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<document id="B28394F4C882214216657CA1037DB754" ID-DOI="10.11646/phytotaxa.394.3.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13717607" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725651963364" checkinUser="felipe" docAuthor="Shalisko, Viacheslav, Sundue, Michael A., Villalobos-Arámbula, Alma Rosa, Muñiz-Castro, Miguel Ángel &amp; Vázquez-García, J. Antonio" docDate="2019" docId="03DB793CFFC82C54FF5897D544B36F44" docLanguage="en" docName="phytotaxa.394.3.1.pdf" docOrigin="Phytotaxa 394 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.394.3.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Parrisia Shalisko &amp; Sundue 2019, gen. nov." docType="treatment" docVersion="2" lastPageNumber="188" masterDocId="FFE20144FFC32C58FFD0920146466926" masterDocTitle="Taxonomic novelties in grammitid ferns (Polypodiaceae) from the Neotropics and Madagascar supported by molecular data" masterLastPageNumber="208" masterPageNumber="176" pageNumber="187" updateTime="1725660222309" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="5DA6F0171EAF6385A009A9A09CD342E0">Taxonomic novelties in grammitid ferns (Polypodiaceae) from the Neotropics and Madagascar supported by molecular data</mods:title>
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<mods:namePart id="8C671846CDC402883349770D6E2C613A">Shalisko, Viacheslav</mods:namePart>
<mods:affiliation id="48C7C50109E4E06178D5C82CED85FD81">Laboratorio de Ecosistemática, Instituto de Botánica, Departamento de Botánica y Zoología, Centro Universitario de Ciencias Biológicas y Agropecuarias, Universidad de Guadalajara, Camino Ing. Ramón Padilla Sánchez No. 2100, La Venta del Astillero, Zapopan, 45200, Jalisco, Mexico</mods:affiliation>
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<mods:namePart id="CC97D8BF4F34A576E9E1FE3877A819FB">Sundue, Michael A.</mods:namePart>
<mods:affiliation id="BCBCE50DE326B8F9D0D20D2CA7481996">Pringle Herbarium, Dept. of Plant Biology, University of Vermont, Burlington, VT, 50405 USA</mods:affiliation>
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<mods:namePart id="715D09506C5A17785DCCCB1B51B590C3">Villalobos-Arámbula, Alma Rosa</mods:namePart>
<mods:affiliation id="D012C5B834477922579F7423D9CC69D0">Laboratorio de Genética, Ecosistemática Molecular y Funcional, Departamento de Biología Celular y Molecular, Centro Universitario de Ciencias Biológicas y Agropecuarias, Universidad de Guadalajara, Camino Ing. Ramón Padilla Sánchez No. 2100, La Venta del Astillero, Zapopan, 45200, Jalisco, Mexico</mods:affiliation>
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<mods:namePart id="568D21F5CDC2F875A6C13392E9CB7F68">Muñiz-Castro, Miguel Ángel</mods:namePart>
</mods:name>
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<mods:namePart id="A9CFC750E4DC46ED37A0FD680FFF948A">Vázquez-García, J. Antonio</mods:namePart>
<mods:affiliation id="8D6BF72C06530D8294BC9B923B360F11">Laboratorio de Ecosistemática, Instituto de Botánica, Departamento de Botánica y Zoología, Centro Universitario de Ciencias Biológicas y Agropecuarias, Universidad de Guadalajara, Camino Ing. Ramón Padilla Sánchez No. 2100, La Venta del Astillero, Zapopan, 45200, Jalisco, Mexico</mods:affiliation>
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<mods:title id="B92E5D2865DFBC9123F8B3A9CBB0BF2E">Phytotaxa</mods:title>
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<mods:date id="72C0F8362AB17BE114708E10A678B345">2019</mods:date>
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<mods:number id="C55F2AEF9CE7BB7FE23FAF5D58A1979E">394</mods:number>
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<treatment id="03DB793CFFC82C54FF5897D544B36F44" LSID="urn:lsid:plazi:treatment:03DB793CFFC82C54FF5897D544B36F44" httpUri="http://treatment.plazi.org/id/03DB793CFFC82C54FF5897D544B36F44" lastPageId="12" lastPageNumber="188" pageId="11" pageNumber="187">
<subSubSection id="C3689BA1FFC82C53FF5897D544D46CC9" box="[136,658,1492,1519]" pageId="11" pageNumber="187" type="nomenclature">
<paragraph id="8BCDC82AFFC82C53FF5897D544D46CC9" blockId="11.[136,658,1492,1519]" box="[136,658,1492,1519]" pageId="11" pageNumber="187">
<heading id="D0857F46FFC82C53FF5897D544D46CC9" box="[136,658,1492,1519]" fontSize="11" level="2" pageId="11" pageNumber="187" reason="3">
<taxonomicName id="4C72B3A9FFC82C53FF5897D5478E6CC9" ID-CoL="6J82" authority="Shalisko &amp; Sundue" authorityName="Shalisko &amp; Sundue" authorityYear="2019" box="[136,456,1492,1519]" class="Polypodiopsida" family="Polypodiaceae" genus="Parrisia" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="187" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<emphasis id="B9061438FFC82C53FF5897D546A36CC8" bold="true" box="[136,229,1492,1518]" italics="true" pageId="11" pageNumber="187">Parrisia</emphasis>
Shalisko &amp; Sundue
</taxonomicName>
,
<emphasis id="B9061438FFC82C53FE0397D744716CC9" box="[467,567,1494,1519]" italics="true" pageId="11" pageNumber="187">
<taxonomicNameLabel id="A235A943FFC82C53FE0397D744716CC9" box="[467,567,1494,1519]" pageId="11" pageNumber="187" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="1349D4AFFFC82C53FD9697D744CF6CC9" box="[582,649,1494,1519]" captionStart="FIGURE 2" captionStartId="12.[136,229,898,919]" captionTargetBox="[151,1436,281,873]" captionTargetId="figure-64@12.[151,1436,281,873]" captionTargetPageId="12" captionText="FIGURE 2. Parrisia parietina (Klotzsch) Shalisko &amp; Sundue. A–B. Habit (Sundue et al. 3233). Photos by M. Sundue." figureDoi="http://doi.org/10.5281/zenodo.13717612" httpUri="https://zenodo.org/record/13717612/files/figure.png" pageId="11" pageNumber="187">Fig. 2</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3689BA1FFC82C53FF589421440D6F5B" pageId="11" pageNumber="187" type="type_taxon">
<paragraph id="8BCDC82AFFC82C53FF589421422F6F13" blockId="11.[136,1452,1568,1661]" box="[136,1129,1568,1589]" pageId="11" pageNumber="187">
<emphasis id="B9061438FFC82C53FF58942147496F13" bold="true" box="[136,271,1568,1589]" pageId="11" pageNumber="187">
<typeStatus id="54C97688FFC82C53FF58942146FC6F13" box="[136,186,1568,1589]" pageId="11" pageNumber="187">Type</typeStatus>
species:
</emphasis>
—
<taxonomicName id="4C72B3A9FFC82C53FEF7942145466F13" ID-CoL="4DT67" authority="(Klotzsch) Shalisko &amp; Sundue" authorityName="Shalisko &amp; Sundue. A" authorityYear="2019" baseAuthorityName="Shalisko &amp; Sundue. A" baseAuthorityYear="1847" box="[295,768,1568,1589]" class="Polypodiopsida" family="Polypodiaceae" genus="Parrisia" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="187" phylum="Tracheophyta" rank="species" species="parietina">
<emphasis id="B9061438FFC82C53FEF7942147936F13" box="[295,469,1568,1589]" italics="true" pageId="11" pageNumber="187">Parrisia parietina</emphasis>
(Klotzsch) Shalisko &amp; Sundue
</taxonomicName>
(=
<taxonomicName id="4C72B3A9FFC82C53FCCB942142186F13" ID-CoL="7878J" authority="Klotzsch" authorityName="Klotzsch" box="[795,1118,1568,1589]" class="Polypodiopsida" family="Polypodiaceae" genus="Polypodium" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="187" phylum="Tracheophyta" rank="species" species="parietinum">
<emphasis id="B9061438FFC82C53FCCB942145B86F13" box="[795,1022,1568,1589]" italics="true" pageId="11" pageNumber="187">Polypodium parietinum</emphasis>
Klotzsch
</taxonomicName>
).
</paragraph>
<paragraph id="8BCDC82AFFC82C53FF589445440D6F5B" blockId="11.[136,1452,1568,1661]" pageId="11" pageNumber="187">
<emphasis id="B9061438FFC82C53FF58944546BC6F7F" bold="true" box="[136,250,1604,1625]" pageId="11" pageNumber="187">Basyonym:</emphasis>
—
<taxonomicName id="4C72B3A9FFC82C53FEC39445441E6F7F" ID-CoL="7878J" authority="Klotzsch." authorityName="Klotzsch." box="[275,600,1604,1625]" class="Polypodiopsida" family="Polypodiaceae" genus="Polypodium" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="187" phylum="Tracheophyta" rank="species" species="parietinum">
<emphasis id="B9061438FFC82C53FEC3944547B06F7F" box="[275,502,1604,1625]" italics="true" pageId="11" pageNumber="187">Polypodium parietinum</emphasis>
Klotzsch.
</taxonomicName>
<collectingCountry id="F36588BAFFC82C53FD8D944544A96F7F" box="[605,751,1604,1625]" name="Venezuela" pageId="11" pageNumber="187">VENEZUELA</collectingCountry>
.
<collectingRegion id="49B606C8FFC82C53FD26944545066F7F" box="[758,832,1604,1625]" country="Venezuela" name="Aragua" pageId="11" pageNumber="187">Aragua</collectingRegion>
: ad saxor. parietes humidas Colonia Tovar,
<emphasis id="B9061438FFC82C53FB309445430F6F7F" box="[1248,1353,1604,1625]" italics="true" pageId="11" pageNumber="187">Moritz 253</emphasis>
(
<typeStatus id="54C97688FFC82C53FA86944543EA6F7F" box="[1366,1452,1604,1625]" pageId="11" pageNumber="187" type="holotype">holotype</typeStatus>
B!, fragment NY!,
<typeStatus id="54C97688FFC82C53FEA5946947826F5B" box="[373,452,1640,1661]" pageId="11" pageNumber="187" type="isotype">isotypes</typeStatus>
BM!, K!, P!).
</paragraph>
</subSubSection>
<subSubSection id="C3689BA1FFC82C53FF5894AF42916E15" pageId="11" pageNumber="187" type="diagnosis">
<paragraph id="8BCDC82AFFC82C53FF5894AF42916E15" blockId="11.[136,1452,1710,2059]" pageId="11" pageNumber="187">
<emphasis id="B9061438FFC82C53FF5894AF47436FE1" bold="true" box="[136,261,1710,1735]" pageId="11" pageNumber="187">Diagnosis:</emphasis>
—
<taxonomicName id="4C72B3A9FFC82C53FEF194AF47386FE1" authorityName="Shalisko &amp; Sundue" authorityYear="2019" box="[289,382,1710,1735]" class="Polypodiopsida" family="Polypodiaceae" genus="Parrisia" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="187" phylum="Tracheophyta" rank="genus">
<emphasis id="B9061438FFC82C53FEF194AF47386FE1" box="[289,382,1710,1735]" italics="true" pageId="11" pageNumber="187">Parrisia</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C72B3A9FFC82C53FDCB94AF44DE6FE1" authorityName="sensu Bishop and Smith" authorityYear="1992" box="[539,664,1710,1735]" class="Polypodiopsida" family="Polypodiaceae" genus="Enterosora" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="187" phylum="Tracheophyta" rank="genus">
<emphasis id="B9061438FFC82C53FDCB94AF44DE6FE1" box="[539,664,1710,1735]" italics="true" pageId="11" pageNumber="187">Enterosora</emphasis>
</taxonomicName>
, but differs from the latter by radial rhizomes (vs. dorsiventral), the location of sori in the medium of the vein and the presence of setae around the sori. It is distinguished from
<taxonomicName id="4C72B3A9FFC82C53FAE494D343EA6FCD" authorityName="Swartz" authorityYear="1800" box="[1332,1452,1746,1771]" class="Polypodiopsida" family="Polypodiaceae" genus="Grammitis" higherTaxonomySource="GBIF" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="187" phylum="Tracheophyta" rank="genus" sensu="stricto">
<emphasis id="B9061438FFC82C53FAE494D343EA6FCD" box="[1332,1452,1746,1771]" italics="true" pageId="11" pageNumber="187">Grammitis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235A943FFC82C53FF5894F746EA6E29" box="[136,172,1782,1807]" pageId="11" pageNumber="187" sensu="stricto">s.s.</taxonomicNameLabel>
by the absence of a dark-colored sclerotized leaf margin, and by the presence of setae on the blade. It differs from
<taxonomicName id="4C72B3A9FFC82C53FF58951B475B6E15" authorityName="Gaudichaud-Beaupre" authorityYear="1824" box="[136,285,1818,1843]" class="Polypodiopsida" family="Polypodiaceae" genus="Adenophorus" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="187" phylum="Tracheophyta" rank="genus">
<emphasis id="B9061438FFC82C53FF58951B475B6E15" box="[136,285,1818,1843]" italics="true" pageId="11" pageNumber="187">Adenophorus</emphasis>
</taxonomicName>
in lacking the uniseriate reddish glandular paraphyses and hairs, present in the latter.
</paragraph>
</subSubSection>
<subSubSection id="C3689BA1FFC82C54FF6D953F457C6827" lastPageId="12" lastPageNumber="188" pageId="11" pageNumber="187" type="description">
<paragraph id="8BCDC82AFFC82C54FF6D953F457C6827" blockId="11.[136,1452,1710,2059]" lastBlockId="12.[136,1452,160,257]" lastPageId="12" lastPageNumber="188" pageId="11" pageNumber="187">
<emphasis id="B9061438FFC82C53FF6D953F47176E71" bold="true" box="[189,337,1854,1879]" pageId="11" pageNumber="187">Description:</emphasis>
—
<emphasis id="B9061438FFC82C53FEBD953F47F36E71" box="[365,437,1854,1879]" italics="true" pageId="11" pageNumber="187">Plants</emphasis>
epiphytic,
<emphasis id="B9061438FFC82C53FDE4953F44DF6E71" box="[564,665,1854,1879]" italics="true" pageId="11" pageNumber="187">rhizomes</emphasis>
radial, rarely dorsiventral, with golden brown or brown non-clathrate scales.
<emphasis id="B9061438FFC82C53FF0D9563476B6E5D" box="[221,301,1890,1915]" italics="true" pageId="11" pageNumber="187">Fronds</emphasis>
monomorphic, stipes setose, brown or blackish, laminae simple, linear, linear-spathulate or linearelliptic, rounded at the apex, attenuate at the base, sinuate, sometimes lobed up to 1/3 of the distance to midrib, somewhat thick and spongy; fronds covered with septate setae, from castaneous to brown, clustered close to sori, at midrib and on the margin, as well as with simple or branched glandular hairs on both leaf surfaces; hydathodes absent; veins free, from simple to forked, rarely two times forked.
<emphasis id="B9061438FFC82C53FCB795F345D0612D" box="[871,918,2034,2059]" italics="true" pageId="11" pageNumber="187">Sori</emphasis>
round or nearly so, superficial or subimpressed, exindusiate, without waxy or glandular paraphyses, located abaxially on blades in a single row on both sides from the midrib, that start at some distance from blade base, and extend up to leaf apex, sori born medially on the one-time forked vein.
<emphasis id="B9061438FFCF2C54FEC692E947CD6827" box="[278,395,232,257]" italics="true" pageId="12" pageNumber="188">Sporangia</emphasis>
with 13 to 16 thickened annulus cells.
</paragraph>
</subSubSection>
<caption id="DF0D98A2FFCF2C54FF58918343466AB1" ID-DOI="http://doi.org/10.5281/zenodo.13717612" ID-Zenodo-Dep="13717612" box="[136,1280,898,919]" httpUri="https://zenodo.org/record/13717612/files/figure.png" pageId="12" pageNumber="188" startId="12.[136,229,898,919]" targetBox="[151,1436,281,873]" targetPageId="12" targetType="figure">
<paragraph id="8BCDC82AFFCF2C54FF58918343466AB1" blockId="12.[136,1280,898,919]" box="[136,1280,898,919]" pageId="12" pageNumber="188">
<emphasis id="B9061438FFCF2C54FF58918346B86AB1" bold="true" box="[136,254,898,919]" pageId="12" pageNumber="188">FIGURE 2.</emphasis>
<taxonomicName id="4C72B3A9FFCF2C54FED3918344BC6AB1" authority="(Klotzsch) Shalisko &amp; Sundue. A" authorityName="Shalisko &amp; Sundue. A" authorityYear="2019" baseAuthorityName="Shalisko &amp; Sundue. A" baseAuthorityYear="1847" box="[259,762,898,919]" class="Polypodiopsida" family="Polypodiaceae" genus="Parrisia" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="188" phylum="Tracheophyta" rank="species" species="parietina">
<emphasis id="B9061438FFCF2C54FED3918347F76AB1" box="[259,433,898,919]" italics="true" pageId="12" pageNumber="188">Parrisia parietina</emphasis>
(Klotzsch) Shalisko &amp; Sundue. A
</taxonomicName>
–B. Habit (
<emphasis id="B9061438FFCF2C54FCB3918342506AB1" box="[867,1046,898,919]" italics="true" pageId="12" pageNumber="188">Sundue et al. 3233</emphasis>
). Photos by M. Sundue.
</paragraph>
</caption>
<subSubSection id="C3689BA1FFCF2C54FF6D91EA47A16D0E" pageId="12" pageNumber="188" type="etymology">
<paragraph id="8BCDC82AFFCF2C54FF6D91EA47A16D0E" blockId="12.[136,1452,1003,1568]" pageId="12" pageNumber="188">
<emphasis id="B9061438FFCF2C54FF6D91EA470E6D22" bold="true" box="[189,328,1003,1028]" pageId="12" pageNumber="188">Etymology:</emphasis>
—Named after Barbara S. Parris (*1945), honoring her enormous contributions to grammitid ferns, particularly in the paleotropics.
</paragraph>
</subSubSection>
<subSubSection id="C3689BA1FFCF2C54FF6D963244B36F44" pageId="12" pageNumber="188" type="distribution">
<paragraph id="8BCDC82AFFCF2C54FF6D963244AE6D56" blockId="12.[136,1452,1003,1568]" pageId="12" pageNumber="188">
<emphasis id="B9061438FFCF2C54FF6D9632471F6D6A" bold="true" box="[189,345,1075,1100]" pageId="12" pageNumber="188">Distribution:</emphasis>
—
<collectingCountry id="F36588BAFFCF2C54FEA69632478B6D6A" box="[374,461,1075,1100]" name="Mexico" pageId="12" pageNumber="188">Mexico</collectingCountry>
,
<collectingCountry id="F36588BAFFCF2C54FE0A963244106D6A" box="[474,598,1075,1100]" name="Guatemala" pageId="12" pageNumber="188">Guatemala</collectingCountry>
,
<collectingCountry id="F36588BAFFCF2C54FDB3963244A76D6A" box="[611,737,1075,1100]" name="Costa Rica" pageId="12" pageNumber="188">Costa Rica</collectingCountry>
,
<collectingCountry id="F36588BAFFCF2C54FD3E9632450C6D6A" box="[750,842,1075,1100]" name="Panama" pageId="12" pageNumber="188">Panama</collectingCountry>
,
<collectingCountry id="F36588BAFFCF2C54FC86963245F56D6A" box="[854,947,1075,1100]" name="Jamaica" pageId="12" pageNumber="188">Jamaica</collectingCountry>
, Hispaniola,
<collectingCountry id="F36588BAFFCF2C54FB98963242FF6D6A" box="[1096,1209,1075,1100]" name="Colombia" pageId="12" pageNumber="188">Colombia</collectingCountry>
,
<collectingCountry id="F36588BAFFCF2C54FB169632437A6D6A" box="[1222,1340,1075,1100]" name="Venezuela" pageId="12" pageNumber="188">Venezuela</collectingCountry>
,
<collectingCountry id="F36588BAFFCF2C54FA98963243E16D6A" box="[1352,1447,1075,1100]" name="Ecuador" pageId="12" pageNumber="188">Ecuador</collectingCountry>
,
<collectingCountry id="F36588BAFFCF2C54FF58965646F86D56" box="[136,190,1111,1136]" name="Peru" pageId="12" pageNumber="188">Peru</collectingCountry>
,
<collectingCountry id="F36588BAFFCF2C54FF19965647586D56" box="[201,286,1111,1136]" name="Bolivia" pageId="12" pageNumber="188">Bolivia</collectingCountry>
, Africa,
<collectingCountry id="F36588BAFFCF2C54FEAE965644416D56" box="[382,519,1111,1136]" name="Madagascar" pageId="12" pageNumber="188">Madagascar</collectingCountry>
, Mascarene islands.
</paragraph>
<paragraph id="8BCDC82AFFCF2C54FF6D967A42B36F06" blockId="12.[136,1452,1003,1568]" pageId="12" pageNumber="188">
<emphasis id="B9061438FFCF2C54FF6D967A47F26DB2" bold="true" box="[189,436,1147,1172]" pageId="12" pageNumber="188">Generic comparison:</emphasis>
—
<taxonomicName id="4C72B3A9FFCF2C54FE00967A446B6DB2" authorityName="Shalisko &amp; Sundue" authorityYear="2019" box="[464,557,1147,1172]" class="Polypodiopsida" family="Polypodiaceae" genus="Parrisia" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="188" phylum="Tracheophyta" rank="genus">
<emphasis id="B9061438FFCF2C54FE00967A446B6DB2" box="[464,557,1147,1172]" italics="true" pageId="12" pageNumber="188">Parrisia</emphasis>
</taxonomicName>
belongs to grammitid genera without clearly defined hydathodes(anhydathodous), among them are
<taxonomicName id="4C72B3A9FFCF2C54FE94969E479F6D9E" authorityName="Gaudichaud-Beaupre" authorityYear="1824" box="[324,473,1183,1208]" class="Polypodiopsida" family="Polypodiaceae" genus="Adenophorus" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="188" phylum="Tracheophyta" rank="genus">
<emphasis id="B9061438FFCF2C54FE94969E479F6D9E" box="[324,473,1183,1208]" italics="true" pageId="12" pageNumber="188">Adenophorus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C72B3A9FFCF2C54FE38969E45C66D9E" authority="Parris (1998: 909)" authorityName="Parris" authorityPageNumber="909" authorityYear="1998" box="[488,896,1183,1208]" class="Polypodiopsida" family="Polypodiaceae" genus="Chrysogrammitis" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="188" phylum="Tracheophyta" rank="genus">
<emphasis id="B9061438FFCF2C54FE38969E44EC6D9E" box="[488,682,1183,1208]" italics="true" pageId="12" pageNumber="188">Chrysogrammitis</emphasis>
<bibRefCitation id="EFE3B5DBFFCF2C54FD61969E45C66D9E" author="Parris, B. S." box="[689,896,1183,1208]" pageId="12" pageNumber="188" pagination="909 - 918" refId="ref14474" refString="Parris, B. S. (1998) Chrysogrammitis, a new genus of Grammitidaceae (Filicales). Kew Bulletin 53: 909 - 918. https: // doi. org / 10.2307 / 4118878" type="journal article" year="1998">Parris (1998: 909)</bibRefCitation>
</taxonomicName>
,
<emphasis id="B9061438FFCF2C54FC5C969E42CB6D9E" box="[908,1165,1183,1208]" italics="true" pageId="12" pageNumber="188">
<taxonomicName id="4C72B3A9FFCF2C54FC5C969E42436D9E" authorityName="Bishop" authorityYear="1988" box="[908,1029,1183,1208]" class="Polypodiopsida" family="Polypodiaceae" genus="Ceradenia" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="188" phylum="Tracheophyta" rank="genus">Ceradenia</taxonomicName>
,
<taxonomicName id="4C72B3A9FFCF2C54FBC0969E42CB6D9E" authorityName="sensu Bishop and Smith" authorityYear="1992" box="[1040,1165,1183,1208]" class="Polypodiopsida" family="Polypodiaceae" genus="Enterosora" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="188" phylum="Tracheophyta" rank="genus">Enterosora</taxonomicName>
</emphasis>
(including
<taxonomicName id="4C72B3A9FFCF2C54FADF969E43DD6D9E" baseAuthorityName="Bishop &amp; Smith" baseAuthorityYear="1992" box="[1295,1435,1183,1208]" class="Polypodiopsida" family="Polypodiaceae" genus="Zygophlebia" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="188" phylum="Tracheophyta" rank="genus">
<emphasis id="B9061438FFCF2C54FADF969E43DD6D9E" box="[1295,1435,1183,1208]" italics="true" pageId="12" pageNumber="188">Zygophlebia</emphasis>
</taxonomicName>
),
<taxonomicName id="4C72B3A9FFCF2C54FF5896C247466DFA" authorityName="Swartz" authorityYear="1800" box="[136,256,1219,1244]" class="Polypodiopsida" family="Polypodiaceae" genus="Grammitis" higherTaxonomySource="GBIF" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="188" phylum="Tracheophyta" rank="genus" sensu="stricto">
<emphasis id="B9061438FFCF2C54FF5896C247466DFA" box="[136,256,1219,1244]" italics="true" pageId="12" pageNumber="188">Grammitis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235A943FFCF2C54FEDA96C247696DFA" box="[266,303,1219,1244]" pageId="12" pageNumber="188" sensu="stricto">s.s.</taxonomicNameLabel>
,
<taxonomicName id="4C72B3A9FFCF2C54FE9096C247926DFA" authorityName="Smith" authorityYear="1875" box="[320,468,1219,1244]" class="Polypodiopsida" family="Polypodiaceae" genus="Lomaphlebia" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="188" phylum="Tracheophyta" rank="genus">
<emphasis id="B9061438FFCF2C54FE9096C247926DFA" box="[320,468,1219,1244]" italics="true" pageId="12" pageNumber="188">Lomaphlebia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C72B3A9FFCF2C54FE3596C2453E6DFA" authority="C. Presl (1836: 165 - 166)" authorityName="C. Presl" authorityPageNumber="165 - 166" authorityYear="1836" box="[485,888,1219,1244]" class="Polypodiopsida" family="Polypodiaceae" genus="Prosaptia" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="188" phylum="Tracheophyta" rank="genus">
<emphasis id="B9061438FFCF2C54FE3596C244146DFA" box="[485,594,1219,1244]" italics="true" pageId="12" pageNumber="188">Prosaptia</emphasis>
C.
<bibRefCitation id="EFE3B5DBFFCF2C54FDA396C2453E6DFA" author="Presl, C. B." box="[627,888,1219,1244]" pageId="12" pageNumber="188" refId="ref14778" refString="Presl, C. B. (1836) Tentamen Pteridographiae: seu genera filicacearum praesertim juxta venarum decursum et distributionem exposita. Vol. 3. Typis Filiorum Theophili Haase, Prague, 290 pp. https: // doi. org / 10.5962 / bhl. title. 630" type="book" year="1836">Presl (1836: 165–166)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C72B3A9FFCF2C54FC6696C2476C6C26" authority="Alderwerelt van Rosenburgh (1912: 37 - 39)" authorityName="Alderwerelt van Rosenburgh" authorityPageNumber="37 - 39" authorityYear="1912" class="Polypodiopsida" family="Polypodiaceae" genus="Scleroglossum" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="188" phylum="Tracheophyta" rank="genus">
<emphasis id="B9061438FFCF2C54FC6696C2421E6DFA" box="[950,1112,1219,1244]" italics="true" pageId="12" pageNumber="188">Scleroglossum</emphasis>
<bibRefCitation id="EFE3B5DBFFCF2C54FBB196C2476C6C26" author="Alderwerelt van Rosenburgh, C. R. W. K." pageId="12" pageNumber="188" pagination="1 - 41" refId="ref11235" refString="Alderwerelt van Rosenburgh, C. R. W. K. (1912) New or interesting Malayan ferns 4. Bulletin du Jardin Botanique de Buitenzorg Ser. 2, 2 (7): 1 - 41." type="journal article" year="1912">Alderwerelt van Rosenburgh (1912: 37–39)</bibRefCitation>
</taxonomicName>
. Compared to the phylogenetically related genera, it is distinguished from
<taxonomicName id="4C72B3A9FFCF2C54FBB196E6429F6C26" authorityName="Swartz" authorityYear="1800" box="[1121,1241,1255,1280]" class="Polypodiopsida" family="Polypodiaceae" genus="Grammitis" higherTaxonomySource="GBIF" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="188" phylum="Tracheophyta" rank="genus" sensu="stricto">
<emphasis id="B9061438FFCF2C54FBB196E6429F6C26" box="[1121,1241,1255,1280]" italics="true" pageId="12" pageNumber="188">Grammitis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235A943FFCF2C54FB0F96E643456C26" box="[1247,1283,1255,1280]" pageId="12" pageNumber="188" sensu="stricto">s.s.</taxonomicNameLabel>
by the absence of a dark-colored sclerotized leaf margin, and by the presence of setae on the blade.
<taxonomicName id="4C72B3A9FFCF2C54FBF8970A42FB6C02" authorityName="Gaudichaud-Beaupre" authorityYear="1824" box="[1064,1213,1291,1316]" class="Polypodiopsida" family="Polypodiaceae" genus="Adenophorus" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="188" phylum="Tracheophyta" rank="genus">
<emphasis id="B9061438FFCF2C54FBF8970A42FB6C02" box="[1064,1213,1291,1316]" italics="true" pageId="12" pageNumber="188">Adenophorus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C72B3A9FFCF2C54FA9E970A43ED6C02" authorityName="Shalisko &amp; Sundue" authorityYear="2019" box="[1358,1451,1291,1316]" class="Polypodiopsida" family="Polypodiaceae" genus="Parrisia" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="188" phylum="Tracheophyta" rank="genus">
<emphasis id="B9061438FFCF2C54FA9E970A43ED6C02" box="[1358,1451,1291,1316]" italics="true" pageId="12" pageNumber="188">Parrisia</emphasis>
</taxonomicName>
by their unique uniseriate reddish glandular paraphyses and hairs (
<bibRefCitation id="EFE3B5DBFFCF2C54FCB0972E45B46C6E" author="Ranker, T. A." box="[864,1010,1327,1352]" pageId="12" pageNumber="188" pagination="170 - 171" refId="ref15291" refString="Ranker, T. A. (2008) A New Combination in Adenophorus (Polypodiaceae). American Fern Journal 98 (3): 170 - 171. https: // doi. org / 10.1640 / 0002 - 8444 (2008) 98 [170: ANCIAP] 2.0. CO; 2" type="journal article" year="2008">Ranker 2008</bibRefCitation>
) that are lacking
<taxonomicName id="4C72B3A9FFCF2C54FB7E972E434D6C6E" authorityName="Shalisko &amp; Sundue" authorityYear="2019" box="[1198,1291,1327,1352]" class="Polypodiopsida" family="Polypodiaceae" genus="Parrisia" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="188" phylum="Tracheophyta" rank="genus">
<emphasis id="B9061438FFCF2C54FB7E972E434D6C6E" box="[1198,1291,1327,1352]" italics="true" pageId="12" pageNumber="188">Parrisia</emphasis>
</taxonomicName>
.
<taxonomicName id="4C72B3A9FFCF2C54FAC8972E43EA6C6E" authorityName="Smith" authorityYear="1875" box="[1304,1452,1327,1352]" class="Polypodiopsida" family="Polypodiaceae" genus="Lomaphlebia" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="188" phylum="Tracheophyta" rank="genus">
<emphasis id="B9061438FFCF2C54FAC8972E43EA6C6E" box="[1304,1452,1327,1352]" italics="true" pageId="12" pageNumber="188">Lomaphlebia</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C72B3A9FFCF2C54FEAC9752479F6C4A" authorityName="Shalisko &amp; Sundue" authorityYear="2019" box="[380,473,1363,1388]" class="Polypodiopsida" family="Polypodiaceae" genus="Parrisia" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="188" phylum="Tracheophyta" rank="genus">
<emphasis id="B9061438FFCF2C54FEAC9752479F6C4A" box="[380,473,1363,1388]" italics="true" pageId="12" pageNumber="188">Parrisia</emphasis>
</taxonomicName>
by the presence of a commissure vein at the margin of the leaf blade, thus presenting a row of areoles along the margin.
<taxonomicName id="4C72B3A9FFCF2C54FE24977644316CB6" authorityName="Kaulfuss" authorityYear="1820" box="[500,631,1399,1424]" class="Polypodiopsida" family="Polypodiaceae" genus="Cochlidium" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="188" phylum="Tracheophyta" rank="genus">
<emphasis id="B9061438FFCF2C54FE24977644316CB6" box="[500,631,1399,1424]" italics="true" pageId="12" pageNumber="188">Cochlidium</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C72B3A9FFCF2C54FCBC9776458F6CB6" authorityName="Shalisko &amp; Sundue" authorityYear="2019" box="[876,969,1399,1424]" class="Polypodiopsida" family="Polypodiaceae" genus="Parrisia" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="188" phylum="Tracheophyta" rank="genus">
<emphasis id="B9061438FFCF2C54FCBC9776458F6CB6" box="[876,969,1399,1424]" italics="true" pageId="12" pageNumber="188">Parrisia</emphasis>
</taxonomicName>
by the presence of hydathodes, the absence of dark colored setae, and presence of a coenosorus deeply immersed into the lamina in several species.
<taxonomicName id="4C72B3A9FFCF2C54FAE5979A43ED6C92" authorityName="Bishop" authorityYear="1988" box="[1333,1451,1435,1460]" class="Polypodiopsida" family="Polypodiaceae" genus="Ceradenia" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="188" phylum="Tracheophyta" rank="genus">
<emphasis id="B9061438FFCF2C54FAE5979A43ED6C92" box="[1333,1451,1435,1460]" italics="true" pageId="12" pageNumber="188">Ceradenia</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C72B3A9FFCF2C54FE5697BE47A56CFE" authorityName="Shalisko &amp; Sundue" authorityYear="2019" box="[390,483,1471,1496]" class="Polypodiopsida" family="Polypodiaceae" genus="Parrisia" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="188" phylum="Tracheophyta" rank="genus">
<emphasis id="B9061438FFCF2C54FE5697BE47A56CFE" box="[390,483,1471,1496]" italics="true" pageId="12" pageNumber="188">Parrisia</emphasis>
</taxonomicName>
by the presence of waxy deposits of a white, yellowish, or tan color, produced by paraphyses, as well as a lack of circumsoral setae.
<taxonomicName id="4C72B3A9FFCF2C54FD1797E245626CDA" authorityName="Shalisko &amp; Sundue" authorityYear="2019" box="[711,804,1507,1532]" class="Polypodiopsida" family="Polypodiaceae" genus="Parrisia" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="188" phylum="Tracheophyta" rank="genus">
<emphasis id="B9061438FFCF2C54FD1797E245626CDA" box="[711,804,1507,1532]" italics="true" pageId="12" pageNumber="188">Parrisia</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C72B3A9FFCF2C54FBF397E242E66CDA" authorityName="sensu Bishop and Smith" authorityYear="1992" box="[1059,1184,1507,1532]" class="Polypodiopsida" family="Polypodiaceae" genus="Enterosora" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="188" phylum="Tracheophyta" rank="genus">
<emphasis id="B9061438FFCF2C54FBF397E242E66CDA" box="[1059,1184,1507,1532]" italics="true" pageId="12" pageNumber="188">Enterosora</emphasis>
</taxonomicName>
by radial rhizomes (vs. dorsiventral), the location of sori in the medium of the vein and the presence of setae around the sori.
</paragraph>
<paragraph id="8BCDC82AFFCF2C54FF58944844B36F44" blockId="12.[136,757,1609,1634]" box="[136,757,1609,1634]" pageId="12" pageNumber="188">The following species are transferred to the new genus:</paragraph>
</subSubSection>
</treatment>
</document>
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