192 lines
23 KiB
XML
192 lines
23 KiB
XML
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<mods:title id="9572F68727A7C09CF2D4D9EF81E536AD">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<mods:namePart id="6F1929434879CCD1B7CCE2A076D1DF14">Rosa, Cecilia Lozano De La</mods:namePart>
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<mods:affiliation id="004B53478DFEDFF67E70627AD1B8F124">Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela.</mods:affiliation>
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<mods:nameIdentifier id="C98A1446CD5260DC84920E5A3C88AA59" type="email">lozanoceci@gmail.com</mods:nameIdentifier>
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<mods:namePart id="F881E71D4A8DB4436B4D1EF4C1862117">Cupello, Mario</mods:namePart>
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<mods:affiliation id="0C492FA599A9C2B64ED25F7217B7D76D">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A & M University, College Station, TX, 77843, USA.</mods:affiliation>
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<mods:nameIdentifier id="04374F53618ADD6F28A3792AB7D56551" type="email">mcupello@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="52D0F0B3865F8C97BE8A81DCF7EC3347">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:affiliation id="60587C9F93DD8233E26D4D36EAC8C779">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="81780B9BBE7C644FC6F270D670582FF4" type="email">vazdemello@gmail.com</mods:nameIdentifier>
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<mods:date id="83DE8377933E6F61B9D663D0DF9E5215">2024</mods:date>
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<treatment id="1D507666308952B0EF09FB24754500FE" ID-DOI="http://doi.org/10.5281/zenodo.13945930" ID-GBIF-Taxon="246298181" ID-Zenodo-Dep="13945930" LSID="urn:lsid:plazi:treatment:1D507666308952B0EF09FB24754500FE" httpUri="http://treatment.plazi.org/id/1D507666308952B0EF09FB24754500FE" lastPageId="212" lastPageNumber="213" pageId="211" pageNumber="212">
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<subSubSection id="DDE394FB308952B7EF09FB24731905A1" box="[542,1045,1155,1182]" pageId="211" pageNumber="212" type="nomenclature">
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<paragraph id="9546C770308952B7EF09FB24731905A1" blockId="211.[542,1045,1155,1182]" box="[542,1045,1155,1182]" pageId="211" pageNumber="212">
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<heading id="CE0E701C308952B7EF09FB24731905A1" box="[542,1045,1155,1182]" centered="true" fontSize="11" level="2" pageId="211" pageNumber="212" reason="2">
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<taxonomicName id="52F9BCF3308952B7EF09FB24731905A1" ID-CoL="QNB9" authority="(Boheman, 1858)" baseAuthorityName="Boheman" baseAuthorityYear="1858" box="[542,1045,1155,1182]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="trinodosum">
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<emphasis id="A78D1B62308952B7EF09FB24744D05A1" bold="true" box="[542,833,1155,1182]" italics="true" pageId="211" pageNumber="212">Canthidium trinodosum</emphasis>
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(
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<bibRefCitation id="F168BA81308952B7EE47FB24730105A1" author="Boheman C. H." box="[848,1037,1155,1182]" pageId="211" pageNumber="212" pagination="1 - 112" refId="ref111809" refString="Boheman C. H. 1858. Coleoptera. Species novas descripsit. In: Boheman C. H. (1858 - 1859) Kongliga Svenska Fregatten Eugenies Resa omkring Jorden under befȧl af C. A. Virgin, aren 1851 - 1853. Vetenskapliga Iakttagelser pa H. M. Konung Oscar den Forstes befallning utgifna af K. Svenska Vetenskaps Akademien. Andra delen. Zoologi. 1. Insecta. Hafte 4: 1 - 112. P. A. Norstedt & Soner, Stockholm. [Publication date according to Bousquet (2016).] https: // doi. org / 10.5962 / bhl. title. 2467" type="journal article" year="1858">Boheman, 1858</bibRefCitation>
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)
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="DDE394FB308952B7EDAAFB63755C06FD" pageId="211" pageNumber="212" type="reference_group">
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<paragraph id="9546C770308952B7EDAAFB63738A04EA" blockId="211.[189,1400,1220,1986]" pageId="211" pageNumber="212">
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<treatmentCitationGroup id="B5E9E05E308952B7EDAAFB63738A04EA" pageId="211" pageNumber="212">
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<treatmentCitation id="1458E161308952B7EDAAFB6375C205E1" author="Boheman C. H." box="[189,718,1220,1247]" page="46" pageId="211" pageNumber="212" year="1858">
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<taxonomicName id="52F9BCF3308952B7EDAAFB6375C205E1" ID-CoL="7XPDD" authority="Boheman, 1858: 46" authorityName="Boheman" authorityPageNumber="46" authorityYear="1858" box="[189,718,1220,1247]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="trinodosus">
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<emphasis id="A78D1B62308952B7EDAAFB6376D205E1" box="[189,478,1220,1246]" italics="true" pageId="211" pageNumber="212">Onthophagus trinodosus</emphasis>
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<bibRefCitation id="F168BA81308952B7ECF2FB6375C205E1" author="Boheman C. H." box="[485,718,1220,1247]" pageId="211" pageNumber="212" pagination="1 - 112" refId="ref111809" refString="Boheman C. H. 1858. Coleoptera. Species novas descripsit. In: Boheman C. H. (1858 - 1859) Kongliga Svenska Fregatten Eugenies Resa omkring Jorden under befȧl af C. A. Virgin, aren 1851 - 1853. Vetenskapliga Iakttagelser pa H. M. Konung Oscar den Forstes befallning utgifna af K. Svenska Vetenskaps Akademien. Andra delen. Zoologi. 1. Insecta. Hafte 4: 1 - 112. P. A. Norstedt & Soner, Stockholm. [Publication date according to Bousquet (2016).] https: // doi. org / 10.5962 / bhl. title. 2467" type="journal article" year="1858">Boheman, 1858: 46</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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. Type locality:
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<collectingCountry id="EDEE87E0308952B7EE97FB6374C005E1" box="[896,972,1220,1246]" name="Brazil" pageId="211" pageNumber="212">Brazil</collectingCountry>
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:
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<collectingRegion id="573D0992308952B7EEC0FB63738F05E1" box="[983,1155,1220,1246]" country="Brazil" name="Rio de Janeiro" pageId="211" pageNumber="212">Rio de Janeiro</collectingRegion>
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. Name-bearing type: a single known
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<typeStatus id="4A4279D2308952B7ECB1FB4F750D043D" box="[422,513,1256,1282]" pageId="211" pageNumber="212" type="syntype">syntype</typeStatus>
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(NHRS), examined by FZVM. Note: Boheman’s name is a junior primary homonym of
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<taxonomicName id="52F9BCF3308952B7EC9DFAAC74660419" ID-CoL="7XPDD" authority="Fahraeus, 1857" authorityName="Fahraeus" authorityYear="1857" box="[394,874,1290,1318]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="trinodosus">
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||
<emphasis id="A78D1B62308952B7EC9DFAAC75A0041A" box="[394,684,1291,1317]" italics="true" pageId="211" pageNumber="212">Onthophagus trinodosus</emphasis>
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Fåhraeus, 1857
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</taxonomicName>
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. Both names are currently in use, the former for a South American species treated in the genus
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<taxonomicName id="52F9BCF3308952B7EE53FA8974C30477" ID-CoL="3GVY" authorityName="Erichson" authorityYear="1847" box="[836,975,1326,1352]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="genus">
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<emphasis id="A78D1B62308952B7EE53FA8974C30477" box="[836,975,1326,1352]" italics="true" pageId="211" pageNumber="212">Canthidium</emphasis>
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</taxonomicName>
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since
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<treatmentCitation id="1458E161308952B7E90CFA8973D90477" author="Harold E." box="[1051,1237,1326,1352]" pageId="211" pageNumber="212" year="1867">
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||
<bibRefCitation id="F168BA81308952B7E90CFA8973D90477" author="Harold E." box="[1051,1237,1326,1352]" pageId="211" pageNumber="212" pagination="60 - 93" refId="ref123189" refString="Harold E. 1867 b. Nachtrag zur Bearbeitung der Gattung Canthidium. Coleopterologische Hefte 2: 60 - 93. Available from https: // www. biodiversitylibrary. org / page / 16069403 [accessed 16 Feb. 2024]." type="journal article" year="1867">Harold (1867b)</bibRefCitation>
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</treatmentCitation>
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, the latter for an African species of
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<taxonomicName id="52F9BCF3308952B7ECFFFAF6758A0454" ID-CoL="68DQ" authorityName="Latreille" authorityYear="1802" box="[488,646,1361,1387]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="genus">
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<emphasis id="A78D1B62308952B7ECFFFAF6758A0454" box="[488,646,1361,1387]" italics="true" pageId="211" pageNumber="212">Onthophagus</emphasis>
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</taxonomicName>
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. Since the two primary homonyms are in use and have not been applied to taxa considered congeneric after 1899, the junior homonym must not be automatically replaced as it would be in other situations. The case must instead be referred to the Commission under Article 23.9.5 for a ruling. In the meantime, current usage is maintained.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="9546C770308952B7EDAAFA79755C06FD" blockId="211.[189,1400,1220,1986]" pageId="211" pageNumber="212">
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<treatmentCitationGroup id="B5E9E05E308952B7EDAAFA79755C06FD" pageId="211" pageNumber="212">
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||
<treatmentCitation id="1458E161308952B7EDAAFA7975ED04C7" author="Harold E." box="[189,737,1502,1528]" page="403" pageId="211" pageNumber="212" year="1862">
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||
<taxonomicName id="52F9BCF3308952B7EDAAFA7975ED04C7" ID-CoL="49Q2D" authority="Harold, 1862: 403" authorityName="Harold" authorityPageNumber="403" authorityYear="1862" box="[189,737,1502,1528]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="subarmatus">
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||
<emphasis id="A78D1B62308952B7EDAAFA7976FD04C7" box="[189,497,1502,1528]" italics="true" pageId="211" pageNumber="212">Onthophagus subarmatus</emphasis>
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||
<bibRefCitation id="F168BA81308952B7ECE8FA7975ED04C7" author="Harold E." box="[511,737,1502,1528]" pageId="211" pageNumber="212" pagination="379 - 403" refId="ref123046" refString="Harold E. 1862. Beitrage zur Kenntnifs einiger Coprophagen Lamellicornien (Viertes Stuck). Berliner Entomologische Zeitschrift 6: 379 - 403. https: // doi. org / 10.1002 / mmnd. 47918620310" type="journal article" year="1862">Harold, 1862: 403</bibRefCitation>
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||
</taxonomicName>
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||
</treatmentCitation>
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||
(new replacement name for
|
||
<treatmentCitation id="1458E161308952B7E958FA7976A40723" author="Boheman C. H." pageId="211" pageNumber="212" year="1858">
|
||
<taxonomicName id="52F9BCF3308952B7E958FA7976A40723" ID-CoL="7XPDD" authority="Boheman, 1858" authorityName="Boheman" authorityYear="1858" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="trinodosus">
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||
<emphasis id="A78D1B62308952B7E958FA79727B04C7" box="[1103,1399,1502,1528]" italics="true" pageId="211" pageNumber="212">Onthophagus trinodosus</emphasis>
|
||
<bibRefCitation id="F168BA81308952B7EDFDF9A676A40723" author="Boheman C. H." box="[234,424,1537,1564]" pageId="211" pageNumber="212" pagination="1 - 112" refId="ref111809" refString="Boheman C. H. 1858. Coleoptera. Species novas descripsit. In: Boheman C. H. (1858 - 1859) Kongliga Svenska Fregatten Eugenies Resa omkring Jorden under befȧl af C. A. Virgin, aren 1851 - 1853. Vetenskapliga Iakttagelser pa H. M. Konung Oscar den Forstes befallning utgifna af K. Svenska Vetenskaps Akademien. Andra delen. Zoologi. 1. Insecta. Hafte 4: 1 - 112. P. A. Norstedt & Soner, Stockholm. [Publication date according to Bousquet (2016).] https: // doi. org / 10.5962 / bhl. title. 2467" type="journal article" year="1858">Boheman, 1858</bibRefCitation>
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||
</taxonomicName>
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||
</treatmentCitation>
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||
due to its junior primary homonymy with
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<treatmentCitation id="1458E161308952B7EEACF9A572290723" author="Fåhraeus" box="[955,1317,1537,1564]" pageId="211" pageNumber="212" year="1857">
|
||
<taxonomicName id="52F9BCF3308952B7EEACF9A572290723" ID-CoL="7XPDD" authority="Fahraeus, 1857" authorityName="Fahraeus" authorityYear="1857" box="[955,1317,1537,1564]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="trinodosus">
|
||
<emphasis id="A78D1B62308952B7EEACF9A573520724" box="[955,1118,1537,1564]" italics="true" pageId="211" pageNumber="212">O. trinodosus</emphasis>
|
||
<bibRefCitation id="F168BA81308952B7E97EF9A672290723" box="[1129,1317,1537,1564]" pageId="211" pageNumber="212" year="1857">Fåhraeus, 1857</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
).
|
||
<typeStatus id="4A4279D2308952B7E82BF9A5727A0723" box="[1340,1398,1538,1564]" pageId="211" pageNumber="212">Type</typeStatus>
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||
locality and name-bearing
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||
<typeStatus id="4A4279D2308952B7EF3EF982756C0700" box="[553,608,1573,1599]" pageId="211" pageNumber="212">type</typeStatus>
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||
: the same as for
|
||
<taxonomicName id="52F9BCF3308952B7EE38F98373C50701" ID-CoL="7XPDD" authority="Boheman" authorityName="Boheman" authorityYear="1858" box="[815,1225,1572,1598]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="trinodosus">
|
||
<emphasis id="A78D1B62308952B7EE38F983735E0701" box="[815,1106,1572,1598]" italics="true" pageId="211" pageNumber="212">Onthophagus trinodosus</emphasis>
|
||
Boheman
|
||
</taxonomicName>
|
||
in accordance with Article 72.7. Note: when
|
||
<treatmentCitation id="1458E161308952B7EF4CF9EF7408075D" author="Harold E." box="[603,772,1608,1634]" pageId="211" pageNumber="212" year="1862">
|
||
<bibRefCitation id="F168BA81308952B7EF4CF9EF7408075D" author="Harold E." box="[603,772,1608,1634]" pageId="211" pageNumber="212" pagination="379 - 403" refId="ref123046" refString="Harold E. 1862. Beitrage zur Kenntnifs einiger Coprophagen Lamellicornien (Viertes Stuck). Berliner Entomologische Zeitschrift 6: 379 - 403. https: // doi. org / 10.1002 / mmnd. 47918620310" type="journal article" year="1862">Harold (1862)</bibRefCitation>
|
||
</treatmentCitation>
|
||
established this nomen novum, the species was still treated in
|
||
<taxonomicName id="52F9BCF3308952B7EC4AF9CC76F707BA" ID-CoL="68DQ" authorityName="Latreille" authorityYear="1802" box="[349,507,1643,1669]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A78D1B62308952B7EC4AF9CC76F707BA" box="[349,507,1643,1669]" italics="true" pageId="211" pageNumber="212">Onthophagus</emphasis>
|
||
</taxonomicName>
|
||
. However, five years later, after seeing “the
|
||
<typeStatus id="4A4279D2308952B7EEE8F9CC733E07BA" box="[1023,1074,1643,1669]" pageId="211" pageNumber="212">type</typeStatus>
|
||
specimen”,
|
||
<treatmentCitation id="1458E161308952B7E9D5F9CC727A07BA" author="Harold E." box="[1218,1398,1643,1669]" pageId="211" pageNumber="212" year="1867">
|
||
<bibRefCitation id="F168BA81308952B7E9D5F9CC727A07BA" author="Harold E." box="[1218,1398,1643,1669]" pageId="211" pageNumber="212" pagination="60 - 93" refId="ref123189" refString="Harold E. 1867 b. Nachtrag zur Bearbeitung der Gattung Canthidium. Coleopterologische Hefte 2: 60 - 93. Available from https: // www. biodiversitylibrary. org / page / 16069403 [accessed 16 Feb. 2024]." type="journal article" year="1867">Harold (1867b)</bibRefCitation>
|
||
</treatmentCitation>
|
||
transferred it to
|
||
<taxonomicName id="52F9BCF3308952B7ECB3F92975230797" ID-CoL="3GVY" authorityName="Erichson" authorityYear="1847" box="[420,559,1678,1704]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A78D1B62308952B7ECB3F92975230797" box="[420,559,1678,1704]" italics="true" pageId="211" pageNumber="212">Canthidium</emphasis>
|
||
</taxonomicName>
|
||
and resurrected the replaced name, invalidating his own nomen novum. He did so because, in his view, the homonymy had been undone once the species was transferred to another genus and, therefore, nothing prevented Boheman’s name from being used. This is not the interpretation of the current Code: primary homonymies are not undone even if the homonyms are no longer applied to congeneric taxa; the junior homonym is, with a few exceptions, permanently invalid according to Article 57.2 and a substitute name should be adopted following Article 60. But since, in this case, as explained above, the junior primary homonym is in prevailing usage in a different genus as that of its senior homonym, Harold’s decision should be respected until the Commission rules on the case.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DDE394FB308E52B0EDAAFEAF76CA006E" box="[189,454,264,337]" pageId="212" pageNumber="213" type="distribution">
|
||
<paragraph id="9546C770308E52B0EDAAFEAF765A001D" blockId="212.[189,342,264,290]" box="[189,342,264,290]" pageId="212" pageNumber="213">
|
||
<heading id="CE0E701C308E52B0EDAAFEAF765A001D" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="212" pageNumber="213" reason="3">
|
||
<emphasis id="A78D1B62308E52B0EDAAFEAF765A001D" bold="true" box="[189,342,264,290]" pageId="212" pageNumber="213">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="9546C770308E52B0EDAAFE9076CA006E" blockId="212.[189,454,311,337]" box="[189,454,311,337]" pageId="212" pageNumber="213">
|
||
<collectingCountry id="EDEE87E0308E52B0EDAAFE907609006E" box="[189,261,311,337]" name="Brazil" pageId="212" pageNumber="213">Brazil</collectingCountry>
|
||
(Cupello 2018).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DDE394FB308E52B0EDAAFEDF754500FE" pageId="212" pageNumber="213" type="reference_group">
|
||
<paragraph id="9546C770308E52B0EDAAFEDF769500AC" blockId="212.[189,409,376,403]" box="[189,409,376,403]" pageId="212" pageNumber="213">
|
||
<heading id="CE0E701C308E52B0EDAAFEDF769500AC" bold="true" box="[189,409,376,403]" fontSize="11" level="3" pageId="212" pageNumber="213" reason="3">
|
||
<emphasis id="A78D1B62308E52B0EDAAFEDF769500AC" bold="true" box="[189,409,376,403]" pageId="212" pageNumber="213">Literature record</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="9546C770308E52B0EDAAFE0F754500FE" blockId="212.[189,585,423,450]" box="[189,585,423,450]" pageId="212" pageNumber="213">
|
||
<bibRefCitation id="F168BA81308E52B0EDAAFE0F766400FE" author="Roze J. A." box="[189,360,423,450]" pageId="212" pageNumber="213" pagination="39 - 63" refId="ref137914" refString="Roze J. A. 1955. Lista preliminar de la familia Scarabaeidae sensu lato de Venezuela. Boletin del Museo de Ciencias Naturales 1 (1): 39 - 63." type="journal article" year="1955">Roze 1955: 44</bibRefCitation>
|
||
(cited for
|
||
<collectingRegion id="573D0992308E52B0ECF6FE00753100FE" box="[481,573,423,449]" country="Venezuela" name="Aragua" pageId="212" pageNumber="213">Aragua</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |