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<document id="CEF85948F574C5DFC6091AFC003502C9" ID-DOI="10.5281/zenodo.274673" ID-GBIF-Dataset="74bce358-34e4-424f-a9ec-c0493249ed99" ID-ISSN="1175-5326" ID-Zenodo-Dep="274673" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459999065726" checkinUser="plazi" docAuthor="Sharaf, Mostafa R., Taylor, Brian &amp; Klingenberg, Christiana" docDate="2009" docId="62248797FFE1A155FF53F8FDA2FA67CF" docLanguage="en" docName="zt02004p058.pdf" docOrigin="Zootaxa 2004" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Solenopsis cooperi Donisthorpe 1947" docType="treatment" docVersion="9" lastPageNumber="54" masterDocId="9E1DFFEFFFE3A150FFC4FFDFA17E6628" masterDocTitle="Ants of the Genus SOLENOPSIS Westwood, 1840 (Hymenoptera: Formicidae) in Egypt with a description of the worker castes of S. cooperi Donisthorpe, 1947" masterLastPageNumber="58" masterPageNumber="49" pageNumber="51" updateTime="1698233054908" updateUser="plazi">
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<mods:title id="3E60E090AACABE95F993A44FD2657E74">Ants of the Genus SOLENOPSIS Westwood, 1840 (Hymenoptera: Formicidae) in Egypt with a description of the worker castes of S. cooperi Donisthorpe, 1947</mods:title>
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<mods:namePart id="43756CA5F65AD5AD18A9F2F67E971586">Sharaf, Mostafa R.</mods:namePart>
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<treatment id="62248797FFE1A155FF53F8FDA2FA67CF" ID-DOI="http://doi.org/10.5281/zenodo.6219369" ID-GBIF-Taxon="119380618" ID-Zenodo-Dep="6219369" LSID="urn:lsid:plazi:treatment:62248797FFE1A155FF53F8FDA2FA67CF" httpUri="http://treatment.plazi.org/id/62248797FFE1A155FF53F8FDA2FA67CF" lastPageId="5" lastPageNumber="54" pageId="2" pageNumber="51">
<subSubSection id="A297650AFFE1A152FF53F8FDA1846177" pageId="2" pageNumber="51" type="nomenclature">
<paragraph id="EA323681FFE1A152FF53F8FDA31D6114" blockId="2.[151,611,1826,1887]" box="[151,611,1826,1852]" pageId="2" pageNumber="51">
<heading id="B17A81EDFFE1A152FF53F8FDA31D6114" bold="true" box="[151,611,1826,1852]" fontSize="11" level="1" pageId="2" pageNumber="51" reason="1">
<taxonomicName id="2D8D4D02FFE1A152FF53F8FDA31D6114" ID-CoL="4Y6D6" authority="Donisthorpe, 1947" authorityName="Donisthorpe" authorityYear="1947" box="[151,611,1826,1852]" class="Insecta" family="Formicidae" genus="Solenopsis" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="cooperi">
<emphasis id="D8F9EA93FFE1A152FF53F8FDA31D6114" bold="true" box="[151,611,1826,1852]" pageId="2" pageNumber="51">
<emphasis id="D8F9EA93FFE1A152FF53F8FDA0076114" bold="true" box="[151,377,1826,1852]" italics="true" pageId="2" pageNumber="51">Solenopsis cooperi</emphasis>
<bibRefCitation id="8E1C4B70FFE1A152FE44F8FDA31D6114" author="Donisthorpe" box="[384,611,1826,1852]" pageId="2" pageNumber="57" refString="Donisthorpe, H. (1947) Results of the Armstrong college expedition to Siwa Oasis (Libyan desert) 1935: A second list of Formicidae. Bulletin de la Societe Fouad Ier d'Entomologie, 31, 109 - 111." type="journal article" year="1947">Donisthorpe, 1947</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="EA323681FFE1A152FF53F89AA1846177" blockId="2.[151,611,1826,1887]" box="[151,250,1861,1887]" pageId="2" pageNumber="51">(Plate 1)</paragraph>
</subSubSection>
<subSubSection id="A297650AFFE1A153FF53F84DA5536687" lastPageId="3" lastPageNumber="52" pageId="2" pageNumber="51" type="reference_group">
<paragraph id="EA323681FFE1A152FF53F84DA39861E1" blockId="2.[151,1437,1938,2025]" pageId="2" pageNumber="51">
<treatmentCitationGroup id="CA9D11AFFFE1A152FF53F84DA39861E1" pageId="2" pageNumber="51">
<taxonomicName id="2D8D4D02FFE1A152FF53F84DA3016181" ID-CoL="4Y6D6" authority="Donisthorpe, 1947: 110" authorityName="Donisthorpe" authorityPageNumber="110" authorityYear="1947" box="[151,639,1938,1961]" class="Insecta" family="Formicidae" genus="Solenopsis" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="cooperi">
<emphasis id="D8F9EA93FFE1A152FF53F84DA0166181" box="[151,360,1938,1961]" italics="true" pageId="2" pageNumber="51">Solenopsis cooperi</emphasis>
<treatmentCitation id="6B2C1090FFE1A152FEB0F84DA3016181" author="Donisthorpe" box="[372,639,1938,1961]" page="110" pageId="2" pageNumber="51" year="1947">
<bibRefCitation id="8E1C4B70FFE1A152FEB0F84DA33A6181" author="Donisthorpe" box="[372,580,1938,1961]" pageId="2" pageNumber="57" refString="Donisthorpe, H. (1947) Results of the Armstrong college expedition to Siwa Oasis (Libyan desert) 1935: A second list of Formicidae. Bulletin de la Societe Fouad Ier d'Entomologie, 31, 109 - 111." type="journal article" year="1947">Donisthorpe, 1947</bibRefCitation>
: 110
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="35368823FFE1A152FD4BF84DA38C6181" box="[655,754,1938,1961]" pageId="2" pageNumber="51" type="holotype">Holotype</typeStatus>
gyne and
<typeStatus id="35368823FFE1A152FCB0F84DA2AE6181" box="[884,976,1938,1961]" pageId="2" pageNumber="51" type="paratype">paratype</typeStatus>
gynes and males,
<collectingCountry id="929A7611FFE1A152FB69F84DA4786181" box="[1197,1286,1938,1961]" name="Egypt" pageId="2" pageNumber="51">EGYPT</collectingCountry>
: Siwa, 15 &amp;
<date id="9E331041FFE1A152FF03F86DA04E61E1" box="[199,304,1970,1993]" pageId="2" pageNumber="51" value="1935-05-24">24.v.1935</date>
(
<emphasis id="D8F9EA93FFE1A152FEFBF86DA09361E1" box="[319,493,1970,1993]" italics="true" pageId="2" pageNumber="51">J. Omer-Cooper</emphasis>
) (BMNH) [examined].
</treatmentCitationGroup>
</paragraph>
<paragraph id="EA323681FFE1A153FF53F80DA5536687" blockId="2.[151,1437,1938,2025]" lastBlockId="3.[199,1069,151,175]" lastPageId="3" lastPageNumber="52" pageId="2" pageNumber="51">
<taxonomicName id="2D8D4D02FFE1A152FF53F80DA0E961C1" authority="Sharaf" authorityName="Sharaf" box="[151,407,2002,2025]" class="Insecta" family="Formicidae" genus="Solenopsis" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="bakri">
<emphasis id="D8F9EA93FFE1A152FF53F80DA03461C1" box="[151,330,2002,2025]" italics="true" pageId="2" pageNumber="51">Solenopsis bakri</emphasis>
Sharaf
</taxonomicName>
<emphasis id="D8F9EA93FFE1A152FE5BF80DA0CD61C1" box="[415,435,2002,2025]" italics="true" pageId="2" pageNumber="51">in</emphasis>
<bibRefCitation id="8E1C4B70FFE1A152FE78F80DA34A61C1" author="Fadl" box="[444,564,2002,2025]" pageId="2" pageNumber="58" refString="Fadl, H. H., Bakr, R. F., Badawy, R. M. &amp; Sharaf, M. R. (2007) Six new species of ants (Insecta: Hymenoptera: Formicidae) from Egypt. Proceedings of the second International Conference of the Entomological Society of Egypt. 1, 235 - 249" type="journal article" year="2007">Fadl, 2007</bibRefCitation>
: 242.
<typeStatus id="35368823FFE1A152FDB2F80DA3A761C1" box="[630,729,2002,2025]" pageId="2" pageNumber="51" type="holotype">Holotype</typeStatus>
worker, one alate gyne, one male and
<typeStatus id="35368823FFE1A152FB44F80DA5AD61C1" box="[1152,1235,2002,2025]" pageId="2" pageNumber="51" type="syntype">syntype</typeStatus>
workers,
<collectingCountry id="929A7611FFE1A152FA84F80DA4E761C1" box="[1344,1433,2002,2025]" name="Egypt" pageId="2" pageNumber="51">EGYPT</collectingCountry>
: Saloga Islands, Aswan,
<date id="9E331041FFE0A153FE02FF47A3556687" box="[454,555,152,175]" pageId="3" pageNumber="52" value="2002-06-02">2.vi.2002</date>
(
<emphasis id="D8F9EA93FFE0A153FDFEFF47A3836687" box="[570,765,152,175]" italics="true" pageId="3" pageNumber="52">Mostafa R. Sharaf</emphasis>
) (EESC) [examined].
<emphasis id="D8F9EA93FFE0A153FC29FF48A5536687" bold="true" box="[1005,1069,151,175]" pageId="3" pageNumber="52">Syn.n</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="A297650AFFE0A154FF53FF01A3B96312" lastPageId="4" lastPageNumber="53" pageId="3" pageNumber="52" type="description">
<paragraph id="EA323681FFE0A153FF53FF01A50C6737" blockId="3.[151,1435,222,287]" pageId="3" pageNumber="52">
<emphasis id="D8F9EA93FFE0A153FF53FF01A18966D0" bold="true" box="[151,247,222,248]" pageId="3" pageNumber="52">Worker</emphasis>
. TL 1.531.76, HL 0.410.56, HW 0.340.45, SL 0.250.35, EL 0.02, AL 0.420.62, PL 0.120.21, PW 0.120.17, PPL 0.100.14, PPW 0.080.17, SI 7377, CI 8083 (12 measured).
</paragraph>
<paragraph id="EA323681FFE0A153FF53F8AEA47161E6" blockId="3.[151,1436,1905,2036]" pageId="3" pageNumber="52">
<emphasis id="D8F9EA93FFE0A153FF53F8AEA07461A1" bold="true" box="[151,266,1905,1929]" pageId="3" pageNumber="52">PLATE 1.</emphasis>
<taxonomicName id="2D8D4D02FFE0A153FED5F8ADA0A061A1" box="[273,478,1906,1929]" class="Insecta" family="Formicidae" genus="Solenopsis" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="cooperi">
<emphasis id="D8F9EA93FFE0A153FED5F8ADA0A061A1" box="[273,478,1906,1929]" italics="true" pageId="3" pageNumber="52">Solenopsis cooperi</emphasis>
</taxonomicName>
(ac) small worker, (a) head in full-face view, (b) body in profile, (c) dorsal view; (df) large worker, (d) head in full-face view, (e) body in profile, (f) dorsal view; (gi) alate gyne, (g) head in full-face view, (h) body in profile, (i) dorsal view; (jl) male, (j) head in full-face view, (k) body in profile, (l) dorsal view.
</paragraph>
<paragraph id="EA323681FFE0A154FF02F805A03C670E" blockId="3.[151,1436,1905,2036]" lastBlockId="4.[151,1438,152,2034]" lastPageId="4" lastPageNumber="53" pageId="3" pageNumber="52">
The workers examined could be divided into two size groups, suggesting dimorphism. Worker dimorphism has not been reported for Egyptian
<taxonomicName id="2D8D4D02FFE7A154FD01FF47A23C6699" box="[709,834,152,177]" class="Insecta" family="Formicidae" genus="Solenopsis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="D8F9EA93FFE7A154FD01FF47A23C6699" box="[709,834,152,177]" italics="true" pageId="4" pageNumber="53">Solenopsis</emphasis>
</taxonomicName>
, but it is known for
<taxonomicName id="2D8D4D02FFE7A154FBE9FF47A5E66699" box="[1069,1176,152,177]" class="Insecta" family="Formicidae" genus="Solenopsis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="orbula">
<emphasis id="D8F9EA93FFE7A154FBE9FF47A5E66699" box="[1069,1176,152,177]" italics="true" pageId="4" pageNumber="53">S. orbula</emphasis>
</taxonomicName>
from
<collectingCountry id="929A7611FFE7A154FB1AFF47A446669A" box="[1246,1336,152,178]" name="France" pageId="4" pageNumber="53">Corsica</collectingCountry>
(
<bibRefCitation id="8E1C4B70FFE7A154FA83FF47A1A966F0" author="Emery" pageId="4" pageNumber="58" refString="Emery, C. (1916) Fauna Entomologica Italiana. 1. Hymenoptera. Formicidae. Bulletino della Societa Entomologica Italiana, 47, 79 - 275." type="journal article" year="1916">Emery, 1916</bibRefCitation>
). Both major and minor workers were collected from a nest in association with gynes and males. Both groups of workers have a similar head shape, especially the flattened profile. The morphological differences are as follows:
</paragraph>
<paragraph id="EA323681FFE7A154FF02FEEDA58164AA" blockId="4.[151,1438,152,2034]" pageId="4" pageNumber="53">
<emphasis id="D8F9EA93FFE7A154FF02FEEDA0046764" bold="true" box="[198,378,306,332]" pageId="4" pageNumber="53">Minor worker</emphasis>
(Plate 1, Fig. a, b. c). Head relatively long and narrow, with a distinctive narrow profile; CI ca. 78. Mandibles quite narrow with only moderate teeth, the apical tooth somewhat elongated. Clypeus with a weakly impressed anterior margin and distinct but not sharp carinae. Frontal lobes set quite far apart. Frontal notch distinct. SI ca. 80. Head smooth and shiny with very minute hair pits. Promesonotum with a smooth profile, anteriorly rounded but posteriorly flat. Metanotal groove distinct, quite shallow and continuous dorsally. Propodeum with a distinctive dorsal longitudinal impression and posteriorly terminating in angular corners. Propodeal spiracles vertically ovoid and facing posteriorly. Petiole anterior peduncle short and narrow; viewed from above this is quite distinct. Petiole node quite high and narrow. Body with relatively few scattered short hairs. Overall colour dusky yellow, with the posterior of the gaster darkened.
</paragraph>
<paragraph id="EA323681FFE7A154FF02FD51A3D865B8" blockId="4.[151,1438,152,2034]" pageId="4" pageNumber="53">
<emphasis id="D8F9EA93FFE7A154FF02FD51A0076480" bold="true" box="[198,377,654,680]" pageId="4" pageNumber="53">Major worker</emphasis>
(Plate 1, Fig. d, e, f). Head smooth with widely scattered but distinct hair pits; CI ca. 80. Mandibles narrow. Clypeus with only a weakly impressed anterior margin and very weak carinae. Frontal lobes set moderately far apart. Scape with a narrow base broadening after the mid-point and widest at about the two-thirds point, SI ca. 70. Promesonotum profile smooth and uninterrupted, almost flat posterior to the anterior convex half of the pronotum. Metanotal groove distinctly impressed. Propodeum with a small circular spiracle. Petiole node quite low with a domed profile. Gaster distinctly lighter than rest of the body. Overall with sparse, fine, relatively short, erect hairs.
</paragraph>
<paragraph id="EA323681FFE7A154FF02FC42A1896203" blockId="4.[151,1438,152,2034]" pageId="4" pageNumber="53">The workers are readily distinguished from the known circum-Mediterranean species by the long peduncle to the petiole; the relatively high, narrowly rounded petiole profile; the proportionally longer head which is flattened in lateral view; and the angular, near dentate and longitudinally weakly furrowed propodeal dorsum.</paragraph>
<paragraph id="EA323681FFE7A154FF02FBE8A0476250" blockId="4.[151,1438,152,2034]" pageId="4" pageNumber="53">
<emphasis id="D8F9EA93FFE7A154FF02FBE8A0776279" bold="true" box="[198,265,1079,1105]" pageId="4" pageNumber="53">Gyne</emphasis>
(Plate 1, Figs g, h, i).
<typeStatus id="35368823FFE7A154FDD4FBE7A305627A" box="[528,635,1080,1106]" pageId="4" pageNumber="53" type="paratype">Paratype</typeStatus>
. TL 4.0, HL 0.75, HW 0.55, SL 0.50, EL 0.15, AL 1.0, SI 91, CI 73 (1 measured).
</paragraph>
<paragraph id="EA323681FFE7A154FF02FB5AA0A462EE" blockId="4.[151,1438,152,2034]" pageId="4" pageNumber="53">Additional material: TL 4.504.75; HL 0.75; HW 0.570.60; SL 0.470.475; EL 0.170.20; AL 1.15; SI 7883; CI 79 (2 measured).</paragraph>
<paragraph id="EA323681FFE7A154FF02FB0CA5A062C5" blockId="4.[151,1438,152,2034]" box="[198,1246,1235,1261]" pageId="4" pageNumber="53">The large ocelli are very distinctive compared to the other circum-Mediterranean species.</paragraph>
<paragraph id="EA323681FFE7A154FF02FB27A3B96312" blockId="4.[151,1438,152,2034]" pageId="4" pageNumber="53">
<emphasis id="D8F9EA93FFE7A154FF02FB27A078633A" bold="true" box="[198,262,1272,1298]" pageId="4" pageNumber="53">Male</emphasis>
(Plate 1, Figs j, k, l).
<typeStatus id="35368823FFE7A154FDE6FB26A3E6633B" box="[546,664,1273,1299]" pageId="4" pageNumber="53" type="paratype">Paratypes</typeStatus>
. TL 3.0, HL 0.350.45, HW 0.400.45, SL 0.10, EL 0.25, AL 1.101.25, SI 2225, CI 100114 (3 measured).
</paragraph>
</subSubSection>
<subSubSection id="A297650AFFE7A154FF02FA99A5E163D3" pageId="4" pageNumber="53" type="materials_examined">
<paragraph id="EA323681FFE7A154FF02FA99A3C763AF" blockId="4.[151,1438,152,2034]" pageId="4" pageNumber="53">
<emphasis id="D8F9EA93FFE7A154FF02FA99A0B76348" bold="true" box="[198,457,1350,1376]" pageId="4" pageNumber="53">Additional material.</emphasis>
TL 3.03.6, HL 0.450.50, HW 0.400.52, SL 0.0750.20, EL 0.170.25, AL 1.051.22, SI 2143, CI 89104 (7 measured).
</paragraph>
<paragraph id="EA323681FFE7A154FF02FA4BA5E163D3" blockId="4.[151,1438,152,2034]" pageId="4" pageNumber="53">
The newly collected specimens appear to match the rather short description of
<taxonomicName id="2D8D4D02FFE7A154FB62FA4BA45D6385" box="[1190,1315,1428,1453]" class="Insecta" family="Formicidae" genus="Solenopsis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="cooperi">
<emphasis id="D8F9EA93FFE7A154FB62FA4BA45D6385" box="[1190,1315,1428,1453]" italics="true" pageId="4" pageNumber="53">S. cooperi</emphasis>
</taxonomicName>
given by
<bibRefCitation id="8E1C4B70FFE7A154FF53FA64A0F663FD" author="Donisthorpe" box="[151,392,1467,1493]" pageId="4" pageNumber="57" refString="Donisthorpe, H. (1947) Results of the Armstrong college expedition to Siwa Oasis (Libyan desert) 1935: A second list of Formicidae. Bulletin de la Societe Fouad Ier d'Entomologie, 31, 109 - 111." type="journal article" year="1947">Donisthorpe (1947)</bibRefCitation>
, i.e. notably being brown, smooth &amp; shining, with pale yellow appendages. The large ocelli also appear to be distinctive compared to the other circum-Mediterranean species.
</paragraph>
</subSubSection>
<subSubSection id="A297650AFFE7A155FF01F9D7A08366D7" lastPageId="5" lastPageNumber="54" pageId="4" pageNumber="53" type="reference_group">
<paragraph id="EA323681FFE7A154FF01F9D7A2A8617F" blockId="4.[151,1438,152,2034]" pageId="4" pageNumber="53">
<taxonomicName id="2D8D4D02FFE7A154FF01F9D7A0DA6009" ID-CoL="4Y6D6" box="[197,420,1544,1569]" class="Insecta" family="Formicidae" genus="Solenopsis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="cooperi">
<emphasis id="D8F9EA93FFE7A154FF01F9D7A0DA6009" box="[197,420,1544,1569]" italics="true" pageId="4" pageNumber="53">Solenopsis cooperi</emphasis>
</taxonomicName>
has remained little known since the description by
<bibRefCitation id="8E1C4B70FFE7A154FBC1F9D7A593600A" author="Donisthorpe" box="[1029,1261,1544,1570]" pageId="4" pageNumber="57" refString="Donisthorpe, H. (1947) Results of the Armstrong college expedition to Siwa Oasis (Libyan desert) 1935: A second list of Formicidae. Bulletin de la Societe Fouad Ier d'Entomologie, 31, 109 - 111." type="journal article" year="1947">Donisthorpe (1947)</bibRefCitation>
when only the gynes and males were collected from Siwa Oasis in the western desert (previously known as the Libyan Desert). It appears to be restricted to
<collectingCountry id="929A7611FFE7A154FD9DF98AA3DA6047" box="[601,676,1621,1647]" name="Egypt" pageId="4" pageNumber="53">Egypt</collectingCountry>
. During two field trips carried out by the first author to Saloga Island Protectorate (located in the River Nile in Aswan) and to Abu-Swelam village (El-Minyia Governorate, Upper
<collectingCountry id="929A7611FFE7A154FF2CF97CA04C6095" box="[232,306,1699,1725]" name="Egypt" pageId="4" pageNumber="53">Egypt</collectingCountry>
), many workers, males and two alate gynes were collected. These were mistakenly described as a new species by Sharaf
<emphasis id="D8F9EA93FFE7A154FE0CF916A09E60CA" box="[456,480,1737,1762]" italics="true" pageId="4" pageNumber="53">in</emphasis>
<bibRefCitation id="8E1C4B70FFE7A154FE2CF916A30360CB" author="Fadl" box="[488,637,1737,1763]" pageId="4" pageNumber="58" refString="Fadl, H. H., Bakr, R. F., Badawy, R. M. &amp; Sharaf, M. R. (2007) Six new species of ants (Insecta: Hymenoptera: Formicidae) from Egypt. Proceedings of the second International Conference of the Entomological Society of Egypt. 1, 235 - 249" type="journal article" year="2007">Fadl (2007)</bibRefCitation>
. By comparing the sexual castes collected with the
<typeStatus id="35368823FFE7A154FB3FF916A45160CB" box="[1275,1327,1737,1763]" pageId="4" pageNumber="53">type</typeStatus>
material deposited in the ant collection of the British Museum of Natural History, we have confidence in our description of the associated workers and our further notes on the sexuals. Thus, the worker caste of
<taxonomicName id="2D8D4D02FFE7A154FAE2F8C8A4E26118" ID-CoL="4Y6D6" box="[1318,1436,1815,1840]" class="Insecta" family="Formicidae" genus="Solenopsis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="cooperi">
<emphasis id="D8F9EA93FFE7A154FAE2F8C8A4E26118" box="[1318,1436,1815,1840]" italics="true" pageId="4" pageNumber="53">S. cooperi</emphasis>
</taxonomicName>
is described for the first time and measurements of all castes are given.
</paragraph>
<paragraph id="EA323681FFE7A155FF02F8BBA08366D7" blockId="4.[151,1438,152,2034]" lastBlockId="5.[151,1436,152,487]" lastPageId="5" lastPageNumber="54" pageId="4" pageNumber="53">
Very little is known of the biology of
<taxonomicName id="2D8D4D02FFE7A154FD49F8BBA2786155" ID-CoL="4Y6D6" box="[653,774,1892,1917]" class="Insecta" family="Formicidae" genus="Solenopsis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="cooperi">
<emphasis id="D8F9EA93FFE7A154FD49F8BBA2786155" box="[653,774,1892,1917]" italics="true" pageId="4" pageNumber="53">S. cooperi</emphasis>
</taxonomicName>
, except for the colony from which our specimens were collected. This colony was small and in loose sandy soil at the base of an old tree. It is likely the species has a broad distribution and may be present at many other localities in
<collectingCountry id="929A7611FFE7A154FC55F86EA2A961E3" box="[913,983,1969,1995]" name="Egypt" pageId="4" pageNumber="53">Egypt</collectingCountry>
or North Africa, especially in the Nile Valley and its islands, because the abundance of transport on the Nile ensures no place can be considered isolated. In addition, a single worker specimen was collected from Abu-Swelam village in the Nile valley far from Aswan. The original finding (
<bibRefCitation id="8E1C4B70FFE6A155FD83FF61A25166F0" author="Donisthorpe" box="[583,815,190,216]" pageId="5" pageNumber="57" refString="Donisthorpe, H. (1942) The Formicidae of the Armstrong College expedition to the Siwa Oasis. Annals and Magazine of Natural History, (11) 9, 26 - 33." type="journal article" year="1942">Donisthorpe, 1942</bibRefCitation>
) at Siwa Oasis also is a long way from the Aswan location of the second finding.
</paragraph>
</subSubSection>
<subSubSection id="A297650AFFE6A155FF02FED4A2FA67CF" pageId="5" pageNumber="54" type="materials_examined">
<paragraph id="EA323681FFE6A155FF02FED4A2C267B2" blockId="5.[151,1436,152,487]" pageId="5" pageNumber="54">
<emphasis id="D8F9EA93FFE6A155FF02FED4A0C3670D" bold="true" box="[198,445,267,293]" pageId="5" pageNumber="54">Material examined.</emphasis>
<specimenCount id="FC8BFD08FFE6A155FE03FED3A331670E" box="[455,591,268,294]" pageId="5" pageNumber="54" type="worker">32 workers</specimenCount>
, 2 ♀,
<specimenCount id="FC8BFD08FFE6A155FD5AFED3A3B3670E" box="[670,717,268,294]" pageId="5" pageNumber="54" type="generic">7 ♂</specimenCount>
;
<collectingCountry id="929A7611FFE6A155FD1BFED3A256670E" box="[735,808,268,294]" name="Egypt" pageId="5" pageNumber="54">Egypt</collectingCountry>
, Saloga Islands Protectorate, Aswan (Upper
<collectingCountry id="929A7611FFE6A155FA81FED3A4F1670E" box="[1349,1423,268,294]" name="Egypt" pageId="5" pageNumber="54">Egypt</collectingCountry>
),
<date id="9E331041FFE6A155FF53FEEDA1816764" box="[151,255,306,332]" pageId="5" pageNumber="54" value="2002-05-02">2.v.2002</date>
;
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;
<geoCoordinate id="8FB95046FFE6A155FE40FEEDA0976764" box="[388,489,306,332]" direction="east" orientation="longitude" pageId="5" pageNumber="54" precision="555" value="32.56">32.56º E</geoCoordinate>
(
<emphasis id="D8F9EA93FFE6A155FE3FFEEDA3EC6763" box="[507,658,306,331]" italics="true" pageId="5" pageNumber="54">M. R. Sharaf</emphasis>
);
<specimenCount id="FC8BFD08FFE6A155FD68FEEDA2666764" box="[684,792,306,332]" pageId="5" pageNumber="54" type="worker">1 worker</specimenCount>
, Abu-Swelam (El-Minyia Governorate, Upper
<collectingCountry id="929A7611FFE6A155FA81FEEDA4F16764" box="[1349,1423,306,332]" name="Egypt" pageId="5" pageNumber="54">Egypt</collectingCountry>
),
<date id="9E331041FFE6A155FF53FE86A066675B" box="[151,280,345,371]" pageId="5" pageNumber="54" value="2003-06-29">29.vi.2003</date>
;
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;
<geoCoordinate id="8FB95046FFE6A155FE5BFE86A37B675B" box="[415,517,345,371]" direction="east" orientation="longitude" pageId="5" pageNumber="54" precision="555" value="30.45">30.45º E</geoCoordinate>
<emphasis id="D8F9EA93FFE6A155FDC8FE86A3C7675A" box="[524,697,345,370]" italics="true" pageId="5" pageNumber="54">(M. R. Sharaf)</emphasis>
. Three workers were deposited in BTC; five will be placed in the
<collectionCode id="8C9CAE44FFE6A155FF06FE5FA07367B2" box="[194,269,384,410]" pageId="5" pageNumber="54">EESC</collectionCode>
; two will be at
<collectionCode id="8C9CAE44FFE6A155FE05FE5FA36867B2" box="[449,534,384,410]" httpUri="http://grbio.org/cool/tc3a-fmpc" name="Staatliches Museum fuer Naturkunde Karlsruhe (State Museum of Natural History)" pageId="5" pageNumber="54">SMNK</collectionCode>
and the remaining workers in SHC.
</paragraph>
<paragraph id="EA323681FFE6A155FF02FE79A2FA67CF" blockId="5.[151,1436,152,487]" pageId="5" pageNumber="54">
<emphasis id="D8F9EA93FFE6A155FF02FE79A0D567E8" bold="true" box="[198,427,422,448]" pageId="5" pageNumber="54">Previous records.</emphasis>
Burg El-Arab (Alexandria),
<date id="9E331041FFE6A155FCEAFE79A2D067E8" box="[814,942,422,448]" pageId="5" pageNumber="54" value="1956-03-20">20.iii.1956</date>
(
<emphasis id="D8F9EA93FFE6A155FC0FFE79A56567E8" bold="true" box="[971,1051,422,448]" pageId="5" pageNumber="54">
<collectionCode id="8C9CAE44FFE6A155FC0FFE79A56567E8" box="[971,1051,422,448]" pageId="5" pageNumber="54">ALFC</collectionCode>
</emphasis>
); Burg El-Arab (Alexandria),
<date id="9E331041FFE6A155FF53FE12A06867CF" box="[151,278,461,487]" pageId="5" pageNumber="54" value="1956-03-20">20.iii.1956</date>
(
<emphasis id="D8F9EA93FFE6A155FEE3FE12A00C67CF" bold="true" box="[295,370,461,487]" pageId="5" pageNumber="54">
<collectionCode id="8C9CAE44FFE6A155FEE3FE12A00C67CF" box="[295,370,461,487]" pageId="5" pageNumber="54">AINC</collectionCode>
</emphasis>
) (
<bibRefCitation id="8E1C4B70FFE6A155FE4FFE12A20667CF" author="Mohammed" box="[395,888,461,487]" pageId="5" pageNumber="58" refString="Mohammed, A. H. (1979) Taxonomic studies of family Formicidae (Hymenoptera) in A. R. Egypt. Faculty of Science, Ain Shams University, Entomology Dep. Cairo, Egypt. (Unpublished thesis), 288 pp." type="book" year="1979">Mohammed, 1979; based on sexuals only</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>