treatments-xml/data/02/D7/6E/02D76E62717C4B1F102FE1242232BBF9.xml
2024-06-21 12:21:39 +02:00

256 lines
21 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.124.34679" ID-GBIF-Dataset="0ed8fe19-ae7d-4551-b04c-40506ffc4274" ID-PMC="PMC6598937" ID-Pensoft-Pub="1314-2003-124-87" ID-Pensoft-UUID="CD370A37FFD8203C596C8A6F9E10FFC0" ID-PubMed="31274979" ID-Zenodo-Dep="3261874" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1314-2003-124-87" ModsDocOrigin="PhytoKeys 124" ModsDocTitle="Insights on the systematics and morphology of Humiriaceae (Malpighiales): androecial and extrafloral nectary variation, two new combinations, and a new Sacoglottis from Guyana" checkinTime="1561202160728" checkinUser="pensoft" docAuthor="Wurdack, Kenneth J. &amp; Zartman, Charles E." docDate="2019" docId="02D76E62717C4B1F102FE1242232BBF9" docLanguage="en" docName="PhytoKeys 124: 87-121" docOrigin="PhytoKeys 124" docSource="http://dx.doi.org/10.3897/phytokeys.124.34679" docTitle="Duckesia liesneri K. Wurdack &amp; C. E. Zartman 2019, comb. nov." docType="treatment" docVersion="6" id="CD370A37FFD8203C596C8A6F9E10FFC0" lastPageNumber="96" masterDocId="CD370A37FFD8203C596C8A6F9E10FFC0" masterDocTitle="Insights on the systematics and morphology of Humiriaceae (Malpighiales): androecial and extrafloral nectary variation, two new combinations, and a new Sacoglottis from Guyana" masterLastPageNumber="121" masterPageNumber="87" pageNumber="96" updateTime="1668139343810" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Insights on the systematics and morphology of Humiriaceae (Malpighiales): androecial and extrafloral nectary variation, two new combinations, and a new Sacoglottis from Guyana</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wurdack, Kenneth J.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zartman, Charles E.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>124</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>87</mods:start>
<mods:end>121</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.124.34679</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.124.34679</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-124-87</mods:identifier>
<mods:identifier type="Pensoft-UUID">CD370A37FFD8203C596C8A6F9E10FFC0</mods:identifier>
<mods:identifier type="Zenodo-Dep">3261874</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="158363996" LSID="urn:lsid:plazi:treatment:02D76E62717C4B1F102FE1242232BBF9" httpUri="http://treatment.plazi.org/id/02D76E62717C4B1F102FE1242232BBF9" lastPageNumber="96" pageId="9" pageNumber="96">
<subSubSection pageId="9" pageNumber="96" type="nomenclature">
<paragraph pageId="9" pageNumber="96">
<taxonomicName LSID="2C532402-EF13-5431-8723-415C6A0D15E9" authority="(Cuatrec.) K. Wurdack &amp; C. E. Zartman" authorityName="K. Wurdack &amp; C. E. Zartman" authorityYear="2019" baseAuthorityName="K. Wurdack &amp; C. E. Zartman" baseAuthorityYear="2019" class="Magnoliopsida" family="Humiriaceae" genus="Duckesia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Duckesia liesneri" order="Malpighiales" pageId="9" pageNumber="96" phylum="Tracheophyta" rank="species" species="liesneri" status="comb. nov.">Duckesia liesneri (Cuatrec.) K.Wurdack &amp; C.E.Zartman</taxonomicName>
<taxonomicNameLabel pageId="9" pageNumber="96">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="96" type="reference_group">
<paragraph pageId="9" pageNumber="96">
<taxonomicName authorityName="J. Cuatrec" authorityYear="1991" class="Magnoliopsida" family="Humiriaceae" genus="Humiriastrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Humiriastrum liesneri" order="Malpighiales" pageId="9" pageNumber="96" phylum="Tracheophyta" rank="species" species="liesneri">Humiriastrum liesneri</taxonomicName>
Cuatrec., Phytologia 71: 165. 1991. Type: VENEZUELA. Amazonas, Depto. Rio Negro: Cerro Aracamuni, summit, Proa camp, 1400 m, 28 Oct 1987,
<emphasis italics="true" pageId="9" pageNumber="96">R. Liesner &amp; G. Carnevali 22589</emphasis>
(holotype: US-3111383; isotypes: MO-3641994, NY-00329209, US-3118837, VEN-277523).
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="96" type="notes">
<paragraph pageId="9" pageNumber="96">Notes.</paragraph>
<paragraph pageId="9" pageNumber="96">
When
<bibRefCitation author="Cuatrecasas, J" journalOrPublisher="Contributions from the United States National Herbarium" pageId="20" pageNumber="107" refId="B6" refString="Cuatrecasas, J, 1991. Miscellaneous notes on neotropical flora XX. A new species of Humiriastrum. Phytologia 71: 165-166." title="Miscellaneous notes on neotropical flora XX. A new species of Humiriastrum. Phytologia 71: 165 - 166." year="1991">Cuatrecasas (1991)</bibRefCitation>
originally described
<taxonomicName authorityName="J. Cuatrec" authorityYear="1991" class="Magnoliopsida" family="Humiriaceae" genus="Humiriastrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Humiriastrum liesneri" order="Malpighiales" pageId="9" pageNumber="96" phylum="Tracheophyta" rank="species" species="liesneri">
<emphasis italics="true" pageId="9" pageNumber="96">Humiriastrum liesneri</emphasis>
</taxonomicName>
, he expressed uncertainty about its anther structure and relationships due to reliance on young buds for floral details. A second collection (Venezuela. Amazonas, Sierra de
<normalizedToken originalValue="Unturán">Unturan</normalizedToken>
, 1150 m, 3 Feb 1989,
<emphasis italics="true" pageId="9" pageNumber="96">Henderson 933</emphasis>
, US), correctly annotated by Cuatrecasas as
<taxonomicName lsidName="H. liesneri" pageId="9" pageNumber="96" rank="species" species="liesneri">
<emphasis italics="true" pageId="9" pageNumber="96">H. liesneri</emphasis>
</taxonomicName>
in 1994, is in flower but did not cause any classification changes for the subsequent Flora of the Venezuelan Guayana (
<bibRefCitation author="Cuatrecasas, J" editor="Berry, PE" journalOrPublisher="Volume 5 Eriocaulaceae-Lentibulariaceae. Missouri Botanical Garden Press, St. Louis" pageId="20" pageNumber="107" pagination="623 - 641" refId="B7" refString="Cuatrecasas, J, Huber, O, 1999. Humiriaceae. In: Berry, PE, Yatskievych, K, Holst, BK, Eds., Flora of the Venezuelan Guayana. Volume 5 Eriocaulaceae-Lentibulariaceae. Missouri Botanical Garden Press, St. Louis: 623 - 641" title="Humiriaceae." volumeTitle="Flora of the Venezuelan Guayana." year="1999">Cuatrecasas and Huber 1999</bibRefCitation>
). The anther morphology of this species (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Stamen structure of Humiriaceae. A Duckesia verrucosa tetrasporangiate anther, dorsal B Duckesia verrucosa tetrasporangiate anther, lateral C Duckesia liesneri tetrasporangiate anther, lateral D Endopleura uchi disporangiate anther, lateral E Endopleura uchi tetrasporangiate anther, lateral F Duckesia verrucosa sterile anther, lateral G Humiriastrum cuspidatum disporangiate anther, lateral H Humiriastrum cuspidatum androecium, dorsal I Endopleura uchi tetrasporangiate anther, lateral J Humiriastrum dentatum disporangiate anther, lateral K Humiriastrum dentatum disporangiate anther, ventral L Humiriastrum diguense disporangiate anther, lateral M Humiria balsamifera var. imbaimadaiensis stamen cluster with 2 of 3 types, dorsal. f = filament attachment location. Sources: A, B, F Ducke 2108 C Henderson 933 D, E, I Assuncao 605 G, H Cid et al. 4264 J, K Hatschbach 56145 L Quizhpe et al. 612 M Wurdack 4814 (all US.)" figureDoi="10.3897/phytokeys.124.34679.figure1" httpUri="https://binary.pensoft.net/fig/311887" pageId="9" pageNumber="96">1C</figureCitation>
), with its four separate sporangia, suggests an affiliation with
<taxonomicName authorityName="Cuatrecasas" authorityYear="1961" class="Magnoliopsida" family="Humiriaceae" genus="Duckesia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Duckesia" order="Malpighiales" pageId="9" pageNumber="96" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="96">Duckesia</emphasis>
</taxonomicName>
or
<taxonomicName class="Magnoliopsida" family="Humiriaceae" genus="Endopleura" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endopleura" order="Malpighiales" pageId="9" pageNumber="96" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="96">Endopleura</emphasis>
</taxonomicName>
, and the transfer here to the former is supported by more similar connective morphology, absence of disporangiate anthers, and similar small leaves (Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of select diagnostic features for Duckesia and Endopleura." httpUri="http://table.plazi.org/id/C75AAAD181AA802DEC612B974E10A405" pageId="9" pageNumber="96" tableUuid="C75AAAD181AA802DEC612B974E10A405">1</tableCitation>
). The two species of
<taxonomicName authorityName="Cuatrecasas" authorityYear="1961" class="Magnoliopsida" family="Humiriaceae" genus="Duckesia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Duckesia" order="Malpighiales" pageId="9" pageNumber="96" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="96">Duckesia</emphasis>
</taxonomicName>
are otherwise very distinct from each other; fruits remain unknown for
<taxonomicName class="Dicotyledoneae" family="Humiriaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="D. liesneri" order="Malpighiales" pageId="9" pageNumber="96" phylum="Angiospermae" rank="species" species="liesneri">
<emphasis italics="true" pageId="9" pageNumber="96">D. liesneri</emphasis>
</taxonomicName>
and would likely provide additional distinguishing characters. The two collections of
<taxonomicName class="Dicotyledoneae" family="Humiriaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="D. liesneri" order="Malpighiales" pageId="9" pageNumber="96" phylum="Angiospermae" rank="species" species="liesneri">
<emphasis italics="true" pageId="9" pageNumber="96">D. liesneri</emphasis>
</taxonomicName>
are ca. 60 km apart on isolated Venezuelan tepuis near the Brazilian border and north of
<taxonomicName class="Dicotyledoneae" family="Humiriaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="D. verrucosa" order="Malpighiales" pageId="9" pageNumber="96" phylum="Angiospermae" rank="species" species="verrucosa">
<emphasis italics="true" pageId="9" pageNumber="96">D. verrucosa</emphasis>
</taxonomicName>
, which is a lowland Brazilian species.
</paragraph>
<caption ID-Table-UUID="C75AAAD181AA802DEC612B974E10A405" httpUri="http://table.plazi.org/id/C75AAAD181AA802DEC612B974E10A405" pageId="9" pageNumber="96" start="Table 1" startId="T1">
<paragraph pageId="9" pageNumber="96">
<emphasis bold="true" pageId="9" pageNumber="96">Table 1.</emphasis>
Comparison of select diagnostic features for
<emphasis italics="true" pageId="9" pageNumber="96">
<taxonomicName authorityName="Cuatrecasas" authorityYear="1961" class="Magnoliopsida" family="Humiriaceae" genus="Duckesia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Duckesia" order="Malpighiales" pageId="9" pageNumber="96" phylum="Tracheophyta" rank="genus">Duckesia</taxonomicName>
and
<taxonomicName class="Magnoliopsida" family="Humiriaceae" genus="Endopleura" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endopleura" order="Malpighiales" pageId="9" pageNumber="96" phylum="Tracheophyta" rank="genus">Endopleura</taxonomicName>
</emphasis>
.
</paragraph>
</caption>
<paragraph pageId="9" pageNumber="96">
<table pageId="9" pageNumber="96">
<tr pageId="9" pageNumber="96">
<th colspan="1" pageId="9" pageNumber="96" rowspan="1">Characters</th>
<th colspan="1" pageId="9" pageNumber="96" rowspan="1">
<taxonomicName authorityName="Cuatrec" authorityYear="1961" baseAuthorityName="Ducke" class="Magnoliopsida" family="Humiriaceae" genus="Duckesia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Duckesia verrucosa" order="Malpighiales" pageId="9" pageNumber="96" phylum="Tracheophyta" rank="species" species="verrucosa">
<emphasis italics="true" pageId="9" pageNumber="96">Duckesia verrucosa</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="9" pageNumber="96" rowspan="1">
<taxonomicName authorityName="K. Wurdack &amp; C. E. Zartman" authorityYear="2019" baseAuthorityName="K. Wurdack &amp; C. E. Zartman" baseAuthorityYear="2019" class="Magnoliopsida" family="Humiriaceae" genus="Duckesia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Duckesia liesneri" order="Malpighiales" pageId="9" pageNumber="96" phylum="Tracheophyta" rank="species" species="liesneri">
<emphasis italics="true" pageId="9" pageNumber="96">Duckesia liesneri</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="9" pageNumber="96" rowspan="1">
<taxonomicName authorityName="Cuatrec" authorityYear="1961" baseAuthorityName="Huber" class="Magnoliopsida" family="Humiriaceae" genus="Endopleura" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endopleura uchi" order="Malpighiales" pageId="9" pageNumber="96" phylum="Tracheophyta" rank="species" species="uchi">
<emphasis italics="true" pageId="9" pageNumber="96">Endopleura uchi</emphasis>
</taxonomicName>
</th>
</tr>
<tr pageId="9" pageNumber="96">
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Habit</td>
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Tree, 12-30 m (27-67 cm dbh)</td>
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Shrub to small tree, 2-4 m</td>
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Tree, (12)20-40 m (30-70 dbh)</td>
</tr>
<tr pageId="9" pageNumber="96">
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Leaves</td>
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">
Petiole to 1 mm long; blade 5.5-7.5
<normalizedToken originalValue="×">x</normalizedToken>
1.7-2.3 cm, lanceolate, subcoriaceous; teeth setae robust, to 0.2 mm long, sometimes retained at maturity; abaxial glands sparse, 0-3 per leaf, 2/3 exmedial distance
</td>
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">
Petiole to 2 mm long; blade 3.5-4.5
<normalizedToken originalValue="×">x</normalizedToken>
2.2-2.7 cm, elliptic to obovate, coriaceous; teeth setae delicate, to 0.3 mm long, rapidly deciduous; abaxial glands absent
</td>
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">
Petiole 10-30 mm long; blade 17-28
<normalizedToken originalValue="×">x</normalizedToken>
5.5-6.5 cm, oblong to narrowly elliptic, subcoriaceous; teeth setae delicate, to 0.5 mm long, rapidly deciduous; abaxial glands in row near margin
</td>
</tr>
<tr pageId="9" pageNumber="96">
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Inflorescence</td>
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Axillary, small, to 1.5 cm long; bracts persistent</td>
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Axillary and subterminal, to 2.5 cm long; bracts deciduous</td>
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Axillary, to 6.5 cm long; bracts persistent</td>
</tr>
<tr pageId="9" pageNumber="96">
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Flower</td>
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Calyx glabrous with minutely hispid margin; petals glabrous</td>
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Calyx hispid; petals coarsely hirsute</td>
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Calyx centrally coarsely hirsute, margin minutely hispid; petals coarsely hirsute</td>
</tr>
<tr pageId="9" pageNumber="96">
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Androecium</td>
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Filaments papillose, very short-connate; anthers 5 tetrasporangiate and 15+ sterile</td>
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Filaments smooth, connate up to 1/3 length; anthers 20 tetrasporangiate</td>
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Filaments minutely papillose, very short-connate; anthers 10 tetrasporangiate and 10+ disporangiate</td>
</tr>
<tr pageId="9" pageNumber="96">
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Gynoecium</td>
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Pistil longer than ovary</td>
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Pistil shorter than ovary</td>
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Pistil as long as ovary</td>
</tr>
<tr pageId="9" pageNumber="96">
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Fruit</td>
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Ovoid; endocarp corrugated, 13+-radiate with densely packed woody ribs; resinous cavities small; valves conspicuous</td>
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Unknown</td>
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Ellipsoid; endocarp star-shaped, 10-radiate with 5 divided woody ribs; resinous cavities absent; valves inconspicuous</td>
</tr>
<tr pageId="9" pageNumber="96">
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">Distribution &amp; ecology</td>
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">
Brazil (Amazonas especially near Manaus,
<normalizedToken originalValue="Pará">Para</normalizedToken>
); 50-125 m elevation, terra firme
</td>
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">
Venezuela (Amazonas on Cerro Aracamuni and Sierra de
<normalizedToken originalValue="Unturán">Unturan</normalizedToken>
); 1150-1400 m elevation, upland cloud or semi-open tepui forest
</td>
<td colspan="1" pageId="9" pageNumber="96" rowspan="1">
Bolivia (Pando), Brazil (
<normalizedToken originalValue="Amapá">Amapa</normalizedToken>
, Amazonas,
<normalizedToken originalValue="Pará">Para</normalizedToken>
,
<normalizedToken originalValue="Rhondônia">Rhondonia</normalizedToken>
) Guyana, Suriname, Venezuela; 50-620 m elevation, terra firme
</td>
</tr>
</table>
</paragraph>
<caption ID-Zenodo-Dep="3262016" doi="10.3897/phytokeys.124.34679.figure6" httpUri="https://binary.pensoft.net/fig/311892" pageId="9" pageNumber="96" start="Figure 6" startId="F6">
<paragraph pageId="9" pageNumber="96">
<emphasis bold="true" pageId="9" pageNumber="96">Figure 6.</emphasis>
Macromorphology of
<taxonomicName authorityName="K. Wurdack &amp; C. E. Zartman" authorityYear="2019" class="Magnoliopsida" family="Humiriaceae" genus="Sacoglottis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sacoglottis perryi" order="Malpighiales" pageId="9" pageNumber="96" phylum="Tracheophyta" rank="species" species="perryi">
<emphasis italics="true" pageId="9" pageNumber="96">Sacoglottis perryi</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="9" pageNumber="96">A</emphasis>
Dried fruit with central seed embedded in woody endocarp, longitudinal split along carpel wall (dissected by Cuatrecasas)
<emphasis bold="true" pageId="9" pageNumber="96">B</emphasis>
dried fruit with 2 central seeds and endocarp lacunae, transverse section
<emphasis bold="true" pageId="9" pageNumber="96">C</emphasis>
fresh fruit with fleshy yellow-green exocarp and liquid in endocarp lacunae, transverse section
<emphasis bold="true" pageId="9" pageNumber="96">D</emphasis>
young inflorescence with bracts intact (b) or fallen leaving bract scars (bs)
<emphasis bold="true" pageId="9" pageNumber="96">E</emphasis>
mature bud with marginal sepal gland (small red dot in center)
<emphasis bold="true" pageId="9" pageNumber="96">F</emphasis>
partly open flower with intact anthers
<emphasis bold="true" pageId="9" pageNumber="96">G</emphasis>
post-anthetic flower
<emphasis bold="true" pageId="9" pageNumber="96">H</emphasis>
freshly cut trunk
<emphasis bold="true" pageId="9" pageNumber="96">I</emphasis>
type in life just before pressing. Sources:
<emphasis bold="true" pageId="9" pageNumber="96">A</emphasis>
<emphasis italics="true" pageId="9" pageNumber="96">Gillespie 2810</emphasis>
<emphasis bold="true" pageId="9" pageNumber="96">B</emphasis>
<emphasis italics="true" pageId="9" pageNumber="96">Tripp 2984</emphasis>
<emphasis bold="true" pageId="9" pageNumber="96">C, E-I</emphasis>
<emphasis italics="true" pageId="9" pageNumber="96">Redden 7264</emphasis>
<emphasis bold="true" pageId="9" pageNumber="96">D</emphasis>
<emphasis italics="true" pageId="9" pageNumber="96">Hoffman 1600</emphasis>
(all US).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>