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<document id="114401E1F28E22CEE900490B3589E720" ID-DOI="10.11646/phytotaxa.395.4.4" ID-ISSN="1179-3163" ID-Zenodo-Dep="13717718" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725652298113" checkinUser="felipe" docAuthor="Attar, Farideh &amp; Rad, Mohammad Amini" docDate="2019" docId="A8671025FFDFFFD877B59BE80EA5DF03" docLanguage="en" docName="phytotaxa.395.4.4.pdf" docOrigin="Phytotaxa 395 (4)" docSource="http://dx.doi.org/10.11646/phytotaxa.395.4.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Cousinia sharifii Attar &amp; Amini Rad 2019, sp. nov." docType="treatment" docVersion="2" lastPageNumber="290" masterDocId="545E685DFFDFFFDB773D9C5C0C71D87B" masterDocTitle="Cousinia sharifii, a new species of Cousinia sect. Cynaroideae (Asteraceae) from NW Iran" masterLastPageNumber="295" masterPageNumber="287" pageNumber="287" updateTime="1725660927393" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="1AB94D886633D53CDF2F5AAF6962819A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E6B4540662A5EE4F108F83BDA00E0342">
<mods:title id="E4F285348ABBF82EA327903D29607506">Cousinia sharifii, a new species of Cousinia sect. Cynaroideae (Asteraceae) from NW Iran</mods:title>
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<mods:name id="C11A56286136E79B403C547CAA7AE802" type="personal">
<mods:role id="4BB68BA5B8CA93A9DFC84F1140999059">
<mods:roleTerm id="57C7E174AACD27E469E390AC120C5264">Author</mods:roleTerm>
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<mods:namePart id="6C4169B53A99F26D7F879C6582825467">Attar, Farideh</mods:namePart>
<mods:affiliation id="0EE3888ABC05BB10C651D08BF4296669">School of Biology and Center of Excellence in Phylogeny of Living Organisms, College of Science, Univ. of Tehran, Tehran, Iran.</mods:affiliation>
</mods:name>
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<mods:role id="96A6E703267EABC679AAFA992E70EAC0">
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<mods:namePart id="FFAF2955599871F45B32B8942ACE1A2D">Rad, Mohammad Amini</mods:namePart>
</mods:name>
<mods:typeOfResource id="D626E79B3C2BEE53F5DC72445C6AED07">text</mods:typeOfResource>
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<mods:title id="1010C12412E6310A73EB306C13BF922C">Phytotaxa</mods:title>
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<mods:part id="0D9D5B7000A315CF416552AA25C137B2">
<mods:date id="EDB3A33E641CA52122CBB4EB1ACB853D">2019</mods:date>
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<mods:number id="05DA6C76DFD4433D0686FF9CB527031E">2019-03-12</mods:number>
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<mods:number id="26377511F73AF3370770E26F3A4185BD">395</mods:number>
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<mods:classification id="5BF5B3717F6BF186CCA87245D585E9AC">journal article</mods:classification>
<mods:identifier id="E6B5A3D8E0EE404B96BF3F55161A5FAF" type="DOI">10.11646/phytotaxa.395.4.4</mods:identifier>
<mods:identifier id="7DB1BAA46F545347DA4703D72FBDD4CC" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="C173E5F7BB078CD5944F5DFF36EB9E67" type="Zenodo-Dep">13717718</mods:identifier>
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<treatment id="A8671025FFDFFFD877B59BE80EA5DF03" LSID="urn:lsid:plazi:treatment:A8671025FFDFFFD877B59BE80EA5DF03" httpUri="http://treatment.plazi.org/id/A8671025FFDFFFD877B59BE80EA5DF03" lastPageId="3" lastPageNumber="290" pageId="0" pageNumber="287">
<subSubSection id="68D4F2B8FFDFFFDB77B59BE80F5DDFB4" box="[136,812,1972,1999]" pageId="0" pageNumber="287" type="nomenclature">
<paragraph id="2071A133FFDFFFDB77B59BE80F5DDFB4" blockId="0.[136,812,1972,1999]" box="[136,812,1972,1999]" pageId="0" pageNumber="287">
<heading id="7B39165FFFDFFFDB77B59BE80F5DDFB4" box="[136,812,1972,1999]" fontSize="11" level="2" pageId="0" pageNumber="287" reason="3">
<taxonomicName id="E7CEDAB0FFDFFFDB77B59BE80E5CDFB4" ID-CoL="Z2GW" authority="Attar &amp; Amini Rad" authorityName="Attar &amp; Amini Rad" authorityYear="2019" box="[136,557,1972,1999]" class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="0" pageNumber="287" phylum="Tracheophyta" rank="species" species="sharifii" status="sp. nov.">
<emphasis id="12BA7D21FFDFFFDB77B59BE80D3ADFB4" bold="true" box="[136,331,1972,1999]" italics="true" pageId="0" pageNumber="287">Cousinia sharifii</emphasis>
Attar &amp; Amini Rad
</taxonomicName>
,
<emphasis id="12BA7D21FFDFFFDB75059BE80EFFDFB5" bold="true" box="[568,654,1972,1998]" italics="true" pageId="0" pageNumber="287">
<taxonomicNameLabel id="0989C05AFFDFFFDB75059BE80EFFDFB5" box="[568,654,1972,1998]" pageId="0" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="B8F5BDB6FFDFFFDB75A39BEA0E9ADFB4" box="[670,747,1974,1999]" captionStart="FIGURE 1" captionStartId="1.[136,229,2029,2050]" captionTargetBox="[189,1398,187,2001]" captionTargetId="figure-14@1.[189,1398,187,2001]" captionTargetPageId="1" captionText="FIGURE 1. Holotype of Cousinia sharifii." figureDoi="http://doi.org/10.5281/zenodo.13717720" httpUri="https://zenodo.org/record/13717720/files/figure.png" pageId="0" pageNumber="287">Figs. 1</figureCitation>
,
<figureCitation id="B8F5BDB6FFDFFFDB75C49BEA0F78DFB4" box="[761,777,1974,1999]" captionStart="FIGURE 2" captionStartId="2.[136,229,1374,1395]" captionTargetBox="[159,1426,190,1335]" captionTargetId="figure-17@2.[151,1436,190,1350]" captionTargetPageId="2" captionText="FIGURE 2. Cousinia sharifii: Amini Rad &amp; Torabi, 58564-IRAN." figureDoi="http://doi.org/10.5281/zenodo.13717724" httpUri="https://zenodo.org/record/13717724/files/figure.png" pageId="0" pageNumber="287">2</figureCitation>
,
<figureCitation id="B8F5BDB6FFDFFFDB74289BEA0F52DFB4" box="[789,803,1974,1999]" captionStart="FIGURE 3" captionStartId="3.[136,229,1170,1191]" captionTargetBox="[199,1393,193,1145]" captionTargetId="figure-14@3.[194,1394,190,1145]" captionTargetPageId="3" captionText="FIGURE 3. Distribution map of the species: Cousinia sharifii; C. eriocephala; C. qaradaghensis; C. zagrica." figureDoi="http://doi.org/10.5281/zenodo.13717726" httpUri="https://zenodo.org/record/13717726/files/figure.png" pageId="0" pageNumber="287">3</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="68D4F2B8FFDFFFD977B59B800935DF86" lastPageId="2" lastPageNumber="289" pageId="0" pageNumber="287" type="materials_examined">
<paragraph id="2071A133FFDFFFDB77B59B800911DF8A" blockId="0.[136,1376,2012,2033]" box="[136,1376,2012,2033]" pageId="0" pageNumber="287">
<emphasis id="12BA7D21FFDFFFDB77B59B800CB3DF8A" bold="true" box="[136,194,2012,2033]" pageId="0" pageNumber="287">Type:</emphasis>
—
<collectingCountry id="58D9E1A3FFDFFFDB77E69B800D69DF8A" box="[219,280,2012,2033]" name="Iran" pageId="0" pageNumber="287">IRAN</collectingCountry>
.
<collectionCode id="46DF39F6FFDFFFDB761C9B800D46DF8A" box="[289,311,2012,2033]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="0" pageNumber="287" type="Herbarium">W</collectionCode>
Azarbaijan: Piranshahr, Silveh, Mashkan,
<quantity id="E7360CD6FFDFFFDB75ED9B800F6ADF8A" box="[720,795,2012,2033]" metricMagnitude="3" metricUnit="m" metricValue="1.89" pageId="0" pageNumber="287" unit="m" value="1890.0">1890 m</quantity>
,
<date id="547087F3FFDFFFDB74189B800FE6DF8A" box="[805,919,2012,2033]" pageId="0" pageNumber="287" value="2012-07-03">3 July 2012</date>
,
<emphasis id="12BA7D21FFDFFFDB749C9B8008D6DF8A" box="[929,1191,2012,2033]" italics="true" pageId="0" pageNumber="287">Amini Rad &amp; Torabi 58564</emphasis>
(
<typeStatus id="FF751F91FFDFFFDB738E9B800978DF8A" box="[1203,1289,2012,2033]" pageId="0" pageNumber="287" type="holotype">holotype</typeStatus>
<collectingCountry id="58D9E1A3FFDFFFDB72329B800923DF8A" box="[1295,1362,2012,2033]" name="Iran" pageId="0" pageNumber="287">IRAN</collectingCountry>
!).
</paragraph>
<caption id="74B1F1BBFFDEFFDA77B59BB10E57D079" ID-DOI="http://doi.org/10.5281/zenodo.13717720" ID-Zenodo-Dep="13717720" box="[136,550,2029,2050]" httpUri="https://zenodo.org/record/13717720/files/figure.png" pageId="1" pageNumber="288" startId="1.[136,229,2029,2050]" targetBox="[189,1398,187,2001]" targetPageId="1" targetType="figure">
<paragraph id="2071A133FFDEFFDA77B59BB10E57D079" blockId="1.[136,550,2029,2050]" box="[136,550,2029,2050]" pageId="1" pageNumber="288">
<emphasis id="12BA7D21FFDEFFDA77B59BB10C8FD079" bold="true" box="[136,254,2029,2050]" pageId="1" pageNumber="288">FIGURE 1.</emphasis>
Holotype of
<taxonomicName id="E7CEDAB0FFDEFFDA76409BB10E53D079" authorityName="Attar &amp; Amini Rad" authorityYear="2019" box="[381,546,2029,2050]" class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="288" phylum="Tracheophyta" rank="species" species="sharifii">Cousinia sharifii</taxonomicName>
.
</paragraph>
</caption>
<caption id="74B1F1BBFFDDFFD977B599020F7DDD08" ID-DOI="http://doi.org/10.5281/zenodo.13717724" ID-Zenodo-Dep="13717724" box="[136,780,1374,1395]" httpUri="https://zenodo.org/record/13717724/files/figure.png" pageId="2" pageNumber="289" startId="2.[136,229,1374,1395]" targetBox="[159,1426,190,1335]" targetPageId="2" targetType="figure">
<paragraph id="2071A133FFDDFFD977B599020F7DDD08" blockId="2.[136,780,1374,1395]" box="[136,780,1374,1395]" pageId="2" pageNumber="289">
<emphasis id="12BA7D21FFDDFFD977B599020C8FDD08" bold="true" box="[136,254,1374,1395]" pageId="2" pageNumber="289">FIGURE 2.</emphasis>
<emphasis id="12BA7D21FFDDFFD9763E99020DDCDD08" box="[259,429,1374,1395]" italics="true" pageId="2" pageNumber="289">
<taxonomicName id="E7CEDAB0FFDDFFD9763E99020DD9DD08" authorityName="Attar &amp; Amini Rad" authorityYear="2019" box="[259,424,1374,1395]" class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="289" phylum="Tracheophyta" rank="species" species="sharifii">Cousinia sharifii</taxonomicName>
:
</emphasis>
Amini Rad &amp; Torabi, 58564-IRAN.
</paragraph>
</caption>
<paragraph id="2071A133FFDDFFD977B599FA0E78DECC" blockId="2.[136,1452,1446,1719]" pageId="2" pageNumber="289">
<emphasis id="12BA7D21FFDDFFD977B599FA0E78DECC" italics="true" pageId="2" pageNumber="289">
Planta perennis, monocarpica. Caulis usque ad
<quantity id="E7360CD6FFDDFFD9755F99FA0EECDDC0" box="[610,669,1446,1467]" metricMagnitude="-1" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="289" unit="cm" value="45.0">45 cm</quantity>
longus, ramosus, floccoso-araneosus. Folia coriacea, sinuato-dentata, spinosa, indistincte discoloria, supra araneoso-tomentosa, subtus dense albido-tomentosa; folia radicalia ignota; folia caulina decurrentia, parte liberi ca. 9 ×
<quantity id="E7360CD6FFDDFFD976B699B20DCDDE78" box="[395,444,1518,1539]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="289" unit="cm" value="6.0">6 cm</quantity>
(spinis inclusis); folia summa reducta, triangularia, basi rotundata, 5.5 ×
<quantity id="E7360CD6FFDDFFD9739399B2097DDE78" box="[1198,1292,1518,1539]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="2" pageNumber="289" unit="cm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 cm</quantity>
.
<taxonomicName id="E7CEDAB0FFDDFFD9722599B20D59DE5C" class="Magnoliopsida" family="Asteraceae" genus="Synflorescentia" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="289" phylum="Tracheophyta" rank="species" species="corymbosa">Synflorescentia corymbosa</taxonomicName>
; capitula
<quantity id="E7360CD6FFDDFFD976B59A4E0D9CDE5C" box="[392,493,1554,1575]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="2" pageNumber="289" unit="mm" value="27.5" valueMax="30.0" valueMin="25.0">25–30 mm</quantity>
longa, ca. 160-flora; pedunculis
<quantity id="E7360CD6FFDDFFD974119A4E0FF4DE5C" box="[812,901,1554,1575]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="2" pageNumber="289" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">5–10 mm</quantity>
longis; involucrum globosum, araneosum, spinis inclusis
<quantity id="E7360CD6FFDDFFD977809A6A0D53DE30" box="[189,290,1590,1611]" metricMagnitude="-2" metricUnit="m" metricValue="4.55" metricValueMax="4.8" metricValueMin="4.3" pageId="2" pageNumber="289" unit="mm" value="45.5" valueMax="48.0" valueMin="43.0">43–48 mm</quantity>
diametro, usque ad
<quantity id="E7360CD6FFDDFFD976DF9A6A0E53DE30" box="[482,546,1590,1611]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="289" unit="mm" value="30.0">30 mm</quantity>
longum; phylla ca. 100, coriacea, appendiculata; appendices triangulares; exteriora patentia; intermedia imbricata, margine supra medium utrinque 1 spinulis; interiora longiora, spinulis nullis; interiora et intima prominentia, exappendiculata; receptaculi setae scabrae.
<taxonomicName id="E7CEDAB0FFDDFFD975469A220F58DEE8" box="[635,809,1662,1683]" class="Magnoliopsida" family="Asteraceae" genus="Corolla" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="289" phylum="Tracheophyta" rank="species" species="purpurea">Corolla purpurea</taxonomicName>
,
<quantity id="E7360CD6FFDDFFD9740A9A220FD1DEE8" box="[823,928,1662,1683]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="2.4" metricValueMin="2.3" pageId="2" pageNumber="289" unit="mm" value="23.5" valueMax="24.0" valueMin="23.0">23–24 mm</quantity>
longa, tubo limbo
<quantity id="E7360CD6FFDDFFD9735B9A2208D8DEE8" box="[1126,1193,1662,1683]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="289" unit="mm" value="11.0">11 mm</quantity>
longo. Antherarum tubus concolor, glaber. Achaenia ignota.
</emphasis>
</paragraph>
<paragraph id="2071A133FFDDFFD977B59AB40935DF86" blockId="2.[136,1452,1768,2045]" pageId="2" pageNumber="289">
Perennial, monocarpic. Stem up to
<quantity id="E7360CD6FFDDFFD9751E9AB40E1DDF7A" box="[547,620,1768,1793]" metricMagnitude="-1" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="289" unit="cm" value="45.0">45 cm</quantity>
long, branched, floccose-arachnoid. Leaves coriaceous, sinuate-dentate, spiny, indistinctly discolored, arachnoid-tomentose above, densely arachnoid-tomentose below; radical leaves unknown; stem leaves decurrent, free part 9 ×
<quantity id="E7360CD6FFDDFFD9759C9B6C0EABDF32" box="[673,730,1840,1865]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="289" unit="cm" value="6.0">6 cm</quantity>
(spines included); upper leaves reduced, triangular, rounded at base, 5.5 ×
<quantity id="E7360CD6FFDDFFD976309B080D0ADF16" box="[269,379,1876,1901]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="2" pageNumber="289" unit="cm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 cm</quantity>
. Synflorescence corymbose; capitulum
<quantity id="E7360CD6FFDDFFD974039B080FC8DF16" box="[830,953,1876,1901]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="2" pageNumber="289" unit="mm" value="27.5" valueMax="30.0" valueMin="25.0">25–30 mm</quantity>
long, ca. 160-flowered; peduncle
<quantity id="E7360CD6FFDDFFD972029B0809DDDF16" box="[1343,1452,1876,1901]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="2" pageNumber="289" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">5–10 mm</quantity>
long; involucre globose, arachnoid,
<quantity id="E7360CD6FFDDFFD9751A9B240ED2DFEA" box="[551,675,1912,1937]" metricMagnitude="-2" metricUnit="m" metricValue="4.55" metricValueMax="4.8" metricValueMin="4.3" pageId="2" pageNumber="289" unit="mm" value="45.5" valueMax="48.0" valueMin="43.0">43–48 mm</quantity>
in diam. (including spines), up to
<quantity id="E7360CD6FFDDFFD973049B2408FADFEA" box="[1081,1163,1912,1937]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="289" unit="mm" value="30.0">30 mm</quantity>
long; phyllaries ca. 100, coriaceous, appendiculate; appendages triangular; outer ones spreading; middle ones imbricate, with one spine on each side; inner ones longer, spineless; inner and innermost ones exserted; exappendiculate; receptacle bristles scabrous; corolla purple,
<quantity id="E7360CD6FFDDFFD9760F9BB80DDADF86" box="[306,427,2020,2045]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="2.4" metricValueMin="2.3" pageId="2" pageNumber="289" unit="mm" value="23.5" valueMax="24.0" valueMin="23.0">23–24 mm</quantity>
long, tube equaling the limb; anther tube concolored, glabrous. Achene unknown.
</paragraph>
</subSubSection>
<caption id="74B1F1BBFFDCFFD877B598CE08DADCDC" ID-DOI="http://doi.org/10.5281/zenodo.13717726" ID-Zenodo-Dep="13717726" box="[136,1195,1170,1191]" httpUri="https://zenodo.org/record/13717726/files/figure.png" pageId="3" pageNumber="290" startId="3.[136,229,1170,1191]" targetBox="[199,1393,193,1145]" targetPageId="3" targetType="figure">
<paragraph id="2071A133FFDCFFD877B598CE08DADCDC" blockId="3.[136,1195,1170,1191]" box="[136,1195,1170,1191]" pageId="3" pageNumber="290">
<emphasis id="12BA7D21FFDCFFD877B598CE0C8FDCDC" bold="true" box="[136,254,1170,1191]" pageId="3" pageNumber="290">FIGURE 3.</emphasis>
Distribution map of the species:
<taxonomicName id="E7CEDAB0FFDCFFD8750198CE0E90DCDC" authorityName="Attar &amp; Amini Rad" authorityYear="2019" box="[572,737,1170,1191]" class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="sharifii">Cousinia sharifii</taxonomicName>
;
<taxonomicName id="E7CEDAB0FFDCFFD875D198CE0F0CDCDC" box="[748,893,1170,1191]" class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="eriocephala">C. eriocephala</taxonomicName>
;
<taxonomicName id="E7CEDAB0FFDCFFD874B498CE0845DCDC" authorityName="Rechinger" authorityYear="1972" box="[905,1076,1170,1191]" class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="qaradaghensis">C. qaradaghensis</taxonomicName>
;
<taxonomicName id="E7CEDAB0FFDCFFD8737D98CE08D6DCDC" box="[1088,1191,1170,1191]" class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="zagrica">C. zagrica</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="68D4F2B8FFDCFFD8778098A70956DD43" pageId="3" pageNumber="290" type="etymology">
<paragraph id="2071A133FFDCFFD8778098A70956DD43" blockId="3.[136,1452,1275,1912]" pageId="3" pageNumber="290">
<emphasis id="12BA7D21FFDCFFD8778098A70D39DD6F" bold="true" box="[189,328,1275,1300]" pageId="3" pageNumber="290">Etymology:</emphasis>
—The epithet ‘sharifii’ honours Prof. Dr. Ghavamoddin Sharif, a pioneering Iranian mycologist who introduced some fungi and plants to the world and collected a large amount of plants and fungi from
<collectingCountry id="58D9E1A3FFDCFFD873CF99430952DD43" box="[1266,1315,1311,1336]" name="Iran" pageId="3" pageNumber="290">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="68D4F2B8FFDCFFD87780991F0EA5DF03" pageId="3" pageNumber="290" type="distribution">
<paragraph id="2071A133FFDCFFD87780991F0E94DDDF" blockId="3.[136,1452,1275,1912]" pageId="3" pageNumber="290">
<emphasis id="12BA7D21FFDCFFD87780991F0EF0DD27" bold="true" box="[189,641,1347,1372]" pageId="3" pageNumber="290">Distribution, habitat and phenology:</emphasis>
—
<taxonomicName id="E7CEDAB0FFDCFFD875A0991F0F2FDD27" authorityName="Attar &amp; Amini Rad" authorityYear="2019" box="[669,862,1347,1372]" class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="sharifii">
<emphasis id="12BA7D21FFDCFFD875A0991F0F2FDD27" box="[669,862,1347,1372]" italics="true" pageId="3" pageNumber="290">Cousinia sharifii</emphasis>
</taxonomicName>
is known from the
<typeStatus id="FF751F91FFDCFFD8736D991F08F1DD27" box="[1104,1152,1347,1372]" pageId="3" pageNumber="290">type</typeStatus>
locality in W Azarbaijan province of
<collectingCountry id="58D9E1A3FFDCFFD8762D993B0D30DDFB" box="[272,321,1383,1408]" name="Iran" pageId="3" pageNumber="290">Iran</collectingCountry>
. It was found in mountainous steppes at
<quantity id="E7360CD6FFDCFFD87436993B0F10DDFB" box="[779,865,1383,1408]" metricMagnitude="3" metricUnit="m" metricValue="1.89" pageId="3" pageNumber="290" unit="m" value="1890.0">1890 m</quantity>
a.s.l. The new species is a narrow endemic (
<figureCitation id="B8F5BDB6FFDCFFD87265993B09ECDDFB" box="[1368,1437,1383,1408]" captionStart="FIGURE 3" captionStartId="3.[136,229,1170,1191]" captionTargetBox="[199,1393,193,1145]" captionTargetId="figure-14@3.[194,1394,190,1145]" captionTargetPageId="3" captionText="FIGURE 3. Distribution map of the species: Cousinia sharifii; C. eriocephala; C. qaradaghensis; C. zagrica." figureDoi="http://doi.org/10.5281/zenodo.13717726" httpUri="https://zenodo.org/record/13717726/files/figure.png" pageId="3" pageNumber="290">Fig. 3</figureCitation>
). Flowers appear in July and mature achenes in August.
</paragraph>
<paragraph id="2071A133FFDCFFD8778099F30E70DE07" blockId="3.[136,1452,1275,1912]" pageId="3" pageNumber="290">
<emphasis id="12BA7D21FFDCFFD8778099F30D18DDB3" bold="true" box="[189,361,1455,1480]" pageId="3" pageNumber="290">Relationships:</emphasis>
—Taxa of
<taxonomicName id="E7CEDAB0FFDCFFD876DB99F30F52DDB3" authorityName="Bunge" authorityYear="1865" box="[486,803,1455,1480]" class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="section" section="Cynaroideae">
<emphasis id="12BA7D21FFDCFFD876DB99F30E3ADDB3" box="[486,587,1455,1480]" italics="true" pageId="3" pageNumber="290">Cousinia</emphasis>
sect.
<emphasis id="12BA7D21FFDCFFD875AE99F30F52DDB3" box="[659,803,1455,1480]" italics="true" pageId="3" pageNumber="290">Cynaroideae</emphasis>
</taxonomicName>
that are distributed in the northwest of
<collectingCountry id="58D9E1A3FFDCFFD873CB99F30952DDB3" box="[1270,1315,1455,1480]" name="Iran" pageId="3" pageNumber="290">Iran</collectingCountry>
include:
<taxonomicName id="E7CEDAB0FFDCFFD872AF99F30D22DD97" authority="C. A. Mey." authorityName="C. A. Mey." class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="grandis">
<emphasis id="12BA7D21FFDCFFD872AF99F30CAFDD97" italics="true" pageId="3" pageNumber="290">C. grandis</emphasis>
C.A.Mey.
</taxonomicName>
in
<bibRefCitation id="445FDCC2FFDCFFD87645998F0E1ADD97" author="Candolle, A. P. de" box="[376,619,1491,1516]" pageId="3" pageNumber="290" refId="ref1898" refString="Candolle, A. P. de (1838) Prodromus systematicis naturalis regni vegetabilis, vol. 6. Treuttel &amp; Wurtz, Paris, 687 pp." type="book" year="1838">Candolle (1838: 557)</bibRefCitation>
,
<taxonomicName id="E7CEDAB0FFDCFFD8754A998F0F83DD97" authority="Rechinger (1972: 38)" authorityName="Rechinger" authorityPageNumber="38" authorityYear="1972" box="[631,1010,1491,1516]" class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="bobeckii">
<emphasis id="12BA7D21FFDCFFD8754A998F0E86DD97" box="[631,759,1491,1516]" italics="true" pageId="3" pageNumber="290">C. bobeckii</emphasis>
<bibRefCitation id="445FDCC2FFDCFFD875C2998F0F83DD97" author="Rechinger, K. H." box="[767,1010,1491,1516]" pageId="3" pageNumber="290" pagination="1 - 329" refId="ref2239" refString="Rechinger, K. H. (1972) Compositae - Cynareae I: Cousinia. In: Rechinger, K. H. (Ed.) Flora Iranica, vol. 90. Akademische Druck- und Verlagsanstalt, Graz, pp. 1 - 329." type="book chapter" year="1972">Rechinger (1972: 38)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="E7CEDAB0FFDCFFD874C3998F0D12DE6B" authority="(Bieberstein 1798: 118) Meyer (1831: 230)" authorityName="Meyer" authorityPageNumber="230" authorityYear="1831" baseAuthorityName="Bieberstein" baseAuthorityPageNumber="118" baseAuthorityYear="1798" class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="cynaroides">
<emphasis id="12BA7D21FFDCFFD874C3998F08EBDD97" box="[1022,1178,1491,1516]" italics="true" pageId="3" pageNumber="290">C. cynaroides</emphasis>
(
<bibRefCitation id="445FDCC2FFDCFFD87394998F09D2DD97" author="Marschall von Bieberstein, F." box="[1193,1443,1491,1516]" pageId="3" pageNumber="290" refId="ref2068" refString="Marschall von Bieberstein, F. (1798) Tableau des provinces situees sur la cote occidentale de la mer Caspienne entre les fleuves Terek et Kour. St. Petersbourg, 120 pp." type="book" year="1798">Bieberstein 1798: 118</bibRefCitation>
)
<bibRefCitation id="445FDCC2FFDCFFD877B599AB0D12DE6B" author="Meyer, C. A." box="[136,355,1527,1552]" pageId="3" pageNumber="290" refId="ref2149" refString="Meyer, C. A. (1831) Verzeichniss der Pflanzen, welche wahrend der, auf allerhochsten Befehl, in den Jahren 1829 und 1830 unternommenen Reise in Caucasus und in den Provinzen am westlichen Ufer des Caspischen Meeres gefunden und eingesammelt worden sind. Kaiserliche Akademie der Wissenschaften, St. Petersburg." type="book" year="1831">Meyer (1831: 230)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="E7CEDAB0FFDCFFD8764D99AB0F7FDE6B" authority="Rechinger (1964: 344)" authorityName="Rechinger" authorityPageNumber="344" authorityYear="1964" box="[368,782,1527,1552]" class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="algurdina">
<emphasis id="12BA7D21FFDCFFD8764D99AB0E71DE6B" box="[368,512,1527,1552]" italics="true" pageId="3" pageNumber="290">C. algurdina</emphasis>
<bibRefCitation id="445FDCC2FFDCFFD8753799AB0F7FDE6B" author="Rechinger, K. H." box="[522,782,1527,1552]" pageId="3" pageNumber="290" pagination="344 - 354" refId="ref2205" refString="Rechinger, K. H. (1964) Zehn neue Cousinia - Arten aus Afghanistan und Kurdestan. Anzeiger der Osterreichischen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse 101: 344 - 354." type="journal article" year="1964">Rechinger (1964: 344)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="E7CEDAB0FFDCFFD8742199AB08EEDE6B" authority="Rechinger (1972: 244)" authorityName="Rechinger" authorityPageNumber="244" authorityYear="1972" box="[796,1183,1527,1552]" class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="gilliatii">
<emphasis id="12BA7D21FFDCFFD8742199AB0FE3DE6B" box="[796,914,1527,1552]" italics="true" pageId="3" pageNumber="290">C. gilliatii</emphasis>
<bibRefCitation id="445FDCC2FFDCFFD874A699AB08EEDE6B" author="Rechinger, K. H." box="[923,1183,1527,1552]" pageId="3" pageNumber="290" pagination="1 - 329" refId="ref2239" refString="Rechinger, K. H. (1972) Compositae - Cynareae I: Cousinia. In: Rechinger, K. H. (Ed.) Flora Iranica, vol. 90. Akademische Druck- und Verlagsanstalt, Graz, pp. 1 - 329." type="book chapter" year="1972">Rechinger (1972: 244)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="E7CEDAB0FFDCFFD8739099AB0D7ADE4F" authority="Attar &amp; Ghahreman" authorityName="Attar &amp; Ghahreman" class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="shebliensis">
<emphasis id="12BA7D21FFDCFFD8739099AB093ADE6B" box="[1197,1355,1527,1552]" italics="true" pageId="3" pageNumber="290">C. shebliensis</emphasis>
Attar &amp; Ghahreman
</taxonomicName>
in
<bibRefCitation id="445FDCC2FFDCFFD876099A470EF1DE4F" author="Ghahreman, A. &amp; Iranshahr, M. &amp; Attar, F." box="[308,640,1563,1588]" pageId="3" pageNumber="290" pagination="15 - 22" refId="ref1979" refString="Ghahreman, A., Iranshahr, M. &amp; Attar, F. (1999) Introduction two new and a rare species of the genus Cousinia Cass., sect. Cynaroideae (Asteraceae). Iranian Journal of Botany 8: 15 - 22. https: // doi. org / 10.1007 / BF 00936283" type="journal article" year="1999">
Ghahreman
<emphasis id="12BA7D21FFDCFFD876FC9A470D8DDE4F" box="[449,508,1563,1588]" italics="true" pageId="3" pageNumber="290">et al.</emphasis>
(1999: 19)
</bibRefCitation>
,
<taxonomicName id="E7CEDAB0FFDCFFD875B39A470875DE4F" authority="Attar et al. (2002: 5)" authorityName="Attar" authorityPageNumber="5" authorityYear="2002" box="[654,1028,1563,1588]" class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="zagrica">
<emphasis id="12BA7D21FFDCFFD875B39A470F77DE4F" box="[654,774,1563,1588]" italics="true" pageId="3" pageNumber="290">C. zagrica</emphasis>
<bibRefCitation id="445FDCC2FFDCFFD8742D9A470875DE4F" author="Attar, F. &amp; Ghahreman, A. &amp; Assadi, M." box="[784,1028,1563,1588]" pageId="3" pageNumber="290" pagination="5 - 7" refId="ref1797" refString="Attar, F. &amp; Ghahreman, A. &amp; Assadi, M. (2002) Cousinia zagrica (Asteraceae), a new species from West of Iran. Sendtnera 8: 5 - 7." type="journal article" year="2002">
Attar
<emphasis id="12BA7D21FFDCFFD8746E9A470FFFDE4F" box="[851,910,1563,1588]" italics="true" pageId="3" pageNumber="290">et al.</emphasis>
(2002: 5)
</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="E7CEDAB0FFDCFFD8732E9A470CCFDE23" authority="Rechinger (1972: 258)" authorityName="Rechinger" authorityPageNumber="258" authorityYear="1972" class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="qaradaghensis">
<emphasis id="12BA7D21FFDCFFD8732E9A4708ADDE4F" box="[1043,1244,1563,1588]" italics="true" pageId="3" pageNumber="290">C. qaradaghensis</emphasis>
<bibRefCitation id="445FDCC2FFDCFFD873D89A470CCFDE23" author="Rechinger, K. H." pageId="3" pageNumber="290" pagination="1 - 329" refId="ref2239" refString="Rechinger, K. H. (1972) Compositae - Cynareae I: Cousinia. In: Rechinger, K. H. (Ed.) Flora Iranica, vol. 90. Akademische Druck- und Verlagsanstalt, Graz, pp. 1 - 329." type="book chapter" year="1972">Rechinger (1972: 258)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="E7CEDAB0FFDCFFD877F79A630ED8DE23" authority="Attar &amp; Djavadi (2010: 161)" authorityName="Attar &amp; Djavadi" authorityPageNumber="161" authorityYear="2010" box="[202,681,1599,1624]" class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="sahandica">
<emphasis id="12BA7D21FFDCFFD877F79A630D2EDE23" box="[202,351,1599,1624]" italics="true" pageId="3" pageNumber="290">C. sahandica</emphasis>
<bibRefCitation id="445FDCC2FFDCFFD876589A630ED8DE23" author="Attar, F. &amp; Djavadi, S. B." box="[357,681,1599,1624]" pageId="3" pageNumber="290" pagination="130 - 184" refId="ref1650" refString="Attar, F. &amp; Djavadi, S. B. (2010) A taxonomic revision of Cousinia sect. Cynaroideae (Asteraceae, Cardueae) in the Flora of Iran. Iranian Journal of Botany 16: 130 - 184." type="journal article" year="2010">Attar &amp; Djavadi (2010: 161)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="E7CEDAB0FFDCFFD875DC9A6308FCDE23" authority="Attar et al. (2000: 699)" authorityName="Attar" authorityPageNumber="699" authorityYear="2000" box="[737,1165,1599,1624]" class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="sabalanica">
<emphasis id="12BA7D21FFDCFFD875DC9A630F0CDE23" box="[737,893,1599,1624]" italics="true" pageId="3" pageNumber="290">C. sabalanica</emphasis>
<bibRefCitation id="445FDCC2FFDCFFD874B89A6308FCDE23" author="Attar, F. &amp; Ghahreman, A. &amp; Assadi, M." box="[901,1165,1599,1624]" pageId="3" pageNumber="290" pagination="697 - 700" refId="ref1734" refString="Attar, F. &amp; Ghahreman, A. &amp; Assadi, M. (2000) Three new species of the genus Cousinia (Asteraceae, Sect. Cynaroideae) from Iran. Nordic Journal of Botany 20: 697 - 700. https: // doi. org / 10.1111 / j. 1756 - 1051.2000. tb 00755. x" type="journal article" year="2000">
Attar
<emphasis id="12BA7D21FFDCFFD874FB9A630F8EDE23" box="[966,1023,1599,1624]" italics="true" pageId="3" pageNumber="290">et al.</emphasis>
(2000: 699)
</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="E7CEDAB0FFDCFFD873A79A630C81DE07" authority="Boiss. &amp; Hausskn." authorityName="Boiss. &amp; Hausskn." class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="eriocephala">
<emphasis id="12BA7D21FFDCFFD873A79A630930DE23" box="[1178,1345,1599,1624]" italics="true" pageId="3" pageNumber="290">C. eriocephala</emphasis>
Boiss. &amp; Hausskn.
</taxonomicName>
in
<bibRefCitation id="445FDCC2FFDCFFD8762E9A3F0D8CDE07" author="Boissier, E. P." box="[275,509,1635,1660]" pageId="3" pageNumber="290" refId="ref1835" refString="Boissier, E. P. (1875) Flora orientalis, vol. 3. H. Georg, Geneva &amp; Basel, 1033 pp." type="book" year="1875">Boissier (1875: 504)</bibRefCitation>
.
</paragraph>
<paragraph id="2071A133FFDCFFD877809ADB0CD7DF77" blockId="3.[136,1452,1275,1912]" pageId="3" pageNumber="290">
<taxonomicName id="E7CEDAB0FFDCFFD877809ADB0D0DDEDB" authorityName="Attar &amp; Amini Rad" authorityYear="2019" box="[189,380,1671,1696]" class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="sharifii">
<emphasis id="12BA7D21FFDCFFD877809ADB0D0DDEDB" box="[189,380,1671,1696]" italics="true" pageId="3" pageNumber="290">Cousinia sharifii</emphasis>
</taxonomicName>
is morphologically similar to
<taxonomicName id="E7CEDAB0FFDCFFD875E99ADB0F0DDEDB" box="[724,892,1671,1696]" class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="eriocephala">
<emphasis id="12BA7D21FFDCFFD875E99ADB0F0DDEDB" box="[724,892,1671,1696]" italics="true" pageId="3" pageNumber="290">C. eriocephala</emphasis>
</taxonomicName>
and
<taxonomicName id="E7CEDAB0FFDCFFD874899ADB085BDEDB" box="[948,1066,1671,1696]" class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="zagrica">
<emphasis id="12BA7D21FFDCFFD874899ADB085BDEDB" box="[948,1066,1671,1696]" italics="true" pageId="3" pageNumber="290">C. zagrica</emphasis>
</taxonomicName>
, two rare species of NW
<collectingCountry id="58D9E1A3FFDCFFD872739ADB090ADEDB" box="[1358,1403,1671,1696]" name="Iran" pageId="3" pageNumber="290">Iran</collectingCountry>
and E
<collectingCountry id="58D9E1A3FFDCFFD8779D9AF70C80DEBF" box="[160,241,1707,1732]" name="Turkey" pageId="3" pageNumber="290">Turkey</collectingCountry>
, in its habit, densely canescent-arachnoid involucre and globose capitulum. It can be distinguished from the two species by the shape of involucre and phyllaries, and by the number of flowers and phyllaries (
<tableCitation id="6D4C9488FFDCFFD873E49A93095EDE93" box="[1241,1327,1743,1768]" captionStart="TABLE 1" captionStartId="7.[136,224,161,184]" captionText="TABLE 1. Comparisons of morphological characters of Cousinia sharifii and its relatives." pageId="3" pageNumber="290">Table 1</tableCitation>
;
<figureCitation id="B8F5BDB6FFDCFFD872009A9309F8DE93" box="[1341,1417,1743,1768]" captionStart="FIGURE 1" captionStartId="1.[136,229,2029,2050]" captionTargetBox="[189,1398,187,2001]" captionTargetId="figure-14@1.[189,1398,187,2001]" captionTargetPageId="1" captionText="FIGURE 1. Holotype of Cousinia sharifii." figureDoi="http://doi.org/10.5281/zenodo.13717720" httpUri="https://zenodo.org/record/13717720/files/figure.png" pageId="3" pageNumber="290">Figs. 1</figureCitation>
,
<figureCitation id="B8F5BDB6FFDCFFD872AB9A9309D7DE93" box="[1430,1446,1743,1768]" captionStart="FIGURE 4" captionStartId="4.[136,229,2029,2050]" captionTargetBox="[154,1434,190,2004]" captionTargetId="figure-17@4.[154,1434,190,2004]" captionTargetPageId="4" captionText="FIGURE 4. Cousinia eriocephala (Typus)." figureDoi="http://doi.org/10.5281/zenodo.13717728" httpUri="https://zenodo.org/record/13717728/files/figure.png" pageId="3" pageNumber="290">4</figureCitation>
,
<figureCitation id="B8F5BDB6FFDCFFD877B59AAF0CE9DF77" box="[136,152,1779,1804]" captionStart="FIGURE 5" captionStartId="5.[136,229,2029,2050]" captionTargetBox="[194,1393,190,2004]" captionTargetId="figure-14@5.[194,1393,190,2004]" captionTargetPageId="5" captionText="FIGURE 5. Cousinia zagrica (Typus)." figureDoi="http://doi.org/10.5281/zenodo.13717730" httpUri="https://zenodo.org/record/13717730/files/figure.png" pageId="3" pageNumber="290">5</figureCitation>
).
</paragraph>
<paragraph id="2071A133FFDCFFD877809B4B0EA5DF03" blockId="3.[136,1452,1275,1912]" pageId="3" pageNumber="290">
The new species is also related to
<taxonomicName id="E7CEDAB0FFDCFFD875059B4B0EBADF4B" authorityName="Candolle" authorityYear="1838" box="[568,715,1815,1840]" class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="canescens">
<emphasis id="12BA7D21FFDCFFD875059B4B0EBADF4B" box="[568,715,1815,1840]" italics="true" pageId="3" pageNumber="290">C. canescens</emphasis>
</taxonomicName>
and
<taxonomicName id="E7CEDAB0FFDCFFD8743D9B4B0FB4DF4B" authorityName="Rechinger" authorityYear="1972" box="[768,965,1815,1840]" class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="qaradaghensis">
<emphasis id="12BA7D21FFDCFFD8743D9B4B0FB4DF4B" box="[768,965,1815,1840]" italics="true" pageId="3" pageNumber="290">C. qaradaghensis</emphasis>
</taxonomicName>
because of the arachnoid-greyish habit, but differs from these species by some characters (
<tableCitation id="6D4C9488FFDCFFD875B39B670E92DF2F" box="[654,739,1851,1876]" captionStart="TABLE 1" captionStartId="7.[136,224,161,184]" captionText="TABLE 1. Comparisons of morphological characters of Cousinia sharifii and its relatives." pageId="3" pageNumber="290">Table 1</tableCitation>
;
<figureCitation id="B8F5BDB6FFDCFFD875CC9B670F4DDF2F" box="[753,828,1851,1876]" captionStart="FIGURE 1" captionStartId="1.[136,229,2029,2050]" captionTargetBox="[189,1398,187,2001]" captionTargetId="figure-14@1.[189,1398,187,2001]" captionTargetPageId="1" captionText="FIGURE 1. Holotype of Cousinia sharifii." figureDoi="http://doi.org/10.5281/zenodo.13717720" httpUri="https://zenodo.org/record/13717720/files/figure.png" pageId="3" pageNumber="290">Figs. 1</figureCitation>
&amp;
<figureCitation id="B8F5BDB6FFDCFFD874629B670F1EDF2F" box="[863,879,1851,1876]" captionStart="FIGURE 6" captionStartId="6.[136,229,2029,2050]" captionTargetBox="[187,1400,190,2004]" captionTargetId="figure-17@6.[187,1401,190,2004]" captionTargetPageId="6" captionText="FIGURE 6. Cousinia qaradaghensis (Typus)." figureDoi="http://doi.org/10.5281/zenodo.13717732" httpUri="https://zenodo.org/record/13717732/files/figure.png" pageId="3" pageNumber="290">6</figureCitation>
).
<taxonomicName id="E7CEDAB0FFDCFFD874BE9B670810DF2F" authorityName="Candolle" authorityYear="1838" box="[899,1121,1851,1876]" class="Magnoliopsida" family="Asteraceae" genus="Cousinia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="290" phylum="Tracheophyta" rank="species" species="canescens">
<emphasis id="12BA7D21FFDCFFD874BE9B670810DF2F" box="[899,1121,1851,1876]" italics="true" pageId="3" pageNumber="290">Cousinia canescens</emphasis>
</taxonomicName>
is endemic to
<collectingCountry id="58D9E1A3FFDCFFD8723C9B670920DF2F" box="[1281,1361,1851,1876]" name="Turkey" pageId="3" pageNumber="290">Turkey</collectingCountry>
and has not been reported from
<collectingCountry id="58D9E1A3FFDCFFD876AD9B030DCCDF03" box="[400,445,1887,1912]" name="Iran" pageId="3" pageNumber="290">Iran</collectingCountry>
(
<bibRefCitation id="445FDCC2FFDCFFD876F19B030EB9DF03" author="Attar, F. &amp; Djavadi, S. B." box="[460,712,1887,1912]" pageId="3" pageNumber="290" pagination="130 - 184" refId="ref1650" refString="Attar, F. &amp; Djavadi, S. B. (2010) A taxonomic revision of Cousinia sect. Cynaroideae (Asteraceae, Cardueae) in the Flora of Iran. Iranian Journal of Botany 16: 130 - 184." type="journal article" year="2010">Attar &amp; Djavadi 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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