedorowski" authorityYear="1987" box="[353,662,1483,1507]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="aucta">
|
||
<emphasis id="F3E504127F14FFF5FEDDFA33FE23FA68" box="[353,449,1484,1507]" italics="true" pageId="13" pageNumber="14">L. aucta</emphasis>
|
||
Fedorowski, 1987
|
||
</taxonomicName>
|
||
from the Pennsylvanian–Permian transition of southern
|
||
<collectingRegion id="035516E27F14FFF5FD17FA13FD08F988" box="[683,746,1516,1539]" country="United States of America" name="Texas" pageId="13" pageNumber="14">Texas</collectingRegion>
|
||
in the following:
|
||
</paragraph>
|
||
<paragraph id="C12ED8007F14FFF5FF10F9B2FD81F90E" blockId="13.[172,775,1613,1861]" pageId="13" pageNumber="14">• Number of septa are higher and diameter (n:d ratio) is larger in the present species.</paragraph>
|
||
<paragraph id="C12ED8007F14FFF5FF10F972FEC2F94F" blockId="13.[172,775,1613,1861]" pageId="13" pageNumber="14">• The wall of the current species is remarkably thicker.</paragraph>
|
||
<paragraph id="C12ED8007F14FFF5FF10F932FDD9F8CF" blockId="13.[172,775,1613,1861]" pageId="13" pageNumber="14">
|
||
• Minor septa are short and restricted to the corallite wall in the latest mature stages of
|
||
<taxonomicName id="0691A3837F14FFF5FDFBF912FD43F88F" authorityName="Fedorowski" authorityYear="1987" box="[583,673,1773,1796]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="aucta">
|
||
<emphasis id="F3E504127F14FFF5FDFBF912FD43F88F" box="[583,673,1773,1796]" italics="true" pageId="13" pageNumber="14">L. aucta</emphasis>
|
||
</taxonomicName>
|
||
, whereas they are absent in the current species even in the last ephebic stage near the calice floor.
|
||
</paragraph>
|
||
<paragraph id="C12ED8007F14FFF5FCFEFF13FAAAFEC8" blockId="13.[834,1436,235,323]" pageId="13" pageNumber="14">
|
||
• The counter septum in the present species is longer, enters the free axial area, whereas it is only slightly longer than the other major septa in
|
||
<emphasis id="F3E504127F14FFF5FB56FED3FAAAFEC8" box="[1258,1352,300,323]" italics="true" pageId="13" pageNumber="14">
|
||
<taxonomicName id="0691A3837F14FFF5FB56FED3FAA1FEC8" authorityName="Fedorowski" authorityYear="1987" box="[1258,1347,300,323]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="aucta">L. aucta</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="C12ED8007F14FFF5FCFEFE92FABFFDEE" blockId="13.[813,1437,364,613]" pageId="13" pageNumber="14">
|
||
The type species of the genus,
|
||
<taxonomicName id="0691A3837F14FFF5FB2BFE92FC83FE2F" authority="Soshkina, 1925" authorityName="Soshkina" authorityYear="1925" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="asymetrica">
|
||
<emphasis id="F3E504127F14FFF5FB2BFE92FAC9FE0F" box="[1175,1323,365,388]" italics="true" pageId="13" pageNumber="14">L. asymetrica</emphasis>
|
||
<bibRefCitation id="A500A5F17F14FFF5FA8FFE93FC83FE2F" author="Soshkina, E. D." pageId="13" pageNumber="14" pagination="76 - 104" refId="ref29330" refString="Soshkina, E. D. (1925). Les coraux du Permien inferieur (etage d'Artinsk) du versant occidental de l'Oural. Bulletin De La Societe Des Naturalistes De Moscou, Section Geologique, 33, 76 - 104." type="journal article" year="1925">Soshkina, 1925</bibRefCitation>
|
||
</taxonomicName>
|
||
differs by the early appearance of free axial area in the
|
||
<typeStatus id="1E2A66A27F14FFF5FCE7FE53FC59FE4F" box="[859,955,428,452]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
(
|
||
<bibRefCitation id="A500A5F17F14FFF5FC72FE53FB42FE4E" author="Fedorowski, J." box="[974,1184,428,453]" pageId="13" pageNumber="14" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski, 1987a</bibRefCitation>
|
||
, with comparisons to further similar specimens from different taxa) and the appearance of minor septa (Chwieduk, 2013), though n:d ratios are similar.
|
||
<taxonomicName id="0691A3837F14FFF5FC4AFDF2FBB3FDAF" authorityName="Fedorowski" authorityYear="1987" box="[1014,1105,525,548]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="aucta">
|
||
<emphasis id="F3E504127F14FFF5FC4AFDF2FBB3FDAF" box="[1014,1105,525,548]" italics="true" pageId="13" pageNumber="14">L. aucta</emphasis>
|
||
</taxonomicName>
|
||
illustrated in
|
||
<bibRefCitation id="A500A5F17F14FFF5FB4CFDF2FC4CFDCE" author="Kora, M. & Mansour, Y." pageId="13" pageNumber="14" pagination="597 - 614" refId="ref27107" refString="Kora, M., & Mansour, Y. (1991). Late Carboniferous solitary rugose corals from the western side of the Gulf of Suez, Egypt. Neues Jahrbuch Fur Geologie Und Palaontologie Monatshefte, Stuttgart, 10, 597 - 614." type="journal article" year="1991">Kora and Mansour (1991)</bibRefCitation>
|
||
(=
|
||
<taxonomicName id="0691A3837F14FFF5FC6AFDD2FB85FDCF" authority="El-Desouky & Herbig & Kora, 2023" authorityName="El-Desouky & Herbig & Kora" authorityYear="2023" box="[982,1127,557,580]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="paraaucta" status="n. sp.">
|
||
<emphasis id="F3E504127F14FFF5FC6AFDD2FB85FDCF" box="[982,1127,557,580]" italics="true" pageId="13" pageNumber="14">L. paraaucta</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E8D6B9697F14FFF5FBCEFDD2FB52FDCE" box="[1138,1200,557,581]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
|
||
) has smaller dimensions and a lower n:d ratio than the present species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |