180 lines
18 KiB
XML
180 lines
18 KiB
XML
<document id="CB67887EC6B62A61A1778AAB22525298" ID-CLB-Dataset="29267" ID-DOI="10.11646/zootaxa.4496.1.17" ID-GBIF-Dataset="a161220f-7db6-4ccc-bfaa-de528dab984c" ID-ISSN="1175-5326" ID-Zenodo-Dep="1446761" ID-ZooBank="68B7C32E-BDEB-4C7C-A4F8-BC242DE24351" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538747124433" checkinUser="plazi" docAuthor="Nxele, Thembeka Clara, Plisko, Jadwiga Danuta & Ramongalo, Boipelo K." docDate="2018" docId="03F02941C429FFA9FF48FF7AFB61FBD5" docLanguage="en" docName="zootaxa.4496.1.17.pdf" docOrigin="Zootaxa 4496 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tritogenia Kinberg 1867" docType="treatment" docVersion="7" lastPageNumber="215" masterDocId="FFC95139C428FFA8FFDFFFA4FFA3FFC8" masterDocTitle="The first proandric species of Tritogenia Kinberg, 1867 from Botswana, Tritogenia talana sp. n. (Clitellata, Crassiclitellata, Tritogeniidae)" masterLastPageNumber="217" masterPageNumber="214" pageNumber="215" updateTime="1698565648559" updateUser="ExternalLinkService">
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<mods:title id="EF21DAB5EC7711013815728F0A22E403">The first proandric species of Tritogenia Kinberg, 1867 from Botswana, Tritogenia talana sp. n. (Clitellata, Crassiclitellata, Tritogeniidae)</mods:title>
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<mods:namePart id="7A794D2BF8BA5865EFC0ED5B53FEB664">Nxele, Thembeka Clara</mods:namePart>
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<mods:namePart id="7DCDC873F9FAABDE08D95AD1FFD92445">Plisko, Jadwiga Danuta</mods:namePart>
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<mods:namePart id="037002B85FD83E45B2F903667D3C5C78">Ramongalo, Boipelo K.</mods:namePart>
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<mods:title id="773AEE28433A74BB96CF270C2732955F">Zootaxa</mods:title>
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<mods:part id="EF2B5570BACFB06E60295BC7F5010FFE">
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<mods:date id="6803770DB6E97C56F3137F8D61A4A851">2018</mods:date>
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<mods:number id="E0B138D6E5802E5C48E6593B3D23DDBF">2018-10-05</mods:number>
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<mods:number id="FD74CA57518F19CE9A0BD505DA6D587F">4496</mods:number>
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<treatment id="03F02941C429FFA9FF48FF7AFB61FBD5" ID-DOI="http://doi.org/10.5281/zenodo.5968651" ID-GBIF-Taxon="148639810" ID-Zenodo-Dep="5968651" LSID="urn:lsid:plazi:treatment:03F02941C429FFA9FF48FF7AFB61FBD5" httpUri="http://treatment.plazi.org/id/03F02941C429FFA9FF48FF7AFB61FBD5" lastPageNumber="215" pageId="1" pageNumber="215">
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<subSubSection id="C343CBDCC429FFA9FF48FF7AFD86FF30" box="[151,549,222,248]" pageId="1" pageNumber="215" type="nomenclature">
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<paragraph id="8BE69857C429FFA9FF48FF7AFD86FF30" blockId="1.[151,549,222,248]" box="[151,549,222,248]" pageId="1" pageNumber="215">
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<heading id="D0AE2F3BC429FFA9FF48FF7AFD86FF30" bold="true" box="[151,549,222,248]" fontSize="11" level="1" pageId="1" pageNumber="215" reason="1">
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<emphasis id="B92D4445C429FFA9FF48FF7AFD86FF30" bold="true" box="[151,549,222,248]" pageId="1" pageNumber="215">
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Genus
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<taxonomicName id="4C59E3D4C429FFA9FF31FF7AFD86FF30" authority="Kinberg, 1867" authorityName="Kinberg" authorityYear="1867" box="[238,549,222,248]" class="Clitellata" family="Microchaetidae" genus="Tritogenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="215" phylum="Annelida" rank="genus">
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<emphasis id="B92D4445C429FFA9FF31FF7AFECFFF30" bold="true" box="[238,364,222,248]" italics="true" pageId="1" pageNumber="215">Tritogenia</emphasis>
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<bibRefCitation id="EFC8E5A6C429FFA9FEACFF7AFD86FF30" box="[371,549,222,248]" pageId="1" pageNumber="215" refId="ref2096" refString="Kinberg, J. H. G. (1867) Annulata nova. Ofversigt af Kongliga Vetenskaps-Akademiens Forhandlingar, 23, 97 - 103." type="journal article">Kinberg, 1867</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C343CBDCC429FFA9FF48FE81FA80FEB0" pageId="1" pageNumber="215" type="reference_group">
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<treatmentCitationGroup id="AB49BF79C429FFA9FF48FE81FE84FE92" pageId="1" pageNumber="215">
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<taxonomicName id="4C59E3D4C429FFA9FF48FE81FE4AFEF3" authority=": Benham 1888: 72" authorityName="Benham" authorityPageNumber="72" authorityYear="1888" box="[151,489,293,315]" class="Clitellata" family="Microchaetidae" genus="Brachydrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="215" phylum="Annelida" rank="genus">
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<emphasis id="B92D4445C429FFA9FF48FE81FEB8FEF3" box="[151,283,293,315]" italics="true" pageId="1" pageNumber="215">Brachydrilus</emphasis>
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:
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<treatmentCitation id="0AF8BE46C429FFA9FEF3FE81FE4AFEF3" author="" box="[300,489,293,315]" page="72" pageId="1" pageNumber="215">
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<bibRefCitation id="EFC8E5A6C429FFA9FEF3FE81FE63FEF3" box="[300,448,293,315]" pageId="1" pageNumber="215" refId="ref1955" refString="Benham, W. B. (1888) Note on a new earthworm. Zoologischer Anzeiger, 11, 72 - 75." type="journal article">Benham 1888</bibRefCitation>
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: 72
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</treatmentCitation>
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</taxonomicName>
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;
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<treatmentCitation id="0AF8BE46C429FFA9FE28FE81FD42FEF3" author="" box="[503,737,293,315]" page="462" pageId="1" pageNumber="215">
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<bibRefCitation id="EFC8E5A6C429FFA9FE28FE81FD08FEF3" box="[503,683,293,315]" pageId="1" pageNumber="215" refId="ref2122" refString="Michaelsen, W. (1900) Oligochaeta. In: Das Tierreich. Lief. 10. Verlag von R. Friedlander & Sohn, Berlin, pp. 1 - 575." type="book chapter">Michaelsen 1900</bibRefCitation>
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: 462
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</treatmentCitation>
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;
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<treatmentCitation id="0AF8BE46C429FFA9FD30FE81FC7AFEF3" author="" box="[751,985,293,315]" page="332" pageId="1" pageNumber="215">
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<bibRefCitation id="EFC8E5A6C429FFA9FD30FE81FC00FEF3" box="[751,931,293,315]" pageId="1" pageNumber="215" refId="ref2240" refString="Michaelsen, W. (1918) Die Lumbriciden mit besonderer Berucksichtigung der bisher als Familie Glossoscolecidae zusammengefassten Unterfamilien. Zoologische Jahrbucher, Abteilung fur Systematik, 41, 1 - 398." type="journal article">Michaelsen 1918</bibRefCitation>
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: 332
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||
</treatmentCitation>
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||
;
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<treatmentCitation id="0AF8BE46C429FFA9FC38FE81FB72FEF3" author="" box="[999,1233,293,315]" page="900" pageId="1" pageNumber="215">
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<bibRefCitation id="EFC8E5A6C429FFA9FC38FE81FB38FEF3" box="[999,1179,293,315]" pageId="1" pageNumber="215" refId="ref2797" refString="Stephenson, J. (1930) The Oligochaeta. Clarendon Press, Oxford, 978 pp." type="book">Stephenson 1930</bibRefCitation>
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: 900
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</treatmentCitation>
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;
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<treatmentCitation id="0AF8BE46C429FFA9FB01FE81FE80FE92" author="" page="77" pageId="1" pageNumber="215">
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<bibRefCitation id="EFC8E5A6C429FFA9FB01FE81FF5EFE92" pageId="1" pageNumber="215" refId="ref2688" refString="Reynolds, J. W. & Cook, D. G. (1976) Nomenclatura Oligochaetologica. A catalogue of names, descriptions and type specimens of the Oligochaeta. Runge Press, Ottawa, 217 pp." type="book">Reynolds & Cook 1976</bibRefCitation>
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: 77
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</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BE69857C429FFA9FF48FEC6FA80FEB0" blockId="1.[151,1436,293,407]" box="[151,1315,354,376]" pageId="1" pageNumber="215">
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<treatmentCitationGroup id="AB49BF79C429FFA9FF48FEC6FA80FEB0" box="[151,1315,354,376]" pageId="1" pageNumber="215">
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<taxonomicName id="4C59E3D4C429FFA9FF48FEC6FDA6FEB0" authority=": Michaelsen 1902: 33" authorityName="Michaelsen" authorityPageNumber="33" authorityYear="1902" box="[151,517,354,376]" class="Clitellata" family="Microchaetidae" genus="Microchaetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="215" phylum="Annelida" rank="genus">
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<emphasis id="B92D4445C429FFA9FF48FEC6FE82FEB0" box="[151,289,354,376]" italics="true" pageId="1" pageNumber="215">Microchaetus</emphasis>
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:
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<treatmentCitation id="0AF8BE46C429FFA9FEF1FEC6FDA6FEB0" author="" box="[302,517,354,376]" page="33" pageId="1" pageNumber="215">
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<bibRefCitation id="EFC8E5A6C429FFA9FEF1FEC6FE43FEB0" box="[302,480,354,376]" pageId="1" pageNumber="215" refId="ref2156" refString="Michaelsen, W. (1902) Neue Oligochaeten und neue Fundorte alt-bekannter. Mitteilungen aus dem Naturhistorischen Museum in Hamburg, 19, 1 - 54." type="journal article">Michaelsen 1902</bibRefCitation>
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: 33
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||
</treatmentCitation>
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</taxonomicName>
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||
;
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<treatmentCitation id="0AF8BE46C429FFA9FDCFFEC6FD79FEB0" author="" box="[528,730,354,376]" page="6" pageId="1" pageNumber="215">
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<bibRefCitation id="EFC8E5A6C429FFA9FDCFFEC6FD62FEB0" box="[528,705,354,376]" pageId="1" pageNumber="215" refId="ref2184" refString="Michaelsen, W. (1907) Oligochaeten von Natal und dem Zululand. Arkiv for Zoologi, 4 (4), 1 - 12." type="journal article">Michaelsen 1907</bibRefCitation>
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: 6
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</treatmentCitation>
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||
;
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<treatmentCitation id="0AF8BE46C429FFA9FD3AFEC6FC6AFEB0" author="" box="[741,969,354,376]" page="431" pageId="1" pageNumber="215">
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<bibRefCitation id="EFC8E5A6C429FFA9FD3AFEC6FC35FEB0" box="[741,918,354,376]" pageId="1" pageNumber="215" refId="ref2211" refString="Michaelsen, W. (1913) The Oligochaeta of Natal and Zululand. Annals of the Natal Museum, 2 (4), 397 - 457." type="journal article">Michaelsen 1913</bibRefCitation>
|
||
: 431
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||
</treatmentCitation>
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||
;
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||
<treatmentCitation id="0AF8BE46C429FFA9FC0CFEC6FB4EFEB0" author="" box="[979,1261,354,376]" page="77" pageId="1" pageNumber="215">
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<bibRefCitation id="EFC8E5A6C429FFA9FC0CFEC6FB64FEB0" box="[979,1223,354,376]" pageId="1" pageNumber="215" refId="ref2688" refString="Reynolds, J. W. & Cook, D. G. (1976) Nomenclatura Oligochaetologica. A catalogue of names, descriptions and type specimens of the Oligochaeta. Runge Press, Ottawa, 217 pp." type="book">Reynolds & Cook 1976</bibRefCitation>
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: 77
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||
</treatmentCitation>
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||
, 192.
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||
</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C343CBDCC429FFA9FF48FE25FD22FE5F" box="[151,641,385,407]" pageId="1" pageNumber="215" type="materials_examined">
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<paragraph id="8BE69857C429FFA9FF48FE25FD22FE5F" blockId="1.[151,1436,293,407]" box="[151,641,385,407]" pageId="1" pageNumber="215">
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<emphasis id="B92D4445C429FFA9FF48FE25FE81FE5F" bold="true" box="[151,290,385,407]" pageId="1" pageNumber="215">Type species.</emphasis>
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<taxonomicName id="4C59E3D4C429FFA9FEF6FE26FDDEFE5F" authority="Kinberg, 1867" authorityName="Kinberg" authorityYear="1867" box="[297,637,385,407]" class="Clitellata" family="Microchaetidae" genus="Tritogenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="215" phylum="Annelida" rank="species" species="sulcata">
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<emphasis id="B92D4445C429FFA9FEF6FE26FE41FE5F" box="[297,482,385,407]" italics="true" pageId="1" pageNumber="215">Tritogenia sulcata</emphasis>
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||
<bibRefCitation id="EFC8E5A6C429FFA9FE37FE25FDDEFE5F" box="[488,637,385,407]" pageId="1" pageNumber="215" refId="ref2096" refString="Kinberg, J. H. G. (1867) Annulata nova. Ofversigt af Kongliga Vetenskaps-Akademiens Forhandlingar, 23, 97 - 103." type="journal article">Kinberg, 1867</bibRefCitation>
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||
</taxonomicName>
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||
.
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C343CBDCC429FFA9FF48FE60FA92FDA4" pageId="1" pageNumber="215" type="diagnosis">
|
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<paragraph id="8BE69857C429FFA9FF48FE60FA92FDA4" blockId="1.[151,1437,452,1053]" pageId="1" pageNumber="215">
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||
<emphasis id="B92D4445C429FFA9FF48FE60FEB2FE15" bold="true" box="[151,273,452,477]" pageId="1" pageNumber="215">Diagnosis.</emphasis>
|
||
One oesophageal gizzard in 6–7. Four or five septa variably thickened, with thickened septa 6/7 attached externally to the gizzard. Dorsal blood vessel double in segments 4–10, 11, 12; double when crossing septa; single after segment 10, 11 or 12. Two or more small nephridia per segment (may be missing in some posterior segments). Testes and their funnels in the species earlier accredited to this genus in holandric arrangement, in presently described new species these are proandric. Description based on
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||
<bibRefCitation id="EFC8E5A6C429FFA9FB4DFDF0FA8FFDA4" box="[1170,1324,596,621]" pageId="1" pageNumber="215" refId="ref2543" refString="Plisko, J. D. (2013) A new family Tritogeniidae for the genera Tritogenia and Michalakus, earlier accredited to the composite Microchaetidae (Annelida: Oligochaeta). African Invertebrates, 54, 69 - 92. https: // doi. org / 10.5733 / afin. 054.0107" type="journal article">Plisko (2013)</bibRefCitation>
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||
.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C343CBDCC429FFA9FF18FDD3FB61FBD5" pageId="1" pageNumber="215" type="description">
|
||
<paragraph id="8BE69857C429FFA9FF18FDD3FB61FBD5" blockId="1.[151,1437,452,1053]" pageId="1" pageNumber="215">
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||
<emphasis id="B92D4445C429FFA9FF18FDD3FEFAFD58" bold="true" box="[199,345,631,656]" pageId="1" pageNumber="215">Description.</emphasis>
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||
Annulation of pre-clitellar segments present; 1–3 usually simple, 4–9, 10 ringleted, with 2–4 ringlets annulated or not, with an external segmental subdivision; first ringlet may be longer than second. Postclitellar segments may be externally subdivided, or not. Setae: minute and irregularly placed; on preclitellar segments, may be recognizable under high magnification (120–250×) and only as a few pairs of
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||
<emphasis id="B92D4445C429FFA9FAF3FD40FAEBFD35" box="[1324,1352,740,765]" italics="true" pageId="1" pageNumber="215">ab</emphasis>
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||
or
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<emphasis id="B92D4445C429FFA9FAA5FD40FA36FD35" box="[1402,1429,740,765]" italics="true" pageId="1" pageNumber="215">cd</emphasis>
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||
; sometimes
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||
<emphasis id="B92D4445C429FFA9FEC6FCA3FE96FCE8" box="[281,309,775,800]" italics="true" pageId="1" pageNumber="215">ab</emphasis>
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||
seen only on a few segments. As from segment 9,
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<emphasis id="B92D4445C429FFA9FC4FFCA3FC0FFCE8" box="[912,940,775,800]" italics="true" pageId="1" pageNumber="215">ab</emphasis>
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||
may be associated with papillae. On postclitellar segments, setae easier to notice, although variably placed; in four pairs located mostly ventrally, with
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||
<emphasis id="B92D4445C429FFA9FA5FFC88FA3FFC8D" box="[1408,1436,812,837]" italics="true" pageId="1" pageNumber="215">ab</emphasis>
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||
and
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||
<emphasis id="B92D4445C429FFA9FF15FCEBFF46FCA0" box="[202,229,847,872]" italics="true" pageId="1" pageNumber="215">cd</emphasis>
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paired very closely, or distantly, with
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||
<emphasis id="B92D4445C429FFA9FD76FCF5FD66FCA0" box="[681,709,849,872]" italics="true" pageId="1" pageNumber="215">aa</emphasis>
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||
2–3 times as
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<emphasis id="B92D4445C429FFA9FCB2FCEBFC24FCA0" box="[877,903,847,872]" italics="true" pageId="1" pageNumber="215">bc</emphasis>
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||
;
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||
<emphasis id="B92D4445C429FFA9FC46FCEBFC16FCA0" box="[921,949,847,872]" italics="true" pageId="1" pageNumber="215">dd</emphasis>
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||
nearly 0.6–0.7 of U. Occasionally
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<emphasis id="B92D4445C429FFA9FA8CFCEBFACEFCA0" box="[1363,1389,847,872]" italics="true" pageId="1" pageNumber="215">cd</emphasis>
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not noticeable, when
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<emphasis id="B92D4445C429FFA9FEBDFCD0FEDDFC45" box="[354,382,884,909]" italics="true" pageId="1" pageNumber="215">ab</emphasis>
|
||
easily noted. Papillae and genital glands variably shaped, with segments on/in which they occur varying, occasionally associated with genital setae. Male pores occur variably in the area of tubercula pubertatis, although the pores are difficult to detect externally. Spermathecae small, present or absent, often deeply embedded in body tissue; variably shaped, paired or multiple; in the majority of species they are close to septa
|
||
<date id="FFE7BE97C429FFA9FAA5FC44FE88FBD4" pageId="1" pageNumber="215" value="2013-11-12" valueMax="2013-11-12" valueMin="2013-11-12">11/ 12 and 12/13</date>
|
||
, although in some species may be located in more segments, sometimes up to six.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |